Corresponding author: Petar Beron ( beron@mail.bg ) © 2021 Petar Beron.
This is an open access book distributed under the terms of the Creative Commons Attribution License (CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.
Citation:
Beron P (2021) ACARORUM CATALOGUS VIII. Trombidiformes, Prostigmata, Superfamilia Cheyletoidea (Cheyletidae, Psorergatidae, Demodecidae, Harpyrhynchidae, Syringophilidae), Superfamilia Cloacaroidea (Cloacaridae, Epimyodicidae). Advanced Books. https://doi.org/10.3897/ab.e69591
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The year 1955 was remarkable in the history of Acarology. Several attempts of classification of various groups have been published almost simultaneously: of the Acaridiae (C.E. Yunker), of the Trombidiform mites (F. Cunliffe) and of Suborder Mesostigmata (J. H. Camin and F.E. Gorirossi). In the same year Dubinin and al. revised many groups of parasites of rodents, Zumpt & Till wrote their paper on Southafrican mites and many other acarological initiatives took place. Nevertheless, 70 years later the taxonomy of the Acari is still subject of many changes, due to deeper approach and to the new taxa, described from all over the world. From Cheyletidae have been detached several new families, some have been later downgraded to subfamily level (Ophioptinae) and some peculiar new taxa have been described (Cloacaridae, Epimyodicidae and the new superfamily Cloacaroidea). Here the new acquisations are followed with the conviction of the present author that the jugement of the new generation, based on cladistics and DNA analysis, is well founded.
Recently Walter, Lindquist, Smith, Cook & Krantz (2009) presented a new treatment of what was considered order Acariformes. Formerly divided into three suborders: Prostigmata, Acaridida and Oribatida, in the new system only two entities remain (Trombidiformes and Sarcoptiformes), reminiscent of some very old systems. These entities are ranked as orders, and the Trombidiformes are subdivided into two suborders. Sphaerolichida contains only the superfamilies Lordalycoidea and Sphaerolichoidea, Prostigmata (retaining the rank of suborder) – the other 36 superfamilies. The superfamilies considered in this volume (Cheyletoidea and the newly created superfamily Cloacaroidea Bochkov et OConnor, 2008) share the cohort Raphignathina with Raphignathoidea and Tetranychoidea.
The present volume contains data on 1243 species from 177 genera and seven families. The work was strongly facilitated by the revisions and chеcklists of different families, written by prominent specialists: Cheyletoidea (Dubinin, 1958, Bochkov, 2002), Cheyletidae (Volgin, 1969, Summers & Price, 1970, Fain, 1972, Gerson, Fain & Smiley, 1999, Bochkov & Fain, 2001), Psorergatidae (Giesen, 1990), Demodecidae (Izdebska & Rolbiecki, 2020), Harpirhynchidae (Fritsch, 1954; Bochkov & Literák, 2006; Fain, 1964; Bochkov, Mironov & Fain, 1999; Bochkov, OConnor & Klompen, 2015), Syringophilidae (Kethley, 1970; Skoracki, 2011; Skoracki, Zabludovskaya & Bochkov, 2012; Glowska, Chrzanowski & Kaszewska, 2015; Zmudzinski & Skoracki, 2016), and Cloacaridae (Fain, 1968).
The superfamily Cheyletoidea is composed of five families. Part of Cheyletidae and all members of Psorergatidae, Demodecidae, Harpirhynchidae and Syringophilidae are obligate parasites, mostly on vertebrates.
Here is the evolution of the concept of Cheyletoidea in the systems of various authors (after Bochkov, 2002, completed)
Cunliffe, 1955 – Cheyletoidea as a new superfamily of Cheyletidae, Myobiidae, Demodecidae, and Heterocheylidae.
Dubinin, 1957 – Cheyletidae, Syringophilidae, Myobiidae, Harpirhynchidae (incl. Ophioptinae)
Baker et al., 1958 – Cheyletidae, Syringophilidae, Myobiidae, Harpirhynchidae, Ophioptidae, Demodecidae, Psorergatidae, Heterocheylidae
Lawrence, 1959 – Cheyletidae, Syringophilidae, Myobiidae, Harpirhynchidae (incl. Psorergates and Ophioptes)
Volgin, 1969 – Cheyletidae, Syringophilidae, Harpirhynchidae (incl. Psorergates), Ophioptidae, Cloacaridae. Myobiidae is raised in a superfamily Myobioidea
Kethley, 1970 – Cheyletidae, Syringophilidae, Myobiidae, Harpirhynchidae, Demodecidae, Psorergatidae, Ophioptidae, Cloacaridae
Smiley, 1970 – Cheyletiellinae becomes a family (Cheyletiellidae)
Fain et al., 1997 – Cheyletiellinae again downgraded to subfamily rank
Bochkov, 1999 – Ophioptidae is downgraded to subfamily of Harpirhynchidae
Bochkov, 2002 – Complete review of the system and phylogeny of Cheyletoidea. Epimyodicinae is raised to family status. Seven families: Cheyletidae, Harpirhynchidae, Syringophilidae, Psorergatidae, Demodicidae, Cloacaridae, Epimyodicidae
Bochkov & OConnor, 2008 – Epimyodicidae and Cloacaridae form the new superfamily Cloacaroidea Walter et al. (in Krantz & Walter, 2009) – seven families of Cheyletoidea and two families in Cloacaroidea. This system is followed in the present Catalogue.
Superfamily Cheyletoidea Cunliffe, 1955
Fam. Cheyletidae Leach, 1815
Chelonotinae Volgin, 1969
Cheyletinae Leach, 1815
Cheyletiellinae Volgin, 1961
Criokerontinae Smiley, 1978
Metacheyletinae Fain, 1972
Ornithocheyletiinae Smiley, 1978
Fam. Psorergatidae Dubinin, 1955
Fam. Harpirhynchidae Dubinin, 1957
Harpirhynchinae Dubinin, 1957
Harpipalpinae Fain, 1972
Ophioptinae Southcott, 1956
Fam. Demodecidae Nicolet, 1855
Fam. Syringophilidae Lavoipierre, 1953
Picobiinae Johnston et Kethley, 1973
Syringophilinae Lavoipierre, 1953
Superfamily Cloacaroidea Bochkov et O’Connor, 2008
Fam. Cloacaridae Camin, Moss, Oliver et Singer, 1967
Cloacarinae Camin, Moss, Oliver et Singer, 1967
Pneumophaginae Fain et Smiley, 1989
Fam. Epimyodicidae Fain, Lukoschus et Rosmalen, 1982
Cheyletidae Leach, 1815: 399
Large family (acc. to Bochkov, 2009b, 380 sp. of 72 genera; acc. to Xia, Klompen & Childers, 2011, 370 sp. of 73 genera; acc. to the list of Zhang et al., 2011, 75 genera and 438 sp.). Contains both predators and permanent parasites of vertebrates. Some are nest or house-dust inhabitants, others live on plants or in insect colonies (phoretic), or are found (Hoffmannita) on Crustacea (Decapoda) and Arachnida (Scorpiones). There are Cheyletids (Cheyletiella parasitivorax, Ch. yasguri) living on domestic animals (rabbits, cats, dogs) and recorded as vectors of mange on Man. According to Bochkov (2004), 280 species of 54 genera (77.8%) are predators, 80 sp. of 19 gen. (23.2%) are parasites of birds and mammals.
The first Cheyletid was described by Schrank (1781, the well known “Acarus” eruditus, type of the genus Cheyletus Latreille, 1796). The family Cheyletidae was described by Leach (1815). Among the many publications until 1969 should be mentionned the revision (and many other papers) of Oudemans (1906), the paper of Baker (1949) on North American Cheyletidae and the analysis of Dubinin (1957). Other acarologists having contributed to the taxonomy of Cheyletidae before 1969 are Heller, Mégnin, Berlese, Michael, Canestrini, Trouessart, Ewing, Zachvatkin, Rohdendorf, Womersley, Cooreman, Ehara, Elbadry, Hara, Sasa, Muma, De Leon, Samšinak, Gerson, Flechtmann, Domrow, Radford, Pelaez, Lawrence, Smiley, Yunker and others.
By the end of the 60es almost simultaneously appeared two world revisions of Cheyletidae. Most remarkable is the monograph of Volgin (1969 in Russian, translated into English in 1987). In his book (and in some earlier publications since 1949) the prominent Russian specialist described 21 valid new genera and restructured the entire system of the family, with new tribes and species groups. Volgin divided these mites in two subfamilies, Cheyletinae and Cheyletiellinae (the parasites of vertebrates). This last subfamily was raised in family rank in the review of Smiley (1970), but was again downgraded in subfamily by Fain et al., 1997). His tribe Chelonotini was raised in subfamily. The other revision (Summers & Price, 1970) indicates that by this time the Cheyletidae is composed by “close to” 50 genera and 186 species. By now the number of the species is more than double, what is largely the result of the quoted revisions. In the last 50 years many papers appeared on the Cheyletidae of the Philippines (Corpuz-Raros), China-Taiwan (Tseng), Israel and Australia (Gerson), Iran (Saboori, etc.), Pakistan (many authors). Smiley (1978) described the subfamilies Criokerontinae and Ornithocheyletiinae. Other workers who contributed to the study of the taxonomy of Cheyletidae are Thewke, Enns, Klompen, Barilo, Klimov, Mironov, OConnor, Chaudhri, Delfinado, Doğan, Goff, Guilhon, Gupta, Haitlinger, Hassan, Hu, Ide, Kethley, Khaustov, Jeffrey, Kamran, Koç, Ayıldız, Kuznetzov, Lin, Lucza, Mathur, Meyer, Nagar, Olivier, Patxot, Putatunda, Kapil, Qayyum, Ramaraju, Rasool, Shiba, Soliman, Tjying, Uchikawa, Vaivanijkul, Wafa, Xia, Yousef, Zaher and others.
A. Fain started publishing papers on Cheyletidae in 1972. Alone or together with Bochkov, Gerson, Lukoschus, Smiley, Nadchatram and other co-authors, he described the subfamily Metacheyletinae, 7 new genera and many new species, mainly parasitic. In the comprehensive papers of Fain, Smiley & Gerson (1997) and Gerson, Fain & Smiley (1999) the number of accepted species in the family was stated as “more than 400” and the genera as “about 75”. Fain et al. (1999) compiled a list of all taxa in Cheyletidae. Bochkov & Fain (1998) left Cheyletidae with 73 valid genera and two genera incertae sedes. Followed the descriptions of Kenyacheylus Fain et Bochkov, 2001, Granulocheyletus Fain et Bochkov, 2002, Picocheyletus Bochkov et OConnor, 2003, Aztecocheyletus Bochkov et Klimov, 2004, Oconnoricheylus Bochkov et Otto, 2010 and Lanceacheyla Xia, Klompen et Childers, 2011.
In this volume the family is considered as composed of six subfamilies, the largest being the Cheyletinae (375 sp.).
Family Cheyletidae Leach, 1815 – 80 gen., ca. 474 sp. (part of them sp. dubia or inquirendae)[acc. to Zhang et al., 2011 – 438 sp.]
Subfamily Chelonotinae Volgin, 1969 – 1 gen., 1 sp.
Chelonotus Berlese, 1893 – 1 sp.
Subfamily Cheyletiellinae Volgin, 1961 – 4 gen., 33 sp.
Bothrocheyla Volgin, 1964 – 2 sp.
Cheyletiella Canestrini, 1886 – 6 sp.
Eucheyletiella Volgin, 1960 – 7 sp.
Neocheyletiella Baker, 1949 – 18 sp.
Subfamily Cheyletinae Leach, 1815 – 64 gen., 375 sp.
Tribus Cheyletini Leach, 1815 – 23 gen., 189 sp.
Anthribicheyla Thewke, 1980 – 1 sp.
Aztecocheyletus Bochkov et Klimov, 2004 – 1 sp.
Camincheyletus Smiley et Whitaker, 1981 – 1 sp.
Cheletacarus Volgin, 1961 – 5 sp.
Cheletonella Womersley, 1941 – 7 sp.
Cheletomimus Oudemans, 1904 – 68 sp. (many inquir.)
Cheletomimus Oudemans – 19 sp. (incl. 15 inquir.)
Hemicheyletia Volgin, 1969 – 45 sp.
Philippicheyla Corpuz-Raros, 1972 – 4 sp.
Cheletophanes Oudemans, 1904 – 1 sp. + 1 inc. sed.
Cheletophyes Oudemans, 1914 – 13 sp. + 7 inc. sed.
Cheyletus Latreille, 1796 – 48 sp. (35 + 13 inc. sed.)
Eucheyletia Baker, 1949 – 14 sp. + 1 inc. sed.
Granulocheyletus Fain et Bochkov, 2002 – 3 sp.
Kenyacheylus Fain et Bochkov, 2001 – 1 sp.
Ker Muma, 1964 – 7 sp.
Laeliocheyletia Summers et Price, 1970 – 1 sp.
Lanceacheyla Xia, Klompen et Childers, 2011 – 2 sp.
Lepidocheyla Volgin, 1963 – 2 sp.
Nidocheyletus Bochkov et O’Connor, 2010 – 1 sp.
Nodele Muma, 1964 – 5 sp.
Paracheyletia Volgin, 1955 – 1 sp.
Paracheyletiella Kuznetzov, 1977 – 1 sp.
Pavlovskycheyla Volgin, 1965 – 3 sp.
Tutacheyla Corpuz – Raros, 1972 – 1 sp.
Zachvatkiniola Volgin, 1969 – 1 sp.
Tribus Acaropsellini Volgin, 1969 – 7 gen., 47 sp.
Acaropsella Volgin, 1969 – 7 sp.
Acaropsellina Summers, 1976 – 15 sp.
Atarsacheylus Thewke, 1980 – 1 sp.
Chelacheles Baker, 1958 – 19 sp.
Neoacaropsis Volgin, 1962 – 1 sp.
Neochelacheles Smiley et Williams, 1972 – 3 sp.
Paracaropsis Volgin, 1969 – 1 sp.
Tribus Bakini Volgin, 1969 – 34 gen., 161 sp.
Aegyptocheyla Yousef, 1979 – 1 sp.
Alliea Yunker, 1960 – 2 sp.
Bak Yunker, 1961 – 14 sp.
Caudacheles Gerson, 1968 – 3 sp.
Chelacaropsis Baker, 1949 – 7 sp.
Cheletogenes Oudemans, 1905 – 12 sp.
Cheletoides Oudemans, 1904 – 2 sp.
Cheletomorpha Oudemans, 1904 – 7 sp.
Cheletopsis Oudemans, 1904 – 14 sp.
Cheletosoma Oudemans, 1905 – 1 sp.
Cheyletia Heller, 1884 – 5 sp.
Chiapacheylus De Leon, 1962 – 3 sp.
Columbicheyla Thewke et Enns, 1972 – 3 sp.
Cunliffella Volgin, 1969 – 8 sp.
Dubininiola Volgin, 1969 – 3 sp.
Eucheletopsis Volgin, 1969 – 1 sp.
Eutogenes Baker, 1949 – 14 sp.
Grallacheles De Leon, 1962 – 3 sp.
Hoffmannita Pelaez, 1962 – 4 sp.
Hylopecheyla Fain, 1972 – 2 sp.
Hypopicheyla Volgin, 1969 – 2 sp.
Metacheletoides Fain, 1972 – 4 sp.
Mexecheles De Leon, 1962 – 10 sp.
Microcheyla Volgin, 1966 – 5 sp.
Muricheyla Fain, 1972 – 1 sp.
Neoeucheyla Radford, 1950 – 6 sp.
Oconnoricheylus Bochkov et Otto, 2010 – 2 sp.
Oudemansicheyla Volgin, 1969 – 2 sp.
Paramicrocheyla Olivier et Theron, 1989 – 2 sp.
Picocheyletus Bochkov et OConnor, 2003 – 1 sp.
Promuricheyla Fain, 1979 – 1 sp.
Prosocheyla Volgin, 1969 – 9 sp.
Samsinakia Volgin, 1965 – 8 sp.
Thewkacheyla Ide et Kethley, 1977 – 1 sp.
Subfamily Criokerontinae Smiley, 1978 – 4 gen., 12 sp.
Tribus Niheliini Smiley, 1978 – 4 gen., 12 sp.
Criokeron Volgin, 1966 – 2 sp.
Galagocheles Fain, 1979 – 2 sp.
Nihelia Domrow et Baker, 1960 – 3 sp.
Sciurocheyla Volgin, 1969 – 5 sp.
Subfamily Metacheyletinae Fain, 1980 – 1 genus, 5 sp.
Metacheyletia Fain, 1972 – 5 sp.
Subfamily Ornithocheyletiinae Smiley, 1978 – 6 gen., 42 sp.
Tribus Ornithocheyletiini Volgin, 1969 – 5 gen., 40 sp.
Apodicheles Fain, 1979 – 4 sp.
Bakericheyla Volgin, 1966 – 5 sp.
Apodicheyla Fain, 1979 – 1 sp.
Bakericheyla Volgin, 1966 – 4 sp.
Ornithocheyletia Volgin, 1964 – 31 sp.
Tribus Teinocheylini Fain, 1974 – 1 genus, 2 sp.
Teinocheylus Fain, 1974 – 2 sp.
Cheyletidae Leach, 1815: 399 (Kramer, 1877: 245)
(= Cheyletides Leach, 1815)
Type genus: Cheyletus Latreille, 1796
Chelonotini Volgin, 1969: 329
Chelonotinae: Fain, 1979: 408
Type genus: Chelonotus Berlese, 1893
Chelonotus Berlese, 1893: 73
Type species: Chelonotus selenorhynchus Berlese, 1893
Chelonotus selenorhynchus Trouessart in Berlese, 1893: 9, 15, 77; Oudemans, 1906: 159; Baker, 1949: 312; Domrow, 1960: 457 (selenirhynus); 1964: 16; Nadchatram & Domrow, 1966: 135; Volgin, 1969: 330; Summers & Price, 1970: 72
Chelonotus oudemansi Baker, 1949: 310 – type: Sulawesi, on Callosciurus tenuis
Chelonotus ewingi Baker, 1949: 311 – type: Sulawesi, Temboan, on Callosciurus prevosti
Type locality: Pulosembeh, Sulawesi (Indonesia)
Distribution: Indonesia (Sulawesi), Malaysia (Malaya, Borneo, Pulau Tioman)
Hosts: Tupaia glis (Scandentia: Tupaiidae), Sundasciurus hippurus, S. [Baginia] lowii, S. [Baginia] tenuis evidens, Lariscus insignis (Rodentia: Sciuridae)
Cheyletiellinae Volgin, 1961: 255
Type genus: Cheyletiella Canestrini, 1886
Bothrocheyla Volgin, 1964: 94
Type species: Neoeucheyla (Bothrocheyla) pavlovskyi Volgin, 1964
Neoeucheyla (Bothrocheyla) pavlovskyi Volgin, 1964: 94
Neoeucheyla pavlovskyi Volgin: Vysotskaja, 1977: 188
Bothrocheyla pavlovskyi: Volgin, 1969: 262
Neoeucheyla beeri Thewke & Enns, 1972: 455
Type locality of Neoeucheyla (Bothrocheyla) pavlovskyi: Murmanskaya and Zakarpatskaya oblasti (Russia, Ukraine)
Distribution: Czech Rep., Russia, Ukraine
Hosts: nests of rodents; on Heteroptera (Aradus lugubris, A. corticalis)
Neoeucheyla typhosa Summers & Price, 1970: 67
Bothrocheyla typhosa (Summers et Price)
Type locality: Lake Pillsbury, Napa Co., California
Distribution: USA (California)
Host: from soil under pine trees
Cheyletiella Canestrini, 1886: 170
Type species: Cheyletus parasitivorax Mégnin, 1878
Bicheyletiella Fain, 1972: 48 (synonymized by Fain & Bochkov, 2001b: 293)
Type species: Bicheyletiella romerolagi Fain, 1972
Cheyletiella blakei Smiley, 1970: 1072; Keh, 1975: 1; van Bronswijk & de Kreek, 1976: 315; Mehl, 1978; Domrow, 1991: 1305; Fain & Bochkov, 2001c: 90
Type locality: Ithaca, N.Y.
Distribution: Australia (N.S.W., Victoria, Tasmania), France, Norway, Russia, USA (N.Y.)
Hosts: Felis catus (Carnivora: Felidae), Homo sapiens (Primates: Hominidae)
Cheyletiella dengi Hu, 1992: 183
Type locality: Xinxiang (China)
Distribution: China
Cheyletus parasitivorax Mégnin, 1878: 10; Canestrini, 1879: 40; Mégnin, 1880: 241;
Cheyletiella parasitivorax (Mégnin): Canestrini, 1886: 175; Berlese, 1886: 28; Cooper, 1946: 480; Baker, 1949: 270; Starkoff & Starkoff, 1950: 96; Barr, 1955: 323; Volgin, 1960: 242; 1969: 360; Alcaino & Tagle, 1966; Smiley, 1970: 1075; Amaral, 1971: 106; Rak, 1972: 62; 1973: 189; Shepherd & Edmonds, 1977; Mehl, 1978; Wassel, Whitaker & Spicka, 1980: 419; Hoffmann et al., 1994: 210; Fain & Bochkov, 2001: 89; Whitaker et al., 2007: 6; Xia, 2010: 161
[Cheyletiella americana Ewing, 1909b: 77. – Type: Urbana. Illinois, under bark (acc. to Baker, 1949, belongs to Pseudocheylus)]
Cheletiella parasitivorax: Oudemans, 1906: 212; Hirst, 1917: 132; Womersley, 1941: 59
Ewingella americana Vail & Augustson, 1943: 419 – Type from Oryctolagus cuniculus, California, USA
Cheyletiella furmani Smiley, 1970: 1074 – Jacksonville, Fla., USA, from Sylvilagus palustris paludicola (Lagomorpha: Leporidae); synonymized by Fain & Bochkov, 2001c: 88
Cheyletiella katangae Fain, 1972: 48 – Katanga, DR Congo, from Lepus whytei (Lagomorpha: Leporidae)
Cheyletiella dengi Hu & Hou, 1992: 183 – Type: China, Xinjiang, from Oryctolagus cuniculus (Lagomorpha: Leporidae); synonymized by Fain & Bochkov, 2001c: 88
Type locality of Cheyletus parasitivorax Mégnin: Probably France
Distribution: Australia, Brazil, Chile, China, Cuba, Danemark, England, France, Greece, Holland, Iran, Italy, Mexico, Nigeria, Norway, Russia, South Africa, New Zealand, USA
Hosts: Homo sapiens (Primates: Hominidae), Lepus whytei, Oryctolagus cuniculus, Pronolagus randensis, Romerolagus diazi, Sylvilagus bachmanni, S. floridanus (Lagomorpha: Leporidae), Felis catus (Carnivora: Felidae)
Bicheyletiella romerolagi Fain, 1972: 49; 1979: 422
Cheyletiella mexicana Uchikawa & Suzuki, 1979: 21; Hofmann et al., 1994: 211
Cheyletiella romerolagi (Fain): Fain & Bochkov, 2001b: 293; 2001c: 89
Type locality: Mexico
Distribution: Mexico
Host: Romerolagus diazi (Lagomorpha: Leporidae)
Cheyletiella strandtmanni Smiley, 1970: 1072; Fain & Bochkov, 2001c: 89
Type locality: Ma-am Lio
Distribution: Taiwan
Host: Lepus sp. (Lagomorpha: Leporidae)
Cheyletiella yasguri Smiley, 1965: 76; Vercammen-Grandjean & Rak, 1968: 407; Volgin, 1969: 365; Davies & Kyle, 1969: 136; Bakkers & Fain, 1972: 25; Rak, 1973: 181; Alcaino & Calderon, 1976: 40; Mehl, 1978; Fain, Scheepers & de Groot, 1982: 624; Chin, 1996: 267; Fain & Bochkov, 2001c: 90
Type locality: New York
Distribution: Chile, England, France, Germany, Holland, Iran, Ireland, Korea, Norway, USA
Host: Canis lupus familiaris (Carnivora: Canidae); phoretic hyperparasite of a louse-fly (Hippobosca longipennis), collected on dog in Iran
Eucheyletiella Volgin, 1960: 238
Type species: Cheyletiella ochotonae Volgin, 1960
Eucheyletiella faini Bochkov & Mironov, 1999: 148
Type locality: Gorno-Altaysk Province, Chuyskaya steppe, Tashanta Village
Distribution: Kazakhstan, Russia (Tuva, Altai)
Host: Ochotona rufescens (Lagomorpha: Ochotonidae)
Eucheyletiella faini Bochkov & Malikov, 1997: 43; Bochkov & Mironov, 1999: 144
Type locality: Kerman Province, near to Shad-Dad Tunnel
Distribution: Iran
Host: Ochotona rufescens (Lagomorpha: Ochotonidae)
Cheyletiella johnstoni Smiley, 1965: 77
Eucheyletiella johnstoni (Smiley): Volgin, 1969: 374
Type locality: New Mexico (Santa Fe)
Distribution: USA (New Mexico)
Host: Ochotona princeps (Lagomorpha: Ochotonidae)
Cheyletiella ochotonae Volgin, 1960: 238
Eucheyletiella ochotonae Volgin, 1969: 368; Dubinina & Bochkov, 1989: 107; Bochkov & Mironov, 1999: 142
Type locality: Terskey Alatau (Chong Kizilsu), Kirghyzstan
Distribution: Kirghyzstan, Turkmenia
Hosts: Ochotona macrotis, O. roylei, O. rufescens (Lagomorpha: Ochotonidae), Sylvaemus sylvaticus (Rodentia: Muridae)
Eucheyletiella pallasius Bochkov & Mironov, 1999: 146
Type locality: Tuva, Mugir-Aken
Distribution: Russia (Tuva)
Host: Ochotona pallasi (Lagomorpha: Ochotonidae)
Eucheyletiella pusillinus Bochkov & Mironov, 1999: 144
Type locality: Orenburg Prov.
Distribution: Russia
Host: Ochotona pusilla (Lagomorpha: Ochotonidae)
Cheyletiella takahashii Sasa & Kono, 1951: 205
Eucheyletiella takahashii (Sasa et Kono): Volgin, 1969: 372
Eucheyletiella takahasii (Sasa et Kono): Bochkov & Mironov, 1999: 150
Type locality: Hokkaido
Distribution: Japan (Hokkaido), Russia (Asiatic)
Host: Ochotona hyperborea, O. h. yesoensis (Lagomorpha: Ochotonidae)
Neocheyletiella Baker, 1949: 271
Ornithocheyla Lawrence, 1959: 433
Type species: Neocheyletiella rohwery Baker, 1949
Neocheyletiella aegithali Bochkov & OConnor, 2003: 277; Halliday, Kamran & Bashir, 2018: 87
Type locality: NW Frontier Prov., Malakand Div., Swat Dist., Yakh Tangai, 2076 m
Distribution: Pakistan
Host: Aegithalos concinnus (Passeriformes: Aegithalidae)
Neocheyletiella alfortensis Guilhon & Euzeby, 1984: 497
Type locality: unknown
Distribution: unknown
Host: Leiothrix lutea (Passeriformes: Timaliidae)
Neocheyletiella amandavae Fain, 1972: 47; 1980: 33
Type locality: Antwerp Zoo
Distribution: Java
Host: Amandava amandava (Passeriformes: Estrildidae)
Neocheyletiella ardeola Bochkov & Ochoa, 2005: 142
Type locality: Lake Fuhwa
Distribution: Mozambique
Host: Ardeolla ralloides (Pelecaniformes: Ardeidae)
Neocheyletiella artami Domrow, 1965: 210; 1970: 273; Summers & Price, 1970: 77; Smiley, 1978: 236
Type locality: Exeter (Tasmania)
Distribution: Australia (Tasmania, New South Wales)
Host: Artamus cyanopterus (Passeriformes: Artamidae)
Neocheyletiella athene Fain & Bochkov, 2002a: 159
Type locality: India
Distribution: India
Host: Athene brama (Strigiformes: Strigidae)
Neocheyletiella avicola Fain, 1972: 47; 1980: 33
Neocheyletiella vestergaardi Smiley, 1978: 236 – Copenhagen, Denmark, caged Erythrura prasina (Passeriformes: Estrildidae)
Type locality: Antwerp Zoo
Distribution: ? (South America, Australia)
Hosts: Ara sp., Agapornis fisheri (Psittaciformes: Psittacidae), Erythrura prasina (Passeriformes: Estrildidae)(N. vestergaardi)
Neocheyletiella lonchurae Fain & Bochkov, 2002a: 159
Type locality: Akanyaru River
Distribution: Rwanda
Host: Lonchura sp. (Passeriformes: Estrildidae)
Cheyletus macronychus Mégnin, 1878: 427; 1880: 243; Poppe, 1888: 237
Cheyletiella macronycha Berlese & Trouessart, 1889: 136
Cheletiella macronycha: Oudemans, 1906: 212
Neocheyletiella macronycha, Baker, 1949: 272; Fain, 1980: 39; Smiley, 1978: 236
Type locality: Bengal
Distribution: India
Host: birds (Passeriformes)
Neocheyletiella media Fain, 1972: 48; 1980: 28
Type locality: host died in Antwerp
Distribution: S. China
Host: Leiothrix lutea (Passeriformes: Timaliidae)
Ornithocheyla megaphallos Lawrence, 1959: 434; Lindholm, Venter & Ueckermann, 1998
Neocheyletiella megaphallos (Lawrence): Volgin, 1969: 419; Summers & Price, 1970: 77
Type locality: Debeete
Distribution: Botswana, South Africa
Hosts: Estrilda erythbronotos (Passeriformes: Estrildidae), Chrysococcyx caprius (Cuculiformes: Cuculidae)
Cheyletiella microrhyncha Berlese & Trouessart, 1889: 136; Berlese, 1893: 76
Cheletiella microrhyncha: Oudemans, 1906: 212
Neocheyletiella microrhyncha (Berlese & Trouessart): Baker, 1949: 272; Summers & Price, 1970: 77; Smiley, 1978: 234; Bochkov & Galloway, 2001: 2017
Neocheyletiella oudemansi Volgin, 1969: 417 – Belarus, from Delichon urbica and Riparia riparia
Neocheyletiella callawaye Smiley, 1970: 1063; 1978: 236
Type locality: France
Distribution: Belarus, Canada, France, Russia; Rwanda; USA
Hosts: Hirundo rustica, Delichon urbica, Petrochelidon pyrchonota, Cecropis abyssinicus unitatis, Progne subis, Psalidoprocne albiceps, Riparia riparia (Passeriformes: Hirundinidae)
Neocheyletiella nectarinia Bochkov & OConnor, 2003: 277
Type locality: Romblon Prov., Sibuyan Isl., Magdivang, Tampayan
Distribution: Philippines
Host: Nectarinia jugularis (Passeriformes: Nectariniidae)
Neocheyletiella pittae Fain, 1972: 47; 1980: 33
Type locality: Antwerp Zoo
Distribution: ?
Host: Pitta megarhyncha (Passeriformes: Pittidae)
Neocheyletiella queleae Fain & Bochkov, 2002 : 155
Type locality: Rubona, Rwanda
Distribution: Rwanda
Hosts: Quelea quelea (Passeriformes: Ploceidae), Streptopelia semitorquata (Columbiformes: Columbidae)
Neocheyletiella rohweri Baker, 1949: 274; Volgin, 1969: 414; Summers & Price, 1970: 77; Smiley, 1978: 236
Type locality: Guffey, Colorado
Distribution: USA (Colorado)
Host: Sitta pygmaea melanotus (Passeriformes: Sittidae)
Neocheyletiella siva Fain, 1972: 48; 1980: 37
Type locality: Antwerp Zoo
Distribution: India
Host: Siva [Minia] cyanoptera (Passeriformes: Timaliidae)
Neocheyletiella smallwoodae Baker, 1949: 273; Volgin, 1969: 415; Summers & Price, 1970: 77; Smiley, 1978: 236
Type locality: Guffei, Colorado
Distribution: USA (Colorado, Delawar)
Host: Leucosticte australis (Passeriformes: Fringillidae, Carduellinae)
Distribution: Japan, Korea (McClure H. Eliott, N. Ratanaworadhan et al. 1973)
Host: Emberiza cioides (Passeriformes: Emberizidae)
Cheyletinae Leach, 1815: 399
Type genus: Cheyletus Latreille, 1796
Cheyletini Leach, 1815: 399
Type genus: Cheyletus Latreille, 1796
Anthribicheyla Thewke, 1980: 335
Type species: Anthribicheyla bocki Thewke, 1980
Anthribicheyla bocki Thewke, 1980: 337
Type locality: Vera Cruz
Distribution: Mexico
Hosts: Ischnocerus, Ptychoderes (Coleoptera: Anthribidae)
Aztecocheyletus Bochkov & Klimov, 2004: 24
Type species: Nodele conquistador Bochkov et Klimov, 2004
Nodele conquistador Bochkov & Klimov, 2004: 24
Aztecocheyletus conquistador Bochkov & Klimov, 2004: 23
Type locality: Jalisco, Chamela, Est. biologica
Distribution: Mexico
Host: Azthecanthidium tenochtitlanicum (Hymenoptera: Megachilidae)
Camincheyletus Smiley & Whitaker, 1981: 111
Type species: Camincheyletus glaucomys Smiley et Whitaker, 1981
Camincheyletus glaucomys Smiley & Whitaker, 1981: 114
Type locality: Oregon
Distribution: USA (Oregon)
Host: Glaucomys sabrinus (nest)(Rodentia: Pteromyidae)
Cheletacarus Volgin, 1961: 248
Type species: Cheletacarus raptor Volgin, 1961
Cheletacarus gryphus Summers & Price 1970: 36
Type locality: Davis, California
Distribution: USA (California)
Cheletacarus novemdentis Meyer, 1988: 9
Type locality: Mountain Zebra Nat. Park
Distribution: South Africa
Cheletacarus ploceus Gupta & Paul, 1985: 6
Type locality: India
Distribution: India
Cheletacarus raptor Volgin, 1961: 248; 1969: 169; Lucza, Ripka & Saly, 1996; Lin et al., 1997; Ripka, Fain & Bolland, 1999: 366; Xia, 2010: 161; Bochkov & Abramov, 2016: 336; Khaustov, 2021: 234
Type locality: Kishinev (Moldova)
Distribution: China, Hungary, Moldavia, Russia
Hosts: Diaspidiothis otsreaeformis, D. spurcatus, Parthenolecanium corni (Homoptera: Coccodea)
Cheletophanes rugosa Womersley, 1941: 62
Cheletonella rugosa: Baker, 1949: 292
Chelacheles rugosus (Womersley): Volgin, 1961: 253; 1969: 172
Cheletacarus rugosus (Womersley): Summers & Price, 1970: 36
Type locality: Brisbane, Queensland
Distribution: Australia (Queensland)
Host: Calymmaderus sp. (Coleoptera: Anobiidae)
Cheletonella Womersley, 1941: 60
Type species: Cheletonella vespertilionis Womersley, 1941
Cheletonella daddariensis Aheer & Chaudhri, 1987: 104; Halliday, Kamran & Bashir, 2018: 86
Type locality: Pakistan
Distribution: Pakistan
Cheletonella emersus Chaudhri, Akbar & Rasool, 1979: 150; Aheer & Chaudhri, 1987: 103; Halliday, Kamran & Bashir, 2018: 86
Type locality: Pakistan
Distribution: Pakistan
Cheletonella hoffmannae Smiley, 1996: 241
Type locality: Quincy, Owen Co., Indiana
Distribution: USA (Indiana)
Host: guano of Eptesicus fuscus (Chiroptera: Vespertilionidae)
Cheletonella iraniensis Salarzehi, Hajizadeh & Ueckermann, 2019: 189
Type locality: Khomam City
Distribution: Iran
Cheletonella pilosa Tseng, 1977: 240; Xia, 2010: 161
Type locality: Taiwan
Distribution: Taiwan
Cheletonella vespertilionis Womersley, 1941: 61; Baker, 1949: 292; Domrow, 1959: 234; 1974: 18; 1991: 1305; Volgin, 1969: 153; 1987: 191; Summers & Price, 1970: 37; Whitaker, 1973: 1149; Domrow, 1974: 18; Fain & Nadchatram, 1980: 197; Fain & Bochkov, 2001; Ezequiel et al., 2001; Ezequiel et al., 2003; Whitaker et al., 2007: 6; Dogan et al., 2011: 782
Cheletonella caucasica Volgin, 1955: 168; 1956: 210; 1969: 154; Rekk, 1976: 68 –
Algeria, Armenia, Brazil, Georgia, Tajikistan, Ukraine (Cricetulus migratorius, bird’s nest)
Type locality: Glen Osmond (S. Australia)
Distribution: Australia (N.S.W., South Australia), Brazil, Malaysia, USA (California, Indiana, Pennsylvania, Texas); Armenia, Georgia, Tajikistan, Iran, Belgium, Ukraine
Hosts: bat (Chiroptera indet.), Eptesicus fuscus, guano of Myotis lucifugus, M. myotis (Chiroptera: Vespertilionidae), guano of Tadarida punctata (Molossidae), bat guano in Malaysia (Negeri Sembilan), nest of Hylochelidon ariel (Passeriformes: Hirundinidae)
Cheletonella juglandis Xia, Zhu & Ye, 1999: 150
Type locality: Jiangxi, Nanchang
Distribution: China
Cheletomimus Oudemans, 1904b: 163
Type species: Cheletes berlesei Oudemans, 1904
Cheletomimus Oudemans, 1904b: 163
Type species: Cheletes berlesei Oudemans, 1904
Cheyletus ornatus Berlese, 1886: 28 (misidentification)
Cheletes berlesei Oudemans, 1904b: 154
Cheletomimus ornatus (Berlese): Oudemans, 1906: 136;
Cheletomimus berlesei (Oudemans): Baker, 1949: 293; Volgin, 1955: 172; Yunker, 1961: 1032; De Leon, 1962: 134; Summers & Price, 1970: 9; Rekk, 1976: 68; Patxot & Goff, 1985: 160
Cheyletomimus berlesei (Oudemans): Evans, 1954: 806; Bochkov, Hakimitabar & Saboori, 2005: 107; Torre Santana, 2015
Cheletomimus (Ch.) berlesei (Oudemans): Fain, Bochkov & Corpuz-Raros, 2002: 30
Type locality: Italy
Distribution: Angola, Cuba, Georgia, Greece (ornatus), Great Britain, Iran, Italy, Israel, Mexico, Palestina, Russia, USA (Florida, Hawaiian Is.)
Host: Aspidiotus lataniae (Homoptera: Coccodea), bird’s nest
Cheletomimus daltoniensis Corpuz-Raros, 1998: 265
Cheletomimus (Cheletomimus) daltoniensis Corpuz-Raros: Fain, Bochkov & Corpuz-Raros, 2002: 33
Type locality: Luzon, Dalton Pass, Santa Fe, Nueva Vizcaya, 1070 m
Distribution: Philippine Is.
Cheletomimus duosetosus Muma, 1964: 242; Summers & Price, 1970: 9; Patxot & Goff, 1985: 161; Papaioannou-Souliotis, 1994; Feres & Flechtmann, 1995: 535
Cheletomimus (Cheletomimus) duosetosus Muma: Fain, Bochkov & Corpuz-Raros, 2002: 33
Type locality: Florida (USA)
Distribution: Brazil, Greece, USA (Florida, Hawaiian Is.)
Cheletomimus (Cheletomimus) ochoai Fain, Bochkov & Corpuz-Raros, 2002: 33
Type locality: California, San Francisco
Distribution: USA (California)
Host: on Howea posteriana (calix)
Hemicheyletia Volgin, 1969: 201
Type species: Paracheyletia bakeri Ehara, 1962
Andrecheyla Volgin, 1969: 219
Type species: Cheyletia scutellata De Leon, 1962
Dendrocheyla Volgin, 1969: 205
Type genus: Dendrocheyla bregetovae Volgin, 1969
Paracheyletia anarbora De Leon, 1967: 341
Hemicheyletia anarbora (De Leon): Summers & Price, 1970: 18; Corpuz-Raros & Sotto, 1977: 154; Ehara & Abdul Ghani, 1988: 239; Torre Santana, 2015: 49
Hemicheyletia pusillifolium Lin-Jianzhen, Pen-Wenfu et Chen-Yume, 1994: 141; Lin et al., 2000: 86 – China (Fujian)
Cheletomimus (Hemicheyletia) anarbora: Corpuz-Raros, 1977: 154; Fain, Bochkov & Corpuz-Raros, 2002: 49
Cheletomimus anarbora: Xia, 2010: 161
Type locality: Trinidad
Distribution: China, Cuba, Malaysia, Philippine Is., Taiwan, Trinidad
Hemicheyletia asiaticus Volgin, 1978: 213
Cheletomimus (Hemicheyletia) asiatica: Fain, Bochkov & Corpuz-Raros, 2002: 39
Type locality: Sayat Distr., Turkmenistan
Distribution: Turkmenistan (in a burrow of tarantula)
Hemicheyletia athenae Corpuz-Raros, 1988b: 418
Cheletomimus (Hemicheyletia) athenae: Fain, Bochkov & Corpuz-Raros, 2002: 54
Type locality: Mt. Banahaw at Kinabuhayan, Dolores,
Quezon, Luzon Is., the Philippines
Distribution: Philippine Is.
Paracheyletia bakeri Ehara, 1962: 109; Muma, 1964: 245; Gerson, 1967: 360; Kanavel & Selhime, 1967: 107
Hemicheyletia bakeri (Ehara): Volgin, 1969: 202; Summers & Price, 1970: 12; Tseng, 1977; Gerson, 1994: 445; Xia et al., 2000
Hemicheyletia tumidus Qayyum & Chaudhri, 1979: 169. Syn. by Fain, Bochkov & Corpuz-Raros, 2002: 36
Cheletomimus (Hemicheyletia) bakeri (Ehara): Fain, Bochkov & Corpuz-Raros, 2002: 36; Bochkov, Hakimitabar & Saboori, 2005: 107
Cheletomimus bakeri (Ehara): Xia, 2010: 161; Fain, Bochkov & Corpuz-Raros, 2002: 36; Bochkov, Hakimitabar & Saboori, 2005: 107; Halliday, Kamran & Bashir, 2018: 83
Type locality: Sapporo (Hokaido)
Distribution: Algeria, Australia, Canada, China, DR Congo, Egypt (Hemicheletus b.), Galapagos Isl., Iran, Israel, Japan, Pakistan, Panama, Philippines, Taiwan, USA
Hemicheyletia blosis Chaudhri & Akbar, 1985: 172.
Cheletomimus blosis (Chaudhri et Akbar): Halliday, Kamran & Bashir, 2018: 83
Type locality: Pakistan
Distribution: Pakistan
Dendrocheyla bregetovae Volgin, 1969: 207
Hemicheyletia bregetovae (Volgin): Diaz Patxot & Goff, 1985: 161
Cheletomimus (Hemicheyletia) bregetovae: Fain, Bochkov & Corpuz-Raros, 2002: 54; Koç & Ayyildiz, 1996:
Hemicheyletia scitula Corpuz-Raros, 1972: 256. Type from Mt. Makling, the Philippines. Syn. by Fain, Bochkov & Corpuz-Raros, 2002: 54
Type locality of Dendrocheyla bregetovae: Hosta, Sochi, (Russia)
Distribution: Armenia, Bulgaria, Philippines, Russia, Turkey, Crimea, USA (Hawaian Is. – Oahu)
Host: nests of Apodemus spp. (Rodentia: Muridae)
Hemicheyletia chui Tseng, 1977: 218
Cheletomimus (Hemicheyletia) chui (Tseng): Fain, Bochkov & Corpuz-Raros, 2002: 39
Cheletomimus chui (Tseng): Xia, 2010: 161
Type locality: Taiwan
Distribution: China, Malaysia, Taiwan
Host: from nest of Muscicapa grandis (Passeriformes: Muscicapidae)
Paracheyletia congensis Cunliffe, 1962: 197
Hemicheyletia congensis (Cunliffe): Volgin, 1969: 203; Summers & Price, 1970: 1
Hemicheyletia congoensis (Cunliffe): Corpuz-Raros, 1998:
Hemicheyletia lacinia Rasool & Chaudhri, 1979: 2 – Pakistan
Cheletomimus (Hemicheyletia) congensis (Cunliffe): Fain, Bochkov & Corpuz-Raros, 2002: 38; Bochkov, Hakimitabar & Saboori, 2005: 107
Cheletomimus congoensis (Cunliffe): Fain, Bochkov & Corpuz-Raros, 2002: 38; Bochkov, Hakimitabar & Saboori, 2005: 107; Halliday, Kamran & Bashir, 2018: 84
Type locality of Paracheyletia congensis Cunliffe, 1962: Kysenyi (DR Congo)
Distribution: DR Congo, Iran, Pakistan, Philippines
Cheyletia cordovensis De Leon, 1962: 219
Hemicheyletia cordovensis (De Leon): Volgin, 1969: 204; Summers & Price, 1970: 14
Cheletomimus (Hemicheyletia) cordovensis (De Leon): Fain, Bochkov & Corpuz-Raros, 2002: 39
Type locality: Cordoba
Distribution: Mexico (Veracruz)
Host: Phoebe psychotrioides (Passeriformes: Tyrannidae)
Hemicheyletia darwinia Summers & Price, 1970: 16
Cheletomimus (Hemicheyletia) darwinia (Summers et Price): Fain, Bochkov & Corpuz-Raros, 2002: 49
Type locality: Darwin Research Station, Galapagos
Distribution: Ecuador (Galapagos Is.)
Host: From grass near bay shore
Hemicheyletia echis Chaudhri & Akbar, 1985: 187
Cheletomimus echis (Chaudhri & Akbar): Halliday, Kamran & Bashir, 2018: 84
Type locality: Pakistan
Distribution: Pakistan
Hemicheyletia disparis Chaudhri & Akbar, 1985: 180.
Cheletomimus disparis (Chaudhri & Akbar): Halliday, Kamran & Bashir, 2018: 84
Type locality: Pakistan
Distribution: Pakistan
Hemicheyletia glomus Chaudhri & Akbar, 1985: 181;
Cheletomimus glomus (Chaudhri et Akbar): Halliday, Kamran & Bashir, 2018: 84
Type locality: Pakistan
Distribution: Pakistan
Cheletomimus (Hemicheyletia) gracilis Fain, Bochkov & Corpuz-Raros, 2002: 57
Type locality: Hawaii (on Allium cepa from Japan)
Distribution: Peru, the Philippines, USA (Hawaii)
Host: plants
Hemicheyletia granula Summers & Price, 1970: 13
Cheletomimus (Hemicheyletia) granula (Summers et Price): Fain, Bochkov & Corpuz-Raros, 2002: 36
Type locality: Quetzaltepeque
Distribution: El Salvador
Host: from moss and ant nest
Cheletomimus (Hemicheyletia) greenwoodi Fain, Bochkov & Corpuz-Raros, 2002: 54
Type locality: Barstey, Wales, Great Britain
Distribution: Great Britain
Host: from nest of Anthus spinoletta (Passerifpormes: Motacillidae)
Hemicheyletia hastatus Chaudhri & Akbar, 1985: 183; Afzal, Ali & Akbar, 2003: 90; Kamran, Khan & Nazir, 2005: 54
Cheletomimus hastatus (Chaudhri & Akbar): Halliday, Kamran & Bashir, 2018: 84
Type locality: Pakistan
Distribution: Pakistan
Hemicheyletia indica Gupta, 1991: 234; Fain, Bochkov & Corpuz-Raros, 2002: 28
Type locality: India
Distribution: India
Hemicheyletia kysenyiensis Thewke & Enns, 1979: 218
Cheletomimus (Hemicheyletia) kysenyiensis (Thewke et Enns): Fain, Bochkov & Corpuz-Raros, 2002: 38
Type locality: Kysenyi (former Zaire)
Distribution: D. R. Congo
Hemicheyletia lanceolata Corpuz-Raros, 1998: 274; Fain, Bochkov & Corpuz-Raros, 2002: 28
Type locality: Luzon Is., Pintor, Gamu, Isabela
Distribution: Philippine Is.
Hemicheyletia leytensis Corpuz-Raros, 1988: 419; 1998: 275
Cheletomimus (Hemicheyletia) leytensis (Corpuz-Raros): Fain, Bochkov & Corpuz-Raros, 2002: 42
Type locality: Babatngon, Leyte
Distribution: Philippines (Leyte, Luzon)
Hemicheyletia lindquisti Thewke & Enns 1979: 221
Cheletomimus (Hemicheyletia) lindquisti (Thewke et Enns): Fain, Bochkov & Corpuz-Raros, 2002: 57; Bochkov & Abramov, 2016: 336
Type locality: 2,5 miles SE Kananaskis F. E. Station, Alberta
Distribution: Canada (Alberta), Russia
Hemicheyletia makilingensis Corpuz-Raros, 1972: 256
Cheletomimus (Hemicheyletia) makilingensis (Corpuz-Raros): Fain, Bochkov & Corpuz-Raros, 2002: 39
Type locality: Laguna, Philippines
Distribution: Philippines
Host: Plant host Schizostachium
Hemicheyletia mexicana Thewke & Enns, 1979: 223
Cheletomimus (Hemicheyletia) mexicanus: Fain, Bochkov & Corpuz-Raros, 2002: 52
Type locality: three miles north of San Cristobal, Las Casas, Chiapas
Distribution: Mexico (Chiapas)
Hemicheyletia newyorkensis Delfinado & Khaing-Fields, 1976: 192; Fain, Bochkov & Corpuz-Raros, 2002: 28
Type locality: New York, Long Island
Distribution: USA
Hemicheyletia nucella Chaudhri & Akbar, 1985: 166; Haq & Afzal, 2007: 2206
Cheletomimus nucella (Chaudhri et Akbar): Halliday, Kamran & Bashir, 2018: 84
Type locality: Pakistan
Distribution: Pakistan
Hemicheyletia omissa Tseng, 1977: 222
Cheletomimus (Hemicheyletia) omissa (Tseng): Fain, Bochkov & Corpuz-Raros, 2002: 60
Cheletomimus omissus (Tseng): Xia, 2010: 161
Type locality: Taiwan
Distribution: China, Taiwan
Hemicheyletia parcus Afzal, Ali & Akbar, 2003: 89
Cheletomimus parcus (Afzal, Ali et Akbar): Halliday, Kamran & Bashir, 2018: 85
Type locality: Pakistan
Distribution: Pakistan
Hemicheyletia parodos Chaudhri & Akbar, 1985: 173.
Cheletomimus parodos (Chaudhri еt Akbar): Halliday, Kamran & Bashir, 2018: 85
Type locality: Pakistan
Distribution: Pakistan
Paracheyletia recki Volgin, 1966b: 283; 1969: 183; Fain & Bochkov, 2001: 51
Cheletomimus (Hemicheyletia) recki (Volgin): Khaustov, 2021: 214
Type locality: Voronezh Reserve
Distribution: Russia
Host: ant hill
Hemicheyletia reticulata Jeffrey & Campbell, 1975: 103
Cheletomimus (Hemicheyletia) reticulata (Jeffrey et Campbell): Fain, Bochkov & Corpuz-Raros, 2002: 42
Type locality: Scotland, Argyll, Island of Gigha
Distribution: Great Britain
Hemicheyletia rostella Summers & Price, 1970: 17
Cheletomimus (Hemicheyletia) rostellus (Summers et Price): Fain, Bochkov & Corpuz-Raros, 2002: 36
Type locality: California (Dorrington, Calaveras Co., etc.)
Distribution: USA (California)
Host: from rotting logs
Hemicheyletia salvus Chaudhri & Akbar, 1985: 149
Cheletomimus salvus (Chaudhri et Akbar): Halliday, Kamran & Bashir, 2018: 85
Type locality: Pakistan
Distribution: Pakistan
Cheyletia scutellata De Leon, 1962: 130
Paracheyletia scutellata (De Leon): Muma, 1964: 246
Andrecheyla scutellata (De Leon): Volgin, 1969: 219
Hemicheyletia scutellata (De Leon): Summers & Price, 1970: 14
Cheletomimus (Hemicheyletia) scutellatus (De Leon): Fain, Bochkov & Corpuz-Raros, 2002: 42
Type locality: Florida
Distribution: Mexico (Chiapas), USA (Florida)
Hosts: Swietenia mahagoni (Meliaceae), Xylosma elliptica (Salicaceae)
Hemicheyletia serrula Summers & Price, 1970: 15
Cheletomimus (Hemicheyletia) serrula (Summers et Price): Fain, Bochkov & Corpuz-Raros, 2002: 48
Type locality: Darwin Res. Station
Distribution: Galapagos Is.
Hemicheyletia tanosis Kamran, Khan & Nazir, 2005: 52
Cheletomimus tanosis (Kamran, Khan et Nazir): Halliday, Kamran & Bashir, 2018: 84
Type locality: Chak 98/M.L., 14 km East of Karor, Layyah, Pakistan
Distribution: Pakistan
Host: from pomegranate (Punica granatum)
Hemicheyletia transversa Corpuz-Raros, 1972: 257; Corpuz-Raros & Sotto, 1972: 170
Cheletomimus (Hemicheyletia) transversa (Corpuz-Raros): Fain, Bochkov & Corpuz-Raros, 2002: 39
Hemicheyletia morii Ehara & Ghani Ibrahim, 1988: 237 – Kepong, Selangor, Malaysia. Syn. by Fain, Bochkov & Corpuz-Raros, 2002: 39
Type locality: Laguna, the Philippines
Distribution: Malaysia, Philippines, Singapore
Host: Plant hosts Centrostemma multiflora and Bambusa sp.
Hemicheyletia trepsis Chaudhri & Akbar, 1985: 175
Cheletomimus trepsis (Chaudhri & Akbar): Halliday, Kamran & Bashir, 2018: 85
Type locality: Pakistan
Distribution: Pakistan
Cheyletia tropica Shiba, 1976: 169
Hemicheyletia tropica (Shiba): Ehara & Ghani, 1988: 240
Cheletomimus (Hemicheyletia) tropica (Shiba): Fain, Bochkov & Corpuz-Raros, 2002: 48
Type locality: Pasoh Forest Reserve
Distribution: Malaysia
Hemicheyletia uichancoi Corpuz-Raros, 1972: 258
Cheletomimus (Hemicheyletia) uichancoi (Corpuz-Raros): Fain, Bochkov & Corpuz-Raros, 2002: 57
Type locality: Laguna, the Philippines
Distribution: Philippines, Vanuatu
Host: Plant host Leucaena leucocephala (Mimosaceae)
Hemicheyletia umbo Chaudhri & Akbar, 1985: 185
Cheletomimus umbo (Chaudhri et Akbar): Halliday, Kamran & Bashir, 2018: 85
Type locality: Pakistan
Distribution: Pakistan
Hemicheyletia vescus Qayyum & Chaudhri, 1979: 167
Cheletomimus (Hemicheyletia) vescus (Qayyum et Chaudhri): Fain, Bochkov & Corpuz-Raros, 2002: 38; Bochkov, Hakimitabar & Saboori, 2005: 107
Hemicheyletia laguncula Rasool & Chaudhri, 1979: 4 – Pakistan. Syn. by Fain, Bochkov & Corpuz-Raros, 2002: 38
Cheletomimus vescus (Qayyum et Chaudhri): Halliday, Kamran & Bashir, 2018: 85
Type locality: Pakistan
Distribution: Iran, Pakistan
Hemicheyletia vivus Chaudhri & Akbar, 1985: 177
Cheletomimus vivus (Chaudhri et Akbar): Halliday, Kamran & Bashir, 2018: 85
Type locality: Pakistan
Distribution: Pakistan
Paracheyletia volgini Cunliffe, 1963: 197
Dendrocheyla volgini: Volgin, 1969: 210–211
Hemicheyletia volgini (Cunliffe): Summers & Price, 1970: 14
Cheletomimus (Hemicheyletia) volgini (Cunliffe): Fain, Bochkov & Corpuz-Raros, 2002: 48
Type locality: Mulunga (D.R. Congo)
Distribution: D. R. Congo, Rwanda
Cheyletia wellsi Baker, 1949: 300; De Leon, 1962: 132
Paracheyletia wellsi (Baker): Volgin, 1955: 169; Mumma, 1964: 245; Garrett & Haramoto, 1967: 410
Dendrocheyla wellsi (Baker): Volgin, 1969: 211
Hemicheyletia wellsi (Baker): Summers & Price, 1970: 15; Flechtmann, 1971: 31; Corpuz – Raros & Sotto, 1977: 157; Tseng, 1977: 224; Gerson, 1994: 445; Feres & Flechtmann, 1995: 535
Cheletomimus (Hemicheyletia) wellsi: Fain, Bochkov & Corpuz-Raros, 2002: 45;
Doğan & Ayyildiz, 2004: 1; Bochkov, Hakimitabar & Saboori, 2005: 107; Buosi et al., 2006: 7
Cheletomimus wellsi: Xia, 2010: 161
Type locality: Azores, intercepted at Philadelphia, Pa.
Distribution: Argentina, Australia, Azores, Brazil, Chile, China, Cuba, Iran, Jamaica, Japan, Malaysia, Mexico, Panama, Philippine Is., Puerto Rico, Taiwan (may be wellsinus), Trinidad, Turkey, USA (incl. Hawaii)
Paracheyletia wellsina De Leon, 1967: 34
Hemicheyletia wellsina (De Leon): Summers & Price, 1970: 18; Corpuz-Raros & Sotto, 1977: 158; Ehara & Abdul Ghani, 1988
Hemicheyletia arecana Tjying, 1972: 8 – Taiwan. Syn. by Fain, Bochkov & Corpuz-Raros, 2002: 45
Cheletomimus (Hemicheyletia) wellsina (De Leon): Fain, Bochkov & Corpuz-Raros, 2002: 45
Cheletomimus (Hemicheyletia) wellsinus (De Leon): Torre Santana, 2015: 50
Cheletomimus wellsina (De Leon): Xia, 2010: 161
Type locality: Trinidad (Caribbean)
Distribution: Algeria, Benin (Dahomey), Cameroon, China, Cuba, Guatemala, Malaysia, Philippine Is., Rwanda, Taiwan, Trinidad, USA (Florida, Texas)
Philippicheyla Corpuz-Raros, 1972: 265 (genus)
Philippicheyla Corpuz-Raros: Fain & Bochkov, 2001: 299 (subgenus of Hemicheyletia)
Type species: Philippicheyla filipina Corpuz-Raros, 1972
Cheletomimus (Philippicheyla) ascutatus Fain, Bochkov & Corpuz-Raros, 2002: 60
Type locality: Elizabeth La, California
Distribution: USA (California)
Host: from inner tree bark
Cheletomimus (Philippicheyla) crowei Bochkov & Otto, 2010: 9
Type locality: New Zealand
Distribution: New Zealand
Host: from avocado
Philippicheyla notelaeae Gerson, 1994: 437
Cheletomimus (Philippicheyla) notelaeae (Gerson): Fain, Bochkov & Corpuz-Raros, 2002: 60
Type locality: Indooroopilly, Queensland
Distribution: Australia (Queensland)
Host: bud scales on branches of Notelaea longifolia (Oleraceae)
Philippicheyla philipina Corpus-Raroz, 1972: 256
Cheletomimus (Philippicheyla) filipina (Corpus-Raroz): Fain, Bochkov & Corpuz-Raros, 2002: 60
Type locality: Mt. Makiling, Laguna, the Philippines
Distribution: Philippines
Host: on decomposing bark of a fallen log
Species inquirendae:
Cheletomimus binus Tseng, 1973: 76; Salarzehi, 2018:
Type locality: Taiwan
Distribution: Iran, Taiwan
Cheletomimus bisetosus Tseng, 1977
Type locality: Taiwan
Distribution: Taiwan
Cheletomimus cambio Aheer, Akbar & Chaudhri, 1994: 346; Aheer, Akbar & Chaudhri, 1998: 125; Gerson, Fain & Smiley, 1999: 50; Fain, Bochkov & Corpuz-Raros, 2002: 28; Haq & Afzal, 2007: 2206; Halliday, Kamran & Bashir, 2018: 84
Type locality: Pakistan
Distribution: Pakistan
Cheletomimus cantor Rasool, Chaudhri & Akbar, 1980: 28; Chaudhri & Akbar, 1985: 145; Aheer, Akbar & Chaudhri, 1998: 125; Gerson, Fain & Smiley, 1999: 50; Fain, Bochkov & Corpuz-Raros, 2002: 28; Halliday, Kamran & Bashir, 2018: 84
Type locality: Pakistan
Distribution: Pakistan
Cheletomimus citrosinensis Patxot & Goff, 1985: 157
Type locality: Oahu I., Mililani
Distribution: USA (Hawaiian Is.)
Cheletomimus domitus Chaudhri & Akbar, 1985: 150; Aheer, Akbar & Chaudhri, 1998: 126; Halliday, Kamran & Bashir, 2018: 84
Type locality: Pakistan
Distribution: Pakistan
Cheletomimus fissilis Chaudhri & Akbar, 1985: 155; Aheer, Akbar & Chaudhri, 1998: 126; Halliday, Kamran & Bashir, 2018: 84
Type locality: Pakistan
Distribution: Pakistan
Cheletomimus flecto Aheer, Akbar & Chaudhri, 1998: 126; Fain, Bochkov & Corpuz-Raros, 2002: 28; Aheer, Akbar & Chaudhri, 1998: 126; Halliday, Kamran & Bashir, 2018: 84
Type locality: Pakistan
Distribution: Pakistan
Cheletomimus heredis Qayyum & Chaudhri, 1979: 9; Halliday, Kamran & Bashir, 2018: 84
Type locality: Pakistan
Distribution: Pakistan
Cheletomimus larme Aheer, Akbar & Chaudhri, 1994: 348; 1998: 125; Halliday, Kamran & Bashir, 2018: 84
Cheletomimus larmae Aheer, Akbar & Chaudhri: Gerson, Fain & Smiley, 1999: 51; Fain, Bochkov & Corpuz-Raros, 2002: 28
Type locality: Pakistan
Distribution: Pakistan
Cheletominus [sic, err. typogr.] minutus Soliman, 1975: 99
Type locality: Borg el Arab
Distribution: Egypt
Cheletomimus sialkotensis Chaudhri & Akbar, 1985: 147; Halliday, Kamran & Bashir, 2018: 85
Type locality: Pakistan
Distribution: Pakistan
Cheletomimus telma Chaudhri & Akbar, 1985: 149; Aheer, Akbar & Chaudhri, 1998: 130; Halliday, Kamran & Bashir, 2018: 85
Type locality: Pakistan
Distribution: Pakistan
Cheletomimus trema Aheer, Akbar & Chaudhri, 1998: 126; Fain, Bochkov & Corpuz-Raros, 2002: 28; Halliday, Kamran & Bashir, 2018: 85
Type locality: North West Frontier Province, Dargai
Distribution: Pakistan
Cheletomimus zamia Aheer, Akbar & Chaudhri, 1994: 348; 1998: 125; Gerson, Fain & Smiley, 1999: 51; Fain, Bochkov & Corpuz-Raros, 2002: 28; Halliday, Kamran & Bashir, 2018: 85
Type locality: Pakistan
Distribution: Pakistan
Cheletophanes Oudemans, 1904: 162
Type species: Cheyletus montandoni Berlese et Trouessart, 1889
Cheyletus Montandoni Berlese & Trouessart, 1889: 133
Cheletophanes montandoni (Berlese et Trouessart): Oudemans, 1904b: 162; 1906: 140; Baker, 1949 : 303; Volgin, 1969: 165; Summers & Price, 1970: 35
Type locality: Broşteni
Distribution: Romania
Host: Aradus pictus (Heteroptera: Aradidae)
Cheletophanes peregrinus Berlese, 1921: 194; Baker, 1949: 304
Type locality: Florence
Distribution: Italy
Host: „in detritis corticis pinorum, ad terram”
Cheletophyes Oudemans, 1914: 101
Type species: Cheletophyes vitzthumi Oudemans, 1914
Cheletophyes apicola Fain, Lukoschus & Nadchatram, 1980: 311; Fain & Bochkov, 2001: 63
Type locality: Kuala Pilah, Negri Sembilan
Distribution: Malaysia
Host: nest of Xylocopa (Platynopoda) latipes (Hymenoptera: Apidae)
Cheletophyes aurorae Haitlinger, 2000: 86
Type locality: Arusha
Distribution: Tanzania
Host: Xylocopinae (Hymenoptera: Apidae)
Cheletophyes clavipilis Fain, Lukoschus & Nadchatram, 1980: 310
Type locality: Kuala Pilah, Negri Sembilan
Distribution: Malaysia
Host: nest of Xylocopa (Platynopoda) latipes (Hymenoptera: Apidae)
Cheletophyes decorus Bochkov & Klimov, 2005: 19
Type locality: Cam Ranh Peninsula, Khanh Hoa Prov. (Vietnam), Trang Prov., Khao Chong Nature Education Centre (Thailand)
Distribution: Thailand, Vietnam
Host: Xylocopa tranquebarica (Hymenoptera: Apidae)
Cheletophyes eckerti Summers & Price, 1970: 40; Fain & Bochkov, 2001: 64
Type locality: Ludhiana (India)
Distribution: Angola, D.R. Congo, India
Host: Xylocopa (Koptortosoma) aestuans (Hymenoptera: Apidae)
Cheletophyes indiacus Smiley & Whitaker, 1981: 123; Fain & Bochkov, 2001: 63
Cheletophyes xylocopae Ramaraju & Mohanasundaram, 1999: 121 – Tamil Nadu,
Coimbatore, India, from Xylocopa sp. (Hymenoptera: Apidae)
Type locality: India
Distribution: India
Host: Xylocopa sp. (Hymenoptera: Apidae)
Cheletophyes mbomba Klimov, Bochkov & OConnor, 2006: 56
Type locality: Bas-Uele Prov., Bambesa
Distribution: DR Congo
Host: Xylocopa torrida (Hymenoptera: Apidae)
Cheletophyes occisor Bochkov & Klimov, 2005: 18
Type locality: Popondetta Distr., Popondetta
Distribution: Papua New Guinea
Host: Xylocopa aruana (Hymenoptera: Apidae)
Cheletophyes panamensis Klompen, Méndez & Lukoschus, 1984: 249; Fain & Bochkov, 2001: 63
Type locality: Capira, Aguacate (Panama)
Distribution: Costa Rica, Panama
Host: Xylocopa (Megaxylocopa) frontalis (Hymenoptera: Apidae)
Cheletophyes torridae Klimov, Bochkov & OConnor, 2006: 55
Type locality: Sangmelima, Fulasi
Distribution: Cameroon
Host: Xylocopa torrida (Hymenoptera: Apidae)
Cheletophyes vespae Fain & Bochkov, 2001: 65
Type locality: KG. Duson, Tampin, N. Sembilan
Distribution: Malaysia (N. Sembilan)
Host: Vespa tropica (Hymenoptera: Vespidae)
Cheletophyes vitzthumi Oudemans, 1914:101; 1915: 51; Volgin, 1965: 290; 1969: 160; Summers & Price, 1970: 40; Fain & Bochkov, 2001: 63
Type locality: Willowmore, Cape Province
Distribution: South Africa
Host: Xylocopa (Koptorthosoma) caffra (Hymenoptera: Apidae)
Species inquirendae (Fain & Bochkov, 2001: 61):
Cheletophyes deodikari Putatunda & Kapil, 1988: 318
Type locality: Haryana, Hisar
Distribution: India
Host: Xylocopa fenestrata (Hymenoptera: Apidae)
Cheletophyes harnaji Putatunda & Kapil, 1988: 320
Type locality: India
Distribution: India
Host: Xylocopa pubescens (Hymenoptera: Apidae)
Cheletophyes newtoni Putatunda & Kapil, 1988: 322
Type locality: Haryana, Hisar
Distribution: India
Host: Xylocopa fenestrata (Hymenoptera: Apidae)
Cheletophyes orientalis Putatunda & Kapil, 1988: 324
Type locality: Haryana, Hisar
Distribution: India
Host: Xylocopa fenestrata (Hymenoptera: Apidae)
Cheletophyes ruttneri Putatunda & Kapil, 1988: 325
Type locality: Haryana, Hisar
Distribution: India
Host: Xylocopa fenestrata (Hymenoptera: Apidae)
Cheletophyes shendei Putatunda & Kapil, 1988: 326
Type locality: India
Distribution: India
Host: Xylocopa fenestrata (Hymenoptera: Apidae)
Cheyletus Latreille, 1796: 179
Type species: Acarus eruditus Schrank, 1781
Cheyletus ascutatus Bochkov & OConnor, 2003 : 327
Type locality: Leyte Is., 10 km N, 4,5 km E Baybay
Distribution: Philippines (Leyte)
Host: Batomys salomonseni (Rodentia: Muridae)
Cheyletus attiahi Yousef & Issa, 1972: 42
Type locality: Egypt
Distribution: Egypt
Cheyletus batomysi Bochkov & OConnor, 2003: 329
Type locality: Bukidnon Prov., Mindanao Is., Mt. Katangland Range, Camp Phillips, 2250 m
Distribution: Philippines
Host: Batomys salomonseni (Rodentia: Muridae)
Cheyletus bidentatus Fain & Nadchatram, 1980: 194
Type locality: Bukit Lanjan Forest Reserve, Selangor, Segamat, Labis Forest Reserve (Johore)
Distribution: Iran, Malaysia (Selangor, Johore)
Host: nest of Hylopetes spadiceus (Rodentia: Sciuridae)
Cheyletus cacahuamilpensis Baker, 1949: 282; Volgin, 1949: 114; Costa, 1961: 259; 1967: 109; Summers & Price, 1970: 26; Fain, 1982: 89; Fain & Bochkov, 2001; Whitaker et al., 2007: 6; Palyvos et al., 2008: 213; Salarzehi et al., 2018: 465
Eucheyletia mungeri McGregor, 1956: 24 (syn. by Summers & Price, 1970)
Cheyletus baloghi Volgin, 1969: 116. Type: Crimea (Sevastopol), on Spermophilus [Citellus] pygmaeus (nests)
Type locality: Cave of Cacahuamilpa (Mexico, Guerrero)
Distribution: Algeria, Crimea, Greece, Iran, Israel, Mexico, Peru, USA, South Africa
Hosts: bat guano, Myotis velifer (Chiroptera: Vespertilionidae), Spermophilus [Citellus] pygmaeus (Rodentia: Sciuridae, nests), Microtus guentheri (Rodentia: Arvicolidae)
Cheyletus carnifex Zachvatkin, 1935: 27; Rohdendorf, 1940: 89; Volgin, 1969: 112; Torre Santana, 2015
Cheyletus aversor Rohdendorf, 1940: 86; Baker, 1949: 283; Volgin, 1955: 163; 1969: 89; Wharton & Arlian, 1972: 719; Žďárková, 1979; Shen, 1991; Lin et al., 2000; Xia et al., 2000; Lukáš et al., 2007: 97 – Bulgaria, China, Czech Republic, Russia, Tajikistan, Uganda, Ukraine, USA
Cheyletus beauchampi Baker, 1949: 282 – Portugal, intercepted at Boston, Mass. (syn. by Summers & Price, 1970)
Cheyletus acarophagus Zacher & Soliman, 1967: 25
Cheyletus allactaga Fain & Lukoschus, 1981: 122 – Bulgan-Gol, Mongolia, on Allactaga sibirica (Rodentia: Dipodidae)
Cheyletus zaheri Hassan & Rakha, 1982: 89
Type locality of Cheyletus carnifex: Moscow
Type locality of Cheyletus aversor: Ordzhonikidze (Caucasus) and Altai
Type locality of Cheyletus acarophagus: Egypt
Type locality of Cheyletus zaheri: Egypt
Distribution: Bulgaria, China, Czech Republic, Iran, Russia, Tajikistan, Uganda, Ukraine
Hosts: Allactaga sibirica (Rodentia: Dipodidae)
Acarus eruditus Schrank, 1781: 513
Cheyletus robertsoni Brady, 1875: 302
Cheletes eruditus (Schrank): Oudemans, 1903: 118; 1906: 84
Eutarsus cancriformis Hessling, 1852: 258
Cheyletus ferox Berlese et Trouessart, 1889: 14; Banks, 1906: 134 – “Port-Praslin Nieuw Ireland”.
Cheletes ferox (Berlese et Trouessart): Oudemans, 1903a: 83; 1906: 115
Cheyletus eburneus Hardy, 1933: 336
Cheyletus seminivorus, Packard, 1870: 665; Ewing, 1909: 76; 1912: 416
Cheyletus doddi Baker, 1949: 279 – Ithaca, N.Y.
Cheyletus desitus Qayyum & Chaudhri, 1977: 90
Cheyletus mortelmansi Fain, 1972: 37 – Zoo d’Anvers
Cheyletus eruditus (Schrank): Kramer, 1876: 188; Canestrini, 1886: 171; Haller, 1882: 312; Oudemans, 1903: 118; Jablonowski, 1918: 4; Hughes, 1948: 102; Baker, 1949: 278; Starkoff & Starkoff, 1950: 96; Turk, 1953: 18; Franz, 1954: 367; Haarløv, 1957: 26; Cwilich & Hadani, 1965: 54; Volgin, 1969: 78; Shiba, 1969b: 101; 1969a: 159; Shen, 1975; Mumcuoglu, 1976: 365; Kumada et al., 1978: 262; Hallas, 1978: 30; Whitaker et al., 2007: 6; Xia, 2010: 161; Torre Santana, 2015; Halliday, Kamran & Bashir, 2018: 87
Cheletes strennus Oudemans, 1904: 161; 1906: 8.3
Cheyletus strennus: Baker, 1949: 280
Type locality: Austria
Distribution: cosmopolitic
Hosts: Coracopsis nigra barkleyi (Psittaciformes: Psittrichasiidae), Didelphis virginiana (Didelphiomorpha: Didelphidae), Pipistrellus abramus (Chiroptera: Vespertilionidae), Peromyscus leucopus (Rodentia: Cricetidae, Neotominae), Menetes berdmorei, Rattus norvegicus (Rodentia: Muridae), Sciurus carolinensis (Rodentia: Sciuridae), Bassariscus astutus (Carnivora: Procyonidae)
Eucheyletia funisciuri Fain, 1972: 38
Cheyletus funisciuri (Fain): Fain & Bochkov, 2001: 100
Type locality: Rwanda
Distribution: DR Congo, Rwanda
Host: Funisciurus carruthersi (Rodentia: Sciuridae)
Cheyletus gerbillicola Fain & Lukoschus, 1981: 6
Type locality: Askham, South of Kalahari region
Distribution: South Africa
Host: Gerbillurus paeba (Rodentia: Muridae, Gerbillinae)
Eucheyletia kivuensis Fain, 1972: 39
Cheyletus kivuensis (Fain): Fain & Bochkov, 2001: 100
Type locality: Kivu
Distribution: D.R. Congo
Host: Rodentia indet.
Cheyletus kuznetzovi Bochkov & Khaustov, 1999: 116; Dogan, Jalaeian & Kamali, 2011: 782
Type locality: Dagskiy Reserve (Turkmenistan)
Distribution: Iran, Turkmenistan
Host: from ant hill
Cheyletus legendrei Fain, 1982: 83
Type locality: Tananarive
Distribution: Madagascar
Host: nest of Foudia madagascariensis (Passeriformes: Ploceidae)
Cheyletus linsdalei Baker, 1949: 281; Volgin, 1969: 111; Summers & Price, 1970: 29
Type locality: California, Monterey Co.
Distribution: USA (California)
Host: Citellus beecheyi (Rodentia: Sciuridae)
Cheyletus mafekingensis Fain, 1982: 86
Type locality: Mafeking
Distribution: South Africa
Host: Philetarius socius (Passeriformes: Ploceidae)
Cheletes malaccensis Oudemans, 1903a: 84; 1906: 88
Cheyletus malaccensis (Oudemans): Baker, 1949: 284; Hughes, 1961: 197; Gerson, 1967: 361; Shiba, 1969a: 161; Flechtmann, 1971: 30; Regev, 1974 : 85; Shen, 1975; Nakada, 1975: 151; Kumada et al., 1978: 262; Caceres & Fain, 1978: 302; Rosa & Flechtmann, 1979: 196; Žďárková, 1979; Fain & Nadchatram, 1980: 191; Saleh et al., 1986: 37; Whitaker & Dietz, 1987: 194; Rostom, 1993 : 29; Mariana et al., 2000: 714; Lukáš et al., 2007: 97; Palyvos et al., 2008: 213; Xia, 2010: 162; Torre Santana, 2015; Halliday, Kamran & Bashir, 2018: 87
Cheyletus ugandanus Lawrence, 1954: 65 – Karamoja, Uganda, nest of weaver-bird
Cheyletus ayyazi Akbar, Aheer & Ishtiaq, 1993: 293 – Pakistan
Cheyletus polymorphus Volgin, 1949: 584; 1969: 90; Sosnina et al., 1969: 370 – Greece, Bulgaria
Cheyletus caucasicus Zachvatkin, 1949: 288
Cheyletus egypticus Elbadry, 1969: 157 – Egypt
Cheyletus baridos Akbar, Rahi & Chaudhri, 1988: 5 – Pakistan
Cheyletus wahndoensis Akbar & Aheer, 1994: 343 – Pakistan
Cheyletus avidus Qayyum & Chaudhri, 1977: 89 – Pakistan
Cheyletus mianiensis Farooq, Akbar & Qureshi, 2000: 257 – Pakistan
Cheyletus rafiquiensis Farooq, Akbar & Qureshi, 2000: 259 – Pakistan
Cheyletus phantosis Akbar & Aheer, 1994: 342 – Pakistan
Cheletes vorax Oudemans, 1903b: 84; 1906: 122 – Mariana Is., bats (Chiroptera)
Cheyletus vorax (Oud.): Volgin, 1969: 99
Cheyletus rohdendorfi Zachvatkin, 1949: 290; Volgin, 1969: 80; Rekk, 1976: 68 – Baku, Azerbaidjan; Armenia
Cheyletus infensus Akbar, Aheer & Shaudhri, 1993: 295 – Pakistan
Cheletes fortis Oudemans, 1904: 161; 1906: 96 – on parrot skin, New Guinea
Cheyletus fortis (Oud.): Baker, 1949: 280; Tanaka, 1953: 73; Shiba, 1966: 142; 1969a: 161; Volgin, 1969: 87; Summers & Price, 1970: 28; Fain & Nadchatram, 1980: 191; Patxot & Goff, 1985: 162; Mariana et al., 2000: 714; Ezequiel et al., 2001: 913
Type locality: Malacca, Straits Settlements
Distribution: Algeria, Australia (Queensland), Azerbaidzan, Bangladesh, Brazil, Bulgaria, Burma, China, Czech Rep., D. R. Congo, Cuba, Greece, India, Iran, Israel, Japan, Malaysia, New Guinea, Pakistan, Philippines, Peru, Russia, Rwanda, Saudi Arabia, Senegal, Tadjikistan, Taiwan, USA (Hawaian is.)
Hosts: Psittinus incertus (Psittaciformes: Psittacidae), Pipistrellus abramus (Chiroptera: Vespertilionidae), Callosciurus sladeni midas, Petinomys setosus (Rodentia: Sciuridae), Thalpomys [Akodon] reinhardti (Rodentia: Arvicolidae, Sigmodontinae); Microtus arvalis (Rodentia: Arvicolidae), nests of birds and rats; for Cheyletus fortis: rodents (Callosciurus, Ratufa, Rattus). According to Fain & Nadchatram (1980), „accidental contamination“
Cheyletus malayensis Cunliffe, 1962: 201; Patxot & Goff, 1985: 161; Fain & Nadchatram, 1980: 192; Mariana et al., 2000: 714; Torre Santana, 2015
Type locality: Rantau Panjang, 5 mi N of Klang, Selangor, Malaysia
Distribution: Cuba, India, Iran, Malaysia, USA (Hawaian Is. – Oahu)
Hosts: nests of Lonchura malacca (= Munia atricapilla), Munia striata, Pycnonotus goiaver (Passeriformes: Estrildidae)
Cheyletus misonnei Fain & Lukoschus, 1981: 1
Type locality: Aminuis
Distribution: Namibia
Host: Tatera sp. (Rodentia: Arvicolidae)
Cheyletus morinus Barilo, 1986: 27
Type locality: Uzbekistan
Distribution: Uzbekistan
Cheyletus nidicolus Fain, 1972: 37; 1979: 280
Type locality: Butare
Distribution: Rwanda, South Africa
Host: birds’ nests
Cheyletus philippinensis Corpuz-Raros, 1988: 328
Type locality: Philippines
Distribution: Philippines
Hosts: woodpeckers (Picoidea: Picidae), Harpactes ardens (Trogoniformes: Trogonidae)
Cheyletus pluridens Fain & Nadchatram, 1980: 195
Type locality: Kuantan (Pahang), Ulu Langat Forest Reserve (Selangor)
Distribution: Malaysia (Selangor)
Host: Rhinosciurus laticaudatus (Rodentia: Sciuridae)
Cheyletus pseudomalaccensis Fain, 1982: 86
Type locality: Potchefstroom, Transvaal
Distribution: South Africa
Host: Plocepasser mahali (Passeriformes: Ploceidae)
Cheyletus punctulatus Fain & Lukoschus, 1981: 121
Type locality: 20 km N. of Oned Draa, Southern Morocco
Distribution: Morocco
Host: Gerbillus pyramidum (Rodentia: Muridae, Gerbillinae)
Cheyletus rarus Shiba, 1969: 162
Type locality: Kuwabara-- sho, Matsuyama
Distribution: Japan
Cheyletus rashtiensis Salarzehi, Hajizadeh & Ueckermann, 2018: 641; Khaustov, 2021: 228
Type locality: Rasht (Iran)
Distribution: Iran, Russia (Siberia)
Cheyletus rwandae Fain, 1972: 38; 1979: 282
Type locality: Butare
Distribution: D. R. Congo, Rwanda
Host: Tachyoryctes rwandae (Rodentia: Rhizomyidae)
Cheyletus schneideri (Oud.): Oudemans, 1902: 15; Turk, 1946: 805; Baker, 1949: 281; Volgin, 1969: 102
Cheletes schneideri Oudemans, 1904: 16; 1906: 84
Type locality: San Remo (Italy)
Distribution: England, Italy
Cheletes strennus Oudemans, 1904: 161; 1906: 83
Cheyletus strennus: Baker, 1949: 280
Type locality: unknown
Distribution: unknown
Host: unknown
Eucheyletia tanzaniensis Fain, 1972 : 39
Cheyletus tanzaniensis (Fain): Fain & Bochkov, 2001: 101
Type locality: Tanganyika
Distribution: Tanzania
Hosts: Coleura gallarum (Chiroptera: Emballonuridae); nests of rodents (Dasymys incomtus, Pelomys walambae, Textor sp., Lagonosticta senegallus)
Cheletes trouessarti Oudemans, 1902: 16; 1903: 129; 1906: 88
Cheyletus macrocherus Hardy, 1933: 350
Cheyletus furibundus Rohdendorf, 1940: 85 – Kirzhach, Russia (Vladimirskaya obl.)
Cheyletus davisi Baker, 1949: 283 – Italy, intercepted at Boston, Mass. (syn. by Summers & Price, 1970)
Cheyletus truculentus Volgin, 1949: 586; 1969: 113 – Karaganda, Kazakhstan
Cheyletus praedabundus Kuzin, in Rohdendorf, 1940: 85 – Rostov on Don, Russia
Cheyletus woodroffei Jeffrey, 1980: 46 – Upton Grey, Hampshire, England, from a bat roost (probably Plecotus auritus)
Cheyletus trouessarti (Oudemans): Oudemans, 1903: 16; Baker, 1949: 280; Volgin, 1969: 109; Shen, 1975; Caceres & Fain, 1978: 302; Mumcuoglu, 1976: 365; Hallas, 1978: 30; Žďárková, 1979; Haitlinger, 1982: 442; Philips & Dindal, 1990: 177; Lucza, Ripka & Saly, 1996; Bochkov, Hakimitabar & Saboori, 2005: 106; Lukáš et al., 2007: 97; Xia, 2010: 162
Type locality: unknown
Distribution: Bulgaria, China, Czech Rep., Denmark, England, Holland, Hungary, India, Iran, Kazakhstan, Peru, Poland, Russia, Switzerland, USA
Cheyletus trux Rohdendorf, 1940: 87; Baker, 1949: 282; Rekk, 1976: 68; Vysotskaja, 1977: 188; Efremova & Bochkov, 1990: 99; Eliopoulos & Papadoulis, 2001:
Cheyletus hendersoni Baker, 1949: 279 – Arkansas Cave, USA; “Czechoslovakia”
Cheyletus tenuipilis Fain, Feldman-Musham & Mumcuoglu, 1980: 35 – La Louvière (Belgium); Louadi & Robaux, 1992: 183
Type locality: Ivanovo, Tatarstan
Distribution: Algeria, Belarus, Belgium, “Czechoslovakia”, Georgia, Greece, Israel, Russia, Switzerland, Ukraine, USA (Arkansas, New Mexico, Colorado)
Host: nests, stored grain, house dust
Cheletes venator Vitzthum, 1920: 2
Cheyletus venator (Vitzthum) Volgin, 1969: 119
Type locality: Amani
Distribution: Tanzania
Host: nest of Xylocopa (Afroxylocopa) nigrita (Hymenoptera: Apidae)
Cheyletus vivatus Qayyum & Chaudhri, 1977: 92; Halliday, Kamran & Bashir, 2018: 87
Cheyletus spatiosus Qayyum & Chaudhri, 1977: 94
Cheyletus tutela Qayyum & Chaudhri, 1977: 95
Type locality: Pakistan
Distribution: Pakistan
Cheyletus volgini Fain & Bochkov, 2001: 95
Type locality: Russia
Distribution: Russia
Cheyletus zumpti Fain, 1972: 38; 1979: 281
Type locality: Garamba Nat. Park
Distribution: D. R. Congo, South Africa, Nigeria, Rwanda
Host: nests of rodents; type from Tatera (Rodentia: Arvicolidae)
Species inquirendae:
Cheletes acer Oudemans, 1904b:162; 1906: 112
Cheyletus acer (Oudemans): Baker, 1949: 286; Volgin, 1969: 94
Type locality: Colombia
Distribution: Colombia
Host: Aves (Birdskin)
Cheletes alacer Oudemans, 1904b: 162; 1906: 108
Cheyletus alacer Oudemans: Baker, 1949: 285
Type locality: Colombia
Distribution: Colombia
Host: skin of Pyrrhura cruentata (Psittaciformes: Psittacidae)
Cheletes audax Oudemans, 1904: 162; 1906: 99
Cheyletus audax (Oudemans): Baker, 1949: 285; Volgin, 1969: 93
Type locality: New Guinea
Distribution: New Guinea
Host: skin (Aves indet.)
Cheyletus burmiticus Cockerell, 1917: 41; Baker, 1949: 287 (inc. sedis)
Type locality: Burma
Distribution: Burma
Host: Burmese amber
Cheyletus clavispinus Banks, 1902: 172; 1906: 746
Type locality: Indianapolis (Indiana)
Distribution: USA (Indiana)
Host: Beneath wings of an Aradus (Heteroptera)
Cheyletus crassus Qayyum & Chaudhri, 1977: 87; Chaudhri, Akbar & Rasool, 1979: 153; Akbar, Rahi & Chaudhri, 1988: 5; Gerson, Fain & Smiley, 1999: 55; Farooq, Akbar & Qureshi, 2000: 257; Fain & Bochkov, 2001: 84; Halliday, Kamran & Bashir, 2018: 86
Type locality: Pakistan
Distribution: Pakistan
Cheyletus ferox Trouessart, 1889: 134; Volgin, 1969: 96
Cheletes ferox, Oudemans, 1906: 115
Type locality: Praslin
Distribution: Seychelle Is.
Host: Coracopsis nigra barklyi (Psittaciformes: Psittacidae)
Cheletes intrepidus Oudemans, 1903b: 84; 1906: 119
Cheyletus intrepidus, Volgin, 1969: 98
Type locality: Colombia
Distribution: Colombia
Host: Aves indet.
Cheyletus nigriceps Mola, 1907: 43
Type locality: Europe
Distribution: Europe
Cheyletus parumsetosus Karpelles, 1884: 238
Cheletes parumsetosus: Oudemans, 1906: 81
Type locality: Europe
Distribution: Europe
Cheyletus patagiatus Nordenskiold, 1900: 37
Type locality: Finland ?
Distribution: Finland ?
Cheletes promptus Oudemans, 1904b: 161; 1906: 81
Type locality: New Guinea
Distribution: New Guinea
Host: bird’s skin
Cheletes rapax Oudemans, 1903a: 84; 1906: 103
Cheyletus rapax (Oudemans): Baker, 1949: 284; Volgin, 1969: 85
Type locality: Mariana is.
Distribution: Mariana Is.
Hosts: birds and mammals in det.
Cheletes saevus Oudemans, 1904b: 161; 1906: 84
Cheyletus saevus Oudemans: Baker, 1949: 287
Type locality: Netherlands
Distribution: Netherlands
Host: in dust
Eucheyletia Baker, 1949: 294
Type species: Eucheyletia bishoppi Baker, 1949
Eucheyletia bakeri Volgin, 1963a: 62; 1969: 149; Fain & Bochkov, 2001: 49
Type locality: Alaska
Distribution: USA (Alaska)
Host: Bombus sp. (Hymenoptera: Apidae)
Eucheyletia bishoppi Baker, 1949: 295; Volgin, 1963: 57; 1969: 142; Summers & Price,
1970: 32; Fain & Bochkov, 2001: 49; Xia et al., 2004; Xia, 2010: 162
Eucheyletia asiatica Volgin, 1955: 166; 1969: 144; Vysotzkaja, 1974: 136 – Kazakhstan, Ukraine, in nests of rodents
Eucheyletia oregonensis Smiley & Whitaker, 1981: 11 – type from Oregon, nest of Glaucomys sabrinus (Sciuridae); Shen, 1991; Xia, 2010: 162
Type locality: California
Distribution: China, Kazakhstan, Ukraine, USA (California, Oregon)
Hosts: nest of Neotoma fuscipes (Rodentia: Arvicolidae), soil samples; from Tamias striatus and nest of Glaucomys sabrinus (Sciuridae); from Sorex pacificus and S. trowbridgei (Soricomorpha: Soricidae)
Eucheyletia bothrophila Volgin, 1963a: 44; 1969: 126; Vysotskaja, 1977: 188; Efremova & Bochkov, 1990: 102; Fain & Bochkov, 2001: 49
Type locality: Khustovskiy Distr., Zakarpatskaya obl.
Distribution: Belarus, Ukraine
Host: nests of rodents (Myodes = Clethrionomys glareolus, Microtus arvalis, M. nivalis, Sylvaemus flavicollis, Sciurus vulgaris)
Eucheyletia eoa Volgin, 1963a: 52; 1969: 134; Fain & Bochkov, 2001: 49
Type locality: Olginski Region, Imanskiy Region (Primorskiy Krai)
Distribution: Russia (Primorskiy Krai in the Far East)
Host: nests of rodents (Eutamias sibiricus, Apodemus speciosus)
Cheyletus flabellifer Michael, 1878: 135; Mégnin, 1880: 241
Cheyletus (Cheyletia) flabellifer: Berlese, 1893: 64
Cheyletia flabellifera: Oudemans, 1904a: 162
Cheletia flabellifera, Oudemans, 1906: 127
Eucheyletia flabellifera (Michael): Baker, 1949: 295; Volgin, 1963a: 47; 1969: 124;
Kolebinova, 1967: 75; Shen, 1975; Haitlinger, 1982: 442; Fain & Bochkov, 2001: 49; Xia, 2010: 162; Salarzehi et al., 2018: 480
Eucheyletia taurica Volgin, 1963a: 48; 1969: 130 – Gurzuf, Crimea
Type locality: Warwikshire (England)
Distribution: Austria, Belgium, Bulgaria, China, England, Germany, Holland, Iran, Poland, Crimea
Hosts: Myodes [Clethrionomys] glareolus, Microtus arvalis (Rodentia: Arvicolidae), Sylvaemus flavicollis, S. sylvaticus (Muridae), Glis glis (Gliridae), Crocidura russula, C. suaveolens (Soricomorpha: Soricidae), Talpa europaea (Soricomorpha: Talpidae), birds
Eucheyletia hardyi Baker, 1949: 296; Volgin, 1963: 56; 1969: 141; Summers & Price, 1970: 33; Mosquera, 1989: 73; Fain & Bochkov, 2001: 49
Type locality: Harlingen, Texas
Distribution: Argentina, Chile, USA (Texas)
Host: nest of Neotoma micropus (Rodentia: Arvicolidae)
Cheyletia harpyia Rohdendorf, 1940: 90
Eucheyletia harpyia (Rohdendorf): Baker, 1949: 294; Volgin, 1963a: 59; 1969: 148; Shen, 1975; Fain & Bochkov, 2001: 49; Xia, 2010: 162
Type locality: Ivanovskaya Oblast (Russia)
Distribution: China, Russia
Eucheyletia nidicola Definado & Khaing-Fields, 1976: 193
Type locality: New York
Distribution: USA
Eucheyletia omissa Xia, Liang & Zhu, 2004; Xia, 2010: 162
Type locality: China
Distribution: China
Eucheyletia pavlovskyi Volgin, 1963a: 54; 1969: 138; Fain & Bochkov, 2001: 49
Type locality: Karaganda Area
Distribution: Kazahstan
Host: nest of Lagurus lagurus (Rodentia: Arvicolidae)
Eucheyletia sibirica Volgin, 1963a: 61; 1969: 151; Efremova & Bochkov, 1990: 98; Fain & Bochkov, 2001: 49
Type locality: Asinovskiy Region, Tomskaya obl.
Distribution: Belarus, Russia
Eucheyletia sinensis Volgin, 1963a: 50; 1969: 133; Summers & Price, 1970: 33; Whitaker, Baccus, Cleveland & Gu, 1990; Fain & Bochkov, 2001: 49; Xia, 2010: 162
Type locality: Yunnan
Distribution: China (Yunnan), Nepal, Taiwan
Hosts: Niviventer fulvescens, Rattus losea [humiliatus] celsus (Rodentia: Muridae)
Eucheyletia variformis Lin, 2000: 187
Type locality: Fujian
Distribution: China
Eucheyletia womersleyi Volgin, 1963: 59; 1969: 121
Type locality: Australia
Distribution: Australia
Granulocheyletus Fain & Bochkov, 2002: 37
Type species: Granulocheyletus corpuzrarosae Fain et Bochkov, 2002
Granulocheyletus corpuzrarosae Fain & Bochkov, 2002: 38
Type locality: Weelawadjia Cave, near Eneabba, Western Australia
Distribution: Australia
Granulocheyletus gallowayi Fain, 2003: 100
Type locality: Motunau Isl., New Zealand, from the nest of Eudyptula albosignata
(Sphenisciformes: Spheniscidae)
Distribution: New Zealand
Hemicheyletia kureatollensis Goff, 1982: 83
Granulocheyletus kureatollensis (Goff): Fain & Bochkov, 2002: 40; Fain, 2003: 101
Type locality: Kure Atoll
Distribution: Hawaiian Is.
Kenyacheylus Fain & Bochkov, 2001: 56
Type species: Kenyacheylus troglodytes Fain et Bochkov, 2001
Kenyacheylus troglodytes Fain & Bochkov, 2001: 56
Type locality: Kimakia Cave, Kiboko, 140 km SE Nairobi
Distribution: Kenya
Ker Muma, 1964: 250
Type species: Ker palmatus Muma, 1964
Ker acidalia Aheer, Akbar & Chaudhri, 1997: 119; Halliday, Kamran & Bashir, 2018: 87
Type locality: Pakistan
Distribution: Pakistan
Ker afrotropicalis Fain & Bochkov, 2001: 59
Type locality: Musha, Rwanda
Distribution: D.R. Congo, Rwanda
Host: nest of Grammomys sp. (Rodentia: Muridae)
Ker bakeri Zaher & Soliman, 1967: 24; Summers & Price, 1970: 22; Rosa & Flechtmann, 1979: 196; Žďárková, 1979; Fain & Bochkov, 2001: 58; Ezequiel et al., 2001: 913
Type locality: Giza (Egypt)
Distribution: Brazil, Egypt, Malaysia, “Czechoslovakia”
Host: house dust
Ker caeterus Barilo, 1986: 299; Fain & Bochkov, 2001: 58
Type locality: Uzbekistan
Distribution: Uzbekistan
Ker mercedesae Corpuz-Raros, 1998: 279; Fain & Bochkov, 2001: 58
Type locality: Leyte Is., Visayas St. Col. Agriculture, Baybay, Leyte
Distribution: Philippines
Ker palmatus Muma, 1964: 226; Volgin, 1969: 226; Summers & Price, 1970: 22; Patxot & Goff, 1985: 161; Shen, 1991; Lin et al., 2000; Fain & Bochkov, 2001: 58; Xia, 2010: 162
Type locality: Florida (Sebring)
Distribution: China, USA (Florida, Hawaian Is – Nihoa)
Ker pintoriensis Corpuz-Raros, 1998: 281; Fain & Bochkov, 2001: 58
Type locality: Luzon Is., Pintor, Gamu, Isabela
Distribution: Philippines
Laeliocheyletia Summers & Price, 1970: 21
Type species: Laeliocheyletia teretis Summers et Price, 1970
Laeliocheyletia teretis Summers & Price, 1970: 21
Type locality: Quetzaltepeque
Distribution: El Salvador
Host: Tenebrionidae (Coleoptera)
Lanceacheyla Xia, Klompen & Childers, 2011: 30
Type species: Lanceachyla whartoni Xia, Klompen et Childers, 2011
Lanceacheyla filipina Corpuz-Raros & Naredo, 2016: 2
Type locality: Luzon Is., Santol Cave, Biak—na-bato Nat. Park, San Miguel, Bulacan Prov.
Distribution: Philippines
Lanceacheyla whartoni Xia, Klompen & Childers, 2011: 30
Type locality: Florida, Hardee Co, 10 km S of Ona, Lilly
Distribution: USA (Florida)
Host: citrus trees
Lepidocheyla Volgin, 1963: 940
Type species: Lepidocheyla gracilis Volgin, 1963
Lepidocheyla caucasica Volgin, 1978: 940; Fain & Bochkov, 2001: 55
Hemicheyletia hissariensis Mathur & Mathur, 1981: 69 (syn of L. gracilis, acc. to Fain & Bochkov, 2001: 55) – Hissar, India (debris of the wheat straw)
Type locality: Azerbaidjan, Nahichevanskaya obl., Karachuk Village
Distribution: Azerbaidjan, India
Lepidocheyla gracilis Volgin, 1963: 940; 1969: 221; Lin et al., 1998; Fain & Bochkov, 2001: 55; Bochkov, Hakimitabar & Saboori, 2005: 103; Xia, 2010: 162
Type locality: Verkhnaya Terrasa (Tadjikistan), Donezk (Ukraine)
Distribution: China, Iran, Israel, Tadjikistan, Turkmenia, Ukraine
Nidocheyletus Bochkov & OConnor, 2010: 60
Type species: Nidocheyletus kennedyae Bochkov et OConnor, 2010
Nidocheyletus kennedyae Bochkov & OConnor, 2010: 60
Type locality: Michigan
Distribution: USA (Michigan)
Nodele Muma, 1964: 252
Neocheletophyes Volgin, 1965: 296
Type species: Nodele calamondin Muma, 1964
Type species of Neocheletophyes Volgin: Cheletophyes philippinensis Baker, 1949
Nodele calamondin Muma, 1964: 252; Volgin, 1969: 162; Summers & Price, 1970: 41;
Fain & Bochkov, 2001: 54; Bochkov, Hakimitabar & Saboori, 2005: 107
Nodele simplex Wafa et Soliman, 1968: 223 – Giza, Egypt – synonymized by Bochkov et al., 2001: 123
Type locality: Florida, Fort Myers
Type locality of Nodele simplex: Giza, Egypt
Distribution: Egypt, Iran, USA (Florida, California)
Nodele coccineae Thewke & Enns, 1968: 216; Summers & Price, 1970: 42; Fain & Bochkov, 2001: 54
Type locality: Missouri
Distribution: USA (Missouri)
Host: in galleries of bark beetle, Pseudopityophthorus minutissimus, in scarlet oak
Nodele mu Haines, 1988: 362; Fain & Bochkov, 2001: 54
Type locality: ? Bogor, Java
Distribution: Indonesia or Tanzania
Host: Dinoderus minutus (Coleoptera: Bostrychidae)
Cheletophyes philippinensis Baker, 1949: 288
Neocheletophyes philippinensis (Baker): Volgin, 1965: 296
Nodele philippinensis (Baker): Muma, 1964: 252; Volgin, 1969: 163; Summers & Price, 1970: 42; Diaz Patxot & Goff, 1985: 161; Fain & Bochkov, 2001: 54
Type locality: Bataan (Philippines)
Distribution: Philippines, USA (Oahu I., Hawaii Is.)
Nodele superba Kuznetzov, 1977: 923; Fain & Bochkov, 2001: 54
Type locality: Ukraine
Distribution: Ukraine
Paracheyletia Volgin, 1955: 168
Type species: Cheyletus pyriformis Banks, 1904
Cheyletus pyriformis Banks, 1904: 17; 1906: 135
Cheyletus longipalpus Ewing, 1909: 54
Cheyletia pyriformis, Garman, 1948: 1; Baker, 1949: 298
Paracheyletia assimilis Volgin, 1955: 168 – England (syn. by Summers & Price, 1970: 19)
Paracheyletia pyriformis, Volgin, 1969: 177; Summers & Price, 1970: 19; André, 1975: 31; Bochkov, Hakimitabar & Saboori, 2005: 107; Bochkov & Abramov, 2016; Khaustov, 2021: 233
Paracheyletia hortensis Volgin, 1969: 181; (acc. to Fain & Bochkov, 2001, a synonym) – Krasnodarskaiy Kray, Slavyanskiy Rayon, Russia
Paracheyletia samsinaki Volgin, 1966b: 281; 1969: 179 (acc. to Fain & Bochkov, 2001, a synonym) – Prague, Czech Rep., on Scolytus pygmaeus (Coleoptera: Scolytidae)
Type locality: Kirkwood, Mo.
Distribution: Algeria, Belgium, Czech Rep., Germany, Iran, Russia, USA (Montana, Illinois, Massachusetts, New York)
Paracheyletia recki Volgin, 1966b: 283; 1969: 183; Fain & Bochkov, 2001: 51
Type locality: Voronezh Reserve
Distribution: Russia
Host: ant hill
Paracheyletiella Kuznetzov, 1977: 926
Type species: Paracheyletiella volgini Kuznetzov, 1977
Paracheyletiella volgini Kuznetzov, 1977: 926
Type locality: Crimea
Distribution: Crimea
Host: sparrow nest
Pavlovskicheyla Volgin, 1965: 290
Type species: Cheletophyes semenovi Kuzin in Rohdendorf, 1940
Pavlovskicheyla philippicana Corpuz – Raros, 1998: 285; Fain & Bochkov, 2001 : 58
Type locality: Luzon Is., Sta Rosa, Laguna
Distribution: Peru, Philippine Islands
Pavlovskicheyla platydemae Thewke & Enns, 1975: 671
Type locality: Missouri
Distribution: USA (Missouri)
Host: Platydema sp. (Coleoptera: Tenebrionidae)
Cheletophyes semenovi Kuzin in Rohdendorf, 1940: 94; Bakeri, 1949: 290
Pavlovskicheyla semenovi: Volgin, 1965: 292; 1969: 229; Fain & Bochkov, 2001: 57
Type locality: Andizhanskaya Prov.
Distribution: Uzbekistan
Host: from old seeds of the cotton-plant
Tutacheyla Corpuz – Raros, 1972: 266
Indonesicheyla Thewke, 1980: 339
Type species: Tutacheyla robusta Corpuz-Raros, 1972
Type species of Indonesicheyla Thewke: I. buruensis Thewke, 1980
Tutacheyla robusta Corpuz-Raros, 1972: 266
Indonesicheyla buruensis Thewke, 1980: 339 – type: Indonesia (Buru)
Tutacheyla buruensis (Thewke)
Type locality: Philippines
Distribution: Indonesia (Buru), Philippines
Host: pin-hole borer (Coleoptera: Platypodidae)
Zachvatkiniola Volgin, 1969: 156
Type species: Eucheyletia reticulata Cunliffe, 1962
Eucheyletia reticulata Cunliffe, 1962: 200; Volgin, 1963: 63; Volgin, 1963: 63; 1969: 157; Summers & Price, 1970: 34; Shen, 1975; Nishida (Version 2009)
Zachvatkiniola reticulata (Cunliffe): Volgin, 1969: 157; Bochkov, Hakimitabar & Saboori, 2005: 106; Xia, 2010: 162
Type locality: Tahiti (Hawaiian Quarantine)
Distribution: China (Guandong), Georgia, Iran, Japan, French Polynesia (Tahiti)
Acaropseini [sic] Volgin, 1969: 307
Acaropsellini: Bochkov & Fain, 2003: 24
Type genus: Acaropsella Volgin, 1969
Acaropsella Volgin, 1969: 314
Type species: Neoacaropsis rohdendorfi Volgin, 1962
Acaropsella kinshasensis Fain, 1972: 39; Fain & Bochkov, 2001: 68; Xia, 2010: 160
Acaropsella konoi Tseng, 1977: 245
Acaropsella filipina Corpuz-Raros, 1988: 415
Type locality: Kinshasa (Congo)
Distribution: DR Congo, Philippines, Taiwan
Acaropsis kulagini Rohdendorf, 1940: 78; Baker, 1949: 313
Neoacaropsis kulagini: Volgin, 1962: 699
Acaropsella kulagini (Rohdendorf): Volgin, 1969: 319; Summers & Price, 1970: 63; Fain & Bochkov, 2001: 68; Bochkov, Hakimitabar & Saboori, 2005: 105
Type locality: Blagodarnoe Village, Stavropol District (Russia)
Distribution: Iran, Russia, USA (California)
Acaropsella nobilis Rasool, Chaudhri & Akbar, 1980: 33; Chaudhri & Akbar, 1985: 198; Akbar, Jahan & Mughal, 2008: 188; Gerson, Fain & Smiley, 1999: 39; Halliday, Kamran & Bashir, 2018: 80
Type locality: Pakistan
Distribution: Pakistan
Neoacaropsis rohdendorfi Volgin, 1962: 697
Acaropsella rohdendorfi (Volgin): Volgin, 1969: 315; Summers & Price, 1970: 63; Fain & Bochkov, 2001: 68
Type locality: Kishinev, under Lepidosaphes ulni (Homoptera: Coccodea)
Distribution: Moldova
Acaropsella schmidtmanni Price, 1972: 47; Fain & Bochkov, 2001: 68
Type locality: California
Distribution: USA (California)
Acaropsella shaziai Akbar, Jahan & Mughal, 2008: 188; Halliday, Kamran & Bashir, 2018: 80
Type locality: Shekhupura, Punjab
Distribution: Pakistan
Neoacaropsis volgini Gerson, 1967: 361
Acaropsis aegyptiaca Wafa & Soliman, 1968: 221 – Egypt (Giza and Tahreer), nest of birds, debris (synonymized by Fain & Bochkov, 2001: 67)
Acaropsella aegyptiaca (Wafa et Soliman): Summers & Price, 1970: 64
Acaropsella volgini (Gerson): Volgin, 1969: 317; Summers & Price, 1970: 64; Rostom, 1993: 29 (with “?”); Koç, 1998: 195; Fain & Bochkov, 2001: 68; Salarzehi, 2018:
Acaropsella walii Akbar, Jahan & Mughal, 2008: 189; Halliday, Kamran & Bashir, 2018: 80
Type locality: Pakistan
Distribution: Pakistan
Acaropsellina Summers, 1976: 191
Acaropsis Moquin -Tandon, 1863: 314
Type species: Acaropsis sollers Kuzin in Rohdendorf, 1940
Acaropsis altilis Haq & Zia, 2001b: 61; Haq, Khan, Riaz & Rasool, 2004: 182; Haq & Afzal, 2007: 2206
Acaropsellina altilis (Haq et Zia): Halliday, Kamran & Bashir, 2018: 82
Type locality: Pakistan
Distribution: Pakistan
Acaropsellina anarsia Summers, 1976: 193
Type locality: Colorado
Distribution: USA
Acaropsis clamo Qayyum et Chaudhri, 1979: 158; Haq, Khan, Riaz & Rasool, 2004: 182
Acaropsellina clamo (Qayyum & Chaudhri): Gerson, Fain & Smiley, 1999: 39; Halliday, Kamran & Bashir, 2018: 82
Type locality: Pakistan
Distribution: Pakistan
Acaropsis culpa Haq & Zia, 2001b: 64; Haq, Khan, Riaz & Rasool, 2004: 182
Acaropsellina culpa (Haq et Zia): Halliday, Kamran & Bashir, 2018: 82
Type locality: Pakistan
Distribution: Pakistan
Cheyletus doctus Berlese, 1886: 33; 1893: 74
Acaropsis docta (Berlese): Oudemans, 1905: 209; Baker, 1949: 314; Volgin, 1969: 309; Summers & Price, 1970: 62; Shen, 1975; Rizk et al., 1979; Qayyum & Chaudhri, 1979c: 2; Farooq, 2000: 41; Haq & Zia, 2001a: 60; Haq & Zia, 2001b: 68; Haq, Khan, Riaz & Rasool, 2004: 183; Palyvos & Emmanouel, 2006: 25
Acaropsellina docta (Berlese): Summers, 1976: 194; Lucza, Ripka & Saly, 1996; Gerson, Fain & Smiley, 1999: 39;.Xia, 2010: 161; Salarzehi et al., 2018; Halliday, Kamran & Bashir, 2018: 82
Type locality: Florence, Italy
Distribution: China, Egypt, Great Britain, Greece, Holland, Hungary, India, Italy, Iran, Iraq
Acaropsis hirnai Akbar, Jahan & Qazi, 2007: 92
Acaropsellina hirnai (Akbar, Jahan & Qazi): Halliday, Kamran & Bashir, 2018: 82
Type locality: Pakistan
Distribution: Pakistan
Acaropsis humilis Haq & Zia, 2001b: 72; Haq, Khan, Riaz & Rasool, 2004: 183
Acaropsellina humilis (Haq et Zia): Halliday, Kamran & Bashir, 2018: 82
Type locality: Pakistan
Distribution: Pakistan
Neoacaropsis levis Corpuz-Raros, 1972: 263
Acaropsellina levis (Corpuz-Raros): Fain & Bochkov, 2001: 68
Type locality: Laguna
Distribution: Philippines
Host: plant (Morus alba)
Acaropsella nanchangensis Xia & Zhu, 1997: 6; Xia, 2010: 160
Acaropsellina nanchangensis (Xia et Zhu): Fain & Bochkov, 2001: 68
Type locality: Nanchang
Distribution: China
Acaropsis opsis Rasool, Chaudhri & Akbar, 1980: 34
Acaropsellina opsis (Rasool, Chaudhri & Akbar): Gerson, Fain & Smiley, 1999: 40; Halliday, Kamran & Bashir, 2018: 81
Type locality: Pakistan
Distribution: Pakistan
Acaropsis orbis Qayyum & Chaudhri, 1979: 2
Acaropsellina orbis (Qayyum & Chaudhri): Gerson, Fain & Smiley, 1999: 40; Halliday, Kamran & Bashir, 2018: 81
Type locality: Pakistan
Distribution: Pakistan
Acaropsis philippinensis Corpuz-Raros & Sotto, 1977: 144
Acaropsellina philippinensis (Corpuz-Raros & Sotto): Corpuz-Raros, 1988: 415
Type locality: Tanggalian, Piddig, Ilocos Norte
Distribution: Philippine Is.
Acaropsis platessa Aheer, Akbar & Chaudhri, 1991: 336
Acaropsellina platessa (Aheer, Akbar et Chaudhri): Gerson, Fain & Smiley, 1999: 40; Halliday, Kamran & Bashir, 2018: 81
Type locality: Pakistan
Distribution: Pakistan
Acaropsis porta Qayyum & Chaudhri, 1979: 5
Acaropsellina porta (Qayyum et Chaudhri): Gerson, Fain & Smiley, 1999: 40; Halliday, Kamran & Bashir, 2018: 82
Type locality: Pakistan
Distribution: Pakistan
Acaropsis sheikhupuraensis Akbar, Jahan & Qazi, 2007: 90
Acaropsellina sheikhupuraensis (Akbar et al.): Halliday, Kamran & Bashir, 2018: 82
Type locality: Pakistan
Distribution: Pakistan
Acaropsis sigilla Haq, Khan, Riaz & Rasool, 2004: 184
Acaropsellina sigilla (Haq, Khan, Riaz & Rasool): Halliday, Kamran & Bashir, 2018: 82
Type locality: Pakistan
Distribution: Pakistan
Acaropsis shorkotiensis Akbar, Rahi & Chaudhri, 1988: 2; Haq, Khan, Riaz & Rasool, 2004: 184
Acaropsellina shorkotiensis (Akbar et al.): Gerson, Fain & Smiley, 1999: 40; Halliday, Kamran & Bashir, 2018: 82
Type locality: Pakistan
Distribution: Pakistan
Acaropsis sodalis Haq & Zia, 2001b: 70
Acaropsellina sodalis (Haq & Zia): Halliday, Kamran & Bashir, 2018: 82
Type locality: Pakistan
Distribution: Pakistan
Acaropsis callida Kuzin in Rohdendorf, 1940: 78 – Rostov na Don, Russia, in grain
Acaropsis sollers: Kuzin in Rohdendorf, 1940: 78; Baker, 1949: 314; Volgin, 1969: 311; Summers & Price, 1970: 61; Rizk et al., 1979; Efremova & Bochkov, 1990: 102; Rostom, 1993: 29; Athanassiou et al., 2003: 413
Acaropsellina sollers (Kuzin in Rohdendorf): Bochkov, Hakimitabar & Saboori, 2005: 105; Xia, 2010: 161
Type locality: Leningrad, Russia
Distribution: Belarus, Greece, India, Iran, Iraq, Kazakhstan, Russia, Saudi Arabia, Turkmenistan, Uzbekistan
Acaropsis subitus Akbar, Mushtaq & Akbar, 1999: 264
Acaropsellina subitus (Akbar, Mushtaq & Akbar): Halliday, Kamran & Bashir, 2018: 82
Type locality: Pakistan
Distribution: Pakistan
Acaropsis summersi Smiley & Whitaker, 1981: 121
Acaropsellina summersi (Smiley et Whitaker): Whitaker & Maser, 1985: 69; Gerson, Fain & Smiley, 1999: 40
Type locality: Oregon
Distribution: USA
Host: Glaucomys sabrinus (nest)(Rodentia: Sciuridae, Sciurinae)
Acaropsis tyrophagus Elbadry, 1969: 162
Acaropsellina tyrophagus (Elbadry): Gerson, Fain & Smiley, 1999: 40
Type locality: Egypt
Distribution: Egypt
Acaropsis unicus Haq, Khan, Riaz & Rasool, 2004: 186
Acaropsellina unictus (Haq, Khan, Riaz et Rasool): Halliday, Kamran & Bashir, 2018: 82
Type locality: Muzaffarabad, N. Kashmir
Distribution: Pakistan
Acaropsis venutus Akbar, Mushtaq & Akbar, 1999: 163
Acaropsellina venutus (Akbar, Mushtaq et Akbar): Halliday, Kamran & Bashir, 2018: 82
Type locality: Pakistan
Distribution: Pakistan
Acaropsis vitrus Aheer, Akbar & Chaudhri 1991: 336; Akbar, Mushtaq & Akbar, 1999: 263; Farooq, 2000: 41; Haq & Zia, 2001a: 60; Haq, Khan, Riaz & Rasool, 2004: 182; Haq & Afzal, 2007: 2206.
Acaropsellina vitrus (Aheer, Akbar et Chaudhri): Gerson, Fain & Smiley, 1999: 40; Halliday, Kamran & Bashir, 2018: 82
Type locality: Pakistan
Distribution: Pakistan
Atarsacheylus Thewke, 1980: 333
Type species: Atarsacheylus vichii Thewke, 1980
Atarsacheylus vichii Thewke, 1980: 333
Type locality: one mile north of Vichy, Maries Co., Missouri
Distribution: USA (Missouri)
Chelacheles Baker, 1958: 235
Type species: Chelacheles strabismus Baker, 1958
Chelachecaropsis Attiah, 1973: 349
Type species: Chelachecaropsis bakeri Attiah, 1973
Chelacheles aigamuxa Bochkov & OConnor, 2004: 559
Type locality: South Africa
Distribution: South Africa
Chelacheles alexandrinus Hassan & Gomaa 1981: 115; Bochkov et al., 1999: 118; Fain & Bochkov, 2001: 82; Bochkov & OConnor, 2004: 556
Type locality: Egypt, Barg El-Arab, Northern Coast
Distribution: Crimea, Egypt
Host: litter
Chelacheles apanaskevichi Bochkov & Abramov, 2016: 337; Khaustov, 2021: 234
Type locality: Tula Province, near Suvorov town
Distribution: Russia
Chelacheles bacchusi Bochkov, Haustov & Kuznetzov, 1999: 117; Gerson et al., 1999: 7; Eliopoulos & Papadoulis, 2001: 31; Bochkov & OConnor, 2004: 556
Type locality: Sudak
Distribution: Crimea
Chelacheles baiwanganae Corpuz-Raros & Sotto, 1977: 152; Fain & Bochkov, 2001c: 82; Bochkov & Fain, 2001: 82; Bochkov & OConnor, 2004: 551
Type locality: Los Baños, Laguna
Distribution: Philippine Is., Thailand
Host: beetle (Coleoptera: Scolytidae); in Thailand from Sinoxylon crassum (Coleoptera: Bostrichidae)
Chelachecaropsis bakeri Attiah, 1973: 349
Chelacheles bakeri (Attiah): Fain & Bochkov, 2001b: 299; 2001c: 83
Chelacheles humilis Rasool, Chaudhri & Akbar, 1980: 30; Fain & Bochkov, 2001c: 83
Type locality: Cairo
Distribution: Egypt
Host: rice plantation
Chelacheles bipanus Summers & Price, 1970: 79; Shen, 1991; Fain & Bochkov, 2001c: 83; Xia, 2010: 161
Type locality: American Canyon, Solano Co., California
Distribution: China, USA (California)
Chelacheles humilis Rasool, Chaudhri & Akbar, 1980: 30; Chaudhri & Akbar, 1985: 194; Gerson, Fain & Smiley, 1999: 47; Halliday, Kamran & Bashir, 2018: 83
Type locality: Pakistan
Distribution: Pakistan
Chelacheles hellenicus Eliopoulos & Papadoulis, 2001: 31; Fain & Bochkov, 2001c: 83; Bochkov & OConnor, 2004: 573
Type locality: Attiki Co., Athens, Votanikos
Distribution: Greece
Chelacheles indra Bochkov & Otto, 2010: 16
Type locality: India (intercepted on Australian border)
Distribution: India
Host: from Sinoxylon crassum (Coleoptera: Bostrichidae)
Chelacheles klimovi Bochkov & OConnor, 2004: 559
Type locality: NY, Nassau Co., Tobay Beach
Distribution: USA
Host: wrack from beach
Chelacheles lanceolatus Tseng, 1977: 259; Lin & Zhang, 1997: 197; Corpuz-Raros, 1998: 263; Gerson et al., 1999: 47; Fain & Bochkov, 2001: 82; Bochkov & OConnor, 2004: 578; Xia, 2010: 161
Type locality: Taiwan
Distribution: Taiwan
Host: Chicken feathers (accidental?)
Chelacheles michalskii Samšinák, 1962: 183; Volgin, 1969: 305; Liu et al., 1997; Fain & Bochkov, 2001c: 83; Bochkov, Hakimitabar & Saboori, 2005: 105; Xia, 2010: 161; Torre Santana, 2015
Type locality: Prague
Distribution: China, Cuba, Czech Rep., Iran
Host: «In den Gängen von Scolytus multistriatus und S. pygmaeus»
Chelacheles peritremaculatus Thewke, 1974: 33; Hassan & Gomaa, 1981: 115; Corpuz-Raros, 1998: 263; Bochkov et al., 1999: 117; Gerson et al., 1999: 47; Fain & Bochkov, 2001c: 82; Eliopoulos & Papadoulis, 2001: 31; Bochkov & OConnor, 2004: 565
Type locality: Seven miles of West Plains, Howell Co., Missouri, on oak branches
Distribution: USA (Missouri)
Chelacheles robustus Corpuz-Raros, 1998: 263; Eliopoulos & Papadoulis, 2001: 31; Fain & Bochkov, 2001c: 82; Bochkov & OConnor, 2004: 570
Type locality: Leyte I., Baybay, Visayas St., Col. of Agriculture Campus
Distribution: Philippine Is. (Leyte)
Host: rice mill
Chelacheles seminole Bochkov & OConnor, 2004: 570
Type locality: Florida, Lee Co., Punta Rassa
Distribution: USA (Florida)
Host: fungi Daldinia concentrica (Xylariales: Xylariaceae)
Chelachecaropsis stigmaeoides Barilo, 1989: 137; Gerson et al., 1999: 47
Chelacheles stigmaeoides (Barilo): Fain & Bochkov, 2001c: 83; Bochkov & OConnor, 2004: 575
Type locality: Samarkand area
Distribution: Uzbekistan
Host: soil from ant hill (Formica sp.)
Chelacheles strabismus Baker, 1958: 234; Patxot & Goff, 1985: 160; Fain & Bochkov, 2001: 83
Chelacheles algericus Fain & Bochkov, 2001: 83; syn. by Bochkov & OConnor, 2004 – terra typica: Algeria
Type locality: Lisbon (Portugal)
Distribution: Algeria, Portugal, USA (Hawaiian Is. – Oahu)
Chelacheles striola Lin in Lin & Zhang, 1997: 245; Bochkov & OConnor, 2004: 567; Xia, 2010: 161
Type locality: Fujian
Distribution: China
Chelacheles temoak Bochkov & OConnor, 2004: 569
Type locality: Nevada, Washoe Co., Patrick
Distribution: USA (Nevada)
Host: Diadasia sphaeralcearum (Hymenoptera: Apidae)
Chelacheles thomasi Bochkov & Otto, 2010: 12
Type locality: Sydney/Rosebury
Distribution: Australia
Host: from oranges
Neoacaropsis Volgin, 1962: 685
Type species: Neoacaropsis granulatus Volgin, 1962
Neoacaropsis granulatus Volgin, 1962: 685; 1969: 320; Xie et al., 2000; Fain & Bochkov, 2001: 68; Xia, 2010: 161
Type locality: Sochi (Krasnodarskiy Kray)
Distribution: China, Russia
Neochelacheles Smiley & Williams, 1972: 312
Type species: Neochelacheles messersmithi Smiley et Williams, 1972
Neochelacheles corpuzrarosae Bochkov & OConnor, 2004: 582
Type locality: Negros Oriental, Lake Balinsasayao, 3 km N 14 km W Dumaguete City
Distribution: Philippines
Host: from fungi of the superfamily Polyporaceae (Agarycales)
Neochelacheles mendicus Bochkov & OConnor, 2004: 587
Type locality: Luzon Island, Baltalason
Distribution: Philippines
Host: Bysrax tuberculatus (Coleoptera: Tenebrionidae)
Neochelacheles messersmithi Smiley & Williams, 1972: 312; Fain & Bochkov, 2001c: 81
Type locality: College Park, Meryland
Distribution: USA (Meryland, Ohio)
Host: Bolitotherus cornutus (Coleoptera: Tenebrionidae, Bolitophaginae)
Paracaropsis Volgin, 1969: 322
Type species: Acaropsis travisi Baker, 1949
Acaropsis travisi Baker, 1949: 322
Paracaropsis travisi (Baker): Volgin, 1969: 324; Summers & Price, 1970: 39
Paracaropsis strofi Samšinák, 1956 (syn. by Summers & Price, 1970: 39)
Type locality: Newton, Georgia
Distribution: USA (Georgia)
Host: Sceloporus woodi (Sauria: Iguanidae)
Bakini Volgin, 1969: 299
Type species: Bak sanctaehelenae Yunker, 1961
Aegyptocheyla Yousef, 1979: 366
Type species: Aegyptocheyla summersi Yousef, 1979
Aegyptocheyla summersi Yousef, 1979: 366
Type locality: Kalyobia and Giza Governorates
Distribution: Egypt
Alliea Yunker, 1960: 278
Type species: Alliea laruei Yunker, 1960
Alliea laruei Yunker, 1960: 278; Volgin, 1969: 233
Type locality: Tampa (Florida)
Distribution: USA (Florida)
Host: Rattus norvegicus (Rodentia: Muridae)
Alliea prasadi Corpuz-Raros, 1998: 261
Type locality: Luzon Is., UPLB Campus, College, Laguna
Distribution: Philippines
Bak Yunker, 1961:1023
Type species: Bak sanctaehelenae Yunker, 1961
Bak deleoni Yunker, 1961: 1028; Volgin, 1969: 302; Summers & Price, 1970: 80; Patxot & Goff, 1985: 159; Fain & Bochkov, 2001: 80
Type locality: Coral Gables (Florida)
Distribution: D.R. Congo, USA (Florida, Hawaiian Is.)
Bak elongatus Patxot & Goff, 1985: 158; Fain & Bochkov, 2001: 80
Type locality: Oahu I., Honolulu, Waimanalo
Distribution: USA (Hawaiian Is.)
Bak faini Corpuz-Raros, 2000: 323; Fain & Bochkov, 2001: 80; Xia, 2010: 161
Type locality: Philippines
Distribution: China, Philippines
Bak furcatus Gerson & Fain, 1991: 17; Corpuz Raros, 2000; Fain & Bochkov, 2001c: 80
Type locality: Chantaburi (Thailand)
Distribution: Philippines, Thailand
Host: in debris from a hive of Apis cerana (Hymenoptera: Apidae)
Bak gersoni Corpuz-Raros, 2000: 326; Fain & Bochkov, 2001c: 80
Type locality: Philippines
Distribution: Galapagos Is., Philippines
Bak indonesiensis Bochkov & Otto, 2010: 17
Type locality: Indonesia (intercepted on Australian border)
Distribution: Indonesia
Host: beetles (Coleoptera indet.)
Bak iranica Pactinat-Saeij, Bagheri & Skvarla, 2017: 848; Salarzehi, 2018
Type locality: Amol, Mazandaran Prov.
Distribution: Iran
Bak ligyscutatus Flechtmann, 1971: 48 (according to Fain & Bochkov, 2001c: 78, invalid species, published only in a thesis, but actually published regularly)
Type locality: Piracicaba, São Paulo
Distribution: Brazil
Host: decomposing organic material
Bak micidus Summers & Price, 1970: 81; Fain & Bochkov, 2001c: 79; Xia, 2010: 161
Type locality: Santa Cruz, Old Bella Vista Trail
Distribution: China, Galapagos Is. (Ecuador)
Bak nadchatrami Fain et Bochkov, 2001c: 80
Type locality: Belakang Pasar, Lama Kepong, Kuala Lumpur
Distribution: Malaysia
Bak ozarkensis Thewke & Enns, 1974: 43; Fain & Bochkov, 2001c: 79
Type locality: four miles west of Macks Creek, Camden Co., Missouri
Distribution: USA (Missouri, Pennsylvania)
Bak payatus Corpuz-Raros & Sotto, 1977: 146; Fain & Bochkov, 2001c: 79
Type locality: Laguna
Distribution: Philippine Is.
Bak sanctaehelenae Yunker, 1961: 1023; Volgin, 1969: 300; Summers & Price, 1970: 80; Fain & Bochkov, 2001c: 79
Type locality: Napa Co., Mt. St. Helena, California
Distribution: USA (California)
Bak truncatus Corpuz-Raros & Sotto, 1977: 148; Fain & Bochkov, 2001c: 79
Type locality: Makiling Bot, Gardens, Mt. Makiling, Laguna
Distribution: Philippine Is.
Caudacheles Gerson, 1968: 645
Type species: Caudacheles khayae Gerson, 1968
Caudacheles khayae Gerson, 1968: 645; Fain & Bochkov, 2001: 77; Salarzehi, 2018:
Type locality: Rehovot (Israel)
Distribution: Iran, Israel
Caudacheles lieni Tseng, 1977; Fain & Bochkov, 2001: 77; Bochkov, Hakimitabar & Saboori, 2005: 104; Xia, 2010: 162
Type locality: Taiwan
Distribution: Iran, Taiwan
Caudacheles trigintaduae Lin & Zhang, 1997: 136
Type locality: China
Distribution: China
Chelacaropsis Baker, 1949: 315
Type species: Chelacaropsis moorei Baker, 1949
Cheletonata Womersley, 1955 = Cheletacaropsis Baker, 1949 (syn. by Fain & Bochkov, 2001a: 294)
Chelacaropsis apus Fain, 1972: 40
Type locality: Butare
Distribution: Rwanda
Host: Apus affinis, “dejections”(Apodiformes: Apodidae)
Cheletacaropsis kenyensis Fain & Bochkov, 2001b: 70
Type locality: Kimakia Cave, Hunter’s Lodge, Kiboko, 140 km SE Nairobi
Distribution: Kenya
Cheletonata milesi Womersley, 1955: 214
Cheletacaropsis milesi (Womersley): Fain & Bochkov, 2001b: 73
Type locality: Northern Australia
Distribution: Australia
Chelacaropsis moorei Baker, 1949: 315; Volgin, 1969: 326
Type locality: Florida
Distribution: China, USA (Florida)
Host: Glaucomys volans querceti (Rodentia: Pteromyidae)
Chelacaropsis reticulata Soliman, 1975: 97
Type locality: Farskour, Domiatta, North of Deltha
Distribution: Egypt
Chelacaropsis rwandana Fain, 1972: 40
Type locality: Butare
Distribution: Rwanda
Host: Spermestes cucullatus (Passeriformes: Estrildidae)
Chelacaropsis terrestris Corpuz-Raros & Sotto, 1977: 150
Type locality: Los Baños, Laguna
Distribution: Philippine Is.
Cheletogenes Oudemans, 1905: 208
Type species: Cheyletus ornatus Canestrini et Fanzago, 1876
Cheletogenes aceriai Khan, 1970: 278; Fain & Bochkov, 2001: 85; Gerson, Fain & Smiley, 1999: 47; Halliday, Kamran & Bashir, 2018: 82
Type locality: Pakistan
Distribution: Pakistan
Host: mango tree bugs
Cheletogenes carinatus Aheer, Akbar & Chaudhri, 1992: 39; Fain & Bochkov, 2001: 85 (Species dubia); Gerson, Fain & Smiley, 1999: 47; Aheer, Akbar & Chaudhri, 1992: 39; Halliday, Kamran & Bashir, 2018: 83
Type locality: Pirmahal
Distribution: Pakistan
Host: on Vitis vinifera (plant)
Cheletogenes dissitus Akbar, Rahi & Chaudhri, 1988: 4; Gerson, Fain & Smiley, 1999: 47; Fain & Bochkov, 2001: 85; Halliday, Kamran & Bashir, 2018: 83
Type locality: Pakistan
Distribution: Pakistan
Cheletogenes iconis Chaudhri & Akbar, 1985: 128; Fain & Bochkov, 2001: 85; Aheer, Akbar & Chaudhri, 1992: 36; Halliday, Kamran & Bashir, 2018: 83
Type locality: 3 km S Multan
Distribution: Pakistan
Host: on Gossypium hirsutum
Cheletogenes meihuashanense Lin & Liu, 1994: 220 (223); Lin et al., 2000: 86; Xia, 2010: 161
Type locality: Meihuashan, Fujian
Distribution: China (Fujian)
Cheletogenes monosetosus Tseng, 1977: 229; Ehara & Abdul Ghani, 1988: 239; Fain & Bochkov, 2001: 85; Xia, 2010: 161
Type locality: Taiwan
Distribution: Malaysia, Taiwan
Host: from trees (Gigantochloa rostrata, Bambusa glaucescens)
Cheyletus ornatus Canestrini & Fanzago, 1876: 106; 1877: 147; Canestrini, 1886: 173; Starkoff & Starkoff, 1950: 96
Cheyletus saccardianus Berlese, 1886: 32; Oudemans, 1906: 153
Cheyletia ornata, Oudemans, 1904: 154
Cheyletus cocciphilus Banks, 1914: 56. Syn. by Baker, 1949: 305
Cheletogenes ornatus (Canestrini et Fanzago): Oudemans, 1905: 208; 1906: 153; Baker, 1949: 305; Cooreman, 1951: 33; Volgin, 1955: 174; 1969: 296; Gerson, 1967: 359; Wafa & Soliman, 1968: 222; Summers & Price, 1970: 53; Flechtmann, 1971: 30; Shiba, 1976: 173; Nachev & Trenchev, 1987; Yousef, Zaher & Kandil, 1979: 228; Ripka, Fain & Bolland, 1999: 366; Fain & Bochkov, 2001: 85; Buosi et al., 2006: 7
Cheyletogenes ornatus (Canestrini et Fanzago): Namaghi, 2010: 135
Type locality: Italy
Distribution: Algeria, Armenia, Australia, Brazil, Chile, China, Cuba, Egypt, Georgia, Hungary, India, Iran, Italy, Israel, Kirghizstan, Malaysia, Mexico, Morocco, Philippines, Porto Rico, Russia, South Africa, USA (Florida, Louisiana, California, Hawaiian Is.), Venezuela
Cheletogenes petiginis Qayyum & Chaudhri, 1977; Fain & Bochkov, 2001: 85; Rasool & Chaudhri, 1979a: 7; Chaudhri & Akbar, 1985: 134; Akbar, Rahi & Chaudhri, 1988: 4; Gerson, Fain & Smiley, 1999: 47; Halliday, Kamran & Bashir, 2018: 83
Type locality: Pakistan
Distribution: Pakistan
Cheletogenes sagacis Chaudhri & Akbar, 1985; Aheer, Akbar & Chaudhri, 1992: 36; Halliday, Kamran & Bashir, 2018: 83
Type locality: 2 km N Haripur
Distribution: Pakistan
Host: on plants
Cheletogenes scaber Qayyum & Chaudhri, 1977: 111; Fain & Bochkov, 2001: 85; Salarzehi, 2018; Halliday, Kamran & Bashir, 2018: 83
Type locality: Pakistan
Distribution: Iran, Pakistan
Cheletogenes vulgatus Rasool & Chaudhri, 1979: 8; Fain & Bochkov, 2001: 85; Chaudhri & Akbar, 1985: 129; Akbar, Rahi & Chaudhri, 1988: 4; Gerson, Fain & Smiley, 1999: 47; Halliday, Kamran & Bashir, 2018: 83
Type locality: Pakistan
Distribution: Pakistan
Cheletogenes waitei Gerson, 1994: 441; Fain & Bochkov, 2001: 85
Type locality: Nambour (Queensland)
Distribution: Australia (Queensland)
Cheletoides Oudemans, 1904: 154
Type species: Syringophilus uncinatus Heller, 1880
Cheletoides chirunduensis Fain, 1979: 1023
Type locality: Chirundu
Distribution: Zimbabwe
Host: Numida meleagris (Galliformes: Numididae)
Syringophilus uncinatus Heller, 1880: 188
Cheyletus uncinatus (Heller): Poppe, 1888: 239
Cheletoides uncinatus (Heller): Oudemans, 1904: 154; 1906: 204; Volgin, 1969: 336
Cheletoides uncinata (Heller): Oudemans, 1906: 204; Baker, 1949: 275; Summers & Price, 1970: 74
Type locality: Kiel
Distribution: Germany
Host: Pavo cristatus (Galliformes: Phasianidae)
Cheletomorpha Oudemans, 1904: 162
Acheletomorpha Volgin, 1969: 195
Type species: Acarus lepidopterorum Shaw, 1794
Cheletomorpha bakeri Lawrence 1954: 67; Summers & Price, 1970: 51; Fain & Bochkov, 2001: 52
Acheletomorpha bakeri (Lawrence): Volgin, 1969: 196
Type locality: Karamoja (Uganda)
Distribution: D.R. Congo, Rwanda, Uganda
Host: in nests of birds and rodents
Cheletomorpha dolosus Aheer et al., 1997: 11; Fain & Bochkov, 2001: 52; Halliday, Kamran & Bashir, 2018: 86
Type locality: Faisalabad
Distribution: Pakistan
Acarus lepidopterorum Shaw, 1794: 187
Cheyletus venustissimus C.L. Koch, 1839: fasc. 23; Canestrini, 1886: 172; Hull, 1918: 32; Starkoff & Starkoff, 1950: 96
Cheyletus seminivorus Packard, 1878: 665
Cheletomorpha lepidopterorum (Shaw): Oudemans, 1929: 298; Baker, 1949: 302; Volgin, 1969: 190; Summers & Price, 1970: 49; Shen, 1975; Rekk, 1976: 68; Caceres & Fain, 1978: 302; Enghoff, 1988: 49; Peredo et al., 1994: 17; Lucza, Ripka & Saly, 1996; Fain & Bochkov, 2001: 52; Petrova et al., 2004: 91; Bochkov, Hakimitabar & Saboori, 2005: 105; Xia, 2010: 161; Bochkov & Abramov, 2016; Halliday, Kamran & Bashir, 2018: 86
Cheyletus rufus Hardy, in André, 1933: 352
Cheletomorpha venustissima, Oudemans, 1904: 162; 1906: 144; Thor, 1912: 390; Zachvatkin, 1935: 25; Rohdendorf, 1940: 92; Womersley, 1941: 62; Starkoff & Starkoff, 1950: 96;
Cheletomorpha tatami Hara, 1955: 69; Gerson, 1967: 360; Shiba, 1969a: 164
Cheletophyes knowltoni Beer & Dailey, 1956: 409
Cheyletus longipes Banks, 1909: 133 – Canada, Ontario, Guelph
Type locality: England
Distribution: Armenia, Australia, Azerbaidzan, Belgium, Brazil, Canada, Chile, China, Cuba, Egypt, Denmark, England, France, Georgia, Germany, Greece, Holland, Hungary, India, Indonesia, Iran, Israel, Italy, Japan, Latvia, Malaysia, Malta, Mexico, New Guinea, Norway, Pakistan, Peru, the Philippines, Portugal, Rwanda, New Guinea, Russia, South Africa, Spain, Switzerland, Taiwan, Turkey, USA
Host: Proxenus sp. (Lepidoptera: Noctuidae) and other moths
Cheletomorpha obrutus Qayyum & Chaudhri, 1977: 75; Halliday, Kamran & Bashir, 2018: 86
Type locality: Pakistan
Distribution: Pakistan
Cheletomorpha opacus Qayyum & Chaudhri, 1977: 73; Halliday, Kamran & Bashir, 2018: 86
Type locality: Pakistan
Distribution: Pakistan
Cheletomorpha orientalis Oudemans 1928: 343; Baker, 1949: 303; Volgin, 1969: 195; Summers & Price, 1970: 51
Type locality: Java
Distribution: Indonesia (Java)
Host: on leaves of orchid, Palaenopsis sp.
Cheletomorpha tenerum Qayyum & Chaudhri, 1977: 71; Fain & Bochkov, 2001: 52; Halliday, Kamran & Bashir, 2018: 86
Type locality: Pakistan
Distribution: Pakistan
Cheletopsis Oudemans, 1904: 163
Type species: Cheyletus nörneri Poppe, 1888
Cheletopsis anax Oudemans, 1904b: 170; 1906: 195; Baker, 1949: 309; Volgin, 1969: 354; Summers & Price, 1970: 73; Bochkov et al., 2002: 10
Type locality: France
Distribution: France
Host: “Crocethia alba (= Totanus calidris)”, ? Tringa totanus (Charadriiformes: Scolopacidae)
Cheletopsis animosa Oudemans, 1904b: 170; 1906: 189; Baker, 1949: 310; Volgin, 1969: 352; Summers & Price, 1970: 73; Bochkov et al., 2002: 10
Type locality: France
Distribution: France, Eastern Nord America
Hosts: ?Tringa totanus (Charadriiformes: Scolopacidae), Sterna hirundo (Charadriiformes: Laridae, Sterninae)
Cheletopsis basilica Oudemans, 1904: 170; 1906: 186; Baker, 1949: 310; Volgin, 1969: 351; Summers & Price, 1970: 73; Bochkov et al., 2002: 8
Type locality: France
Distribution: France, Poland; USA (Alaska, Michigan)
Hosts: Calidris melanotus, Tringa totanus (Charadriiformes: Scolopacidae), Charadrius hiaticula (Charadriiformes: Charadriidae), Chlidonius niger (Charadriiformes: Laridae)
Cheletopsis charadrii Mironov, Bochkov & Chirov, 1991: 52; Bochkov et al., 2002: 18; Chernichko & Kivganov, 2013: 84
Type locality: Ottuk Village, near Issyk-Kul Lake (Kirghizstan)
Distribution: Kirghizstan, Ukraine
Hosts: Charadrius alexandrinus, Ch. dubius (Charadriiformes: Charadriidae)
Cheletopsis daberti Kivganov & Bochkov, 1994: 39 (also “deberti”, laps. typ.); Bochkov et al., 2002: 15; Chernichko & Kivganov, 2013: 84
Type locality: Tigulskij Res., Odessa Prov., Ukraine
Distribution: Poland, Ukraine; USA (N. Dakota)
Hosts: Calidris temminckii, Tringa glareola, T. nebularia, Micropalama himantopus (Charadriiformes: Scolopacidae)
Cheletopsis ferrugineae Chernichko & Kivganov, 2013: 85
Type locality: Tilihul Lagoon
Distribution: Ukraine
Host: Calidris ferruginea (Charadriiformes: Charadriidae)
Cheletopsis impavida Oudemans, 1904b: 170; 1906: 175; Baker, 1949: 310; Volgin, 1969: 342; Summers & Price, 1970: 73; Mironov, Bochkov & Chirov, 1991: 55; Bochkov et al., 2002: 12; Chernichko & Kivganos, 2013: 83
Type locality: France
Distribution: France, Poland; Russia (Primorie, Crimea), Kazakhstan, Kirghizstan; USA (Kansas, Michigan); Argentina
Hosts: Tringa totanus, T. stagnatilis (Charadriiformes: Scolopacidae), Calidris minutus, C. pusilla, C. ferruginea, C. temminckii, C. ruficollis, Micropalama himantopus (Charadriiformes: Scolopacidae), Rostratula semicollaris (Charadriiformes: Rostratulidae)
Cheletopsis limnodromi Bochkov, Fain & Dabert, 2002 : 18
Type locality: Sapelo Isl., McIntosh Co., Georgia, USA
Distribution: USA (Georgia)
Host: Limnodromus griseus (Charadriiformes: Scolopacidae)
Cheletopsis magnanima Oudemans, 1904: 170; 1906: 193; Volgin, 1969: 350; Summers & Price, 1970: 73; Mironov et al., 1991: 55; Bochkov et al., 2002: 22
Type locality: Chile (in museum in Angers, France)
Distribution: Chile
Host: Tringa flavipes (Charadriiformes: Scolopacidae, museum bird from Chile)
Cheletopsis mariae Mironov, Bochkov & Chirov, 1991: 51; Bochkov et al., 2002: 12
Type locality: Ottuk Village, Issyk-Kul Lake, Kirghizstan
Distribution: Poland; Kirghizstan
Host: Actitis hypoleucos (Charadriiformes: Scolopacidae)
Cheyletus Nörneri Poppe, 1888: 239
Cheletopsis Nörneri (Poppe): Oudemans, 1906: 180
Cheletes noerneri Poppe: Poppe, 1909: 51
Cheletopsis norneri (Poppe): Baker, 1949: 310; Volgin, 1969: 345; Mironov et al., 1991: 55; Bochkov et al., 2002: 8
Type locality: France
Distribution: Bulgaria, France, Germany, Poland, Russia, Ukraine; Kazakhstan, Kirghizstan; Egypt; USA; Paraguay
Host: Tringa totanus (Charadriiformes: Scolopacidae), Sterna hirundo, S. repressa, S. simplex, Gelochelidon nilotica, Chlidonias h. hybrida (Charadriiformes: Laridae)
Cheletopsis prosobonialis Bochkov, Fain & Dabert, 2002: 22
Type locality: Tepoto Isl., Tuamotu Archipelago
Distribution: French Polynesia (Tuamotu)
Host: Prosobonia cancellata (Charadriiformes: Scolopacidae)
Cheletopsis rynchops Bochkov, Fain & Dabert, 2002 : 18
Type locality: South Tica-Buzi River, Distr. Vila Machado, Mozambique
Distribution: Mozambique; USA (N. Carolina)
Hosts: Rynchops flavirostris, R. niger (Charadriiformes: Laridae)
Cheletopsis thalasseus Bochkov & OConnor, 2003: 270
Type locality: Florida, Monroe Co., Dry Tortugas, Garden Key
Distribution: USA (Florida)
Host: Thalasseus [Sterna] sandviciensis (Charadriiformes: Laridae, Sterninae)
Cheletosoma Oudemans, 1905: 207
Type species: Cheletosoma tyrannus Oudemans, 1905
Cheletosoma tyrannus Oudemans, 1905: 207; 1906: 168; Baker, 1949: 315; Volgin, 1969: 332
Type locality: «Tropical America»
Distribution: «Tropical America»
Host: Aramus scolopaceus (Gruiformes: Aramidae)
Cheyletia Haller, 1884: 233
Type species: Acarus squamosus De Geer, 1776
Cheyletia squamosa Baker, 1949: 301
Cheyletia americana Volgin, 1969: 240
Type locality: Birnham wood (Wisconsin)
Distribution: USA
Cheyletia aradiphila Volgin, 1966b: 279; 1969: 238; Summers & Price, 1970: 65
Type locality: Regions of Murmansk, Leningrad, Vologda (Totma) and Irkutsk
Distribution: Russia
Host: Aradus lugubris (Heteroptera: Aradidae)
Cheyletia laureata Haller, 1884: 234; Karpelles, 1893: 124; Volgin, 1969: 237; Summers & Price, 1970: 65 (syn. of Cheyletia squamosa)
Type locality: Germany
Distribution: Germany, Hungary
Cheyletus squamosus Oudemans, 1897: 128 (not Acarus squamosus De Geer)
Cheyletia papillifera Volgin, 1955: 171; 1969: 240; Žďárková, 1979; Summers & Price, 1970: 65; Solarz, 1989: 243
Type locality: Utrecht, Arnhem
Distribution: Holland, “Czechoslovakia”, Poland
Host: Chloris chloris (Passeriformes: Fringillidae)
Acarus squamosus De Geer, 1778: 116
Cheyletia squamosa (De Geer): Summers & Price, 1970: 65
Cheyletia laureata Haller, 1884: 223
Type locality: Austria
Distribution: Austria
Chiapacheylus De Leon, 1962: 135
Type species: Chiapacheylus edentatus De Leon, 1962
Chiapacheylus edentata De Leon, 1962: 135; Torre Santana, 2015: 49
Chiapacheylus edentatus De Leon: Volgin, 1969: 278; Summers & Price, 1970: 52
Type locality: Tuxtla Gutierrez (Chiapas)
Distribution: Cuba, Mexico (Chiapas)
Hosts: Xylosoma elliptica, Citharexylum fructicosum
Chiapacheylus desertorum Zaher & Soliman, 1967: 22
Type locality: El-Fayoum and Bury El-Arab
Distribution: Egypt
Chiapacheylus macrocorneus Zaher & Soliman, 1967: 23
Type locality: Giza and Tahrcer Proc.
Distribution: Egypt
Columbicheyla Thewke & Enns, 1972: 450
Type species: Columbicheyla macroflabellata Thewke et Enns, 1972
Columbicheyla bicicri Lin & Zhang, 1997: 133; Fain & Bochkov, 2001: 76; Xia, 2010: 161
Type locality: Fudjian
Distribution: China (Fudjian)
Columbicheyla macroflabellata Thewke & Enns, 1972: 450; Lin & Zhang, 1997; Fain & Bochkov, 2001: 76; Xia, 2010: 161
Type locality: Missouri
Distribution: China, Malaysia, Philippine Is., USA (Missouri)
Eucheyletia nindota Corpuz-Raros, 1988b: 417
Columbicheyla nindota (Corpuz-Raros): Corpuz-Raros, 1998: 266
Type locality: Leyte Is.
Distribution: Philippines
Type species: Neoeucheyla (Bothrocheyla) pavlovskyi Volgin, 1964
Cunliffella Volgin, 1969: 255 (as subgenus)
Bothrocheyla Volgin 1964: 93
Type species: Neoeucheyla tuberculicoxa Volgin, 1964
Cheyletia bulgarica Volgin, 1955: 171
Neoeucheyla bulgarica (Volgin): Volgin, 1964: 89; 1969: 252; Summers & Price, 1970: 69; Corpus-Raros & Sotto, 1977: 161
Cunliffella bulgarica (Volgin): Fain & Bochkov, 2001 : 72; Bochkov, Hakimitabar & Saboori, 2005: 105
Type locality: Pazardzhik Distr. (Bulgaria)
Distribution: Bulgaria, China, Iran, Philippine Is.
Neoeucheyla dua Corpuz-Raros, 1998: 282
Cunliffella dua (Corpuz-Raros): Fain & Bochkov, 2001 : 72
Type locality: Mt. Makiling, Makiling Bot. Gardens, UPLB Forestry Campus, College, Laguna
Distribution: Philippines
Neoeucheyla maysa Corpuz-Raros, 1998: 284
Cunliffella maysa (Corpuz-Raros): Fain & Bochkov, 2001 : 72
Type locality: Luzon Is., Putting Lupa, Calamba, Laguna
Distribution: Philippines
Neoeucheyla mumai Volgin, 1969: 254; Summers & Price, 1970: 69
Cunliffella mumai (Volgin): Fain & Bochkov, 2001 : 72
Type locality: Sebastian (Florida)
Distribution: U. S. A. (Florida)
Host: from Pinus clausa
Neoeucheyla ornata Wafa & Soliman, 1968: 228; Summers & Price, 1970: 69
Cunliffella ornata (Wafa et Soliman): Fain & Bochkov, 2001 : 72
Type locality: Giza
Distribution: Egypt
Eucheyla panamensis Baker, 1949: 291; Patxot & Goff, 1985: 161
Neoeucheyla (Cunliffella) panamensis (Baker): Volgin, 1969: 260
Cunliffella panamensis: Summers & Price, 1970: 69; Patxot & Goff, 1985: 161; Fain & Bochkov, 2001: 72
Type locality: Panama
Distribution: Panama, USA (Colorado, California, Hawaii Is., Oahu)
Host: among eggs of termites
variegata Barilo, 1985: 45
Cunliffella variegata (Barilo): Fain & Bochkov, 2001: 72; Salarzehi, 2018:
Type locality: Uzbekistan
Distribution: Iran, Tadjikistan, Uzbekistan
Eucheyla whartoni Baker, 1949: 69; Volgin, 1964: 93
Neoeucheyla (Cunliffella) whartoni (Baker) Volgin, 1969: 255
Cunliffella tuberculicoxa Volgin, 1964: 90 – type: Murmansk Distr., Russia
Neoeucheyla (Cunliffella) tuberculicoxa Volgin, 1969: 256
Cunliffella tuberculicoxa (Volgin): Summers & Price, 1970: 71; Fain & Bochkov, 2001: 72 (syn.)
Cunliffella whartoni: Summers & Price, 1970: 70; Khaustov, 2021: 235
Type locality: Birnamwood, Wisconsin
Distribution: Estonia, Kyrgyzstan, Russia (as C. tuberculicoxa), USA (Wisconsin)
Hosts (of Cunliffella tuberculicoxa): Aradus lugubris, A. aterrimus, A. nemtschinovi (Heteroptera: Aradidae); C. whartoni is recorded by Khaustov (2021) from ant hill of Formica rufa (Formicidae).
Dubininiola Volgin, 1969: 242
Type species: Dubininiola polylepis Volgin, 1969
Polycheyletus Vaivanijkul, 1979: 251
Type species: Polycheyletus boonkongae Vaivanijkul, 1979
Oudemansicheyla batangeniae Corpuz-Raros & Sotto, 1977: 163
Polycheyletus batangenius (Corpuz-Raros et Sotto): Corpuz-Raros, 1980: 63
Dubininiola batangensis (Corpuz-Raros et Sotto): Fain & Bochkov, 2001 : 300; 2001 : 77
Type locality: Tanauan, Batangas
Distribution: Philippine Is.
Polycheyletus boonkongae Vaivanijkul, 1979: 251
Dubininiola boonkongae (Vaivanijkul): Fain & Bochkov, 2001 : 300; 2001: 77
Type locality: Thailand
Distribution: Thailand
Dubininiola polylepis Volgin, 1969: 242; Fain & Bochkov, 2001 : 300; 2001 : 77
Type locality: Deynausskiy Rayon
Distribution: Turkmenistan
Host: rodent’s burrow
Eucheletopsis Volgin, 1969: 356
Type species: Cheyletus major Trouessart, 1893 (Oudemans, 1904)
Cheyletus major Trouessart in Berlese, 1893: 74 (nomen nudum)
Cheletopsis major (Trouessart): Oudemans, 1904b: 163; 1906: 200; Baker, 1949: 309
Eucheletopsis major (Trouessart): Volgin, 1969: 356; Summers & Price, 1970: 73
Type locality: New Guinea
Distribution: New Guinea
Host: Hemiprocne (= Dendrochelidon) mystacea (Apodiformes: Hemiprocnidae)
Eutogenes Baker, 1949: 304
Type species: Eutogenes foxi Baker, 1949
Eutogenes bakeri Corpuz-Raros, 1998: 268
Type locality: Leyte Is., Visayas St. College of Agriculture, Baybay
Distribution: the Philippines
Eutogenes bicornis Khaustov, 2021: 200
Type locality: Tyumen Province, Tyumen Region, Tyumen, Zatyumenskiy park
Distribution: Russia
Eutogenes citri Gerson, 1967: 363; Volgin, 1969: 285; Summers & Price, 1970: 60
Type locality: Karmon
Distribution: Israel
Eutogenes cornutus Corpuz-Raros, 1998: 270
Type locality: Luzon Is., Mt. Makiling, Putting Lupa, Calamba, Laguna, 300 m
Distribution: the Philippines
Eutogenes foxi Baker, 1949: 304; Volgin, 1969: 281; Summers & Price, 1970: 58
Type locality: Brownsville (Texas)
Distribution: USA (Texas, Florida), Puerto Rico
Eutogenes frater Volgin, 1958: 460; 1969: 282; Summers & Price, 1970: 60; Fain & Bochkov, 2001: 87; Bochkov, Hakimitabar & Saboori, 2005: 104;
Eutogenes africanus Wafa & Soliman, 1968: 225 – type: Giza, Burg El Arab and the north of the Delta, associated with scale insects, also collected from the bird nests; synonymized by Fain & Bochkov, 2001: 87
Type locality: Bulgaria
Distribution: Algeria, Bulgaria, Egypt, Iran, Turkey
Eutogenes makilingiensis Corpuz-Raros, 1998: 271
Type locality: Mt. Makiling, Luzon Is., Putting Lupa, Calamba, Laguna, 300 m
Distribution: Philippines
Eutogenes narashinoensis Hara & Hanada, 1960: 25; Volgin, 1969: 286; Summers & Price, 1970: 60
Type locality: Narashino, Chiba Prefecture, free living
Distribution: Japan
Cheletogenes onoi Shiba, 1976: 171
Eutogenes onoi (Shiba): Ehara & Abdul Ghani, 1988: 240
Type locality: Malaysia
Distribution: Malaysia
Eutogenes pinicola Thewke & Enns, 1972: 453
Type locality: Missouri
Distribution: USA (Missouri)
Eutogenes punctata Zaher & Soliman, 1966: 65; Wafa & Soliman, 1968: 225; Summers & Price, 1970: 60
Eutogenes punctatus Volgin, 1969: 283
Type locality: Giza
Distribution: Egypt
Cheletogenes quadrisetatus Berlese 1913: 79; Wafa & Soliman, 1968: 224
Cheletogenes quadrisetosus: Baker, 1949: 305
Eutogenes quadrisetatus (Berlese); Volgin, 1969: 284; Summers & Price, 1970: 60; Xia, 2010: 161
Type locality: Java
Distribution: China, Indonesia (Java)
Eutogenes reticularis Olivier & Theron, 1988: 253
Type locality: Transvaal
Distribution: South Africa
Eutogenes vicinus Summers & Price, 1970: 59; Moser & Roton, 1971: 1775; Smiley & Moser, 1975: 406
Type locality: San Luis Obispo, California
Distribution: U.S.A. (California, Louisiana)
Grallacheles De Leon, 1962: 135
Type species: Grallacheles bakeri De Leon, 1962
Grallacheles bakeri De Leon, 1962: 135; Muma, 1964: 248; Volgin, 1969: 275; Summers & Price, 1970: 49; Corpuz – Raros & Sotto, 1977: 154; Tseng, 1977; Rosa & Flechtmann, 1979: 196; Žďárková, 1979; Fain & Bochkov, 2001 : 78; Xia, 2010: 161; Torre Santana, 2015
Paracheyletia woolfordi Cunliffe, 1962: 200 – type from Japan, intercepted in Hawaii; Tahiti; Nishida (Version 2009)
Type locality: Coral Gables (Florida), Mante (Tamaulipas)
Distribution: Angola, Brazil, D.R. Congo, Cuba, Japan, Malaysia, Mexico, Philippines Is., Surinam, Tahiti, Taiwan, USA (Florida, Hawaii), ‘Czechoslovakia’
Grallacheles indicus Podder, Gupta & Saha, 2006: 333
Type locality: West Bengal
Distribution: India
Grallacheles nanfengensis Xia, Ye & Zhu, 1997: 173; Xia et al., 2000; Xia, 2010: 161
Type locality: Jiangxi
Distribution: China (Jiangxi)
Hoffmannita Pelaez, 1962: 72
Myrmicocheyla Volgin, 1963: 935
Type species: Hoffmannita mexicana Pelaez, 1962
Myrmicocheyla clavipes Volgin, 1963: 935
Hoffmannita navicula: Lin & Zhang, 1997
Hoffmannita clavipes (Volgin): Volgin, 1969: 266; Xia, 2010: 161; Khausov, 2021: 235 (Kurgan and Tyumen Provinces)
Type locality: Ushkovo, Russia (Leningradskaya Prov.), ant hills of Formica rufa (Hymenoptera: Formicidae)
Distribution: China, Czech Rep., Russia
Hosts: Aradus betulae, A. corticalis (Heteroptera: Aradidae), ant hills
Hoffmannita gersoni Fain & Bochkov, 2001 : 73
Type locality: Curacao
Distribution: Netherlands-Antilles
Host: Coenobita clypeata (Decapoda: Coenobitidae)
Hoffmannita mexicana Pelaez, 1962: 75; Fain & Bochkov, 2001 : 73
Type locality: Estado de Oaxaca
Distribution: Mexico
Host: Centruroides flavopictus (Scorpiones: Buthidae)
Hoffmannita rimandoi Corpuz-Raros, 1972: 262
Type locality: Philippines
Distribution: Philippines
Hylopecheyla Fain, 1972: 41
Type species: Hylopecheyla bunguranensis Fain, 1972
Hylopecheyla bunguranensis Fain, 1972: 42; 1979: 420; Fain & Bochkov, 2001: 60
Type locality: Bunguran Is., Natunas
Distribution: Indonesia (Natunas, Java)
Hosts: Hylopetes everetti, H. sagitta (Rodentia: Pteromyidae)
Hylopecheyla malayi Fain & Nadchatram, 1980: 197; Fain & Bochkov, 2001: 59
Type locality: Jeugka Triangle Feedacheme (Pahang)
Distribution: Malaysia
Host: Tupaia glis (Scandentia: Tupaiidae)
Hypopicheyla Volgin, 1969: 242
Type species: Hypopicheyla elongata Volgin, 1969
Hypopicheyla elongata Volgin, 1969: 243; Summers & Price, 1970: 65–66; Doğan & Ayyildiz, 2004: 1; Bayram & Cobanoglu, 2006; Salarzehi, 2018
Type locality: Russia (many areas in European and Asiatic Russia); Estonia (Tartu)
Distribution: Estonia, Russia, Iran, Turkey; ?USA (California)
Hosts: Aradus aterrimus, A. crenaticollis, A. lugubris, A. nemtschinovi, etc. (Heteroptera: Aradidae)
Cheyletia mirabilis Volgin, 1955: 170; 1956: 212
Hypopicheyla mirabilis (Volgin): Volgin, 1969: 245; Summers & Price, 1970: 65; Corpuz-Raros, 1998; Bochkov, Hakimitabar & Saboori, 2005: 104
Type locality: Hiva (Uzbekistan)
Distribution: Iran, Philippines, Uzbekistan
Host: Gonocephalum rusticum (Coleoptera: Tenebrionidae)
Metacheletoides Fain, 1972: 40
Type species: Metacheletoides numidae Fain, 1972
Cheletoides akanyaruensis Fain, 1972: 40
Metacheletoides akanyaruensis (Fain): Fain, 1979: 1017
Type locality: Akanyaru River
Distribution: Rwanda
Host: Numida meleagris intermedia (Galliformes: Numididae)
Metacheletoides crinifer Fain, 1979: 1018
Type localites: Butare, Gisagara (Rwanda), Marona (Cameroons), Lama Kara (Togo), Lotanoko (D. R. Congo)
Distribution: Cameroons, D. R. Congo, Rwanda, Togo
Hosts: Crinifer piscator, C. zonurus, Corythaixoides leucogaster (Musophagiformes: Musophagidae)
Metacheletoides gisagarensis Fain, 1979: 1021
Type locality: Butare (Rwanda), Molegbe (D. R. Congo), Lama Kara (Togo)
Distribution: D. R. Congo, Rwanda, Togo
Hosts: Crinifer piscator, C. zonurus (Musophagiformes: Musophagidae)
Metacheletoides numidae Fain, 1972: 41; 1979: 1013
Type locality: n. Akanyaru River (Rwanda)
Distribution: Burundi, Ethiopia, Rwanda
Hosts: Numida meleagris intermedia, N. meleagris (Galliformes: Numididae)
Mexecheles De Leon, 1962: 132
Acarocheyla Volgin, 1965: 293 (syn. by Summers & Price, 1970: 43)
Type species: Mexecheles cunliffei De Leon, 1962
Mexecheles aztecorum De Leon, 1962: 133; Summers & Price, 1970: 45
Type locality: Oaxaca, Tuxtla Gutierez (Chiapas)
Distribution: Mexico (Oaxaca, Chiapas)
Host: on Quercus conzattii and Cocos nucifera
Mexecheles cunliffei De Leon, 1962: 132; Volgin, 1969: 185; Summers & Price, 1970: 44
Type locality: Huajuapan de León, Oaxaca
Distribution: Mexico (Oaxaca)
Host: Ipomea murucoides (Convolvulaceae)
Cheletophyes hawaiiensis Baker, 1949: 289
Mexecheles hawaiiensis (Baker): Muma, 1964: 248; Volgin, 1969: 198; Summers & Price, 1970: 44; Flechtmann, 1971: 31; Torre Santana, 2015
Mexecheles intermedius De Leon, 1962: 133; Muma, 1964: 248
Acarocheyla hawaiiensis: Volgin, 1965: 293
Type locality: Kailua, Oahu, Hawaii
Distribution: Brazil, Mexico, Panama, USA (Hawaiian Is.)
Acarocheyla impolita Smiley & Moser, 1970: 231
Mexecheles impolitus (Smiley et Moser): Summers & Price, 1970: 48; Fain & Bochkov, 2001: 55
Type locality: Elizabeth, Louisiana
Distribution: USA (Louisiana)
Host: in boring dust and in galleries of Dendroctonus spp. (Coleoptera: Scolytidae)
Cheletophyes marshalli Baker, 1949: 290
Mexecheles aztecorum De Leon, 1962: 133; Summers & Price, 1970: 45
Acarocheyla marshalli: Volgin, 1965: 294
Mexecheles marshalli (Baker): Gerson, Fain & Smiley, 1999: 70
Type locality: Imboden (Arkansas)
Distribution: Mexico, USA (Arkansas)
Mexecheles panneus Summers & Price, 1970: 46
Type locality: Lehman Creek, 8 miles W of Baker, Nevada
Distribution: USA (Nevada)
Mexecheles shiva Bochkov & Ochoa, 2005: 137
Type locality: Bombay
Distribution: India
Mexecheles thailandensis Fuangarworn & Lekprayoon, 2010: 60
Type locality: Sasaloy Subdistrict, Tharua District, Ayutthaya Province
Distribution: Thailand
Host: from tree bark infested by scolytid beetles
Cheyletia virginiensis Baker, 1949: 299
Paracheyletia virginiensis (Baker): Volgin, 1955: 169
Mexecheles virginiensis (Baker): De Leon, 1962: 132; Volgin, 1969: 187; Summers & Price, 1970: 47; Fain & Bochkov, 2001: 54; Bochkov & Abramov, 2016; Diumina & Abramov, 2018: 213; Khaustov, 2021: 234
Acarocheyla virginiensis (Baker): Smiley & Moser, 1970: 233
Type locality: Gloucester Co., Virginia (USA)
Distribution: India (Kashmir), U.S.A. (Virginia, Louisiana, Nevada, Utah), Russia
Host: from boring dust of Ips calligraphus (Coleoptera: Scolytidae)
Mexecheles votandinii Jeffrey, 1976: 668; 1977: 654
Type locality: Girvan, Ayrshire (Scotland)
Distribution: Scotland (U.K.)
Microcheyla Volgin, 1966b: 277
Type species: Microcheyla parvula Volgin, 1966
Microcheyla bengalensis Gupta & Paul, 1992: 76
Type locality: West Bengal, Medinipur
Distribution: India
Host: Ploceus philippinus (Passeriformes: Ploceidae)
Microcheyla granifera Kuznetzov, 1977: 925
Microcheyla ozkani Koç & Ayyldiz, 1995: 226 – Turkey (syn. by Fain & Bochkov, 2001 : 74)
Type locality: Yalta, Crimea
Distribution: Turkey, Crimea
Microcheyla parvula Volgin, 1966b: 278; 1969: 270; Summers & Price, 1970: 72; Salarzehi, 2018:
Type locality: Ramit (Tadjikistan)
Distribution: Iran, Tadjikistan
Microcheyla paraparvula Olivier & Theron, 1989b: 49
Type locality: Transvaal
Distribution: South Africa (Transvaal)
Type locality: Transvaal
Distribution: South Africa (Transvaal)
Muricheyla Fain, 1972: 42
Type species: Muricheyla sicista Fain, 1972
Muricheyla sicista Fain, 1972: 43
Type locality: Caucase
Distribution: Caucase
Host: Sicista subtilis (Rodentia: Zapodidae)
Neoeucheyla Radford, 1950: 82
Type species: Cheyletia (Eucheyla) loricata Berlese, 1913
Neoeucheyla bochkovi Fain, 2003: 97
Type locality: Algiers
Distribution: Algeria
Host: from a flower in a garden
Neoeucheyla iranica Fain & Ardeshir, 2000: 329; Fain & Bochkov, 2001 : 72; Bochkov, Hakimitabar & Saboori, 2005: 105
Type locality: Iran
Distribution: Iran
Host: in a silo
Cheyletia (Eucheyla) loricata Berlese, 1913: 79; Volgin, 1955: 170
Neoeucheyla loricata (Berlese): Volgin, 1969: 25; Summers & Price, 1970: 67; Fain &
Bochkov, 2001 : 72; Bochkov & Mironov, 2007: 75
Type locality: Palermo («in Agri Panormitani muscis»)
Distribution: Italy (Sicily)
Neoeucheyla minuta Barilo, 1986: 28; Fain & Bochkov, 2001 : 72; Bochkov & Mironov, 2007: 75
Type locality: Uzbekistan
Distribution: Uzbekistan
Neoeucheyla macrocorneus Soliman, 1975: 101
Type locality: Giza
Distribution: Egypt
Neoeucheyla ploceus Gupta & Paul, 1992: 74; Bochkov & Mironov, 2007: 75
Type locality: West Bengal, Medinpur
Distribution: India
Host: Ploceus philippinus (Passeriformes: Ploceidae)
Oconnoricheylus Bochkov & Otto, 2010: 2
Type species: Oconnoricheylus speciosus Bochkov et Otto, 2010
Alliea chimaera Bochkov & Ochoa, 2005: 140
Oconnorichelus chimaera: Bochkov & Otto, 2010: 3
Type locality: Queensland
Distribution: Australia (Queensland)
Oconnoricheylus speciosus Bochkov & Otto, 2010: 3
Type locality: USA
Distribution: USA
Host: from oranges
Oudemansicheyla Volgin, 1969: 272
Type species: Cheletomimus denmarki Yunker, 1961
Oudemansicheyla coprosomae Thewke & Enns, 1976: 369
Type locality: Cobb L., Nelson Co.
Distribution: New Zealand
Cheletomimus denmarki Yunker, 1961: 1032; De Leon, 1962: 134
Oudemansicheyla denmarki (Yunker): Volgin, 1969: 272; Summers & Price, 1970: 10;
Tseng, 1977: 254; Gerson, 1994: 439; Lin et al., 2000: 86; Xia, 2010: 161
Type locality: Florida (USA)
Distribution: Australia, China (Fujian), Taiwan, USA (Florida, Louisiana)
Paramicrocheyla Olivier & Theron, 1989: 238; Fain & Bochkov, 2001 : 299 (genus incertae sedis)
Type species: Paramicrocheyla spinula Olivier et Theron, 1989
Paramicrocheyla spinula Olivier & Theron, 1989: 238
Type locality: Transvaal
Distribution: South Africa (Transvaal)
Paramicrocheyla ungulina Olivier & Theron, 1989: 242
Type locality: Transvaal
Distribution: South Africa (Transvaal)
Picocheyletus Bochkov & OConnor, 2003: 268
Type species: Picocheyletus tricholaemae Bochkov et OConnor, 2003
Picocheyletus tricholaemae Bochkov & OConnor, 2003: 268
Type locality: Ogooue – Maritime, 25.2 km NW Doussala
Distribution: Gabon
Host: Tricholaema hirsuta flavipunctata (Piciformes: Capitonidae)
Promuricheyla Fain, 1979: 413
Type species: Promuricheyla lukoschusi Fain, 1979
Promuricheyla lukoschusi Fain, 1979: 413
Type species: Mt. Katanglad, Mindanao
Distribution: Philippines (Mindanao)
Host: Nannosciurus surrutilus (Rodentia: Sciuridae)
Prosocheyla Volgin, 1969: 288
Type species: Cheletogenes oaklandia Baker, 1949
Prosocheyla Volgin, 1969: 288
Type species: Cheletogenes oaklandia Baker, 1949
Prosocheyla acanthus Smiley & Moser, 1970: 229; Fain & Bochkov, 2001: 85
Prosocheyla acantha: Summers & Price, 1970: 57
Type locality: Maurepas (Louisiana)
Distribution: USA (Louisiana)
Host: outer bark of loblolly pine
Cheletogenes oaklandia Baker, 1949: 306
Prosocheyla (Prosocheyla) oaklandia (Baker): Volgin, 1969: 289; Fain & Bochkov, 2001: 85
Prosocheyla oaklandia (Baker): Summers & Price, 1970: 55 (many localities in California); Lucza, Ripka & Saly, 1996 (oaklandi)
Type locality: Oakland, California
Distribution: Hungary, USA (California)
Prosocheyla ripkai Fain & Bochkov, 2001: 85
Type locality: Budapest
Distribution: Hungary
Host: from a tree Populus simonii
Prosocheyla (P.) sinensis Xia, Liang & Zhu, 2005: 205; Xia, 2010: 161
Type locality: Henan
Distribution: China
Cheletogenes traubi Baker, 1949: 307
Prosocheyla traubi (Baker): Volgin, 1969: 291; Summers & Price, 1970: 56; Lucza, Ripka & Saly, 1996; Ripka, Fain & Bolland, 1999: 366; Fain & Bochkov, 2001: 85;
Type locality: Assam, Stillwell Road, near Ledo, India
Distribution: Burma, Hungary, India
Prosocheyla villosa Bochkov & Haustov, 1999: 151; Fain & Bochkov, 2001: 85
Type locality: Crimea, Aj-Danil’
Distribution: Crimea
Host: Xylocleptes bispinus (Coleoptera: Curculionidae)
Reckiana Volgin, 1969: 291
Type species: Cheletogenes buckneri Baker, 1949
Cheletogenes buckneri Baker, 1949: 308
Prosocheyla (Reckiana) buckneri (Baker): Volgin, 1969: 293; Fain & Bochkov, 2001: 85
Prosocheyla buckneri (Baker): Summers & Price, 1970: 56
Type locality: Santa Paula, California
Distribution: USA (California)
Cheletogenes citrifoliata Muma, 1964: 241
Prosocheyla (Reckiana) citrifoliata (Muma): Volgin, 1969: 294; Fain & Bochkov, 2001: 85; syn. by Summers & Price, 1970: 56 with Prosocheyla buckneri
Type locality: Florida
Distribution: USA (Fla)
Cheletogenes hepburni Lawrence, 1954: 70
Prosocheyla (Reckiana) hepburni (Lawrence): Volgin, 1969: 292
Prosocheyla hepburni (Lawrence): Summers & Price, 1970: 58
Type locality: Hamburg, Peddie Distr., South Africa
Distribution: South Africa
Host: Lepidosaphes beckii (Homoptera: Coccodea: Diaspididae)
Samsinakia Volgin, 1965: 296
Cryptocheyla Fain, 1972: 41
Type species: Cheletophyes theodoridis Samšinak, 1959
Samsinakia carabae Ramaraju & Mohanasundram, 1999: 125; Fain & Bochkov, 2001: 76
Type locality: Tamil Nadu, Trichy
Distribution: India (Tamil Nadu)
Host: Carabidae (Coleoptera)
Samsinakia charanasriae Fuangarworn & Lekprayoon, 2010: 64
Type locality: Buang-Chawak Park, Supan Buri Province
Distribution: Thailand
Host: from soil and leaf-litter
Samsinakia gonocephalum Fain, 1984: 686; Bochkov & Mironov, 1998: 265; Fain & Bochkov, 2001: 76
Type locality: DR Congo
Distribution: D. R. Congo (former Zaire), India
Host: Gonocephalum simplex (Coleoptera: Tenebrionidae)
? Samsinakia pagongae Corpuz-Raros & Sotto, 1977: 165; Fain & Bochkov, 2001: 76
Type locality: Cale, Tanauan, Batangas
Distribution: Philippine Is.
Cheletophyes theodoridis Samšinák, 1959: 81
Samsinakia theodoridis (Samšinák): Volgin, 1965: 298; 1969: 247; Summers & Price, 1970: 71; Fain & Bochkov, 2001: 76
Type locality: Tananarive
Distribution: Madagascar
Host: Selinus abacoides (Coleoptera: Tenebrionidae)
Samsinakia tilae Bochkov & Klimov, 2005: 15
Type locality: Chiapas, 39 km on road between Tuxtla Gutierrez and Revolutión Mexicana
Distribution: Mexico (Chiapas)
Host: Blastinus sp. (Coleoptera: Tenebrionidae)
Samsinakia trilobitus Bochkov & Mironov, 1998: 266; Fain & Bochkov, 2001: 76; Kazemi & Paktinat-Saeij. 2021: 107
Type locality: Anamalai Hills, Cinchora
Distribution: India, Iran
Host: beetles (Coleoptera: Tenebrionidae)
Cryptocheyla volgini Fain, 1972: 41
Samsinakia volgini (Fain): Fain, 1980: 105; Gerson, 1994: 444; Fain & Bochkov, 2001: 76
Type locality: Butare (Rwanda)
Distribution: Australia, D. R. Congo, Rwanda
Host: nest of Spermestes cucullatus (Passeriformes); on Gonocephalum simplex (Coleoptera: Tenebrionidae)
Thewkachela Ide & Kethley, 1977: 559
Type species: Thewkachela ratufi Ide et Kethley, 1977
Thewkachela ratufi Ide & Kethley, 1977: 559; Fain & Nadchatram, 1980: 199
Type locality: Sabah
Distribution: Malaysia (Sabah), Thailand
Hosts: Ratufa affinis, R. bicolor (Rodentia: Sciuridae)
Criokerontinae Smiley, 1978: 239
Type genus: Criokeron Volgin, 1966
Niheliinae Smiley, 1978: 239
Type genus: Nihelia Domrow et Baker, 1960
Criokerontini Smiley, 1978: 239
Type genus: Criokeron Volgin, 1966
Criokeron Volgin, 1966: 293
Type species: Nihelia quinta Domrow et Baker, 1963
Nihelia quinta Domrow & Baker, 1963: 230
Criokeron quinta: Fain & Nadchatram, 1980: 199
Criokeron quintus (Domrow et Baker); Volgin, 1966: 219; 1969: 386; Fain & Lukoschus, 1985: 264
Type locality: Subang, Selangor
Distribution: Malaysia, Thailand
Hosts: Tupaia belangeri, T. glis (Scandentia: Tupaiidae)
Criokeron thailandicus Fain & Lukoschus, 1985: 266
Type locality: Nakhon Kachisima, Pak Thong Chai, Ban Sakaerat
Distribution: Thailand
Host: Tupaia glis (Scandentia: Tupaiidae)
Galagocheles Fain, 1979: 629
Type species: Cheletiella lemuricola Lawrence, 1948
Galagocheles lawrencei Bochkov, Mironov & OConnor, 2006: 189
Type locality: 3 km SE Nayaoure (Burkina Faso = Upper Volta)
Distribution: Burkina Faso, DR Congo, Rwanda
Hosts: Galago moholi, G. senegalensis (Primates: Galagonidae)
Cheletiella lemuricola Lawrence, 1948: 39
Hemicheyletus lemuricola: Lawrence, 1954: 74; 1961: 114
Hemicheyletus (H.) lemuricola: Volgin, 1969: 378
Nihelia lemuricola (Lawrence): Domrow & Baker, 1963: 226; Summers & Price, 1970: 75
Galagocheles lemuricola (Lawrence): Fain, 1979: 630; Bochkov, Mironov & OConnor, 2006: 185
Type locality: Kwa Zulu-Natal, Cato Ridge, nr. Pietermaritzburg
Distribution: D. R. Congo (former Zaire), Rwanda, South Africa
Hosts: Otolemur garnetti, O. [Galago] crassicaudata monteri, Galago moholi (Primates: Galagonidae)
Hemicheyletus Lawrence, 1954: 74
Nihelia Domrow & Baker, 1960: 194
Type species: Cheletiella curvidens Lawrence, 1948
Nihelia calcarata Domrow & Baker, 1960: 194; 1963: 226
Hemicheyletus calcaratus (Domrow et Baker): Volgin, 1969: 382
Hemicheyletus calcarata: Nagar, 1978: 102
Hemicheyletus herpestesi: Nagar & Saxena, 1977: 35
Type locality: Kanchunaburi, Muang, Thailand
Distribution: India, Thailand
Hosts: Herpestes sp. (Thailand), H. edwardsii (India)(Carnivora: Herpestidae)
Cheletiella curvidens Lawrence, 1948: 40
Hemicheyletus curvidens, Lawrence, 1954: 74; 1956: 338; 1961: 114
Hemicheyletus (Hemicheyletus) curvidens: Volgin, 1969: 380
Nihelia curvidens (Lawrence): Domrow & Baker, 1963: 226; Summers & Price, 1970: 75
Type locality: Natal
Distribution: Angola, South Africa
Hosts: Galerella [Herpestes] sanguinea, Herpestes [Myonax cauui] punctulatus, H. bocagei (Carnivora: Herpestidae)
Hemicheyletus cynictis Fain, 1979: 625
Nihelia cynictis (Fain)
Type locality: Bloemhof (Transvaal)
Distribution: South Africa (Transvaal)
Host: Cynictis penicillata (Carnivora: Viverridae)
Hemicheyletus (Sciurocheyla) Volgin, 1969: 384
Thryonomycheyla Fain, 1972: 42 (syn. by Bochkov & OConnor, 2004)
Smileycheles Fain, 1979: 627 (syn. by Bochkov & OConnor, 2004)
Type species: Nihelia squamosa Domrow et Baker, 1963
Type species of Smileycheles Fain: S. camerounensis Fain, 1979
Type species of Thryonomycheyla Fain: T. congolensis Fain, 1972
Thryonomycheyla angolensis Fain & Bochkov, 2002: 148
Sciurocheyla angolensis (Fain et Bochkov):
Type locality: Angola
Distribution: Angola
Host: (?) Thryonomys swinderianus (Rodentia: Thryonomyidae)(contamination ?)
Smileycheles camerounensis Fain, 1979: 627
Sciurocheyla camerounensis (Fain): Bochkov & OConnor, 2004: 316
Type locality: Foulassi (Cameroon)
Distribution: Cameroon, Tanzania
Hosts: Heliosciurus undulatus (Rodentia: Sciuridae), Zenkerela insignis (Rodentia: Anomaluridae)(cont.?)
Thryonomycheyla congolensis Fain, 1972: 42
Sciurocheyla congolensis (Fain): Bochkov & OConnor, 2004: 309
Type locality: D.R. Congo
Distribution: D.R. Congo
Host: (?) Thryonomys swinderianus (Rodentia: Thryonomyidae)(cont.?)
Sciurocheyla funisciurus Bochkov & OConnor, 2004: 311
Type locality: Cibitoke Prov., Nat. Park, Ndora Zone, E Kirumura, 2100 m
Distribution: Burundi
Host: Funisciurus pyrrhopus (Rodentia: Sciuridae)
Nihelia squamosa Domrow & Baker, 1963: 227; Summers & Price, 1970: 75
Hemicheyletus (Sciurocheyla) squamosus (Domrow et Baker): Volgin, 1969: 384
Sciurocheyla squamosus (Domrow et Baker): Fain & Lukoschus, 1985: 264
Sciurocheyla squamosa (Domrow et Baker): Bochkov & Oconnor, 2004: 309
Type locality: Nan Prov., Pang Nam NU, Bun Yun Distr., betw. Ban Rong Kwang and Ban Klang Wiang
Distribution: Thailand
Host: Menetes sp. (Rodentia: Sciuridae)
Metacheyletiinae Fain, 1980: 107
Type genus: Metacheyletia Fain, 1972
Metacheyletia Fain, 1972: 43
Type species: Metacheyletia obesa Fain, 1972
Metacheyletia amazonae Bochkov & OConnor, 2003: 266
Type locality: Guyana
Distribution: Guyana
Host: Amazona amazonica (Psittaciformes: Psittacidae)
Metacheyletia degenerata Fain & Bochkov, 2003: 119
Type locality: Central Africa (Western Tanzania)
Distribution: Central Africa
Host: Serinus mozambicus (Passeriformes: Fringillidae)
Metacheyletia longisetosa Atyeo, Kethley & Pérez, 1984: 125
Type locality: Jalisco, Puerto Vallarta
Distribution: Mexico
Host: Amazona finsehi (Psittaciformes: Psittacidae)
Metacheyletia ngaii Bochkov & Skoracki, 2011: 93
Type locality: Tanganyika
Distribution: Tanzania
Host: Corythaixoides leucogaster (Musophagiformes: Musophagidae)
Metacheyletia obesa Fain, 1972: 43; 1980: 107
Type locality: Antwerp Zoo
Distribution: ?
Host: Psittacula sp. (Psittaciformes: Psittaculidae)
Ornithocheyletiinae Smiley, 1978: 228
Type genus: Ornithocheyletia Volgin, 1964
Ornithocheyletiini Volgin, 1969: 387
Apodichelini Fain, Smiley & Gerson, 1997: 81
Neocheyletiellini Fain, Smiley & Gerson, 1997: 81
Type genus: Ornithocheyletia Volgin, 1964
Apodicheles Fain, 1979: 253
Type species: Apodicheles cypsiurus Fain, 1979
Apodicheles apus Fain, 1979: 255; Fain & Bochkov, 2002a: 159
Type locality: Butare
Distribution: Rwanda
Host: Apus caffer (Apodiformes: Apodidae)
Apodicheles collocalia Bochkov & OConnor, 2003: 273
Type locality: Leyte Is., 4 km S, 1 km E Inopacan
Distribution: Philippines (Leyte)
Host: Collocalia esculenta marginata (Apodiformes: Apodidae)
Apodicheles cypsiurus Fain, 1979: 253
Type locality: Tzaneen, Transvaal
Distribution: South Africa
Host: Cypsiurus p. parvus (Apodiformes: Apodidae)
Cheyletus heteropalpus Mégnin, 1878: 11; 1880: 242
Cheyletiella heteropalpus: Berlese, 1886: fasc. 28, 2
Cheyletiella heteropalpa: Oudemans, 1906: 213
Neocheyletiella heteropalpus (Mégnin): Baker, 1949: 272
Cheyletiella heteropalpa: Starkoff & Starkoff, 1950: 96
Neocheyletiella heteropalpa: Brown, 1971: 1380
Apodicheles heteropalpus (Mégnin)
Type locality: Probably France
Distribution: France (?), Italy, USA
Host: Columba livia (Columbiformes: Columbidae)
Bakericheyla Volgin, 1966: 297
Type species: Cheyletiella chanayi Berlese et Trouessart, 1889
Apodicheyla Fain, 1979: 255
Type species: Bakericheyla (Apodicheyla) africana Fain, 1979
Bakericheyla (Apodicheyla) africana Fain, 1979: 256; Fain & Bochkov, 2001: 88
Type locality: Tzaneen, Transvaal
Distribution: South Africa
Host: Cypsiurus parvus (Apodiformes: Apodidae)
Bakericheyla Volgin, 1966: 297
Type species: Cheyletiella chanayi Berlese et Trouessart, 1889
Bakericheyla (Bakericheyla) benoiti Fain, 1980: 135
Type locality: Galim
Distribution: Cameroons
Host: Merops [Melittophagus] bullocki (Coraciiformes: Meropidae) on the skin
Cheyletiella Chanayi Berlese & Trouessart, 1889: 135; Berlese, 1889: fasc. 56, No 3
Cheletiella chanayi (Berlese et Trouessart): Oudemans, 1906: 213
Neocheyletiella chanayi, Baker, 1949: 274; Dubinin, 1957: 90; Volgin & Nikolaeva, 1965: 300
Bakericheyla chanayi (Berlese et Trouessart): Furman & Sousa, 1969: 858; Volgin, 1969: 400; 1987: 492; Smiley, 1970: 1060; Summers & Price, 1970: 77; Domrow, 1991: 1303; Fain & Bochkov, 2001: 88
Neocheyletiella faini Lawrence, 1959: 431 – Mossel Bay, Cape Province, from Cossypha dichroa (Passeriformes: Muscicapidae); synonymized by Fain & Bochkov, 2001: 88
Bakericheyla faini (Lawrence): Volgin, 1969: 405
Bakericheyla (B.) faini (Lawrence): Fain, 1980: 134 – from Cossypha dichroa, Cape Prov., South Africa
Bakericheyla (Bakericheyla) chanayi (Berlese et Trouessart): Fain, 1980: 133
Bakericheyla chanayi latior Fain, 1972: 46 – on Paroaria gularis
Type locality: Lyon, France
Distribution: Australia (Northern Territory), Holland, South Africa
Hosts: Motacilla alba (Passeriformes: Motacillidae), Poephila acuticauda, P. gouldiae (Passeriformes: Estrildidae), Carduelis carduelis, Fringilla coelebs (Passeriformes: Fringillidae), Paroaria gularis (Passeriformes: Thraupidae), Cossypha dichroa (Passeriformes: Muscicapidae)
Neocheyletiella subquadrata Lawrence, 1959: 427
Bakericheyla subquadrata (Lawrence): Volgin, 1969: 411; Summers & Price, 1970: 78; Fain & Bochkov, 2001: 88
Bakericheyla (B.) subquadrata (Lawrence): Fain, 1980: 134
Type locality: Kanji (Botswana)
Distribution: Botswana, South Africa
Host: Merops [Melittophagus pusillus] (Coraciiformes: Meropidae)
Neocheyletiella transvaalica Lawrence, 1959: 430
Bakericheyla transvaalica (Lawrence): Volgin, 1969: 406; Summers & Price, 1970: 78; Fain & Bochkov, 2001: 88
Bakericheyla (B.) transvaalica Fain, 1980: 134
Type locality: Mabelikwa (Transvaal)
Distribution: former U. S. S. R., South Africa, Rwanda, Zimbabwe
Hosts: Melittophagus pusillus, M. bullockoides, Merops apiaster, M. persicus, M. nubicoides (Coraciiformes: Meropidae)
Ornithocheyletia Volgin, 1964: 28
Type species: Ornithocheyletia dubinini Volgin, 1964
Ornithocheyletia aitkeni Fain, 1972: 46; 1981: 201
Type locality: Guama
Distribution: Brazil
Host: Turdus fumigatus (Passeriformes: Turdidae)
Ornithocheyletia argentinensis Fain, 1972: 45; 1981: 198
Type locality: Antwerp Zoo
Distribution: South America, (Argentina)
Hosts: Forpus passerinus, Nandays nanday (Psittaciformes: Psittacidae)
Ornithocheyletia boeroi Brandetti & Bischoff-de-Alzuet, 1981: 236
Type locality: Argentina
Distribution: Argentina
Host: Cyanocorax chrysops (Passeriformes: Corvidae)
Cheyletiella canadensis Banks, 1909: 133
Neocheyletiella canadensis (Banks): Baker, 1949: 272; Volgin, 1964: 32
Ornithocheyletia canadensis (Banks), Volgin, 1969: 393; Summers & Price, 1970: 78
Type locality: Guelph (Ontario)
Distribution: Bulgaria, Canada
Host: Picus viridis (Piciformes: Picidae)
Ornithocheyletia columbigallinae Fain & Bochkov, 2002: 148
Type locality: Piracicaba
Distribution: Brazil
Host: Columbigallina sp. (Columbifomes: Columbidae)
Ornithocheyletia dubinini Volgin, 1964: 28; 1969: 390; Summers & Price, 1970: 78; Smiley, 1970: 1065; 1977: 229; Fain, 1981: 186; Skoracki, Bochkov & Sikora, 2004: 87
Ornithocheyletia barri Smiley, 1977: 234 – type: Davis, California, on Sturnus vulgaris
Type locality: Kishinev (Moldavia)
Distribution: Holland, Moldavia, Poland; USA (California)
Host: Sturnus vulgaris (Passeriformes: Sturnidae)
Ornithocheyletia eulabes Fain, 1981: 200
Type locality: Antwerp Zoo
Distribution: Asia (Antwerp Zoo)
Host: Eulabes javana (Passerifomes: Eulabetidae)
Ornithocheyletia francolini Fain, 1972: 44
Type locality: Waterport, N. Transvaal
Distribution: South Africa
Host: Francolinus natalensis (Galliformes: Phasianidae)
Ornithocheyletia garrulax Fain, 1972: 46; 1981: 202
Type locality: Antwerp Zoo
Distribution: ?
Host: Garrulax lencolophus bicolor (Passeriformes: Timaliidae)
Ornithocheyletia geopeliae Fain, 1981: 192
Type locality: Antwerp Zoo
Distribution: ?
Host: Geopelia striata (Columbiformes: Columbidae)
Ornithocheyletia gersoni Smiley, 1970: 1065
Type locality: Wooster
Distribution: USA (Ohio)
Host: Dendrocopos pubescens (Piciformes: Picidae)
Ornithocheyletia granatina Fain, 1972: 46; 1981: 204
Type locality: Antwerp Zoo
Distribution: ?
Host: Granatina ianthinogaster (Passeriformes: Estrildidae)
Ornithocheyletia hallae hallae Smiley, 1970: 1068; Fain, 1981: 190;
Ornithocheyletia hallae: Haarløv & Mørch, 1975: 284; Smiley, 1978: 229; Hallas, 1978: 32; Rosa & Flechtmann, 1979: 121; Fain & Bochkov, 2002: 151; Valim & Gazeta, 2007: 518: Bilal, 2012: 169
Type locality: Brownsville, Texas
Distribution: Brazil, Denmark, England, South Africa, USA (Texas, California), Turkey, Iran
Hosts: Columba livia, Columbigallina sp. (Columbiformes: Columbidae); phoretic on Pseudolynchia canariensis (Diptera: Hippoboscidae)
Ornithocheyletia lamprocolius Fain, 1972: 45; 1981: 204
Type locality: Central Africa
Distribution: Central Africa
Host: Lamprocolius chloropterus (Passeriformes: Sturnidae)
Ornithocheyletia lawrenceae Smiley, 1970: 1065
Type locality: San Antonio
Distribution: USA (Texas)
Host: Psittacula sp. (Psittaciformes: Psittaculidae)
Ornithocheyletia leiotrix Fain, 1972: 45; 1981: 198
Type locality: Antwerp Zoo
Distribution: Eastern Asia
Host: Leiothrix lutea (Passeriformes: Timaliidae)
Ornithocheyletia lepidus Fain, 1981: 204
Type locality: Antwerp Zoo
Distribution: ?
Host: Garrulax leucolophus bicolor (Passeriformes: Leiothricidae)
Ornithocheyletia lichmerae Smiley, 1984: 239 (lapsus)
Ornithocheyletia lichmerae Domrow, 1991: 1313
Type locality: Western Australia
Distribution: Australia (W. A.)
Hosts: Geiciphila indistincta, Grallina cyanoleuca (Passeriformes: Monarchidae)
Ornithocheyletia lonchurae Smiley, 1984: 240; Domrow, 1991: 1313
Type locality: Western Australia
Distribution: Australia (W. A.)
Host: Donacola castaneothorax (Passeriformes: Estrildidae)
Ornithocheyletia lukoschusi Smiley, 1970: 1069; Fain, 1981: 198
Type locality: Nijmegen
Distribution: Holland
Host: Hirundo rustica (Passeriformes: Hirundinidae)
Ornithocheyletia mironovi Chirov & Bochkov, 1998: 35
Type locality: Lake Issyk – Kul Region, Ottuk Village
Distribution: Kyrgyzstan
Host: Riparia riparia (Passeriformes: Hirundinidae)
Ornithocheyletia orioli Fain & Bochkov, 2002: 151
Type locality: Akanyaru River, Rwanda
Distribution: Rwanda
Hosts: Oriolus oriolus (Passeriformes: Oriolidae), Lybius sp. (Piciformes: Capitonidae),
Indicator indicator (Piciformes: Indicatoridae), Pachyphantes superciliosa (Passeriformes: Ploceidae), Chrysococcyx klaasi (Cuculiformes: Cuculidae)
Ornithocheyletia phylloscopi Bochkov, Mironov & Gorgol, 1995: 74
Type locality: Leningradskaya oblast
Distribution: Russia, Ukraine
Host: Phylloscopus trochilis (Passeriformes: Phylloscopidae)
Cheyletiella pinguis Berlese, 1889: 56; 1893: 75
Cheletiella pinguis, Oudemans, 1906: 213; Poppe, 1909: Womersley, 1941: 59
Neocheyletiella pinguis, Baker, 1949: 275
Ornithocheyletia pinguis, Volgin, 1964: 34; 1969: 396; Smiley, 1970: 1065; 1977: 229; Fain, 1981: 186
Type locality: Florence
Distribution: Germany, Italy
Hosts: Columba livia domestica (Columbiformes: Columbidae), Sturnus vulgaris (Passeriformes: Sturnidae), Turdus merula (Passeriformes: Turdidae)
Ornithocheyletia prognae Bochkov & Galloway, 2001: 2017
Type locality: Winnipeg, Manitoba
Distribution: Canada
Host: Progne subis (Passeriformes: Hirundinidae)
Ornithocheyletia psittaci poicephali Fain, 1981: 194
Type locality: Western Africa
Distribution: Western Africa
Host: Poicephalus senegalus (Psittaciformes: Psittacidae)
Ornithocheyletia psittaci Fain, 1972: 43
Type locality: Antwerp Zoo
Distribution: Western Africa
Host: Psittacus erythacus (Psittaciformes: Psittacidae)
Ornithocheyletia smileyi Fain, 1972: 45; 1981: 196
Type locality: Antwerp Zoo
Distribution: ?
Host: Myopsitta monachus (Psittaciformes: Psittacidae)
Ornithocheyletia turdi Bochkov & Galloway, 2001: 2015
Type locality: Winnipeg, Manitoba
Distribution: Canada
Host: Turdus migratorius (Passeriformes: Turdidae)
Ornithocheyletia volgini Smiley, 1970: 1069; Fain, 1981: 190
Type locality: Wayne Co., Ohio
Distribution: USA (Ohio)
Host: Petrochelidon pyrrhonota (Passeriformes: Hirundinidae)
Ornithocheyletia wauthyi Fain & Bochkov, 2002: 152
Type locality: Rwanda
Distribution: Rwanda
Host: Lamprocolius sp. (Passeriformes: Sturnidae)
Teinocheylini Fain, 1974: 273
Teinocheylinae Fain
Type genus: Teynocheylus Fain, 1974
Teinocheylus Fain, 1974: 271
Type species: Teinocheylus longissimus Fain, 1974
Teinocheylus gundii Fain, Gerrits & Lukoschus, 1982: 450
Type locality: Chebikar
Distribution: Tunisia
Hosts: Ctenodactylus gundi, C. vali, Massoutiera mzabi (Rodentia: Ctenodactylidae)
Teinocheylus longissimus Fain, 1974: 271
Type locality: Berbera, N. Somalia
Distribution: Somalia
Host: Pectinator spekei (Rodentia: Ctenodactylidae)
EUROPE
Armenia (Volgin, 1969) – Cheletogenes ornatus, Cheletomimus (Hemicheyletia) bregetovae, Cheletomorpha lepidopterorum, Cheyletus malaccensis [syn. rohdendorfi]
Austria (Franz, 1954; Willmann, 1951) – Cheyletia squamosa, Cheyletus eruditus
Azerbaidjan (Zachvatkin, 1949; Halilova, 1967; Volgin, 1963, 1969, 1978; Abdullaeva, 1978; Abdullaeva & Aslanov, 1989; Aslanov & Mehtieva, 1992) – Cheyletus eruditus, Ch. malaccensis (= polymorphus, rohdendorfi), Ch. carnifex [syn. aversor], Cheletomorpha lepidopterorum, Lepidocheyla caucasica
Belarus (Efremova & Bochkov, 1990) – Acaropsellina sollers, Cheyletus eruditus, Ch. trux, Eucheyletia bothrophila, E. sibirica, Neocheyletiella microrhyncha (= oudemansi)
Belgium (Dufait, 1968; André, 1975; Fain & Bochkov, 2001) – Cheyletiella (?) parasitivorax, Eucheyletia flabellifera, Cheletonella vespertilionis, Cheletomorpha lepidopterorum, Paracheyletia pyriformis
Bulgaria (Volgin, 1955, 1958, 1964a, 1964b, 1969; Zankov, 1955; Kolebinova, 1967; Sosnina et al., 1969; Nachev & Trenchev, 1987; Beron, 2011; Trencheva & Trenchev, 2018) – Cheletogenes ornatus, Cheletopsis norneri, Cheyletus carnifex [syn. aversor], Ch. malaccensis (= polymorphus), Eucheyletia flabellifera, Eutogenes frater, Cheletomimus (Hemicheyletia) bregetovae, Neoeucheyla bulgarica, Ornithocheyletia canadensis
Croatia (Pagliarini, 1979) – Cheyletus eruditus
Czech Rep. (Samšinak, 1956; Volgin, 1966; Fain & Bochkov, 2001; Lukáš et al., 2007) – Chelacheles michalskii, Cheyletus carnifex (= aversor), Ch. eruditus, Ch. malaccensis, Ch. trouessarti, Paracaropsis strofi, Paracheyletia pyriformis (= samsinaki),
“Czechoslovakia” (Žďárková, 1979) – Acaropsellina docta, Cheletomorpha lepidopterorum, C. hendersoni, Cheyletia papillifera, Cheyletus carnifex (= aversor), C. eruditus, C. malaccensis, C. trouessarti, Grallacheles bakeri, Ker bakeri
Denmark (Haarløv & Mørch, 1975; Haarløv, 1957; Enghoff, 1988; Hallas, 1978) – Cheletomorpha lepidopterorum, Cheyletiella parasitivorax, Ch. yasguri, Cheyletus eruditus, Ch. trouessarti, Ornithocheyletia hallae
Estonia (Volgin, 1964) – Cunliffella whartoni (= tuberculicoxa)
Finland (Harju, Husman, Merikoski & Pennanen, 2006) – Mexecheles virginiensis
France (contin.)(Megnin, 1883; Guilhon & Marchand, 1972, 1974) – Cheletomorpha lepidopterorum, Cheyletiella blakei, Ch. parasitivorax, Ch. yazguri, Neocheyletiella microrhyncha
Georgia (Volgin, 1963; Rekk, 1976; Fain, Bochkov & Corpuz-Raros, 2002) – Cheletogenes ornatus, Cheletomimus (Ch.) berlesei, Cheletomorpha lepidopterorum, Cheletonella caucasica, Cheyletus eruditus, Ch. malaccensis [syn. rohdendorfi], Ch. trux, Eucheyletia reticulata
Germany (Heller, 1880; Haller, 1882; Poppe, 1909; Rack, 1971; Fain & Bochkov, 2001) – Cheletoides uncinatus, Cheletomorpha lepidopterorum, Cheletopsis noerneri, Cheyletiella yazguri, Cheyletus eruditus, Eucheyletia flabellifera, Ornithocheyletia pinguis, Paracheyletia pyriformis
Great Britain (Michael, 1878; Hull, 1918; Turk, 1946, 1953; Evans, 1954; Griffiths, 1960; Beesley, 1963; Walton, 1974; Hughes, 1976; Jeffrey, 1976, 1978, 1979, 1980; Fain & Bochkov, 2001; Fain, Bochkov & Corpuz-Raros, 2002; Baker & Craven, 2003) – Acaropsellina docta, A. sollers, Cheletogenes ornatus, Cheletomimus (Hemicheyletia) berlesei, Cheletomimus greenwoodi, Cheletomorpha lepidopterorum, Cheyletiella parasitivorax, Ch. yasguri, Cheyletus eruditus, Ch. trouessarti (= Ch. woodroffei), Eucheyletia flabellifera, Ornithocheyletia hallae
Greece (Papaioannou-Souliotis, 1994; Eliopoulos & Papadoulis, 2001; Athanassiou et al., 2003; Palyvos & Emmanouel, 2006; Palyvos, Emmanouel & Saitanis, 2008) – Acaropsellina docta, A. sollers, Chelacheles hellenicus, Cheletogenes berlesei (= ornatus), Cheletomimus duosetosus, Cheletomorpha lepidopterorum, Cheyletiella parasitivorax, Cheyletus aversor, Ch. cacahuamilpaensis, Ch. eruditus, Ch. malaccensis (= Ch. polymorphus), Ch. trouessarti, Ch. trux (= tenuipilis),
Holland (Oudemans, 1903; Smiley, 1970; Bronswijk et al., 1972; Volgin, 1987; Fain & Bochkov, 2001) – Acaropsellina docta, Bakericheyla chanayi, Cheletomorpha lepidopterorum, Cheyletiella parasitivorax, Ch. yasguri, Cheyletus eruditus, Ch. trouessarti, Eucheyletia flabellifera, Ornithocheyletia lukoschusi
Hungary (Dudich, Kolosváry & Szalay, 1940; Komlovszky, 1980, 1984; Lucza, Ripka & Saly, 1996; Ripka, Fain & Bolland, 1999; Fain & Bochkov, 2001) – Acaropsellina docta, Cheletacarus raptor, Cheletogenes ornatus, Cheyletus aversor, Ch. eruditus, Ch. trouessarti, Prosocheyla ripkai, P. traubi
Ireland (Baker, 1969; Dodd, 1970) – Cheyletiella yasguri
Iceland (Sellnick, 1940) – Cheyletus eruditus
Italy (contin.) (Berlese, 1886, 1889; Oudemans, 1904; Starkoff & Starkoff, 1950; Volgin, 1987;) – Acaropsellina docta, Apodicheles heteropalpus, Cheletogenes ornatus, Cheletomimus (Ch.) berlesei, Cheyletiella parasitivorax, Cheletomorpha lepidopterorum (= Ch. venustissima), Cheyletus eruditus, Ch. schneideri, Ornithocheyletia pinguis
Sicily (Berlese, 1913) – Neoeucheyla loricata
Latvia (Petrova & Salmane, 2000; Petrova, Salmane & Čudare, 2004) – Cheletomorpha lepidopterorum
Malta (Pfliegler et al., 2017) – Cheletomorpha lepidopterorum
Moldavia (Volgin, 1961) – Cheletacarus raptor
Norway (Thor, 1912; Austad, 1968; Dotterud, Korsgaard & Falk, 1995) – Cheyletiella parasitivorax, Cheletomorpha lepidopterorum (= Ch. venustissima), Cheyletus eruditus
Poland (Willmann, 1949; Wegner & Przyborowski, 1958; Haitlinger, 1982, 2000; Kivganov & Bochkov, 1994; Haitlinger & Jankowska, 2005; Solarz, 1989; Skoracki, Bochkov & Sikora, 2004) – Acaropsis docta, Cheletophyes aurorae, Cheletopsis daberti, Cheyletia papillifera, Cheyletus eruditus, Ch. trouessarti, Eucheyletia flabellifera, Ornithocheyletia dubinini
Portugal (cont.) (Baker, 1949, 1958, Bochkov & OConnor, 2004) – Chelacheles strabismus, Cheletomorpha lepidopterorum, Cheyletus eruditus
Madeira (Willmann, 1939) – Cheyletus eruditus
Russia (all, continent.)(Rohdendorf, 1940; Volgin, 1962, 1964, 1966, 1969; Fain, 1972; Hughes, 1976; Bochkov, Mironov & Gorgol, 1994; Bochkov & Mironov, 1999; Fain & Bochkov, 2001; Chernichko & Kivganov, 2013; Bochkov & Abramov, 2016; Diumina & Abramov, 2018; Khaustov et al., 2019; Khaustov, 2021) – Acaropsella kulagini, Acaropsellina sollers, Chelacheles apanaskevichi, Cheletacarus raptor, Cheletogenes ornatus, Cheletomimus berlesei, Ch. lindquisti, Ch, recki, Cheletomorpha lepidopterorum, Cheletopsis impavida, Cheyletia aradiphila, Cheyletiella blakei, Ch. parasitivorax, Cheyletus eruditus, Ch. malaccensis (= polymorphus), Ch. rashtiensis, Ch. trouessarti (= truculentus Volgin), Ch. volgini, Cunliffella tuberculicoxa, Eucheyletia eoa, E. harpyia, E. sibirica, Eucheyletiella daurica, E. pallasius, E. takahasii, Eutogenes frater, Mexecheles virginiensis, Muricheyla sicista, Neoacaropsis granulatus, Neocheyletiella microrhyncha, Ornithocheyletia phylloscopi, Paracheyletia pyriformis
Crimea (Volgin, 1963a; Kuznetsov, 1977; Bochkov, Khaustov & Kuznetsov, 1999; Fain & Bochkov, 2018) – Chelacheles alexandrinus, Cheletopsis impavida, Cheyletus cacahuamilpensis, Eucheyletia flabellifera (= taurica), Mictrocheyla granigera, Paracheyletia volgini, Prosochela villosa
Slovakia (Žďárková, 1979) – see Czechoslovakia
Spain (cont.) (Mihelcic, 1958; Zapatero Ramos et al., 1982; Cordero del Campillo et al., 1994) – Cheletonella vespertilionis, Cheyletus eruditus
Sweden (Boström et al., 1997) – Cheyletidae (Gotland I.)
Switzerland (Schweizer, 1922, 1951; Schweizer & Bader, 1963; Mumcuoglu, 1976) – Cheletomorpha lepidopterorum, Cheyletus eruditus, Ch. trouessarti, Eucheyletia flabellifera
Turkey (all) (Özer et al., 1986, 1989; Özkan, Ayyıldız & Soysal. 1988; Koç & Ayyıldız, 1995, 1996; Koç, 1998; Fain & Bochkov, 2001; Dogan & Ayyıldız, 2002, 2004; Bayram & Çobanoğlu, 2006; Bilal, 2012; Akpınar, Çobanoğlu & Öğreten, 2017; Çobanoğlu, Cilbircioğlu & Zabludovskaya, 2020; Doğan S. et al., 2019; Doğan, Doğan & Erman, 2019) – Acaropsella volgini, Bothrocheyla pavlovskyi, Cheletomimus (Hemicheyletia) bregetovae, Ch. (H.) wellsi, Cheletomorpha lepidopterorum, Cheyletus eruditus, Ch. malaccensis, Eutogenes frater, Hypopicheyla elongata, Microcheyla granifera (= ozkani), Ornithocheyletia hallae
Ukraine (Volgin, 1969; Vysotskaja [= Vysotzkaja], 1974, 1977; Kuznetsov, 1977; Kivganov & Bochkov, 1994; Bochkov & Khaustov, 1999; Bochkov, Khaustov & Kuznetsov, 1999; Fain & Bochkov, 2001; Chernichko & Kivganov, 2013) – Bothrocheyla pavlovskyi, Chelacaropsis moorei, Chelacheles alexandrinus, Ch. bacchusi, Cheletopsis charadrii, Ch. daberti, Ch. ferruginea, Cheyletus cacahuamilpensis [syn. baloghi], Ch. eruditus, Ch. trux, Eucheyletia bishoppi [= asiatica], E. bothrophila, E. taurica, Microcheyla granifera, Nodele superba, Paracheyletiella volgini, Prosocheyla villosa
“Yugoslavia” (Wikerhauser & Pinter, 1987) – Cheyletiella blakei
AFRICA
Algeria (Louadi & Robaux, 1992; Fain & Bochkov, 2001; Fain, Bochkov & Corpuz-Raros, 2002; Fain, 2003) – Chelacheles strabismus (= algericus), Cheletogenes ornatus, Cheletomimus (Hemicheyletia) bakeri, Cheletonella vespertilionis, Cheyletus cacahuamilpensis, Ch. malaccensis, Ch. trux (sub tenuipilis), Eutogenes frater, Neoeucheyla bochkovi, Paracheyletia pyriformis
Angola (Fain & Bochkov, 2001, 2002) – Cheletomimus (Ch.) berlesei, Cheletophyes eckerti, Nihelia curvidens, Sciurocheyla angolensis
Botswana (Lawrence, 1959) – Neocheyletiella megaphallos
Burkina Faso (Bochkov, Mironov & OConnor, 2006) – Galagocheles lawrencei
Burundi (Fain & Bochkov, 2001; Bochkov & OConnor, 2004) – Chelacaropsis apus, Sciurocheyla funisciurus, Bakericheyla benoiti, Cheletophyes torridae, Sciurocheyla [Smileycheles] camerounensis
Cameroon (Fain, Bochkov & Corpuz-Raros, 2002) – Cheletomimus wellsina
Congo DR (Cunliffe, 1962, Fain, 1972; Fain & Bochkov, 2001; Fain, Bochkov & Corpuz-Raros, 2002; Klimov, Bochkov & OConnor, 2006; Bochkov, Mironov & OConnor, 2006) – Acaropsella kinshasensis, Bak deleoni, Cheletomimus (Hemicheyletia) bakeri, Ch. (H.) congensis, Cheletomorpha bakeri, Cheletophyes eckerti, Ch. mbomba, Cheyletiella parasitivorax (= katangae), Cheyletus funisciuri, Ch. kivuensis, Galagocheles lawrencei, G. lemuricola, Grallacheles bakeri, Ker afrotropicalis, Samsinakia volgini, Sciurocheyla congolensis
Egypt (El Badry & Zaher, 1960; Zaher & El Badry, 1962; Zaher & Soliman, 1966, 1967; Wafa & Soliman, 1968; Elbadry, 1969; Rasmi et al., 1972; Yousef & Issa, 1972; Attiah, 1973; El Tawab & Yousef, 1975; Soliman, 1975, 1977; Yousef, 1975, 1979; Hassan & Gomaa, 1982; Yousef, Zaher & Kandil, 1979; Saleh et al., 1985, 1986; Fain & Bochkov, 2001; Negm & Mesbah, 2014; Metwally et al., 2016) – Acaropsella aegyptiaca, Acaropsellina docta, Aegyptocheyla summersi, Chelacaropsis moorei, Ch. reticulata, Chelacheles alexandrinus, Ch. bakeri, Cheletogenes ornatus, Cheletomorpha lepidopterorum, Cheyletus egypticus, Ch. eruditus, Ch. malaccensis (incl. fortis), Eutogenes frater (= africanus), E. punctata, Ker bakeri, Neoeucheyla macrocorneus, N. ornata, Nodele calamondin [=simplex]; Genus incertae sedis: Aegyptocheyla; Sp. inquirenda: Cheletomimus minutus
Gabon (Bochkov & OConnor, 2003) – Picocheyletus tricholaemae
Kenya (Fain & Bochkov, 2001) – Kenyacheylus troglodytes, Chelacaropsis kenyensis
Madagascar (Samšinak, 1959; Volgin, 1965; Fain, 1982) – Cheyletus legendrei; Samsinakia theodoridis
Morocco (Cooreman, 1951; Fain & Lukoschus, 1981) – Cheletogenes ornatus, Cheyletus punctulatus
Mozambique (Rodriges, 1967a, 1967b, 1968; Bochkov & Ochoa, 2005) – Neocheyletiella ardeola
Nigeria (George et al., 1992) – Cheyletiella parasitivorax
Rwanda (Fain, 1972; Fain & Bochkov, 2001, 2002a; Fain, Bochkov & Corpuz-Raros, 2002; Bochkov, Mironov & OConnor, 2006) – Apodicheles apus, Bakericheyla transvaalica, Chelacaropsis apus, Ch. rwandana, Cheletomimus wellsina, Cheletomorpha bakeri, Ch. lepidopterorum, Cheyletus funisciuri, Ch. tanzaniensis, Galagocheles lawrencei, G. lemuricola, Ker afrotropicalis, Neocheyletiella lonchurae, N. megaphallos, N. microrhyncha, N. queleae,
Sainte-Helene (Summers & Price, 1970) –
Senegal (Gueye-N’Diaye & Fain, 1987) – Cheyletus malaccensis
Somalia (Fain, 1974) – Teinocheylus longissimus
South Africa (Lawrence, 1948, 1954; Meyer & Ryke, 1960; Fain, 1982; Meyer, Loots, van Pletzen, Engelbrecht & Walker, 1973; Meyer, 1988; Fain & Lukoschus, 1981; Olivier & Theron, 1988, 1989a, 1989b; Lindholm, Venter & Ueckermann, 1998; Bochkov & OConnor, 2004) – Apodicheles cypsiurus, Bakericheyla africana, B. faini, B. subquadrata, B. transvaalica, Chelacheles aigamuxa, Cheletacarus novemdentis, Cheyletiella parasitivorax, Cheyletus cacahuamilpensis, Ch. mafekingensis, Ch. gerbillicola, Ch. pseudomalaccensis, Eutogenes reticularis, Galagocheles lemuricola, Microcheyla paraparvula, M. squamosa, Neocheyletiella megaphallos, Nihelia curvidens, N. cynictis, Paramicrocheyla spinula, P. ungulina, Prosocheyla (Reckiana) hepburni
Tanzania (Vitzthum, 1920; Fain, 1972; Haitlinger, 2000; Fain & Bochkov, 2001; Bochkov & OConnor, 2004; Bochkov & Skoracki, 2011) – Cheletophyes aurorae, Cheyletus venator, Ch. tanzaniensis, Metachyletia ngaii, Sciurocheyla camerounensis
Tunisia (Fain, Gerrits & Lukoschus, 1982) – Teinocheylus gundii
Uganda (Lawrence, 1954) – Cheletomorpha bakeri, Cheyletus malaccensis (= ugandanus)
Zambia (Mwase & Baker, 2006) – Cheyletus spp.
Zimbabwe (Fain, 1979) – Bakericheyla transvaalica, Cheletoides chirunduensis
ASIA
Bangladesh (Asmat & Abdul Kader, 1990) – Cheyletus malaccensis
Burma (Myanmar)(Summer & Price, 1970) – Cheyletus malaccensis (= fortis), Prosocheyla
China (Volgin, 1963a; Shen, 1975, 1984, 1991; Lung-Shut, 1984; Whitaker, Shen, 1975, 1991; Wang, 1989; Baccus, Cleveland & Gu, 1990; Shen, 1991; Fang Lin, Qian Kaiming & Gao Cheng, 1991; Hu & Hou, 1992; Lin, 1997; Lin & Liu, 1994; Xia, Ye & Zhu, 1997a, 1997b; Xia & Zhu, 1997; Lin & Zhang, 1992, 1997; Lin, Zhang & Yang, 1997, 1998; Xia, Rong & Zhimin, 1997; Lin et al., 2000; Zhu et al., 2000; Fain & Bochkov, 2001; Xia, Liang, Zeng & Zhu, 2004; Xia, 2010) – Acaropsellina docta, A. nanchangensis, A. sollers, Bak faini, Caudacheles trigintaduae, Chelacaropsis moorei, Ch. bipanus, Cheletacarus raptor, Chelacheles bipanus, Ch. michalskii, Ch. striola, Cheletogenes meihuashanense, Ch. ornatus, Cheletomimus (Hemicheyletia) anarbora (= Hemicheyletia pusillifolium), Ch. bakeri, Ch. chui, Ch. omissus, Ch. wellsi, Ch. wellsinus, Cheletomorpha lepidopterorum, Cheyletiella parasitivorax (= dengi), Cheyletus carnifex, Ch. eruditus, Ch. malaccensis, Ch. trouessarti, Columbicheyla bicicri, C. macroflabellata, Eucheyletia bishoppi, E. flabellifera, E. harpyia, E. omissa, E. sinensis, E. variformis, Eutogenes quadrisetatus, Grallacheles bakeri, G. nanfengensis, Hoffmannita clavipes (= H. navicula), Ker palmatus, Lepidocheyla gracilis, Neoacaropsis granulatus, Oudemansicheyla denmarki, Prosocheyla sinensis, Zachvatkiniola reticulate. Sp. inquirendae: Cheletonella juglandis
India (Baker, 1949; Summers & Price, 1970; Fain, 1972; Nagar & Saxena, 1977; Nagar, 1979; gupta & Chattopadhyay, 1978; Smiley & Whitaker, 1981; Mathur & Mathur, 1981; gupta & paul, 1985, 1992; Putatunda & Kapil, 1988; Gupta, 1991, 2002; Bochkov & Mironov, 1998; Ramaraju & Mohanasundaram, 1999; Fain & Bochkov, 2001, 2002a; putatunda, Kumar & Banerjee, 2004; Bochkov & Ochoa, 2005; Podder, Gupta & Saha, 2006; Bochkov & Otto, 2010) – Chelacaropsis moorei, Chelacheles indra, Cheletophyes deodikari, Ch. eckerti, Ch. harnaji, Ch. haryanaensis, Ch. Indiacus (= xylocopae), Ch. newtoni, Ch. orientalis, Ch. ruttneri, Ch. shendei, Cheyletus eruditus, Grallacheles indicus, Lepidocheyla caucasica [= hissariensis], Mexecheles shiva, M. virginiensis, Microcheyla bengalensis, Neocheyletiella athene, N. siva, Nihelia calcarata, Prosocheyla (P.) traubi, Samsinakia carabae, S. gonocephalum, S. trilobitus; Neoeucheyla ploceus (Sp. inquir.)
Andaman and Nicobar Is. (Gupta & Ghosh, 1980) – Chelacaropsis moorei
Indonesia (Berlese, 1913; Domrow, 1960; 1964: 17; Fain, 1972; Thewke, 1980; Haines, 1988; Fain & Bochkov, 2001; Bochkov & Otto, 2010) – Bak indonesiensis, Chelonotus selenirhynchus [= Ch. oudemansi = Ch. ewingi], Cheletophyes apicola, Eutogenes quadrisetatus, Hylopecheyla bunguranensis, Neocheyletiella amandavae, Nodele mu, Tutacheyla buruensis
Iran (Vercammen-Grandjean & Rak, 1968; Rak, 1972, 1973; Sepasgozarian, 1978; Kamali, 1990; Faraji & Kamali, 1993; Yazdani & Ebrahimi, 1993; Bochkov & Malikov, 1996; Fathipour, Kamali & Ostovan, 1999; Ardeshir et al., 2000; Fain & Ardeshir, 2000; Bochkov, Fain & Ardeshir, 2001; Fain & Bochkov, 2001; Kamali, Ostovan & Atamehr, 2001; Merhnejad & Ueckermann, 2001; Bochkov, Hakimitabar & Saboori, 2005; Ostovan & Kamali, 1997; Sahraeian et al., 2006; Ardeshir & Nematollahi, 2008; Namaghi, 2010; Beyzavi & Ostovan, 2011; Doğan, Jalaeian & Kamali, 2011; Modarres, 2012; Pactinat-Saeij, Bagheri & Skvarla, 2017; Ardeshir, 2017; Salarzehi, Hajizadeh, Hakimitabar & Ueckermann, 2018; Salarzehi, Hajizadeh & Ueckermann, 2018, 2019) – Acaropsella kulagini, A. volgini, Acaropsellina docta, A. sollers, Bak iranica, Caudacheles khayae, C. lieni, Chelacheles michalskii, Ch. strabismus, Cheletogenes ornatus, Ch. scaber, Cheletomimus (Ch.) berlesei, Ch. binus, Ch. (Hemicheyletia) bakeri, Ch. (H.) congensis, Ch. (H.) vescus, Ch. (H.) wellsi, Ch. parasitivorax, Ch. yasguri, Cheyletus bidentatus, Ch. cacahuamilpensis, Ch. carnifex (= Ch. acarophagus = allactaga = zacheri), Ch. eruditus, Cheyletus kuznetzovi, Ch. malayensis, Ch. malaccensis, Ch. rashtiensis, Ch. trouessarti, Cheletomorpha lepidopterorum, Cheletonella iraniensis, C. vespertilionis, Cunliffella bulgarica, C. variegata, Eucheyletiella faini, E. flabellifera, Eutogenes frater, Hypopicheyla elongata, H. mirabilis, Lepidocheyla gracilis, Microcheyla parvula, Neoeucheyla iranica, Nodele calamondin, Paraheyletia pyriformis, Samsinakia trilobites, Zachvatkiniola reticulata
Iraq (Rizk et al., 1979) – Acaropsellina docta, A. sollers
Israel and Palestine (Shulov, 1957; Cwilich & Hadani, 1965; Costa, 1961: 259; 1967: 109; Gerson, 1967, 1968; Feldman-Musham et al., 1985; Fain & Bochkov, 2001 Fain, Bochkov & Corpuz-Raros, 2002) – Acaropsella volgini, Caudacheles khayae, Cheletogenes ornatus, Cheletomimus (Ch.) berlesei, Cheletomimus (Hemicheyletia) bakeri, Cheletomorpha lepidopterorum, Ch. tatami, Cheyletiella sp., Cheyletus cacahuamilpensis, Ch. eruditus, Ch. malaccensis, Ch. trux (= tenuipilis), Eutogenes citri, Lepidocheyla gracilis, Neoacaropsis volgini
Japan (Sasa & Kono, 1951; Tanaka, 1953; Hara, 1955; Hara & Hanada, 1960; Ehara, 1962; Shiba, 1969a, 1969b; Summers & Price, 1970; McClure, Ratanaworadhan et al. 1973; Kumada et al., 1978; Razzaq, Shiraishi, Manabe & Ohbayashi, 2001) – Chelacaropsis moorei, Cheletomorpha lepidopterorum [Cheletophyes tatami Hara], Cheyletus eruditus, Ch. malaccensis (= fortis), Ch. rarus, Cheletomimus (Hemicheyletia) bakeri, Ch. (H.) wellsi, Eucheyla favosa, Eucheyletiella takahashii, Eutogenes narashinoensis, Neocheyletiella sp., Zachvatkiniola reticulata
Kazakhstan (Kuzin, 1940; Volgin, 1955, 1963; Bochkov & Mironov, 1999) – Acaropsellina sollers, Cheyletus trouessarti [sub Ch. praedabundus Kuzin], [Cheyletus rabiosus], Eucheyletia asiatica, E. pavlovskyi, Eucheyletiella pallasianus
Korea (Nord and South) (McClure, Eliott, Ratanaworadhan et al. 1973; Shin, 1996; Sang-Hun Kim et al., 2008) – Cheyletiella parasitivorax, Ch. yasguri, Neocheyletiella sp.
Kyrghyzstan (Volgin, 1969; Dubinina & Bochkov, 1989; Chirov & Mironov, 1998; Chirov & Bochkov, 1998) – Cheyletus carnifex, Cunliffella tuberculicoxa, Eucheyletiella ochotonae, Ornithocheyletia mironovi
Malaysia (continent.)(Oudemans, 1903; Nadchatram & Domrow, 1966; Shiba, 1976; Yunus & Ho, 1980; Fain & Nadchatram, 1980; Fain, Lukoschus & Nadchatram, 1980; Ehara & Abdul Ghani, 1988; Mariana et al., 2000: 714; Fain & Bochkov, 2001) – Cheletogenes monosetosus, Ch. ornatus, Cheletomimus (Hemicheyletia) anarbora, Ch. (H.) transversus (= Hemicheyletia morii), Ch. (H.) tropicus, Ch. (H.) wellsi, Ch. (H.) wellsinus, Cheletomorpha lepidopterorum, Cheletophyes apicola, Ch. clavipilis, Ch. vespae, Cheletonella vespertilionis, Chelonotus selenirhynchus, Cheyletus malaccensis (= Ch. fortis), Ch. malayensis, Ch. trouessarti, Criokeron quintus, Eutogenes onoi, Grallacheles bakeri, Hylopecheyla malayi, Ker bakeri
Sarawak & Sabah (Domrow, 1964; Ide & Kethley, 1977) – Chelonotus selenirhynchus, Thewkachela ratufi
Mongolia (Fain & Lukoschus, 1981) – Cheyletus carnifex [syn. allactaga]
Nepal (Summers & Price, 1970) – Eucheyletia sinensis
“New Guinea” (Summers & Price, 1970)
Pakistan (khan, 1970; qayyum & chaudhri, 1977a, 1977b, 1977c, 1979a, 1979b; Rasool & Chaudhri, 1979; Rasool, Chaudhri & Akbar, 1980; Chaudhri & Akbar, 1985; Akbar, Rahi & Chaudhri, 1988; Aheer, Akbar & Chaudhri, 1988, 1991, 1992, 1993, 1994, 1998; Baloch, 1991; Akbar, Aheer & Ishtiaq, 1993; Aheer et al., 1997; Farooq, 2000; Farooq, Akbar & Qureshi, 2000; Haq & Zia, 2001a, 2001b;, Haq et al., 2004; Fain, Bochkov & Corpuz-Raros, 2002; Bochkov & OConnor, 2003; Afzal, Ali & Akbar, 2003; Kamran, Khan & Nazir, 2005; Haq & Afzal, 2007; Akbar, Jahan & Mughal, 2008; Halliday, Kamran & Bashir, 2018) – Acaropsella nobilis, A. shaziai, A. walii, Acaropsellina campusensis, A. clamo, A. opsis, A. orbis, A. platessa, A. porta, A. shorkotiensis, A. subitus, A. venustus, A. vitrus, Chelacheles humilis, Cheletogenes aceriai, Ch. carinatus, Ch. dissitus, Ch. iconis, Ch. petigenis, Ch. sagacis, Ch. scaber, Ch. vulgatus, Cheletomimus (Hemicheyletia) bakeri, Ch. (H.) tanosus, Ch. (H.) vescus, Cheletomorpha dolosus, Ch. lepidopterorum, Ch. obrutus, Ch. opacus, Ch. tenerum, Cheletonella daddaciensis, Ch. emersus, Cheyletus crassus, Ch. eruditus, Ch. malaccensis (= ayyazi, = baridos = infensus = phantosis = wahndoensis), Ch. vivatus, Ker acidalia, Neocheyletiella aegithali; Sp. inquirendae: Cheletomimus cambio, Ch. cantor, Ch. larmae, Ch. telma, Ch. trema, Ch. zamia, Cheletomorpha obrutus
Philippines (Summers & Price, 1970; Corpuz – Raros, 1972, 1980, 1988a, 1988b, 1998, 2000; Corpuz – Raros & Sotto, 1977; Malaban & Corpuz-Raros, 1998; Fain, Bochkov & Corpuz-Raros, 2002; Bochkov & OConnor, 2003, 2004; Corpuz-Raros L. A., J.C.B. Naredo. 2016) – Acaropsella filipina, Acaropsellina levis, A. philippinensis, Alliea prasadi, Apodicheles collocalia, Bak faini, B. furcatus, B. gersoni, B. payatus, B. truncatus, Chelacaropsis terrestris, Chelacheles baiwanganae, Ch. robustus, Cheyletus ascutatus, Ch. batomysi, Ch. malaccensis (= fortis), Ch. malayensis, Cheletogenes ornatus, Cheletomimus (Ch.) daltoniensis, Cheletomimus (Hemicheyletia) athenae, Ch. (H.) bakeri, Ch. (H.) leytensis, Ch. (H.) transversa, Ch. (H.) uichancoi, ?Ch. (H.) wellsi, Ch. (H.) wellsinus, Cheletomimus (Philippicheyla) philipina, Cheletomorpha dolosus, Ch. tenerum, Cunliffella duo, C. maysa, Eucheyletia nindota, Eutogenes makilingensis, Hoffmannita rimandoi, Ker mercedesae, K. pintoriensis, Neochelacheles corpuzrarosae, N. mendicus, Pavlovskicheila philippicana, Samsinakia pagongae. Species inquirendae: Hemicheyletia lanceolata
Saudi Arabia (soliman & Al-Yousif, 1980; el-Bahrawy & al-Dakhil, 1993; Rostom, 1993) – Acaropsella ? volgini, Acaropsellina sollers, Cheyletus eruditus, Ch. malaccensis
Singapore (Fain, Bochkov & Corpuz-Raros, 2002) – Cheletomimus (Hemicheyletia) asiaticus
Sri Lanka (Parameswaran Pillai, 1957) – Cheyletus munroi [var. bituberculatus]
Tadjikistan (Volgin, 1963) – Cheyletus carnifex, Ch. malaccensis (= polymorphus), Lepidocheyla gracilis
Taiwan (Sugimoto, 1942; Summers & Price, 1970; tjying I. – S. (= Tseng), 1971, 1972; Tseng, 1973, 1977, 1982, 1984) – Acaropsella kinshasensis (= konoi), Bak micidus, Caudacheles lieni, Chelacheles lanceolatus, Cheletogenes monosetosus, Cheletomorpha lepidopterorum, Cheyletiella strandtmanni, Cheletomimus anarbora, Ch. (H.) bakeri, Ch. chui, Ch. omissus, Ch. (H.) wellsinus, Cheletonella pilosa, Cheyletus eruditus, Ch. malaccensis, Ch. trouessarti, Eucheyletia sinensis, Oudemansicheyla denmarki, Zachvatkiniola reticulata
Thailand (vaivanijkul, 1979; Uchikawa & Suzuki, 1980; Fain & Lukoschus, 1985; Lekprayoon & Smiley, 1986; Gerson & Fain, 1991; Bochkov & OConnor, 2004; Bochkov & Klimov, 2005; Fuangarworn & Lekprayoon, 2010) – Bak furcatus, Chelacaropsis moorei, Criokeron quintus, C. thailandicus, Chelacheles baiwanganae, Cheletophyes decorus, Cheyletus eruditus, Dubininiola boonkongae, Mexecheles thailandensis, Samsinakia charanasriae, Sciurocheyla squamosa
Turkmenia (Volgin, 1969, 1978) – Acaropsellina sollers, Dubininiola polylepis, Cheletomimus (Hemicheyletia) asiaticus, Lepidocheyla gracilis
Uzbekistan (Kuzin, 1940; Volgin, 1955; Barilo, 1985, 1986a, 1986b, 1989) – Acaropsellina sollers, Chelacheles [Chelachecaropsis] stigmaeoides, Cheyletus carnifex, Ch. morinus, Hypopicheyla mirabilis, Ker caeterus, Neoeucheyla minuta, Pavlovskicheyla semenovi
Vietnam (Bochkov & Klimov, 2005) – Cheletophyes decorus
NORTH AMERICA
Canada (Banks, 1909; Ayalew & Vaillancourt, 1976; Thewke & Enns, 1979; Bochkov & Galloway, 2001; Fain, Bochkov & Corpuz-Raros, 2002) – Cheletomimus (Hemicheyletia) bakeri, Ch. (H.) lindquisti, Cheletomorpha lepidopterorum, Cheyletiella yasguri, Mexecheles virginiensis, Neocheyletiella microrhyncha, Ornithocheyletia canadensis, O. prognae, O. turdi
Mexico (Baker, 1949; De Leon, 1962; Pelaez, 1962; Fain, 1972; Thewke & Enns, 1979; Uchikawa & Suzuki, 1979; Thewke, 1980; Atyeo, Kethley & Pérez, 1984; Hoffmann, Cervantes & Morales-Malacara, 1994; Estébanes – González, 1997; Fain & Bochkov, 2001; Bochkov & Klimov, 2004, 2005) – Anthribicheyla bocki, Cheletogenes ornatus, Cheletomimus berlesei, Ch. cordovensis, Ch. scutellatus, Cheyletiella parasitivorax, Cheyletiella [syn. Bicheyletiella] romerolagi [= mexicana], Cheyletus cacahuamilpensis, Ch. eruditus, Chiapacheylus edentatus, Hoffmannita mexicana; Metacheyletia longisetosa, Mexecheles hawaiiensis, M. marshalli, Nodele (Aztecocheyletus) conquistador, Samsinakia tilae
USA (Banks, 1902, 1904, 1906; Ewing, 1909a, 1909b, 1912; Miller, 1925; Cooper, 1946; Baker, 1949; Yunker, 1960, 1961; De Leon, 1962; Muma, 1964; Vercammen-Grandjean & Rak, 1968; Summers & Price, 1970; Smiley & Moser, 1970, 1975; Hughes, 1976; Smiley, 1970, 1996; Moser & Roton, 1971; Smiley & Williams, 1972; Whitaker, 1973; Whitaker & Wilson, 1974; Thewke & Enns, 1972, 1974, 1975; Keh, 1973, 1975; Delfinado & Khaing – Fields, 1976; Thefke, 1974, 1977; Whitaker, Pascal & Mumford, 1979; Whitaker, Tieben & Maser, 1980; Smiley & Whitaker, 1981; Whitaker & Maser, 1985; Philips & Dindal, 1990; Cudmore, 1986; Fain & Bochkov, 2001; Fain, Bochkov & Corpuz-Raros, 2002; Bochkov & OConnor, 2003, 2004; Whitaker et al., 2007; Bochkov & Otto, 2010; Xia, Klompen & Childers, 2011) – Acaropsella kulagini, Acaropsellina sollers, Alliea laruei, Anthribicheyla bocki, Atarsacheylus vichii, Bak deleoni, B. ozarkensis, B. sanctaehelenae, Bothrocheyla typhosa, Camincheyletus glaucomys, Chelacheles bipanus, Ch. peritremaculatus, Ch. seminole, Ch. strabismus, Ch. temoak, Cheletacarus gryphus, Cheletogenes ornatus, Cheletomimus (Ch.) berlesei, Ch. duosetosus, Cheletomimus (Hemicheyletia) bakeri, Ch.(H.) newyorkensis, Ch. (H.) rostella, Cheletonella vespertilionis, Ch. hoffmannae, Cheletopsis thalasseus, Cheyletiella blakei, Ch. parasitivorax (= furmani = americana), Ch. yasguri, Cheyletus cacahuamilpensis (nr), Ch. carnifex (= aversor), Ch. eruditus, Ch. linsdalei, Ch. malaccensis, Ch. trouessarti, Ch. trux (= hendersoni), Cunliffella whartoni, Eucheyletia bishoppi (= E. oregonensis), E. hardyi, Eucheyletiella johnstoni, Eutogenes pinicola, E. vicinus, Grallacheles bakeri, Ker palmatus, Lanceacheyla whartoni, Mexecheles impolitus, M. panneus, M. virginiensis, Modele calamondin, M. coccineae, Neochelacheles messersmithi, Neocheyletiella roweri, N. smallwoodae, Ornithocheyletia gersoni, O. hallae hallae, O. lawrenceae, Oudemansicheyla denmarki, Paracaropsis travisi, Paracheyletia pyriformis (= Cheyletus longipalpus), Pavlovskicheyla platydemae, Prosocheyla (P.) acantha, P. (P.) oaklandia, P. (Reckiana) buckneri
Alaska (Volgin, 1963) – Eucheyletia bakeri
Hawaiian Is. (Baker, 1949; Garrett & Haramoto, 1967; Haramoto, 1969; Summers & Price (1970); Goff, 1982, 1983, 1987; Paxot & Goff, 1985; Fain, Bochkov & Corpuz-Raros, 2002) – Bak deleoni, B. elongatus, Chelacheles strabismus, Cheletogenes ornatus, Cheletomimus (Ch.) berlesei, Ch. (Ch.) duosetosus, Ch. (Hemicheyletia) bakeri, Ch. (H.) bregetovae, Ch. (H.) gracilis, Ch. (H.) granulae, Ch. (H.) kureatollensis, Ch. (H.) wellsi, Cheyletus eruditus, Ch. malaccensis (= Ch. fortis), Ch. malayensis, Cunliffella panamensis, Grallacheles bakeri, Ker palmatus, Mexecheles hawaiiensis, Nodele philippinensis; Incertae sedis: Cheletomimus citrosinensis
SOUTH and CENTRAL AMERICA
Argentina (Brandetti & Bischoff-de-Alzuet, 1981; Mosquera, 1989) – Cheletomimus (Hemicheyletia) wellsi, Eucheyletia hardyi, Ornithocheyletia boeroi
Brazil (Flechtmann, 1968, 1971; Reis & Paschoal, 1968; Arruda et al., 1969; Amaral, 1971; Fain, 1972; Rosa & Flechtmann, 1979; Whitaker & Dietz, 1987; Bagio et al., 1988a, 1988b; Feres & Flechtmann, 1995; Ezequiel et al., 2001, 2003; Buosi et al., 2006; Valim & Gazeta, 2007) – Bak ligyscutatus, Cheletogenes ornatus, Cheletomimus duosetosus, Ch. wellsi, Cheletomorpha lepidopterorum, Cheletonella vespertilionis (= caucasica), Cheyletiella parasitivorax, Cheyletus eruditus, Ch. malaccensis (= fortis), Eutogenes sp., Ker bakeri, Mexecheles hawaiiensis, Ornithocheyletia aitkeni, O. hallae
Chile (contin.)(Baker, 1949; Alcaino & Tagle, 1966; Alcaino & Calderon, 1976; Artigas & Casanueva, 1983; Casanueva & Artigas, 1985; Peredo et al., 1994; Fain & Bochkov, 2001) – Cheletogenes ornatus, Cheletomorpha lepidopterorum, Cheyletiella parasitivorax, Ch. yasguri, Cheletomimus (Hemicheyletia) wellsi, Cheyletus eruditus
Columbia (Oudemans, 1903; Baker, 1949) – Cheyletus eruditus, Ch. intrepidus
Costa Rica – Cheletophyes panamensis
Cuba (de la Cruz, 1971; Cuervo, Dusbábek, de la Cruz & Abreu, 1983; Socarras, 1987; Torre Santana, 2015) – Chelacheles michalskii, Cheyletiella parasitivorax, Cheletogenes ornatus, Cheletomimus (Ch.) berlesei, Cheletomimus anarbora, Cheletomimus (Hemicheyletia) wellsi, Ch. wellsinus, Cheletomorpha lepidopterorum, Cheyletus carnifex, Ch. eruditus, Ch. malaccensis, Ch. malayensis, Chiapacheylus edentatus, Grallacheles bakeri, Mexecheles hawaiiensis
Ecuador (contin.)
Galapagos Is. (Summers & Price, 1970; Fain & Bochkov, 2001; Fain, Bochkov & Corpuz-Raros, 2002) – Bak gersoni, B. micidus, Cheletomimus (Hemicheyletia) bakeri, Ch. (H.) darwinia, Ch. (H.) serrula
El Salvador (Summers & Price, 1970) – Cheletomimus (Hemicheyletia) granulus, Laeliocheyletia teretis
Guatemala (Fain, Bochkov & Corpuz-Raros, 2002) – Cheletomimus wellsina
Honduras – Mexecheles virginiensis
Jamaica (Corpuz-Raros & Sotto, 1977) – Cheletomimus (Hemicheyletia) wellsi
Netherlands-Antilles (Fain & Bochkov, 2001) – Hoffmannita gersoni
Panama (Thewke, 1980; Klompen, Mendez & Lukoschus, 1984; Fain, Bochkov & Corpuz-Raros, 2002) – Anthribicheyla bocki, Cheletomimus (Hemicheyletia) bakeri, Ch. (H.) wellsinus, Cheletophyes panamensis, Cunliffella panamensis, Mexecheles hawaiiensis
Peru (Caceres & Fain, 1975, 1978a, 1978b; Fain & Bochkov, 2001) – Cheletomorpha lepidopterorum, Cheyletus cacahuamilpensis, Ch. malaccensis, Ch. trouessarti, Pavlovskycheyla philippicana
Porto Rico (Banks, 1914)
Suriname (Fain & Bochkov, 2001) – Grallacheles bakeri
Trinidad (De Leon, 1967) – Cheletomimus anarbora, Ch. wellsina
Venezuela (Fain & Bochkov, 2001) – Cheletogenes ornatus
OCEANIA
Australia (continent)(Womersley, 1941, 1942a, 1942b, 1956; Champ, 1965, 1966; Shepherd & Edmonds, 1977; Smiley, 1984; Koch, 1986; Domrow, 1991; Gerson, 1994; Fain & Bochkov, 2000; 2002b; Bochkov & Ochoa, 2005; Bochkov & Otto, 2010) – Chelacheles thomasi, Cheletacarus rugosus, Cheletogenes ornatus, Ch. waitei, Cheletomimus (Hemicheyletia) bakeri, Cheletomimus (Philippicheyla) notelaeae, Cheletomorpha lepidopterorum, Chelacaropsis [syn. Cheletonata] milesi, Cheletonella vespertilionis, Cheletophanes rugosus, Cheyletiella blakei, Ch. parasitivorax, Ch. yasguri, Cheyletus eruditus, Ch. malaccensis, Eucheyletia womersleyi, Granulocheyletus corpuzrarosae, Mexecheles virginiensis, Neocheyletiella ardeola, N. artami, Oconnoricheylus chimaera, Oudemansicheyla denmarki, Ou. coprosomae, Ornithocheyletia lawrenceae, O. lichmerae, O. lonchurae, Prosocheyla traubi, Samsinakia volgini
Tasmania (Domrow, 1991) – Cheyletiella blakei
Papua New Guinea (Oudemans, 1904; Bochkov & Klimov, 2005) – Eucheletopsis major, Cheletophyes occisor
French Polynesia (Tahiti)(Cunliffe, 1962; Nishida, Version 2009) – Grallacheles bakeri (= Paracheyletia woolfordi), Zachvatkiniola reticulata
Marianne Is. (Oudemans, 1903a) – Cheyletus malaccensis (= vorax), Ch. rapax
New Guinea (Fain & Bochkov, 2001) – Cheletomorpha lepidopterorum
New Zealand (Womersley, 1942a; Lamb, 1952; Bull, 1953; Davies & Kyle, 1969; Thewke & Enns, 1976; Gerson, 1994; Fain, 2003; Bochkov & Otto, 2010) – Cheletomimus crowei, Cheyletiella parasitivorax, Granulocheyletus gallowayi, Oudemansicheyla coprosomae
Vanuatu (Fain, Bochkov & Corpuz-Raros, 2002) – Cheletomimus (Hemicheyletia) uichancoi
Arthropoda
Crustacea
Decapoda
Fam. Coenobitidae
Coenobita clypeata – Hoffmannita gersoni
Arachnida
Scorpiones
Fam. Buthidae
Centruroides flavopictus – Hoffmannita mexicana
Insecta
Coleoptera
Fam. Scolytidae
Dendroctonus frontalis – Mexecheles impolitus, M. virginiensis, Prosocheyla acanthus
Fam. Tenebrionidae
Blastinus sp. – Samsinakia tilae
Bolitotherus cornutus – Neochelacheles messersmithi
Gonocephalum simplex – Samsinakia gonocephalum, G. volgini
Platydema – Pavlovskicheyla platydemae
Selinus abacoides – Samsinakia theodoridis
Fam. Bostrychidae
Dinoderus minutus – Nodele mu
Sinoxylon crassum – Chelacheles baiwanganae
Xyloperthoides nidipennis – Chelacheles aigamuxa
Heteroptera
Fam. Aradidae
A. aterrimus – Cunliffella tuberculicoxa, Hypopicheyla elongata
A. betulae – Hoffmanita clavipes
A. corticalis – Hoffmanita clavipes
A. crenaticollis – Hypopicheyla elongata
A. lugubris – Cheyletia aradiphila, Cunliffella tuberculicoxa, Hypopicheyla elongata
A. nemtschinovi – Cunliffella tuberculicoxa, Hypopicheyla elongata
Hymenoptera
Fam. Apidae
Diadasia sphaeralcearum – Chelacheles temoak
Xylocopa aruana – Cheletophyes occisor
Xylocopa (Megaxylocopa) frontalis – Cheletophyes panamensis
Xylocopa caffra – Cheletophyes vitzthumi
X. fenestata – Cheletophyes deodicari, Ch. newtoni, Ch. orientalis, Ch. ruttneri, Ch. shendei
X. pubescens – Cheletophyes harnaji
Xylocopa tranquebarica – Cheletophyes decorus
Xylocopa sp. – Cheletophyes xylocopae
Xylocopinae – Cheletophyes aurorae
Fam. Megachilidae
Azthecanthidium tenochtitlanicum – Nodele (Aztecocheyletus) conquistador
Fam. Vespidae
Vespa tropica – Cheletophyes vespae
Diptera
Fam. Hippoboscidae
Pseudolynchia canariensis – Ornithocheyletia hallae
Vertebrata
Reptilia
Sauria
Fam. Iguanidae
Sceloporus woodi – Paracaropsis travisi
Aves
Apodiformes
Fam. Apodidae
Apus affinis – Chelacaropsis apus
A. caffer – Apodicheles apus
Collocalia esculenta – Apodicheles collocalia
Fam. Hemiprocnidae
Hemiprocne (= Dendrochelidon) mystacea – Eucheletopsis major
Charadriiformes
Fam. Charadriidae
Charadrius dubius – Cheletopsis charadrii
Ch. hiaticula – Cheletopsis basilica
Fam. Rostratulidae
Rostratula semicollaris – Cheletopsis impavida
Fam. Scolopacidae
Actitis hypoleucos – Cheletopsis mariae
Calidris ferruginea – Cheletopsis impavida
C. melanotis – Cheletopsis basilica
C. minutus – Cheletopsis impavida
C. pusilla – Cheletopsis impavida
C. ruficollis – Cheletopsis impavida
C. temminckii – Cheletopsis daberti, Ch. impavida
Limnodromus griseus – Cheletopsis limnodromi
Micropalama himantopus – Cheletopsis daberti, Ch. impavida
Prosobonia cancellata – Cheletopsis prosobonialis
Tringa flavipes – Cheletopsis magnanima
T. glareola – Cheletopsis daberti
T. nebularia – Cheletopsis daberti
T. stagnatilis – Cheletopsis impavida
T. totanus – Cheletopsis basilica, Ch. norneri
Fam. Laridae
Subfam. Sterninae
Thalasseus [Sterna] sandvicensis – Cheletopsis thalasseus
Columbiformes
Fam. Columbidae
Columba livia – Ornithochyletia hallae, O. pinguis
Columbigallina sp. – Ornithochyletia columbigallinae, O. hallae
Streptopelia semitorquata – Neocheyletiella quleae
Coraciiformes
Fam. Meropidae
Melithophagus bullockoides – Bakericheyla (B.) transvaalica
M. pusillus – Bakericheyla transvaalica, B. subquadrata
Merops apiaster – Bakericheyla (B.) transvaalica
M. nubicoides – Bakericheyla (B.) transvaalica
M. persicus – Bakericheyla (B.) transvaalica
Merops [Melithophagus] bullocki – Bakericheyla benoiti
Cuculiformes
Fam. Cuculidae
Chrysococcyx caprius – Neocheyletiella (= Ornithocheyla) megaphallos
Ch. klaasi – Ornithochyletia orioli
Galliformes
Fam. Numididae
Numida meleagris meleagris – Metacheletoides numidae
N. meleagris intermedia – Metacheletoides akanyaruensis, M. numidae
Fam. Phasianidae
Francolinus natalensis – Ornithocheyletia francolini
Gruiformes
Fam. Aramidae
Aramus scolopaceus – Cheletosoma tyrannus
Musophagiformes
Fam. Musophagidae
Corythaxoides leucogaster – Metacheyletia ngaii
Crinifer piscator – Metacheletoides gisagarensis, M. crinifer
C. zonurus – Metacheletoides gisagarensis, M. crinifer
Passeriformes
Fam. Aegithalidae
Aegithalos concinnus – Neocheyletiella aegithali
Fam. Artamidae
Artamus cyanopterus – Neocheyletiella artami
Fam. Corvidae
Cyanocorax chrysops – Ornithochyletia boeroi
Fam. Estrildidae
Donacola castaneothorax – Ornithochyletia lonchurae
Erythrura prasina – Neocheyletiella vestergaardi
Estrilda erythronotos – Neocheyletiella megaphallos
Granatina ianthinogaster – Ornithochyletia granatina
Lonchura striata – Chelacaropsis moorei
Poephila acuticauda – Bakericheyla chanayi
P. gouldiae – Bakericheyla chanayi
Spermestes cucullatus – Chelacaropsis rwandana
Fam. Fringillidae
Carduelis carduelis – Bakericheyla chanayi
Chloris chloris – Cheyletia papillifera
Leucosticte australis – Neocheyletiella vestergaardi
Serinus mozambicus – Metacheyletia degenerata
Fam. Hirundinidae
Cecropis abyssinicus unitatis – Neocheyletiella microrhyncha
Delichon urbica – Neocheyletiella microrhyncha
Hirundo rustica – Neocheyletiella microrhyncha
Petrochelidon pyrchonota – Neocheyletiella microrhyncha
Progne subis – Neocheyletiella microrhyncha
Psalidoprocne albiceps – Neocheyletiella microrhyncha
Riparia riparia – Neocheyletiella microrhyncha
Fam. Monarchidae
Geiciphila indistincta – Ornithocheyletia lichmerae
Grallina cyanoleuca – Ornithocheyletia lichmerae
Fam. Motacillidae
Motacilla alba – Bakericheyla chanayi
Fam. Muscicapidae
Cossypha dichroa – Bakericheyla (B.) faini
Fam. Nectariniidae
Nectarinia jugularis – Neocheyletiella nectarinia
Fam. Oriolidae
Oriolus oriolus – Ornithochyletia orioli
Fam. Phylloscopidae
Phylloscopus trochilis – Ornithocheyletia phylloscopi
Fam. Pittidae
Pitta megarhyncha – Neocheyletiella pittae
Fam. Ploceidae
Foudia madagascariensis – Cheyletus legendrei
Pachyphantes superciliosa – Ornithochyletia orioli
Philetarius socius – Cheyletus mafekingensis
Plocepasser mahali – Cheyletus pseudomalaccensis
Ploceus philippinus – Microcheyla bengalensis, Neoeucheyla ploceus
Quelea quelea – Neocheyletiella queleae
Fam. Sittidae
Sitta pygmaea melanotis – Neocheyletiella rohweri
Fam. Sturnidae
Lamprocolius chloropterus – Ornithochyletia lamprocolius
Lamprocolius sp. – Ornithochyletia wauthyi
Sturnus vulgaris – Ornithocheyletia dubinini, O. pinguis
Fam. Timaliidae
Leiothrix lutea – Neocheyletiella media, N. alfortensis, Ornithocheyletia leiothris
Fam. Turdidae
Turdus fumigatus – Ornithocheyletia aitkeni
T. merula – Ornithocheyletia pinguis
Fam. Tyrannidae
Phoebe psychotrioides – Hemicheyletia cordovensi
Piciformes
Fam. Capitonidae
Lybius sp. – Ornithochyletia orioli
Tricholaema hirsuta flavipunctata – Picocheyletus tricholaemae
Fam. Indicatoridae
Indicator indicator – Ornithochyletia orioli
Fam. Leiothricidae
Garrulax leuolophus bicolor – Ornithocheyletia garrulax
Fam. Picidae
Dendrocopos pubescens – Ornithocheyletia gersoni
Picus viridis – Ornithocheyletia canadensis
Pelecaniformes
Fam. Ardaeidae
Ardeolla ralloides – Neocheyletiella ardeola
Psittaciformes
Fam. Psittacidae
Amazona finschi – Metacheyletia longisetosa
Ara sp. – Neocheyletiella avicola
Forpus passerinus – Ornithocheyletia argentinensis
Nandays nanday – Ornithocheyletia argentinensis
Psittacus erythacus – Ornithocheyletia psittaci psittaci
Pyrrhura cruentata – Cheyletus alacer
Fam. Psittaculidae
Agapornis fisheri – Neocheyletiella avicola
Fam. Psittrichasidae
Coracopsis nigra barkleyi – Cheyletus eruditus (= ferox)
Strigiformes
Fam. Strigidae
Athene brama – Neocheyletiella athene
Trogoniformes
Fam. Trogonidae
Harpactes ardens – Cheyletus philippinensis
Mammalia
Didelphiomorpha
Fam. Didelphidae
Didelphis virginiana – Cheyletus eruditus
Soricomorpha
Fam. Soricidae
Sorex fumeus – Eucheyletia bishoppi
S. pacificus – Eucheyletia bishoppi
S. trowbridgei – Eucheyletia bishoppi
Scandentia
Fam. Tupaiidae
Tupaia belangeri – Criokeron quintus
T. glis – Criokeron quintus, C. thailandicus, Hylopecheyla malayi
Chiroptera
Fam. Emballonuridae
Coleura gallarum – Cheyletus tanzaniensis
Fam. Vespertilionidae
Eptesicus fuscus – Cheletonella vespertilionis
Myotis velifer – Cheyletus cacahuamilpensis (nr)
Rodentia
Fam. Anomaluridae
(?) Zenkerella insignis – Sciurocheyla camerounensis
Fam. Ctenodactylidae
Ctenodactylus gundi – Teynocheylus gundii
C. vali – Teynocheylus gundii
Massoutiera mzabi – Teynocheylus gundii
Pectinator spekei – Teinocheylus longissimus
Fam. Muridae
Batomys salomonseni – Cheyletus ascutatus, Ch. batomysi
Menetes berdmorei – Cheyletus eruditus
Niviventer fulvescens – Eucheyletia sinensis
Rattus norvegicus – Cheyletus eruditus, Ch. trouessarti
R. humiliatus celsus – Eucheyletia sinensis
R. whiteheadi – Cheletophyes apicola
Subfam. Gerbillinae
Gerbillurus paeba – Cheyletus gerbillicola
Gerbillus pyramidum – Cheyletus punctulatus
Fam. Arvicolidae
Microtus agrestis – Cheyletus trouessarti
M. arvalis – Cheyletus malaccensis (= polymorphus)
M. guentheri – Cheyletus cacahuamilpensis
M. pinetorum – Eucheyletia bishoppi
Fam. Cricetidae
Neotoma cinerea – Eucheyletia bishoppi
N. floridana – Cheyletus malaccensis
N. magister – Cheyletus malaccensis
Peromyscus leucopus – Cheyletus eruditus
Thalpomys [Akodon] reinhardti – Cheyletus malaccensis
Fam. Sciuridae
Callosciurus sladeni midas – Cheyletus malaccensis
Citellus beecheyi – Cheyletus lindsdalei
Funisciurus carruthersi – Cheyletus funisciuri
Glaucomys sabrinus (nest) – Acaropsellina summersi, Camincheyletus glaucomys, Eucheyletia bishoppi (= oregonensis)
Heliosciurus undulatus – Sciurocheyla camerounensis
Hylopetes spadiceus – Hylopecheyla bunguranensis
Lariscus insignis – Chelonotus selenirhynchus
Nannosciurus surrutilus – Promuricheyla lukoschusi
Petinomys setosus – Cheyletus malaccensis
Ratufa affinis – Thewkachela ratufi
R. bicolor – Thewkachela ratufi
Rhinosciurus laticaudatus – Cheyletus pluridens
Spermophilus beldingi – Cheyletus lindsdalei
Sundasciurus hippurus – Chelonotus selenirhynchus
S. lowii – Chelonotus selenirhynchus
S. tenuis – Chelonotus selenirhynchus
Tamias striatus – Eucheyletia bishoppi
Lagomorpha
Fam. Leporidae
Lepus whytei – Cheyletiella parasitivorax (= katangae)
Lepus sp. – Cheyletiella strandtmanni
Oryctolagus cuniculus – Cheyletiella parasitivorax
Romerolagus diazi – Cheyletiella [=Bicheyletiella] romerolagi [= mexicana], Ch. parasitivorax
Sylvilagus bachmanni – Cheyletiella parasitivorax
S. floridanus – Cheyletiella parasitivorax
S. palustris paludicola – Cheyletiella parasitivorax (= furmani)
Fam. Ochotonidae
Ochotona hyperborea yesoensis – Eucheyletiella takahasii
O. hyperborea – Eucheyletiella takahasii
O. pallasi – Eucheyletiella pallasius
O. princeps – Eucheyletiella johnstoni
O. pusillinus – Eucheyletiella pusillinus
O. roylei – Eucheyletiella ochotonae
O. rufescens – Eucheyletiella faini, E. ochotonae
Carnivora
Fam. Canidae
Canis familiaris – Cheyletiella yasguri (phoretic)
Fam. Felidae
Felis catus – Cheyletiella blakei, Ch. parasitivorax
Fam. Herpestidae
Herpestes bocagei – Nihelia curvidens
H. punctulatus – Nihelia curviden
H sanguineus – Nihelia curvidens
H. edwardsii – Nihelia calcarata
Fam. Mustelidae
Spilogale putorius – Eucheyletia bishoppi
Fam. Procyonidae
Bassariscus astutus – Cheyletus eruditus, Eucheyletia hardyi
Fam. Viverridae
Cynictis penicillata – Nihelia cynictis
Primates
Fam. Galagonidae
Galago senegalensis – Galagocheles lawrencei
G. moholi – Galagocheles lawrencei, G. lemuricola
Otolemur garnetti – Galagocheles lemuricola
O. crassicaudata monteri – Galagocheles lemuricola
Fam. Hominidae
Homo sapiens – Cheyletiella blakei, Ch. parasitivorax
Psorergatidae Dubinin, 1955: 150
The itch-mites of the family Psorergatidae are tiny inhabitants of the skin of Soricomorpha, Macroscelidea, Scandentia, Rodentia, Lagomorpha, Carnivora, Primates, and Artiodactyla.
The first species of this peculiar mites was described as early as 1883 by J.B. Tyrrell in Canada. For the next 70 years very little attention was given to the tiny intradermal Psorergatid mites. In Europe the example of Michael (1889) who described from England the second Psorergatid species (under the “new” genus Goniomerus), was not followed until the time of Dubinin and Fain. In his paper on rodent parasites (1955) and in his revision of Cheyletoidea (1957) Dubinin described, among other new families, Psorergatidae with only two species.
Meanwhile, several new species have been described from countries outside Europe, mainly from bigger mammals (now in subgenus Psorobia): P. ovis Womersley, 1941 (Australia), P. cercopitheci Zumpt et Till, 1955 (South Africa), P. bos Johnston, 1964 (USA).
The papers of Fain and co-authors from 1959 to 1973 laid down the structure of the family, but it was F. Lukoschus who, between 1967 and 1979, together with his co-authors Beaujean, Driessen, Fain, De Cock, Fauran, Giesen, Maa, Hallmann, Rack, Rosmalen, described 47 (?) of all 77 species in the family. Giesen (1990) made the comprehensive revision of the family. Other modern authors are R. Haitlinger (1979–1997), A. Baker (2005) and others. As Psorergatids seem to be host-specific, many new species are to be discovered.
For the time being the family Psorergatidae consists of 77 species [73, acc. to Zhang et al., 2011], distributed among three genera, as pointed by Fain (1959a, b):
Psorergates Tyrrell, 1883 – 49 sp.
Psorergatoides Fain, 1959 – 18 sp.
Psorobia Fain, 1959 – 10 sp.
Psorergatidae Dubinin, 1955: 150
Type genus: Psorergates Tyrrell, 1883
Psorergates Tyrrell, 1883: 337
Type species: Psorergates simplex Tyrrell, 1883
Goniomerus Michael, 1889: 400
Type species: Goniomerus musculinus Michael, 1889
Psorergates (Psorergates) agrestis Lukoschus, Fain & Beaujean, 1967: 149
Psorergates agrestis, Giesen, 1990: 56
Type locality: Hoensbroek (Limburg, Holland)
Distribution: Holland
Host: Microtus agrestis bailloni (Rodentia: Arvicolidae)
Psorergates (Psorergates) apodemi Fain, Lukoschus & Hallmann, 1966: 261; Sosnina, 1970: 538; Beron, 1973: 182
Psorergates apodemi: Haitlinger, 1980: 700; Giesen, 1990: 56
Type locality: Hatert (Holland)
Distribution: Austria, Bulgaria, Germany, Holland, Poland, Romania, Switzerland, Ukraine, f. Tchecoslovakia
Hosts: Sylvaemus flavicollis, S. sylvaticus, S. microps, S. uralensis (Rodentia: Muridae, Murinae)
Psorergates (Psorergates) arvalis Lukoschus, Fain & Beaujean, 1967: 171
Psorergates arvalis, Giesen, 1990: 56
Type locality: Parque Nacional de Aigües Tortes, Lerida
Distribution: Spain
Host: Microtus arvalis meridianus (Rodentia: Arvicolidae)
Psorergates auricola Lukoschus, Fain & Beaujean, 1967: 167
Type locality: Jaca
Distribution: Spain
Host: Microtus [Pitymys] duodecimcostatus flavescens (Rodentia: Arvicolidae)
Psorergates baueri Lukoschus, de Cock & Driessen, 1971: 190
Type locality: St. Michael, Salzburg
Distribution: Austria
Host: Neomys fodiens (Soricomorpha: Soricidae)
Psorergates (Psorergates) callipidis Lukoschus, Fain & Beaujean, 1967: 156; Lukoschus, 1967: 203
Type locality: El Puerto, Asturia
Distribution: Spain
Host: Sylvaemus sylvaticus callipides (Rodentia: Muridae, Murinae)
Psorergates canadensis Kok, Lukoschus & Clulow, 1971: 1243
Type locality: Sudbury (Ont.)
Distribution: Canada (Ontario, Manitoba)
Host: Microtus pennsylvanicus (Rodentia: Arvicolidae)
Psorergates cinereus Kok, Lukoschus & Clulow, 1971: 1239
Type locality: Sudbury (Ont.)
Distribution: Canada (Ontario, Manitoba)
Host: Sorex cinereus (Soricomorpha: Soricidae)
Psorergates (Psorergates) crocidurae Lukoschus, 1968a: 81; Giesen & Lukoschus, 1983: 410
Psorergates crocidurae Giesen, 1990: 54
Type locality: Nijmegen (Holland)
Distribution: China, Germany, France, Holland, Spain
Host: Crocidura russula russula (Soricomorpha: Soricidae)
Psorergates cryptotis Giesen & Lukoschus, 1983: 404
Type locality: Cerro Punta
Distribution: Panama
Host: Cryptotis n. nigrescens (Soricomorpha: Soricidae)
Psorergates (Psorergates) deomydis Lukoschus, Fain & Beaujean, 1967: 159
Psorergates deomydis Giesen, 1990: 56
Type locality: Albert (= Virunga) Nat. Park
Distribution: DR Congo
Host: Deomys ferrugineus chrystyi (Rodentia: Muridae, Deomyinae)
Psorergates desmanae Lukoschus, 1968b: 125
Type locality: Moulis (Ariège)
Distribution: France
Host: Galemys pyrenaicus (Soricomorpha: Talpidae, Desmaninae)
Psorergates (Psorergates) dissimilis Fain, Lukoschus & Hallmann, 1966: 267
Psorergates dissimilis, Giesen, 1990: 56; Haitlinger, 1978: 143
Type locality: Hamert, Limburg Prov.
Distribution: Austria, Holland, Poland, Switzerland, Crimea
Hosts: Microtus arvalis, Myodes [Clethrionomys] glareolus (Rodentia: Arvicolidae)
Psorergates doriae Giesen, Lukoschus & Nadchatram, 1982: 319
Type locality: Kampong Taii, Sarawak
Distribution: Malaysia (Sarawak)
Host: Crocidura [doriae] foetida (Soricomorpha: Soricidae)
Psorergates dremomydis Giesen, Lukoschus & Nadchatram, 1982: 316
Type locality: Gunong (Mt.) Brinchang, 1500 m, Pahang
Distribution: Malaysia (Pahang)
Host: Dremomys rufigenis (Rodentia: Sciuridae, Sciurinae)
Psorergates (Psorergates) eliomydis Lukoschus, Fain & Beaujean, 1967: 156
Psorergates eliomydis, Giesen, 1990: 56
Type locality: Ntra. Sra. del Pilar (Formentera, Ibiza)
Distribution: Spain (Ibiza)
Host: Eliomys quercinus ophiusae (Rodentia: Gliridae)
Psorergates etruscus de Cock, Lukoschus & Ariani, 1970: 1
Type locality: Naples (Italy)
Distribution: Italy
Host: Suncus etruscus (Soricomorpha: Soricidae)
Psorergates fritzi Giesen, Nutting & Nadchatram, 1989: 133
Type locality: Fraser’s Hill, Pahang, 1200 m
Distribution: Malaysia (Pahang), Singapore
Host: Tupaia glis (Scandentia: Tupaiidae)
Psorergates glaucomys Ah, Peckham & Ateyo, 1973: 369
Type locality: Savannah, Chatham Co., Georgia
Distribution: U.S.A. (Georgia)
Host: Glaucomys v. volans (Rodentia: Sciuridae)
Psorergates hispanicus Lukoschus, Fain & Beaujean, 1967: 163; Giesen, 1990: 56
Type locality: Jaca
Distribution: Spain
Hosts: Mus musculus commissarius, M. spretus (Rodentia: Muridae, Murinae)
Psorergates meati ? – Giesen, 1990: 56
Type locality: Spain
Distribution: Spain
Host: Sylvaemus sylvaticus callipides (Rodentia: Muridae, Murinae)
Psorergates mexicanus Giesen, Lukoschus, Whitaker & Gettinger, 1983: 169
Type locality: Socorro Co. (New Mexico)
Distribution: USA (New Mexico)
Host: Notiosorex crawfordi (Soricomorpha: Soricidae)
Psorergates micromydis Lukoschus, Fain & Beaujean, 1967: 143
Type locality: Hellevoetsluis
Distribution: Holland
Host: Micromys minutus soricinus (Rodentia: Muridae, Murinae)
Psorergates microti Fain, Lukoschus & Hallmann, 1966: 266
Type locality: Hatert
Distribution: Holland
Hosts: Microtus agrestis, Myodes [Clethrionomys] glareolus (Rodentia: Arvicolidae)
Psorergates (Psorergates) muricola Fain, 1961: 66; Fain, Lukoschus & Hallmann, 1966: 259; Lukoschus, Fain & Beaujean, 1967: 175; Beron, 1970: 146; 1973: 182
Psorergates muricola: Haitlinger, 1987a: 82; Sosnina, 1970: 539; Giesen, 1990: 56
Type locality: Kawa, Lake Albert, DR Congo
Distribution: Belgium, Bulgaria, DR Congo, France, Holland, Italy, Poland, Russia
Hosts: Sylvaemus flavicollis, S. sylvaticus, Hybomys univittatus, Lophuromys aquilus, Mus musculus, Otomys [irroratus elgonis] tropicalis (Rodentia: Muridae, Murinae)
Psorergates muscardinus Lukoschus, de Cock & Driessen, 1971: 185
Type locality: München
Distribution: Germany
Host: Muscardinus avellanarius (Rodentia: Gliridae)
Goniomerus musculinus Michael, 1889: 405; Canestrini, 1894: 812
Psorergates simplex var. musculinus, Neumann, 1893: 21
Psorergates musculinus (Michael): Oudemans, 1906: 78; Hirst, 1919: pl. XII, fig. 52; Vitzthum, 1929: 54; Turk, 1953; Zumpt & Till, 1955: 269; Dubinin, 1955: 152; Fain, Lukoschus & Hallmann, 1966: 254; Rioux & Golvan, 1961:
Type locality: England
Distribution: England, Russia
Hosts: Microtus agrestis, Myodes [Clethrionomys] glareolus (Rodentia: Arvicolidae), Sylvaemus sylvaticus (Muridae)
Psorergates neerlandicus Lukoschus, de Cock & Driessen, 1971: 192
Type locality: Texel Island, the Netherlands
Distribution: Holland
Host: Microtus oeconomus (Rodentia: Arvicolidae)
Psorergates oeconomi Lukoschus, Fain & Beaujean, 1967: 139
Type locality: Zuidland, Rockanje (Holland)
Distribution: Holland
Host: Microtus oeconomus arenicola (Rodentia: Arvicolidae)
Psorergates simplex, Lavoirierre, 1946: 81 (nec Tyrrell, 1883)
Psorergates oettlei Till, 1960: 75
Psorergates (Psorergates) oettlei (Till): Fain, 1961: 61; Fain, Lukoschus & Hallmann, 1966: 258
Type locality: Johannesburg
Distribution: South Africa
Host: Mastomys [Rattus] natalensis (Rodentia: Muridae, Murinae)
Psorergates olawaensis Haitlinger, 1987: 539
Type locality: Wroclaw
Distribution: Poland
Host: Crocidura suaveolens (Soricomorpha: Soricidae)
Psorergates paraxeri Giesen & Lukoschus, 1982: 2
Type locality: Zoutpansberg (22° 58’ S, 29° 56’ E), Mahalapye (Botswana)
Distribution: Botswana, South Africa
Host: Paraxerus cepapi (Rodentia: Sciuridae)
Psorergates peromysci Giesen, Lukoschus, Whitaker & Gettinger, 1983: 164
Type locality: Lively (Ontario), Abercorn (Quebec), Lincoln Co. (Oregon), Oconne Co. (Georgia)
Distribution: Canada (Ontario, Quebec), USA (Oregon, Georgia)
Hosts: Peromyscus leucopus, P. maniculatus (Rodentia: Cricetidae)
Psorergates pinetorum Giesen, Lukoschus, Whitaker & Gettinger, 1983: 166
Type locality: Huntingdon Co. (Pennsylvania), Bicknell (Indiana)
Distribution: USA (Pennsylvania, Indiana)
Host: Microtus pinetorum (Rodentia: Arvicolidae)
Psorergates (Psorergates) pitymydis Lukoschus, Fain & Beaujean, 1967: 151
Psorergates pitymydis, Giesen, 1990: 56
Type locality: Arganda nr. Madrid, Jaca (Huesca)
Distribution: Spain
Host: Microtus [Pitymys] duodecimcostatus flavescens (Rodentia: Arvicolidae)
Psorergates polonicus: Haitlinger, 1986: 425
Type locality: Wroclaw – Swojczyce
Distribution: Poland
Host: Microtus [Pitymys] subterraneus (Rodentia: Arvicolidae)
Psorergates quercinus Lukoschus, de Cock & Driessen, 1971: 195
Type locality: Madrid
Distribution: Spain
Host: Eliomys quercinus (Rodentia: Gliridae)
Psorergates rattus Fain & Goff, 1986: 197; Cierocka & Izdebska, 2019: 314
Type locality: Oahu I., Nuuanu (Hawaiian Is.)
Distribution: USA (Hawaiian Islands), Poland
Host: Rattus norvegicus (Rodentia: Muridae, Murinae)
Psorergates simplex Tyrrell, 1883: 337; Neumann, 1893: 13; Canestrini, 1894: 809; Oudemans, 1906: 77; Vitzthum, 1929: 54; Dubinin, 1953: 276; Zumpt & Till, 1955: 269; Dubinin, 1955: 151; Till, 1960: 75; Fain, 1961: 61; Spicka, 1976: 418; Kamali, Ostovan & Atamehr, 2001; Izdebska & Fryderyk, 2012: 20
Psorergates (Psorergates) simplex (Tyrrell): Fain, Lukoschus & Hallmann, 1966: 252
Type locality: Canada
Distribution: Canada, France, Holland, Iran, Russia, USA (Indiana)
Hosts: Mus musculus (Rodentia: Muridae, Murinae); Arvicola terrestris, Microtus arvalis (Arvicolidae)
Psorergates sorici Lukoschus, 1968: 85
Type locality: Nijmegen (Holland)
Distribution: Holland
Host: Sorex araneus araneus (Soricomorpha: Soricidae)
Psorergates squamipes Lukoschus, Louppen & Maa, 1974: 51
Type locality: Arisan, 2300 m
Distribution: Taiwan
Host: Anourosorex squamipes (Soricomorpha: Soricidae)
Psorergates (Psorergates) talpae Lukoschus, 1968: 75
Type locality: Nijmegen (Holland)
Distribution: Holland, Spain
Host: Talpa europaea europaea (Soricomorpha: Talpidae, Talpinae)
Psorergates townsendi Giesen, Lukoschus, Whitaker & Gettinger, 1983: 169
Type locality: Lincoln Co. (Oregon)
Distribution: USA (Oregon)
Host: Microtus townsendi (Rodentia: Arvicolidae)
Psorergates tupaiae Giesen & Lukoschus, 1982: 259
Type locality: Mt. Kenepai (Borneo)
Distribution: Malaysia (Borneo)
Host: Tupaia dorsalis (Scandentia: Tupaiidae)
Psorergates urotrichi Giesen & Lukoschus, 1983: 405
Type locality: Hakuba, Nagano
Distribution: Japan
Host: Urotrichus talpoides (Soricomorpha: Talpidae)
Psorergates watsoni Kok, Lukoschus & Clulow, 1971: 1243
Type locality: Sudbury (Ont.)
Distribution: Canada (Ontario)
Host: Peromyscus maniculatus (Rodentia: Cricetidae)
Psorergates (Psorergates) zibethicalis Lukoschus, Fain & Beaujean, 1967: 134
Psorergates zibethicalis,
Type locality: Prümzurlay in der Eifel, Germany
Distribution: [Germany]
Host: Ondatra zibethica (Rodentia: Arvicolidae)
Psorergates sp.
Distribution: Canada
Host: Microsorex koyi (Soricomorpha: Soricidae)
Psorergatoides Fain, 1959b: 232 (Subgenus of Psorergates)
Psorergatoides: Giesen, 1990: 22 (Genus)
Type species: Psorergates (Psorergatoides) kerivoulae Fain, 1959
Psorergatoides artibei Lukoschus, Rosmalen & Fain, 1973: 70
Type locality: Paramaribo
Distribution: Suriname
Host: Artibeus lituratus fallase (Chiroptera: Phyllostomatidae, Stenodermatinae)
Psorergatoides australiensis Giesen, Lukoschus & Fain, 1982: 315; Domrow, 1991: 1336; Halliday, 1998: 79
Type locality: Geikie Gorge, W. A. (18° 05’ S; 125° 43’ E)
Distribution: Australia (Western Australia)
Hosts: Vespadellus [Eptesicus] pumilus, V. [E.] [douglasi] douglasorum, Nyctophilus arnhemensis, N. gouldi, N. walkeri (Chiroptera: Vespertilionidae)
Psorergatoides desmodus Lukoschus, Louppen & Fauran, 1979: 311
Type locality: Cayenne
Distribution: French Guyana
Host: Desmodus rotundus (Chiroptera: Phyllostomatidae, Desmodontinae)
Psorergates (Psorergatoides) emballonurae Fain, 1959b: 241
Psorergatoides emballonurae Fain
Type locality: Buin (Bougainville Is.)
Distribution: Papua New Guinea (Bougainville)
Host: Mosia [Emballonura] nigrescens (Chiroptera: Emballonuridae)
Psorergatoides glossophagae Lukoschus, Rosmalen & Fain, 1973: 63
Type locality: Leonsberg
Distribution: Suriname
Host: Glossophaga soricina (Chiroptera: Phyllostomatidae, Glossophaginae)
Psorergatoides guyanensis Lukoschus, Louppen & Fauran, 1979: 322
Type locality: Rochambeau
Distribution: French Guyana
Host: Rhinophylla pumilio (Chiroptera: Phyllostomatidae)
Psorergates (Psorergatoides) hipposideros Fain, 1959b: 237
Psorergatoides hipposideros (Fain): Giesen, 1990: 55
Type locality: Bunia (Ituri); Grotte de Kakontwe
Distribution: DR Congo
Hosts: Hipposideros abae, H. centralis (Chiroptera: Hipposideridae)
Psorergatoides indicicola Lukoschus, Rosmalen & Fain, 1973: 67
Type locality: Lelydorp
Distribution: Suriname
Hosts: Saccopteryx bilineata, S. canescens (Chiroptera: Emballonuridae)
Psorergatoides kerivoulae Fain, 1959a: 61
Psorergates (Psorergatoides) kerivoulae Fain, 1959b: 239
Psorergates kerivoulae Fain: Haitlinger, 1979b: 109
Type locality: Bokuma, Pawa (DR Congo)
Distribution: Belgium, DR Congo, Ivory Coast, Malaysia (Borneo), Poland
Hosts: Kerivoula cuprosa, K. [harrisoni] lanosa, Myotis bocagei, M. muricola, M. myotis, M. mystacinus, Plecotus auritis (Chiroptera: Vespertilionidae)
Psorergates laviae Fain, 1959a: 64
Psorergates (Psorergatoides) laviae Fain, 1959b: 247
Psorergatoides laviae (Fain): Giesen, 1990: 54
Type locality: Akanyaru River, Gabiro, Kagera Nat. Park
Distribution: Rwanda
Host: Lavia frons (Chiroptera: Megadermatidae)
Psorergatoides (Psorergatoides) lonchorhinae Fain, 1959b: 243
Psorergatoides lonchorhina Fain, Giesen, 1990: 54
Type locality: San Esteban
Distribution: Venezuela
Hosts: Saccopteryx canescens [canina] (Chiroptera: Emballonuridae), Lonchorhina aurita (Chiroptera: Phyllostomatidae)
Psorergatoides molossi Lukoschus, Rosmalen & Fain, 1973: 72
Type locality: Lelydorp
Distribution: Suriname
Host: Molossus molossus (Chiroptera: Molossidae)
Psorergatoides nyctali Baker, 2005: 67; Heddergott & Claussen, 2008: 57
Type locality: Dartington Hall, near Totnes, Devon, England
Distribution: England, Germany
Host: Nyctalus noctula (Chiroptera: Vespertilionidae)
Psorergates (Psorergatoides) nycteris Fain, 1959a: 56
Psorergatoides nycteris (Fain): Giesen, 1990: 54
Type locality: Musha, nr. Akanyaru; Elisabethville (= Lubumbashi)
Distribution: DR Congo, Rwanda
Host: Nycteris macrotis (Chiroptera: Nycteridae)
Psorergatoides peropteryx Lukoschus, Louppen & Fauran, 1979: 316
Type locality: Cayenne
Distribution: French Guyana
Hosts: Cormura brevirostris, Peropteryx macrotis (Chiroptera: Emballonuridae)
Psorergates rhinolophi Fain, 1959a: 59
Psorergates (Psorergatoides) rhinolophi Fain, 1959b: 235
Psorergatoides rhinolophi (Fain): Giesen, 1990: 54; Heddergott & Claussen, 2008: 57
Type locality: Lubudi, Uvira (DR Congo), Kambisa (Angola)
Distribution: Angola, Belgium, Burma, DR Congo, France, Germany, Italy, Spain
Hosts: Rhinolophus aethiops, Rh. affinis, Rh. zuluensis, Rh. euryale, Rh. ferrumequinum, Rh. fumigatus, Rh. hipposideros, Rh. hildebrandti, Rh. mehelyi (Chiroptera: Rhinolophidae)
Psorergatoides surinamensis Lukoschus, Louppen & Fauran, 1979: 319
Type locality: Wageningen
Distribution: Suriname
Hosts: Lophostoma [Tonatia] carrikeri, Lophostoma brasiliense [T. nicaraguae] (Chiroptera: Phyllostomatidae)
Psorergatoides tadaridae Giesen, Lukoschus & Nadchatram, 1982: 325
Type locality: Gombak Forest Reserve, Selangor
Distribution: Malaysia (Selangor)
Host: Mops [Tadarida] mops (Chiroptera: Molossidae)
Psorobia Fain, 1959a: 55
Psorergates (Psorobia): Fain, 1961: 61
Type species: Psorergates ovis Womersley, 1941
Psorergates bos Johnston, 1964: 1; Oberem & Malam, 1983: 63
Psorobia bos (Johnston): Fain, 19; Giesen & Yunker, 1985: 37; Baker, Andrews & Fox, 1996; Andrews, Baker, Fox & Brizuela, 1997: 183.
Type locality: New Mexico
Distribution: Great Britain, South Africa, USA (New York, New Mexico)
Host: Bos taurus (Artiodactyla: Bovidae)
Psorobia castoris Kok, Lukoschus & Clulow, 1970: 1419
Type locality: Blezard Valley (Ontario)
Distribution: Canada (Ontario)
Host: Castor canadensis (Rodentia: Castoridae)
Psorergates cercopitheci Zumpt & Till, 1955: 272; Seier, 1985: 236
Psorobia cercopitheci (Zumpt & Till): Fain, 19...
Type locality: Bedford, Cape Province
Distribution: South Africa
Host: Cercopithecus aethiops pygerythus (Primates: Cercopithecidae)
Psorobia elephantuli Giesen, Spicka & Whitaker, 1985: 147
Type locality: Mezuna, Tunis
Distribution: Tunisia
Host: Elephantulus rozeti (Macroscelidea: Macroscelididae)
Psorergates (Psorobia) foinae Fain & Lukoschus, 1968: 3
Psorobia foinae (Fain et Lukoschus): Giesen, 1990: 56
Type locality: Torgny, Belgium
Distribution: Belgium
Host: Martes foina (Carnivora: Mustelidae)
Psorergates hystrici Till, 1957: 329
Psorobia hystrici (Till): Giesen, 1990: 56
Type locality: Winburg, Orange Free State
Distribution: South Africa
Host: Hystrix africaeaustralis (Rodentia: Hystricidae)
Psorobia sp.: Wassel, Whitaker & Spicka, 1980: 419
Psorobia lagomorphae Giesen, Spicka & Whitaker, 1985: 148
Type locality: Parke Co., 8 mi. S. Rockville, Indiana
Distribution: USA (Indiana)
Host: Sylvilagus floridanus (Lagomorpha: Leporidae)
Psorergates (Psorobia) mustelae Lukoschus, 1969: 111
Psorobia mustelae (Lukoschus): Giesen, 1990: 56
Type locality: Nijmegen (Holland)
Distribution: Holland
Hosts: Mustela erminea, M. nivalis (Carnivora: Mustelidae)
Psorergates ovis Womersley, 1941: 56; Carter, 1941: 193; Davis, 1954: 255; Fiedler & du Toit, 1954: 21; Zumpt & Till, 1955: 270; Whitten & Elliott, 1956: 19; Sinclair & Girson, 1970; Sinclair, 1990: 59; Whitten, 1962: 9; Pomroy, 1985: 18
Psorergates (Psorobia) ovis (Womersley): Fain, 1961: 62
Psorobia ovis (Womersley): Fain, 1959a: 56
Type locality: Australia
Distribution: Australia, New Zealand, South Africa, USA
Host: Ovis aries (Artiodactyla: Bovidae)
Psorergates (Psorobia) zumpti Fain, 1965: 295
Psorobia zumpti (Fain): Giesen, 1990: 56
Type locality: Bloemfontein
Distribution: South Africa
Host: Cryptomys hottentotus (Rodentia: Bathyergidae)
N.B.: In the general paper of Giessen (1990) are mentioned also Psorergates alleni and P. ramai.
EUROPE
Austria (Lukoschus, de Cock & Driessen, 1971) – Psorergates apodemi, P. baueri, P. dissimilis
Azerbaidzhan (Gadjiev et al., 1988) – Psorobia ovis
Belgium (Fain, 1959; Fain & Lukoschus, 1968) – Psorergates muricola, Psorergatoides kerivoulae, P. rhinolophi; Psorobia foinae
Bulgaria (Beron, 1970, 1973) – Psorergates apodemi, P. muricola
Denmark
Groenland (Giesen, 1990) – Psorergates sp.
France (Neumann, 1893; Lukoschus, 1968a, 1968b) – Psorergates desmanae, P. simplex, Psorergatoides rhinolophi
Germany (Lukoschus, Fain & Beaujean, 1967; Lukoschus, de Cock & Driessen, 1971; Heddergott & Claussen, 2008) – Psorergates apodemi, P. muscardinus, P. zibethicalis, Psorergatoides nyctali, P. rhinolophi
Great Britain (Michael, 1889; Andrews, Baker, Fox & Brizuela, 1997; Baker, 2005) – Psorergates musculinus, Psorergatoides nyctali, Psorobia bos
Holland (Fain, Lukoschus & Hallmann, 1966; Lukoschus, Fain & Beaujean, 1967; Lukoschus, 1968a, 1969; Lukoschus, de Cock & Driessen, 1971; Giessen, 1990) – Psorergates agrestis, P. apodemi, P. crocidurae, P. dissimilis, P. micromydis, P. microti, P. neerlandicus, P. oeconomi, P. simplex, P. sorici, P. talpae; Psorobia mustelae
Italy (Piana, 1886; de Cock, Lukoschus & Ariani, 1970) – Psorergates etruscus; Psorergatoides rhinolophi
Poland (Haitlinger, 1978, 1979b, 1986, 1987, 1997; Izdebska & Fryderyk, 2012; Cierocka & Izdebska, 2019) – Psorergates apodemi, P. dissimilis, P. microti, P. muricola, P. musculinus, P. olawaensis, P. polonicus, P. rattus, P. simplex, Psorergatoides kerivoulae
Romania (Haitlinger, 1980) – Psorergates apodemi
Russia (all) (Dubinin, 1953, 1955; Sosnina, 1970) – Psorergates apodemi, P. musculinus, P. simplex
Spain (cont.) (Lukoschus, Fain & Beaujean, 1967; Lukoschus, 1967, 1968a; Lukoschus, de Cock & Driessen, 1971; Zapatero Ramos et al., 1982) – Psorergates arvalis, P. auricola, P. callipidis, P. crocidurae, P. eliomydis [Ibiza], P. hispanicus, P. meati, P. pitymydis, P. quercinus, P. talpae; Psorergatoides rhinolophi
Sweden (Giesen, 1990) – Psorergates sp.
Switzerland (Sosnina, 1970) – Psorergates apodemi
Ukraine (Dubinin, 1955; Sosnina, 1970) – Psorergates apodemi, P. dissimilis
AFRICA
Angola (Giesen, 1990) – Psorergatoides rhinolophi
Botswana (Giesen & Lukoschus, 1982) – Psorergates paraxeri
Congo DR (formerly Zaire)(Fain, 1959a, 1959b, 1961; Fain, Lukoschus & Hallmann, 1966; Lukoschus, Fain & Beaujean, 1967;) – Psorergates deomydis; Psorergatoides kerivoulae, P. nycteris, P. rhinolophi
Ivory Coast (Giesen, 1990) – Psorergatoides kerivoulae
Madagascar (Giesen, 1990) – Psorergatidae gen. sp.
Rwanda (Fain, 1959a) – Psorergatoides laviae, P. nycteris
South Africa (Lavoipierre, 1946; Fiedler & du Toit, 1954; Zumpt & Till, 1955; Till, 1960; Fain, 1965; Oberem & Malan, 1983) – Psorergates cepapi, P. oettlei, P. paraxeri; Psorobia bos, P. cercopitheci, P. hystrici, P. ovis, P. zumpti
Tunisia (Giesen, Spicka & Whitaker, 1985) – Psorobia elephantuli
ASIA
Burma (Giesen, 1990) – Psorergatoides rhinolophi
China (Giesen, 1990) – Psorergates crocidurae
Iran (Kamali, Ostovan & Atamehr, 2001) – Psorergates simplex
Japan (Giesen & Lukoschus, 1983) – Psorergates urotrichi
Malaysia (cont.) (Beck, 1971; Giesen, Lukoschus & Nadchatram, 1982; Giesen, Nutting & Nadchatram, 1989; Giesen & Lukoschus, 1982) – Psorergates doriae, P. dremomydis, P. fritzi; Psorergatoides tadaridae
Sarawak and Sabah (Giesen & Lukoschus, 1982) – Psorergates tupaiae; Psorergatoides kerivoulae
Singapore (Giesen, Nutting & Nadchatram, 1989) – Psorergates fritzi
Taiwan (Lukoschus, Louppen & Maa, 1974) – Psorergates squamipes
NORTH AMERICA
Canada (Tyrrell, 1883; Kok, Lukoschus & Clulow, 1970, 1971; Giesen, Lukoschus, Whitaker & Gettinger, 1983) – Psorergates canadensis, P. cinereus, P. peromysci, P. simplex, P. watsoni; Psorobia castoris
Mexico (Giesen, 1990) – Psorergates mexicanus
USA (Ah, Peckham & Ateyo, 1973; Spicka, 1976; Wassel, Whitaker & Spicka, 1980; Giesen, Lukoschus, Whitaker & Gettinger, 1983; Giesen, Spicka & Whitaker, 1985; Giesen & Yunker, 1985;) – Psorergates glaucomys, P. mexicanus, P. peromysci, P. pinetorum, P. simplex, P. townsendi; Psorobia bos, P. lagomorphae, P. ovis
Hawaii Is. (Fain & Goff, 1986) – Psorergatus rattus
SOUTH and CENTRAL AMERICA
French Guyana (Lukoschus, Louppen & Fauran, 1979) – Psorergatoides desmodus, P. guyanensis, P. peropteryx
Panama (Giesen & Lukoschus, 1983) – Psorergates cryptotis
Suriname (Lukoschus, Rosmalen & Fain, 1973) – Psorergatoides artibei, P. glossophagae, P. molossi, P. nidicicola, P. surinamensis
Venezuela (Fain, 1959b) – Psorergates sp., Psorergatoides lonchorhinae
OCEANIA
Australia (continent) (Womersley, 1941; Carter, 1941; Toop, 1948; Murray, 1961; Downing & Mort, 1962; Sutherland, 1965; Giesen, Lukoschus & Fain, Sinclair & Girson, 1970; 1982; Sinclair, 1990; Domrow, 1991) – Psorergatoides australiensis; Psorobia ovis
New Zealand (Whitten & Elliott, 1956; Whitten, 1962; Pomroy, 1985) – Psorobia ovis
Papua New Guinea (Fain, 1959b) – Psorergatoides emballonurae
Mammalia
Soricomorpha
Fam. Soricidae
Anourosorex squamipes – Psorergates squamipes
Crocidura doriae – Psorergates [doriae] foetida
C. lasiura – Psorergates crocidurae
C. russula russula – Psorergates crocidurae
C. suaveolens – Psorergates olawaensis
Cryptotis n. nigrescens – Psorergates cryptotis
Microsorex koyi – Psorergates sp.
Neomys fodiens – Psorergates baueri
Notiosorex crawfordi – Psorergates mexicanus
Sorex araneus – Psorergates sorici
S. cinereus – Psorergates cinereus
Suncus etruscus – Psorergates etruscus
Fam. Talpidae
Galemys pyrenaicus – Psorergates desmanae
Neurotrichus gibbsi – Psorergates sp.
Talpa europaea europaea – Psorergates talpae
Urotrichus talpoides – Psorergates urotrichi
Afrosoricida
Fam. Tenrecidae
Echinops telfairi – Psorergatidae sp.
Macroscelidea
Fam. Macroscelididae
Elephantulus rozeti – Psorobia elephantuli
Scandentia
Fam. Tupaiidae
Tupaia dorsalis – Psorergates tupaiae
T. glis – Psorergates fritzi
Chiroptera
Fam. Emballonuridae
Cormura brevirostris – Psorergatoides peropteryx
Mosia [Emballonura] nigrescens – Psorergatoides emballonurae
Peropteryx macrotis – Psorergatoides peropteryx
Saccopteryx canina – Psorergatoides lonchorhinae
S. bilineata – Psorergatoides indicicola
S. canescens – Psorergatoides indicicola
Fam. Hipposideridae
Hipposideros caffer centralis – Psorergatoides hipposideros
H. abae – Psorergatoides hipposideros
Fam. Megadermatidae
Lavia frons – Psorergatoides laviae
Fam. Molossidae
Molossus ater – Psorergatoides molossi
M. molossus – Psorergatoides molossi
Mops [Tadarida] mops – Psorergatoides tadaridae
Fam. Nycteridae
Nycteris macrotis – Psorergatoides nycteris
Fam. Phyllostomatidae
Artibeus lituratus fallase – Psorergatoides artibei
Desmodus rotundus – Psorergatoides desmodus
Glossophaga soricina – Psorergatoides glossophagae
Lonchorhina aurita – Psorergatoides lonchorhinae
Rhynophylla pumilio – Psorergatoides gyuanensis
Lophostoma [Tonatia] carrikeri – Psorergatoides surinamensis
Lophostoma brasiliense [Tonatia nicaraguae] – Psorergatoides surinamensis
Fam. Rhinolophidae
Rhinolophus aethiops – Psorergatoides rhinolophi
Rh. affinis – Psorergatoides rhinolophi
Rh. zuluensis – Psorergatoides rhinolophi
Rh. euryale – Psorergatoides rhinolophi
Rh. fumigatus – Psorergatoides rhinolophi
Rh. hildebrandti – Psorergatoides rhinolophi
Rh. hipposideros – Psorergatoides rhinolophi
Rh. mehelyi – Psorergatoides rhinolophi
Fam. Vespertilionidae
Kerivoula cuprosa – Psorergatoides kerivoulae
K. [harrisoni bellula] lanosa – Psorergatoides kerivoulae
Myotis bocagei – Psorergatoides kerivoulae
M. muricola – Psorergatoides kerivoulae
M. myotis – Psorergatoides kerivoulae
M. mystacinus – Psorergatoides kerivoulae
Nyctalus noctula – Psorergatoides nyctali
Nyctophilus arnhemensis – Psorergatoides australiensis
N. gouldi – Psorergatoides australiensis
N. walkeri – Psorergatoides australiensis
Plecotus auritis – Psorergatoides kerivoulae
Vespadellus [Eptesicus] pumilus – Psorergatoides australiensis
Vespadellus douglasorum [E. douglasi] – Psorergatoides australiensis
Rodentia
Fam. Arvicolidae
Arvicola terrestris – Psorergates simplex
Myodes [Clethrionomys] glareolus – Psorergates microti, P. dissimilis
M. [C.] gapperi atabascae – Psorergates sp.
M. [C.] gapperi proteus – Psorergates sp.
M. [C.] rufocanus – Psorergates sp.
M. [C.] rutilus – Psorergates sp.
Dicrostonyx groenlandicus – Psorergates sp.
Microtus agrestis bailloni – Psorergates agrestis
M. agrestis – Psorergates microti
M. arvalis – Psorergates simplex, P. dissimilis
M. arvalis meridianus – Psorergates arvalis
M. [Pitymys] duodecimcostatus flavescens – Psorergates auricola, P. pitymydis
M. nivalis – Psorergates sp.
M. oeconomus – Psorergates neerlandicus
M. oeconomus arenicola – Psorergates oeconomi
M. pennsylvanicus – Psorergates canadensis
M. [Pitymys] pinetorum – Psorergates pinetorum
M. townsendi – Psorergates townsendi
Microtus townsendi – Psorergates townsendi
Ondatra zibethica – Psorergates zibethicalis
Fam. Bathyergidae
Cryptomys hottentotus – Psorobia zumpti
Fam. Castoridae
Castor canadensis – Psorobia castoris
Fam. Cricetidae
Peromyscus maniculatus – Psorergates watsoni, P. peromysci
P. leucopus – Psorergates peromysci
Reithrodontomys megalotis – Psorergates sp.
Sigmodon hispidus – Psorergates sp.
Fam. Gliridae (Myoxidae)
Eliomys quercinus – Psorergates pitymydis
E. quercinus ophiusae – Psorergates eliomydis
Muscardinus avellanarius – Psorergates muscardinus
Fam. Heteromyidae
Heteromys anomalus – Psorergates sp.
Fam. Hystricidae
Hystrix africaeaustralis – Psorobia hystrici
Fam. Muridae
Deomys ferrugineus chrystyi – Psorergates deomydis
Hybomys univittatus – Psorergates muricola
Lophuromys aquilus – Psorergates muricola
Micromys minutus soricinus – Psorergates micromydis
Mus musculus – Psorergates muricola, P. simplex
M. musculus commissarius – Psorergates hispanicus
M. spretus – Psorergates hispanicus
Otomys [irroratus elgonis] tropicalis – Psorergates muricola
Rattus norvegicus – Psorergates rattus
R. natalensis – Psorergates oettlei
Sylvaemus sylvaticus – Psorergates apodemi, P. muricola
S. sylvaticus callipides – Psorergates callipidis, P. meati
S. flavicollis – Psorergates apodemi, P. muricola
S. microps – Psorergates apodemi
S. uralensis – Psorergates apodemi
Fam. Sciuridae
Dremomys rufigenis – Psorergates dremomydis
Glaucomys v. volans – Psorergates glaucomys
Paraxerus cepapi – Psorergates paraxeri
Lagomorpha
Fam. Leporidae
Sylvilagus floridanus – Psorobia lagomorphae
Carnivora
Fam. Mustelidae
Martes foina – Psorobia foinae
Mustela erminea – Psorobia mustelae
M. nivalis – Psorobia mustelae
Primates
Fam. Cercopithecidae
Cercopithecus aethiops pygerythrus – Psorobia cercopitheci
C. mona mona – Psorobia sp.
Macaca arctoides (USA, captured) – Psorobia sp.
Artiodactyla
Fam. Bovidae
Bos taurus – Psorobia bos
Ovis aries – Psorobia ovis
Demodecidae Nicolet, 1855: 383
The history of the study of Demodecidae starts with Owen (1843), who described the genus Demodex. Hirst (1919) published a comprehensive review of Demodex – the only known genus and mentioned that already there are more than 230 publications on the group, mostly on veterinary cases (D. canis, etc.). Nutting (1964) added information about 650 more titles, and since there have been hundreds of publications – the literature on Demodecidae is well over 1000 titles. Most of them are of little taxonomical or host – related interest and represent case reports. After Hirst (1918a, 1918b, 1919, 1921, 1923) particularly important were the contributions of Fain (1959, 1960) who described the new genera Rhinodex and Stomatodex and 4 of the valid species in the family. Banks (1915) and Dubinin (1958) raised the superfamily Demodicoidea, but this arrangement was not followed. Among the acarologists having contributed to the taxonomy of Demodecidae should be noted C.E. Desch (since 1969), W.B. Nutting (since 1950), V. Bukva (since 1982, described the genera Apodemodex and Soricidex), Lukoschus and co-authors (described the genera Ophthalmodex and Pterodex) and Izdebska and co-authors (since 2000), described the genus Glossicodex and many new species.
According Bochkov, 2009b, this family includes 100 sp., including 86 valid sp. of Demodex. Izdebska & Rolbiecki (2020) presented a checklist containing 122 valid species of Demodecidae. We have listed here 122 species of Demodecidae, six species are not described according to ICZN and two are nomina nuda. They live in the skin glands of mammals (32 families and 14 orders, fide Bochkov, 2009b). The representatives of Ophthalmodex live in the epithelium of the eyelids, cornea, and lacrimal canals of bats (Hipposideridae, Vespertilionidae, Molossidae, Phylostomatidae) and rodents (Muridae). The Stomatodex prefer the oral cavity and esophagus of bats, rodents and primates. The Rhinodex have been found in the nose cavity of Galagidae (Primates). The small genera Soricidex, Apodemodex and Pterodex inhabit the skin follicles or the epithelium of Soricidae, Muridae and Pteropodidae (fruit bats). Most of the demodecids are monoxenous parasites. Some are important for the animal health, causing demodectic mange, and even to humans (Demodex folliculorum, D. brevis).
Here efforts were made to include in the bibliography most papers containing taxonomic and faunistic information and some of the medical papers (others could be seen in the bibliography of Izdebska & Rolbiecki, 2020).
Fam. Demodecidae Nicolet, 1855 – 8 genera, 122 species
Apodemodex Bukva, 1996 – 1 sp.
Demodex Owen, 1843 – 106 sp.
Glossicodex Izdebska et Rolbiecki, 2016 – 1 sp.
Ophthalmodex Lukoschus et Nutting, 1979 – 7 sp.
Pterodex Lukoschus, Woeltjes, Desch et Nutting, 1980 – 1 sp.
Rhinodex Fain, 1959 – 1 sp.
Soricidex Bukva, 1982 – 1 sp.
Stomatodex Fain, 1959 – 4 sp.
Demodicidae Owen, 1843
Demodecidae Nicolet, 1855: 383
Demodicoidea Banks, 1915 (superfamily)
Demodicoidea Dubinin, 1954: 73; 1955: 141; 1957: 124 (superfamily)
Type genus: Demodex Owen, 1843
Apodemodex Bukva, 1996: 312
Type species: Apodemodex cornutus Bukva, 1996
Apodemodex cornutus Bukva, 1996: 312; Izdebska & Rolbiecki, 2020: 10
Type locality: České Budějovice, South Bohemia
Distribution: Czech Rep.
Host: Neomys anomalus (Soricomorpha: Soricidae)
Demodex Owen, 1843
Type species: Acarus folliculorum Simon, 1842
Demodex acutipes Bukva & Preisler, 1988: 69; Izdebska & Rolbiecki, 2020: 10
Type locality: Czech Rep.
Distribution: Czech Rep., Poland
Host: Cervus elaphus (Artiodactyla: Cervidae)
Demodex aelleni Fain, 1960: 81; Izdebska & Rolbiecki, 2020: 10
Type locality: Geneva
Distribution: Switzerland
Host: Myotis daubentoni (Chiroptera: Vespertilionidae)
Demodex agrarii Bukva, 1994: 306; Izdebska & Cydzik, 2010: 59; Izdebska & Rolbiecki, 2020: 10
Type locality: Kokšov-Bakša, Eastern Slovakia
Distribution: Poland, Slovakia
Host: Apodemus agrarius (Rodentia: Muridae)
Demodex ailuropodae Xu, Xie, Liu, Zhou & Shi, 1986: 163; Izdebska & Rolbiecki, 2020: 10
Type locality: Shanghai Zool. Garden
Distribution: China
Host: Ailuropoda melanoleuca (Carnivora: Ailuropodidae)
Demodex antechini Nutting & Sweatman, 1970: 425; Domrow, 1991: 1339; Desch & Holz, 2006: 81; Izdebska & Rolbiecki, 2020: 10
Type locality: Canberra
Distribution: Australia
Hosts: Antechinus agilis, A. stuartii (Dasyuromorphia: Dasyuridae, Dasyuridae)
Demodex apodemi Hirst, 1918: 145; 1919: 35; Izdebska, 2012: 1101; Izdebska & Rolbiecki, 2020: 10
Demodex arvicolae var. apodemi Hirst: Dubinin, 1953: 276
Demodex arvicolae apodemi Hirst: Izdebska & Cydzik, 2010: 59
Type locality: England
Distribution: England, Poland, Russia
Hosts: Apodemus agrarius, Sylvaemus sylvaticus (Rodentia: Muridae). “Specimens on Apodemus agrarius probably belong to the separate species” (Izdebska et al., 2020).
Demodex araneae Nutting, 1950: Thesis; Izdebska & Rolbiecki, 2020: 10
Type locality: laboratory
Distribution: South America
Host: Ateles sp. (Primates: Atelidae). Note by Izdebska et al., 2020: “Nom. nud.; description not published within the meaning of the ICZN”].
Demodex aries Desch, 1986: 367; Bukva, 1990: 82; Domrow, 1991: 1343 (synonymised the species with Demodex ovis Railliet, 1893); Izdebska & Rolbiecki, 2020: 11
Type locality: Wellington City (New Zealand)
Distribution: Czech Rep., New Zealand
Host: Ovis aries (Artiodactyla: Bovidae)
Demodex artibei Vargas, Bassols, Desch, Quintero & Polaco, 1995: 75; Izdebska & Rolbiecki, 2020: 11
Type locality: Mexico
Distribution: Mexico
Host: Artibeus aztecus (Chiroptera: Phyllostomatidae)
Demodex arvicolae Zschokke, 1888: 69; Dubinin, 1953: 276; Izdebska & Rolbiecki, 2020: 11
Type locality: Europe
Distribution: England, Poland, Russia
Hosts: Microtus arvalis (?), M. agrestis, Mus musculus (?)(Rodentia: Arvicolidae)
Demodex aurati Nutting, 1961: 515; Sarashina & Sato, 1986: 619; Izdebska & Rolbiecki, 2020: 11
Type locality: laboratory Golden Hamsters
Distribution: from laboratory animals
Host: Mesocricetus auratus (Rodentia: Cricetidae)
Demodex auricularis Izdebska, Rolbiecki & Fryderyk, 2014: 215; Izdebska & Rolbiecki, 2020: 11
Type locality: Poland
Distribution: Poland
Host: Sylvaemus sylvaticus (Rodentia: Muridae)
Demodex bandicotae Izdebska, Rolbiecki, Morand & Ribas, 2017; Izdebska & Rolbiecki, 2020: 11
Type locality: Laos
Distribution: Laos
Host: Bandicota indica (Rodentia: Muridae)
Demodex bantengi Firda, Nutting & Sweatman, 1987: 227; Izdebska & Rolbiecki, 2020: 11
Type locality: Indonesia (Bali)
Distribution: Indonesia
Host: Bos javanicus (Artiodactyla: Bovidae)
Demodex bicaudatus Kniest & Lukoschus, 1981: 111; Domrow, 1991: 1339; Izdebska & Rolbiecki, 2020: 11
Type locality: Mitchell Plateau
Distribution: Australia
Host: Macroglossus minimus [lagochilus] (Chiroptera: Pteropodidae)
Demodex bisonianus Kadulski & Izdebska, 1996: 103; Izdebska, 2006: 171; Izdebska & Rolbiecki, 2020: 11
Type locality: Prevalence
Distribution: Poland
Host: Bison bonasus (Artiodactyla: Bovidae, Bovinae)
Demodex bonapartei Nutting, 1950: Thesis; Izdebska & Rolbiecki, 2020: 11.
Type locality: ?
Distribution: North America (North and East of the Great Lakes)
Host: Mustela erminea cicognanii (Carnivora: Mustelidae). Note by Izdebska & Rolbiecki, 2020: “Nom. nud.; description not published within the meaning of the ICZN”].
Demodex folliculorum: Faxon, 1878: 11
Demodex folliculorum var. bovis Stiles, 1892: 289; Railliet, 1893: 638; Cleland, 1913: 591
Demodex bovis Stiles: Canestrini & Kramer, 1899: 3; Hirst, 1919: 28; 1921: 371; Freire & Dias, 1947: 247; Desch & Nutting, 1971; 1973: 500; Esuruoso, 1977: 65; Tashiro et al., 1979: 30; Noorudin & Rahman, 1985; Matthes & Bukva, 1993; Candona, Vargas & Perdomo, 2013; Izdebska & Rolbiecki, 2020: 12.
Type locality: ?
Distribution: cosmopolitic
Hosts: Bos taurus, Bos taurus indicus, accid. Bison bonasus in farm (Artiodactyla: Bovidae, Bovinae)
Acarus foliculorum Simon, 1842: 231 (partim, fig. 3, 4)
Demodex folliculorum brevis Akbulatova, 1963: 40
Demodex brevis Akbulatova: Desch & Nutting, 1972: 171; László & Zoltán, 1973: 55; Nutting & Green, 1974: 10; 1976: 307; Hallas, 1978: 30; English & Nutting, 1981: 362; Andrews, 1988: 97; Tenquist & Charleston, 2001: 495; Özdemir, 2003; Zeytun & Karakurt, 2019: 2; Izdebska & Rolbiecki, 2020: 12
Type locality: Russia
Distribution: cosmopolitan
Host: Homo sapiens (Primates: Hominidae)
Demodex buccalis Bukva, Vítovec & Vlček, 1985: 158; Izdebska & Rolbiecki, 2020: 12
Type locality: South Bohemia
Distribution: Czech Rep., Poland
Host: Myodes [Clethrionomys] glareolus (Rodentia: Arvicolidae)
Demodex folliculorum var. equi Railliet, 1895
Demodex equi Railliet, 1895: Gmeiner, 1908
Demodex caballi (Railliet): Izdebska & Rolbiecki, 2020: 12
Type locality: Europe
Distribution: USA, New Zealand (cosmopolitan?)
Host: Equus caballus (Perissodactyla: Equidae)
Demodex cafferi Nutting & Guilfoy, 1979: 9; Dräger & Paine, 1980: 521; Izdebska & Rolbiecki, 2020: 12
Type locality: South Africa
Distribution: South Africa
Host: Syncerus caffer (Artiodactyla: Bovidae, Bovinae)
Demodex canis Leydig, 1859: 345; Canestrini & Kramer, 1899: 2; Hirst, 1919: 3; Cooreman, 1946: 2; Starkoff & Starkoff, 1950: 92; Roberts, 1952: 189; Desch & Nutting, 1973: 501; Nutting, 1976: 214; Nutting & Desch, 1978: 140; Desch & Hillier, 2003: 148; Domrow, 1991: 1340; Izdebska & Rolbiecki, 2020: 12, many other authors
Demodex folliculorum var. canis: Railliet, 1893: 635
Demodex folliculorum var. caninus Mégnin, 1877: 111
Demodex caninus: Canestrini, 1899: 23; Hirst, 1917: 232
Type locality: Europe
Distribution: cosmopolitic
Host: Canis lupus familiaris (Carnivora: Canidae)
Demodex folliculorum var. caprae: Railliet, 1893; 1895: 638; Tenquist & Charleston, 2001: 495
Demodex caprae: Canestrini & Kramer, 1899: 3; Hirst, 1919: 38; Cooreman, 1946: 2; Dubinin, 1957: 132; Desch & Nutting, 1973: 501; Lebel & Nutting, 1973b: 517; Nutting, 1976: 214; Lebel & Desch, 1979; Tsedev & Grebenyuk, 1975: 127; Domrow, 1991: 1341; Izdebska & Rolbiecki, 2020: 12
Type locality: Europe
Distribution: range of host
Host: Capra hircus (Artiodactyla: Bovidae, Caprinae)
Demodex carolliae Desch, Lebel, Nutting & Lukoschus, 1971: 303; Izdebska & Rolbiecki, 2020: 13
Type locality: Lelydorp (Suriname)
Distribution: Suriname
Host: Carollia perspicillata (Chiroptera: Phyllostomatidae)
Demodex castoris Izdebska, Fryderyk & Rolbiecki, 2016: 1; Izdebska & Rolbiecki, 2020: 13
Type locality: Poland
Distribution: Poland
Host: Castor fiber (Rodentia: Castoridae)
Demodex folliculorum var. cati Mégnin, 1877: 112; Railliet, 1893: 637
Demodex cati: Canestrini & Kramer, 1899: 3; Hirst, 1919: 26; Dubinin, 1957: 132; Nutting, 1976: 214; Desch & Nutting, 1979b: 281; Domrow, 1991: 1341; Yathiraj et al., 1994: 393; Löwenstein, Beck, Bessmann & Mueller, 2005: 290; Izdebska & Rolbiecki, 2020: 13
Type locality: Europe
Distribution: range of hosts
Host: Felis catus (Carnivora: Felidae)
Demodex caviae Bacigalupo & Roveda, 1954: 149; Izdebska & Rolbiecki, 2020: 13
Type locality: laboratory animal
Distribution: Argentina
Host: Cavia porcellus (Rodentia: Caviidae)
Demodex cervi Prietsch, 1886; Vanselow, 1910; Kutzer & Grunberg, 1972: 9; Izdebska & Rolbiecki, 2020: 13
Type locality: Austria
Distribution: Austria
Host: Cervus elaphus (Artiodactyla: Cervidae)
Demodex chiropteralis Hirst, 1921: 367; Fain, 1960: 81; Baker & Craven, 2003: 11; Izdebska & Rolbiecki, 2020: 13
Type locality: England
Distribution: England, Poland
Host: Plecotus auritus (Chiroptera: Vespertilionidae)
Demodex conicus Izdebska & Rolbiecki, 1915: 168
Type locality: Poland
Distribution: Poland
Host: Mus musculus (Rodentia: Muridae)
Demodex cornei Izdebska & Rolbiecki, 2018: 346 (Mason. 1993: 92 - nomen nudum)
Type locality: Poland
Distribution: Poland, India, probably cosmopoliotic
Host: Canis lupus familiaris (Carnivora: Canidae)
Demodex criceti Nutting & Rauch, 1958: 328; Nutting, 1961: 515; Sarashina & Sato, 1986: 619; Izdebska & Rolbiecki, 2020: 13
Type locality: laboratory Golden Hamsters, Amherst, Mass.
Distribution: ? [USA]
Host: Mesocricetus auratus (Rodentia: Cricetidae)
Demodex cricetuli Hurley & Desch, 1994: 529; Izdebska & Rolbiecki, 2020: 13
Type locality: Massachsetts
Distribution: USA (Massachusetts)
Host: Cricetulus migratorius (Rodentia: Cricetidae)
Demodex cuniculi Pfeiffer, 1903; Izdebska & Rolbiecki, 2020: 14
Type locality: Europe
Distribution: cosmopolite
Host: Oryctolagus cuniculus (Lagomorpha: Leporidae)
Demodex cyonis Morita, Ohmi, Kiwaki, Ike & Nagata, 2018: 323; Izdebska & Rolbiecki, 2020: 14
Type locality: Japan
Distribution: Japan
Host: Canis lupus familiaris (Carnivora: Canidae)
Demodex dasypodi Desch & Stewart, 2002: 169; Izdebska & Rolbiecki, 2020: 14
Type locality: Louisiana
Distribution: USA
Host: Dasypus novemcinctus (Cingulata: Dasypodidae)
Demodex desmodi Desch, 1994: 39; Izdebska & Rolbiecki, 2020: 14
Type locality: Suriname
Distribution: Suriname
Host: Desmodus rotundus (Chiroptera: Phyllostomatidae, Desmodontinae)
Demodex folliculorum var. equi Railliet, 1893: 639; Bennison, 1943: 36
Demodex equi: Canestrini & Kramer, 1899: 3; Hirst, 1919; Dubinin, 1957: 132; Kolev, 1958: 299; Domrow, 1991: 1341; Izdebska & Rolbiecki, 2020: 14
Demodex sp. – Nutting, 1976: 214
Demodex caballi Railliet, 1895; Desch & Nutting, 1979: 235; Tenquist & Charleston, 2001: 495. Syn. by Domrow, 1991: 1341
Type locality: Europe
Distribution: cosmopolitic
Host: Equus caballus (Perissodactyla: Equidae)
Demodex folliculorum var. erinacei Hirst, 1919: 25; Izdebska & Rolbiecki, 2020: 14
Type locality: England
Distribution: England
Host: Erinaceus europaeus (Erinaceomorpha: Erinaceidae)
Demodex ermineae Hirst, 1919: 32
Type locality: Exeter (England)
Distribution: range of host
Host: Mustela erminea (Carnivora: Mustelidae)
Host: Felis catus (Carnivora: Felidae)]
Demodex flagellurus Bukva, 1985: 73; Izdebska, 2000; Izdebska & Rolbiecki, 2006: 175; 2020: 14
Type locality: České Budějovice, South Bohemia
Distribution: Czech Rep., Poland
Host: Mus musculus (Rodentia: Muridae)
Acarus folliculorum Simon, 1842: 231 (partim, figs. 1, 2, 8)
Demodex folliculorum (Simon): Haller, 1882: 296 (ex Canis familiaris!); Canestrini & Kramer, 1899: 2; Poppe, 1909: 66; Starkoff & Starkoff, 1950: 92; Lamb, 1952: 372; Desch & Nutting, 1972: 169; 1978: 422; Xie, Liu, Hsu & Hseu, 1982: 268; Candotti et al., 1996; Domrow, 1991: 1342; Zeytun & Karakurt, 2019: 2; Izdebska & Rolbiecki, 2020: 14 (also many other authors)
Demodex folliculorum longus Akbulatova, 1963: 40
Demodex hominis Leydig, 1859: 345
Type locality: Europe
Distribution: cosmopolitan
Host: Homo sapiens (Primates: Hominidae)
Demodex folliculorum sinensis Xie, Liu, Hsu & Hsu, 1982: 267; Izdebska & Rolbiecki, 2020: 14
Type locality: Shanghai
Distribution: China
Host: Homo sapiens (Primates: Hominidae)
Demodex foveolator Bukva, 1984: 45; Izdebska & Rolbiecki, 2020: 15
Type locality: České Budĕjovice, South Bohemia
Distribution: Czech Rep.
Host: Crocidura suaveolens (Soricomorpha: Soricidae)
Demodex fusiformis Izdebska & Rolbiecki, 2015: 608; Izdebska & Rolbiecki, 2020: 15
Type locality: Poland
Distribution: Poland
Host: Mus musculus (Rodentia: Muridae)
Demodex gapperi Nutting, Emejuaiwe & Tisdel, 1971: 660; Nutting, Beekman & Snyder, 1978: 646; Izdebska & Rolbiecki, 2020: 15
Type locality: New York State, Massachusetts
Distribution: USA
Host: Myodes [Clethrionomys] gapperi (Rodentia: Arvicolidae)
Demodex gatoi Desch & Stewart, 1999: 167
Type locality: Baton Rouge (USA)
Distribution: USA (Louisiana), Austria, Finland, Poland, Spain
Host: Felis catus [silvestris] (Carnivora: Felidae)
Demodex ghanensis Oppong, Lee & Yasin, 1975: 39; Abu-Samra & Shuaib, 2014: 77; Izdebska & Rolbiecki, 2020: 15
Type locality: Ghana
Distribution: Ghana, Sudan
Host: Bos taurus (Artiodactyla: Bovidae)
Demodex arvicolae var. glareoli Hirst, 1919: 35
Demodex glareoli Hirst: Izdebska, Kozina & Gólcz, 2013: 129; Izdebska & Rolbiecki, 2020: 15
Type locality: England
Distribution: England
Host: Myodes [Clethrionomys] glareolus britannicus (Rodentia: Arvicolidae)
Demodex gliricolens Hirst, 1921: 368; Izdebska & Rolbiecki, 2020: 15
Type locality: England (?)
Distribution: England (?)
Host: Arvicola terrestris (Rodentia: Arvicolidae)
Demodex gracilentus Izdebska & Rolbiecki, 2013: 1203; 2020: 15
Type locality: Poland
Distribution: Poland
Host: Apodemus agrarius (Rodentia: Muridae)
Demodex huttereri Mertens, Lukoschus & Nutting, 1983: 489; Izdebska & Rolbiecki, 2020: 15
Type locality: laboratory animal captured near Fulda, Germany
Distribution: Germany, Poland
Host: Apodemus agrarius (Mammalia: Muridae)
Demodex injai Desch & Hillier, 2003: 146; Izdebska & Rolbiecki, 2020: 15
Type locality: Columbus, Ohio
Distribution: USA (Ohio)
Host: Canis lupus familiaris (Carnivora: Canidae)
Demodex intermedius Lukoschus, Mertens, Nutting & Nadchatram, 1984: 234; Izdebska & Rolbiecki, 2020: 16
Type locality: Bukit Fraser
Distribution: Malaysia
Host: Tupaia glis (Scandentia: Tupaiidae)
Demodex kutzeri Bukva, 1987: 175; Kadulski, 1996a: 204; 1996b: 353; Desch et al., 2010: 585; Izdebska & Rolbiecki, 2020: 16
Demodex cervi sensu Kutzer & Grünberg, 1972, part.; nec D. cervi Vanselow, 1910
Type locality: Wattens/Tirol
Distribution: Austria, USA
Hosts: Alces alces, Capreolus capreolus, Cervus elaphus, C. nelsoni, C. nippon pseudaxis, Dama dama, Odocoileus h. hemionus, O. virginianus (Artiodactyla: Cervidae)
Demodex lacrimalis Lukoschus & Jongman, 1974: 274; Izdebska & Rolbiecki, 2020: 16
Type locality: Nijmegen
Distribution: Holland, Italy, Poland
Host: Sylvaemus sylvaticus (Rodentia: Muridae, Murinae)
Demodex leucogasteri Hughes & Nutting, 1981: 182; Izdebska & Rolbiecki, 2020: 16
Type locality: Washington State, Tennessee
Distribution: USA (Washington State, Tennessee)
Host: Onychomys leucogaster (Rodentia: Arvicolidae, Neotominae)
Demodex longior Hirst, 1918: 145; 1919: 25; Izdebska & Rolbiecki, 2020: 16
Type locality: England
Distribution: England
Host: Sylvaemus sylvaticus (Rodentia: Muridae)
Demodex longissimus Desch, Nutting & Lukoschus, 1972: 35; Izdebska & Rolbiecki, 2020: 16
Type locality: Brokopondo, Brownsweg, Lelydorp, Onverwacht, Zandery
Distribution: Suriname
Host: Carollia perspicillata (Chiroptera: Phyllostomatidae)
Demodex lutrae Izdebska & Rolbiecki, 2014: 784; Izdebska & Rolbiecki, 2020: 17
Type locality: Poland
Distribution: Poland
Host: Lutra lutra (Carnivora: Mustelidae)
Demodex macaci Karjala, Desch & Starost, 2005: 948; Izdebska & Rolbiecki, 2020: 17
Type locality: Laboratory colony
Distribution: laboratory colony
Host: Macaca mulatta (Primates: Cercopithecidae)
Demodex macroglossi Desch, 1981: 41; Domrow, 1991: 1342; Izdebska & Rolbiecki, 2020: 17
Type locality: Western Australia
Distribution: Australia (W.A.)
Host: Macroglossus minimus (Chiroptera: Pteropodidae)
Demodex marculus Izdebska & Rolbiecki, 2015: 607, 2020: 17
Type locality: Poland
Distribution: Poland
Host: Mus musculus (Rodentia: Muridae)
Demodex marsupiali Nutting, Lukoschus & Desch, 1980: 84; Izdebska & Rolbiecki, 2020: 17
Type locality: Lelydorp, Paramaribo, Coronie
Distribution: Suriname
Host: Didelphis marsupialis (Didelphimorphia: Didelphidae)
Demodex melanopteri Lukoschus, Jongman & Nutting, 1972: 54; Izdebska & Rolbiecki, 2020: 17
Type locality: Meerzorg, Lelydorp
Distribution: Suriname
Host: Eptesicus brasiliensis melanopterus (Chiroptera: Vespertilionidae)
Demodex melesinus Hirst, 1921: 372; Izdebska & Rolbiecki, 2020: 17
Type locality: England
Distribution: England
Host: Meles meles (Carnivora: Mustelidae)
Host: Meriones spp. (Rodentia: Muridae)]
Demodex mexicanus Vargas, Bassols, Desch, Quintero & Polaco, 1995: 76; Izdebska & Rolbiecki, 2020: 17
Type locality: Mexico
Distribution: Mexico
Host: Plecotus [Corynorhinus] mexicanus (Chiroptera: Vespertilionidae)
Demodex microti Izdebska & Rolbiecki, 2013: 189; Izdebska & Rolbiecki, 2020: 17
Type locality: Poland
Distribution: Poland
Host: Microtus arvalis (Rodentia: Arvicolidae)
Demodex mollis Izdebska, Rolbiecki, Fryderyk & Mierzyński, 2017; Izdebska & Rolbiecki, 2020: 18
Type locality: Poland
Distribution: Poland
Host: Sylvaemus flavicollis (Rodentia: Muridae)
Demodex molossi Desch, Nutting & Lukoschus, 1972 : 43; Izdebska & Rolbiecki, 2020: 18
Type locality: Lelydorp, Paramaribo
Distribution: Suriname
Host: Molossus molossus (Chiroptera: Molossidae)
Demodex muscardini Hirst, 1917: 433; 1919: 30; Bregetova et al., 1955; Izdebska & Rolbiecki, 2020: 18
Type locality: England
Distribution: Armenia, Great Britain
Host: Muscardinus avellanarius (Rodentia: Gliridae)
Demodex folliculorum var. musculi Oudemans, 1897: 269
Demodex musculi Oudemans, 1905: 12; 1909: 318; Hirst, 1917: 233; Hill et al., 1999: 13; Izdebska & Rolbiecki, 2020: 18
Demodex arvicolae var. musculi: Hirst, 1919: 34; Dubinin, 1953: 276
Demodex arvicolae musculi Oudemans: Izdebska & Rolbiecki, 2006: 175
Type locality: Europe
Distribution: England, Poland, Russia, Spain
Host: Mus musculus (Rodentia: Muridae)
Demodex myotidis Di Benedetto, 1961 (MA Thesis); Izdebska & Rolbiecki, 2020: 18
Type species: USA
Distribution: USA
Hosts: Eptesicus fuscus, Myotis l. lucifugus, M. septentrionalis (Chiroptera: Vespertilionidae). Note by Izdebska & Rolbiecki, 2020: “Nom. nud.; description not published within the meaning of the ICZN”].
Demodex mystacina Desch, 1989: 221; Izdebska & Rolbiecki, 2020: 18
Type locality: Omahuta Kauri Sanctuary
Distribution: New Zealand
Host: Mystacina tuberculata (Chiroptera: Mystacinidae)
Demodex nanus Hirst, 1918: 146; Desch, 1987: 19; 1989: 228; Bukva, 1993; Haitlinger & Jankowska, 2005: 38; Izdebska & Rolbiecki, 2020: 18
Type locality: unknown
Distribution: Great Britain, Poland, Russia
Hosts: Rattus norvegicus, R. rattus (Rodentia: Muridae, Murinae)
Demodex neomydis Bukva, 1995: 299; Izdebska & Rolbiecki, 2020: 18
Type locality: České Budĕjovice, South Bohemia
Distribution: Czech Rep.
Host: Neomys anomalus (Soricomorpha: Soricidae)
Demodex neoopisthosomae Desch, Lukoschus & Nadchatram, 1986: 14; Izdebska & Rolbiecki, 2020: 19
Type locality: Malaysia
Distribution: Malaysia
Host: Eonycteris spelaea (Chiroptera: Pteropodidae, Macroglossinae)
Demodex norvegicus Bukva, 1995: 152; Haitlinger & Jankowska, 2005: 38; Izdebska & Rolbiecki, 2020: 19
Type locality: České Budĕjovice, South Bohemia
Distribution: Czech Rep., Poland
Host: Rattus norvegicus (Rodentia: Muridae, Murinae)
Demodex novazelandica Desch, 1989: 226; Izdebska & Rolbiecki, 2020: 19
Type locality: Omahuta Kauri Sanctuary
Distribution: New Zealand
Host: Mystacina tuberculata (Chiroptera: Mystacinidae)
Demodex nycticeii Desch, 1996: 187; Izdebska & Rolbiecki, 2020: 19
Type locality: Florida
Distribution: USA (Florida)
Host: Nycticeius humeralis (Chiroptera: Vespertilionidae)
Demodex odocoilei Desch & Nutting, 1974: 785; Jasques et al., 2001: 221; Gentes, Proctor & Wobeser, 2007: 758; Turner & Cano, 2008: 572; Izdebska & Rolbiecki, 2020: 19
Type locality: Dougherty County (Georgia), Eucha (Oklahoma), Stafford County (Virginia)
Distribution: Canada, USA (Georgia, Oklahoma, Virginia, Texas)
Host: Odocoileus hemionus columbianus, O. virginianus (Artiodactyla: Cervidae)
Demodex folliculorum var. ovis Railliet, 1895: 638; Hirst, 1919: 24; Bukva, 1990: 82; Tenquist & Charleston, 2001: 497
Demodex ovis Railliet: Hallas, 1978: 30; Izdebska & Rolbiecki, 2020: 19
Type locality: Europe
Distribution: range of host (incl. Australia, New Zealand, Czech Rep., Israel, Poland)
Host: Ovis aries (Artiodactyla: Bovidae)
Demodex peromysci Lombert, Lukoschus & Whitaker, 1983: 377; Izdebska & Rolbiecki, 2020: 19
Type locality: Pennsylvania, Huntingdon Co.
Distribution: USA (Pennsylvania)
Host: Peromyscus leucopus (Rodentia: Cricetidae)
Demodex phocidi Desch, Dailey & Tuomi, 2003: 231; Izdebska et al., 2020: 49; Izdebska & Rolbiecki, 2020: 19
Type locality: Alaska SeaLife Center, Seward
Distribution: USA (Alaska), Poland
Host: Phoca vitulina (Carnivora Pinnipedia: Phocidae)
Demodex phodopi Desch, Davis & Klompen, 2006: 75; Izdebska & Rolbiecki, 2020: 19
Type locality: laboratory colony
Distribution: [USA](actually, Asia)
Host: Phodopus sungorus (Rodentia: Cricetidae)
Demodex phylloides Csokor, 1879: 133; Canestrini, 1899: 968; Canestrini & Kramer, 1899: 3; Hirst, 1919: 27; Lamb, 1952: 372; Dubinin, 1957: 132; Desch & Nutting, 1973: 500; Rak, 1976: 509; Quintero, 1977: 42; Tenquist & Charleston, 2001: 497; Desch & Stewart, 2002: 173; Reeves et al., 2007: 47; Izdebska & Rolbiecki, 2020: 19
Demodex folliculorum var. suis Railliet, 1895: 637; Ligrain & Régulato, 1903: 371
Distribution: Algeria, Argentina, Australia, Brazil, Canada, France, Germany, Iran, Madagascar, Mexico, Poland, Russia, New Zealand, Spain, Sweden, USA
Host: Sus scrofa (Artiodactyla: Suidae)
Demodex phyllostomatis Leydig, 1859: 345; Hirst, 1919: 39; Fain, 1960: 80; Izdebska & Rolbiecki, 2020: 20
Type locality: Suriname
Distribution: Suriname
Host: Phyllostoma hastatum (Chiroptera: Phyllostomatidae)
Demodex plecoti Izdebska, Rolbiecki & Mienzynski, 2019: 377.
Type locality: Poland
Distribution: Poland
Host: Plecotus auritus (Chiroptera: Vespertilionidae)
Demodex ponderosus Izdebska & Rolbiecki, 2014: 740
Type locality: Poland
Distribution: Poland
Host: Rattus norvegicus (Rodentia: Muridae)
Demodex pseudaxisi Shpringol’ts – Shmidt, 1937: 133
Type locality: Far East of Russia
Distribution: Russia
Host: Cervus nippon hortulorum (Arthiodactyla: Cervidae)
Demodex ratti Hirst, 1917: 233; 1919: 37; Dubinin, 1953: 276; Bukva, 1995: 149; Haitlinger & Jankowska, 2005: 38
Type locality: Europe; redescription from Vranín, South Bochemia
Distribution: Czech Rep., England, Poland, Russia
Host: Rattus norvegicus (Rodentia: Muridae)
Demodex ratticola Bukva, 1995: 156; Izdebska & Rolbiecki. 2012b: 71
Type locality: České Budějovice
Distribution: Czech Rep., Poland
Host: Rattus norvegicus (Rodentia: Muridae)
Demodex rosus Bukva, Vítovec & Vlček, 1985: 151
Type locality: Lišov, distr. of České Budějovice
Distribution: Czech Rep.
Host: Sylvaemus flavicollis (Rodentia: Muridae)
Demodex sabani Desch, Lukoschus & Nadchatram, 1984: 55
Type locality: Kuala Selangor Road, 35 km from Selangor, Malaysia
Distribution: Malaysia
Hosts: Leopoldamys sabanus, L. edwardsi, Niviventer cremoriventer, N. rapit, Rattus annandalei, R. jalorensis, Sundamys mulleri (Rodentia: Muridae)
Demodex saimiri Lebel & Nutting, 1973: 719; Izdebska & Rolbiecki, 2020: 21
Type locality: Zoo
Distribution: South America
Host: Saimiri sciureus (Primates: Cebidae, Saimiriinae)
Demodex sciurei Lebel, 1970: Master’s Thesis
Type locality: Zoo
Distribution: South America
Host: Saimiri sciureus (Primates: Cebidae, Saimiriinae). Note by Izdebska et al., 2020: “Nom. nud.; description not published within the meaning of the ICZN”].
Demodex sciurinus Hirst, 1923: 994; Izdebska & Rolbiecki, 2020: 3
Type locality: Inverness (Great Britain)
Distribution: Great Britain, Poland
Host: Sciurus vulgaris (Rodentia: Sciuridae, Sciurinae)
Demodex sinocricetuli Desch & Hurley, 1997: 317; Izdebska & Rolbiecki, 2020: 21
Type locality: [Maryland, Beltsville]
Distribution: [USA (Maryland), laboratory animals]
Host: Cricetulus barabensis (Rodentia: Cricetidae)
Demodex soricinus Hirst, 1918: 145; 1919: 37; 1921: 368; Dubinin, 1953: 276; Fain, 1960: 81; van Eyndhoven, 1965: 103; Bukva, 1993: 243; Izdebska, 2004: 47; Izdebska & Rolbiecki, 2020: 21
Type locality: Great Britain
Distribution: Czech Republic, Great Britain, Holland, Poland, Russia
Hosts: Sorex araneus (Soricomorpha: Soricidae), Plecotus auritus (?, probably wrong host record)(Chiroptera: Vespertilionidae)
Demodex spelaea Desch, Lukoschus & Nadchatram, 1986: 20; Desch, 1989: 226; Izdebska & Rolbiecki, 2020: 21
Type locality: Malaysia
Distribution: Malaysia
Host: Eonycteris spelaea (Chiroptera: Pteropodidae, Macroglossinae)
Demodex sungori Desch, Davis & Klompen, 2006: 77; Izdebska & Rolbiecki, 2020: 21
Type locality: laboratory colony
Distribution: [USA](actually, Asia)
Host: Phodopus sungorus (Rodentia: Cricetidae)
Demodex sylvilagi Maravelas, 1962: MA Thesis
Type locality: New England
Distribution: USA
Host: Sylvilagus transitionalis (Lagomorpha: Leporidae). Note by Izdebska et al., 2020: “Nom. nud.; description not published within the meaning of the ICZN”].
Demodex talpae Hirst, 1921: 370; Izdebska & Rolbiecki, 2003: 97, 2020: 21
Type locality: Great Britain
Distribution: Great Britain, Poland
Host: Talpa europaea (Soricomorpha: Talpidae)
Demodex tauri Bukva, 1986: 364; Izdebska & Rolbiecki, 2020: 21
Type locality: České Budĕjovice
Distribution: Czech Rep.
Host: Bos taurus (Artiodactyla: Bovidae, Bovinae)
Demodex tigris Shi, Xie & Hsu, 1985: 385; Izdebska & Rolbiecki, 2020: 22
Type locality: Shanghai Zool. Garden, Shanghai
Distribution: China
Host: Panthera tigris amoyensis (Carnivora: Felidae)
Demodex tortellinioides Desch & Holz, 2006: 81; Izdebska & Rolbiecki, 2020: 22
Type locality: Victoria
Distribution: Australia
Host: Antechinus agilis (Dasyuromorphia: Dasyuridae)
Demodex transitionalis Moravelas, 1962
Type locality: USA
Distribution: USA
Host: Sylvilagus transitionalis (Lagomorpha: Leporidae). Note by Izdebska et al., 2020: “Nom. nud.; description not published within the meaning of the ICZN”].
Demodex uncii Desch, 1993: 63; Izdebska & Rolbiecki, 2020: 22
Type locality: Zoo
Distribution: Asia (collected in Zoo in USA)
Host: Uncia uncia (Carnivora: Felidae)
Demodex ursi Desch, 1995: 23; Foster, Cames & Forrester, 1998: 161; Izdebska & Rolbiecki, 2020: 22
Type locality: Florida
Distribution: USA (Florida)
Host: Ursus americanus (Carnivora: Ursidae)
Demodex vibrissae Izdebska, Rolbiecki & Fryderyk, 2016; Izdebska & Rolbiecki, 2020: 22
Type locality: Poland
Distribution: Poland
Host: Mus musculus (Rodentia: Muridae)
Demodex zalophi Dailey & Nutting, 1980: 424; Izdebska & Rolbiecki, 2020: 22
Type locality: San Diego (California), Arlington (Texas) and Australia
Distribution: captive sea lions in California, Texas and Australia
Host: Zalophus californianus (Carnivora Pinnipedia: Otariidae)
Glossicodex Izdebska & Rolbiecki, 2016: 135
Type species: Glossicodex musculi Izdebska et Rolbiecki, 2016
Glossicodex musculi Izdebska & Rolbiecki, 2016: 135; Izdebska & Rolbiecki, 2020: 22
Type locality: Poland
Distribution: Poland
Host: Mus musculus (Rodentia: Muridae)
Ophthalmodex Lukoschus & Nutting, 1979: 299
Type species: Ophthalmodex artibei Lukoschus et Nutting, 1979
Ophthalmodex apodemi Bukva, Nutting & Desch, 1992: 269; Izdebska & Rolbiecki, 2020: 22
Type locality: Česke Budejovice, South Bohemia
Distribution: Czech Rep.
Host: Sylvaemus sylvaticus (Rodentia: Muridae, Murinae)
Ophthalmodex artibei Lukoschus & Nutting, 1979: 301; Izdebska & Rolbiecki, 2020: 22
Type locality: Lelydorp
Distribution: Suriname
Host: Artibeus lituratus (Chiroptera: Phyllostomatidae, Stenodermatinae)
Opthalmodex australiensis Woeltjes & Lukoschus, 1981: 308 (lapsus)
Ophthalmodex australiensis: Domrow, 1991: 1344; Izdebska & Rolbiecki, 2020: 22
Type locality: Western Australia
Distribution: Australia
Host: Rhinonicteris aurantius (Chiroptera: Hipposideridae)
Ophthalmodex carolliae Lukoschus, Woeltjes, Desch & Nutting, 1980b: 45; Izdebska & Rolbiecki, 2020: 22
Type locality: Lelydorp, Zanderij, Onverwacht, Brownsweg
Distribution: Suriname
Host: Carollia perspicillata (Chiroptera: Phyllostomatidae, Carolliinae)
Ophthalmodex juniatae Veal, Giesen & Whitaker, 1984: 348; Izdebska & Rolbiecki, 2020: 23
Type locality: Entriken, Huntingdon Co, Penn.
Distribution: USA (Pennsylvania)
Host: Myotis lucifugus (Chiroptera: Vespertilionidae, Vespertilioninae)
Ophthalmodex molossi Lukoschus, Woeltjes, Desch & Nutting, 1980b: 47; Izdebska & Rolbiecki, 2020: 23
Type locality: Paramaribo
Distribution: Suriname
Host: Molossus molossus (Chiroptera: Molossidae)
Ophthalmodex wilsoni Woeltjes & Lukoschus, 1981: 311; Domrow, 1991: 1344; Izdebska & Rolbiecki, 2020: 23
Type locality: Western Australia
Distribution: Australia
Host: Vespadelus pumilus (Chiroptera: Vespertilionidae, Vespertilioninae)
Pterodex Lukoschus, Woeltjes, Desch & Nutting, 1980: 9
Type species: Pterodex carolliae Lukoschus, Woeltjes, Desch et Nutting, 1980
Pterodex carolliae Lukoschus, Woeltjes, Desch & Nutting, 1980a: 10; Izdebska & Rolbiecki, 2020: 23
Type locality: Lelydorp, Zanderij, Onverwacht, Brownsweg
Distribution: Suriname
Host: Carollia perspicillata (Chiroptera: Phyllostomatidae, Carolliinae)
Rhinodex Fain, 1959: 264
Type species: Rhinodex baeri Fain, 1959
Rhinodex baeri Fain, 1959: 264; Izdebska & Rolbiecki, 2020: 23
Type locality: Bugesera
Distribution: Rwanda
Host: Galago moholi (Primates: Galagonidae)
Soricidex Bukva, 1982 : 343
Type species: Soricidex dimorphus Bukva, 1982
Soricidex dimorphus Bukva, 1982: 344; Izdebska, 2004a: 47; Izdebska & Rolbiecki, 2020: 23
Type locality: Česke Budejovice, South Bohemia
Distribution: Czech Rep., Poland
Host: Sorex a. araneus (Soricomorpha: Soricidae, Soricinae)
Soricidex sp.: Bukva, 1996: 315
Locality: Česke Budejovice, South Bohemia
Distribution: Czech Rep.
Host: Neomys anomalus (Soricomorpha: Soricidae)
Stomatodex Fain, 1959: 267
Type species: Stomatodex galagoensis Fain, 1959
Stomatodex cercarteti Desch, 1991: 239; Domrow, 1992: 1606; Halliday, 1998: 49; Izdebska & Rolbiecki, 2020: 23
Type locality: New South Wales
Distribution: Australia
Host: Cercartetus nanus (Diprotodontia: Burramyidae)
Stomatodex corneti Fain, 1960: 84; Baker & Craven, 2003: 12; Izdebska & Rolbiecki, 2020: 23
Type locality: Grottes de Han (Belgium)
Distribution: Belgium, Great Britain, Rwanda
Host: Barbastella barbastellus (Chiroptera: Vespertilionidae), ?Nycteris sp. (Nycteridae)
Stomatodex corneti myotis Fain, 1960: 86; Izdebska & Rolbiecki, 2020: 23
Type locality: Grottes de Han (Belgium)
Distribution: Belgium
Hosts: Myotis dasycneme, M. myotis (Chiroptera: Vespertilionidae)
Stomatodex galagoensis Fain, 1959: 268; Izdebska & Rolbiecki, 2020: 23
Type locality: Bugesera
Distribution: Rwanda
Host: Galago moholi (Primates: Galagonidae)
Stomatodex rousetti Fain, 1960: 83; Izdebska & Rolbiecki, 2020: 24
Type locality: Mahyusha Cave, nr. Katana
Distribution: DR Congo
Host: Rousettus aegyptiacus leachi (Chiroptera: Pteropodidae)
EUROPE
Armenia (Bregetova et al., 1955) – Demodex muscardini
Austria (Kutzer & Grünberg, 1972; Bukva, 1987) – Demodex cervi, D. kutzeri
Azerbaidzhan (Gadzhiev et al., 1988) – Demodex bovis, D. canis, D. cuniculi, D. folliculorum, D. phylloides
Belgium (Fain, 1960; de Bosschere et al., 2007) – Stomatodex c. corneti, S. corneti myotis
Bulgaria (Drenski, 1921; Kolev, 1958; Hristov & Mihailov, 1988; Nedelchev, Rusev & Vassileva, 2001; Iliev, Zhelev, Ivanov & Prelezov, 2019) – Demodex aurati, D. canis, D. cati, D. caviae, D. equi
Czech Rep. (Daniel, Bozdech & Moucka, 1959; Bukva, 1982, 1984, 1985a, 1985b, 1986, 1993a, 1993b, 1994, 1995a, 1995b, 1996; Bukva, Nutting & Desch, 1992; Fleischer et al., 1996) – Apodemodex cornutus, Demodex aries, D. buccalis, D. flagellurus, D. folliculorum, D. foveolator, D. neomydis, D. norvegicus, D. ovis, D. ratti, D. ratticola, D. rosus, D. soricinus, D. tauri; Ophthalmodex apodemi, Soricidex dimorphus
Denmark (Hallas, 1978) – Demodex brevis, D. canis, D. equi, D. folliculorum, D. ovis
Finland (Saari et al., 2009) – Demodex gatoi
France (contin.)(Hirst, 1919; 1921) – Demodex bovis, D. equi, D. phylloides
Germany (Haller, 1882; Poppe, 1909; Hirst, 1919; Hoffman & Hiepe, 1987) – Demodex bovis, D. equi, D. folliculorum, D. phylloides
Great Britain (Hirst, 1917, 1919; 1921, 1923; Brownlee, 1935; Turk, 1953; Baker & Craven, 2003) – Demodex arvicolae, D. bovis, D. canis, D. cati, D. chiropteralis, D. cuniculi, D. equi, D. erinacei, D. ermineae, D. folliculorum, D. melesinus, D. ovis, D. ratti, D. sciurinus, D. soricinus, Stomatodex corneti
Greece (Himonas, Theodorides & Alexakis, 1975) – Demodex spp.
Holland (van Eyndhoven, 1965; Lukoschus & Jongman, 1974) – Demodex lacrimalis, D. soricinus
Hungary (Dudich et al., 1940) – Demodex folliculorum
Ireland (Baker, 1970, 1973) – Demodex canis
Italy (continent.) (Starkoff & Starkoff, 1950; Lukoschus & Jongman, 1974; Candotti et al., 1996) – Demodex canis, D. folliculorum, D. lacrimalis
Norway (Mehl, 1979) – Demodex bovis, D. canis, D. cati, D. caprae, D. equi, D. folliculorum
Poland (Kadulski, 1978, 1996a, 1996b, 1997; Kadulski & Izdebska, 1996; Kadulski, Izdebska & Konczyk, 1996; Fryderyk & Izdebska, 2001; Izdebska & Fryderik, 2000, 2002, 2011, 2012a, 2012b; Izdebska & Rolbiecki, 2002, 2003, 2004, 2006, 2012a, 2012b, 2013a, 2013b, 2014, 2015a, 2015b, 2016, 2018, 2020; Izdebska, 2000a, 2000b, 2001a, 2001b, 2001c, 2001d, 2002, 2004a, 2004b, 2005, 2006a, 2006b, 2006c, 2007, 2008, 2009, 2012; Izdebska, Rolbiecki & Fryderik, 2011, 2014, 2016; Haitlinger & Jankowska, 2005; Izdebska & Jankowski, 2006; Czepita et al., 2007; Izdebska & Cydzik, 2010; Izdebska & Krawczyk, 2012) – Demodex acutipes, D. agrarii, D. apodemi, D. auricularis, D. musculi, D. bisonianus, D. bovis, D. brevis, D. buccalis, D. canis, D. caprae, D. castoris, D. cati, D. chiropteralis, D. conicus, D. cornei, D. corniculatus, D. equi, D. flagellurus, D. folliculorum, D. fusiformis, D. gatoi, D. glareoli, D. gracilentus, D. injai, D. kutzeri, D. lacrimalis, D. longior, D. lutrae, D. marculus, D. marsupiali, D. melesinus, D. microti, D. mollis, D. musculi, D. nanus, D. norvegicus, D. ovis, D. phocidi, D. phylloides, D. plecoti, D. ratti, D. ratticola, D. sciurinus, D. soricinus, D. talpae, D. vibrissae, Glossicodex musculi, Soricidex dimorphus
Romania (Stoenescu et al., 1972; May, Putin & Aciocîrlánoaia, 1975) – Demodex canis
Russia (all, continent.)(Isaychikov, 1928; Chernobayev, 1928; Rozgon, 1930; Shpringol’ts – Shmid, 1937; Dubinin, 1953; Klinski, 1956, 1959; Polyakov, 1956, 1977; Dubinina, 1987; Moskvina, 2017) – Demodex apodemi, D. aries, D. arvicolae, D. bovis, D. brevis, D. caballi, D. canis, D. caprae, D. cati, D. equi, D. folliculorum, D. musculi, D. nanus, D. ovis, D. phylloides, D. pseudaxisi
Slovakia (Bukva, 1994) – Demodex agrarii
Spain (cont.)(Hirst, 1919) – Demodex phylloides
Canary Islands (Ollé et al., 2020) – Demodex musculi
Sweden (Hist, 1919) – Demodex phylloides
Switzerland (Fain, 1960) – Demodex aelleni
Turkey (all)(Özdemir et al., 2003; Karaer et al., 2009; Pekmezci, Pekmezci & Bolukbas, 2018; Zeytun & Karakurt, 2019; Sari, Zeytun, Doğan & Karakurt, 2019) – Demodex brevis, D. canis, D. folliculorum
AFRICA
Algeria (Legrain & Régulato, 1903; Hist, 1919) – Demodex phylloides
Angola (Freire & Dias, 1947) – Demodex bovis
Botswana (Dräger & Paine, 1980) – Demodex cafferi
Congo DR (Hirst, 1919; Fain, 1960) – Demodex bovis, Stomatodex rousetti
Egypt (Salib, 2018) – Demodex
Ethiopia (Mulugeta, Yacob & Ashenafi, 2010; Chanie, Tadesse & Bogale, 2013) – Demodex bovis
Ghana (Oppong, 1970; Oppong, Lee & Yasin, 1975) – Demodex ghanensis
Libya (Gabaj et al., 1992) – Demodex sp.
Madagascar (Hist, 1919) – Demodex phylloides
Malawi (Griffiths, 1945) – Demodex bovis
Nigeria (Esuruoso, 1977; Slingerberg, Mohammed & Bida, 1980; George et al., 1992; Ugbomoiko, Ariza & Heukelbach, 2008) – Demodex canis, D. folliculorum
Rwanda (Fain, 1959, 1960) – Rhinodex baeri, Rh. galagoensis, Stomatodex c. corneti
South Africa (Du Toit & Fiedler, 1960; Nutting & Guilfoy, 1979; Wolhuter et al., 2009) – Demodex bovis, D. cafferi, D. canis, D. caprae, D. folliculorum, D. phylloides
Sudan (Abu-Samra, Hago & Musa, 1981; Abu-Samra, Imbabi & Mahgoub, 1981; Abu-Samra & Shuaib, 2014) – Demodex canis, D. ghanensis
Zambia (Mwase & Baker, 2006) – Demodex bovis, D. canis, D. folliculorum, D. phylloides
ASIA
Bangladesh (Noorudin & Rahman, 1985) – Demodex bovis
China (Zhang et al., 1982; Xie, Liu, Ye & Hsu, 1982; Shi, Xie & Hsu, 1985; Liu, Xu, Xie & Xu, 1988; Yehua Xu, Hexiu Xie, Sulan Liu, Zhongyong Zhou, Xinquan Shi, 1986; Chen, 199?; Fan et al., 1990; Lin, Wang & Hong, 2000; Liu, 2007; Zhao, Guo & Wu, 2009; Zhao, Cheng, Hu & Ma, 2014) – Demodex ailuropodae, D. brevis, D. caprae, D. folliculorum, D. sinensis, D. tigris
Cyprus (Cooreman, 1946) – Demodex canis, D. caprae
India (Hirst, 1919; Sreenivasan & Rizvi, 1946; Venkatesan, Nandy & Krishnan, 1979; Chakrabarti & Pardhan, 1985; Yathiraj et al., 1994; Nayak et all., 1997; putatunda, Kumar & Banerjee, 2004, Sivajothi et al., 2013) – Demodex canis, D. cati, D. caprae, D. cornei, D. ovis
Indonesia (Sangvaranond, 1979) – Demodex
Iran (Rafyi, Alavi & Rak, 1967; Rak, 1969, 1973, 1976; Rak & Rahgozar, 1975a, 1975b) – Demodex canis, D. ovis, D. phylloides
Israel and Palestina (Yeruham, Rosen & Hadani, 1986) – Demodex ovis
Japan (Tashiro et al., 1979; Sarashina & Sato, 1986; Morita, 2018) – Demodex aurati, D. criceti [cage], D. cyonis, D. bovis
Korea (Chee, Jeong-Hyun et al., 2008) – Demodex canis
Kuwait (Tarello, 2007) – Demodex sp.
Laos (Izdebska, 2017) – Demodex bandicotae
Malaysia (Beck, 1971; Lukoschus, Mertens, Nutting & Nadchatram, 1984; Desch, Lukoschus & Nadchatram, 1984, 1986; Dorny et al., 1994) – Demodex caprae, D. intermedius, D. neoopisthosomae, D. sabani, D. spelaea
Mongolia (Tsedev & Grebenyuk, 1975; Matthes & Bukva, 1993) – Demodex bovis, D. caprae
Saudi Arabia (Tarello, 2007) – Demodex sp.
Thailand (Tarello, 2007) – Demodex sp.
NORTH AMERICA
Canada (Hist, 1919; Smith, 1960, 1961a, 1961b; Gentes, Proctor & Wobeser, 2007) – Demodex odocoilei, D. phylloides
Mexico (Hoffmann et al., 1972; Quintero, 1977; Vargas, Bassols, Desch, Quintero & Polaco, 1995; Rodriguez-Vivas et al., 2003) – Demodex artibei, D. canis, D. folliculorum, D. mexicanus, D. phylloides
USA (Faxon, 1878; Stiles, 1892; Hirst, 1919; Cram, 1925; Baker & Nutting, 1950; Besch & Griffiths, 1956; Carpenter, Freeny & Patton, 1972; Desch & Nutting, 1974; Nutting, Beekman & Snyder, 1978; Dailey & Nutting, 1980; Hughes & Nutting, 1981; Lombert, Lukoschus & Whitaker, 1983; Veal, Giesen & Whitaker, 1984; Forrester, Spalding & Woodling, 1993; Hurley & Desch, 1994; Desch, 1995, 1996; Waggie & Marion, 1997; Foster, Cames & Forrester, 1998; Desch & Bonner, 1999; Jacques, Jenks, Hildreth, Schauer & Johnson, 2001; Desch & Stewart, 2002; Desch & Hillier, 2003; Bildfell et al., 2004; Jekl, Hauptman, Jeklova & Knotek, 2006; Reeves et al., 2007; Turner & Cano, 2008; Prasad, 2009; Desch et al., 2010; Yabsley et al., 2013) – Demodex bovis, D. brevis, D. canis, D. caprae, D. cricetuli, D. dasypodi, D. equi, D. folliculorum, D. gapperi, D. gatoi, D. injai, D. kutzeri, D. leucogasteri, D. nycticeii, D. odocoilei, D. peromysci, D. phylloides, D. ursi, D. zalophi, Ophthalmodex juniatae
SOUTH and CENTRAL AMERICA
Argentine (Hirst, 1919) – Demodex phylloides
Aruba (Sutmöller, 1957) – Demodex sp.
Brazil (Hirst, 1919; Rocha & Pardi, 1954; Faccini, Santos & Bechara, 2004; Santarém, Farias & Tostes, 2005; Cardoso & Franco, 2003) – Demodex bovis, D. phylloides
Chile (Crossley, Gorman & Alcaino, 1970; Cekalović, 1976: 37; Alcaino & Gorma, 1999) – Demodex canis
Colombia (Cardona, Vargas & Perdomo, 2013) – Demodex bovis
Cuba (Guerra et al., 2010) – Demodex spp.
Suriname (Leydig, 1859; Fain, 1960; Desch, Lebel, Nutting & Lukochus, 1971; Desch & Nutting, 1972; Lukoschus, Jongman & Nutting, 1972; Desch, Nutting & Lukoschus, 1972; Lukoschus & Nutting, 1979; Nutting, Lukoschus & Desch, 1980; Lukoschus, Woeltjes, Desch & Nutting, 1980a, 1980b) – Demodex carolliae, D. desmodi, D. longissimus, D. marsupiali, D. melanopteri, D. molossi, D. phyllostomatis, Ophthalmodex artibei, O. carolliae, O. molossi, Pterodex carolliae
Venezuela (Alvarez et al., 2007) – Demodex sp.
South America (Lebel & Nutting, 1973) – Demodex saimiri
OCEANIA
Australia (continent)(Carter, 1942; Nutting & Wooley, 1965; English, 1969; Nutting & Sweatman, 1970; Nutting & Green, 1976; Woeltjes & Lukoschus, 1981; Veal, Giesen & Whitaker, 1984; Kniest & Lukoschus, 1981; Desch, 1981, 1991; Holz, 1998; Domrow, 1991, 1992; Vogelnest et al., 2000; Desch & Holz, 2006) –Demodex antechini, D. bicaudatus, D. bovis, D. brevis, D. canis, D. caprae, D. cati, D. equi, D. folliculorum, D. macroglossi, D. ovis, D. phylloides, D. tortellinioides, D. zalophi, Demodex sp., Ophthalmodex australiensis, O. wilsoni, Stomatodex cercarteti
New Zealand (Thomson, 1922; Nelson, 1956; Sweatman, 1962; Whitten, 1962; Nutting, Kettle, Tenquist & Whitten, 1975; Andrews, 1982; Desch, 1986, 1989; Tenquist & Charleston, 2001; Hill, McKenna & Mirams, 2008) – Demodex aries, D. brevis, D. canis, D. caprae, D. ermineae, D. folliculorum, D. mystacina, D. novazelandica, D. ovis, D. phylloides
Tokelau (Andrews, 1988) – Demodex
Mammalia
Metatheria
Dasiuromorphia
Fam. Dasiuridae
Antechinus agilis – Demodex antechini, D. tortellinioides
A. stuarti – Demodex antechini
Didelphimorphia
Fam. Didelphidae
Didelphis marsupialis – Demodex marsupiali
Diprotodontia
Fam. Phascolarctidae
Phascolarctos cinereus – Demodex sp.
Fam. Burramyidae
Cercartetus nanus – Stomatodex cercarteti
Eutheria
Soricomorpha
Fam. Soricidae
Crocidura suaveolens – Demodex foveolator
Neomys anomalus – Apodemodex cornutus, Demodex neomydis, Soricidex sp.
Sorex araneus – Demodex soricinus, Soricidex dimorphus
Fam. Talpidae
Talpa europaea – Demodex talpae
Chiroptera
Fam. Pteropodidae
Eonycteris spelaea – Demodex neoopistosomae, D. spelaea
Macroglossus minimus – Demodex bicaudatus, D. macroglossi
Rousettus aegyptiacus leachi – Stomatodex rousetti
Fam. Hipposideridae
Rhinonicteris aurantia – Ophthalmodex australiensis
Fam. Phyllostomatidae
Artibeus aztecus – Demodex artibei lituratus – Ophthalmodex artibei
Carollia perspicillata – Demodex carolliae, D. longissimus, Ophthalmodex carolliae, Pterodex carolliae
Desmodus rotundus – Demodex desmodi
Phyllostoma hastatum – Demodex phyllostomati
Fam. Molossidae
Molossus molossus – Demodex molossi, Ophthalmodex molossi
Fam. Mystacinidae
Demodex mystacina, D. novazelandica
Fam. Vespertilionidae
Barbastella barbastellus – Stomatodes corneti corneti
Eptesicus melanopterus – Demodex melanopteri
Myotis daubentoni – Demodex aelleni
M. lucifugus – Ophthalmodex juniatae
M. myotis – Stomatodes corneti myotis
M. dasycneme – Stomatodes corneti myotis
Nycticeius humeralis – Demodex nycticeii
Plecotus auritus – Demodex chiropteralis, D. plecoti, D. soricinus
P. [Corynorhinus] mexicanus – Demodex mexicanus
Vespadelus pumilus – Ophthalmodex wilsoni
Rodentia
Fam. Arvicolidae
Arvicola terrestris – Demodex gliricolens
Microtus agrestis – Demodex agrestis
M. arvalis – Demodex arvicolae
Myodes [Clethrionomys] gapperi – Demodex gapperi
M. [C.] glareolus – Demodex buccalis, D. glareoli
M. [Clethrionomys, Evotomys] glareolus britannicus – Demodex glareoli
Fam. Caviidae
Cavia – Demodex caviae
Fam. Cricetidae
Cricetulus migratorius – Demodex cricetuli
C. barabensis – Demodex sinocricetuli
Mesocricetus auratus – Demodex aurati, D. criceti
Peromyscus leucopus – Demodex peromysci
Phodopus sungarus – Demodex phodopi, D. sungori
Fam. Gliridae (Myoxidae)
Muscardinus avellanarius – Demodex muscardini
Fam. Muridae
Apodemus agrarius – Demodex agrarii, D. apodemi, D. gracilentus, D. huttereri
Bandicota indica – Demodex bandicotae
Sylvaemus flavicollis – Demodex corniculatus, D. mollis, D. rosus
S. sylvaticus – Demodex apodemi, D. auricularis, D. lacrimalis, D. longior, Ophthalmodex apodemi
Leopoldamys sabanus – Demodex sabani
L. edwardsi – Demodex sabani
Mus musculus – Demodex conicus, D. flagellurus, D. fusiformis, D. marculus, D. musculi, D. vibrissae, Glossicodex musculi
Niviventer cremoriventer – Demodex sabani
N. rapit – Demodex sabani
Rattus rattus – Demodex nanus
R. norvegicus – Demodex nanus, D. norvegicus, D. ponderosus, D. ratti, D. ratticola
R. annandalei – Demodex sabani
R. jalorensis – Demodex sabani
Sundamys mulleri – Demodex sabani
Fam. Sciuridae
Cynomys ludovicianus – Demodex
Sciurus vulgaris – Demodex sciurinus
Fam. Castoridae
Castor fiber – Demodex castoris
Lagomorpha
Fam. Leporidae
Oryctolagus cuniculus – Demodex oryctolagi
Carnivora
Fam. Ailuropodidae
Ailuropoda melanoleuca – Demodex ailuropoda
Fam. Canidae
Canis lupus familiaris – Demodex canis, D. cornei, D. cyonis, D. injai
Fam. Felidae
Felis catus – Demodex gatoi, D. cati
Panthera tigris amoyensis – Demodex tigris
Uncia uncia – Demodex uncii
Fam. Mustelidae
Lutra lutra – Demodex lutrae
Meles meles – Demodex melesinus
Mustela nivalis vulgaris – Demodex sp.
M. erminea – Demodex erminea
Fam. Procyonidae
Procyon lotor – Demodex sp.
Fam. Ursidae
Ursus americanus – Demodex ursi
Carnivora Pinnipedia
Fam. Otariidae
Zalophus californianus – Demodex zalophi
Fam. Phocidae
Phoca vitulina – Demodex phocidi
Primates
Fam. Galagonidae
Galago moholi – Rhinodex baeri, Stomatodex galagoensis
Fam. Cebidae
Saimiri sciureus – Demodex saimiri
Fam. Cercopithecidae
Macaca mulatta – Demodex macaci
Fam. Callithricidae
Saguinus geoffroyi – Demodex sp.
S. midas – Demodex sp.
Fam. Atelidae
Ateles sp. – Demodex araneae
Fam. Hominidae
Homo sapiens – Demodex brevis, D. folliculorum, D. f. sinensis
Perissodactyla
Fam. Equidae
Equus caballus – Demodex caballi, D. equi
Artiodactyla
Fam. Bovidae
Bison bonasus – Demodex bisonianus, D. bovis
Bos javanicus – Demodex bantengi
B. taurus – Demodex bovis, D. ghanensis, D. tauri
Capra hircus – Demodex caprae
Odocoileus hemionus – Demodex kutzeri, D. odocoilei
O. virginianus – Demodex kutzeri, D. odocoilei
Ovis aries – Demodex aries, D. ovis
Syncerus caffer – Demodex cafferi
Taurotragus oryx – Demodex sp. (nr. D. kutzeri)
Fam. Cervidae
Alces alces – Demodex kutzeri
Capreolus capreolus – Demodex kutzeri
Cervus elaphus – Demodex acutipes, D. kutzeri
C. nelsoni – Demodex kutzeri
C. nippon pseudaxis – Demodex kutzeri, D. pseudaxisi
Dama dama – Demodex kutzeri
Odocoileus h. hemionus – Demodex kutzeri
O. virginianus – Demodex kutzeri, D. odocoilei
Fam. Suidae
Sus scrofa – Demodex folliculorum, D. phylloides
Harpirhynchidae Dubinin, 1957: 94
Two of the three subfamilies of Harpirhynchidae include skin parasites of birds.
Harpypalpinae are living on Passeriform birds, Harpirhynchinae – also on Passeriformes, but also on birds of other orders – Anseriformes, Apodiformes, Charadriiformes, Ciconiiformes, Coliiformes, Columbiformes, Galliformes, Pelecaniformes. Harpirhynchids were shown to be highly host specific, with 69.5% being monoxenous species and 23% species are associated with several closely related hosts (Bochkov, Klimov & Skoracki, 2017).
In his description of fam. Harpirhynchidae Dubinin (1957) included the specialized subfamily Ophioptinae (parasites of snakes) as subfamily of this family. Many subsequent researchers (Fain, Beron, Lizaso, etc.) followed Southcott (1956) who described the family Ophioptidae, but Bochkov (2002), based on cladistic analysis, restored the subfamilial status of the snake parasites.
According to Bochkov (2009b), the family contains 84 sp. of 14 genera. In line with Bochkov & Literák (2006), the family Harpirhynchidae is considered here to contain three subfamilies with 16 genera and ca. 135 species [93 acc. to Zhang et al., 2011], as follows:
Fam. Harpyrhynchidae Dubinin, 1957 – 16 genera, 135 species
Subfam. Harpypalpinae Fain, 1972 – 2 genera, 19 species
Harpypalpoides Lombert et Moss, 1983 – 9 species
Harpypalpus Dubinin, 1957 – 10 species
Subfam. Harpirhynchinae Dubinin, 1957 – 12 genera, 98 species
Tribus Harpirhynchini Fain, 1972
Anharpyrhynchus Fain, 1995 – 4 species
Crassacarus Bochkov, OConnor et Klompen, 2015 – 6 species
Fainharpirhynchus Bochkov et Galloway, 2013 – 3 species
Harpyrhynchiella Fain, 1972 – 3 species
Harpyrhynchiella Fain, 1972 – 1 species
Cypsharpirhynchus Fain, 1995 – 2 species
Harpyrhynchoides Fain, 1972 – 58 species
Harpirhynchus Mégnin, 1878 – 5 species
Harpirhynchus s. str. Mégnin, 1878 – 4 species
Pseudoharpirhynchus Fain, Bochkov et Mironov, 1999 – 1 species
Neharpyrhynchus Fain, 1995 – 9 species
Ralliharpirhynchus Fain, 1995 – 2 species
Trichorhynchiella Fain, 1995 – 1 species
Tribus Metharpyrhynchini Fain, 1972
Metharpyrhynchus Fain, 1972 – 4 species
Tribus Perharpyrhynchini Fain, 1972
Perharpyrhynchus Fain, 1972 – 2 species
Harpirhynchinae incertae sedes – 1 species
Subfam. Ophioptinae Southcott, 1956 – 2 genera, 18 species
Afrophioptes Fain, 1962 – 2 species
Ophioptes Sambon, 1928 – 16 species
The history of the research on Harpirhynchidae starts with the description of “Sarcoptes” nidulans Nitsch, 1818. The same specied is being described under various names all along 19th century by C.L. Koch (1836), Giebel (1871), Murray (1877), Canestrini (1886), Heim (1892), Railliet (1893), Berlese (1895) and others. In the paper of Berlese & Trouessart (1889) are described three more species.
In his general paper Mégnin (1877) first uses the name Harpirhynchus and under it these mites find place in the family Cheyletidae and remain there for the long 80 years, until the new classification of Dubinin (1957). He motivates the placement of genus Harpirhynchus, together with his new genus Harpypalpus, in a family of it’s own, including also the snake parasites of the new subfamily Ophioptinae. Shortly before Dubinin’s revision of the family Fritsch (1954) published his revision of these mites, leaving them (plus eight newly described species) in the genus Harpirhynchus. New hiatus follows and in 1972 and 1995 A. Fain described nine more genera and the subfamily Harpypalpidae, thus laying the foundation of the modern system of Harpirhynchidae. Many species known to belong to this family (without Ophioptinae, treated here separately) are described by Fain or with his participation.
Since 1999 begin a series of contributions of A. Bochkov (partly together with A. Fain, S.V. Mironov, H. Klompen, I. Literák, B. Sikora, M. Skoracki, T.D. Galloway and other co-authors). Based on solid analysis, using cladistic and other modern methods, they greatly completed the knowledge on Harpirhynchidae. Other specialists having contributed to the studies on the family are R. Domrow, M. Jeleva, N. Banks, H.E. Ewing, W. W. Moss, A.C. Oudemans, J.H. Oliver, R.F. Lawrence, H.A.P.M. Lombert, P.-Y. Henry, V. Černy, W. Sixl, E.M. Boyd, F. Dusbábek, J. Martinu, and others.
The recent review of Bochkov & Literák (2006) summarizes the knowledge on the European Harpirhynchidae (28 species, incl. the four described by Martinu, Dusbábek & Literák, 2008). According to Moss (1979a), the number of species belonging to this family is theoretically more than 2500. The Ophioptinae was downgraded from family to subfamily status, but is treated here on it’s own, as it comprises specific parasites of snakes.
(Harpirhynchinae and Harpypalpinae)
Harpirhynchidae Dubinin, 1957: 94
Type genus: Harpirhynchus Mégnin, 1877
Harpypalpinae Fain, 1972: 49
Type genus: Harpypalpus Dubinin, 1957
Harpypalpoides Lombert & Moss, 1983: 165
Type species: Harpypalpoides lukoschusi Lombert et Moss, 1983
Harpypalpus hirundinis Fain, 1972: 58
Harpypalpoides hirundinis (Fain): Fain, Bochkov & Mironov, 1999: 53; Bochkov & Klompen, 2014а: 469
Type locality: Kinshasa
Distribution: DR Congo
Host: Psalidoprocne sp. (Passeriformes: Hirundinidae)
Harpypalpoides hirundinoides Bochkov & Klompen, 2014a: 468
Type locality: Noord Brabant, Schaijk
Distribution: Holland
Host: Hirundo rustica (Passeriformes: Hirundinidae)
Harpypalpoides johnstoni Bochkov & Klompen, 2014а: 470
Type locality: Holland, Gelderland, Arnhem
Distribution: Holland, Kazakhstan
Hosts: Fringilla coelebs, F. montifringilla (Passeriformes: Fringillidae)
Harpypalpoides lesickii Skoracki, Bochkov & Sikora, 2004: 84; Bochkov & Literák, 2006: 140; Bochkov & Klompen, 2014а: 469
Type locality: Poznań environs, Jeziory village
Distribution: Kazakhstan, Poland
Hosts: Carduelis cannabina, Coccothraustes coccothraustes (Passeriformes: Fringillidae)
Harpypalpoides lukoschusi Lombert & Moss, 1983: 165; Skoracki, Bochkov & Sikora, 2004: 85; Bochkov & Literák, 2006: 140; Bochkov & Klompen, 2014а: 467
Type locality: Utrecht (Holland)
Distribution: Holland, Poland
Hosts: Turdus merula, T. pilaris (Passeriformes: Turdidae)
Harpypalpoides namibiensis Fain, Bochkov & Mironov, 1999: 53; Bochkov & Klompen, 2014а: 469
Type locality: Karas, Aroab
Distribution: Namibia
Host: Emberiza impetuani (Passeriformes: Emberizidae)
Harpypalpoides regulus Bochkov & Klompen, 2014а: 471
Type locality: Noord Brabant, Schaijk
Distribution: Holland
Host: Regulus regulus (Passeriformes: Regulidae)
Harpypalpoides sitta Bochkov & Klompen, 2014a: 474
Type locality: California, Monterey Co.
Distribution: USA (Arizona, California)
Host: Sitta pygmaea (Passeriformes: Sittidae)
Harpypalpoides sylvia Bochkov & Klompen, 2014: 472; Bochkov & Klompen, 2014а: 472
Type locality: Amsterdam
Distribution: Holland
Host: Sylvia atricapilla (Passeriformes: Sylviidae)
Harpypalpus Dubinin, 1957: 100
Type species: Harpirhynchus longipes Fritsch, 1954 (? syn. of Harpirhynchus holopus Berlese et Trouessart, 1889)
Harpypalpus dubinini Fain, 1972: 58; Bochkov & Klompen, 2014а: 465
Type locality: Antwerp Zoo
Distribution: ? Central Africa
Host: Ploceus [Sitagra] intermedius (Passeriformes: Ploceidae)
Harpirhynchus holopus Berlese & Trouessart, 1889: 18
Sarcoborus holopus: Oudemans, 1906: 76
Harpypalpus holopus (Berlese et Trouessart): Fain, Bochkov & Mironov, 1999: 46; Bochkov & Literák, 2006: 140; Bochkov & Klompen, 2014a: 462
Type locality: Italy
Distribution: Belgium, Italy
Hosts: Passer domesticus (Passeriformes: Passeridae), ? Luscinia svecica (Passeriformes: Muscicapidae)(? Emberiza citronella – Emberizidae – see Bochkov & Klompen, 2014a)
Harpypalpus lonchura Bochkov & Klompen, 2014а: 460
Type locality: Western Australia, Mount Hart
Distribution: Australia
Host: Lonchura castaneothorax (Passeriformes: Estrildidae)
Harpyrhynchus longipes Fritsch, 1954: 183
Harpypalpus longipes (Fritsch): Moss, 1979; Skoracki, Bochkov & Sikora, 2004: 86; Bochkov & Klompen, 2014a: 458
Type locality: Germany, Bavaria, Middle Franconia, Kreis Freie Stadt Erlangen
Distribution: Germany, Holland, Poland
Hosts: Cyanistes caeruleus, P. major, Periparus ater (Passeriformes: Paridae), Troglodytes troglodytes (Pass.: Troglodytidae), Nucifraga caryocatactes (Pass.: Corvidae), Carduelis cannabina, Spinus spinus, Pyrrhula pyrrhula, Serinus mozambicus (Pass.: Fringillidae)
Harpypalpus pyrrhula Bochkov & Klompen, 2014a: 462
Type locality: Gelderland, Nijmegen
Distribution: Holland
Host: Pyrrhula pyrrhula (Passeriformes: Fringillidae)
Harpypalpus serini Fain, 1972: 58; Bochkov & Klompen, 2014a: 465
Type locality: Antwerp Zoo
Distribution: Central Africa
Host: Serinus mozambicus (Passeriformes: Fringillidae)
Harpypalpus spermestes Fain, Bochkov & Mironov, 1999: 50
Type locality: Luluabourg (Kananga, Kasai Occidental)
Distribution: DR Congo
Host: Ploceus cucullatus (Passeriformes: Ploceidae)
Harpypalpus sturnus Bochkov & Klompen, 2014a: 463
Type locality: Gelderland, Nijmegen
Distribution: Holland
Host: Sturnus vulgaris (Passeriformes: Sturnidae)
Harpypalpus taeniopygia Bochkov & Klompen, 2014a: 461
Type locality: Western Australia, Brooking Spring
Distribution: Australia
Host: Taeniopygia guttata (Passeriformes: Estrildidae)
Harpypalpus tiarae Fain, Bochkov & Mironov, 1999: 49; Bochkov & Klompen, 2014a: 465
Type locality: Antwerp Zoo
Distribution: Central Africa
Host: Tiaris canora (Passeriformes: Thraupidae)
Harpirhynchinae Dubinin, 1957: 100
Type genus: Harpirhynchus Mégnin, 1877
Harpyrhynchini Fain, 1972: 50
Type genus: Harpirhynchus Mégnin, 1877
Harpyrhynchus (Anharpyrhynchus) Fain, 1972: 50
Harpirhynchus (Anharpyrhynchus) Fain, 1994: 111
Anharpyrhynchus Fain, 1995: 75
Type species: Harpyrhynchus monstrosus Fritsch, 1954
Anharpyrhynchus apodus Bochkov & Klompen, 2014: 320
Type locality: New South Wales, Cordeaux Dam, 34.37°S 150.75°E
Distribution: Australia
Hosts: Lichmera indistincta, Phylidonyris novaehollandiae (Passeriformes: Meliphagidae)
Anharpyrhynchus elizae Bochkov & Klompen, 2014b: 317; Bochkov, OConnor & Klompen, 2015: 112
Type locality: Florida, Columbia Co., O’Leno State Park, 29.92°N 82.58°W
Distribution: USA
Hosts: Cyanocitta cristata, C. stelleri (Passeriformes: Corvidae), Colaptes auratus (Piciformes: Picidae)
Anharpyrhynchus lukoschusi Bochkov & Klompen, 2014b: 320
Type locality: Western Australia, Mount Hart, 17.02 °S 125.07 °E
Distribution: Australia
Hosts: Manorina flavigula, Certhionyx pectoralis (Passeriformes: Meliphagidae)
Harpyrhynchus monstrosus Fritsch, 1954: 188; Dubinin, 1957: 107; Lawrence, 1959a: 425
Harpyrhynchus (Anharpyrhynchus) monstrosus, Fain, 1972: 55
Harpyrhynchus (Anharpyrhynchus) fritschi Fain, 1972: 55 – type from Xanthoura luxuosa, Anvers Zoo
Harpirhynchus monstrosus Fritsch: Domrow, 1991: 1317
Anharpyrhynchus monstrosus (Fritsch): Fain, 1995: 97; Bochkov & Literák, 2006: 140
Type locality: Germany
Distribution: Australia, Belgium, Germany
Hosts: Corvus monedula, Garrulus glandarius, Xanthoura yncas luxuosa (= Cyanocorax yncas)(Passeriformes: Corvidae); recorded from Australia on: Gliciphila indistincta, Meliornis novaehollandiae, Myzantha flavigula, Myzomela pectoralis (Passeriformes: Meliphagidae)
Crassacarus Bochkov, OConnor & Klompen, 2015: 67
Type species: Crassacarus alexfaini Bochkov, OConnor et Klompen, 2015
Crassacarus alexfaini Bochkov, OConnor & Klompen, 2015: 67
Type locality: Michigan, Livingston Co., 4 mi S. Howell, 42.5495394°N, 83.9409112°W
Distribution: USA
Hosts: Cardinalis cardinalis (Passeriformes: Cardinalidae), Passerina ciris (Pass.: Cardinalidae), Setophaga petechia (Pass.: Parulidae)
Harpyrhynchus cylindripalpus: Fritsch, 1954: 183; Dubinin, 1957: 108
Harpirhynchus (Pseudoharpirhynchus) cylindripalpus Fritsch: Fain, Bochkov & Mironov, 1999: 39
Harpyrhynchoides cylindripalpus (Fritsch): Bochkov & Literák, 2006: 140
Crassacarus cylindripalpus (Fritsch) – Bochkov, OConnor & Klompen, 2015: 67
Type locality: Germany
Distribution: Germany
Hosts: Fringilla coelebs (Passeriformes: Fringillidae), Passerina ciris (Pass., Cardinalidae)
Crassacarus fritschi Bochkov, OConnor & Klompen, 2015: 71
Type locality: Michigan, Muskegon Co., Muskegon State game Area, along US route 31,
43.2961989°N, 86.0668945°W
Distribution: USA
Host: Bombycilla cedrorum (Passeriformes: Bombycillidae)
Crassacarus melanerpes Bochkov, OConnor & Klompen, 2015: 75
Type locality: Kansas, Johnson Co., Craig, 38.97°N 94.82°W
Distribution: USA
Host: Melanerpes formicivorus (Piciformes: Picidae)
Crassacarus sialia Bochkov, OConnor & Klompen, 2015: 73
Type locality: Wyoming, Campbell Co., Gillette, 19.3 km E of, 44.29°N 105.5°W
Distribution: USA
Host: Sialia currucoides (Passeriformes: Turdidae)
Crassacarus tinae Bochkov, OConnor & Klompen, 2015: 70
Type locality: Wyoming, Campbell Co., Gillette, 19.3 km E of, 44.29°N 105.5°W
Distribution: USA
Host: Carduelis tristis (Passeriformes: Fringillidae)
Fainharpirhynchus Bochkov & Galloway, 2013: 414
Type species: Fainharpirhynchus contopus Bochkov et Galloway, 2013
Fainharpirhynchus contopus Bochkov & Galloway, 2013: 418; Bochkov, OConnor & Klompen, 2015: 83
Type locality: Manitoba, Winnipeg
Distribution: Canada, USA (Michigan)
Hosts: Contopus cooperi, C. virens (Parasitiformes: Tyrannidae)
Fainharpirhynchus legatus Bochkov & Klompen, 2014: 307
Type locality: Sangre Grande, Cumuto, 1.6 km E on Cumuto Road, 10.58°N, 61.2°W
Distribution: Trinidad & Tobago
Host: Legatus leucophaius (Parasitiformes: Tyrannidae)
Fainharpirhynchus mossi Bochkov & Klompen, 2014: 311
Type locality: North America
Distribution: North America
Host: Myiarchus crinitus (Parasitiformes: Tyrannidae)
Harpyrhynchiella Fain, 1972: 56
Type species: Harpyrhynchus reductus Fritsch, 1954
Cypsharpirhynchus Fain, 1995: 75. Syn. by Bochkov (2015)
Type species: Metharpyrhynchus cypsiuri Fain, 1972
Cypsharpirhynchus Fain, 1995: 75. Syn. by Bochkov (2015)
Type species: Metharpyrhynchus cypsiuri Fain, 1972
Harpyrhynchiella (Cypsharpirhynchus) apus: Bochkov, 2015: 8
Type locality: Kaliningrad Province, Curonian spit
Distribution: Russia
Host: Apus apus (Apodiformes: Apodidae)
Metharpyrhynchus cypsiuri Fain, 1972: 57
Cypsharpirhynchus cypsiuri (Fain): Fain, 1995: 75
Harpyrhynchiella (Cypsharpirhynchus) cypsiuri (Fain): Bochkov, 2015: 7
Type locality: Kananga (= Luluabourg), Prov. Kasai, DR Congo; Togo
Distribution: DR Congo, Togo
Hosts: Apus affinis, Cypsiurus parvus (Apodiformes: Apodidae)
Harpyrhynchiella Fain, 1972: 56
Harpyrhynchiella (Harpyrhynchiella): Bochkov, 2015: 4
Type species: Harpyrhynchus reductus Fritsch, 1954
Harpyrhynchus reductus Fritsch, 1954: 196; Dubinin, 1957: 108
Harpyrhynchiella reductus (Fritsch): Fain, 1972: 56; 1995: 94; Bochkov & Literák, 2006: 140
Harpyrhynchiella (Harpyrhynchiella) reductus (Fritsch): Bochkov, 2015: 4
Type locality: Germany
Distribution: Germany, Holland
Host: Apus apus (Apodiformes: Apodidae)
Harpyrhynchus (Harpyrhynchoides) Fain, 1972: 50
Harpyrhynchoides (genus): Fain, Bochkov & Mironov, 1999: 42
Type species: Harpyrhynchus (Harpyrhynchoides) squamosus Fain, 1972
Pseudoharpirhynchus Fain, Bochkov et Mironov, 1999 is syn. by Bochkov, OConnor & Klompen, 2015. Type: Harpirhynchus agapornis Fain, 1972
Harpyrhynchoides accipiter Bochkov & OConnor, 2013: 503; Bochkov, OConnor & Klompen, 2015: 35
Type locality: Michigan, Chippewa Co., Whitefish Point
Distribution: USA (Michigan, New York)
Host: Accipiter striatus (Accipitriformes: Accipitridae)
Harpyrhynchoides actitis Bochkov, OConnor & Klompen, 2015: 23
Type locality: British Columbia, Vancouver, 49.16°N, 122.86°W
Distribution: Canada
Host: Actitis macularius (Charadriiformes: Scolopacidae)
Harpyrhynchoides aegolius Bochkov & OConnor, 2013: 501
Type locality: Michigan, Oakland Co., Blissfield
Distribution: USA (Michigan)
Host: Aegolius acadicus (Strigiformes: Strigidae)
Harpyrhynchus (Harpyrhynchus) agapornis Fain, 1972: 50; 1995: 77
Harpirhynchus (Pseudoharpirhynchus) agapornis Fain: Fain, Bochkov & Mironov, 1999: 39
Harpyrhynchoides agapornis (Fain): Bochkov, OConnor & Klompen, 2015: 66
Type locality: South Africa
Distribution: South Africa
Hosts: Agapornis nigrigenis, A. pullarius, A. roseicollis (Psittaciformes: Psittaculidae)
Harpyrhynchus nidulans: Mégnin, 1877 : 429; Oudemans, 1939 (? misidentification)
Harpyrhynchoides alaudinus Bochkov, 2000b: 91; Bochkov & Literák, 2006: 140
Type locality: Okskiy Reservation, Moskow Prov.
Distribution: France (?), Russia
Host: Alauda arvensis (Passeriformes: Alaudidae)
Harpyrhynchus (Harpyrhynchoides) alectoris Fain, 1972: 54
Harpirhynchus (Harpyrhynchoides) alectoris, Fain, 1994: 117
Harpyrhynchoides alectoris (Fain): Fain, Bochkov & Mironov, 1999: 45; Bochkov & Literák, 2006: 140
Type locality: Antwerp Zoo
Distribution: Spain
Hosts: Alectoris graeca, A. rufa (Galliformes: Phasianidae)
Harpyrhynchus (Harpyrhynchoides) amazonae Fain, 1972: 52; 1994: 113
Harpyrhynchoides amazonae (Fain): Bochkov, OConnor & Klompen, 2015: 59
Type locality: Antwerp Zoo
Distribution: South America
Host: Amazona aestiva (Psittaciformes: Psittacidae)
Harpyrhynchus (Harpyrhynchoides) anatum Fain, 1972: 54; 1994: 113; Bochkov & Literák, 2006: 140
Type locality: Antwerp Zoo
Distribution: Europe
Host: Anas querquedula (Anseriformes: Anatidae)
Harpyrhynchoides artamus Bochkov & Klompen, 2015: 24
Type locality: SE Asia
Distribution: SE Asia
Host: Artamus fuscus (Passeriformes: Artamidae)
Harpyrhynchus (Harpyrhynchoides) asio Fain, 1972: 52; 1994: 128
Harpyrhynchoides asio (Fain): Bochkov & Literák, 2006: 140
Type locality: Baelen Neet
Distribution: Belgium
Host: Asio otus otus (Strigiformes: Strigidae)
Harpyrhynchoides athene Bochkov, OConnor & Klompen, 2015: 56
Type locality: Texas, Lubbock Co., Lubbock, 33.58°N, 101.86°W
Distribution: USA
Host: Athene canicularia (Strigiformes: Strigidae)
Harpyrhynchoides botaurus Bochkov & Galloway, 2013: 409; Bochkov, OConnor & Klompen, 2015: 33
Type locality: Manitoba, Winnipeg
Distribution: Canada
Host: Botaurus lentiginosus (Pelecaniformes: Ardeidae)
Harpirhynchus brevis Ewing, 1911: 38; Moss et al., 1968: 391; Fain, 1995: 99; Bochkov & Literák, 2006: 140
Harpyrhynchoides brevis (Ewing): Bochkov & Klompen, 2014c
Type locality: ?
Distribution: ?
Hosts: Turdus migratorius (Passeriformes: Turdidae), Coccothraustes vespertina, Loxia curvirostra (Pass.: Fringillidae)
Harpyrhynchoides bubulcus Bochkov, OConnor & Klompen, 2015: 28
Type locality: Florida, Orange Co., Orlando, Lake Adaire, 28.5666208°N, 81.2608791°W
Distribution: USA
Host: Bubulcus ibis (Pelecaniformes: Ardeidae)
Harpyrhynchus butorides Boyd, 1968: 19
Harpirhynchus (Harpyrhynchoides) ?herodius, Fain 1994: 119
Harpyrhynchoides butorides (Boyd): Bochkov, OConnor & Klompen, 2015: 31
Type locality: Massachusetts, Hampshire, 42.38°N, 72.52°W
Distribution: USA
Host: Butorides virescens (Pelecaniformes: Ardeidae)
Harpyrhynchoides calidris Bochkov, OConnor & Klompen, 2015: 17
Type locality: Kansas
Distribution: USA
Host: Calidris minutilla (Charadriiformes: Scolopacidae)
Harpyrhynchus capellae Fritsch, 1954: 179; Dubinin, 1957: 104
Harpirhynchus (Harpyrhynchoides) capellae, Fain, 1994: 117
Harpyrhynchoides capellae (Fritsch): Fain, Bochkov & Mironov, 1999: 45; Bochkov & Literák, 2006: 140
Type locality: Erlangen (Germany)
Distribution: France, Germany
Hosts: Gallinago gallinago, Lymnocryptes minimus (Charadriiformes: Scolopacidae)
Harpirhynchus (Harpyrhynchoides) capitatus Fain, 1976: 129; 1994: 119
Type locality: Antwerp Zoo
Distribution: ?
Host: Columbigallina talpacoti (Columbiformes: Columbidae)
Harpyrhynchoides charadrius Bochkov, OConnor & Klompen, 2015: 20
Type locality: Texas, Lubbock Co., 33.6°N, 101.83°W
Distribution: USA
Host: Charadrius vociferus (Charadriiformes: Charadriidae)
Harpyrhynchoides clamator Bochkov, 2014: 480
Type locality: Limpopo, Waterberg District Municipality, Bela-Bela, Little Kariba
Distribution: South Africa
Host: Clamator jacobinus (Cuculiformes: Cuculidae)
Harpyrhynchoides coccyzus Bochkov, OConnor & Klompen, 2015: 44
Type locality: Michigan, Washtenaw Co., Ann Arbor, UM Campus, 42.2793409°N, 83.7386084°W
Distribution: USA
Hosts: Coccyzus americanus, C. erythrophthalmus (Cuculiformes: Cuculidae)
Harpyrhynchus (Harpyrhynchoides) columbae Fain, 1972: 51; 1994
Harpyrhynchoides columbae (Fain): Bochkov & Literák, 2006: 140
Type locality: Bouillon
Distribution: Belgium
Host: Columba livia domestica (Columbiformes: Columbidae)
Harpyrhynchus (Harpyrhynchoides) coturnix Fain, 1972: 54
Harpirhynchus (Harpyrhynchoides) coturnix, Fain, 1994: 116
Harpyrhynchoides coturnix (Fain): Bochkov, OConnor & Klompen, 2015: 34
Type locality: Antwerp Zoo
Distribution: North America (unknown locality)
Host: Coturnix delegorguei (Galliformes: Phasianidae)
Harpyrhynchus (Harpyrhynchoides) coxatus Fain, 1972: 54; 1994: 133
Type locality: Antwerp Zoo
Distribution: ?
Hosts: Columbigallina talpacoti, Columbina cruziana, Scardafella squamata (Columbiformes: Columbidae)
Harpirhynchus crista – galli Berlese & Trouessart, 1889: 121; Lawrence, 1959b: 108
Sarcoborus crista – galli: Oudemans, 1906: 77
Harpyrhynchus crista – galli: Lawrence, 1959a: 418
Harpirhynchus (Harpyrhynchoides) cristagalli, Fain, 1994: 130
Harpyrhynchoides cristagalli: Bochkov, OConnor & Klompen, 2015: 66
Type locality: South Africa
Distribution: Namibia, South Africa
Hosts: Colius colius, C. striatus (Coliiformes: Coliidae)
Harpyrhynchoides crotophaga Bochkov, OConnor & Klompen, 2015: 61
Type locality: Paramaribo, Wegnaar Zee, 05.82°N 55.17°W
Distribution: Suriname
Host: Crotophaga ani (Cuculiformes: Cuculidae)
Harpyrhynchoides gallowayi Bochkov, OConnor & Klompen, 2015: 51
Type locality: Manitoba, Winnipeg
Distribution: Canada
Host: Columba livia (Columbiformes: Columbidae)
Harpyrhynchoides heatherae Bochkov & Galloway, 2013: 407
Type locality: Manitoba, Winnipeg, Sommerland
Distribution: Canada
Host: Junco hyemalis (Passeriformes: Emberizidae)
Harpyrhynchus herodius Boyd, 1968: 18
Harpyrhynchus (Harpyrhynchoides) herodius Fain, 1994: 112
Harpyrhynchoides herodius (Boyd): Bochkov & Literák, 2006: 140; Bochkov, OConnor & Klompen, 2015: 28
Type locality: Massachusetts, USA
Distribution: Belgium, Holland, USA (Massachusetts)
Hosts: Ardea cinerea, A. herodius (Pelecaniformes: Ardeidae)
Harpyrhynchoides ixobrychus Bochkov, OConnor & Klompen, 2015: 32
Type locality: Michigan, Oakland Co.
Distribution: USA
Host: Ixobrychus exilis (Pelecaniformes: Ardeidae)
Harpyrhynchus (Harpyrhynchoides) kakatoe Fain, 1972: 52; 1994: 120
Harpirhynchus kakatoe: Domrow, 1991: 1317
Harpyrhynchoides kakatoe (Fain): Bochkov, OConnor & Klompen, 2015: 64
Type locality: Antwerp Zoo
Distribution: Australia
Host: Kakatoe galerita (Psittaciformes: Cacatuidae)
Harpyrhynchus (Harpyrhynchus) lawrencei Fain, 1972: 51
Harpirhynchus (Harpyrhynchoides) lawrencei, Fain, 1994: 113
Harpyrhynchoides lawrencei (Fain): Bochkov, OConnor & Klompen, 2015: 60
Type locality: Antwerp Zoo
Distribution: South America
Host: Aratinga nenday (Psittaciformes: Psittacidae)
Harpyrhynchoides megascops Bochkov, OConnor & Klompen, 2015: 39
Type locality: Michigan, Michigan, Oakland Co., Commerce Twp., Marble Ct., 42.5644286°N, 83.4637356°W
Distribution: USA
Host: Megascops asio (Strigiformes: Strigidae)
Harpyrhynchus (Harpyrhynchoides) metropeliae Fain, 1972: 53
Harpyrhynchoides metropeliae (Fain): Bochkov, OConnor & Klompen, 2015: 47
Type locality: Antwerp Zoo
Distribution: South America (unknown locality)
Host: Metropelia caeciliae (Columbiformes: Columbidae)
Harpyrhynchus (Harpyrhynchoides) modestus Fain, 1976: 126; 1994: 134; Bochkov & Galloway, 2013: 413
Harpyrhynchoides modestus (Fain): Bochkov, OConnor & Klompen, 2015: 54
Type locality: Antwerp Zoo
Distribution: Canada
Host: Metropelia caeciliae (Columbiformes: Columbidae)
Harpyrhynchus numidae Lawrence, 1959b: 110
Harpirhynchus (Harpyrhynchoides) numidae, Fain, 1994: 117
Harpyrhynchoides numidae (Fain): Bochkov, OConnor & Klompen, 2015: 64
Type locality: Africa
Distribution: Africa
Host: Numida meleagris (Galliformes: Numididae)
Harpyrhynchus (Harpyrhynchoides) oenae Fain, 1972: 53
Harpyrhynchus (Harpyrhynchoides) lamorali Fain, 1972: 53 – on Oena capensis.
Harpyrhynchus (Harpyrhynchoides) oenae lamorali, Fain, 1994: 125. Syn. by Bochkov et al., 1915
Type locality: Antwerp Zoo
Distribution: ?
Host: Tympanistria tympanistria (Columbiformes: Columbidae)
Harpyrhynchoides parazumpti Fain, Bochkov & Mironov, 1999: 43; Bochkov & Literák, 2006: 140; Bochkov, OConnor & Klompen, 2015: 54
Type locality: Kishinev (Moldavia)
Distribution: Germany, Moldavia, USA
Hosts: Corvus brachyrhynchos, C. frugilegus, C. monedula (Passeriformes: Corvidae)
Harpyrhynchus pectinifer Lawrence, 1959a: 419
Harpirhynchus (Harpyrhynchoides) pectinifer Lawrence: Fain, 1994: 113
Harpyrhynchoides pectinifer (Lawrence): Bochkov, OConnor & Klompen, 2015: 60
Type locality: Newington, East Transvaal, South Africa
Distribution: Rwanda, South Africa, USA, Zimbabwe
Hosts: Campethera abingoni, C. cailliauti, Colaptes auratus (Piciformes: Picidae)
Harpyrhynchoides phalaropus Bochkov & Galloway, 2013: 410; Bochkov, OConnor & Klompen, 2015: 28
Type locality: Manitoba, Winnipeg
Distribution: Canada
Host: Phalaropus lobatus (Charadriiformes: Scolopacidae)
Harpyrhynchoides pluvialis Bochkov, OConnor & Klompen, 2015: 25
Type locality: Ohio, Wayne Co., Wooster, 40.83°N, 81.88°W
Distribution: USA
Host: Pluvialis dominica (Charadriiformes: Charadriidae)
Harpyrhynchus (Harpyrhynchoides) psittaci Fain, 1972: 55; 1994: 129
Harpyrhynchoides psittaci (Fain): Bochkov, OConnor & Klompen, 2015: 66
Type locality: Antwerp Zoo
Distribution: ?
Hosts: Poicephalus senegalus, Psittacus erithacus (Psittaciformes: Psittacidae)
Harpyrhynchus (Harpyrhynchoides) psittaculae Fain, 1972: 52; 1994: 126
Type locality: Antwerp Zoo
Distribution: ?
Hosts: Psittacula alexandri, P. cyanocephala (Psittaciformes: Psittaculidae)
Harpyrhynchoides puffinus Mertins in Bochkov, OConnor & Klompen, 2015: 35
Type locality: Florida, Atlantic Ocean adjacent to east cost of Martin Co.
Distribution: USA
Host: Puffinus gravis (Procellariiformes: Procellariidae)
Harpyrhynchus rosellacinus Lawrence, 1959: 238
Harpyrhynchus (Harpyrhynchoides) rosellacinus, Fain, 1994: 139
Harpyrhynchoides rosellacinus (Lawrence):
Type locality: Sydney
Distribution: Australia (incl. Tasmania)
Hosts: Glossopsitta concinna, Lathamus discolor, Platycercus eximius, Trichoglossus chlorolepidotus, T. moluccanus (Psittaciformes: Psittaculidae)
Harpyrhynchus rubeculinus Černy & Sixl, 1971: 388
Harpyrhynchus (Harpyrhynchoides) rubeculinus (Černy et Sixl): Fain, 1994: 138
Harpyrhynchoides rubeculinus (Černy et Sixl): Fain, Bochkov & Mironov, 1999: 45; Bochkov & Literák, 2006: 140; 2008: 227; Bochkov & OConnor, 2013: 505; Bochkov, OConnor & Klompen, 2015: 51
Type locality: Brunnsee (Austria)
Distribution: Austria, Brazil, Czech Republic, France, USA
Hosts: Erithacus rubecula (Passeriformes: Muscicapidae, Saxicolinae), Catharus ustulatus, Turdus amaurochalinus (Pass.: Turdidae), Pheucticus ludovicianus (Pass.: Cardinalidae)
Harpyrhynchoides setophaga Bochkov & Klompen, 2014c: 406; Bochkov, OConnor & Klompen, 2015: 50
Type locality: Winnipeg, Manitoba
Distribution: Canada
Host: Setophaga ruticilla (Passeriformes: Parulidae)
Harpyrhynchoides spizella Bochkov & Klompen, 2014a: 415
Type locality: California
Distribution: USA
Host: Spizella passerina (Passeriformes: Passerellidae)
Harpyrhynchus (Harpyrhynchoides) squamosus Fain, 1972: 51; 1976: 126
Harpyrhynchoides squamosus (Fain): Bochkov, OConnor & Klompen, 2015: 65
Type locality: Antwerp Zoo
Distribution: ?
Host: Psittacula cyanocephala (Psittaciformes: Psittaculidae)
Harpyrhynchus tracheatus Fritsch, 1954: 180; Dubinin, 1957: 103
Harpirhynchus (Harpyrhynchoides) tracheatus, Fain, 1994: 120
Harpyrhynchoides tracheatus (Fritsch); Bochkov & Literák, 2006: 140; Bochkov & Galloway, 2013: 412; Bochkov & OConnor, 2013: 505; Bochkov, OConnor & Klompen, 2015: 36
Type locality: Germany
Distribution: Canada, Germany, Luxemburg, USA
Hosts: Buteo buteo, B. jamaicensis, B. lineatus (Accipitriformes: Accipitridae)
Harpyrhynchus (Harpyrhynchoides) tyto Fain, 1972: 51; 1994: 112
Harpyrhynchoides tyto (Fain): Bochkov & Literák, 2006: 140
Type locality: Crupet, Namur
Distribution: Belgium
Host: Tyto alba (Strigiformes: Tytonidae)
Harpyrhynchus vercammeni Lawrence, 1959a: 423
Harpyrhynchus (Harpyrhynchoides) vercammeni Lawrence: Fain, 1994: 138
Harpyrhynchoides vercammeni (Lawrence): Bochkov, OConnor & Klompen, 2015: 63
Type locality: Luvungi, Ruzizi, Kivu
Distribution: DR Congo
Host: Centropus toulou grilli (Cuculiformes: Cuculidae)
Harpyrhynchoides vulgaris Bochkov & Galloway, 2004: 27; Bochkov & Klompen 2014b: 409; Bochkov, OConnor & Klompen, 2015: 50
Type locality: Manitoba
Distribution: Canada
Hosts: Haemorhous [Carpodacus] purpureus (Passeriformes: Fringillidae), Setophaga ruticilla (Passeriformes: Parulidae)
Harpyrhynchoides xanthocephalus Bochkov & Klompen, 2014a: 212; Bochkov, OConnor & Klompen, 2015: 51
Type locality: Nebraska: Cherry Co., Valentine
Distribution: USA
Host: Xanthocephalus xanthocephalus (Passeriformes: Icteridae)
Harpyrhynchoides zenaida Bochkov, OConnor & Klompen, 2015: 43
Type locality: Michigan, Washtenaw Co., Ann Arbor, UM Campus, 42.2793409°N,
83.7386084°W
Distribution: USA
Host: Zenaida macroura (Columbiformes: Columbidae)
Harpyrhynchus (Harpyrhynchoides) zumpti Fain, 1972: 54; 1994: 136; Bochkov & Galloway, 2001: 2017
Harpyrhynchus (Harpyrhynchoides) diuca Fain, 1972: 53. Type from Antwerp Zoo, on Diuca diuca (Passeriformes: Thraupidae)
Harpyrhynchoides zumpti (Fain): Bochkov & Galloway, 2001, 2004; Bochkov, OConnor & Klompen, 2015: 48
Harpyrhynchoides kirgizorum Fain, Bochkov & Mironov, 1999: 42. Type loc.: Bishkek, Kirghizia, on Emberiza calandra. Syn. By Bochkov & Klompen, 2014c: 410
Type locality: Bloemenhof, Mafeking
Distribution: Canada, Kirghizia, South Africa
Hosts: Eremopteryx leucotis (Passeriformes: Alaudidae), Diuca diuca (Passeriformes: Thraupidae), Quiscalus quiscula (Passeriformes: Icteridae), Emberiza calandra, Junco hyemalis (Passeriformes: Emberizidae)
Harpirhynchus Mégnin, 1878: 8 (written also Harpyrhinchus and Harpyrynchus)
Type species: Sarcoptes nidulans Nitzsch, 1818
Harpicephalus Canestrini, 1886: 170
Type species: Sarcoptes nidulans Nitzsch, 1818
Pseudoharpirhynchus Fain, Bochkov & Mironov, 1999: 39 (subgenus)
Type species: Harpirhynchus agapornis Fain, 1972
Harpyrhinchus nidulans Nitzsch, 1818: Henry et al., 2004: 34 (misidentification), Literák et al., 2005: 26 (misidentification)
Harpirhynchus dusbabeki Bochkov & Literák, 2006: 138
Type locality: Pusté Úl’any (Slovakia)
Distribution: Austria, Croatia, Czech Rep., France, Poland, Slovakia
Host: Panurus biarmicus (Passeriformes: Panuridae)
Harpirhynchus (Harpirhynchus) galeridae Fain, Bochkov & Mironov, 1999: 39; Bochkov & Literák, 2006: 140
Type locality: Azerbaijan
Distribution: Azerbaijan
Host: Galerida cristata (Passeriformes: Alaudidae)
Harpirhynchus longipilus Banks, 1905a: 20; Fain, 1995: 99
Sarcopterus longipilis: Banks, 1905b: 135
Type locality: USA
Distribution: USA
Host: Loxia sp. (Passeriformes: Fringillidae)
Sarcoptes nidulans Nitzsch, 1818, in Ersch-Gruber, Allg. Encyclop. Wissensch., 1: 250; synonymy in Oudemans, 1906: 75 – 76; 1936: 1132 – 1134
Sarcoptes palumbinus C.L. Koch, 1836: 5, 12; Lucas, 1840: 487; synonymy in Oudemans, 1936: 1134 – 1135
Sarcopterus nidulans: Giebel, 1871: 29; Murray, 1877: 314; Berlese, 1895: 5
Harpirhynchus nidulans Mégnin, 1877: 13; Literák et al., 2005: 26; Bochkov & Literák, 2006: 140
Harpicephalus nidulans: Canestrini, 1886: 170
? Harpirhynchus megnini Heim, 1892: 132
Sarcopterinus nidulans: Railliet, 1893: 697; Berlese, 1913: 130
Sarcoborus nidulans: Oudemans, 1906: 75
Harpyrhynchus nidulans (Nitzsch): Turk, 1953: 18; Jeleva, 1970: 63
Type locality: Germany
Distribution: Bulgaria, Czech Rep., England, Germany, Italy, Slovakia, Switzerland
Hosts: Coccothraustes coccothraustes, Chloris chloris, Fringilla coelebs (Passeriformes: Fringillidae), Alauda arvensis, Galerida cristata (Passeriformes: Alaudidae), Locustella lanceolata (Passeriformes: Locustellidae), Panurus biarmicus (Passeriformes: Panuridae), Corvus corone (Passeriformes: Corvidae), Columba palumbus (Columbiformes: Columbidae)
Harpirhynchus quasimodo Bochkov & Mertins, 2010: 83
Type locality: Florida
Distribution: USA (Florida)
Host: Molothrus ater (Passeriformes: Icteridae)
Distribution: Turkey
Host: Perdix (Galliformes: Phasianidae)
Distribution: SE Asia (McClure H. Eliott, N. Ratanaworadhan et al., 1973)
Host: Eurystomus orientalis (Coraciiformes: Coraciidae)
Harpirhynchus (Neharpyrhynchus) Fain, 1972: 50
Neharpyrhynchus Fain, 1995: 78
Type species: Harpirhynchus plumaris Fritsch, 1954
Neharpyrhynchus agelaius Bochkov, OConnor & Klompen, 2015: 102
Type locality: Michigan, Oakland Co.
Distribution: USA
Host: Agelaius phoeniceus (Passeriformes: Icteridae)
Neharpyrhynchus baile: Bochkov & Literák, 2007: 35; Martinu et al. 2008: 207; Bochkov, OConnor & Klompen, 2015: 98
Type locality: Brazil
Distribution: Brazil, USA
Hosts: Turdus leucomelas (Passeriformes: Turdidae), Dumetella carolinensis (Passeriformes: Mimidae)
Neharpyrhynchus bochkovi Martinu, Dusbábek & Literák, 2008: 198
Type locality: Jinacovice
Distribution: Czech Rep.
Host: Turdus merula (Passeriformes: Turdidae)
Neharpyrhynchus bombycilla Bochkov, OConnor & Klompen, 2015: 96
Type locality: Michigan, Muskegon Co., Muskegon State game Area, US route 31, 43.2961989°N, 86.0668945°W
Distribution: USA
Host: Bombycilla cedrorum (Passeriformes: Bombycillidae)
Neharpyrhynchus campylorhynchus Bochkov, OConnor & Klompen, 2015: 87
Type locality: USA
Distribution: USA
Host: Campylorhynchus brunneicapillus (Passeriformes: Troglodytidae)
Neharpyrhynchus chlorospingus Bochkov & Literák, 2011: 20; Bochkov, OConnor & Klompen, 2015: 108
Type locality: Cerro de la Mueste, 9°34‘N, 83°45‘W
Distribution: Costa Rica
Host: Chlorospingus pileatus (Passeriformes, Passerellidae)
Neharpyrhynchus domrowi Bochkov & Klompen, 2015: 25
Type locality: New South Walles, Loddon Falls, 34.28 S, 150.9 E
Distribution: Australia, Papua New Guinea
Hosts: Acanthorhynchus tenuirostris, Ptiloprora perstriata, Myzomela rodenbergii (Passeriformes: Meliphagidae)
Neharpyrhynchus hippolae Bochkov, 2000a: 534; Bochkov & Literák, 2006: 140; Martinu, Dusbábek & Literák, 2008: 200
Type locality: Oskui, Novgorodskaya Oblast, Russia
Distribution: Czech Rep., Russia
Host: Hippolais icterina (Passeriformes: Sylviidae)
Neharpyrhynchus icterus Bochkov & OConnor, 2013: 506; Bochkov, OConnor & Klompen, 2015: 101
Type locality: California
Distribution: USA
Host: Icterus galbula (Passeriformes: Icteridae)
Neharpyrhynchus loxia Bochkov & Galloway, 2013: 413; Bochkov, OConnor & Klompen, 2015: 108
Type locality: Manitoba, Winnipeg
Distribution: Canada
Host: Loxia curvirostra (Passeriformes: Fringillidae)
Neharpyrhynchus mironovi Bochkov & Literák, 2011: 23; Bochkov, OConnor & Klompen, 2015: 104
Type locality: Minas Gerais, Belo Horizonte, Nova Lima
Distribution: Brazil
Host: Dacnis cayana (Passeriformes, Thraupidae)
Harpyrhynchus novoplumaris Moss, Oliver & Nelson, 1968: 377
Neharpyrhynchus novoplumaris: Fain, 1995: 85 (syn. of N. plumaris)
Neharpyrhynchus novoplumaris: Bochkov, 2000a: 537; Bochkov & OConnor, 2013: 508 (separate species); Bochkov, OConnor & Klompen, 2015: 108
Type locality: Hopland Field Station, Mendocino Co., California
Distribution: USA (California, Arizona, Kansas, Michigan, Missouri, Maryland, Nebraska, Wyoming, New Mexico), Mexico
Hosts: Certhia familiaris (Certhiidae), Baeolophus [Parus] bicolor (Paridae), Kieneria [Pipilo, Melozone] fusca, Amphispiza bilineata, Spizella passerina ….(Passerellidae), Campylorhynchus brunneicapillus (Troglodytidae), Cardinalis cardinalis (Cardinalidae), Junco hyemalis, J. phaeonotus (Emberizidae)(all Passeriformes)
Neharpyrhynchus oenanthe Bochkov, 2014: 483
Type locality: Akanyaru
Distribution: Rwanda
Host: Oenanthe oenanthe (Passeriformes: Muscicapidae)
Neharpyrhynchus pari Martinu, Dusbábek & Literák, 2008: 201
Type locality: Czech Rep., Brno, Sobesice
Distribution: Czech Rep., USA ?
Hosts: Baeolophus bicolor, Cyanistes caeruleus, Parus major, Periparus ater, Poecile montanus, P. palustris (Passeriformes: Paridae)
Neharpyrhynchus picidarum Bochkov, OConnor & Klompen, 2015: 88
Type locality: Michigan, Oakland Co., Pontiac, 42.6110959°N, 83.2877398°W
Distribution: USA
Hosts: Colaptes auratus, Melanerpes formicivorus, M. uropygidialis, Picoides pubescens (Piciformes: Picidae)
Harpirhynchus pilirostris Berlese & Trouessart, 1889: 137 – 138; Dubinin, 1957: 104; Lawrence, 1959b: 108
Sarcoborus pilirostris: Oudemans, 1906: 76
Sarcopterus pilirostris (Berlese et Trouessart) Berlese, 1894: fasc. 73, tav. 5, figs.1–5; nec Sarcopterus nidulans Nitzsch, 1818
Harpyrhynchus ovalis Fritsch, 1957: 193; Lawrence, 1959a: 424 (deutonymph)
Harpirhynchus (Neharpyrhynchus) Fain, 1972: 50
Neharpyrhynchus pilirostris (Berlese et Trouessart): Fain, 1995: 81; Bochkov, 2000a: 534; Bochkov & Literák, 2006: 140; Martinu, Dusbábek & Literák, 2008: 203; Bochkov & Galloway, 2013: 414; Bochkov, OConnor & Klompen, 2015: 101
Type locality: Blanc (Indre, France)
Distribution: Canada, Czech Rep., France, Germany, South Africa, USA
Hosts: Passer domesticus (Passeriformes: Passeridae), ? Aegithalos caudatus (Passeriformes: Aegithalidae)
Harpirhynchus plumaris Fritsch, 1954: 193
Harpyrhynchus plumaris: Dubinin, 1957: 108
Harpyrhynchus ovalis Fritsch, 1954: 192; Lawrence, 1959b: 108
? Harpirhynchus novoplumaris Moss et al., 1968: 377 (Fain, 1995)
Neharpyrhynchus novoplumaris, Fain, 1995: 85
Neharpyrhynchus plumaris (Fritsch): Bochkov, 2000a: 536; Martinu, Dusbábek & Literák, 2008: 203; Bochkov, OConnor & Klompen, 2015: 111
Type locality: Germany
Distribution: Czech Rep., Germany, Russia
Hosts: Fringilla coelebs, Chloris chloris, Carduelis cannabina (Passeriformes: Fringillidae), Muscicapa striata (Pass.: Muscicapidae), Parus ater, P. palustris (Passeriformes: Paridae), ? Aythya ferina (Anseriformes: Anatidae)
Neharpyrhynchus quiscalus Bochkov, OConnor & Klompen, 2015: 98
Type locality: Michigan, Mecosta Co., 1.3 mi E Big Rapids, 43.698604°N,
85.4576146°W
Distribution: USA
Host: Quiscalus quiscula (Passeriformes: Icteridae)
Neharpyrhynchus schoenobaenus Martinu, Dusbábek & Literák, 2008: 203
Type locality: Czech Rep., Sedlec
Distribution: Czech Rep.
Host: Acrocephalus schoenobaenus (Passeriformes: Acrocephalidae)
Neharpyrhynchus spinus Martinu, Dusbábek & Literák, 2008: 204
Type locality: Česke Budejovice
Distribution: Czech Rep.
Host: Spinus spinus (Passeriformes: Fringillidae)
Type locality: Czech Rep.
Distribution: Paraguay (Literák, Bochkov & Franco, 2014)
Host: Setophaga pitiayumi (Passeriformes: Parulidae)
Neharpyrhynchus spizella Bochkov, OConnor & Klompen, 2015: 105
Type locality: California, Mendocino Co., Hopland Field Station, 39°N 123.08°W
Distribution: USA (California, Wyoming)
Host: Spizella passerina (Passeriformes: Passerellidae)
Harpyrhynchus (Neharpyrhynchus) squamiferus Fain, 1972: 55
Neharpyrhynchus squamiferus (Fain): Fain, 1995: 80
Neharpyrhynchus squamifer (Fain): Bochkov, 2000a: 534
Type locality: India
Distribution: India
Host: Temenuchus pagodarum (Passeriformes: Sturnidae)
Neharpyrhynchus tangara Bochkov & Literák, 2011: 20; Literák, Bochkov & Franco, 2014; Bochkov, OConnor & Klompen, 2015: 104
Type locality: Minas Gerais, Belo Horizonte, Nova Lima, Água Limpa
20°13‘S, 43°56‘W (Brazil)
Distribution: Brazil, Paraguay, Peru
Hosts: Paroaria capitata, P. capitata, Tangara cayana, Thraupis episcopus (Passeriformes: Thraupidae)
Harpirhynchus (Neharpyrhynchus) trochilinus Fain, 1972: 55
Neharpyrhynchus trochilinus (Fain): Fain, 1995: 80; Bochkov, 2000a: 534; Bochkov & Literák, 2011: 28; Literák et al., 2012: 109; Literák, Bochkov & Franco, 2014: 184; Bochkov, OConnor & Klompen, 2015: 94
Type locality: Antwerp Zoo
Distribution: Brazil, Costa Rica, Paraguay, Peru
Hosts: Amazilia lactea, Chlorostilbon lucidus, Chrysolampis moschitus, Eugenes fulgens, Panterpe insignis (Trochiliformes: Trochilidae)
Neharpyrhynchus turdus Bochkov, OConnor & Klompen, 2015: 92
Type locality: Michigan, Washtenaw Co., 4.4 mi ESE Saline, 42.1561503°N, 83.7180519°W
Distribution: USA
Host: Turdus migratorius (Passeriformes: Turdidae)
Neharpyrhynchus vireo Bochkov, OConnor & Klompen, 2015: 104
Type locality: Florida, Leon Co., 30.49°N, 84.21°W
Distribution: USA
Host: Vireo olivaceus (Passeriformes: Vireonidae)
Ralliharpirhynchus Fain, 1995: 74
Type species: Metharpyrhynchus porphyrio Fain, 1972
Metharpyrhynchus limnocorax Fain, 1972: 56
Ralliharpirhynchus limnocorax: Fain, 1995: 74
Type locality: Kinshasa
Distribution: DR Congo
Host: Limnocorax flaviceps (Ralliformes: Rallidae)
Metharpyrhynchus porphyrio Fain, 1972: 56
Ralliharpirhynchus porphyrio, Fain, 1995: 74
Type locality: Antwerp Zoo
Distribution: ?
Host: Porphyrio porphyrio (Ralliformes: Rallidae)
Trichorhynchiella Fain, 1995: 75
Type species: Trichorhynchiella paddae Fain, 1995
Trichorhynchiella myiarchus Bochkov & Klompen, 2014b: 304; Bochkov, OConnor & Klompen, 2015: 113
Type locality: unknown
Distribution: North America
Host: Myiarchus crinitus (Passeriformes: Tyrannidae)
Trichorhynchiella paddae Fain, 1995: 99
Type locality: Antwerp Zoo
Distribution: ?
Host: Padda oryzivora (Passeriformes: Estrildidae)
Trichorhynchiella phleocryptes Bochkov & OConnor, 2014a: 324; Bochkov, OConnor & Klompen, 2015: 117
Type locality: Argentina
Distribution: Argentina
Host: Phleocryptes melanops (Passeriformes: Furnariidae)
Metharpyrhynchini Fain, 1972: 56
Type genus: Metharpyrhynchus Fain, 1972
Metharpyrhynchus Fain, 1972: 56
Type species: Metharpyrhynchus macrophallus Fain, 1972
Metharpyrhynchus jynx Fain, 1972: 57
Type locality: Akanyaru
Distribution: Rwanda
Host: Jynx ruficollis (Piciformes: Picidae)
Metharpyrhynchus macrophallus Fain, 1972: 56
Type locality: Antwerp Zoo
Distribution: ?
Host: Padda oryzivora (Passeriformes: Estrildidae)
Metharpyrhynchus mossi Fain, 1995: 87
Type locality: Akanyaru, Eastern region of Rwanda
Distribution: Rwanda
Host: Prinia subflava graueri (Passeriformes: Cisticolidae)
Metharpyrhynchus namibiensis Fain, 1995: 87
Type locality: Arab
Distribution: Namibia
Host: Philetairus socius (Passeriformes: Ploceidae)
Tribus Perharpyrhynchini Fain, 1972: 57
Type genus: Perharpyrhynchus Fain, 1972
Perharpyrhynchus Fain, 1972: 57
Type species: Perharpyrhynchus jacana Fain, 1972
Perharpyrhynchus caprimulgus Bochkov & Klompen, 2014: 313
Type locality: Mozambique
Distribution: Mozambique
Host: Caprimulgus fossii (Caprimulgiformes: Caprimulgidae)
Perharpyrhynchus charadrius Bochkov, OConnor & Klompen, 2015: 80
Type locality: Michigan, Washtenaw Co., Ann Arbor, UM campus, 42.2793409°N, 83.7386084°W
Distribution: USA
Host: Charadrius vociferus (Charadriiformes: Charadriidae)
Perharpyrhynchus elseyornis Bochkov & Klompen, 2014: 314
Type locality: Western Australia, Brooking Springs, 18.12°S, 125.58°E
Distribution: Australia
Host: Elseyornis melanops (Charadriiformes: Charadriidae)
Perharpyrhynchus jacana Fain, 1972: 57
Type locality: Antwerp Zoo
Distribution: South America
Host: Jacana spinosa intermedia (Charadriiformes: Jacanidae)
Perharpyrhynchus recurvirostra Fain, 1976: 130
Type locality: Antwerp Zoo
Distribution: ?
Host: Recurvirostra avocetta (Charadriiformes: Scolopacidae)
EUROPE
Austria (Černy & Sixl, 1971; Bochkov & Literák, 2006) – Harpirhynchus dusbabeki, Harpyrhynchoides rubeculinus
Azerbaidjan (Fain, Bochkov & Mironov, 1999) – Harpirhynchus galeridae
Belgium (Fain, 1972; Fain, Bochkov & Mironov, 1999) – Anharpyrhynchus monstrosus, Harpypalpus holopus, Harpyrhynchoides asio, H. columbae, H. herodius, H. tyto
Bulgaria (Jeleva, 1970) – Harpirhynchus nidulans
Croatia (Bochkov & Literák, 2006) – Harpirhynchus dusbabeki
Czech Rep. (Literák et al., 2005; Bochkov & Literák, 2006, 2008; Martinu, Dusbábek & Literák, 2008) – Harpyrhynchoides rubeculinus, Harpirhynchus dusbabeki, H. nidulans, Neharpyrhynchus bochkovi, N. hippolae, N. pari, N. pilirostris, N. plumaris, N. schoenobaenus, N. spinus
France (contin.)(Berlese & Trouessart, 1889; Fain, Bochkov & Mironov, 1999; Bochkov & Literák, 2006) – Harpyrhynchoides alaudinus, H. capellae, H. rubeculinus, Harpirhynchus dusbabeki, Neharpyrhynchus pilirostris
Germany (Fritsch, 1954; Bochkov, OConnor & Klompen, 2015) – Anharpyrhynchus monstrosus, Harpyrhinchiella reductus, Harpyrhinchoides capellae, H. cylindripalpis, H. parazumpti, H. tracheatus, Harpirynchus nidulans, Neharpyrhynchus pilirostris, N. plumaris
Great Britain (Turk, 1953) – Harpirhynchus nidulans
Holland (the Netherlands)(Lombert & Moss, 1983; Bochkov & Klompen, 2014a) – Harpypalpoides hirundinoides, H. johnstoni, H. lukoschusi, H. regulus, H. sylvia, Harpypalpus longipes, H. pyrrhula, H. sturnus, Harpirhynchiella reductus, Harpyrinchoides herodius
Italy (contin.)(Canestrini, 1886; Berlese & Trouessart, 1889) – Harpirhynchus nidulans, Harpypalpus holopus
Luxemburg (Fain, 1994) – Harpyrhynchoides tracheatus
Moldavia (Fain, Bochkov & Mironov, 1999) – Harpyrhynchoides parazumpti
Poland (Skoracki, Bochkov & Sikora, 2004; Bochkov & Literák, 2006) – Harpypalpoides lesickii, H. lukoschusi, Harpypalpus longipes, Harpirhynchus dusbabeki
Russia (all, continent.)(Bochkov, 2000a, 2000b, 2015) – Harpyrhynchiella apus, Harpyrhynchoides alaudinus, Neharpyrhynchus hippolae, N. plumaris
Slovakia (Literák et al., 2005; Bochkov & Literák, 2006) – Harpirhynchus dusbabeki, H. nidulans
Spain (cont.)(Fain, Bochkov & Mironov, 1999) – Harpyrhynchoides alectoris
Switzerland (Fain, 1995) – Harpirhynchus nidulans
Turkey (Aksin & Erdoğmuş, 2005) – Harpirhynchus sp.
AFRICA
Congo DR (Fain, 1972; Fain, Bochkov & Mironov, 1999) – Harpyrhinchiella [Cypsharpirhynchus] cypsiuri, Harpypalpoides hirundinis, Harpypalpus spermestes, Harpyrhynchoides vercammeni, Ralliharpirhynchus limnocorax,
Mozambique (Bochkov & Klompen, 2014) – Perharpyrhynchus caprimulgus
Namibia (Fain, 1995; Fain, Bochkov & Mironov, 1999) – Metharpyrhynchus namibiensis, Harpypalpoides namibiensis, Harpyrhinchoides cristagalli
Rwanda (Fain, 1994, 1995; Bochkov, 2015) – Harpyrhynchoides pectinifer, Metharpyrhynchus jynx, M. mossi, Neharpyrhynchus oenanthe
South Africa (Lawrence, 1959; Fain, 1972; Bochkov, 2015) – Harpyrhynchoides clamator, H. cristagalli, H. pectinifer, H. pilirostris, H. zumpti, Harpirhynchus agapornis
Togo (Fain, 1972) – Cypsharpirhynchus cypsiuri
Zimbabwe (Fain, 1994) – Harpyrhynchoides pectinifer
Central Africa (Fain, Bochkov & Mironov, 1999) – Harpypalpus tiarae
Africa (Lawrence, 1959b) – Harpyrhynchoides numidae
ASIA
India (Fain, 1972) – Neharpyrhynchus squamiferus
Kazakhstan (Bochkov & Klompen, 2014a) – Harpypalpoides johnstoni, H. lesickii
Kirghizstan (Fain, Bochkov & Mironov, 1999) – Harpyrhynchoides kirgizorum
Malaysia (Berlese & Trouessart, 1889) – Harpypalpus sp. pr. holopus
SE Asia (Bochkov & Klompen, 2015) – Harpyrhynchoides artamus
NORTH AMERICA
Canada (Bochkov & Galloway, 2001, 2004; 2013; Bochkov & Klompen, 2014; Bochkov, OConnor & Klompen, 2015) – Fainharpyrhynchus contopus, Harpyrhynchoides actitis, H. botaurus, H. gallowayi, H. heatherae, H. modestus, H. phalaropus, H. setophaga, H. tracheatus, H. vulgaris, H. zumpti, Neharpyrhynchus loxia, N. pilirostris
Mexico (Bochkov, OConnor & Klompen, 2015) – Junco phaeonotus
USA (Banks, 1905; Chaddock, 1941; Boyd, 1968; Moss, Oliver & Nelson, 1968; Moss, Mitchell & Johnston, 1970; Bochkov & Mertins, 2010; Spelding et al., 2010; Bochkov & OConnor, 2013; Bochkov & Klompen, 2014a, b; Bochkov, OConnor & Klompen, 2015) – Anharpyrhynchus elizae, Crassacarus alexfaini, C. fritschi, C. melanerpes, C. sialia, C. tinae, Fainharpirhynchus contopus, Harpypalpoides sitta, Harpyrhynchoides accipiter, H. aegolius, H. athene, H. bubulcus, H. butorides, H. calidris, H. charadrius, H. coccyzus, H. herodius, H. ixobrychus, H. megascops, H. parazumpti, H. pectinifer, H. pluvialis, puffinus, H. rubeculinus, H. spizella, H. tracheatus, H. xanthocephalus, H. zenaida, Harpirhynchus quasimodo, H. longipilis, Neharpyrhynchus agelaius, N. baile, N. bombycilla, N. campylorhynchus, N. icterus, N. novoplumaris, ? N. pari, N. quiscalus, N. picidarum, N. pilirostris, N. spizella, N. turdus, N. vireo, Perharpyrhynchus charadrius
North America (Bochkov & Klompen, 2014b) – Trichorhynchiella myiarchus
SOUTH and CENTRAL AMERICA
Argentina (Bochkov & OConnor, 2014) – Trichorhynchiella phleocryptes
Brazil (Bochkov & Literák, 2007, 2011; Bochkov, OConnor & Klompen, 2015) – Neharpyrhynchus baile, N. mironovi, N. rubeculinus, N. tangara, N. trochilinus
Costa Rica (Bochkov & Literák, 2011) – Neharpyrhynchus chlorospingus, N. trochilinus
Paraguay (Literák, Bochkov & Franco, 2014) – Neharpyrhynchus tangara, N. trochilinus, N. aff. spinus
Peru (Literák et al., 2012) – Neharpyrhynchus tangara, N. trochilinus
Suriname (Bochkov, OConnor & Klompen, 2015) – Harpyrhynchoides crotophaga
Trinidad and Tobago (Bochkov & Klompen, 2014) – Fainharpirhynchus legatus
South America (Fain, 1972) – Harpyrhynchoides amazonae, H. metropeliae, Neharpirhynchus trochilinus, Perharpyrhynchus jacana
OCEANIA
Australia (continent)(Fain, 1972; Moss, 1979a; Filippich & Domrow, 1985; Domrow, 1991; Bochkov & Klompen, 2014a, 2015) – Anharpyrhynchus apodus, A. lukoschusi, A. monstrosus, Harpypalpus lonchura, H. taeniopygia, Harpyrhynchoides kakatoe, H. rosellacinus, Neharpyrhynchus domrowi, Perharpyrhynchus elseyornis
Tasmania – Harpyrhynchoides rosellacinus
Papua New Guinea (Bochkov & Klompen, 2015) – Neharpyrhynchus domrowi
Unknown distribution (Zoo birds): Harpypalpus dubinini, H. serini, Harpyrhinchoides amazonae, H. anatum, H. capitatus, H. coturnix, H. coxatus, H. diuca, H. lawrencei, H. leptopilus, H. metropeliae, H. modestus, H. oenae, H. oenae, H. lamorali, H. psittaci, H. psittaculae, H. squamosus, H. trochilinus, Ralliharpirhynchus porphyrio, Trichorhinchiella paddae, Metharpyrhynchus macrophallus, Perharpyrhynchus recurvirostra, P. jacana
Aves
Accipitriformes
Fam. Accipitridae
Accipiter striatus – Harpyrhynchoides accipiter
Buteo buteo – Harpyrhynchoides tracheatus
B. jamaicensis – Harpyrhynchoides tracheatus
B. lineatus – Harpyrhynchoides tracheatus
Anseriformes
Fam. Anatidae
Anas querquedula – Harpyrhynchoides anatum
Aythya ferina – Neharpyrhynchus plumaris (doubtful – see Bochkov & Literák, 2006)
Apodiformes
Fam. Apodidae
Apus affinis – Harpyrhynchiella [Cypsharpirhynchus] cypsiuri
A. apus – Harpyrhynchiella reductus
Cypsiurus parvus – Harpyrhynchiella [Cypsharpirhynchus] cypsiuri
Fam. Trochilidae
Amazilia lactea – Neharpyrhynchus trochilinus
A. chionogaster – Neharpyrhynchus trochilinus
Chlorostilbon lucidus – Neharpyrhynchus trochilinus
Chrysolampis moschitus – Neharpyrhynchus trochilinus
Eugenes fulgens – Neharpyrhynchus trochilinus
Panterpe insignis – Neharpyrhynchus trochilinus
Caprimulgiformes
Fam. Caprimulgidae
Caprimulgus fossii – Perharpyrhynchus caprimulgus
Charadriiformes
Fam. Charadriidae
Charadrius vociferus – Harpyrhynchoides charadrius, Perharpyrhynchus charadrius
Elseyornis melanops – Perharpyrhynchus elseyornis
Pluvialis dominica – Harpyrhynchoides pluvialis
Fam. Jacanidae
Jacana spinosa intermedia – Perharpyrhynchus jacana
Fam. Scolopacidae
Actitis macularius – Harpyrhynchoides actitis
Calidris minutilla – Harpyrhynchoides calidris
Gallinago gallinago – Harpyrhynchoides capellae
Lymnocryptes minimus – Harpyrhynchoides capellae
Phalaropus lobatus – Harpyrhynchoides phalaropus
Recurvirostra avocetta – Perharpyrhynchus recurvirostra
Ciconiiformes
Fam. Ciconiidae
Leptoptilus crumeniferus – Harpyrhynchoides leptoptilus
Coliiformes
Fam. Coliidae
Colius colius – Harpyrhynchoides cristagalli
C. striatus – Harpyrhynchoides cristagalli
Columbiformes
Fam. Columbidae
Columba livia domestica – Harpyrhynchoides columbae
C. livia – Harpyrhynchoides gallowayi, H. modestus
C. palumbus – Harpirhynchus nidulans (= palumbinus)
Columbigallina talpacoti – Harpyrhynchoides capitatus, H. coxatus
Columbina cruziana – Harpyrhynchoides coxatus
Metropelia caeciliae – Harpyrhynchoides metropeliae, H. modestus
Oena capensis – Harpyrhynchoides oenae
Scardafella squamata – Harpyrhynchoides coxatus
Tympanistria tympanistria – Harpyrhynchoides oenae [lamorali]
Zenaida macroura – Harpyrhynchoides zenaida
Cuculiformes
Fam. Cuculidae
Clamator jacobinus – Harpyrhynchoides clamator
Coccyzus americanus – Harpyrhynchoides coccyzus
C. erythrophthalmus – Harpyrhynchoides coccyzus
Crotophaga cardinalis – Harpyrhynchoides crotophaga
Galliformes
Fam. Phasianidae
Alectoris graeca – Harpyrhynchoides alectoris
A. rufa – Harpyrhynchoides alectoris
Coturnix delegorguei – Harpyrhynchoides coturnix
Fam. Numididae
Numida meleagris – Harpyrhynchoides numidae
Charadriiformes
Fam. Charadriidae
Charadrius vociferus – Perharpyrhynchus charadrius
Elseyornis melanops – Perharpyrhynchus elseyornis
Fam. Scolopacidae
Phalaropus lobatus – Harpyrhynchoides phalaropus
Passeriformes
Fam. Acrocephalidae
Acrocephalus schoenobaenus – Neharpyrhynchus schoenobaenus
Fam. Aegithalidae
Aegithalos caudatus – Neharpyrhynchus pilirostris (?)
Fam. Alaudidae
Alauda arvensis – Harpyrhynchoides alaudinus, Harpirhynchus nidulans
Galerida cristata – Harpirhynchus galeridae, H. nidulans
Eremopterix leucotis – Harpyrhynchoides zumpti
Fam. Artamidae
Artamus fuscus – Harpyrhynchoides artamus
Fam. Bombycillidae
Bombycilla cedrorum – Crassacarus fritschi, Neharpyrhynchus bombycilla
Fam. Cardinalidae
Cardinalis cardinalis – Crassacarus alexfaini, Neharpyrhynchus novoplumaris
Passerina ciris – Crassacarus cylindripalpus
Pheucticus ludovicianus – Harpyrhynchoides rubeculinus
Fam. Certhiidae
Certhia familiaris – Neharpyrhynchus novoplumaris
Fam. Cisticolidae
Prinia subflava graueri – Metharpyrhynchus mossi
Fam. Corvidae
Corvus brachyrhinchos – Harpyrhynchoides parazumpti
C. corone – Harpirhynchus nidulans
C. frugilegus – Harpyrhynchoides parazumpti
C. monedula – Anharpyrhynchus monstrosus, Harpyrhynchoides parazumpti
Cyanocitta cristata – Anharpyrhynchus elizae
C. stelleri – Anharpyrhynchus elizae
Garrulus glandarius – Anharpyrhynchus monstrosus
Nucifraga caryocatactes – Harpypalpus longipes
Xanthoura yncas luxuosa – Anharpyrhynchus monstrosus
Fam. Emberizidae
Emberiza calandra – Harpyrhynchoides kirgizorum
E. cioides – Harpirhynchus sp.
E. impetuani – Harpypalpoides namibiensis
Junco hyemalis – Harpyrhynchoides heatherae, H. zumpti, Neharpyrhyncus novoplumaris
J. phaeonotus – Neharpyrhyncus novoplumaris
Fam. Estrildidae
Lonchura castaneothorax – Harpypalpus lonchura
Padda oryzivora – Metharpyrhynchus macrophallus, Trichorhynchiella paddae
Taeniopygia guttata – Harpypalpus taeniopygia
Fam. Fringillidae
Amphispiza bilineata – Neharpyrhynchus novoplumaris
Chloris chloris – Harpirhynchus nidulans, Neharpyrhynchus plumaris
Carduelis cannabina – Harpypalpoides lesickii, Harpypalpus longipes, Neharpyrhynchus plumaris
C. tristis – Crassacarus tinae
Coccothraustes coccothraustes – Harpirhynchus nidulans, Harpypalpoides lesickii
C. vespertina – Harpyrhynchoides brevis
Fringilla coelebs – Harpypalpoides johnstoni, Harpirhynchus cylindripalpus, H. nidulans, Neharpyrhynchus plumaris
F. montifringilla – Harpypalpoides johnstoni
Loxia curvirostra – Harpyrhynchoides brevis, Neharpyrhynchus loxia
Loxia sp. – “Harpirhynchus longipilus”
Pyrrhula pyrrhula – Harpypalpus pyrrhula
Serinus mozambicus – Harpypalpus serini
Spinus spinus – Harpypalpus longipes, Neharpyrhynchus spinus
Fam. Furnariidae
Phleocryptes melanops – Trichorhynchiella phleocryptes
Fam. Hirundinidae
Hirundo rustica – Harpypalpoides hirundinoides
Psalidoprocne sp. – Harpypalpoides hirundinis
Fam. Icteridae
Agelaius phoeniceus – Neharpyrhynchus agelaius
Icterus galbula – Neharpyrhynchus icterus
Molothrus ater – Harpirhynchus quasimodo
Quiscalus quiscula – Neharpyrhynchus quiscula
Xanthocephalus xanthocephalus – Harpypalpoides xanthocephalus
Fam. Locustellidae
Locustella lanceolata – Harpirhynchus nidulans
Fam. Meliphagidae
Acanthorhynchus tenuirostris – Neharpyrhynchus domrowi
Certhionyx pectoralis – Anharpyrhynchus lukoschusi
Gliciphila indistincta – Anharpyrhynchus monstrosus
Lichmera indistincta – Anharpyrhynchus apodus
Manorina flavigula – Anharpyrhynchus lukoschusi
Meliornis novaehollandiae – Anharpyrhynchus monstrosus
Myzantha flavigula – Anharpyrhynchus monstrosus
Myzomela pectoralis – Anharpyrhynchus monstrosus
M. rosenbergii – Neharpyrhynchus domrowi
Phylidonyris novaehollandiae – Anharpyrhynchus apodus
Ptiloprora perstriata – Neharpyrhynchus domrowi
Fam. Mimidae
Dumetella carolinensis – Neharpyrhynchus baile
Fam. Muscicapidae
Erithacus rubecula – Harpyrhynchoides rubeculinus
Luscinia svecica – Harpypalpus holopus
Muscicapa striata – Neharpyrhynchus plumaris
Oenanthe oenanthe – Neharpyrhynchus oenanthe
Fam. Panuridae
Panurus biarmicus – Harpirhynchus nidulans
Fam. Paridae
Baeolophus bicolor – Neharpyrhynchus pari [novoplumaris ?]
Cyanistes caeruleus – Harpypalpus longipes, Neharpyrhynchus pari
P. major – Harpypalpus longipes, Neharpyrhynchus pari
Periparus ater – Neharpyrhynchus pari, [N. plumaris]
Poecile atricapillus – Harpypalpus longipes
P. montanus – Neharpyrhynchus pari
P. palustris – Neharpyrhynchus pari, [N. plumaris]
Fam. Parulidae
Setophaga petechia – Crassacarus alexfaini
S. pitiayumi – Neharpyrhynchus aff. spinus
S. ruticilla – Harpyrhynchoides setophaga, H. vulgaris
Fam. Passerellidae
Amphispiza bilineata – Neharpyrhynchus novoplumaris
Chlorospingus pileatus – Neharpyrhynchus chlorospingus
Kieneria [Pipilo, Melozone] fusca – Neharpyrhynchus novoplumaris
Passerina ciris – Crassacarus alexfaini
Spizella passerina – Neharpyrhynchus novoplumaris, N. spizella
Fam. Passeridae
Passer domesticus – Harpypalpus holopus, Neharpyrhynchus piliristris
Fam. Ploceidae
Ploceus [Sitagra] intermedius – Harpypalpus dubinini
Ploceus [Spermestes] cucullatus – Harpypalpus spermestes
Fam. Regulidae
Regulus regulus – Harpypalpoides regulus
Fam. Sittidae
Sitta pygmaea – Harpypalpoides sitta
Fam. Sturnidae
Temenuchus pagodarum – Neharpyrhynchus squamiferus
Fam. Sylviidae
Hippolais icterina – Neharpyrhynchus hippolae
Sylvia atricapilla – Harpypalpoides sylvia
Fam. Thraupidae
Dacnis cayana – Neharpyrhynchus mironovi
Diuca diuca – Harpyrhynchoides diuca, H. zumpti
Paroaria capitata – Neharpyrhynchus tangara
P. coronata – Neharpyrhynchus tangara
Tangara cayana – Neharpyrhynchus tangara
Thraupis episcopus – Neharpyrhynchus tangara
Tiaris canora – Harpypalpus tiarae
Fam. Timaliidae
Panurus biarmicus – Harpirhynchus dusbabeki
Fam. Troglodytidae
Campylorhynchus brunneicapillus – Neharpyrhynchus campylochirus, N. novoplumaris
Troglodytes troglodytes – Harpypalpus longipes
Fam. Turdidae
Catharus ustulatus – Harpyrhynchoides rubeculinus
Sialia currucoides – Crassacarus sialia
Turdus leucomelas – Neharpyrhynchus baile
T. merula – Harpypalpoides lukoschusi, Neharpyrhynchus bochkovi
T. migratorius – Neharpyrhynchus turdus
T. pilaris – Harpypalpoides lukoschusi
T. amaurochalinus – Harpypalpoides rubeculinus
Fam. Tyrannidae
Contopus cooperi – Fainharpirhynchus contopus
C. virens – Fainharpirhynchus contopus
Legatus leucophaius – Fainharpirhynchus legatus
Myiarchus crinitus – Fainharpirhynchus mossi, Trichorhynchiella myiarchus
Fam. Vireonidae
Vireo olivaceus – Neharpyrhynchus vireo
Pelecaniformes
Fam. Ardeidae
Ardea cinerea – Harpyrhynchoides herodius
A. herodius – Harpyrhynchoides herodius
Botaurus lentiginosus – Harpyrhynchoides botaurus
Bubulcus ibex – Harpyrhynchoides bubulcus
Butorides virescens – Harpyrhynchoides butorides
Ixobrychus exilis – Harpyrhynchoides ixobrychus
Piciformes
Fam. Picidae
Campethera abingoni – Harpyrhynchoides pectinifer
C. cailliauti – Harpyrhynchoides pectinifer
Colaptes auratus – Anharpyrhynchus elizae, Harpyrhynchoides pectinifer, Neharpyrhynchus picidarum
Jynx ruficollis – Metharpyrhynchus jynx
Melanerpes formicivorus – Crassacarus melanerpes, Neharpyrhynchus picidarum
M. uropygydialis – Neharpyrhynchus picidarum
Picoides pubescens – Neharpyrhynchus picidarum
Procellariiformes
Fam. Procellariidae
Puffinus gravis – Harpyrhynchoides gravis
Psittaciformes
Fam. Cacatuidae
Kakatoe galerita – Harpyrhynchoides kakatoe
Fam. Psittacidae
Amazona aestiva – Harpyrhynchoides amazonae
Nandays nanday – Harpyrhynchoides lawrencei
Poicephalus senegalus – Harpyrhynchoides psittaci
Psittacus erithacus – Harpyrhynchoides psittaci
Fam. Psittaculidae
Agapornis roseicollis – Harpyrhynchoides agapornis
A. pullarius – Harpyrhynchoides agapornis
A. nigrigenis – Harpyrhynchoides agapornis
Glossopsitta concinna – Harpyrhynchoides rosellacinus
Lathamus discolor – Harpyrhynchoides rosellacinus
Platycercus eximius – Harpyrhynchoides rosellacinus
Psittacula alexandri – Harpyrhynchoides psittaculae
P. cyanocephala – Harpyrhynchoides psittaculae
Trichoglossus chlorolepidotus – Harpyrhynchoides rosellacinus
T. moluccanus – Harpyrhynchoides rosellacinus
Ralliformes
Fam. Rallidae
Limnocorax flaviceps – Ralliharpirhynchus limnocorax
Porphyrio porphyrio – Ralliharpirhynchus porphyrio
Strigiformes
Fam. Tytonidae
Centropus toulou grillii – Harpyrhynchoides vercammeni
Tyto alba – Harpyrhynchoides tyto
Fam. Strigidae
Aegolius acadicus – Harpyrhynchoides aegolius
Asio otus otus – Harpyrhynchoides asio
Athene canicularia – Harpyrhynchoides athene
Megascops asio – Harpyrhynchoides megascops
Mites living all their live inside the scales of snakes of the families Colubridae, Elapidae and Viperidae. Their study started by Sambon (1928), followed by Ewing (1933), Allred (1958), and Lisazo (1983), again concerning South American or Cuban snakes. Radford (1947) recorded the group from Asia and Southcott (1956), from Australia, describing these mites as a new family. Fain (1962, 1964, 1965) described the new genus Afrophioptes from African snakes, nine new species and revised the group, still as a family. Beron (1972) recorded from Bulgaria Ophioptes for the first time in Europe (confirmed by Fain, who prospected thousands of preserved snakes from Europe, without detecting any Ophioptid). It is clear that these mites prefer the warmer climate. Revising Cheyletoidea Bochkov (2002) followed Dubinin (1957), ascribing to them subfamily status in Harpirhynchidae. Still only 18 species in two genera are known.
Ophioptidae Southcott, 1956: 147
Ophioptinae (subf. of Harpirhynchidae): Dubinin, 1957: 98; Bochkov, 2002
Type genus: Ophioptes Sambon, 1928
Afrophioptes Fain, 1962: 157
Type species: Afrophioptes whartoni Fain, 1962
Afrophioptes rhodesiensis Fain, 1962: 158
Type locality: Abercorn (now Mbala)
Distribution: “Rhodesia” (Zambia)
Host: “Cerastes” t. tritaeniatus (Serpentes: Viperidae)
Afrophioptes whartoni Fain, 1962: 158; 1964: 48; 1965: 112
Type locality: Musosa, Katanga
Distribution: Angola, DR Congo
Host: Psammophis s. sibilans (Serpentes: Colubridae)
Ophioptes Sambon, 1928: 138
Type species: Ophioptes parkeri Sambon, 1928
Ophioptes beshkovi Beron, 1972: 689
Type locality: Frolosh Village, Kyustendil Distr.
Distribution: Bulgaria
Host: Platyceps najadum (=Coluber dahli) (Serpentes: Colubridae)
Ophioptes boaedoni Fain, 1962: 156; 1964: 35
Type locality: Teturi, Kivu
Distribution: DR Congo
Host: Boaedon fuliginosus (Serpentes: Colubridae)
Ophioptes brevipilis Lizaso, 1983: 380
Type locality: Brazil
Distribution: Brazil
Host: Chironius flavolineatus (Serpentes: Colubridae)
Ophioptes coluber Radford, 1947: 229; Fain, 1964: 43
Type locality: Imphal, Manipur State
Distribution: India
Host: Coelognathus [Coluber] radiatus (Serpentes: Colubridae)
Ophioptes congolensis Fain, 1962: 155; 1964: 37
Type locality: Upemba, Mpala
Distribution: DR Congo
Host: Philothamnus hoplogaster (Serpentes: Colubridae)
Ophioptes dromicus Allred, 1958b: 107; Fain, 1964: 33
Type locality: Prov. Oriente, Cuba
Distribution: Cuba
Host: Dromicus andreae orientalis (Serpentes: Colubridae)
Ophioptes ekans Mendoza-Roldan & Barros-Battesti, 2017: 2
Type locality: Campo Limpo Paulista
Distribution: Brazil
Host: Crotalus durissus terrificus (Serpentes: Viperidae)
Ophioptes longipilis Lizaso, 1983: 378
Type locality: Brazil
Distribution: Brazil
Host: Oxyrhopus trigeminus (Serpentes: Colubridae)
Ophioptes lycodontis Fain, 1964: 38
Type locality: Java
Distribution: Indonesia (Java)
Host: Lycodon subcinctus (Serpentes: Colubridae)
Ophioptes machadoi Fain, 1965: 110
Type locality: Mwaoka (45 km S Dundo), Dundo
Distribution: Angola
Host: Dispholidus typus punctatus (Serpentes: Colubridae)
Ophioptes najae Fain, 1962: 156; 1964: 41; 1965: 109
Type locality: Kai Bumba, Mayumbe (DR Congo)
Distribution: Angola, DR Congo
Host: Naja melanoleuca (Serpentes: Elapidae)
Ophioptes parkeri Sambon, 1928: 141; Fain, 1964: 21
Ophioptes oudemansi Sambon, 1928: 141
Type locality: Buenavista, Bolivia
Distribution: Argentina, Bolivia, Brazil, Paraguay
Hosts: Erythrolaprus aesculapii, Leimadophis poecilogyrus, L. pictostriatus, Liophis anomalus, Clelia rustica (Serpentes: Colubridae)
Ophioptes samboni Southcott, 1956: 142; Fain, 1964: 47
Type locality: Queensland
Distribution: Australia (Queensland)
Host: Rhynchoelaps fasciolatus (Serpentes: Elapidae)
Ophioptes schoutedeni Fain, 1962: 154; 1964: 35
Type locality: Abercorn (now Mbala)
Distribution: “Rhodésie” (Zambia)
Host: Boaedon lineatus (Serpentes: Colubridae)
Ophioptes southcotti Fain, 1962: 157; 1964: 47
Type locality: Thailand
Distribution: Thailand
Host: Macropisthodon rhodomelas (Serpentes: Colubridae)
Ophioptes tropicalis Ewing, 1933: 53; Allred, 1958a: 287; Fain, 1964: 47
Type locality: Zoo in Washington D.C.
Distribution: Guiana
Host: Erpetodryas carinatus (Serpentes: Colubridae)
EUROPE
Bulgaria (Beron, 1974) – Ophioptes beshkovi
AFRICA
Angola (Fain, 1965) – Afrophioptes whartoni, Ophioptes machadoi, O. najae
Congo (DR)(Fain, 1962) – Afrophioptes whartoni, Ophioptes boaedoni, O. congolensis, O. najae
Zambia (Fain, 1962) – Afrophioptes rhodesiensis, Ophioptes schoutedeni
ASIA
India (Radford, 1947) – Ophioptes coluber
Indonesia (Fain, 1964) – Ophioptes lycodontis
Thailand (Fain, 1962) – Ophioptes southcotti
CENTRAL and SOUTH AMERICA
Argentina (Sambon, 1928) – Ophioptes parkeri
Bolivia (Sambon, 1928, Fain, 1964) – Ophioptes parkeri
Brazil (Fain, 1964, Lizaso, 1983; Mendoza-Roldan et al., 2017) – Ophioptes brevipilis, O. ekans, O. longipilis, O. parkeri
Cuba (Allred, 1958b; Fain, 1964) – Ophioptes dromicus
Guyana (Ewing, 1933) – Ophioptes tropicalis
Paraguay (Sambon, 1928, Fain, 1964) – Ophioptes parkeri
OCEANIA
Australia (Southcott, 1956) – Ophioptes samboni
Serpentes
Fam. Colubridae
Boaedon fuliginosus – Ophioptes boaedoni
B. lineatus – Ophioptes schoutedeni
Chironius flavolineatus – Ophioptes brevipilis
Clelia rustica – Ophioptes parkeri
Platyceps najadum (=Coluber dahli) – Ophioptes beshkovi
Coelognathus [Coluber] radiatus – Ophioptes coluber
Erpetodryas carinatus – Ophioptes tropicalis
Erythrolaprus aesculapii – Ophioptes parkeri
Leimadophis poecilogyrus – Ophioptes parkeri
L. pictostriatus – Ophioptes parkeri
Liophis anomalus – Ophioptes parkeri
Lycodon subcinctus – Ophioptes lycodontis
Macropisthodon rhodomelas – Ophioptes southcotti
Oxyrhopus trigeminus – Ophioptes longipilis
Philothamnus hoplogaster – Ophioptes congolensis
Psammophis s. sibilans – Afrophioptes whartoni
Fam. Elapidae
Naja melanoleuca – Ophioptes najae
Rhyncoelaps fasciolatus – Ophioptes samboni
Fam. Viperidae
“Cerastes” t. tritaeniatus – Afrophioptes rhodesiensis
Crotalus durissus terrificus – Ophioptes ekans
Syringophilidae Lavoipierre, 1953
The quill mites (Syringophilidae) were considered part of Cheyletidae until Lavoipierre (1953) and Dubinin (1957) made of them separate family. The number of the known genera and species in this family literally exploded after the revision of Kethley (1970) who added to the two genera known by this time (Picobia Haller, 1878 and Syringophilus Heller, 1880) another 15 genera. In 1973 he described another one – Colinophilus. Before him Fritsch (1958) revised the genus Syringophilus and some other authors (Oudemans, Poppe, Banks, Berlese, Hancock, Hirst, Lawrence, Womersley, Clark) added to the knowledge of the family, but mostly in general acarological papers. Johnston & Kethley (1973) separated the Picobia in a new subfamily Picobiinae.
The bulk of modern information on Syringophilidae comes from the many important papers of the Polish specialist Skoracky and his co-authors (OConnor, Bochkov, Sikora, Glowska, Fain, Mironov, Hebda, Wauthy, Magowsky, Dabert, Spicer, Hendricks, Møller, Tryjanowski, Hromada and others). They described 17 more genera and many species from the entire world. According to Bochkov (2009b), the family includes 167 sp. of 36 genera. In his monograph on Palearctic Syringophilidae Skoracky (2011) analysed all 118 Palearctic species, describing 6 new genera, 25 new species and many new combinations. Chirov & Kravtsova (1995) described the new genus Mironovia, Kivganov in Kivganov & Sharafat (1995) – the genus Kethleyana, Philips & Norton (1978) – the genus Bubophilus. New information is credited also to Russians (Bochkov, Mironov and others, after 2000), Americans (Casto, 1973–1980, OConnor), Chinese (Liu Bai-li) and other acarologists. Now (end December 2020) the family includes more then 420 described species of 63 genera and many more (at least 5000) should be expected, as many species of birds are not examined for Syringophilidae.
The quill mites, as their name shows, are obligatory and permanent ectoparasites living inside the different types of quills. They feed on soft-tissue fluids of their hosts.
So far quill mites have been recorded on more than 440 species of birds, belonging to 109 bird families (including 60 of Passeriformes) and 28 orders (Accipitriformes, Anseriformes, Apodiformes, Bucerotiformes, Ciconiiformes, Charadriiformes, Coliiformes, Columbiformes, Coraciiformes, Cuculiformes, Falconiformes, Galbuliformes, Galliformes, Gruiformes, Musophagiformes, Otidiformes, Passeriformes, Pelecaniformes, Phoenicopteriformes, Piciformes, Procellariiformes, Psittaciformes, Pteroclidiformes, Strigiformes, Suliformes, Tinamiformes, Trogoniformes, Upupiformes), but many other birds may harbour new genera and species of these rather specific mites.
The system of Syringophilidae is accepted here as follows:
Family Syringophilidae Lavoipierre, 1953 – 63 genera, 415 sp.
Subfamily Syringophilinae Lavoipierre, 1953 – 51 gen., 320 sp.
Apodisyringiana Skoracki, 2005 – 3 sp.
Apodisyringophilus Skoracki et OConnor, 2010 – 1 sp.
Ascetomylla Kethley, 1970 – 2 sp.
Aulobia Kethley, 1970 – 12 sp.
Aulonastus Kethley, 1970 – 17 sp.
Betasyringophiloidus Skoracki, 2011 – 6 sp.
Blaszakia Skoracki et Sikora, 2008 – 2 sp.
Bochkovia Skoracki et OConnor, 2010 – 1 sp.
Bubophilus Philips et Norton, 1978 – 4 sp.
Charadriphilus Bochkov et Chistyakov, 2001 (incl. Charadriaulobia Bochkov, Fain et Skoracki, 2004) – 5 sp.
Chenophila Kethley, 1970 – 5 sp.
Colinophilus Kethley, 1973 – 1 sp.
Colisyringophilus Skoracki, Unsoeld et Ozminski, 2013 – 1 sp.
Corvitorotroglus Skoracki et Bochkov, 2010 – 1 sp.
Creagonycha Kethley, 1970 – 3 sp.
Crotophagisyringophilus Skoracki, 2008 – 1 sp.
Cuculisyringohilus Skoracki, 2008 – 2 sp.
Fritschisyringophilus Bochkov, Fain et Skoracki, 2004 – 1 sp.
Galliphilopsis Skoracki et Sikora, 2004– 5 sp.
Ixobrychiphilus Skoracki, Zmudzinski et Solarczyk, 2017 – 1 sp.
Kalamotrypetes Casto, 1980 – 2 sp.
Kethleyana Kivganov, in Kivganov & Sharafat, 1995 – 1 sp.
Krantziaulonastus Skoracki, 2011 – 5 sp.
Megasyringophilus Fain, Bochkov et Mironov, 2000 – 11 sp.
Meitingsunes Glowska et Skoracki, 2010 – 8 sp.
Mironovia Chirov et Kravtsova, 1995 – 5 sp.
Neoaulobia Fain, Bochkov et Mironov, 2000 – 8 sp.
Neoaulonastus Skoracki, 2004 (= Dissonus Skoracki, 1999, praeoccup.) – 18 sp.
Neobubophilus Skoracki et Unsoeld, 2016 – 2 sp.
Neoperisterophila Skoracki, 2005 – 1 sp.
Neosyringophilopsis Skoracki et Sikora, 2005 – 8 sp.
Niglarobia Kethley, 1970 – 7 sp.
Paraniglarobia Skoracki, 2011 – 2 sp.
Peristerophila Kethley, 1970 (= Castosyringophilus Bochkov et Perez, 2002) – 13 sp.
Phalarophilus Skoracki, Bochkov et OConnor, 2011 – 1 sp.
Philoxanthornea Kethley, 1970 – 3 sp.
Picineoaulonastus Skoracky, Klimovičová, Mucha et Hromada, 2014 – 1 sp.
Picisyringophilus Skoracki et OConnor, 2010 – 1 sp.
Procellariisyringophilus Schmidt et Skoracki, 2007 (= Syringonomus Kethley, 1970, praeoccup.) – 1 sp.
Psittaciphilus Fain, Bochkov et Mironov, 2000 – 4 sp.
Pteroclidisyringophilus Skoracki, Unsoeld et Skorupski, 2011 – 2 sp.
Selenonycha Kethley, 1970 – 3 sp.
Stibarokris Kethley, 1970 – 11 sp.
Syringophiloidus Kethley, 1970 – 48 sp.
Syringophilopsis Kethley, 1970 – 50 sp.
Syringophilus Heller, 1880 – 2 sp.
Terratosyringophilus Bochkov et Perez, 2002 – 5 sp.
Tinamiphylopsis Skoracki et Sikora, 2004 – 2 sp.
Torotrogla Kethley, 1970 – 20 sp.
Trypetoptila Kethley, 1970 – 1 sp.
Subfamily Picobiinae Johnston et Kethley, 1973 – 12 gen., 95 sp.
Calamincola Casto, 1978 [= Cuculiphilus Casto, 1977, praeocc.] – 1 sp.
Charadriineopicobia Skoracki, Spicer et OConnor, 2014 – 4 sp.
Columbiphilus Kivganov et Sharafat, 1995 – 4 sp.
Gunabopicobia Skoracki et Hromada, 2013 – 7 sp.
Lawrencipicobia Skoracki et Hromada, 2013 – 3 sp.
Neopicobia Skoracki, 2011 – 12 sp.
Picobia Haller, 1878 – 42 sp.
Phipicobia Glowska et Schmid, 2014a – 1 sp.
Pipicobia Glowska et Schmid, 2014b – 4 sp.
Pseudopicobia Scoracki, Scibek et Sikora, 2012 – 3 sp.
Rafapicobia Skoracki, 2011 – 13 sp.
Tanopicobia Skoracki, Hromada, Jerzak et Sikora, 2020 – 1 sp.
Syringophilidae Lavoipierre, 1953; Dubinin, 1957: 93
Type genus: Syringophilus Heller, 1880
Syringophilinae Lavoipierre, 1953
Type genus: Syringophilus Heller, 1880
Apodisyringiana Skoracki, 2005: 337
Type species: Apodisyringiana haszprunari Skoracki, 2005
Apodisyringiana haszprunari Skoracki, 2005: 338
Type locality: Sumatra
Distribution: Indonesia (Sumatra)
Host: Hemiprocne comata (Apodiformes: Hemiprocnidae)
Apodisyringiana mystaceae Skoracki, 2005: 341
Type locality: Papua New Guinea, New Britain Isl.
Distribution: Papua New Guinea
Hosts: Hemiprocne mystacea mystacea, H. m. aeroplanes (Apodiformes: Hemiprocnidae)
Apodisyringiana hirundapi Skoracki, Kaszewska & Kavetska, 2015: 584
Type locality: Yokohama
Distribution: Japan
Host: Hirundapus caudacutus (Apodiformes: Apodidae)
Apodisyringophilus Skoracki & OConnor, 2010: 5
Type species: Apodisyringophilus collocalius Skoracki et OConnor, 2010
Apodisyringophilus collocalius Skoracki & OConnor, 2010: 5
Type locality: Philippines, Leyte Isl., Inopacan
Distribution: Philippines
Host: Collocalia esculenta (Apodiformes: Apodidae)
Ascetomylla Kethley, 1970: 24
Type species: Ascetomylla gallinula Kethley, 1970
Ascetomylla gallinula Kethley, 1970: 25
Type locality: Johannesburg, Transvaal
Distribution: South Africa
Host: Gallinula chloropus (Gruiformes: Rallidae)
Syringophilopsis porzanae Bochkov & Galloway, 2001: 31
Ascetomylla porzanae (Bochkov et Galloway): Skoracki, Flannery & Spicer, 2008: 299
Type locality: Manitoba: Delta Marsh
Distribution: Canada
Host: Porzana carolina (Gruiformes: Rallidae)
Aulobia Kethley, 1970: 41
Type species: Syringophilus dendroicae Clark, 1964
Aulobia anthreptes Skoracki & Glowska, 2008: 184; Skoracki, Hendricks & Spicer, 2010: 40
Type locality: unknown
Distribution: Southeast Asia (distribution of the host)
Host: Anthreptes malacensis (Passeriformes: Nectariniidae)
Aulobia afroanthreptes Skoracki & Zmudzinski, 2018: 2
Type locality: Tanzania
Distribution: Tanzania
Hosts: Anthreptes longuemarei, A. neglectus (Passeriformes: Nectariniidae)
Aulobia cardueli Skoracki, Hendricks & Spicer, 2010: 36; Skoracki & Bochkov, 2010: 65; Skoracki, 2011: 104
Type locality: Texas, Johnson Co., Garen Ranch
Distribution: Germany, Kazakhstan, Poland, Russia, Slovakia, USA (Texas)
Hosts: Cardinalis cardinalis (Passeriformes: Cardinalidae), Acanthis flammea, Carduelis carduelis, C. flavirostris, Serinus citrinella, Spinus psaltria, S. spinus (Passeriformes: Fringillidae)
Aulobia cisticolae Skoracki & Sikora, 2003: 2; Skoracki et al., 2009: 132; Klimovičova et al., 2014a
Type locality: Kodegbe (Togo)
Distribution: Cameroon, DR Congo, Kenya, Rwanda, Togo
Hosts: Cisticola cantans, C. chubbi, C. erythrops, C. hunteri, C. lateralis (Passeriformes: Cisticolidae)
Syringophilus dendroicae Clark, 1964: 86
Aulobia dendroicus (Clark): Kethley, 1970: 41; Skoracki, Hendricks & Spicer, 2010: 33
Aulobia dendroicae (Clark): Skoracki, 2011: 99
Type locality: Maryland
Distribution: USA (Maryland, Florida, Georgia, California, Texas)
Hosts: Setophaga [Dendroica] nigrescens, S. coronata (Passeriformes: Parulidae)
Aulobia erythroptera Skoracki & Dabert, 2001b: 208
Type locality: Kamina (Togo)
Distribution: Togo
Host: Prinia [Heliolais] erythroptera (Passeriformes: Cisticolidae)
Aulobia leucostictus Skoracki, 2011: 110
Type locality: Europe
Distribution: Europe
Host: Leucosticte arctoa (Passeriformes: Fringillidae)
Aulobia nectariniae Skoracki & Glowska, 2008: 179
Type locality: New Guinea
Distribution: New Guinea
Hosts: Cinnyris jugularis, Leptocoma sericea (Passeriformes: Nectariniidae)
Syringophilopsis stachyris Bochkov, Mironov & Skoracki, 2001: 237
Aulobia stachyris (Bochkov, Mironov et Skoracki): Skoracki, 2011: 107
Type locality: Thirkhed-hunga
Distribution: Nepal
Host: Cyanoderma [Stachyris] pyrrhops (Passeriformes: Timaliidae)
Aulobia sylviae Bochkov & Mironov, 199[8]9: 9; Skoracki, 2011: 99
Type locality: Rybachiy village, Kurshskaya Spit, Kaliningrad Province
Distribution: Croatia, France, Jordan, Kazakhstan, Poland, Russia
Hosts: Sylvia atricapilla, S. curruca, S. nisoria (Passeriformes: Sylviidae)
Syringophilopsis sylviettae Fain, Bochkov & Mironov, 2000: 57
Syringophilopsis sylvietta: Skoracki & Dabert, 2002: 138
Aulobia sylviettae Fain, Bochkov et Mironov): Skoracky, Hendricks & Spicer, 2010: 39; Skoracki, 2011: 99 Skoracki, 2011: 99; Klimovičova et al., 2014a
Type locality: Rwanda
Distribution: Kenya, Rwanda
Host: Sylvietta whytii johnstoni (Passeriformes: Macrosphenidae)
Aulobia virens Skoracki & Dabert, 2001b: 211; Skoracki, 2011: 99
Type locality: Adina (Togo)
Distribution: Togo
Host: Sylvietta virens flaviventris (Passeriformes: Macrosphenidae)
Aulonastus Kethley, 1970: 47
Type species: Aulonastus pipili Kethley, 1970
Aulonastus albus Skoracki, 2002: 304; 2011: 136; Klimovičová & Hromada, 2014: 33
Type locality: Zachodniopomorskie, near Swinoujscie
Distribution: Poland, Slovakia
Hosts: Motacilla alba, M. flava (Passeriformes: Motacillidae)
Aulonastus anthus Skoracki, 2011: 150
Type locality: Wielkopolska, Katy
Distribution: Poland, Slovakia
Hosts: Anthus cervinus, A. trivialis (Passeriformes: Motacillidae)
Aulonastus emberizicus Skoracki, Hendricks & Spicer, 2010c: 132; Skoracki, 2011: 136
Type locality: Jaff Davis Co., Texas
Distribution: USA (California, Texas)
Hosts: Ammodramus savannarum, Passerculus sandwichensis, Zonotrichia atricapilla (Passeriformes: Emberizidae)
Aulonastus euphagus Skoracki, Hendricks & Spicer, 2010c: 134; Skoracki, 2011: 136
Type locality: Mono Co., California
Distribution: USA (California)
Host: Euphagus cyanocephalus (Passeriformes: Icteridae)
Aulonastus fringillus Skoracki, 2011: 152
Type locality: Wielkopolska, Katy
Distribution: Germany, Poland
Host: Fringilla coelebs (Passeriformes: Fringillidae)
Aulonastus henicorhina Sikora & Skoracki, 2012: 60
Type locality: Ecuador, near Quito
Distribution: Ecuador
Host: Henicorhina leucosticta (Passeriformes: Troglodytidae)
Aulonastus lanius Skoracki, 2011: 146
Type locality: Europe
Distribution: Europe
Host: Lanius senator (Passeriformes: Laniidae)
Aulonastus loxius Skoracki, 2011: 148
Type locality: Zachodniopomorskie, Darlowko Wschodnie, Bukowo – Kopan
Distribution: Poland
Host: Loxia curvirostra (Passeriformes: Fringillidae)
Aulonastus lusciniae Skoracki, 2002: 301; 2011: 136
Type locality: Slovakia
Distribution: Slovakia
Host: Luscinia luscinia (Passeriformes: Muscicapidae)
Aulonastus nectariniphilus Skoracki & Zmudzinski, 2018: 1
Type locality: Tanzania
Distribution: Tanzania
Host: Anthreptes reichenowi (Passeriformes: Nectariniidae)
Aulonastus paridus Sikora, Unsöld & Skoracki, 2016: 151
Type locality: 80 mi. S. Arusha
Distribution: Tanzania
Hosts: Melaniparus albiventris, M. fringillinus (Passeriformes: Paridae)
Aulonastus pipili Kethley, 1970: 48; Skoracki, 2011: 136
Type locality: Lake Placid, Highlands Co., Florida
Distribution: USA (Florida)
Host: Pipilo erythrophthalmus (Passeriformes: Passerellidae)
Aulonastus pirangus Skoracki, Hendricks & Spicer, 2010c: 138; Skoracki, 2011: 136
Type locality: Sierra Co., California
Distribution: USA (California)
Host: Piranga ludoviciana (Passeriformes: Cardinalidae)
Aulonastus prunellae Bochkov & Mironov, 1999: 38; Skoracki, 2011: 140
Type locality: Novgorod Prov., Chudovo Distr., Oskuy Vill.
Distribution: Russia
Host: Prunella modularis (Passeriformes: Prunellidae)
Aulonastus pteroptochos Sikora, Kaszewska & Skoracki, 2014: 422
Type locality: Chile, Valdivia
Distribution: Chile
Host: Pteroptochos tarnii (Passeriformes: Rhinocriptidae)
Aulonastus similis Broda, Dabert & Glowska, 2016: 716; Skoracki, 2017:180
Type locality: Los Tuxtlas Tropical Biological Station(UNAM), 18°350700N, 95°403100W, Veracruz
Distribution: Mexico
Hosts: Myiozetetes similis (Passeriformes: Tyrannidae), Habia fuscicauda, H. rubica (Pass., Cardinalidae)
Aulonastus sturnellus Skoracki, Hendricks & Spicer, 2010c: 141; Skoracki, 2011: 136
Type locality: Johnson Co., Texas
Distribution: USA (Texas)
Host: Sturnella magna (Passeriformes: Icteridae)
Betasyringophiloidus Skoracki, 2011: 86
Type species: Betasyringophiloidus phoenicurus Skoracki, 2011
Betasyringophiloidus emberizae Glowska, Broda & Dabert, 2019: 4
Type locality: Novolitovsk, Partizanskiy District, Primorskiy Kray
Distribution: Russia
Host: Emberiza tristrami (Passeriformes: Emberizidae)
Syringophiloidus motacillae Bochkov & Mironov, 1998: 14; Bochkov & Galloway, 2001: 2025; Skoracki, Flannery & Spicer, 2009: 72
Betasyringophiloidus motacillae (Bochkov et Mironov): Skoracki, 2011: 92; Klimovičová & Hromada, 2014: 33
Type locality: Rybachiy village, Kurshskaya Spit, Kaliningrad Province
Distribution: Canada, Poland, Russia, Slovakia, USA (California)
Host: Motacilla flava (Passeriformes: Motacillidae)
Betasyringophiloidus phoenicurus Skoracki, 2011: 87; Pidhorna et al., 2020: 215
Type locality: Zachodniopomorskie, Weltyn (Poland)
Distribution: Poland, Ukraine
Host: Phoenicurus ochruros (Passeriformes: Muscicapidae)
Betasyringophiloidus saxicolus Skoracki, 2011: 90; Pidhorna et al., 2020: 215
Type locality: Zachodniopomorskie, Weltyn (Poland)
Distribution: Poland, Ukraine
Host: Saxicola rubetra (Passeriformes: Muscicapidae)
Syringophiloidus schoeniclus Skoracki, 2002: 309
Syringophiloidus schoeniclusi: Skoracki, 2004: 296; Skoracki & Bochkov, 2010: 65 Betasyringophiloidus schoeniclus (Skoracki): Skoracki, 2011: 96; Skoracki & Mironov, 2013; Glowska, Broda & Dabert, 2019: 6
Type locality: Slovenská Kajňa (Slovakia)
Distribution: Kazakhstan, Mongolia, Poland, Russia, Slovakia
Hosts: Emberiza rutila, E. schoeniclus, E. tristrami (Passeriformes: Emberizidae)
Syringophilus seiuri Clark, 1964: 81
Syringophiloidus seiurus (Clark): Kethley, 1970: 50; Skoracki & Dabert, 2002: 138
Syringophiloidus seiuri: Bochkov & Galloway, 2001: 2025; Skoracki, 2011: 38
Betasyringophiloidus seiuri (Clark, 1964): Skoracki et al., 2016: 458; Skoracki, Spicer & OConnor, 2016: 458; Glowska, Broda & Dabert, 2019; Grossi & Proctor, 2020: 82; Skoracki, 2017: 180
Type locality: Maryland (USA)
Distribution: Canada (Manitoba), DR Congo, Mexico, Togo, USA (Maryland, Florida, S. Carolina, Massachusets, Texas), Kenya
Hosts: Geothlypis philadelphia, Helmitheros vermivorus, Oreothlypis peregrina, O. ruficapilla, Parkesia noveboracensis, Seiurus aurocapilla, Setophaga palmarum, S. pensylvanica (Passeriformes: Parulidae), Melospiza georgiana, M. lincolnii, M. melodia (Passeriformes: Passerellidae), Ploceus bagdafecht (= Othyphantes stuhlmanni), Ploceus [Pachyphantes] superciliosus, Ploceus xanthops, Quelea erythrops (Passeriformes: Ploceidae), Icterus pustulatus (Passeriformes: Icteridae)
Blaszakia Skoracki & Sikora, 2008a: 328
Type species: Galliphilopsis francolinus Skoracki et Sikora, 2004
Blaszakia rossae Skoracki & Sikora, 2008a: 330
Type locality: Katanga, Kongolo
Distribution: DR Congo
Host: Musophaga rossae (Musophagiformes: Musophagidae)
Blaszakia tauracos Skoracki & Sikora, 2008a: 328
Type locality: Uluguru (Tanzania)
Distribution: Guinea, Tanzania
Hosts: Tauraco livingstoni, T. schalowi (Musophagiformes: Musophagidae)
Bochkovia Skoracky & OConnor, 2010: 6
Type species: Bochkovia phalaropi Skoracky et OConnor, 2010
Bochkovia phalaropi Skoracky & OConnor, 2010: 8
Type locality: San Diego Co., San Diego
Distribution: USA
Host: Phalaropus fulicarius (Charadriiformes: Scolopacidae)
Bubophilus Philips & Norton, 1978: 900
Type species: Bubophilus ascalaphus Philips et Norton, 1978
Bubophilus aluconis Nattress et Skoracki, 2009: 49; Skoracki, 2011: 321; Skoracki et al., 2012: 37; Skoracki, Unsoeld, Marciniak & Sikora, 2016: 821
Type locality: England
Distribution: England, Poland, Tanzania
Hosts: Asio otus, Strix aluco, S. woodfordii (Strigiformes: Strigidae)
Bubophilus ascalaphus Philips & Norton, 1978: 900; Skoracki & Dabert, 2002: 146; Skoracki, 2011: 319; Skoracki, Unsoeld, Marciniak & Sikora, 2016: 821
Type locality: Central New York (USA)
Distribution: Cameroon, Nepal, South Africa, USA
Hosts: Bubo africanus, B ascalaphus, B. bubo, B. virginianus (Strigiformes: Strigidae), Tyto alba affinis (Strigiformes: Tytonidae)
Bubophilus asiobius Skoracki & Bochkov, 2002: 142; Skoracki, 2011: 319; Skoracki, Unsoeld, Marciniak & Sikora, 2016: 821
Type locality: Darłówko Wschodnie (Poland)
Distribution: Germany, Poland
Host: Asio otus (Strigiformes: Strigidae)
Bubophilus tytonis Skoracki & Unsold, 2002: 817
Type locality: Cameroon, Wasa
Distribution: Cameroon
Host: Tyto alba affinis (Strigiformes: Tytonidae)
Charadriphilus Bochkov & Chistyakov, 2001: 295
Type species: Charadriphilus lyudmilae Bochkov et Chistyakov, 2001
Charadriaulobia Bochkov, Fain & Skoracki, 2004: 136
Type species: Charadriaulobia vanelli Bochkov, Fain et Skoracki, 2004
Aulobia gallinago Bochkov et Mironov, 1998: 8
Charadriaulobia gallinago: Bochkov, Fain & Skoracki, Fain & Skoracki, 2004: 138
Charadriphilus gallinago (Bochkov et Mironov): Skoracki, Dabert & Schmäschke, 2006: 61; Skoracki, 2011: 259
Type locality: St. Malo (France)
Distribution: France, Poland
Host: Gallinago gallinago (Charadriiformes: Scolopacidae)
Charadriphilus lyudmilae Bochkov & Chistyakov, 2001: 297; Skoracki, 2011: 256
Type locality: Pochi Village, Tosno Distr., Leningrad Distr.
Distribution: Russia
Host: Scolopax rusticola (Charadriiformes: Scolopacidae)
Aulobia paraguaiae Skoracki & Sikora, 2002: 5
Charadriphilus paraguaiae (Skoracki et Sikora): Skoracki, 2011: 256
Type locality: El Bolson – Rio Negro
Distribution: Argentina
Host: Gallinago paraguaiae (Charadriiformes: Scolopacidae)
Charadriphilus ralli Skoracki & Bochkov, 2010: 57; Skoracki, 2011: 260; Skoracki et al., 2014: 227
Type locality: Ongtustik Qazaqstan, Vysokoe village, Vysochanskiy Pond (Kazakhstan)
Distribution: Chile, Kazakhstan, Kirghizia, Paraguay, USA
Hosts: Fulica americana, Gallinula melanops, Laterallus melanophaius, Porzana parva, Rallus aquaticus (Gruiformes: Rallidae)
Charadriaulobia vanelli Bochkov, Fain & Skoracki, 2004: 136
Charadriphilus vanelli (Bochkov, Fain et Skoracki): Skoracki, 2011: 256
Type locality: Brazil (Antverp Zoo)
Distribution: Brazil
Host: Vanellus chilensis (Charadriiformes: Charadriidae)
Chenophila Kethley, 1970: 37
Type species: Chenophila branta Kethley, 1970
Chenophila branta Kethley, 1970: 38; Skoracki, 2011: 267
Type locality: Kensington Park, Oakland Co., Michigan
Distribution: USA (Michigan)
Host: Branta canadensis (Anseriformes: Anatidae)
Chenophila kanduli Bochkov, in Bochkov & Mironov, 1999: 7; Skoracki, 2011: 267
Type locality: Rybachiy village, Kurshskaya Spit, Kaliningrad Province
Distribution: Russia, USA
Hosts: Anas clypeata, A. querquedula (Anseriformes: Anatidae)
Chenophila marmaronetta Zmudzinski & Unsoeld, 2017: 477
Type locality: Turkey, Erciş
Distribution: Turkey
Host: Marmaronetta angustirostris (Anseriformes: Anatidae)
Chenophila nanseni Skoracki & Zawierucha, 2016: 123
Type locality: Svalbard, Nissenfjella
Distribution: Svalbard
Host: Branta leucopsis (Anseriformes: Anatidae)
Chenophila platyrhynchos Tymcio, Kavetska, Czyz & Skoracki, 2013: 660
Type locality: Szczecin
Distribution: Poland
Host: Anas platyrhynchos (Anseriformes: Anatidae)
Colinophilus Kethley, 1973: 2
Type species: Colinophilus wilsoni Kethley, 1970
Colinophilus wilsoni Kethley, 1973: 2
Type locality: Tall Timbers Res. Station, Leon Co., Florida
Distribution: USA (Florida, Kentucky)
Hosts: Calipepla squamata, Colinus virginianus (Galliformes: Odontophoridae)
Colisyringophilus Skoracki, Unsoeld & Ozminski, 2013: 214
Type species: Colisyringophilus tanzanicus Skoracki, Unsoeld et Ozminski, 2013
Colisyringophilus tanzanicus Skoracki, Unsoeld & Ozminski, 2013: 214
Type locality: Tanzania, Soga, Kibaha District
Distribution: Tanzania
Hosts: Colius striatus, Urocolius macrourus (Coliiformes: Coliidae)
Corvitorotroglus Skoracki & Bochkov, 2010: 54
Type species: Corvitorotroglus alpha Skoracki et Bochkov, 2010
Corvitorotroglus alpha Skoracki & Bochkov, 2010: 54
Type locality: Ongtustik Qazaqstan, Chockpak Ornithological Station
Distribution: Kazakhstan
Host: Corvus frugilegus (Passeriformes: Corvidae)
Creagonycha Kethley, 1970: 34
Type species: Creagonycha lara Kethley, 1970
Creagonycha lara Kethley, 1970: 36; Skoracki, 2011: 271
Type locality: Coffee Bluff, Chatham Co., Georgia
Distribution: USA (Georgia)
Host: Larus delawarensis (Charadriiformes: Laridae)
Creagonycha sternae Kivganov, in Kivganov & Sharafat, 1995: 86; Skoracki, 2011: 274
Type locality: Odessa Prov., Tiligulski liman
Distribution: Ukraine
Host: Sternula albifrons (Charadriiformes: Laridae, Sterninae)
Syringophilus totani Oudemans, 1904: 171; 1906: 57
Creagonycha totana (Oudemans): Kethley, 1970: 37; Bochkov & Mironov, 1999: 10; Glowska, Chrzanowski & Kaszewska, 2015: 15
Creagonycha totani (Oudemans): Skoracki et al., 2006; Skoracki, 2011: 271
Type locality: Arcachon, France
Distribution: Canada, France, Kazakhstan, Poland, Togo
Hosts: Calidris alba, C. himantopus, C. minuta, C. temminckii, Tringa glareola, T. totanus (Charadriiformes: Scolopacidae)
Crotophagisyringophilus Skoracki, 2008a: 76
Type species: Crotosyringophilus io Skoracki, 2008
Crotophagisyringophilus io Skoracki, 2008a: 76; Hromada et al., 2016: 532; Skoracki, 2017:
Type locality: Prov. of Formosa (Argentina)
Distribution: Argentina, Bolivia, Mexico
Hosts: Crotophaga ani, C. sulcirostris (Cuculiformes: Cuculidae, Crotophaginae)
Cuculisyringophilus Skoracki, 2008b: 155
Type species: Cuculisyringophilus crotophaginus Skoracki, 2008
Niglarobia chirovi Bochkov & Mironov, 1999: 11
Cuculisyringophilus chirovi (Bochkov et Mironov): Skoracki, 2011: 324; Hromada et al., 2016: 531
Type locality: Ottuk village, Issyk-Kul Lake (Kyrghyzstan)
Distribution: Indonesia, Kyrghyzstan, Papua New Guinea, Tanzania
Hosts: Centropus goliath, Cercococcyx montanus, Cuculus canorus, C. solitarius, Pachycoccyx audeberti, Scythrops novaehollandiae (Cuculiformes: Cuculidae)
Cuculisyringophilus crotophaginus Skoracki, 2008b: 156; Hromada et al., 2016: 532
Type locality: Cambyrila (Paraguay)
Distribution: Colombia, Mexico, Paraguay
Hosts: Crotophaga sulcirostris, Guira guira (Cuculiformes: Cuculidae, Crotophaginae)
Fritschisyringophilus Bochkov, Fain & Skoracki, 2004: 139
Type species: Fritschisyringophilus lonchurae Bochkov, Fain et Skoracki, 2004
Fritschisyringophilus lonchurae Bochkov, Fain & Skoracki, 2004: 139
Type locality: India (Antwerp Zoo)
Distribution: India
Host: Lonchura punctulata (Passeriformes: Estrildidae)
Galliphilopsis Skoracki & Sikora, 2004: 380
Type species: Galliphilopsis francolinus Skoracki et Sikora, 2004
Galliphilopsis bochkovi Skoracki & Sikora, 2004: 382; Skoracki, 2011: 310
Type locality: Himalaya
Distribution: Himalaya (no other data)
Host: Tragopan sp. (Galliformes: Phasianidae). Note by Glowska, Chrzanowski & Kaszewska, 2015: 16: “The specimens recorded on Alectoris barbara and determined as G. bochkovi belong to G. szeptyckii (Skoracki & Sikora 2011)”.
Galliphilopsis colinus Skoracki & Sikora, 2011: 20
Type locality: Zanderij (Suriname)
Distribution: Colombia, Suriname
Hosts: Colinus cristatus, C. leucopogon (Galliformes: Odontophoridae)
Galliphilopsis francolinus Skoracki & Sikora, 2004: 380; Skoracki, 2011: 310
Type locality: South Africa
Distribution: South Africa
Host: Francolinus levaillantoides (Galliformes: Phasianidae)
Galliphilopsis lophurus Skoracki & Sikora, 2004: 383; Skoracki, 2011: 310
Type locality: unknown
Distribution: unknown; host distribution in the Himalaya
Host: Lophura leucomelanos (Galliformes: Phasianidae)
Galliphilopsis szeptyckii Skoracki & Sikora, 2011: 20; Glowska, Chrzanowski & Kaszewska, 2015: 16
Type locality: Adelaide (Australia)
Distribution: Australia, Indonesia, Italy (Sicily), Japan
Hosts: Alectoris barbara, Coturnix chinensis, C. coturnix, C. pectoralis (Galliformes: Phasianidae)
Ixobrychiphilus Skoracki, Zmudzinski & Solarczyk, 2017: 270
Type species: Ixobrychiphilus wallacei Skoracki, Zmudzinski et Solarczyk, 2017
Ixobrychiphilus wallacei Skoracki, Zmudzinski & Solarczyk, 2017: 270
Type locality: China, Middle Course of Yangtze River
Distribution: China
Host: Ixobrychus cinnamomeus (Pelecaniformes: Ardeidae)
Kalamotrypetes Casto, 1980: 233
Type species: Kalamotrypetes Casto, 1980
Kalamotrypetes colinastes Casto, 1980: 233
Type locality: Texas, La Salle Co.
Distribution: USA
Hosts: Callipepla squamata, Colinus virginianus (Galliformes: Odontophoridae)
Kalamotrypetes cracidus Skoracki et Sikora, 2011: 16
Type locality: Guyana
Distribution: Guyana, Paraguay
Hosts: Ortalis canicollis, Penelope sp. (Galliformes: Cracidae)
Kethleyana Kivganov, in Kivganov & Sharafat, 1995: 84
Type species: Kethleyana gelochelodoni Kivganov, 1995
Kethleyana gelochelodoni Kivganov, in Kivganov & Sharafat, 1995: 86; Skoracki, 2011: 275
Type locality: Odessa Prov., Tiligulski liman
Distribution: Ukraine
Host: Gelochelidon nilotica (Charadriiformes: Laridae)
Krantziaulonastus Skoracki, 2011: 128
Type species: Krantziaulonastus Skoracki, 2011
Aulonastus buczekae Skoracki, 2002: 300
Krantziaulonastus buczekae (Skoracki): Skoracki, 2011: 129
Type locality: Pomorskie, Kuznica-Hel
Distribution: Poland
Host: Sturnus vulgaris (Passeriformes: Sturnidae)
Krantziaulonastus dubinini Skoracki & Sikora, 2014: 197
Type locality: Russia, Tula District, Suvorov
Distribution: Russia
Host: Passer domesticus (Passeriformes: Passeridae)
Aulonastus galbulicus Skoracki, 2008: 48
Krantziaulonastus galbulicus (Skoracki): Skoracki, 2011: 128
Type locality: South America
Distribution: South America
Host: Jacamaralcyon tridactyla (Galbuliformes: Galbulidae)
Krantziaulonastus yoyomi Glowska & Skoracki, 2011: 64
Type locality: unknown
Distribution: unknown
Host: Dinemellia dinemelli (Passeriformes: Ploceidae)
Megasyringophilus Fain, Bochkov & Mironov, 2000: 35.
Type species: Megasyringophilus kethleyi Fain, Bochkov et Mironov, 2000
Megasyringophilus aquilus Skoracki, Lontkowski & Stawarczyk, 2010: 1204; Zmudzinski et al., 2015: 292
Type locality: South Africa
Distribution: Kazakhstan, France, Poland, South Africa, Spain
Hosts: Accipiter nisus, Aquila rapax, Clanga pomarina, Hieraaetus pennatus (Accipitriformes: Accipitridae)
Megasyringophilus cacatua Glowska & Laniecka, 2013: 386
Type locality: Australia, Mount Hart
Distribution: Australia
Host: Cacatua galerita (Psittaciformes: Cacatuidae)
Megasyringophilus cyanocephala Fain, Bochkov & Mironov, 2000: 40; Bochkov & Fain, 2003: 3; Skoracki, 2005: 128
Type locality: India (Zoo of Antwerp)
Distribution: India
Hosts: Psittacula cyanocephala, P. eupatria, P. krameri (Psittaciformes: Psittaculidae)
Megasyringophilus dalmas Skoracki & Unsoeld, 2016: 822
Type locality: Venezuela
Distribution: Venezuela
Host: Megascops choliba (Strigiformes: Strigidae)
Megasyringophilus dubinini Bochkov & Fain, 2003: 3
Type locality: Sulawesi
Distribution: Indonesia
Host: Trichoglossus ornatus (Psittaciformes: Psittaculidae)
Megasyringophilus eos Skoracki, 2005b: 132
Type locality: Ceram I.
Distribution: Indonesia (Ceram)
Host: Eos bornea (Psittaciformes: Psittaculidae)
Megasyringophilus geoffroyus Skoracki, 2005b: 132
Type locality: New Guinea
Distribution: New Guinea
Host: Geoffroyus geoffroyi (Psittaciformes: Psittaculidae)
Megasyringophilus kethleyi Fain, Bochkov & Mironov, 2000: 35; Bochkov & Fain, 2003
Type locality: Brazil (Zoo of Antwerp)
Distribution: Brazil
Hosts: Aratinga jandaya, A. pertinax, Brotogeris versicolurus (Psittaciformes: Psittacidae)
Megasyringophilus platycercus Bochkov & Fain, 2003: 4
Type locality: Australia
Distribution: Australia
Host: Platycercus eximius (Psittaciformes: Psittaculidae)
Megasyringophilus rhynchopsittae Bochkov & Perez, 2002: 157
Type locality: Mesa de las Guacamayas, Mpio. Janos
Distribution: Mexico
Host: Rhynchopsitta pachyrhyncha (Psittaciformes: Psittacidae)
Megasyringophilus trichoglossus Fain, Bochkov & Mironov, 2000: 37; Skoracki, 2005b: 132
Type locality: New Guinea (Zoo of Antwerp)
Distribution: Australia, Indonesia (Timor), New Guinea
Hosts: Trichoglossus chlorolepidotus, T. euteles (Psittaciformes: Psittaculidae)
Meitingsunes Glowska & Skoracki, 2010: 61
Type species: Meitingsunes aldwelles Glowska et Skoracki, 2010
Meitingsunes aldwelles Glowska & Skoracki, 2010: 61; Skoracki, 2011: 301; Kaszewska, Skoracki & Kavetska, 2016: 480
Type locality: Choco
Distribution: Colombia
Host: Geotrygon frenata (Columbiformes: Columbidae)
Meitingsunes caprimulgus Skoracki, Kaszewska, Unsoeld & Skorupski, 2015: 128
Type locality: Colombia
Distribution: Colombia
Host: Chordeiles minor (Caprimulgiformes: Caprimulgidae)
Meitingsunes chalcophaps Kaszewska, Skoracki & Kavetska, 2016: 480
Type locality: West Timor
Distribution: Indonesia
Host: Chalcophaps indica (Columbiformes: Columbidae)
Meitingsunes columbicus Skoracki, 2011: 302; Kaszewska, Skoracki & Kavetska, 2016: 480
Type locality: Wielkopolska, Poznan
Distribution: Cameroon, Kazakhstan, Poland, Russia
Hosts: Columba livia, C. oenas, C. palumbus, Treron waalia (Columbiformes: Columbidae)
Meitingsunes ptilinopus Kaszewska, Skoracki & Kavetska, 2020: 2
Type locality: Australia
Distribution: Australia and New Guinea
Hosts: Ptilinopus magnificus, P. rivoli (Columbiformes: Columbidae)
Meitingsunes turacoensis Kaszewska, Skoracki & Kavetska, 2016: 483
Type locality: Sulawesi, Manado
Distribution: Indonesia (Sulawesi, West Timor)
Hosts: Turacoena manadensis, T. modesta (Columbiformes: Columbidae)
Peristerophila tympanistria Skoracki & Dabert, 2002: 141
Meitingsunes tympanistria (Skoracki et Dabert): Glowska & Skoracki, 2010: 61; Skoracki, 2011: 301; Kaszewska, Skoracki & Kavetska, 2016: 480
Type locality: Missahohe
Distribution: Togo
Host: Turtur tympanistria (Columbiformes: Columbidae)
Syringophilus zenadurae Clark, 1964: 83
Peristerophila zenadoura (Clark): Kethley, 1970: 56; Bochkov & Mironov, 1998: 15; Skoracki & Glowska, 2008: 159
Peristerophila zenadourae (Clark): Skoracki & Sikora, 2002: 8; Skoracki & Bochkov, 2010: 66
Meitingsunes zenadourae (Clark): Glowska & Skoracki, 2010: 61; Skoracki, 2011: 301
Type locality: Maryland
Distribution: Argentina, N. Africa, Kazakhstan, USA (Maryland)
Hosts: Columba livia, Zenaida auriculata, Z. asiatica, Z. macroura (Columbiformes: Columbidae)
Mironovia Chirov & Kravtsova, 1995: 370
Type species: Mironovia phasiani Chirov et Kravtsova, 1995
Mironovia coturnae Bochkov, Fain & Skoracki, 2004: 142; Skoracki, 2011
Type locality: Belgium
Distribution: Belgium, Poland
Host: Coturnix coturnix (Galliformes: Phasianidae)
Mironovia lagopus Bochkov & Skirnisson, 2011: 715; Skoracki & Sikora, 2011: 13
Type locality: Iceland, Lake Mývatn
Distribution: Iceland, Norway
Hosts: Lagopus lagopus, L. muta (Galliformes: Phasianidae)
Kalamotrypetes pavodaptes Casto, 1980: 237
Mironovia pavodaptes (Casto): Skoracki & Sikora, 2011: 19
Type locality: Texas, Ft. Hood Reservation, Bell Co.
Distribution: USA
Host: Meleagris gallopavo [pavodaptes] (Galliformes: Phasianidae)
Mironovia phasiani Chirov & Kravtsova, 1995: 370; Skoracki, 2011: 315
Type locality: Tokmak, Kyrghyzstan
Distribution: Kyrghyzstan
Host: Phasianus colchicus (Galliformes: Phasianidae)
Mironovia rouloul Skoracki & Sikora, 2004 : 385; Skoracki, 2011
Type locality: Sumatra
Distribution: Indonesia (Sumatra)
Host: Rollulus rouloul (Galliformes: Phasianidae)
Neoaulobia Fain, Bochkov & Mironov, 2000: 40
Type species: Neoaulobia aratingae Fain, Bochkov et Mironov, 2000
Neoaulobia agapornis Fain, Bochkov & Mironov, 2000: 44; Bochkov & Fain, 2003: 151; Skoracki, 2005: 128
Neoaulobia agaporni: Skoracki & Dabert, 2002: 138
Type locality: Zambia
Distribution: Ethiopia, Namibia, Tanzania, Zambia
Hosts: Agapornis fisheri, A. nigrigenis, A. personatus, A. roseicollis, A. taranta (Psittaciformes: Psittaculidae)
Neoaulobia aratingae Fain, Bochkov & Mironov, 2000: 40; Skoracki, 2005b: 128
Type locality: Brazil (Antwerp Zoo)
Distribution: Brazil
Host: Aratinga jandaya (Psittaciformes: Psittacidae)
Neoaulobia cacatua Marciniak, Skoracki & Hromada, 2019: 192
Type locality: Australia, Victoria State, Melbourne
Distribution: Australia, Papua New Guinea
Hosts: Calyptorhynchus funereus, Probosciger aterrimus (Psittaciformes: Cacatuidae)
Neoaulobia krafti Skoracki, 2005b: 129
Type locality: Australia, New Holland
Distribution: Australia
Host: Cacatua tenuirostris (Psittaciformes: Cacatuidae)
Neoaulobia mexicana Bochkov & Perez, 2002: 149; Bochkov & Fain, 2003: 3; Skoracki, 2005b: 128
Type locality: Huacana, Piaxtla, Sinaloa
Distribution: Brazil, Mexico
Hosts: Aratinga canicularis, A. pertinax (Psittaciformes: Psittacidae)
Neoaulobia mironovi Bochkov & Perez, 2002: 146; Skoracki, 2005b: 129
Type locality: Piaxtla, Sinaloa
Distribution: Mexico
Host: Amazona finschi (Psittaciformes: Psittacidae)
Neoaulobia psittaculae Fain, Bochkov & Mironov, 2000: 44; Skoracki, 2005b: 129
Type locality: India (Zoo of Antwerp)
Distribution: India
Host: Psittacula cyanocephala (Psittaciformes: Psittaculidae)
Aulobia puylaerti Skoracki & Dabert, 1999: 523
Neoaulobia puylaerti (Skoracki et Dabert): Skoracki & Dabert, 2002: 138; Skoracki, 2005b: 129; Glowska et al., 2015: 21
Type locality: Namound joga (Togo)
Distribution: Indonesia (Java), Philippines (Luzon), Togo
Hosts: Loriculus philippensis, L. pusillus, (Psittaciformes: Psittaculidae), Poicephalus senegalus (Psittaciformes: Psittacidae)
Dissonus Skoracki, 1999a: 155 (praeoccup.)
Neoaulonastus Skoracki, 2004: 445
Type species: Dissonus scirpaceus Skoracki, 1999
Neoaulonastus aegithalos Skoracki, 2011: 122
Type locality: Opolskie, Turawa Lake
Distribution: Poland
Host: Aegithalos caudatus (Passeriformes: Aegithalidae)
Neoaulonastus apalis Klimovičova, Mikula, Kahure & Hromada, 2014: 573
Type locality: Kenya
Distribution: Kenya
Host: Apalis porphyrolaema (Passeriformes: Cisticolidae)
Syringophilus bisetatus Fritsch, 1958: 238
Dissonus scirpaceus Skoracki, 1999a: 157; Skoracki & Hebda, 2004: 5
Neoaulonastus scirpaceus (Skoracki): Skoracki, 2004: 445; Skoracki & Bochkov, 2010: 65
Neoaulonastus bisetatus (Fritsch, 1958): Skoracki, 2011: 113; Klimovičová & Hromada, 2014: 33; Pidhorna et al., 2020: 215
Type locality: Germany
Type locality of Dissonus scirpaceus Skoracki: Mierzeja Wiślana (Poland)
Distribution: Germany, Jordan, Kazakhstan, Kenya, Poland, Russia, Slovakia, Ukraine
Hosts: Acrocephalus arundinaceus, A. baeticatus, A. dumetorum, A. paludicola, A. palustris, A. scirpaceus (Passeriformes: Acrocephalidae), Sylvia atricapilla, S. communis, S. curruca (Passeriformes: Sylviidae)
Neoaulonastus caligatus Skoracki, 2011: 120
Type locality: Europe
Distribution: Europe
Host: Iduna caligata (Passeriformes: Sylviidae)
Neoaulonastus chrysocolaptes Skoracky & OConnor, 2010: 22; Skoracki, 2011: 127; Glowska et al., 2015: 21
Type locality: Luzon Is., Cagayan Prov., Penablanca Munic., betw. Bagio and Iland Rivers
Distribution: Philippines
Host: Chrysocolaptes lucidus (Piciformes: Picidae)
Neoaulonastus cinnyris Klimovičova, Smolak, Njiroge & Hromada, 2014: 256
Type locality: Tanzania, Magamba
Distribution: Tanzania
Host: Cinnyris mediocris (Passeriformes: Nectariniidae)
Neoaulonastus granatina Skoracki, Hromada & Unsoeld, 2013: 372
Type locality: Tanzania, Simangiro
Distribution: Tanzania
Host: Granatina ianthinogaster (Passeriformes: Estrildidae)
Neoaulonastus grewlingi Glowska, 2013: 20
Type locality: Australia
Distribution: Australia
Host: Falcunculus frontatus (Passeriformes: Falcunculidae)
Krantziaulonastus lonchurus Skoracki, 2011: 133
Neoaulonastus lonchurus (Skoracki): Skoracki, Hromada, Prevuznakova & Wamiti, 2019: 1806
Type locality: China
Distribution: China, Kenya, Uganda
Hosts: Estrilda astrild, E. atricapilla, E. nonnula, E. erythronotus, E. rhodopyga, E. troglodytes, Lonchura punctulata (Passeriformes: Estrildidae)
Neoaulonastus malaconotus Klimovičova, Skoracki, Njoroge & Hromada, 2016: 187
Type locality: Liberia, Grassfield (78300N; 88350W, altitude 165 m.a.s.l.).
Distribution: Kenya, Liberia, Somalia
Hosts: Laniarius aethiopicus, Nilaus afer, Telophorus nigrifrons (Passeriformes: Malaconotidae)
Krantziaulonastus oryzivorus Skoracki, 2011: 131
Neoaulonastus oryzivorus (Skoracki): Skoracki et al., 2019; Skoracki, Hromada, Prevuznakova & Wamiti, 2019: 1804
Type locality: Japan
Distribution: Japan, Kenya, Uganda
Hosts: Estrilda astrild, E. atricapilla, Lonchura oryzivora (Passeriformes: Estrildidae)
Neoaulonastus picidus Skoracki, 2011: 124
Type locality: Bardejov (Slovakia)
Distribution: Germany, Poland, Slovakia
Hosts: Dendrocopos leucotos, D. major, Picus canus (Piciformes: Picidae)
Neoaulonastus prionops Klimovičova, Skoracki & Hromada, 2016: 579
Type locality: Meru Forest, 1676 m
Distribution: Kenya
Host: Prionops sulcifrons (Passeriformes: Vangidae)
Neoaulonastus quelea Skoracki, Hromada & Unsoeld, 2013: 372
Type locality: Tanzania, Arusha, Usa-River
Distribution: Tanzania
Host: Quelea quelea (Passeriformes: Ploceidae)
Neoaulonastus remizus Skoracki, 2011: 117
Type locality: Opolskie, Turawa Lake
Distribution: Poland
Host: Remiz pendulinus (Passeriformes: Remizidae)
Neoaulonastus riparius Skoracki, 2011: 126
Type locality: Wielkopolska, Katy
Distribution: Poland
Host: Riparia riparia (Passeriformes: Hirundinidae)
Neoaulonastus sidorchukae Zmudzinski, Skoracki & Hromada, 2019: 84
Type locality: Sri Lanka
Distribution: Sri Lanka
Host: Leptocoma zeylonica (Passeriformes: Nectariniidae)
Neoaulonastus tanzanicus Skoracki, Hromada & Unsoeld, 2013: 369; Skoracki et al., 2016: 1544
Type locality: Tanzania
Distribution: Madagascar, Tanzania
Hosts: Euplectes axillaris, Ploceus nelicourvi (Passeriformes: Ploceidae)
Neoaulonastus zosterops Skoracki, Antczak & Riegert, 2009: 129; Skoracki & Mironov, 2013: 129
Type locality: Cameroon
Distribution: Cameroon, Russia (Primorskiy Krai)
Hosts: Zosterops erythropleurus, Z. senegalensis (Passeriformes: Zosteropidae)
Neobubophilus Skoracki & Unsoeld, 2016: 816
Type species: Neobubophilus atheneus Skoracki et Unsoeld, 2016
Neobubophilus atheneus Skoracki & Unsoeld, 2016: 816
Type locality: India
Distribution: India
Hosts: Athene brama, A. noctua (Strigiformes: Strigidae)
Neobubophilus cunicularius Skoracki & Unsoeld, 2016: 817
Type locality: San Pedro Dept., New Germania Distr.
Distribution: Paraguay
Host: Athene cunicularia (Strigiformes: Strigidae)
Neoperisterophila Skoracki, 2005c: 155
Type species: Neoperisterophila regiusi Skoracki, 2005
Neoperisterophila regiusi Skoracki, 2005c: 156
Type locality: New Guinea (no more data, presumed to come from Papua New Guinea)
Distribution: New Guinea
Host: Cicinnurus regius (Passeriformes: Paradisaeidae)
Neosyringophilopsis Skoracki & Sikora, 2005: 22
Type species: Neosyringophilopsis acanthizus Skoracki et Sikora, 2005
Neosyringophilopsis acanthizus Skoracki & Sikora, 2005: 23; Skoracki, 2011: 206
Type locality: Australia, New Holland
Distribution: Australia
Host: Acanthiza inornata (Passeriformes: Acanthizidae)
Syringophilopsis aegythali Bochkov, Mironov & Skoracki, 2001: 235; Skoracki, 2004c: 57; Skoracki & Hebda, 2004: 5
Neosyringophilopsis aegythali Skoracki & Sikora, 2005: 26
Neosyringophilopsis aegithali: Nattress & Skoracki, 2009: 53; Skoracki, 2011: 215; Klimovičová & Hromada, 2014: 34
Type locality: Oskuy Village, Chudovo Distr., Novgorod Prov., Russia
Distribution: England, Poland, Russia, Slovakia
Host: Aegithalos caudatus (Passeriformes: Aegithalidae)
Syringophilopsis garrulaxi Bochkov, Mironov & Skoracki, 2001: 236
Neosyringophilopsis garrulaxi (Bochkov, Mironov & Skoracki): Skoracki & Sikora, 2005: 26; Skoracki, 2011: 206
Type locality: Kolopani, S. Mustang
Distribution: Nepal
Host: Trochalopteron affine [Garrulax affinis] (Passeriformes: Leiotrichidae)
Syringophilopsis locustellus Skoracki, 2004: 51
Neosyringophilopsis locustellus: Skoracki & Sikora, 2005: 26; Skoracki, 2011: 218
Type locality: Zachodniopomorskie, Swinoujstie
Distribution: Poland
Hosts: Locustella luscinioides, L. naevia (Passeriformes: Locustellidae)
Neosyringophilopsis lonchurae Skoracki, 2008: 45 (also as “lonchurus”)
Neosyringophilopsis lonchurus: Skoracki, 2011: 206; Skoracki et al., 2019: 1806
Type locality: Africa
Distribution: Kenya, Uganda
Hosts: Estrilda astrild, E. atricapilla, E. nonnula, E. erythronotus, E. rhodopyga, E. troglodytes, Spermestes cucullata [Lonchurus cucullatus] (Passeriformes: Estrildidae)
Neosyringophilopsis lybidus Skoracki, Klimovičová, Muchai et Hromada, 2014: 180
Type locality: Kenya, Ewaso Nigro River, W.Naro Moru
Distribution: Kenya
Host: Pogoniulus bilineatus (Piciformes: Lybiidae)
Syringophilopsis phylloscopi Bochkov, Mironov & Skoracki, 2001: 234; Skoracki, 2004c: 57
Neosyringophilopsis phylloscopi: Skoracki & Sikora, 2005: 26
Type locality: St. Malo (France)
Distribution: France, Poland, Slovakia
Hosts: Phylloscopus collybita, Ph. trochilus (Passeriformes: Phylloscopidae)
Syringophilus troglodytis Fritsch, 1958: 238; Kethley, 1970: 65; Skoracki, 1999: 184 Syringophilopsis troglodytis: Bochkov & Mironov, 1998: 5; Bochkov & Galloway, 2001: 2023; Skoracki, 2004c: 57
Neosyringophilopsis troglodytis Skoracki & Sikora, 2005: 26; Nattress & Skoracki, 2007: 143; Skoracki, 2011: 213; Skoracki, Spicer & OConnor, 2016: 460
Type locality: Germany
Distribution: Canada, England, Germany, Poland
Hosts: Troglodytes aedon, T. troglodytes (Passeriformes: Troglodytidae)
Niglarobia Kethley, 1970: 43
Type species: Niglarobia ereuneti Kethley, 1970
Niglarobia cursoriae Skoracki, Dabert & Schmäschke, 2006: 55; Skoracki, 2011: 288
Type locality: Ahlenhorst Distr.
Distribution: Namibia
Host: Cursorius temminckii (Charadriiformes: Glareolidae)
Niglarobia ereuneti Kethley, 1970: 44; Skoracki, 2011: 288
Type locality: Sapelo Island, Georgia
Distribution: USA (Georgia)
Host: Calidris [Ereunetes] pusilla (Charadriiformes: Scolopacidae)
Syringophilus helleri Oudemans, 1904: 190; 1906: 62; Dubinin, 1957: 94; Fritsch, 1958: 229
Niglarobia helleri (Oudemans): Kethley, 1970: 47; Bochkov & Mironov, 1998: 12; Skoracki & Sikora, 2002: 7; Skoracki, Dabert & Schmäschke, 2006: 56; Skoracki, 2011: 288
Type locality: France
Distribution: Argentina, France, Poland, USA
Hosts: Arenaria interpres, Tringa flavipes, T. ochropus, T. totanus, T. stagnatilis (Charadriiformes: Scolopacidae)
Niglarobia lapponica Zmudzinski & Unsoeld, 2019: 367
Type locality: Germany
Distribution: Germany
Host: Limosa lapponica (Charadriiformes: Scolopacidae)
Niglarobia rhinoptili Fain, Bochkov & Mironov, 2000: 61; Skoracki & Dabert, 2002: 138; Skoracki, 2011: 288
Type locality: Bloemhof
Distribution: South Africa
Host: Smutsornis [Rhinoptilus] africanus (Charadriiformes: Glareolidae)
Syringophilus trouessarti Oudemans, 1904: 190; 1906: 67; Dubinin, 1957: 94
Niglarobia trouessarti (Oudemans): Kethley, 1970: 47; Skoracki, 2011: 288
Type locality: South America (“Amérique chaude”)
Distribution: South America
Host: Aramus guarauma [scolopaceus] (Gruiformes: Aramidae)
Niglarobia vanelli Zmudzinski & Skoracki, 2018: 704
Type locality: Germany
Distribution: Germany, Russia, Tanzania
Hosts: Vanellus albiceps, V. coconatus, V. crassirostris, V. gregarius, V. lugubris, V. senegallus, V. spinosus, V. vanellus (Charadriiformes: Charadriidae)
Paraniglarobia Skoracki, 2011: 290
Type species: Paraniglarobia Skoracki, 2011
Niglarobia calidridis Bochkov & Mironov, 1999: 12
Paraniglarobia calidridis (Bochkov et Mironov): Skoracki, 2011: 291
Type locality: Sary-Su river, Kzyl Orda Prov.
Distribution: Kazakhstan
Host: Calidris minuta (Charadriiformes: Scolopacidae)
Niglarobia skorackii Bochkov & Galloway, 2004: 40
Paraniglarobia skorackii (Bochkov et Galloway): Skoracki, 2011: 291
Type locality: Manitoba: Whitewater Lake
Distribution: Canada
Host: Tringa melanoleuca (Charadriiformes: Scolopacidae)
Peristerophila Kethley, 1970: 53
Type species: Syringophilus columbae Hirst, 1920
Castosyringophilus Bochkov & Perez, 2002: 151 (syn. by Skoracki, Hromada, Kaszewska & Sikora, 2020)
Type species: Peristerophila mucuya Casto, 1980
Peristerophila accipitridicus Skoracki, Lontkowski & Stawarczyk, 2010: 1207; Skoracki & Bochkov, 2010: 66; Skoracki, 2011: 297; Zmudzinski et al., 2015: 292; Skoracki et al., 2016: 1545
Type locality: bird imported from Africa in Wroclaw Zoo
Distribution: Africa, France, Germany, Kazakhstan, Madagascar
Hosts: Accipiter francesiae, A. nisus, Buteo lagopus, Circaetus gallicus, Terathopius ecaudatus (Accipitriformes: Accipitridae)
Castosyringophilus claravis Skoracki & Glowska, 2008b: 155
Peristerophila claravis (Skoracki et Glowska): Skoracki et al., 2020: 1803; Kaszewska et al., 2020: 358
Type locality: Sierra de Chiquitos
Distribution: Bolivia, Colombia, Ethiopia, Panama, Paraguay, Sudan, Tanzania
Hosts: Claravis pretiosa, Oenas capensis (Columbiformes: Columbidae)
Syringophilus columbae Hirst, 1920: 121; 1922: fig. 41; Dubinin, 1957: 94; Brown, 1971: 1380
Peristerophila columba (Hirst): Kethley, 1970: 56; Casto, 1976; Bochkov & Mironov, 1998: 14
Peristerophila columbae (Hirst): Nattress & Skoracki, 2009: 53; Skoracki, 2011: 294; Kaszewska et al., 2020: 355
Type locality: Tennessee (USA)
Distribution: Angola, Argentina, Canada, China, D.R. Congo, England, Germany, Greece, Hungary, India, Iran, Japan, Jordan, Kenya, Nepal, N. Macedonia, Panama, Poland, Portugal, South Africa, Suriname, Tanzania, Turkey, USA (Tennessee, Kansas, etc.)
Hosts: Columba arguatrix, C. guinea, C. leuconota, C. livia, C. oenas, C. palumbus, C. trocaz, Geotrygon chiriquensis, Patagioenas speciosa, Streptopelia capicola, S. decaocto, S. decipiens, S. orientalis, S. semitorquata, S. transquebarica, S. turtur, Zenaida auriculata (Columbiformes: Columbidae); Buteo jamaicensis (Accipitriiformes: Accipitridae)
Peristerophila coraciidus Skoracki, Hromada & Sikora, 2020: 175
Type locality: Papua New Guinea
Distribution: Papua New Guinea
Host: Eurystomus orientalis (Passeriformes: Coraciidae)
Peristerophila falcophila Skoracki, Hromada, Kaszewska, Kavetska & Unsoeld, 2018: 744
Type locality: Germany
Distribution: Germany, Kenya, Nepal, Romania, Colombia
Hosts: Falco cherrug, F. peregrinus, F. sparverius (Falconiformes: Falconidae)
Castosyringophilus forpi Bochkov & Perez, 2002: 152; Skoracki, 2005: 128
Peristerophila forpi (Bochkov et Perez): Skoracki et al., 2020: 1803
Type locality: Ruiz, Nayarit
Distribution: Mexico
Host: Forpus cyanopygius (Psittaciformes: Psittacidae)
Peristerophila geopelis Kaszewska, Skoracki, Kosicki & Hromada, 2020: 353
Type locality: Java, Trinil
Distribution: Indonesia (Java, Sumatra, Sulawesi), Australia
Hosts: Geopelia cuneata, G. placida, G. striata, Ocyphaps lophotes
(Columbiformes: Columbidae)
Peristerophila lature Kaszewska, Kavetska & Skoracki, 2014: 296; Kaszewska et al., 2020: 358
Type locality: Sulawesi (Indonesia)
Distribution: Indonesia (Sulawesi, Java, Sumatra), Mariana Is., Papua New Guinea
Hosts: Ducula luctuosa, D. spilorrhoa, Ptilinopus jambu, P. melanospilus, P. porphyreus, P. regina (Columbiformes: Columbidae: Treroninae)
Peristerophila leucomela Kaszewska, Skoracki, Kosicki & Hromada, 2020: 355
Type locality: Australia
Distribution: Australia
Host: Columba leucomela (Columbiformes: Columbidae)
Castosyringophilus meropis Skoracki, Hromada & Sikora, 2017: 24
Peristerophila meropis (Skoracki, Hromada et Sikora): Skoracki, Hromada, Kaszewska & Sikora, 2020: 1803
Type locality: Sevilla (Spain)
Distribution: Bosnia & Herzegovina, France (Corsica), Gibraltar, Greece, Italy (Sardinia), Kenya, N. Macedonia, Morocco, Pakistan, Romania (N. Dobroudja), Russia, Tanzania, Turkey
Hosts: Merops apiaster (Coraciiformes: Meropidae), Todiramphus chloris (Cor.: Alcedinidae)
Peristerophila mucuya Casto, 1980: 1; Skoracki, Hromada, Kaszewska & Sikora, 2020: 1803; Kaszewska et al., 2020: 359
Castosyringophilus mucuya (Casto): Bochkov & Perez, 2002: 6; Skoracki, 2005b: 128; Skoracki & Glowska, 2008b: 158
Type locality: Texas, La Sella
Distribution: Argentina, Australia, Brazil, Colombia, Indonesia (Sumbava Is.), Jordan, Monaco, Paraguay, Peru, Suriname, Trinidad & Tobago, USA (Texas)
Hosts: Bolborhynchus aymara, Brotogeris versicolurus (Psittaciformes: Psittacidae), Trichoglossus haematodus (Psittaciformes: Psittaculidae), Columbina minuta, C. passerina, C. squammata, C. talpacoti, Geophaps plumifera, Metropelia ceciliae, M. melanoptera, Scardafella squamata, Streptopelia decaocto, Petrophassa plumifera (Columbiformes: Columbidae)
Peristerophila nestoriae Marciniak, Skoracki & Hromada, 2019: 2
Type locality: New Zealand
Distribution: New Zealand
Host: Nestor meridionalis (Psittaciformes: Strigopidae)
Peristerophila upupi Klimovičova, Mikula, Kahure & Hromada, 2014: 575
Type locality: Nyahururu
Distribution: Kenya
Host: Upupa epops (Upupiformes: Upupidae)
Phalarophilus Skoracki, Bochkov & OConnor, 2011: 325
Type species: Phalarophilus fulicarius Skoracki, Bochkov et OConnor, 2011
Phalarophilus fulicarius Skoracki, Bochkov & OConnor, 2011: 328
Type locality: San Diego Co., San Diego
Distribution: USA
Host: Phalaropus fulicarius (Charadriiformes: Scolopacidae)
Philoxanthornia Kethley, 1970: 50
Philoxanthornea (Zool. Record, misspelling)
Type species: Philoxanthornia anoa Kethley, 1970
Philoxanthornia anoa Kethley, 1970: 53; Skoracki, 2011: 282
Type locality: Sand Island
Distribution: Midway Atoll (USA)
Host: Anous tenuirostris (Charadriiformes: Laridae, Sterninae)
Philoxanthornea clarki Kivganov, in Kivganov & Sharafat, 1995: 87; Skoracki, 2011: 282
Type locality: Odessa Prov., Tiligulski liman
Distribution: Russia, Ukraine
Hosts: Sterna hirundo, Sternula albifrons (Charadriiformes: Laridae, Sterninae), Larus canus (Charadriiformes: Laridae)
Philoxanthornea dubinini Bochkov & Mironov, 1999: 14; Skoracki, 2011: 285
Type locality: Sorbulak Lake, near Alma-Ata
Distribution: Egypt, Kazakhstan
Hosts: Chlidonias leucopterus, Gelochelidon nilotica (Charadriiformes: Laridae)
Picineoaulonastus Skoracky, Klimovičová, Mucha & Hromada, 2014: 180
Type species: Picineoaulonastus pogoniulus Skoracky, Klimovičová, Mucha et Hromada, 2014
Picineoaulonastus pogoniulus Skoracky, Klimovičová, Muchai & Hromada, 2014: 180
Type locality: Kenya
Distribution: Ethiopia, Kenya, Tanzania
Hosts: Pogoniulus bilineatus, P. pusillus (Piciformes: Lybiidae)
Picisyringophilus Skoracky & OConnor, 2010: 12
Type species: Picisyringophilus kratos Skoracky et OConnor, 2010
Picisyringophilus kratos Skoracky & OConnor, 2010: 15
Type locality: Michigan, Washtenaw Co., 2mi NW Dexter
Distribution: USA (Michigan)
Host: Picoides pubescens (Piciformes: Picidae)
Syringonomus Kethley, 1970: 56 (praeoccup.)
Procellariisyringophilus Schmid & Skoracki, 2007: 545
Type species: Syringonomus bulwerius Kethley, 1970
Syringonomus bulwerius Kethley, 1970: 59
Procellariisyringophilus bulwerius (Kethley): Schmid & Skoracki, 2007: 545
Type locality: Nihoa Island, Hawaiian Is.
Distribution: Greece, Indonesia, USA (Hawaiian Is.)
Hosts: Ardenna pacifica, Bulwerius bulwerius, Calonectris diomedea (Procellariiformes: Procellariidae)
Psittaciphilus Fain, Bochkov & Mironov, 2000: 44
Type species: Psittaciphilus amazonae Fain, Bochkov et Mironov, 2000
Psittaciphilus amazonae Fain, Bochkov & Mironov, 2000: 47; Bochkov & Fain, 2003; Skoracki, 2005b: 128
Type locality: Colombia
Distribution: Brazil, Colombia
Hosts: Amazona aestiva, A. amazonica (Psittaciformes: Psittacidae)
Psittaciphilus fritschi Fain, Bochkov & Mironov, 2000: 47; Skoracki, 2005b: 128
Type locality: Zoo of Antwerp
Distribution: ?
Host: parrot (Psittaciformes)
Psittaciphilus montanus Kaszewska & Skoracki, 2018: 1
Type locality: Trinidad I., Aripo Mt.
Distribution: Brazil, Panama, Trinidad & Tobago
Host: Geotrygon montanus (Columbiiformes: Columbidae)
Psittaciphilus patagioenas Kaszewska & Skoracki, 2018: 3
Type locality: Near San Juan River, Chocó Department Lama Hermosa
Distribution: Colombia
Hosts: Patagioenas fasciata, P. speciosa (Columbiiformes: Columbidae)
Pteroclidisyringophilus Skoracki, 2011: 31
Type species: Charadriiphilus re Skoracky et OConnor, 2010
Pteroclidisyringophilus otididus Skoracky, Unsoeld & Skorupski, 2014: 349
Type locality: South Tunisia
Distribution: Tunisia
Host: Chalamydotis undulata (Otidiformes: Otididae)
Charadriiphilus re Skoracky & OConnor, 2010: 18
Pteroclidisyringophilus re (Skoracky et OConnor): Skoracki, 2011: 264
Type locality: Red Sea Governate, Wadi el Metuli
Distribution: Egypt
Host: Pterocles senegallus (Pterocliformes: Pteroclidae)
Selenonycha Kethley, 1970: 31
Type species: Selenonycha baltoda Kethley, 1970
Selenonycha baltoda Kethley, 1970: 34; Bochkov & Mironov, 1998: 11
Type locality: Cameron Parish, Louisiana
Distribution: USA
Hosts: Charadrius placidus, Ch. w. wilsonia, Chlidonius niger (Charadriiformes: Charadriidae)
Selenonycha charadriiformicus Skoracki, 2011: 276
Type locality: Lake Manzala area (Egypt)
Distribution: Egypt, Germany, Poland, Russia
Hosts: Chroicocephalus genei, Ch. ridibundus (Charadriiformes: Laridae), Charadrius placidus (Charadriiformes: Charadriidae), Philomachus pugnax (Charadriiformes: Scolopacidae)
Selenonycha insperata Zmudzinski & Skoracki, 2017: 594
Type locality: Misiones Prov., Laguna San Jose de Posadas
Distribution: Argentina
Host: Phimosus infuscatus (Pelecaniformes: Threskiornithidae)
Stibarokris Kethley, 1970: 62
Type species: Stibarokris phalacrus Kethley, 1970
Ciconichenophilus Skoracky & OConnor, 2010: 12
Type species: Ciconichenophilus phoeniconaias Skoracky et OConnor, 2010
Stibarokris annae Glowska, 2015a: 74
Type locality: Uruguay
Distribution: Uruguay
Host: Ciconia maguari (Ciconiiformes: Ciconiidae)
Stibarokris brevisetosus Skoracki & Zmudzinski [in Skoracki et al., 2016], 2016: 1319
Type locality: Turkey, Ekmekci
Distribution: Turkey
Host: Plegadis falcinellus (Pelecaniformes: Threskiornithidae)
Stibarokris dastychi Glowska & Skoracky, 2011: 64
Type locality: Germany
Distribution: Germany
Host: Phalacrocorax carbo (Suliformes: Phalacrocoracidae)
Stibarokris geronticus Skoracki & Zmudzinski [in Skoracki et al., 2016], 2016: 1313
Type locality: South Africa
Distribution: South Africa
Host: Geronticus calurus (Pelecaniformes: Threskiornithidae)
Stibarokris hydrobates Zmudzinski, Solarczyk & Skoradski, 2016: 432
Type locality: Malta, Filfla
Distribution: Malta
Host: Hydrobates pelagicus (Procellariiformes: Hydrobatidae)
Stibarokris langei Bochkov & Mironov, 1999: 35; Glowska & Skoracky, 2011: 68
Type locality: Novgorod Prov., Chudovo Distr., Oskuy Vill.
Distribution: Russia
Host: Ciconia alba (Ciconiiformes: Ciconiidae)
Stibarokris mariasi Glowska, 2015b: 616
Type locality: Johnston Atoll
Distribution: Johnston Atoll
Host: Puffinus pacificus (Procellariiformes: Procellariidae)
Stibarokris phalacrus Kethley, 1970: 62; Glowska & Skoracky, 2011: 67
Type locality: Pigeon Key, Monroe Co., Florida
Distribution: USA (Florida)
Host: Phalacrocorax auritus (Suliformes: Phalacrocoracidae)
Ciconichenophilus phoeniconaias Skoracky & OConnor, 2010: 13
Stibarokris phoeniconaias Skoracky & OConnor, 2010: 25; Glowska & Skoracky, 2011: 66; Zmudzinski, Solarczyk & Skoradski, 2016: 434; Skoracki, Zmudzinski, Unsoeld & Sikora, 2016: 9
Type locality: Botswana
Distribution: Botswana, Turkey, South America
Hosts: Phoeniconaias minor, Phoenicopterus ruber, Ph. chilensis (Phoenicopteriformes: Phoenicopteridae)
Stibarokris plataleus Skoracki & Zmudzinski, [in Skoracki et al., 2016]. 2016: 1316
Type locality: Neusiedl See
Distribution: Austria
Host: Platalea leucorodia (Pelecaniformes: Threskiornithidae)
Stibarokris theristicus Skoracki, Zmudzinski & Unsoeld [in Skoracki et al., 2016]. 2016: 1313
Type locality: Marajo Island, Pacoval
Distribution: Brazil
Host: Theristicus caudatus (Pelecaniformes: Threskiornithidae)
Syringophiloidus Kethley, 1970: 48
Type species: Syringophilus minor Berlese, 1887
Syringophiloidus agelaius Bochkov, Skoracki, Hendricks & Spicer, 2011: 209
Type locality: Cibola National Wildlife Refuge (Arizona)
Distribution: USA (Arizona)
Host: Agelaius phoeniceus (Passeriformes: Icteridae)
Syringophiloidus amazilia Skoracki, 2017: 181
Тype locality: Veracrux, Los Tuxlas
Distribution: Mexico
Host: Amazilia candida (Passeriformes: Trochilidae)
Syringophiloidus apus Skoracki, Kaszewska & Kavetska, 2015: 584
Type locality: Chile
Distribution: Chile
Host: Apus melba (Apodiformes: Apodidae)
Syringophiloidus artamus Skoracki, 2004: 288; 2011: 36
Type locality: Sumatra
Distribution: Indonesia (Sumatra)
Hosts: Artamus fuscus, A. leucorhynchus (Passeriformes: Artamidae)
Syringophiloidus auduboni Skoracki, Spicer & OConnor, 2016: 462
Type locality: California, Riverside Co.
Distribution: USA
Host: Spizella breweri (Passeriformes: Passerellidae)
Syringophiloidus bombycillae Skoracki, 2002: 305; 2004: 296; 2011: 36; Skoracki, Spicer & OConnor, 2016: 468
Type locality: Bardejov (Slovakia)
Distribution: Poland, Slovakia, USA
Hosts: Bombycilla cedrorum, B. garrulus (Passeriformes: Bombycillidae)
Syringophiloidus bucerotidus Skoracki, 2017: 39
Тype locality: D.R.Congo
Distribution: Cameroon, D.R. Congo, Tanzania
Hosts: Horizocerus hartlaubi, H. albocristatus, Lophoceros pallidirostris, Tockus flavirostris, T. fasciatus, T. deckeni (Bucerotiformes: Bucerotidae)
Syringophiloidus carolae Skoracki et al., 2009: 66; Skoracki, 2011; Skoracki, Spicer & OConnor, 2016: 464
Тype locality: Texas, Hood Co., Jones Ranch
Distribution: USA (Texas)
Hosts: Cardinalis cardinalis, Passerina caerulea (Passeriformes: Cardinalidae), Melanerpes formicivorus (Piciformes: Picidae)
Syringophiloidus carpodaci Bochkov & Apanaskevich, 2001: 323; Skoracki, 2011: 50
Type locality: Southwestern Altai, Zyryanovskii Distr., Shumovsk Village (Kazakhstan)
Distribution: Kazakhstan, Poland, Russia
Host: Carpodacus erythrinus (Passeriformes: Fringillidae)
Syringophiloidus cincli Skoracki, 2017: 184
Тype locality: Puebla, Scapa, 3 Mi NE Huauchinango
Distribution: Mexico
Host: Cinclus mexicanus (Passeriformes: Cinclidae)
Syringophiloidus coccothraustes Skoracki, 2011: 82
Тype locality: Wielkopolskie, Katy near Poznan
Distribution: Germany, Poland
Host: Coccothraustes coccothraustes (Passeriformes: Fringillidae)
Syringophiloidus cypsiuri Fain, Bochkov & Mironov, 2000: 51; Skoracki, 2011: 36
Type locality: Kasai Prov., DR Congo
Distribution: DR Congo
Host: Cypsiurus parvus (Apodiformes: Apodidae)
Syringophiloidus daberti Bochkov, Fain & Skoracki, 2004: 144; Bochkov, Skoracki, Hendricks & Spicer, 2011: 210; Skoracki et al., 2011; Skoracki, Spicer & OConnor, 2016: 465
Type locality: Mexico (Anwerp Zoo)
Distribution: Mexico, USA (Texas)
Host: Passerina ciris (Passeriformes: Cardinalidae)
Syringophiloidus delichonum Bochkov, 2001: 151; Skoracki, 2004: 297; 2011: 69; Skoracki, Spicer & OConnor, 2016: 465
Type locality: Kaliningradskaya Oblast, Kurshskaya Kosa, Ribachiy Vill. (Russia)
Distribution: Poland, Russia, USA
Hosts: Delichon urbicum, Hirundo rustica (Passeriformes: Hirundinidae)
Syringophiloidus dendrocitta Fain, Bochkov & Mironov, 2000: 55; Skoracki, 2011: 36
Type locality: East Asia (Zoo of Antwerp)
Distribution: East Asia
Hosts: Dendrocitta vagabunda (= rufa rufa) (Passeriformes: Corvidae, Sturnus vulgaris (Passeriformes: Sturnidae), [Cyanocitta stelleri (Passeriformes: Corvidae)]
Syringophiloidus erythrurus Skoracki, 2004: 293
Type locality: Tenerife
Distribution: Spain (Tenerife)
Hosts: Erythrura trichroa, Lonchura sp. (Passeriformes: Estrildidae)
Syringophiloidus estrildus Skoracki, Hromada, Prevuznakova & Wamiti, 2019: 1802
Type locality: Masindi
Distribution: Uganda
Host: Estrilda rhodopyga (Passeriformes: Estrildidae)
Syringophiloidus Skoracki & Schmidt, 2014: 61
Type locality: Ogooue Maritime Prov., Moukalaba-Doudou National Park, NW corner of N’dogo Lagoon
Distribution: Gabon
Host: Terpsiphone rufiventer (Passeriformes: Monarchidae)
Syringophilus minor glandarii Fritsch, 1958: 235
Syringophiloidus glandarii (Fritsch): Bochkov & Mironov, 1998: 14; Skoracki, 2011: 57; Skoracki, Spicer & OConnor, 2016: 465
Type locality: Germany
Distribution: Germany, Japan, Kazakhstan, Russia, USA
Hosts: Corvus brachyrhynchos, C. frugilegus, C. monedula, Cyanocitta stelleri, Garrulus glandarius, Pica pica (Passeriformes: Corvidae). For other hosts see the notes in Glowska, Chrzanowski & Kaszewska, 2015: 32.
Syringophiloidus graculae Fain, Bochkov & Mironov, 2000: 51
Type locality: SE Asia
Distribution: SE Asia
Host: Gracula religiosa (Passeriformes: Sturnidae)
Syringophiloidus hirundinis Skoracki, Skoracki, Moller & Tryjanowski, 2003: 17; Pidhorna, 2020: 216
Type locality: Milano (Italy)
Distribution: Denmark, Italy, Poland, Spain, Ukraine
Host: Hirundo rustica (Passeriformes: Hirundinidae)
Syringophiloidus jackowiaki Bochkov, Skoracki, Hendricks & Spicer, 2011: 208
Type locality: Texas, Trinity Co., Davy Crocket Nat. Forest
Distribution: USA (Texas)
Host: Poecile carolinensis (Passeriformes: Paridae)
Syringophiloidus klimovi Skoracki & Bochkov, 2010: 61
Type locality: Ongtustik, Qazaqstan
Distribution: Germany, Kazakhstan
Host: Chloris chloris (Passeriformes: Fringillidae)
Syringophiloidus microcerculus Sikora & Skoracki, 2012: 58
Type locality: Para, Peixe-Boi
Distribution: Brazil
Host: Microcerculus marginatus (Passeriformes: Troglodytidae)
Syringophilus bipectinatus minor Berlese, 1887: 10
Syringophilus minor minor (Berlese): Fritsch, 1958: 235
Syringophiloidus minor (Berlese): Kethley, 1970; Bochkov & Mironov, 1998; Bochkov & Galloway, 2001; Skoracki & Dabert, 2002; Glowska et al., 2007; Skoracki, 2011: 39; Skoracki, Spicer & OConnor, 2016: 467
Type locality: Europe
Distribution: Bosnia and Herzegovina, Canada, China, Congo, Croatia, England, Germany, Jordan, Kirghizia, Poland, Russia, Slovakia, USA
Hosts: Passer domesticus, P. hispaniolensis, P. montanus (Passeriformes: Passeridae), Serinus capistratus (Passeriformes: Fringillidae)
Syringophiloidus molothrus Skoracki, Flannery & Spicer, 2009: 66; Skoracki, Spicer & OConnor, 2016: 466
Type locality: Texas, Hood Co.
Distribution: USA (Texas, Michigan)
Host: Icterus galbula, Molothrus ater (Passeriformes: Icteridae)
Syringophiloidus montanus Skoracki, 2002: 310
Type locality: Bardejov
Distribution: Slovakia
Host: Passer montanus (Passeriformes: Passeriformes)
Syringophiloidus nectariniae Skoracki & Zmudzinski, 2018: 1
Type locality: Tanzania
Distribution: Tanzania
Host: Hedydipna collaris (Passeriformes: Nectariniidae)
Syringophiloidus nkaii Klimovičova, Skoracki, Njoroge & Hromada, 2016: 188
Type locality: Kenya, Archers Post (altitude 1,200 m.a.s.l.).
Distribution: Kenya
Host: Nilaus afer (Passeriformes: Malaconotidae)
Syringophiloidus oriolus Skoracki, 2004: 283
Type locality: Western Europe
Distribution: China, Western Europe
Hosts: Oriolus chinensis, O. oriolus (Passeriformes: Oriolidae)
Syringophiloidus parapresentalis Skoracki, 2011: 63; Skoracki, Spicer & OConnor, 2016: 466; Skoracki, Mironov, Hernandes et Valim, 2016: 5
Type locality: Wielkopolskie, Jeziory, near Poznan
Distribution: Brazil, Germany, Jordan, Kazakhstan, Poland, Russia, Slovakia, USA (California)
Hosts: Turdus atrogularis, T. iliacus, T. merula, T. migratorius, T. pilaris, T. rufiventris (Passeriformes: Turdidae)
Syringophiloidus petronicus Skoracki, 2011: 45
Type locality: Italy
Distribution: Italy
Host: Petronia petronia (Passeriformes: Passeridae)
Syringophiloidus philomelosus Skoracki, 2011: 43
Type locality: Al-Dhleil near Amman
Distribution: Jordan
Host: Turdus philomelos (Passeriformes: Turdidae)
Syringophiloidus picidus Skoracki, Klimičova, Muchai & Hromada, 2014: 184
Type locality: Mutanda (Kenya)
Distribution: Germany, Kenya, Tanzania, Uganda
Host: Dendropicus fuscescens (Piciformes: Picidae)
Syringophiloidus plocei Glowska, Broda & Dabert, 2016: 563
Type locality: Ogooue Maritime Province, Gamba Complex of Protected Areas, near
mouth of Nyanga River
Distribution: Gabon
Hosts: Ploceus aurantius, P. cucullatus, P. nigerrimus (Passeriformes: Ploceidae)
Syringophiloidus presentalis Chirov & Kravtsova, 1995: 376; Skoracki, 2011: 60; Skoracki, Spicer & OConnor, 2016: 466
Type locality: Bishkek (Kirghizia)
Distribution: France, Kirghizia, Poland, Slovakia, USA
Host: Sturnus vulgaris (Passeriformes: Sturnidae)
Syringophiloidus pseudonigritae Glowska, Dragun-Damian & Dabert, 2012b: 65
Type locality: Tanzania
Distribution: Tanzania
Host: Pseudonigrita arnaudi (Passeriformes: Ploceidae)
Syringophiloidus quetzali Skoracki, Mironov & Unsoeld, 2013: 292
Type locality: Guatemala
Distribution: Ecuador, Guatemala
Hosts: Pharomachrus antisianus, Ph. mocinno (Trogoniformes: Trogonidae)
Syringophiloidus serini Bochkov, Fain & Skoracki, 2004: 144; Skoracki, 2011: 38
Type locality: Central Africa (Antwerp Zoo)
Distribution: Central Africa
Host: Serinus mozambicus (Passeriformes: Fringillidae)
Syringophiloidus sialius Skoracki, Flannery & Spicer, 2009: 67; Bochkov, Skoracki, Hendricks & Spicer, 2011: 211; Skoracki, 2011: 38
Type locality: Mendocino Co., Hopland
Distribution: USA (California)
Hosts: Sialia mexicana (Passeriformes: Turdidae), Spizella breweri (Passeriformes: Passerellidae)
Syringophiloidus sporophila Skoracki, 2017: 185
Тype locality: Veracruz, Los Tuxlas
Distribution: Mexico
Host: Sporophila torqueola (Passeriformes: Thraupidae)
Syringophiloidus stawarczyki Skoracki, 2004: 291; 2011: 38; Skoracki, Mironov, Hernands & Valim, 2016: 5
Type locality: Brazil
Distribution: Brazil
Hosts: Euphonia cyanocephala (Passeriformes: Emberizidae), Dacnis cayana, Tachyphonus rufus (Pass., Thraupidae)
Syringophiloidus tarnii Skoracki & Sikora, 2002: 2 (and tarni, p.1); Skoracki, 2011: 38
Type locality: El Bolson, Rio Negro (Argentina)
Distribution: Argentina
Host: Pteroptochos tarnii (Passeriformes: Rhinocryptidae)
Syringophiloidus teledromas Sikora, Kaszewska & Skoracki, 2014: 420
Type locality: Argentina – RioNegro, Cipolletti, Neuquén
Distribution: Argentina
Host: Teledromas fuscus (Passeriformes: Rhinocryptidae)
Syringophiloidus thryothorus Skoracki, Flannery & Spicer, 2009: 69; Skoracki, 2011:
Type locality: Texas, Hood Co., Jones Ranch
Distribution: USA (Texas)
Host: Thryothorus ludovicianus (Passeriformes: Troglodytidae)
Syringophiloidus weiszii Skoracki, Hromada & Tryjanowski, 2001: 30; Skoracki, 2011: 79
Type locality: Bardejov
Distribution: Germany, Slovakia, USA
Hosts: Lanius excubitor, L. ludovicianus (Passeriformes: Laniidae)
Syringophiloidus xanthocephalus Bochkov, Skoracki, Hendricks & Spicer, 2011: 208
Type locality: Cibola National Wildlife Refuge (Arizona)
Distribution: USA (Arizona)
Host: Xantocephalus xanthocephalus (Passeriformes: Icteridae)
Syringophiloidus motacillae of Bochkov & Galloway (2001: 2025) and Skoracki et al. (2009: 72) (incorrect determinations)
Syringophiloidus zonotrichia Bochkov, Skoracki, Hendricks & Spicer, 2011: 203
Type locality: Garen Ranch, Johnson Co. Texas,
Distribution: Canada, USA (Texas)
Host: Zonotrichia albicolis (Passeriformes: Emberizidae)
Host: Campylorhynchus brunneicapillum (Paseriformes: Troglodytidae)
Syringophilopsis Kethley, 1970: 18
Type species: Syringophilus elongatus Ewing, 1911
Syringophilopsis acrocephali Skoracki, 1999a: 160; 2011: 179; Klimovičová & Hromada, 2014: 34; Pidhorna, 2020: 216
Type locality: Mierzeja Wiślana (Poland)
Distribution: Egypt, Poland, Russia, Slovakia, Ukraine
Hosts: Acrocephalus palustris, A. schoenobaenus, A. scirpaceus (Passeriformes: Acrocephalidae)
Syringophilopsis albicollisi Skoracki & Dabert, 2000: 59; Skoracki, 2011: 155
Type locality: Odjolo
Distribution: Togo
Host: Merops albicollis (Coraciiformes: Meropidae)
Syringophilopsis bartrami Skoracki, Spicer & OConnor, 2016: 477
Type locality: Michigan, Wayne Co., Plymouth
Distribution: USA (Michigan)
Host: Spizella passerina (Passeriformes: Passerellidae)
Syringophilopsis blaszaki Skoracki & Dabert, 1999: 89; Skoracki, 2011: 195; Klimovičová & Hromada, 2014: 34; Pidhorna, 2020: 216
Type locality: 6 km NW from Darlowko Wschodnie (Poland)
Distribution: Poland, Russia, Slovakia, Ukraine
Hosts: Anthus pratensis, P. trivialis (Passeriformes: Motacillidae)
Syringophilopsis bochkovi Skoracki, Mironov & Bermúdez, 2019: 231
Type locality: Panama, Darien Prov., Parque Nacional Darien, Rancho Frio
Distribution: Panama
Hosts: Corapipo altera, Manacus vitellinus (Passeriformes: Pipridae)
Syringophilopsis bonariensis Skoracki et al., 2016: 4
Type locality: Minas Gerais, Nova Lima, Agua Limpa, 20°13’05.07”S 43°57’00.2”W
Distribution: Brazil
Host: Molothrus bonariensis (Passeriformes: Icteridae)
Syringophilopsis borini Bochkov & Mironov, 1999: 43; Skoracki, 2004; 2011: 201
Type locality: Novgorod Prov., Chudovo Distr., Oskuy Vill.
Distribution: Poland, Russia, Turkey
Host: Sylvia borin (Passeriformes: Sylviidae)
Syringophilopsis catesbyi Skoracki, Spicer & OConnor, 2016: 470; Skoracki, 2017: 186
Type locality: Florida, Leon Co.
Distribution: Mexico, USA (Florida)
Hosts: Vireo hypochryseus, V. olivaceus (Passeriformes: Vireonidae)
Syringophilopsis certhiae Skoracki, Hendricks & Spicer, 2011: 6
Type locality: California, Monterey
Distribution: USA
Host: Certhia americana (Passeriformes: Certhiidae)
Syringophilopsis corvinae Skoracki & Sykora, 2003: 5; Skoracki, 2011: 155
Type locality: Galim
Distribution: Cameroon
Host: Corvinella corvina (Passeriformes: Laniidae)
Syringophilopsis davidi Glowska & Laniecka, 2012: 385
Type locality: Hurghada
Distribution: Egypt
Host: Calandrella brachydactyla (Passeriformes: Alaudidae)
Syringophilopsis dendroicae Bochkov & Galloway, 2001: 2023; Skoracki, Flannery & Spicer, 2008: 299; Skoracki, 2011: 155; Skoracki, Spicer & OConnor, 2016: 479
Type locality: Winnipeg, Manitoba, Canada
Distribution: Canada, USA
Hosts: Protonotaria citrea, Setophaga [Dendroica] coronata, S. caerulescens, S. petechia, S. [Dendroica] graciae, Cardellina [Wilsonia] pusilla (Passeriformes: Parulidae)
Syringophilopsis dicruri Skoracki, Hromada & Wamiti, 2011: 35
Type locality: Tsavo West National Park, Komboyo Campside
Distribution: Kenya
Host: Dicrurus adsimilis (Passeriformes: Dicruridae)
Syringophilus elongatus Ewing, 1911: 39
Syringophilus icteridae Clark, 1964: 89 – Laurel, Maryland, from Agelaius phoenicius, Euphagus carolinus and Quiscalus quiscula (syn. by Kethley, 1970)
Syringophilopsis elongatus (Ewing): Kethley, 1970: 21; Bochkov & Galloway, 2001: 2023; Skoracki, Flannery & Spicer, 2008: 299; Skoracki, 2011: 155; Skoracki, Hendricks & Spicer, 2011: 19; Skoracki, Spicer & OConnor, 2016: 473
Type locality: Winnipeg, Manitoba
Distribution: Canada, USA (California, Maryland, Georgia, Nebraska, Arizona)
Hosts: Agelaius phoeniceus, Euphagus carolinus, E. cyanocephalus, Quiscalus mexicanus, Q. quiscula (Passeriformes: Icteridae)
Syringophilopsis emberizae Fain, Bochkov & Mironov, 2000: 57; Skoracki, 2011: 156
Type locality: Rwanda (Zoo of Antwerp)
Distribution: Rwanda
Host: Sicalis [Emberiza] luteola (Passeriformes: Thraupidae)
Syringophilopsis empidonax Skoracki, Flannery & Spicer, 2008: 294; Skoracki, 2011: 156
Type locality: Texas, Johnson Co., Garen Ranch
Distribution: USA (Texas)
Hosts: Empidonax hammondii, E. wrightii (Passeriformes: Tyrannidae)
Syringophilopsis faini Bochkov & Apanaskevich, 2001: 321
Type locality: South – western Altai, Zyryanovskii Distr., Shumovsk Village
Distribution: Kazakhstan
Host: Sylvia curruca (Passeriformes: Sylviidae)
Syringophilus fringillae Fritsch, 1958: 230; Skoracki & Bochkov, 2010: 65; Skoracki, 2011: 185; Klimovičová & Hromada, 2014; Pidhorna, 2020: 217
Syringophilopsis fringilla (Fritsch): Kethley, 1970: 21; Bochkov & Mironov, 1998: 5; Skoracki & Dabert, 1999: 89; 2000: 59; Nattress & Skoracki, 2009; Skoracki, 2004c: 54; Klimovičová & Hromada, 2014: 35
Type locality: Germany
Distribution: England, Germany, Kazakhstan, [Morocco], Poland, Russia, Slovakia, Ukraine
Hosts: [Carduelis carduelis], Fringilla coelebs (Passeriformes: Fringillidae), [Passer montanus (Passeridae), Hirundo rustica (Hirundinidae)]. According to Skoracki (2011: 185), S. fringillae is monoxenic parasite of Fringilla coelebs. The records from other birds concern different other species. Zmudzinski & Unsoeld (2019) have recorded it from Fringilla montifringilla.
Syringophilopsis garrulus Skoracki & Dabert, 2002: 137; Skoracki, 2011: 163
Type locality: Africa
Distribution: Africa, Algeria, Germany
Host: Garrulus glandarius (Passeriformes: Corvidae)
Syringophilopsis hirundus Skoracki, 2004c: 60; Nattress & Skoracki, 2007: 143; Skoracki & Bochkov, 2010: 66; Skoracki, 2011: 193
Type locality: Świnoujscie (Poland)
Distribution: England, Kazakhstan, Poland
Host: Hirundo rustica (Passeriformes: Hirundinidae)
Syringophilopsis hunanensis Liu Bai-li, 1988: 277; Skoracki, 2011: 201 (sp. inquirenda
Type locality: Hunan
Distribution: China (Hunan)
Host: Passer domesticus (Passeriformes: Passeridae)
Syringophilopsis icteri Bochkov & Galloway, 2001: 2019; Skoracki, 2011: 156; 2017: 186; Skoracki, Hendricks & Spicer, 2011: 17; Skoracki, Spicer & OConnor, 2016: 473
Type locality: Winnipeg, Manitoba
Distribution: Canada, Mexico, USA
Hosts: Icterus bullocki, I. galbula, I. pustulatus, I. spurius (Passeriformes: Icteridae)
Syringophilopsis idunae Skoracki, 2011: 174
Type locality: Europe
Distribution: Europe
Host: Iduna caligata (Passeriformes: Acrocephalidae)
Syringophilopsis kazmierski Skoracki, 2004c: 46; Skoracki, 2011: 166; Klimovičová & Hromada, 2014: 35; Pidhorna, 2020: 217
Type locality: Darlówko Wsch. – Kopán (Poland)
Distribution: Germany, Jordan, Poland, Romania, Slovakia, Ukraine
Hosts: Ficedula hypoleuca, F. parva, Phoenicurus phoenicurus (Passeriformes: Muscicapidae)
Syringophilopsis kirgizorum Bochkov, Mironov & Kravtsova, 2000: 354; Skoracki, 2004: 51; Glowska, Skoracki & Khourly, 2007: 67; Nattress & Skoracki, 2007: 143; Skoracki, Antczak, Riegert, Fainová & Mikeš, 2009: 128; Skoracki, 2011: 182
Type locality: Bishkek City (Kyrghyzstan)
Distribution: Cameroon, England, Germany, Jordan, Kyrghyzstan, Poland, Russia
Hosts: Carduelis canabina, C. carduelis, Chloris chloris, Linurgus olivaceus, Rhodospiza obsoleta (Passeriformes: Fringillidae)
Syringophilopsis kristini Skoracki, Tryjanowski & Hromada, 2002: 11; Skoracki, 2011: 198
Type locality: Kurov
Distribution: Slovakia
Host: Lanius minor (Passeriformes: Laniidae)
Syringophilopsis lagonostictus Skoracki & Dabert, 2002: 138; Skoracki, 2011: 157
Type locality: 6 km S Mango
Distribution: Togo
Hosts: Lagonosticta rufopicta, L. senegala (Passeriformes: Estrildidae)
Syringophilopsis melittophagi Skoracki & Dabert, 2001b: 108; Skoracki, 2011: 157
Syringophilopsis melittophagii Skoracki & Dabert, 2002: 138
Type locality: 6 km S Mango
Distribution: Togo
Host: Merops (= Melittophagus) bullocki (Coraciiformes: Meropidae)
Syringophilopsis mimidus Sikora, Fajfer & Skoracki, 2012: 30; Skoracki, Spicer & OConnor, 2016: 473
Type locality: Central America
Distribution: Central America, USA (Pennsylvania, Florida)
Host: Dumetella carolinensis, Margarops fuscatus (Passeriformes: Mimidae)
Syringophilopsis muscicapus Skoracki, 2011: 161
Type locality: Japan
Distribution: Japan
Host: Muscicapa dauurica (Passeriformes: Muscicapidae)
Syringophilopsis nitens Skoracki & Dabert, 2001b: 107; Skoracki & Sikora, 2003: 9; Skoracki et al., 2011
Type locality: Atakpame (Togo)
Distribution: Kenya, Togo
Hosts: Hyphanturgus nigricollis brachypterus, Malimbus nitens, Ploceus nigricollis brachypterus, P. vitellinus, Quelea erythrops (Passeriformes: Ploceidae)
Syringophilopsis nucifragus Skoracki, 2011: 188
Type locality: Europe
Distribution: Europe
Host: Nucifraga caryocatactes (Passeriformes: Corvidae)
Syringophilopsis pari Skoracki & Mironov, 2013: 557
Type locality: Primorsky Krai
Distribution: Russia (Far East)
Host: Poecile palustris (Passeriformes: Paridae)
Syringophilopsis passericus Skoracki, 2011: 176
Type locality: Bishkek (Kirghizstan)
Distribution: Kirghizstan
Host: Passer domesticus (Passeriformes: Passeridae)
Syringophilus passerinae Clark, 1964: 91; Skoracki, Flannery & Spicer, 2008: 292; Skoracki, 2011: 157; Skoracki, Hendricks & Spicer, 2011: 17; Skoracki, Spicer & OConnor, 2016: 474
Syringophilopsis passerina (Clark): Kethley, 1970: 23; Bochkov & Mironov, 1999: 7
Type locality: Beltsville, Maryland
Distribution: USA (Maryland, Florida, Texas)
Hosts: Passerina ciris, P. cyanea (Passeriformes: Cardinalidae)
Syringophilopsis philemonis Glowska & Laniecka, 2013: 386
Type locality: Australia, Beagle Bay
Distribution: Australia
Host: Philemon citreogularis (Passeriformes: Meliphagidae)
Syringophilopsis polioptilus Skoracki, Flannery & Spicer, 2008: 292; Skoracki, 2011: 157
Type locality: Texas, Johnson Co., Garen Ranch
Distribution: USA (Texas)
Host: Polioptila caerulea (Passeriformes: Polioptilidae)
Syringophilopsis rusticus Skoracki, 2004: 49; Pidhorna, 2020: 218
Type locality: Olszyn (Poland)
Distribution: Poland, Ukraine
Hosts: Hirundo rustica, Riparia riparia (Passeriformes: Hirundinidae)
Syringophilopsis sialiae Skoracki, Flannery & Spicer, 2008: 298; Skoracki, 2011: 157
Type locality: Texas, Jeff Davis Co., Fort Davis
Distribution: USA (Texas)
Host: Sialia mexicana (Passeriformes: Turdidae)
Syringophilopsis sittae Skoracki, Hendricks & Spicer, 2011: 9; Skoracki & Mironov, 2013:
Type locality: USA (California), Mendocino Co., UC Hopland
Distribution: Russia, USA
Hosts: Sitta carolinensis, S. europaea (Passeriformes: Sittidae)
Syringophilopsis spinolettus Skoracki, 2004c: 57; Skoracki & Bochkov, 2010: 66; Skoracki, 2011: 190
Type locality: Darlówko-Wsch. – Kopań (Poland)
Distribution: Kazakhstan, Poland, Slovakia
Host: Anthus spinoletta (Passeriformes: Motacillidae)
Syringophilopsis sturnellus Skoracki, Hendricks & Spicer, 2011: 13
Type locality: Arizona, Mohave Co.
Distribution: USA (Arizona)
Host: Sturnella neglecta (Passeriformes: Icteridae)
Syringophilopsis sturni Chirov & Kravtsova, 1995: 374; Bochkov & Mironov, 1998: 5; Skoracki, 2004: 51; 2011: 171
Type locality: Telek, Chuy Valley, Kyrghyzstan
Distribution: Kazakhstan, Kyrghyzstan, Poland, Crimea
Host: Sturnus vulgaris (Passeriformes: Sturnidae)
Syringophilopsis szeffleri Glowska, 2014: 142
Type locality: Peru, El Caucho Biol. Station
Distribution: Peru
Host: Lophotriccus pileatus (Passeriformes: Tyrannidae)
Syringophilopsis trogoni Skoracki, Mironov et Unsoeld, 2013: 295
Type locality: Jalisco, Chamela Biol. Station of UNAM (Mexico)
Distribution: Costa Rica, Mexico
Hosts: Trogon citreolus, T. melanocephalus (Trogoniformes: Trogonidae)
Syringophilus turdi Fritsch, 1958: 233
Syringophilopsis turdi (Fritsch): Bochkov & Mironov, 1998: 6; Skoracki, 2011: 158; Skoracki, Spicer & OConnor, 2016: 480
Syringophilopsis turdus: Kethley, 1970: 24; Skoracki & Sikora, 2002: 7; Skoracki, 2004: 51; Skoracki, Flannery & Spicer, 2008: 298; Skoracki & Bochkov, 2010: 65;
Type locality: Germany
Syringophilus hylocichlae Clark, 1964: 87
Syringophilopsis hylocichlae (Clark): Kethley, 1970: 21; Skoracki, 2011: 156 (syn. by Skoracki, Spicer & OConnor, 2016: 480)
Syringophilopsis hylocichla (Clark): Kethley, 1970: 21; Skoracki & Dabert, 2000: 59; Glowska et al., 2015: 37
Type locality: Beltsville, Maryland, USA
Distribution: Argentina, Germany, Kazakhstan, Poland, Russia, Slovakia, USA (California)
Hosts: Catharus (= Hylocichla) fuscescens (sub S. hylocichlae), Turdus albicollis, T. atrogularis, T. iliacus, T. migratorius, T. philomelos, T. pilaris (Passeriformes: Turdidae)
Syringophilopsis tyranni Bochkov & Galloway, 2004: 35; Skoracki, Flannery & Spicer, 2008: 298; Skoracki, 2011: 161; Sikora, Kaszewska & Skoracki, 2014; Skoracki, Spicer & OConnor, 2016: 474; Skoracki, 2017: 188
Type locality: Winnipeg, Manitoba
Distribution: Brazil, Canada, Mexico, USA (California, Texas, Florida), Peru
Hosts: Myiarchus cinerascens, M. crinitus, Myiopagis subplacens, Satrapa icterophrys, Tyrannus tyrannus, T. dominicensis, T. forficatus, T. verticalis (Passeriformes: Tyrannidae)
Syringophilopsis veselovsky Skoracki, Antczak & Riegert, 2009: 125; Skoracki, 2011: 157
Type locality: Cameroon
Distribution: Cameroon, Kenya
Hosts: Chlorocichla flaviventris, Eurillas latirostris, Pycnonotus barbatus (Passeriformes: Pycnonotidae)
Syringophilopsis wilsoni Skoracki, Spicer & OConnor, 2016: 474
Type locality: California, Mono Co.
Distribution: USA (California)
Host: Pheucticus melanocephalus (Passeriformes: Cardinalidae)
Syringophilopsis yosefi Skoracki, Tryjanowski & Hromada, 2002: 14; Skoracki, 2011: 157; Glowska, Chrzanowski & Kaszewska, 2015: 41
Type locality: Yagoua
Distribution: Cameroon
Host: Lanius sp. (Passeriformes: Laniidae)
Syringophilus Heller, 1880: 186
Type species: Syringophilus bipectinatus Heller, 1880
Syringophilus bipectinatus Heller, 1880: 186; Oudemans, 1906: 47–57; Banks, 1907: 597; Baker & Wharton, 1952: 229 – 231; Lavoipierre, 1953: 7; Dubinin, 1957: 94; Fritsch, 1958: 229 – 230; Clark, 1964: 77; Jones, 1968: 28; Kethley, 1970: 18; Gritsenko, 1973: 515; Bochkov & Mironov, 1998: 5; Skoracki & Sikora, 2003: 9; 2004: 385; Glowska, Chrzanowski & Kaszewska, 2015: 41
Type locality: Germany
Distribution: Australia, Belarus, Cameroon, China, Germany, France (Corsica), Kyrghyzstan, Poland, Russia, South Africa (Zululand), Ukraine, USA (Kansas, Colorado)
Hosts: Alectoris rufa, Gallus gallus domesticus, (Galliformes: Phasianidae), Ptilopachus petrosus (Odontophoridae), [Corvus brachyrhynchos (Passeriformes: Corvidae)]
Syringophilus numidae Bochkov, 2000: 103; Skoracki, 2011: 309
Type locality: Transvaal
Distribution: South Africa
Host: Numida meleagris (Galliformes: Numididae)
Terratosyringophilus Bochkov & Perez, 2002: 154
Type species: Peristerophila longisoma Casto, 1979
Terratosyringophilus geotrygonus Skoracki & Glowska, 2008b: 152
Type locality: Cumbre de Valencia
Distribution: Venezuela
Host: Geotrygon linearis (Columbiformes: Columbidae)
Peristerophila longisoma Casto, 1979: 225
Terratosyringophilus longisoma (Casto): Bochkov & Fain, 2003: 154; Glowska, Chrzanowski & Kaszewska, 2015: 42
Type locality: Texas, Presidio Co.
Distribution: USA (Texas)
Hosts: Zenaida asiatica, Z. macroura (Columbiformes: Columbidae)
Terratosyringophilus loricinus Bochkov & Fain, 2003: 6
Type locality: Halmahera Is.
Distribution: Indonesia
Hosts: Lorius garrulus, Trichoglossus haematodus (Psittaciformes: Psittaculidae)
Terratosyringophilus pioni Bochkov & Perez, 2002: 155
Type locality: Near Cd. del Maiz, S.L.P.
Distribution: Mexico
Host: Pionus senilis (Psittaciformes: Psittacidae)
Terratosyringophilus reichholfi Skoracki & Sikora, 2008b: 196; Glowska, Chrzanowski & Kaszewska, 2015: 42
Type locality: Astrolabe
Distribution: Papua New Guinea
Host: Lorius lory (Psittaciformes: Psittaculidae)
Tinamiphylopsis Skoracki & Sikora, 2004a: 348
Type species: Tinamiphylopsis elegans Skoracki et Sikora, 2004
Tinamiphylopsis ariconte Skoracki, Sikora & Ozminski, 2012: 110; Glowska, Chrzanowski & Kaszewska, 2015: 42
Type locality: Paraguay
Distribution: Brazil, Paraguay
Hosts: Nothura boraquira, N. minor, Rhynchotus rufescens (Tinamiformes: Tinamidae)
Tinamiphylopsis elegans Skoracki & Sikora, 2004a: 350
Type locality: Camarones
Distribution: Argentina
Host: Eudromia elegans (Tinamiformes: Tinamidae)
Torotrogla Kethley, 1970: 28
Type species: Torotrogla mima Kethley, 1970
Torotrogla aphelocoma Bochkov, Flannery & Spicer, 2009: 184; Skoracki, 2011: 227
Type locality: Jeff Davis Co., Fort Davis
Distribution: USA (Texas)
Host: Aphelocoma californica (Passeriformes: Corvidae)
Torotrogla calcarius Skoracki, 2004d: 310; 2011: 247
Type locality: Lapland, Finland
Distribution: Finland
Host: Calcarius lapponicus (Passeriformes: Emberizidae)
Torotrogla cardinalis Bochkov, Flannery & Spicer, 2009: 189; Skoracki, 2011: 227
Type locality: Texas, Hood Co., Jones Ranch
Distribution: USA
Host: Cardinalis cardinalis (Passeriformes: Cardinalidae)
Torotrogla cardueli Bochkov & Mironov, 1999: 41; Skoracki, 2004d: 314; 2011: 242; Glowska, Chrzanowski & Kaszewska, 2015: 43
Type locality: Novgorod Prov., Chudovo Distr., Oskuy Vill. (Russia)
Distribution: Finland, Germany, Poland, Russia, Slovakia
Hosts: Linaria cannabina, Carduelis carduelis, Loxia curvirostra, L. leucoptera, L. pytyopsittacus, Serinus canaria, Spinus spinus (Passeriformes: Fringillidae)
Torotrogla coccothraustes Bochkov, Flannery & Spicer, 2009: 191; Skoracki, 2011: 228
Type locality: California, Madera Co., Sierra National Forest
Distribution: USA (California)
Host: Coccothraustes vespertinus (Passeriformes: Fringillidae)
Torotrogla cyanocitta Bochkov, Flannery & Spicer, 2009: 187; Skoracki, 2011: 227
Type locality: Lake Co., Lake Pillsbury; paratypes: Sierra Co., Bear Trap Meadow
Distribution: USA (California)
Host: Cyanocitta stelleri (Passeriformes: Corvidae)
Torotrogla emberizae Glowska, Laniecka & Romanowska, 2018: 791
Type locality: Japan
Distribution: Japan
Host: Emberiza fucata (Passeriformes: Emberizidae)
Torotrogla gaudi Bochkov & Mironov, 1999: 10; Skoracki & Skoracka, 1999: 185; Skoracki, 2004d: 313; 2011: 239
Type locality: Rybachiy village, Kurshskaya Spit, Kaliningrad Province
Distribution: Poland, Russia, Slovakia
Hosts: Fringilla coelebs, F. montifringilla, Pyrrhula pyrrhula (Passeriformes: Fringillidae)
Torotrogla lullulae Skoracki, Hromada & Kuczynski, 2001: 87; Skoracki, 2011: 244
Type locality: Zlate (Slovakia)
Distribution: Russia, Slovakia
Hosts: Eremophila alpestris, Lullula arborea (Passeriformes: Alaudidae)
Torotrogla lusciniae Skoracki, 2004d: 310; 2011: 236
Type locality: Italy
Distribution: Italy
Hosts: Luscinia megarhynchos, L. svecica (Passeriformes: Muscicapidae)
Torotrogla merulae Skoracki, Dabert & Ehrnsberger, 2000: 192; Skoracki, 2004d: 313; Glowska, Skoracki & Khourly, 2007: 67; Skoracki, 2011: 228; Skoracki, Spicer & OConnor, 2016: 483
Type locality: Kuznica – Hel (Poland)
Distribution: Germany, Jordan, Poland, Romania, Russia, Slovakia, USA
Hosts: Turdus merula, T. migratorius, T. philomelos, T. torquatus, T. viscivorus (Passeriformes: Turdidae)
Torotrogla mima Kethley, 1970: 31; Bochkov, Flannery & Spicer, 2009: 184; Skoracki, 2011: 228; Sikora et al., 2011; Skoracki, Fajfer, Kavetska & Sikora, 2012
Type locality: Odum, Wayne Co., Georgia (USA)
Distribution: Argentina, USA (Georgia)
Hosts: Mimus patagonicus, M. polyglottos, M. triurus (Passeriformes: Mimidae)
Torotrogla modularis Nattress & Skoracki, 2007: 140; Skoracki, 2011: 234; Klimovičová & Hromada, 2014: 35
Type locality: West Yorkshire (England)
Distribution: England, Mongolia, Poland, Slovakia
Hosts: Anthus pratensis, Prunella modularis (Passeriformes: Prunellidae)
Torotrogla paenae Glowska, Laniecka, Schmidt & Romanowska, 2018: 865
Type locality: Omaheke, Aminuis
Distribution: Namibia
Host: Cercotrichas paena (Passeriformes: Muscicapidae)
Torotrogla piranga Bochkov, Flannery & Spicer, 2009: 192; Skoracki, 2011: 228
Type locality: California, Sierra Co., Bear Trap Meadow
Distribution: USA (California)
Host: Piranga ludoviciana (Passeriformes: Cardinalidae)
Torotrogla pycnonotus Skoracki, 2004d: 304; 2011: 250
Type locality: Al-Dhleil near Amman
Distribution: Jordan
Host: Pycnonotus xanthopygos (Passeriformes: Pycnonotidae)
Torotrogla rubeculi Skoracki, 2004d: 304; 2011: 231; Klimovičová & Hromada, 2014: 35
Type locality: Zachodniopomorskie, Darlowko Wschodnie, Bukowo Kopań (Poland)
Distribution: Poland, Russia, Slovakia
Host: Erithacus rubecula (Passeriformes: Muscicapidae)
Torotrogla synallaxis Skoracki, Mironov, Hernandes & Valim, 2016: 252
Type locality: Minas Gerais, Nova Lima, APP do Condominio, Miguelso
Distribution: Brazil
Host: Synallaxis spixi (Passeriformes: Furnariidae)
Picobia villosa Hancock, 1895: 382; 1898: 886
Syringophilus villosus (Hancock): Oudemans, 1906: 72
Torotrogla villosa (Hancock): Kethley, 1970: 30; Bochkov, Flannery & Spicer, 2009: 184; Skoracki, 2011: 228
Type locality: Arizona
Distribution: USA (Arizona)
Host: Phainopepla nitens (Passeriformes: Ptilogonatidae)
Torotrogla volgini Skoracki & Mironov, 2013: 560
Type locality: Primorskiy Krai, Partizanskiy Distr., Novolitovsk
Distribution: Russia
Host: Emberiza elegans (Passeriformes: Emberizidae)
Trypetoptila Kethley, 1970: 25
Type species: Trypetoptila casmerodia Kethley, 1970
Trypetoptila casmerodia Kethley, 1970: 28
Type locality: McIntosh Co., Georgia
Distribution: USA (Georgia)
Host: Ardea alba egretta (Pelecaniformes: Ardeidae)
Picobiinae Johnston & Kethley, 1973: 529
Type genus: Picobia Haller, 1878
Lobatinae Casto, 1977: 169
Cuculiphilinae [as nom. nov.](praeoccupied)
Type genus: Cuculiphilus Casto, 1977
Cuculiphilus Casto, 1977: 170 (praeocc. from Uchida, 1926)
Calamincola Casto, 1978: 158
Type species: Cuculiphilus lobatus Casto, 1977
Cuculiphilus lobatus Casto, 1977: 170
Calamincola lobatus (Casto): Casto, 1978: 158; Hromada et al., 2016: 532; Skoracki, 2017: 188
Type locality: South Texas
Distribution: Colombia, Mexico, USA (Texas)
Host: Crotophaga sulcirostris (Cuculiformes: Cuculidae)
Charadriineopicobia Skoracki, Spicer & OConnor, 2014: 102
Type species: Charadriineopicobia calidris Skoracki, Spicer et OConnor, 2014
Charadriineopicobia apricaria Skoracki, Sikora & Spicer, 2016: 12
Type locality: St. Malo
Distribution: France
Host: Pluvialis apricaria (Charadriiformes: Charadriidae)
Charadriineopicobia calidris Skoracki, Spicer & OConnor, 2014: 103; Skoracki, Sikora & Spicer, 2016: 18
Type locality: Texas, Calhoun Co., Matagorda Island
Distribution: USA
Host: Calidris alba (Charadriiformes: Scolopacidae)
Picobia leucophaeus Skoracki, Hendericks & Spicer, 2010: 730
Charadriineopicobia leucophaeus (Skoracki, Hendericks et Spicer): Skoracki, Spicer & OConnor, 2014: 104
Type locality: Texas, Calhoun Co.
Distribution: USA
Host: Leucophaeus atricilla (Charadriiformes: Laridae)
Charadriineopicobia ristowi Zmudzinski & Unsoeld, 2019: 373
Type locality: Germany
Distribution: Germany
Host: Limosa lapponica (Charadriiformes: Scolopacidae)
Columbiphilus Kivganov & Sharafat, 1995: 82
Type species: Columbiphilus khushakhani Kivganov et Sharafat, 1995
Picobia alectoris Fain, Bochkov & Mironov, 2000: 65; Skoracki, Bochkov & Wauthy, 2004: 164
Picobia alectori: Skoracki & Dabert, 2002: 138
Columbiphilus alectoris (Fain, Bochkov et Mironov): Skoracki, 2011: 374; Skoracki & Sikora, 2011; Skoracki & Hromada, 2013; Skoracki, Sikora & Spicer, 2016: 25
Type locality: France (see the notes in Skoracki, Sikora & Spicer, 2016).
Distribution: France, Rwanda
Hosts: Alectoris rufa, Alectoris sp. (Galliformes: Phasianidae)
Columbiphilus khushalkhani Kivganov & Sharafat, 1995: 83; Skoracki, 2011: 374; Skoracki, Sikora & Spicer, 2016: 19
Picobia khushalkhani (Kivganov et Sharafat): Bochkov & Mironov, 1999: 16; Skoracki, Bochkov & Wauthy, 2004: 172
Picobia polonica Skoracki, Magowski & Dabert, 2001: 154 – Type loc. Wielkopolska, Zasutowo, Distr. Poland, India (syn. by Skoracki, Sikora & Spicer, 2016)
Columbiphilus polonica (Skoracki, Magowski et Dabert): Skoracki, 2011: 375; Skoracki & Sikora, 2011
Type locality of C. khushalkhani: Kabul (Afghanistan)
Distribution: Afghanistan, Belgium, Germany, India, Japan, Poland, USA, Central, Eastern and South Africa, USA
Hosts: Columba livia (Columbiformes: Columbidae – accidental host), Coturnix coturnix, Gallus gallus domesticus, G. sonneratii, Francolinus clappertoni (Galliformes: Phasianidae), Acryllium vulturinum, Numida meleagris (Galliformes: Numididae)
Columbiphilus odontophoridus Skoracki & Sikora, 2011: 28
Type locality: Bogota (Colombia)
Distribution: Colombia, Suriname
Hosts: Colinus cristatus, C. leucopogon (Galliformes: Odontophoridae)
Picobia pteroclesi Skoracki & OConnor, 2010: 27
Columbiphilus pteroclesi (Skoracki et OConnor): Skoracki, 2011: 379; Skoracki, Sikora & Spicer, 2016: 26
Type locality: Egypt, Red Sea Governate, Wadi el Metuli
Distribution: Egypt
Hosts: Pterocles coronatus, P. senegallus (Pterocliformes: Pteroclidae)
Gunabopicobia Skoracki & Hromada, 2013: 196
Type species: Picobia zumpti Lawrence, 1959
Gunabopicobia claravis Kaszewska, Skoracki & Hromada, 2018: 292
Type locality: Cauca Province, Mercaderes
Distribution: Colombia
Host: Claravis pretiosa (Columbiformes: Columbidae)
Gunabopicobia geotrygoni Kaszewska, Skoracki & Hromada, 2018: 292
Type locality: Venezuela, Silla de Caracas
Distribution: Colombia, Martinique, Paraguay, Venezuela
Hosts: Geotrygon chrysiei, G. frenata, G. linearis, G. montana (Columbiformes: Columbidae)
Gunabopicobia lathami Kaszewska, Skoracki & Hromada, 2018: 290
Type locality: Australia
Distribution: Australia, Indonesia, Papua New Guinea
Hosts: Leucosarcia melanoleuca, Caloenas nicobarica (Columbiformes: Columbidae)
Gunabopicobia leptotila Kaszewska, Skoracki & Hromada, 2018: 291
Type locality: Formosa Province, Tapiciolé
Distribution: Argentina
Host: Leptotila verreauxi (Columbiformes: Columbidae)
Gunabopicobia masalaje Kaszewska, Kavetska & Skoracki, 2014: 294; Skoracki, Sikora & Spicer, 2016: 26
Type locality: Papua New Guinea
Distribution: Indonesia (Sulawesi), Papua New Guinea
Hosts: Ducula luctuosa, D. pistrinaria, D. rosacea, D. rufigaster, D. spilorrhoa, Ptilinopus iozonus (Columbiformes: Columbidae: Treroninae)
Gunabopicobia metropelia Kaszewska, Skoracki & Hromada, 2018: 295
Type locality: Neuquén Province, Patagonia
Distribution: Argentina
Hosts: Metropelia melanoptera (Columbiformes: Columbidae)
Syringophilus zumpti Lawrence, 1959: 425
Picobia zumpti: Kethley, 1970: 65; Skoracki & Dabert, 2002: 144; Skoracki, Bochkov & Wauthy, 2004: 171; Bochkov, Williams & Proctor, 2005: 122
Neopicobia zumpti: Skoracki, 2011: 384
Gunabopicobia zumpti: Skoracki & Hromada, 2011: 197; Skoracki, Sikora & Spicer, 2016: 27; Kaszewska, Skoracki & Hromada, 2018: 294
Type locality: South Africa
Distribution: South Africa, USA
Hosts: Columba livia, Streptopelia capicola, S. senegalensis, Zenaida macroura (Columbiformes: Columbidae)
Lawrencipicobia Skoracki & Hromada, 2013: 199
Type species: Picobia poicephali Skoracki et Dabert, 2002
Lawrencipicobia calyptorhyncha Marciniak, Skoracki & Hromada, 2019: 194
Type locality: Australia
Distribution: Australia
Host: Calyptorhyncha lathami (Psittaciformes: Cacatuidae)
Picobia poicephali Skoracki & Dabert, 2002: 143, Skoracki, Bochkov & Wauthy, 2004: 171; Skoracki, 2005: 128; 2011: 329
Lawrencipicobia poicephali (Skoracki et Dabert): Skoracki & Hromada, 2014: 199; Skoracki, Sikora & Spicer, 2016: 29; Skoracki, Sikora & Hromada, 2019: 1610
Type locality: Galim
Distribution: Cameroon
Host: Poicephalus senegalus versteri (Psittaciformes: Psittacidae)
Lawrencipicobia sulphurea Marciniak, Skoracki & Hromada, 2019: 196
Type locality: West Timor, East Nusa Tenggara Province, Kupang Regency
Fatulaeu District, Camplong I (Tjamplong)
Distribution: Indonesia (West Timor)
Host: Cacacua sulphurea (Psittaciformes: Cacatuidae)
Neopicobia Skoracki, 2011: 383
Type species: Syringophilus anthi Fritsch, 1958
Syringophilus anthi Fritsch, 1958: 240
Picobia anthi: Fain, Bochkov & Mironov, 2000: 65; Skoracki, Bochkov & Wauthy, 2004: 172
Neopicobia anthi (Fritsch): Skoracki, 2011: 384
Type locality: “Schlesien”
Distribution: Germany, Poland
Hosts: Anthus cervinus, A. trivialis (Passeriformes: Motacillidae)
Picobia cardinalis Skoracki, Hendricks & Spicer, 2010: 733
Neopicobia cardinalis (Skoracki, Hendricks et Spicer): Skoracki, 2011: 384
Type locality: Texas, Johnson Co.
Distribution: USA (Texas)
Host: Cardinalis cardinalis (Passeriformes: Cardinalidae)
Picobia carpodacus Skoracki, Hendricks & Spicer, 2010: 737
Neopicobia carpodacus (Skoracki, Hendricks et Spicer): Skoracki, 2011: 384; Skoracki, Sikora & Spicer, 2016: 35
Type locality: California, Monterey County
Distribution: USA (California)
Host: Haemorhous [Carpodacus] purpureus (Passeriformes: Fringillidae)
Neopicobia ea Skoracki & Unsoeld, in: Skoracki, Unsoeld, Kaszewska & Kavetska, 2014: 78
Type locality: Venezuela, Rio Caura
Distribution: Brazil, Peru, Venezuela
Hosts: Celeus elegans, C. flavus, C. torquatus (Piciformes: Picidae)
Picobia epthianura Skoracki, Glowska & Sikora, 2008: 62
Neopicobia epthianura (Skoracki, Glowska et Sikora): Skoracki, 2011: 384
Type locality: Australia
Distribution: Australia
Host: Epthianura aurifrons (Passeriformes: Meliphagidae)
Neopicobia freya Skoracki & Unsoeld, in: Skoracki, Unsoeld, Kaszewska & Kavetska, 2014: 74; Klimovičova et al., 2014a: 6
Type locality: Paraguay
Distribution: Kenya, Panama, Paraguay, Tanzania
Hosts: Dryocopus galeatus, Dendropicos goerthae, D. griseocephalus, Colaptes [Piculus] rubiginosus (Piciformes: Picidae)
Neopicobia hepburni Glowska & Laniecka, 1914: 636; Skoracki, Sikora & Spicer, 2016: 37
Type locality: Peru
Distribution: Ecuador, Paraguay, Peru, Guyana
Hosts: Picumnus exilis, P. sclateri, P. olivaceus, P. temminckii (Piciformes: Picidae)
Neopicobia herbicolae Glowska, Laniecka & Milensky, 1915: 636
Type locality: Guyana
Distribution: Guyana
Host: Emberizoides herbicola (Passeriformes: Thraupidae)
Picobia ictericus Skoracki & Glowska, in: Skoracki, Glowska, Lontkowski & Stawarczyk, 2010: 143
Neopicobia ictericus (Skoracki et Glowska): Skoracki, Sikora & Spicer, 2016: 33
Type locality: Brazil
Distribution: Brazil
Hosts: Cacicus chrysopterus, Pseudoleistes guirahuro (Passeriformes: Icteridae)
Picobia modularis Skoracki & Magowski, 2001: 114; Skoracki, Bochkov & Wauthy, 2004: 171
Neopicobia modularis (Skoracki et Magowski): Skoracki, 2011: 386; Klimovičová & Hromada, 2014: 36
Type locality: Kuznica – Hel (Poland)
Distribution: Mongolia, Poland, Slovakia
Hosts: Prunella atrogularis, P. modularis (Passeriformes: Prunellidae), Pyrrhula pyrrhula (Pass., Fringillidae)
Neopicobia pari Skoracki, Sikora & Spicer, 2016: 33
Type locality: China, Fuijan Prov.
Distribution: China, Finland, Japan, N. Macedonia,
Hosts: Parus major, Periparus venustulus, Poecile varius (Passeriformes: Paridae)
Picobia troglodytes Skoracki, Hendricks & Spicer, 2010: 731
Neopicobia troglodytes (Skoracki et al.): Skoracki, 2011: 384
Type locality: California, Monterey Co.
Distribution: USA (California)
Hosts: Thryomanes bewickii, Troglodytes aedon (Passeriformes: Troglodytidae), Cardellina pusilla (Pass.: Parulidae)
Phipicobia Glowska & Schmidt, 2014a: 194
Type species: Phipicobia pygiptilae Glowska et Schmidt, 2004
Phipicobia pygiptilae Glowska & Schmidt, 2004: 19
Type locality: Guyana
Distribution: Guyana
Host: Pygiptila stellaris (Passeriformes: Thamnophilidae)
Picobia Haller, 1878: 81 – 91
Type species: Picobia heeri Haller, 1878
Picobia bernieriarum Skoracki, OConnor & Goodman, 2016: 1541
Type locality: Tulear Prov., Fivondronana de Tolagnaro, Forêt de Marosohy along
Enakara-Antseva trail, 15.5 km WNW Ranomafana (~24°34‘S, 46°48‘E)
Distribution: Madagascar
Hosts: Bernieria madagascariensis, Xanthomixis zosterops (Passeriformes: Bernieridae)
Picobia biarmicus Skoracki, Bochkov & Wauthy, 2004: 165; Skoracki, 2011: 328
Type locality: Miedzyodrze
Distribution: Poland
Host: Panurus biarmicus (Passeriformes: Panuridae)
Picobia caudati Skoracki & Hebda, 2004: 2; Zmudzinski & Unsoeld, 2016: 355
Type locality: Popielow near Opole (Poland)
Distribution: Germany, Poland
Host: Aegithalos caudatus (Passeriformes: Aegithalidae)
Picobia cetti Skoracki, 2011: 357
Type locality: Aras
Distribution: Turkey
Host: Cettia cetti (Passeriformes: Cettidae)
Picobia chloris Bochkov, Mironov & Kravtsova, 2000: 352; Skoracki, Bochkov & Wauthy, 2004: 172; Skoracki, 2011: 328
Type locality: Bishkek City
Distribution: Kyrghyzstan
Host: Chloris chloris (Passeriformes: Fringillidae)
Picobia cichladusa Skoracki, Solarczyk & Sikora, 2012: 124; Skoracki & Hromada, 2013: 201
Type locality: Soga
Distribution: Tanzania
Host: Cichladusa arcuata (Passeriformes: Muscicapidae)
Picobia cissa Skoracki, Bochkov & Wauthy, 2004: 168; Skoracki, 2011: 328; Skoracki, Sikora & Spicer, 2016: 52
Type locality: North Indochina
Distribution: North Indochina
Hosts: Cissa chinensis, Urocissa erythrorhyncha (Passeriformes: Corvidae)
Picobia corcoracus Skoracki, Glowska & Sikora, 2008: 64; Skoracki, 2011: 328
Type locality: Australia
Distribution: Australia
Host: Corcorax melanorhamphos (Passeriformes: Corcoracidae)
Picobia currucae Skoracki & Magowski, 2001: 116; Glowska, Skoracky & Khourly, 2007: 64; Skoracki, 2011: 348
Type locality: Poland, Kuznica-Hel
Distribution: Jordan, Poland, Turkey
Hosts: Sylvia curruca, S. hortensis, S. nisoria (Passeriformes: Sylviidae)
Picobia dinemellia Glowska & Skoracki, 2011: 64; Skoracki & Hromada, 2013: 201
Type locality: Tanzania
Distribution: Tanzania
Host: Dinemellia dinemelli (Passeriformes: Ploceidae)
Syringophilus dryobatis Fritsch, 1958: 242
Picobia dryobatis (Fritsch): Kethley, 1970: 65; Skoracki, Bochkov & Wauthy, 2004: 159; Nattress & Skoracki, 2009: 53; Skoracki, Hendricks & Spicer, 2010: 739; Skoracki, 2011: 334; Klimovičova et al., 2014a: 5; Glowska, Chrzanowski & Kaszewska, 2015: 50; Skoracki, Sikora & Spicer, 2016: 62
Picobia dryobates: Bochkov & Mironov, 1998: 15
Type locality: Germany; neotype from Poland (Darlowko Wschodnie, Bukovo – Kopan)
Distribution: Brazil, Germany, England, Ecuador, Ethiopia, Kenya, Moldavia, Nepal, Paraguay, Poland, Slovakia, Suriname, Tanzania, Trinidad and Tobago, USA, Venezuela, Zambia
Hosts: Campethera nubica, Dendrocopos borealis, D. darjellensis, D. leucotos, D. macei, D. major, D. hortorum, D. namaquus, D. stierlingi, D. syriacus, Dendropicos fuscescens, D. goerthae, D. namaquus, D. stierlingi, Dryocopus pileatus, Jynx torquilla, Melanerpes carolinus, M. erythrocephalus, Picoides mixtus, P. pubescens, P. scalaris, P. tridactylus, Veneliornis cassini, V. dignus, V. fumigatus, V. kirkii, V. maculifrons, V. passerines, V. sanguineus, V. spilogaster (Piciformes: Picidae); [Sturnus vulgaris (Passeriformes: Sturnidae)]. Note by Glowska, Chrzanowski & Kaszewska (2015): “Specimens from Sturnus vulgaris determined as P. dryobatis (Bochkov & Mironov 1998) belong to Picobia sturni (Skoracki et al. 2004d)”.
Picobia dziabaszewskii Glowska, Dragun-Damian & Dabert, 2012: 59; Klimovičova et al., 2014a: 5
Type locality: China
Distribution: China, Kenya, Namibia, Tanzania, Zambia
Hosts: Trochalopteron [Garrulax] formosus, Turdoides jardineii, T. leucopygia, T. rubiginosa (Passeriformes: Leiothrichidae)
Picobia echo Skoracki, Solarczyk & Sikora, 2012: 126; Skoracki & Hromada, 2013: 202
Type locality: South Kivu, Bukavu
Distribution: DR Congo, Indonesia, Nepal
Host: Cossypha heuglini, Copsychus malabricus, C. saularis (Passeriformes: Muscicapidae)
Picobia eremophila Skoracki, 2011: 363; Skoracki & Hromada, 2013: 202
Type locality: Egypt
Distribution: Egypt
Host: Eremophila bilopha (Passeriformes: Alaudidae)
Picobia galerida Skoracki, 2011: 367
Type locality: Zachodniopomorskie, near Swinoujscie
Distribution: Poland
Host: Galerida cristata (Passeriformes: Alaudidae)
Picobia Heeri Haller, 1878: 81–91; Oudemans, 1906: 73; Dubinin, 1957: 94; Kethley, 1970: 64; Skoracki, Bochkov & Wauthy, 2004: 158; Skoracki, 2011: 329; Skoracki et al., 2014: 71; Skoracki, Sikora & Spicer, 2016: 62
Syringophilus heeri: Poppe, 1888: 238
Picobia helleri [sic]: Bochkov & Mironov, 1999: 15
Type locality: Zurich (Switzerland)
Distribution: Germany, India, Indonesia (Sumatra), Japan, Nepal, Poland, Switzerland, Thailand
Hosts: Picus awokera, P. canus, P. chlorolophus, P. erythropygius, P. flavinucha, P. punicus, P. squamatus, P. viridis (Piciformes: Picidae)
Picobia illadopsae Skoracki & Hromada, 2013: 202
Type locality: Kakamega, Buyangu, Udo’s campsite, 0°21’11’’N, 34°51’50’’E, 1598 m a.s.l.
Distribution: Kenya
Host: Illadopsis rufipennis (Passeriformes: Pellorneidae)
Picobia indonesiana Skoracki & Glowska, 2008: 281; Skoracki, 2011: 328
Type locality: Indonesia
Distribution: Indonesia
Hosts: Aplonis panayensis, Enodes erythrophris, Mino dumontii (Passeriformes: Sturnidae)
Picobia lamprotornis Klimovičova, Skoracki, Wamiti & Hromada, 2014: 395; Klimovičova et al., 2014a: 2
Type locality: Soysambu Conservancy
Distribution: Kenya
Hosts: Lamprotornis superbus (Passeriformes: Sturnidae), Ploceus ocularis (Passeriformes: Ploceidae)
Picobia lemi Skoracki, Glowska & Sikora, 2008: 60; Skoracki, 2011: 328
Type locality: New Guinea
Distribution: Papua New Guinea
Hosts: Manucodia ater, M. chalybatus (Passeriformes: Paradisaeidae)
Picobia lonchura Skoracki, Sikora & Spicer, 2016: 53
Picobia lonchurae: Skoracki, Hromada, Prevuznakova & Wamiti, 2019: 1804, 2019: 1805
Type locality: Java
Distribution: Indonesia, Uganda
Hosts: Estrilda atricapilla, Lonchura leucogastra (Passeriformes: Estrildidae)
Picobia magellani Skoracki, Sikora & Spicer, 2016: 43
Type locality: Bogota
Distribution: Colombia
Host: Scytalopus magellanicus (Passeriformes: Rhinocriptidae)
Picobia makoli Skoracki, Sikora & Spicer, 2016: 45
Type locality: Leticia
Distribution: Colombia
Host: Xiphocolaptes promeropirhynchus (Passeriformes: Furnariidae)
Picobia mentalis Skoracki & Unsoeld, in: Skoracki, Unsoeld, Kaszewska & Kavetska, 2014: 71
Type locality: Malacca
Distribution: Indonesia (Java), Malaysia (Malacca)
Host: Picus mentalis (Piciformes: Picidae)
Picobia meru Klimovičova, Skoracki & Hromada, 2016: 582
Type locality: Arabuko Sokoke Forest; Ngaye Forest, Meru, 1220 m
Distribution: Kenya
Host: Prionops sulcifrons (Passeriformes: Vangidae)
Picobia myiopagi Glowska & Milensky, 2014: 378
Type locality: Guyana
Distribution: Guyana, Peru
Hosts: Elaenia ruficeps, Myiopagis gaimardii, M. subplacens (Passeriformes: Tyrannidae)
Picobia myrmecocichla Skoracki, Solarczyk & Sikora, 2012: 132; Skoracki & Hromada, 2013: 205
Type locality: Tanzania
Distribution: Tanzania
Host: Myrmecocichla arnotti (Passeriformes: Muscicapidae)
Picobia neomixis Skoracki, OConnor & Goodman, 2016: 1538
Type locality: Tulear Prov., Toliara Province, Fivondronana de
Tolagnaro, 1.5 km NW Manafiafy (~24°46’S, 47°11’E)
Distribution: Madagascar
Host: Neomixis tenella (Passeriformes: Cisticolidae)
Picobia ochoi Glowska & Milensky, 2014: 374
Type locality: Guyana, Cuyuni-Mazarini, Paruima
Distribution: Guyana
Host: Tolmomyias sulphurescens (Passeriformes: Tyrannidae)
Picobia oritis Skoracki, Antczak & Riegert, 2009: 133; Skoracki, 2011: 328; Skoracki & Hromada, 2013: 205; Klimovičova et al., 2014a; Skoracki, Sikora & Spicer, 2016: 52
Type locality: Cameroon, North-Western Province, Bamenda Highlands,
Mendongbuo, 06°05’N, 10°18’E, 2200 m a.s.l.
Distribution: Angola, Botswana, Cameroon, Kenya, Somalia, South Africa, Uganda, Zambia
Hosts: Cinnyris erythrocercus, C. oustaleti, C. talatala, C. venustus, Cyanomitra olivacea, C. oritis (Passeriformes: Nectariniidae)
Picobia paludicola Skoracki & Kiljan, 2002: 2; Skoracki, Bochkov & Wauthy, 2004: 171; Skoracki, 2011: 328
Type locality: Swinoujscie
Distribution: Poland
Host: Acrocephalus paludicola (Passeriformes: Acrocephalidae)
Picobia passeri Skoracki & Sikora, 2014: 195
Type locality: Russia, Tula District, Suvorov
Distribution: Russia
Host: Passer domesticus (Passeriformes: Passeridae)
Picobia phoeniculi Fain, Bochkov & Mironov, 2000: 69; Skoracki & Dabert, 2002: 138; Skoracki, Bochkov & Wauthy, 2004: 168; Skoracki & Hromada, 2013: 206; Skoracki, Sikora & Spicer, 2016: 62
Type locality: Rwanda
Distribution: N. Macedonia, Kazakhstan, Pakistan, Rwanda, Tanzania
Hosts: Phoeniculus purpureus ruwenzoriae (Upupiformes: Phoeniculidae), Upupa epops africana, U. epops (Upupiformes: Upupidae)
Picobia phoenicuri Skoracki & Hromada, 2013: 208
Type locality: Tunisia
Distribution: Tunisia
Host: Phoenicurus moussieri (Passeriformes: Muscicapidae)
Picobia ploceus Klimovičova, Skoracki, Wamiti & Hromada, 2014: 395
Type locality: Kenya
Distribution: Kenya
Host: Ploceus ocularis (Passeriformes: Ploceidae)
Picobia psaltriparus Skoracki, Hendricks & Spicer, 2010: 738
Type locality: Texas, Jeff Davis Co.
Distribution: USA
Host: Psaltriparus minimus (Passeriformes: Aegithalidae)
Picobia pycnonoti Glowska, Skoracki & Khourly, 2007: 64; Skoracki, 2011: 346; Klimovičova et al., 2014a: 6; Skoracki, Sikora & Spicer, 2016: 48
Type locality: Al-Dhleil (Jordan)
Distribution: Jordan, Kenya, Philippines
Hosts: Pycnonotus barbatus, P. goiavier, P. xanthopygus (Passeriformes: Pycnonotidae)
Picobia riparius Skoracki, 2011: 360
Type locality: Lubuskie, Zielona Gora
Distribution: Poland
Host: Riparia riparia (Passeriformes: Hirundinidae)
Picobia sayornis Glowska, 2014: 140
Type locality: Peru, Tumbes, El Caucho Biol. Station
Distribution: Peru
Host: Sayornis nigricans (Passeriformes: Tyrannidae)
Picobia schmidti Glowska & Milensky, 2014: 378
Type locality: Peru, Tumbes, El Caucho Biol. Station
Distribution: Guyana, Peru
Hosts: Lophotriccus pilatus, Hemitriccus margaritaceiventer (Passeriformes: Tyrannidae)
Picobia sturni Skoracki, Bochkov & Wauthy, 2004: 159; Bochkov & Mironov, 1998; Skoracki, 2011b; Skoracki, Sikora & Spicer, 2016: 61
Type locality: Poland, Bukowo – Kopan
Distribution: Japan, Moldavia, Morocco, Poland, Slovakia
Hosts: Sturnus cineraceus, S. unicolor, S. vulgaris (Passeriformes: Sturnidae)
Picobia troglodytidus Sikora & Skoracki, 2012: 62
Type locality: Costa Rica, Cartago
Distribution: Costa Rica
Host: Henicorhina leucophrys (Passeriformes: Troglodytidae)
Argentina, Skoracki & Sikora, 2002: 8
Host: Aphrastura spinicauda (Passeriformes: Furnariidae)
Pipicobia Glowska & Schmidt, 2014b: 58
Type species: Picobia locustella Skoracki, Bochkov et Wauthy, 2004
Picobia glossopsitta Skoracki, Glowska & Sikora, 2008: 67
Neopicobia glossopsitta: Skoracki, 2011: 384
Pipicobia glossopsitta: Glowska & Schmidt, 2014: 59
Type locality: Australia
Distribution: Australia
Host: Glossopsitta porphyrocephala (Psittaciformes: Psittaculidae)
Picobia locustella Skoracki, Bochkov & Wauthy, 2004: 164
Neopicobia locustella Scoracki, 2011: 389
Pipicobia locustella (Skoracki, Bochkov et Wauthy): Glowska & Schmidt, 2014: 58
Type locality: Poland, Swinoujscie
Distribution: Poland
Hosts: Locustella luscinioides, L. naevia (Passeriformes: Locustelliidae)
Picobia pyrrholaemus Skoracki & Glowska, 2008: 284
Neopicobia pyrrholaemus Scoracki, 2011: 384
Pipicobia pyrrholaemus (Skoracki et Glowska): Glowska & Schmidt, 2014: 59
Type locality: Australia
Distribution: Australia
Host: Pyrrholaemus sagittatus (Passeriformes: Acanthizidae)
Pipicobia terpsiphoni Glowska & Schmidt, 2014: 59; Skoracki et al., 2016: 1544
Type locality: Ogooue Maritime Province (Moukalaba-Doudou National Park, Gabon)
Distribution: Gabon, Madagascar
Hosts: Terpsiphone mutata, T. rufiventer (Passeriformes: Monarchidae)
Pseudopicobia Skoracki, Scibek & Sikora, 2012: 230
Type species: Pseudopicobia nonnula Skoracki, Scibek et Sikora, 2012
Pseudopicobia hapaloptila Skoracki, Scibek & Sikora, 2012: 234
Type locality: Riobamba
Distribution: Ecuador
Host: Hapaloptila castanea (Galbuliformes: Bucconidae)
Pseudopicobia malacoptila Skoracki, Scibek & Sikora, 2012: 233
Type locality: Antioquia, Mutatá
Distribution: Colombia
Host: Malacoptila panamensis (Galbuliformes: Bucconidae)
Pseudopicobia nonnula Skoracki, Scibek & Sikora, 2012: 230
Type locality: Bogota
Distribution: Colombia
Host: Nonnula frontalis (Galbuliformes: Bucconidae)
Rafapicobia Skoracki, 2011: 391
Type species: Rafapicobia zirnitra Skoracki, 2011
Rafapicobia automoli Glowska, Laniecka & Milensky, 2015: 489
Type locality: Guyana
Distribution: Guyana
Hosts: Automolus ochralaemus, A. rufipileatus (Passeriformes: Furnariidae)
Rafapicobia brachypteraci Skoracki, Hromada & Sikora, 2020: 171
Type locality: Madagascar
Distribution: Madagascar
Host: Atelornis crossleyi (Coraciiformes: Brachypteracidae)
Picobia brotogeris Fain, Bochkov & Mironov, 2000: 62; Skoracki, Bochkov & Wauthy, 2004: 163; Skoracki, 2005: 128; 2011: 328
Picobia brotogeri: Skoracki & Sikora, 2002: 1
Rafapicobia brotogeris (Fain, Bochkov et Mironov): Skoracki, Sikora & Spicer, 2016: 73
Type locality: Upper Amazonia (Zoo of Antwerp)
Distribution: Brazil
Host: Brotogeris jugularis cyanopterus (Psittaciformes: Psittacidae)
Rafapicobia copsychus Skoracki, OConnor & Goodman, 2016: 1536
Type locality: Tulear Province, Fivondronana de Tolagnaro, Forêt de Marovony, 19 km NE Manantenina (~24°06’S, 47°22’E)
Distribution: Madagascar
Host: Copsychus albospecularis (Passeriformes: Turdidae)
Rafapicobia dendrocolaptesi Skoracki & Solarczyk, 2012: 60
Type locality: Paraguay, Apa-Bergland
Distribution: Argentina, Paraguay
Hosts: Dendrocolaptes platyrostris, D. picumnus (Passeriformes: Furnariidae)
Rafapicobia lepidocolaptesi Skoracki & Solarczyk, 2012: 62; Skoracki, Sikora & Spicer, 2016: 78
Type locality: Ecuador, Guale
Distribution: Colombia, Ecuador, Venezuela
Hosts: Lepidocolaptes affinis, L. souleyetii (Passeriformes: Furnariidae)
Rafapicobia melzeri Skoracki, Unsoeld, Skorupski & Kavetska, 2014: 228; Skoracki, Sikora & Spicer, 2016: 79
Type locality: Germany, Bayern, Ismaninger Lake
Distribution: Chile, France, Germany, Japan, Kirghizia
Hosts: Pardirallus sanguinolentus, Porzana fusca, P. parva, P. porzana, Rallus aquaticus (Gruiformes: Rallidae)
Rafapicobia milenskyi Glowska & Schmidt, 2014: 198
Type locality: Upper Takutu-Upper Essequibo, Lower Rewa River
Distribution: Guyana
Host: Conopophaga aurita (Passeriformes: Conopophagidae)
Rafapicobia momoti Skoracki, Hromada & Sikora, 2020: 171
Type locality: Brazil
Distribution: Brazil
Host: Baryphthengus ruficapillus (Coraciiformes: Momotidae)
Rafapicobia olszanowskii Skoracki, Zmudzinski & Sikora, 2020: 449
Type locality: Ecuador
Distribution: Ecuador
Host: Semnornis ramphastinus (Piciformes: Semnornithidae)
Picobia ramphastos Fain, Bochkov & Mironov, 2000: 33; Skoracki, Bochkov & Wauthy, 2004: 163
Rafapicobia ramphastos (Fain, Bochkov and Mironov): Skoracki, Sikora & Spicer, 2016: 73
Type locality: Guatemala (Zoo of Antwerp)
Distribution: Brazil, Guatemala
Hosts: Ramphastos sulfuratus, R. vietellinus (Piciformes: Ramphastidae)
Rafapicobia thamnophili Glowska & Schmidt, 2014: 196
Type locality: Guyana, Cuyuni Mazarini, Paruima, ca.19 km N, at Mt. Holitipu
Distribution: Guyana
Hosts: Hypocnemis cantator, Myrmeciza ferruginea, Myrmoborus leucophrys, Myrmotherula longipennis, Thamnophilus insignis (Passeriformes: Thamnophilidae)
Rafapicobia toxostoma Sikora, Fajfer & Skoracki, 2012: 30
Type locality: USA
Distribution: USA
Host: Toxostoma curvirostre (Passeriformes: Mimidae)
Rafapicobia zirnitra Skoracki, 2011: 392; Skoracki, Sikora & Spicer, 2016: 77
Type locality: Zachodniopomorskie, near Swinoujscie
Distribution: China, Japan, Poland
Hosts: Ficedula hypoleuca, Luscinia akahige, L. cyanura, L. sibilans, Phoenicurus auroreus, Saxicola rubetra (Passeriformes: Muscicapidae)
Tanopicobia Skoracki, Sikora, Jerzak & Hromada, 2020: 4
Type species: Tanopicobia trachyphoni Skoracki, Sikora, Jerzak et Hromada, 2020
Tanopicobia trachyphoni Skoracki, Sikora, Jerzak & Hromada, 2020: 4
Type locality: Arusha
Distribution: Tanzania
Host: Trachyphonus erythrocephalus (Piciformes: Lybiidae)
EUROPE
Austria (Skoracki, Zmudzinski, Unsoeld & Sikora, 2016) – Stibarokris plataleus
Azerbaidzhan (Amirhanov, 1970, 1973) – Syringophiloidus minor, Syringophilus bipectinatus
Belarus (Bochkov, 2000; Skoracki, 2011) – Syringophilus bipectinatus
Belgium (Bochkov, Fain & Skoracki, 2004; Skoracki, Sikora & Spicer, 2016) – Columbiphilus khushalkhani, Mironovia coturnae
Bosnia and Herzegovina (Skoracki, 2011; Skoracki, Sikora & Spicer, 2016; Skoracki, Hromada & Sikora, 2017) – Peristerophila meropis, Picobia dryobatis, Syringophiloidus minor
Croatia (Skoracki, 2011) – Aulobia sylviae, Syringophiloidus minor
Denmark (Skoracki, 2011) – Syringophiloidus hirundinis
Finland (Skoracki, 2004d; 2011) – Torotrogla calcarius, T. cardueli
France (Oudemans, 1904, 1906; Bochkov & Mironov, 199[8]9; Bochkov, Mironov & Skoracki, 2001; Skoracki, 2011; Skoracki & Sikora, 2011; Skoracki et al., 2014; Zmudzinski et al., 2015; Skoracki, Sikora & Spicer, 2016; Skoracki, Spicer & OConnor, 2016) – Aulobia sylviae, Charadriineopicobia apricaria, Charadriphilus gallinago, Columbiphilus alectoris, Creagonycha totani, Megasyringophilus aquilus, Neosyringophilopsis phylloscopi, Niglarobia helleri, Peristerophila accipitridicus, Rafapicobia melzeri, Syringophiloidus minor, S. presentalis
Corsica (Skoracki, 2011; Skoracki, Hromada & Sikora, 2017) – Peristerophila meropis, Syringophilus bipectinatus
Germany (Heller, 1880; Fritsch, 1958; Skoracki, 2011; Skoracki et al., 2014; Zmudzinski et al., 2015; Skoracki et al., 2018; Zmudzinski & Unsoeld, 2019; Kaszewska et al., 2020) – Aulobia cardui, Aulonastus fragilis, Bubophilus asiobius, Charadriineopicobia ristowi, Columbiphilus khushalkhani, Neoaulonastus bisetatus, N. picidus, Neosyringophilopsis trogloditis, Neopicobia anthi, Niglarobia lapponica, N. vanelli, Picobia caudati, P. dryobatis, P. heeri, Peristerophila accipitridicus, P. columbae, P. falcophila, Rafapicobia melzeri, Selenonycha charadriiformes, Stibarokris dastychi, Syringophiloidus coccothraustes, S. glandarii, S. klimovi, S. minor, S. parapresentalis, S. picidus, S. weiszii, Syringophilopsis fringillae, S. garrulus, S. kazmierski, S. kirgizorum, S. troglodytis, S. turdi, Syringophilus bipectinatus, Torotrogla cardueli, T. merulae
Gibraltar (Skoracki, Hromada & Sikora, 2017) – Peristerophila meropis
Great Britain (Kethley, 1970; Nattress & Skoracki, 2007, 2009; Kaszewska et al., 2020) – Bubophilus aluconis, Neosyringophilopsis aegithali, N. troglodytis, Picobia dryobatis, Peristerophila columbae, Syringophiloidus minor, Syringophilopsis fringillae, S. hirundus, S. kirgizorum, Torotrogla modularis
Greece (Skoracki, Hromada & Sikora, 2017; Kaszewska et al., 2020) – Peristerophila columbae, C. meropis, Procellariisyringophilus bulwerius
Hungary (Kaszewska et al., 2020) – Peristerophila columbae
Iceland (Bochkov & Skirnisson, 2011) – Mironovia lagopus
Italy (contin.)(Skoracki, Møller & Tryjanowski, 2003; Skoracki, 2004, 2011) – Syringophiloidus hirundinis, S. petronicus, Torotrogla lusciniae
Sardinia (Skoracki, Hromada & Sikora, 2017) – Peristerophila meropis
Sicily (Skoracki & Sikora, 2011) – Galliphilopsis szeptyckii [recorded as «bochkovi»]
Malta (Zmudzinski, Solarczyk & Skoracki, 2016) – Stibarokris hydrobates
Moldavia (Bochkov & Mironov, 1999) – Picobia dryobatis, P. sturni
Monaco (Kaszewska et al., 2020) – Peristerophila mucuya
North Macedonia (Skoracki, Sikora & Spicer, 2016: 62; Skoracki, Hromada & Sikora, 2017; Kaszewska et al., 2020) – Peristerophila columbae, Picobia phoeniculi, Peristerophila meropis
Norway (Bochkov & Skirnisson, 2011; Skoracki & Sikora, 2011) – Mironovia lagopus
Svalbard (Skoracki & Zawierucha, 2016) – Chenophila nanseni
Poland (Skoracki, 1999a, 1999b, 2002, 2004a, c, d, 2011; Skoracki & Skoracka, 1999; Skoracki & Dabert, 1999; Skoracki & Magowski, 2001; Skoracki & Bochkov, 2002; Skoracki & Kiljan, 2002; Skoracki & Hebda, 2004; Skoracki, Bochkov & Sikora, 2004; Skoracki, Bochkov & Wauthy, 2004; Skoracki, Dabert & Schmäschke, 2006; Skoracki, Michalik & Sikora, 2010; Skoracki et al., 2012; Skoracki, Sikora & Spicer, 2016) – Aulobia cardueli, A. sylviae, Aulonastus albus, Betasyringophyloidus phoenicurus, B. saxicolus, B. schoeniclus, Bubophilus aluconis, B. asiobius, Charadriphilus gallinago, Chenophila platyrhynchos, Columbiphilus Columbiphilus khushalkhani (= polonica), Cregonycha totani, Krantziaulonastus buczekae, Meitingsunes columbicus, Mironovia coturnae, Neoaulonastus [Dissonus] scirpaceus, Neopicobia anthi, Neosyringophilopsis aegithali, N. locustellus, N. phylloscopi, N. troglodytis, Niglarobia helleri, Neopicobia modularis, Peristerophila columbae, Philoxanthornea clarki, Picobia biarmicus, P. caudati, P. currucae, P. dryobatis, P. galerida, P. heeri, P. paludicola, P. riparius, P. sturni, Pipicobia locustella, Selenonycha charadriiformes, Stibarokris langei, Syringophiloidus bombycillae, S. carpodaci, S. coccothraustes, S. delichonum, S. hirundinis, S. minor, S. parapresentalis, S. presentalis, Syringophilopsis acrocephali, S. blaszaki, S. borini, S. fringillae, S. hirundus, S. kazmierski, S. kirgizorum, S. rusticus, S. spinolettus, S. sturni, S. troglodytis, S. turdi, Syringophilus bipectinatus, Torotrogla cardueli, T. gaudi, T. merulae, T. modularis, T. rubeculi
Portugal (cont.)
Madeira (Kaszewska et al., 2020) – Peristerophila columbae
Romania (Skoracki, 2011; Skoracki, Hromada & Sikora, 2017; Skoracki, 2018) – Peristerophila meropis, Peristerophila falcophila, Syringophilopsis kazmierski, S. kristini, Torotrogla merulae
Russia (all, continent.)(Dubinina, 1987; Bochkov, 2000, 2001; Bochkov & Apanaskevich, 2001; Bochkov & Chystiakov, 2001; Bochkov & Mironov, 1998, 1999; Bochkov, Mironov & Skoracki, 2001; Bochkov & Chistyakov, 2001; Skoracki, 2011; Skoracki & Mironov, 2013; Skoracki & Sikora, 2014; Skoracki, Hromada & Sikora, 2017) – Aulobia cardueli, A. galinago, A. sylviae, Aulonastus prunellae, Betasyringophiloidus motacillae, B. emberizae, B. schoeniclus, Peristerophila meropis, Charadriphilus lyudmilae, Chenophila canduli, Krantziaulonastus dubinini, Melingsunes columbicus, Neoaulonastus zosterops, Neosyringophilopsis aegithali, Niglarobia vanelli, Philoxanthornea clarki, Picobia passeri, Selenonycha charadriiformicus, Stibarokris langei, Syringophiloidus carpodaci, S. delichonum, S. glandarii, S. minor, S. presentatus, Syringophilopsis blascaki, S. borini, S. fringillae, S. kirgizorum, S. pari, S. sittae, S. sturmi, S. troglodytis, S. turdus, Syringophilus bipectinatus, S. parapresentalis, Torotrogla cardueli, T. gaudi, T. lullulae, T. merulae, T. rubeculi, T. volgini
Slovakia (Skoracki, Hromada & Kuczynski, 2001; Skoracki, 2002; Skoracki, Tryjanowski & Hromada, 2002; Skoracki, 2011; Klimovičová & Hromada, 2014) – Aulobia cardueli, Aulonastus albus, Betasyringophiloidus motacillae, B. schoeniclus, Neoaulonastus bisetatus, N. picidus, Neosyringophylopsis aegithali, N. phylloscopi, Niglarobia helleri, Picobia dryobatis, P. sturni, Syringophiloidus bombycillae, S. minor, S. montanus, S. parapresentalis, S. presentalis, Syringophilopsis acrocephali, S. blaszaki, S. fringillae, S. kazmierski, S. kristini, S. spinolettus, Torotrogla cardueli, T. gaudi, T. lullulae, T. merulae, T. modularis, T. rubeculi
Spain (cont.) (Skoracki, 2011; Zmudzinski et al., 2015; Skoracki, Hromada & Sikora, 2017) – Peristerophila meropis, Megasyringophilus aquilus, Syringophiloidus hirundinis
Canary Isl. (Skoracki, 2004) – Syringophiloidus erythrurus
Switzerland (Haller, 1878) – Picobia heeri
Turkey (entire)(Bochkov & Mironov, 1998; Skoracki, 2011; Skoracki, Zmudzinski, Unsoeld & Sikora, 2016; Skoracki, Hromada & Sikora, 2017; Zmudzinski, Solarczyk & Skoradski, 2016) – Peristerophila meropis, Chenophila marmaronetta, Peristerophila columbae, Picobia cetti, P. currucae, Stibarokris brevisetosus, S. phoeniconaias
Ukraine (Kivganov & Sharafat, 1995; Bochkov, 2000; Skoracki, 2011; Pidhorna et al., 2020) – Betasyringophiloidus phoenicurus, B. saxicolus, Creagonycha sternae, Kethleyana gelochelidoni, Neoaulonastus bisetatus, Philoxanthornea clarki, Syringophiloidus hirundinis, Syringophilopsis acrocephali, S. blaszaki, S. fringillae, S. hirundus, S. kazmierski, S. rusticus, S. sturni, Syringophilus bipectinatus, Torotrogla merulae, T. rubeculi
West Europe (Skoracki, 2004) – Syringophiloidus oriolus
Europe (Skoracki, 2011) – Aulobia leucosticte, Aulonastus caligatus, Syringophilopsis S. idunae, S. nucifragus
AFRICA
Angola (Klimovičova et al., 2014c; Kaszewska et al., 2020) – Picobia oritis
Botswana (Skoracky & OConnor, 2010; Klimovičova et al., 2014c) – Picobia oritis, Stibarokris phoeniconaias
Cameroon (Skoracki & Dabert, 2002; Skoracki, Tryjanowski & Hromada, 2002; Skoracki & Sikora, 2003; Skoracki, Antczak, Riegert, Fainová & Mikeš, 2009; Skoracki & Hromada, 2013; Skoracki, Unsoeld, Marciniak & Sikora, 2016; Skoracki, Sikora, Marciniak & Zmudzinski, 2016) – Aulobia cisticolae, Bubophilus ascalaphus, B. tytonus, Lawrencipicobia poicephali, Meitingsunes columbicus, Neoaulonastus zosterops, Picobia oritis, Syringophiloidus bucerotidus, Syringophilopsis kirgisorum, S. veselovsky, S. yosefi, Syringophilus bipectinatus
Congo DR (Fain, Bochkov & Mironov, 2000; Skoracki & Sikora, 2008; Skoracki et al., 2009; Skoracki, Solarczyk & Sikora, 2012; Skoracki, Sikora, Marciniak & Zmudzinski, 2016; Kaszewska et al., 2020) – Aulobia cisticolae, Blaszakia schalowi, Peristerophila columbae, Picobia echo, Syringophiloidus bucerotidus, S. cypsiuri, S. minor, S. seiuri
Egypt (Skoracki & OConnor, 2010; Skoracki et al., 2006; Skoracki, 2011; Glowska & Laniecka, 2012) – Columbiphilus pteroclesi, Philoxanthornea dubinini, Picobia eremophila, Pteroclidisyringophus re, Selenonycha charadriiformes, Syringophilopsis davidi
Ethiopia (Bochkov & Fain, 2003; Skoracki, 2014; Skoracki, Hromada, Zmudzinski, Unsoeld & Sikora, 2018; Kaszewska et al., 2020) – Aulonastus nectariniphilus, Neoaulobia agapornis, Peristerophila claravis, Picineoaulonastus pogoniulus, Picobia dryobatis
Gabon (Glowska & Schmidt, 2014; Glowska, Broda, Gebhard & Dabert, 2016) – Pipicobia terpsiphoni, Syringophiloidus furthi, S. plocei
Guinea (Skoracki & Sikora, 2008) – Blaszakia tauracos
Kenya (Skoracki, Hromada & Wamiti, 2011; Skoracki & Hromada, 2013; Klimovičova et al., 2014a, 2016; Skoracki et al., 2014; Skoracki, Hromada & Sikora, 2017; Skoracki et al., 2018; Skoracki, Hromada, Pervuznakova & Wamiti, 2019; Kaszewska et al., 2020) – Aulobia cisticolae, A. sylviettae, Betasyringophiloidus seiuri, Peristerophila columbae, P. meropis, Neoaulonastus apalis, N. bisetatus, N. lonchurus, N. malaconotus, N. oryzivorus, N. prionops, Neopicobia freya, Neosyringophilopsis lybidus, Peristerophila columbae, P. falcophila, P. upupi, Picineoaulonastus pogoniulus, Picobia dryobatis, P. dziabaszewskii, P. hedydipnae, P. illanopsae, P. lamprotornis, P. meru, P. ploceus, P. pycnonoti, Syrigophiloidus picidus, Syringophilopsis dicruri, S. veselovsky
Liberia (Klimovičova et al., 2014) – Neoaulonastus malaconotus
Madagascar (Skoracki, OConnor, Goodman & Marciniak-Musial, 2016; Skoracki, Hromada & Sikora, 2020) – Neoaulonastus tanzanicus, Peristerophila accipitridicus, Picobia bernieriarum, P. neomixis, Pipicobia terpsiphoni, Rafapicobia brachypteraci, R. copsychus
Morocco (Kethley, 1970; Skoracki, Sikora & Spicer, 2016; Skoracki, Hromada & Sikora, 2017) – Peristerophila meropis, Picobia sturni, Syringophilopsis fringillae
Namibia (Bochkov & Fain, 2003; Skoracki, Dabert & Schmaschke, 2006; Klimovičova et al., 2014; Glowska, Romanowska, Schmidt & Dabert, 2018) – Neoaulobia agapornis, Niglarobia cursoriae, Picobia dziabaszewskii, Torotrogla paenae
Rwanda (Fain, Bochkov & Mironov, 2000; Skoracki & Sikora, 2003) – Aulobia cisticolae, Columbiphilus alectoris, Syringophilopsis emberizae, S. sylviettae
Somalia (Klimovičova et al., 2014c) – Neoaulonastus malaconotus, Picobia oritis
South Africa (Lawrence, 1959; Kethley, 1970; Skoracki & Dabert, 2002; Skoracki, Bochkov & Wauthy, 2004; Skoracki, Zmudzinski, Unsoeld & Sikora, 2016; Skoracki, Sikora & Spicer, 2016; Kaszewska et al., 2020) – Bubophilus ascalaphus, Columbiphilus khushalkhani, Neopicobia zumpti, Galliphilopsis francolinus, Peristerophila columbae, Picobia oritis, Stibarokris geronticus, Syringophilus bipectinatus
Sudan (Kaszewska et al., 2020) – Peristerophila claravis
Tanzania (Bochkov & Fain, 2003; Skoracki & Sikora, 2008; Glowska et al., 2012; Skoracki, Solarczyk & Sikora, 2012; Skoracki, Hromada & Unsoeld, 2013; Skoracki & Hromada, 2013; Klimovičova et al., 2014a; Skoracki et al., 2014; Skoracki, Unsoeld, Marciniak & Sikora, 2016; Skoracki, Sikora, Marciniak & Zmudzinski, 2016; Hromada, Klimovičova, Unsöld & Skoracki, 2016; Skoracki, Hromada & Sikora, 2017; Skoracki et al., 2018; Skoracki, Hromada, Zmudzinski, Unsoeld & Sikora, 2018; Skoracki, Sikora, Jerzak & Hromada, 2020; Kaszewska et al., 2020) – Aulobia afroanthreptes, A. nectariniae, Aulonastus nectariniphilus, A. paridus, Blaszakia tauracos, Bubophilus aluconis, Peristerophila meropis, Colisyringophilus tanzanicus, Cuculisyringophilus chirovi, Neoaulobia agapornis, Neoaulonastus cinnyris, N. granatina, N. quelea, N. tanzanicus, Neopicobia freya, Niglarobia vanelli, Peristerophila claravis, P. columbae, Picineoaulonastus pogoniulus, Picobia cichladusa, P. dinemellia, P. dryobatis, P. dziabaszewskii, P. myrmecocichla, P. phoeniculi, Syringophiloidus bucerotidus, S. pseudonigritae, S. picidus, Syringophilus nectarinae, Tanopicobia trachyphoni
Togo (Skoracki & Dabert, 1999, 2000, 2001b; Skoracki & Sikora, 2003; Glowska & Skoracki, 2010) – Aulobia cisticolae, A. erythroptera, A. virens, Creagonycha totani, Meitingsunes [Peristerophila] tympanistria, Neoaulobia puylaerti, Syringophiloidus seiuri, Syringophilopsis albicollisi, S. lagonostictus, S. melittophagi, S. nitens
Tunisia (Skoracki, 2011; Skoracki & Hromada, 2013; Skoracki, Unsoeld & Skorupski, 2014) – Picobia phoenicuri, Pteroclidisyringophilus otididus
Uganda (Skoracki, 2014; Klimovičova et al., 2014c; Skoracki, Hromada, Pervuznakova & Wamiti, 2019) – Neoaulonastus lonchurae, N. oryzivorus, Picobia lonchurae, P. oritis, Syrigophiloidus estrildus, S. picidus
Zambia (Fain, 2000; Klimovičova et al., 2014c) – Neoaulobia agapornis, Picobia dryobatis, P. dziabaszewskii, P. oritis
Central Africa (Skoracki & Dabert, 2000; Skoracki, Sikora & Spicer, 2016) – Columbiphilus khushalkhani, Syringophilopsis albicollisi
Eastern Africa (Skoracki, Sikora & Spicer, 2016) – Columbiphilus khushalkhani
Continental Africa (Skoracki, 2008) – Neosyringophilopsis lonchurus
ASIA
Afghanistan (Kivganov & Sharafat, 1995) – Columbiphilus khushalkhani
China (Liu Bai-li, 1988; Sun – undated; Skoracki, 2011; Glowska, Gragun-Damian & Dabert, 2012; Skoracki, Sikora & Spicer, 2016; Skoracki, Zmudzinski & Solarczyk, 2017; Kaszewska et al., 2020) – Ixobrychiphilus wallacei, Krantziaulonastus lonchurus, Rafapicobia zirnitra, Syringophiloidus minor, S. oriolus, Peristerophila columbae, Picobia dziabaszewskii
India (Bochkov & Mironov, 1998; Fain, Bochkov & Mironov, 2000; Bochkov, Fain & Skoracki, 2004; Skoracki & Sikora, 2011; Skoracki, Unsoeld, Marciniak & Sikora, 2016; Skoracki, Sikora & Spicer, 2016) – Columbiphilus khushalkhani (= polonica), Fritschisyringophilus lonchurae, Neobubophilus atheneus, Peristerophila columbae, Picobia dryobatis
Indonesia (Skoracki, 2005; Skoracki & Glowska, 2008c; Skoracki & Sikora, 2011; Kaszewska, Kavetska & Skoracki, 2014; Skoracki et al., 2014; Kaszewska, Skoracki & Kavetska, 2016; Hromada, Klimovičova, Unsöld & Skoracki, 2016; Kaszewska, Skoracki & Hromada, 2018; Marciniak, Skoracki & Hromada, 2019; Kaszewska et al., 2020) – Apodisyringiana haszprunari [Sumatra], Cuculisyringophilus chirovi, Galliphilopsis szeptyckii, Gunabopicobia lathami, G. masalaje, Lawrencipicobia sulphurea, Meitingsunes chalcophaps, M. turacoenas, Peristerophila geopelis, P. lature, Picobia echo, P. heeri, P. mentalis, Procellariisyringophilus bulwerius
Iran (Bochkov & Mironov, 1998; Skoracki, 2011; Skoracki, Sikora & Spicer, 2016) – Peristerophila columbae, Picobia dryobatis
Japan (Skoracki, 2004; 2011; Skoracki & Sikora, 2011; Skoracki et al., 2014; Skoracki, Kaszewska & Kavetska, 2015; Skoracki, Sikora & Spicer, 2016; Glowska, Laniecka & Romanowska, 2018; Kaszewskaet al., 2020) – Apodisyringiana hirundapi, Columbiphilus khushalkhani, Galliphilopsis szeptyckii, Krantziaulonastus oryzivorus, Peristerophila columbae, Picobia dryobatis P. heeri, P. sturni, Rafapicobia zirnitra, Syringophiloidus glandarii, Syringophilopsis muscicapus, Torotrogla emberizae
Jordan (Glowska, Skoracki & Khourly, 2007; Skoracki, 2011) – Aulobia sylviae, Peristerophila columbae, P. mucuya, Picobia currucae, P. pycnonoti, Syringophiloidus minor, S. parapresentalis, S. philomelosus, S. presentalis, Syringophilopsis kazmierski, S. kirgizorum, Torotrogla merulae, T. pycnonotus
Kazakhstan (Bochkov & Mironov, 1999; Bochkov & Apanaskevich, 2001; Skoracki & Bochkov, 2010; Skoracky, 2011; Skoracki, Sikora & Spicer, 2016) – Aulobia cardueli, A. sylviae, Betasyringophyloidus phoenicurus, Charadriphilus ralli, Corvitorotroglus alpha, Creagonycha totani, Megasyringophilopsis aquilus, Meitingsunes columbicus, Neoaulonastus scirpaceus, Niglarobia calidridis, Paraniglarobia calidridis, Peristerophila accipitridicus, Picobia phoeniculi, Philoxanthornea dubinini, Syringophyloidus carpodaci, S. glandarii, S. klimovi, S. parapresentalis, S. presentalis, S. schoeniclusi, Syringophilopsis faini, S. fringillae, S. hirundus, S. spinolettus, S. sturni, S. turdi
Kyrghyzstan (Kirghizia)(Chirov & Kravtsova, 1995; Bochkov & Mironov, 1998; Bochkov, Mironov & Kravtsova, 2000; Bochkov, 2000; Skoracki, 2011; Skoracki et al., 2014, 2016) – Picobia chloris, Charadriphilus ralli, Creagonycha totana, Cuculisyringophilus chirovi, Mironovia phasiani, Picobia dyobatis, Rafapicobia melzeri, Syringophiloidus minor, S. presentalis, S. parapresentalis, Syringophilopsis kirgizorum, S. passericus, S. sturni, Syringophilus bipectinatus
Malaysia (Skoracki & Unsoeld, 2014) – Picobia mentalis
Mongolia (Skoracky, 2011) – Betasyringophyloidus phoenicurus, Neopicobia modularis, Torotrogla modularis
Nepal (Bochkov, Mironov & Skoracki, 2001; Skoracki, 2011; Skoracki et al., 2014; Skoracki, Unsoeld, Marciniak & Sikora, 2016; Skoracki, Sikora & Spicer, 2016; Skoracki et al., 2018; Kaszewska et al., 2020) – Aulobia [Syringophilopsis] stachyris, Bubophilus ascalaphus, Neosyringophilopsis garrulaxi, Peristerophila columbae, P. falcophila, Picobia dryobatis, P. echo, P. heeri
Pakistan (Skoracki, Sikora & Spicer, 2016; Skoracki, Hromada & Sikora, 2017) – Peristerophila meropis, Picobia phoeniculi
Philippines (Skoracky & OConnor, 2010; Skoracki, Sikora & Spicer, 2016) – Apodisyringophilus collocalius, Neoaulonastus chrysocolaptes, Picobia pycnonoti
Sri Lanka (Zmudzinski et al., 2019) – Neoaulonastus sidorchukae
Thailand (Skoracki et al., 2014) – Picobia heeri
S.E. Asia (Skoracki & Glowska, 2008; Fain, Bochkov & Mironov, 2000) – Aulobia anthreptes, Syringophiloidus graculae
East Asia (Fain, Bochkov & Mironov, 2000) – Syringophiloidus dendrocitta
NORTH AMERICA
Canada (Bochkov & Galloway, 2001, 2004; Skoracki, Flannery & Spicer, 2008, 2009; Zmudzinski et al., 2015; Kaszewska et al., 2020) – Ascetomylla [Syringophilopsis] porzanae, Betasyringophyloidus motacillae, Megasyringophilus aquilus, Neosyringophilopsis troglodytis, Paraniglarobia skorackii, Peristerophila columbae, Syringophiloidus minor, S. seiuri, S. zonotrichia, Syringophilopsis dendroicae, S. elongatus, S. icteri, S. tyranni
Mexico (Casto, 1977; Bochkov & Perez, 2002; Bochkov, Fain & Skoracki, 2004; Skoracki, 2008, 2017; Broda, Dabert & Glowska, 2016) – Aulonastus similis, Betasyringophiloidus seiuri, Calamincola lobatus, Crotophagisyringophilus io, Cuculisyringophilus crotophaginus, Megasyringophilus kethleyi, Neoaulobia mironovi, Peristerophila forpi, Syringophiloidus amazilia, S. daberti, Syringophilopsis trogoni, S. tyranni, Terratosyringophilus pioni
USA (Hankock, 1895, 1898; Banks, 1907; Ewing, 1911; Hirst, 1920; Rebrassier & Martin, 1932; Hwang, 1959; Brown, 1971; Casto, 1976, 1977, 1978, 1980a, 1980b; Clark, 1964; Kethley, 1970; Philips & Norton, 1978; Bochkov, Williams & Proctor, 2005; Skoracki, Dabert & Schmäschke, 2006; Skoracki, Flannery & Spicer, 2008; Bochkov, Flannery & Spicer, 2009; Skoracki, Hendricks & Spicer, 2010a, b, c; 2011; Glowska & Skoracki, 2010; Skoracky & OConnor, 2010; Bochkov, Skoracki, Hendricks & Spicer, 2011; Skoracki, Bochkov OConnor, 2011; Skoracki, Fajfer & Sikora, 2012; Skoracki, Spicer & OConnor, 2014, 2016; Skoracki, Sikora & Spicer, 2016) – Aulobia cardueli, A. dendroicae, A. erythroptera, Aulonastus emberizicus, A. euphagus, A. pirangus, A. sturnellus, Betasyringophyloidus motacillae, B. seiuri, Bochkovia phalaropi, Bubophilus ascalaphus, Calamincola lobatus, Peristerophila mucuya, Charadriineopicobia leucophaeus, Charadriphilus ralli, Chenophila branta, Ch. canduli, Columbiphilus khushalkhani, Creagonycha lara, Kalamotrypetes colinastes, Meitingsunes [Peristerophila] zenaidurae, Mironovia [Kalamotrypetes] pavodaptes, Neopicobia cardinalis, N. carpodacus, N. zumpti, Niglarobia ereuneti, N. helleri, Peristerophila columbae, P. mucuya, Phalarophilus fulicarius, Philoxanthornea anoa, Picisyringophilus kratos, Picobia dryobatis, P. psaltriparus, Rafapicobia toxostoma, Selenonycha baltoda, Syringophiloidus agelaius, S. bombycillae, S. carolae, S. daberti, S. delichonum, S. glandarii, S. jackowiaki, S. minor, S. molothrus, S. parapresentalis, S. presentalis, S. sialius, S. thryothorus, S. weiszii, S. xanthocephalus, S. zonotrichia, Syringophilopsis bartrami, S. catesbyi, S. certhiae, S. elongatus [= icteridae], S. empidonax, S. hylocichlae, S. icteri, S. mimidus, S. passerinae, S. polioptilus, S. sialiae, S. sittae, S. sturnellus, S. tyranni, S. wilsoni, Syringophilus bipectinatus, Terratosyringophilus longimana, Torotrogla aphelocoma, T. cardinalis, T. coccothraustes, T. cyanocitta, T. merulae, T. mima, T. piranga, T. villosa
Hawaiian Is. (Kethley, 1970; Schmid & Skoracki, 2007) – Procellariisyringophilus bulwerius
SOUTH and CENTRAL AMERICA
Argentina (Skoracki & Sikora, 2002, 2004a, 2008; Skoracki, 2008a; Skoracki & Glowska, 2008b; Sikora et al., 2011; Skoracki & Solarczyk, 2012; Sikora, Kaszewska & Skoracki, 2014; Kaszewska, Skoracki & Hromada, 2018) – Charadriphilus paraguaiae, Crotophagisyringophilus io, Gunabopicobia leptotila, G. metriopelia, Niglarobia helleri, Meitingsunes [Peristerophila] zenaidurae, Peristerophila mucuya, Picobia sp., Rafapicobia dendrocolaptesi, Syringophiloidus tarni, S. teledromas, Syringophilopsis turdi, Tinamiphilopsis elegans, Torotrogla mima
Bolivia (Skoracki, 2008a; Skoracki & Glowska, 2008) – Peristerophila claravis, Crotophagisyringophilus io
Brazil (Fain, Bochkov & Mironov, 2000; Bochkov & Fain, 2003; Bochkov, Fain & Skoracki, 2004; Pires & Daemon, 2007; Skoracki & Glowska, 2008b; Skoracki et al., 2010; Sikora et al., 2012; Skoracki, Sikora & Ozminski, 2012; Skoracki et al., 2014; Skoracki et al., 2016; Kaszewska & Skoracki, 2018; Kaszewska et al., 2020) – Charadriaulobia vanelli, Megasyringophilus kethleyi, Neoaulobia aratingae, Peristerophila mucuya, Picobia dryobatis, P. ictericus, Rafapicobia brotogeris, R. ramphastos, Psittaciphilus montanus, Stibarokris theristicus, Syringophiloidus microcerculus, Syringophilopsis bonariensis, S. tyranni, Tinamiphilopsis ariconte, Torotrogla synallaxis
Chile (Sikora, Kaszewska & Skoracki, 2014; Scoracki et al., 2014; Skoracki, Kaszewska & Kavetska, 2015) – Aulonastus pteroptochos, Rafapicobia melzeri, Syringophiloidus apus
Colombia (Fain, Bochkov & Mironov, 2000; Skoracki, 2008; Glowska & Skoracki, 2010; Skoracki & Sikora, 2011; Skoracki, Scibek & Sikora, 2012; Skoracki & Solarczyk, 2012; Skoracki, Kaszewska, Unsoeld & Skorupski, 2015; Hromada, Klimovičova, Unsöld & Skoracki, 2016; Kaszewska, Skoracki & Hromada, 2018; Kaszewska & Skoracki, 2018; Skoracki et al., 2018; Kaszewska et al., 2020) – Calamincola lobatus, Columbiphilus odontophoridus, Cuculisyringophilus crotophaginus, Galliphilopsis colinus, Gunabopicobia claravis, G. geotrygoni, Meitingsunes aldwelles, M. caprimulgus, Peristerophila claravis, P. falconis, P. mucuya, Picobia magellani, P. makoli, Pseudopicobia malacoptila, P. nonnula, Psittaciphilus amasonae, P. patagioenas, Rafapicobia lepidocolaptesi
Costa Rica (Sikora et al., 2012; 2013) – Henicorhina leucosticta, Syringophilopsis trogoni
Ecuador (Sikora et al., 2012; Skoracki & Solarczyk, 2012; Skoracki, Scibek & Sikora, 2012; Skoracki, Mironov & Unsoeld, 2013; Glowska & Laniecka, 2014; Skoracki, Zmudzinski & Sikora, 2020) – Aulonastus henicorhina, Picobia dryobatis, Picumnus sclateri, Pseudopicobia hapaloptila, Rafapicobia lepidocolaptesi, R. olszanowskii, Syringophiloidus quetzali
Guatemala (Skoracki, Bochkov & Wauthy, 2004; Skoracki, Mironov & Unsoeld, 2013) – Picobia ramphastos, Syringophiloidus quetzali
Guyana (Skoracki & Sikora, 2011; Glowska & Milensky, 2014; Glowska, Laniecka & Milensky, 2014) – Kalamotrypetes cracidus, Neopicobia herbicolae, Picobia myiopagi, P. ochoi, P. schmidti, Rafapicobia automoli
Martinique (Kaszewska, Skoracki & Hromada, 2018) – Gunabopicobia geotrygoni
Panama (Skoracki et al., 2014; Kaszewska & Skoracki, 2018; Skoracki, Mironov & Bermúdez, 2019; Kaszewska et al., 2020) – Neopicobia freya, Peristerophila claravis, P. columbae, Psittaciphilus montanus, Syringophilopsis bochkovi
Paraguay (Skoracki, 2008b; Skoracki & Sikora, 2011; Skoracki, Sikora & Ozminski, 2012; Skoracki & Solarczyk, 2012; Skoracki et al., 2014; Skoracki, Sikora & Spicer, 2016; Skoracki, Unsoeld, Marciniak & Sikora, 2016; Kaszewska, Skoracki & Hromada, 2018; Kaszewska et al., 2020) – Charadriphilus ralli, Cuculisyringophilus crotophaginus, Gunabopicobia geotrygoni, Kalamotrypetes cracidus, Neobubophilus cunicularius, Neopicobia freya, N. hepburni, Peristerophila claravis, P. mucuya, Picobia dryobatis, Rafapicobia dendrocolaptesi, Tinamiphilopsis ariconte
Peru (Glowska & Milensky, 2014; Glowska, 2014; Skoracki et al., 2014; Glowska & Laniecka, 2014; Kaszewska et al., 2020) – Neopicobia ea, Peristerophila mucuya, Picobia myiopagi, P. sayornis, P. schmidti, Picumnus olivaceus, Syringophilopsis szeffleri, S. tyranni
Suriname (Skoracki & Sikora, 2011, Skoracki et al., 2014; Kaszewska & Skoracki, 2018; Kaszewska et al., 2020) – Columbiphilus odontophoridus, Galliphilopsis colinus, Peristerophila columbae, P. mucuya, Picobia dryobatis, Psittaciphilus patagioenas
Trinidad & Tobago (Kaszewska & Skoracki, 2018; Skoracki et al., 2014; Kaszewska et al., 2020) – Peristerophila mucuya, Picobia dryobatis, Psittaciphilus montanus
Uruguay (Glowska, 2015a) – Stibarokris annae
Venezuela (Skoracki & Glowska, 2008; Skoracki & Solarczyk, 2012; Skoracki et al., 2014; Skoracki, Unsoeld, Marciniak & Sikora, 2016; Kaszewska, Skoracki & Hromada, 2018) – Gunabopicobia geotrygoni, Megasyringophilus dalmas, Neopicobia ea, Picobia dryobatis, Rafapicobia lepidocolaptesi, Terratosyringophilus geotrygonus
South America (Oudemans, 1904; Skoracki, 2008; Zmudzinski, Solarczyk & Skoradski, 2016) – Krantziaulonastus galbulicus, Niglarobia trouessarti, Stibarokris phoeniconaias
OCEANIA
Australia (Domrow, 1991; Skoracki, 2005b; Skoracki, Glowska & Sikora, 2008, 2011; Skoracki & Glowska, 2008c; Glowska & Laniecka, 2013; Glowska, 2013; Kaszewska, Skoracki & Hromada, 2018; Marciniak, Skoracki & Hromada, 2019; Kaszewska et al., 2020) – Galliphilopsis szeptyckii, Gunabopicobia lathami, Lawrencipicobia calyptorhyncha, Megasyringophilus cacatua, Meitingsunes ptilinopus, Neoaulobia cacatua, N. krafti, Neopicobia epthianura, N. glossopsitta, Neoaulonastus grewlingi, Peristerophila geopelis, P. leucomela, Picobia corcoracus, Syringophilopsis philemonis, Syringophilus bipectinatus
Johnston Atoll (Glowska 2015) – Stibarokris mariasi
Mariana Islands (Kaszewska, Kavetska & Skoracki, 2014) – Peristerophila lature
Midway Atoll (Kethley, 1970) – Philoxanthornea anoa
New Zealand (Marciniak, Skoracki & Hromada, 2019) – Peristerophila nestoriae
Papua New Guinea (Skoracki, 2005c; Skoracki & Sikora, 2008b; Skoracki, Glowska & Sikora, 2008; Kaszewska, Kavetska & Skoracki, 2014; Hromada, Klimovičova, Unsöld & Skoracki, 2016; Marciniak, Skoracki & Hromada, 2019; Kaszewska, Skoracki & Hromada, 2018; Skoracki, Hromada & Sikora, 2020) – Apodisyringiana mystaceae [New Britain], Aulobia nectariniae, Cuculisyringophilus chirovi, Gunabopicobia lathami, G. masalaje, Lawrencipicobia sulphurea, Meitingsunes ptilinopus, Neoaulobia cacatua, Neoperisterophila regiusi, Peristerophila coraciidus, P. lature, Picobia lemi, Terratosyringophilus reichholfi
Locality unknown (Skoracki & Dabert, 2002) – Syringophilopsis garrulus
Aves
Accipitriformes
Fam. Accipitridae
Accipiter francesiae – Peristerophila accipitridicus
A. nisus – Megasyringophilopsis aquilus, Peristerophila accipitridicus
A. rapax – Megasyringophilus aquilus
Buteo jamaicensis – Peristerophila columbae
B. lagopus – Peristerophila accipitridicus
Circaetus gallicus – Peristerophila accipitridicus
Clanga pomarina – Megasyringophilus aquilus
Hieraaetus pennatus – Megasyringophilus aquilus
Terathopius ecaudatus – Peristerophila accipitridicus
Anseriformes
Fam. Anatidae
Anas clypeata – Chenophila canduli
A. platyrhynchos – Chenophila platyrhynchos
A. querquedula – Chenophila canduli
Branta canadensis – Chenophila branta
B. leucopsis – Chenophila nanseni
Marmaronetta angustirostris – Chenophila marmaronetta
Apodiformes
Fam. Apodidae
Apus melba – Syringophiloidus apus
Collocalia esculenta – Apodisyringophilus collocalius
Cypsiurus parvus – Syringophiloidus cypsiuri
Hirundapus caudacutus – Apodisyringiana hirundapi
Fam. Hemiprocnidae
Hemiprocne comata – Apodisyringiana haszprunari
H. mystacea mystacea – Aposyringiana mystaceae
Fam. Trochilidae
Amazilia candida – Syringophiloidus amazilia
Bucerotiformes
Fam. Bucerotidae
Horizocerus hartlaubi – Syringophiloidus bucerotidus
H. albocristatus – Syringophiloidus bucerotidus
Lophoceros pallidirostris – Syringophiloidus bucerotidus
Tockus flavirostris – Syringophiloidus bucerotidus
T. fasciatus – Syringophiloidus bucerotidus
T. deckeni – Syringophiloidus bucerotidus
Caprimulgiformes
Fam. Caprimulgidae
Chordeiles minor – Meitingsunes caprimulgus
Charadriiformes
Fam. Charadriidae
Charadrius w. wilsonia – Selenonycha baltoda
Ch. placidus – Selenonycha charadriiformicus
Pluvialis apricaria – Charadriineopicobia apricaria
P. dominica – Selenonycha baltoda
Vanellus albiceps – Niglarobia vanelli
V. chilensis – Charadriphiaulobia vanelli
V. coconatus – Niglarobia vanelli
V. crassirostris – Niglarobia vanelli
V. gregarius – Niglarobia vanelli
V. lugubris – Niglarobia vanelli
V. senegallus – Niglarobia vanelli
V. spinosus – Niglarobia vanelli
V. vanellus – Niglarobia vanelli
Fam. Glareolidae
Cursorius temminckii – Niglarobia cursoriae
Smutsornis [Rhinoptilus] africanus – Niglarobia rhinoptili
Fam. Laridae
Chroicocephalus genei – Selenonycha charadriiformicus
Ch. ridibundus – Selenonycha charadriiformicus
Larus californicus – Selenonycha baltoda
L. canus – Philoxanthornia clarki
L. delawarensis – Creagonycha lara
Leucophaeus atricilla – Charadriineopicobia leucophaeus, Selenonycha baltoda
Subfam. Sterninae
Anous tenuirostris – Philoxanthornia anoa
Chlidonias leucopterus – Philoxanthornia dubinini
Ch. niger – Selenonycha baltoda
Gelochelidon nilotica – Kethleyana gelochelidoni, Philoxanthornia dubinini
Sterna hirundo – Philoxanthornia clarki
Sternula albifrons – Philoxanthornia clarki, Creagonycha sternae
Fam. Scolopacidae
Arenaria interpres – Niglarobia helleri
Calidris alba – Charadriineopicobia calidris, Creagonycha totani
C. himantopus – Creagonycha totani
C. minuta – Creagonycha totana, Paraniglarobia calidridis
C. pugnax – Selenonycha charadriiformicus
C. pusilla – Niglarobia ereuneti
C. temminckii – Creagonycha totani
Galinago gallinago – Aulobia galinago, Charadriphilus galinago
G. paraguaiae – Charadriphilus paraguaiae
Limosa lapponica – Niglarobia lapponica, Charadriineopicobia risbowi
Phalaropus fulicarius – Bochkovia phalaropi, Phalarophilus fulicarius
Philomachus pugnax – Selenonycha charadriiformicus
Scolopax rusticola – Charadriphilus lyudmilae
Tringa flavipes – Niglarobia helleri
T. glareola – Creagonycha totani
T. melanoleuca – Niglarobia skorackii
T. ochropus – Niglarobia helleri
T. stagnatilis – Niglarobia helleri
T. totanus – Creagonycha totani, Niglarobia helleri
Ciconiiformes
Fam. Ciconiidae
Ciconia alba – Stibarokris langei
C. maguari – Stibarokris annae
Coliiformes
Fam. Coliidae
Colius striatus – Colisyringophilus tanzanicus
Urocolius macrourus – Colisyringophilus tanzanicus
Columbiformes
Fam. Columbidae
Chalcophaps indica – Meitingsunes chalcophaps
Chordeiles minor – Meitingsunes caprimulgus
Claravis pretiosa – Peristerophila claravis, Gunabopicobia claravis
Columba arguatrix – Peristerophila columbae
C. guinea – Peristerophila columbae
C. leucomela – Peristerophila leucomela
C. leuconota – Peristerophila columbae
C. livia – Columbiphilus khushalkhani, Meitingsunes columbicus, M. zenaidurae, Gunabopicobia zumpti, Peristerophila columbae
C. oenas – Meitingsunes [Peristerophila] zenadourae, Peristerophila columbae
C. palumbus – Meitingsunes columbicus, Peristerophila columbae
C. trocaz – Peristerophila columbae
Columbina minuta – Peristerophila mucuya
C. passerina – Peristerophila mucuya
C. squammata – Peristerophila mucuya
C. talpacoti – Peristerophila mucuya
Ducula luctuosa – Gunabopicobia masalaje, Peristerophila lature
D. pistrinaria – Gunabopicobia masalaje
D. rosacea – Gunabopicobia masalaje
D. rufigaster – Gunabopicobia masalaje
D. spilorrhoa – Gunabopicobia masalaje, Peristerophila lature
Geopelia cuneata – Peristerophila geopelis
G. placida – Peristerophila geopelis
G. striata – Peristerophila geopelis
Geophaps plumifera – Peristerophila mucuya
Geotrygon chiriquensis – Peristerophila columbae
G. chrysie – Gunabopicobia geotrygoni
G. frenata – Gunabopicobia geotrygoni, Meitingsunes aldwelles
G. linearis – Terratosyringophilus geotrygonus, Gunabopicobia geotrygoni
G. montana – Gunabopicobia geotrygoni, Psittaciphilus montanus
Leptotila verreauxi – Gunabopicobia leptotila
Leucosarcia melanoleuca – Gunapicobia lathami
Metriopelia ceciliae – Peristerophila mucuya
M. melanoptera – Peristerophila mucuya, Gunabopicobia metropelia
Ocyphaps lophotes – Peristerophila geopelis
Patagioenas fasciata – Psittaciphilus patagioenas
P. speciosa – Psittaciphilus patagioenas, Peristerophila columbae
Petrophassa plumifera – Peristerophila mucuya
Ptilinopus iozonus – Gunabopicobia masalaje
P. jambu – Peristerophila lature
P. magnificus – Meitingsunes ptilinopus
P. melanospilus – Peristerophila lature
P. porphyreus – Peristerophila lature
P. regina – Peristerophila lature
P. rivoli – Meitingsunes ptilinopus
Stigmatopelia senegalensis – Gunabopicobia zumpti
Streptopelia capicola – Gunabopicobia zumpti, Peristerophila columbae
S. decaocto – Peristerophila columbae
S. decipiens – Peristerophila columbae
S. orientalis – Peristerophila columbae
S. semitorquata – Peristerophila columbae
S. tranquebarica – Peristerophila columbae
S. turtur -Peristerophila columbae
Treron waalia – Meitingsunes columbicolus
Turacoena manadensis – Meitingsunes turacoenas
T. modesta – Meitingsunes turacoenas
Turtur tympanistria – Meitingsunes [Peristerophila] tympanistria
Zenaida auriculata – Meitingsunes [Peristerophila] zenadourae, Peristerophila columbae
Z. asiatica – Meitingsunes [Peristerophila] zenadourae, Terratosyringophilus longisoma
Z. macroura – Gunabopicobia zumpti, Meitingsunes [Peristerophila] zenadourae,
Terratosyringophilus longisoma ?
Coraciiformes
Fam. Alcedinidae
Todiramphus chloris – Peristerophila meropis
Fam. Brachypteracidae
Atelornis crossleyi – Rafapicobia brachypteraci
Fam. Meropidae
Melittophagus bullocki – Syringophilopsis melittophagi
Merops albicollis – Syringophilopsis albicollisi
Fam. Momotidae
Baryphthengus ruficapillus – Rafapicobia momoti
Cuculiformes
Fam. Cuculidae
Centropus goliath – Cuculisyringophilus chirovi
Cercococcyx montanus – Cuculisyringophilus chirovi
Cuculus canorus – Cuculisyringophilus chirovi
Crotophaga ani – Crotophagisyringophilus io
C. solitarius – Cuculisyringophilus chirovi
C. sulcirostris – Cuculiphilus lobatus, Crotophagisyringophilus sulcirostris, Cuculisyringophilus crotophaginus
Guira guira – Cuculisyringophilus crotophaginus
Pachycoccyx audeberti – Cuculisyringophilus chirovi
Scythrops novaehollandiae – Cuculisyringophilus chirovi
Falconiformes
Fam. Falconidae
Falco cherrug – Peristerophila falcophila
F. peregrinus – Peristerophila falcophila
F. sparverius – Peristerophila falcophila
Galbuliformes
Fam. Bucconidae
Hapaloptila castanea – Pseudopicobia hapaloptila
Malacoptila panamensis – Pseudopicobia malacoptila
Nonnula frontalis – Pseudopicobia nonnula
Fam. Galbulidae
Jacamaralcyon tridactyla – Krantziaulonastus galbulicus
Galliformes
Fam. Cracidae
Ortalis canicollis – Kalamotrypetes cracidus
Penelope sp. – Kalamotrypetes cracidus
Fam. Odontophoridae
Calipepla squamata – Colinophilus wilsoni, Kalamotrypetes colinastes
Colinus cristatus – Columbiphilus odontophoridus, Galliphilopsis colinus
C. leucopogon – Columbiphilus odontophoridus, Galliphilopsis colinus
C. virginianus – Colinophilus wilsoni, Kalamotrypetes colinastes
Fam. Phasianidae
Alectoris barbara – Gallophilopsis szeptickii [publ. as «bochkovi»]
A. rufa – Columbiphilus alectoris, Syringophilus bipectinatus
[Alectoris sp]. – Columbiphilus alectoris, Picobia alectori
Coturnix chinensis – Galliphilopsis szeptyckii
C. coturnix – Columbiphilus khushalkhani, Galliphilopsis szeptyckii, Mironovia coturnae
C. pectoralis – Galliphilopsis szeptyckii
Francolinus levaillantoides – Gallophilopsis francolinus
F. clappertoni – Columbiphilus khushalkhani
Gallus gallus domestica – Columbiphilus khushalkhani (=polonica), Syringophilus bipectinatus
G. sonneratii – Columbiphilus khushalkhani (=polonica)
Lagopus lagopus – Mironovia lagopus
L. muta – Mironovia lagopus
Lophura leucomelanos – Gallophilopsis lophurus
Phasianus colchicus – Mironovia phasiani
Ptilopachus petrosus – Syringophilus bipectinatus
Rollulus rouloul – Mironovia rouloul
Tragopan sp. – Gallophilopsis bochkovi
Meleagris gallopavo [pavodaptes] – Mironovia [Kalamotrypetes] pavodaptes
Fam. Numididae
Acryllium vulturinum – Columbiphilus khushalkhani
Numida meleagris –Syringophilus numidae
Numida meleagris mitratus – Columbiphilus khushalkhani
Gruiformes
Fam. Aramidae
Aramus guarauna (= scolopaceus) – Niglarobia trouessarti
Fam. Rallidae
Fulica americana – Charadriphilus ralli
Gallinula chloropus – Ascetomylla gallinula
G. melanops – Charadriphilus ralli
Laterallus melanophaius – Charadriphilus ralli
Pardirallus sanguinolentus – Rafapicobia melzeri
Rallus aquaticus – Charadriphilus ralli, Rafapicobia melzeri
Porzana carolina – Ascetomylla porzanae
P. parva – Charadriphilus ralli, Rafapicobia melzeri
P. porzana – Rafapicobia melzeri
Musophagiformes
Fam. Musophagidae
Musophaga rossae – Blaszakia rossae
Tauraco livingstoni – Blaszakia tauracos
T. schalowi – Blaszakia tauracos
Otidiformes
Fam. Otididae
Chlamydotis undulata – Pteroclidisyringophilus otididus
Passeriformes
Fam. Acanthizidae
Acanthiza inornata – Neosyringophilopsis acanthizus
Pyrrholaemus sagitatus – Neopicobia pyrrholaemus
Fam. Acrocephalidae
Acrocephalus arundinaceus – Neoaulonastus bisetatus
A. baeticatus – Neoaulonastus bisetatus
A. dumetorum – Neoaulonastus bisetatus (= scirpaceus)
A. paludicola – Neoaulonastus bisetatus, Picobia paludicola
A. palustris – Neoaulonastus bisetatus
A. schoenobaenus – Syringophilopsis acrocephali
A. scirpaceus – Neoaulonastus bisetatus (= scirpaceus), Syringophilopsis acrocephali
Iduna caligata – Neoaulonastus caligatus, Syringophilopsis idunae
Fam. Aegithalidae
Aegithalos caudatus – Neoaulonastes aegithalos, Neosyringophilopsis aegythali, Picobia caudati
Psaltriparus minimus – Picobia psaltriparus
Fam. Alaudidae
Calandrella brachydactyla – Syringophilopsis davidi
Eremophila alpestris – Torotrogla lullulae
Galerida cristata – Picobia galerida
Lullula arborea – Torotrogla lullulae
Fam. Artamidae
Artamis fuscus – Syringophiloidus artamis
A. leucorhynchus – Syringophiloidus artamis
Fam. Bernieridae
Bernieria madagascariensis – Picobia bernieriarum
Xanthomixis zosterops – Picobia bernieriarum
Fam. Bombycillidae
Bombycilla cedrorum – Syringophiloidus bombycillae
B. garrulus – Syringophiloidus bombycillae
Fam. Calcariidae
Calcarius lapponicus – Torotrogla calcarius
Fam. Cardinalidae
Cardinalis cardinalis – Neopicobia cardinalis, Syringophiloidus carolae, Torotrogla cardinalis
Habia fuscicauda – Aulonastus similis
H. rubica – Aulonastus similis
Passerina ciris – Syringophiloidus daberti, Syringophilopsis passerinae
. cyanea – Syringophilopsis passerinae, Syringophiloidus carolae
Piranga ludoviciana – Aulonastus pirangus, Torotrogla piranga
Fam. Certhiidae
Certhia americana – Syringophilopsis certhiae
Fam. Cettiidae
Cettia cetti – Picobia cetti
Fam. Cinclidae
Cinclus mexicanus – Syringophiloidus cincli
Fam. Cisticolidae
Apalis porphyrolaema – Neoaulonastus apalis
Cisticola cantans – Aulobia cisticolae
C. chubbi – Aulobia cisticolae
C. erythrops – Aulobia cisticolae
C. hunteri – Aulobia cisticolae
C. lateralis – Aulobia cisticolae
Neomixis tenella – Picobia neomixis
Prinia [Heliolais] erythroptera – Aulobia erythroptera
Fam. Conopophagidae
Conopophaga aurita – Rafapicobia milenskyi
Fam. Corcoracidae
Corcorax melanorhamphos – Picobia corcoracus
Fam. Corvidae
Aphelocoma californica – Torotrogla aphelocoma
Cissa chinensis – Picobia cissa
Corvus brachyrhynchos – Syringophilus bipectinatus, Syringophiloidus glandarii
C. frugilegus – Corvitorotroglus alpha, Syringophiloidus glandarii
C. monedula – Syringophiloidus glandarii
Cyanocitta stelleri – Syringophiloidus glandarii [also published as S. presentalis], Torotrogla cyanocitta
Dendrocitta vagabunda [rufa rufa] – Syringophiloidus dendrocittae
Garrulus glandarius – Syringophiloidus glandarii, S. presentalis, Syringophilopsis garrulus
Nucifraga caryocatactes – Syringophilopsis nucifragus
Pica pica – Syringophiloidus glandarii
Urocissa erythrorhyncha – Picobia cissa
Fam. Dicruridae
Syringophilopsis dicruri – Dicrurus adsimilis
Fam. Emberizidae
Ammodramus savanarum – Aulonastus emberizicus
Emberiza elegans – Torotrogla volgini
E. rutila – Betasyringophiloidus schoeniclus
E. schoeniclus – Betasyringophiloidus schoeniclus
E. tristrami – Betasyringophiloidus schoeniclus
Passerculus sandwichensis – Aulonastus emberizicus
Tachyphonus rufus – Syringophiloidus stawarczyki
Fam. Estrildidae
Erythrura trichroa – Syringophiloidus erythrurus
Estrilda astrild – Neoaulonastus lonchurus, N. oryzivorus
E. atricapilla – Neoaulonastus lonchurus, N. oryzivorus, Picobia lonchurae
E. nonnula – Neoaulonastus lonchurus
E. erythronotus – Neoaulonastus lonchurus
E. rhodopyga – Neoaulonastus lonchurus, Syringophiloidus estrildus
E. troglodytes – Neoaulonastus lonchurus
Granatina ianthinogaster – Neoaulonastes granatina
Lagonosticta rufopicta – Syringophilopsis lagonostictus
L. senegala – Syringophilopsis lagonostictus
Lonchura leucogastra – Picobia lonchura
L. oryzivora – Neoaulonastus oryzivorus
L. punctulata – Fritschsyringophilus lonchurae, Neoaulonastus lonchurus
Lonchura sp. – Syringophiloidus erythrurus
Spermestes cucullatus [Lonchura cucullata] – Neosyringophilopsis lonchurus
Fam. Falcunculidae
Falcunculus frontatus – Neoaulonastus grewlingi
Fam. Fringillidae
Acanthis flammea – Aulobia cardueli
Carduelis cannabina – Syringophilopsis kirgizorum, Torotrogla cardueli
C. cardinalis – Aulobia cardueli
C. carduelis – Aulobia cardueli, Syringophilopsis kirgizorum, Torotrogla cardueli
C. citrinella – Aulobia cardueli
C. flavirostris – Aulobia cardueli
Carpodacus erythrinus – Syringophiloidus carpodaci
Chloris chloris – Picobia chloris, Syringophilopsis kirgizorum, Syringophiloidus klimovi
Coccothraustes coccothraustes – Syringophiloidus coccothraustes
C. vespertinus – Torotrogla coccothraustes
Euphonia cyanocephala – Syringophiloidus stawarczyki
Fringilla coelebs – Syringophilopsis fringillae, Torotrogla gaudi, Aulonastus fringillus
F. montifringilla – Syringophilopsis fringillae, Torotrogla gaudi
Haemorhous purpureus – Neopicobia carpodacus
Leucosticte arctoa – Aulobia leucostictus
Linurgus olivaceus – Syringophilopsis kirgizorum
Loxia curvirostra – Torotrogla cardueli, Aulonastus loxius
L. leucoptera – Torotrogla cardueli
L. pytiopsittacus – Torotrogla cardueli
Pyrrhula pyrrhula – Neopicobia modularis, Torotrogla gaudi
Rhodospiza obsoleta – Syringophilopsis kirgizorum
Serinus canaria – Torotrogla cardueli
S. capistratus – Syringophiloidus minor
S. mozambicus – Syringophiloidus serini
Spinus psaltria – Aulobia cardueli
S. spinus – Aulobia cardueli, Torotrogla cardueli
Fam. Furnariidae
Aphrastura spinicauda – Picobia sp.
Automolus ochralaemus – Rafapicobia automoli
A. rufipileatus – Rafapicobia automoli
Dendrocolaptes platyrostris – Rafapicobia dendrocolaptesi
D. picumnus – Rafapicobia dendrocolaptesi
Lepidocolaptes affinis – Rafapicobia lepidocolaptesi
L. souleyetii – Rafapicobia lepidocolaptesi
Synallaxis spixi – Torotrogla synallaxis
Xiphocolaptes promeropirhynchus – Picobia makoli
Fam. Hirundinidae
Delichon urbicum – Syringophiloidus delichonum, S. glandarii
Hirundo rustica – Syringophiloidus delichonum, S. hirundinis, Syringophilopsis fringillae, S. rusticus
Riparia riparia – Picobia riparius, Syringophilopsis rusticus
Fam. Icteridae
Agelaius phoeniceus – Syringophiloidus agelaius, Syringophilopsis elongatus [= S. icteridae]
Cacicus chrysopterus – Picobia ictericus
Euphagus carolinus – Syringophilopsis elongatus
E. cyanocephalus – Aulonastus euphagus, Syringophilopsis elongatus
Icterus bullockii – Syringophilopsis icteri
I. galbula – Syringophiloidus molothrus, Syringophilopsis icteri
I. pustulatus – Betasyringophiloidus seiuri, Syringophilopsis icteri
I. spurius – Syringophilopsis icteri
Molothrus ater – Syringophiloidus molothrus
M. bonariensis – Syringophilopsis bonariensis
Pheucticus melanocephalus – Syringophilopsis wilsoni
Pseudoleistes guirahuro – Picobia ictericus
Quiscalus mexicanus – Syringophilopsis elongatus
Q. quiscula – Syringophilopsis elongatus
Sturnella magna – Aulonastus sturnellus
S. neglecta – Syringophilopsis sturnellus
Xanthocephalus xanthocephalus – Syringophiloidus xanthocephalus
Fam. Laniidae
Corvinella corvina – Syringophillopsis corvinae
Lanius excubitor – Syringophiloidus weiszii
L. ludovicianus – Syringophiloidus weiszii
L. minor – Syringophilopsis kristini
L. senator – Aulonastus lanius
Lanius sp. – Syringophilopsis yosefi
Fam. Leiothrichidae
Trochalopteron formosum [Garrulax formosus] – Picobia dziabaszewskii
Trochalopteron affine [Garrulax affinis] – Neosyringophilopsis garrulaxi
Turdoides jardineii – Picobia dziabaszewskii
T. leucopygia – Picobia dziabaszewskii
T. rubiginosa – Picobia dziabaszewskii
Fam. Locustellidae
Locustella luscinioides – Neosyringophilopsis locustellus, Neopicobia locustella
L. naevia – Neosyringophilopsis locustellus, Neopicobia locustella
Fam. Macrosphenidae
Sylvietta virens flaviventris – Aulobia virens
S. whytii johnstoni – Aulobia sylviettae
Fam. Malaconotidae
Laniarius aethiopicus – Neoaulonastus malaconotus
Nilaus afer– Neoaulonastus malaconotus
Telophorus nigrifrons– Neoaulonastus malaconotus
Fam. Meliphagidae
Epthianura aurifrons – Neopicobia epthianura
Philemon citreogularis – Syringophilopsis philemonis
Fam. Mimidae
Dumetella carolinensis – Syringophilopsis mimidus
Margarops fuscatus – Syringophilopsis mimidus
Mimus patagonicus – Torotrogla mima
M. polyglottos – Torotrogla mima
M. triurus – Torotrogla mima
Toxostoma curvirostre – Rafapicobia toxostoma
Fam. Monarchidae
Terpsiphone mutata – Pipicobia terpsiphone
T. rufiventer – Pipicobia terpsiphone, Syringophiloidus furthi
Fam. Motacillidae
Anthus cervinus – Aulonastes anthus, Neopicobia anthi
A. pratensis – Syringophilopsis blaszaki, Torotrogla modularis
A. spinoletta – Syringophilopsis spinolettus
A. trivialis – Aulonastes anthus, Neopicobia anthi, Syringophilopsis blaszaki
Motacilla alba – Aulonastus albus
M. flava – Aulonastus albus, Syringophiloidus motacillae
Fam. Muscicapidae
Cichladusa arquata – Picobia cichladusa
Cossypha heuglini – Picobia echo
Copsychus malabricus – Picobia echo
C. saularis – Picobia echo
Erithacus rubecula – Torotrogla rubeculi
Ficedula hypoleuca – Rafapicobia zirnitra, Syringophilopsis kazmierski
F. parva – Syringophilopsis kazmierski
Luscinia akahige – Rafapicobia zirnitra
L. cyanura – Rafapicobia zirnitra
L. megarhynchos – Torotrogla lusciniae
L. sibilans – Rafapicobia zirnitra
L. svecica – Torotrogla lusciniae
Muscicapa dauurica (= latirostris) – Syringophilopsis muscicapus
Myrmecocichla arnotti – Picobia myrmecocichla
Phoenicurus auroreus – Rafapicobia zirnitra
Ph. moussieri – Picobia phoenicuri
Ph. ochruros – Betasyringophyloidus phoenicurus
Ph. phoenicurus – Syringophilopsis kazmierski
Saxicola rubetra – Betasyringophyloidus saxicolus, Rafapicobia zirnitra
Fam. Nectariniidae
Anthreptes longuemarei – Aulobia afroanthreptes, Neoaulonastus cinnyris
A. malacensis – Aulobia anthreptes
A. neglectus – Aulobia afroanthreptes, Neoaulonastus cinnyris
A. reichenowi – Aulonastus nectariniphilus
Cinnyris mediocris – Neoaulonastes cinnyris
C. shelleyi – Aulobia nectariniae
C. chalybeus – Picobia oritis
C. erythrocercus – Picobia oritis
C. jugularis – Aulobia nectariniae
C. mariquensis – Aulobia nectariniae
C. oustaleti – Picobia oritis
C. regius – Aulobia nectariniae
C. shelleyi – Aulobia nectariniae
C. talatala – Picobia oritis
C. venustus – Picobia oritis
Cyanomitra olivacea – Picobia oritis
C. oritis – Picobia oritis
C. verreauxi – Picobia oritis
Hedydipna collaris – Picobia hedydipna
Leptocoma sericea – Aulobia nectariniae
L. zeylonica – Neoaulonastus sidorchukae
Nectarinia takazze – Aulonastus nectariniphilus
Fam. Oriolidae
Oriolus chinensis – Syringophiloidus oriolus
O. oriolus – Syringophiloidus oriolus
Fam. Panuridae
Panurus biarmicus – Picobia biarmicus
Fam. Paradisaeidae
Cicinnurus regius – Neoperisterophila regiusi
Manucodia ater – Picobia lemi
M. chalybatus – Picobia lemi
Fam. Paridae
Parus major – Neopicobia pari
Periparus venustulus – Neopicobia pari
Poecile carolinensis – Syringophiloidus jackowiaki
P. palustris – Syringophilopsis palustris
P. varius – Neopicobia pari
Fam. Parulidae
Geothlypis philadelphia – Betasyringophiloidus seiuri
Helmitheros vermivorum – Betasyringophiloidus seiuri
Oreothlypis peregrina – Betasyringophiloidus seiuri
O. [Vermivora] rufiapilla – Betasyringophiloidus seiuri
Parkesia noveboracensis – Betasyringophiloidus seiuri
Phainopepla nitens – Torotrogla villosa
Protonotaria citrea – Syringophilopsis dendroicae
Seiurus aurocapillus – Betasyringophiloidus seiuri
Setophaga coronata – Aulobia dendroicae, Syringophilopsis dendroicae
S. carulescens – Syringophilopsis dendroicae
S. [Dendroica] graciae – Syringophilopsis dendroicae
S. [D.] nigrescens – Aulobia dendroicae
S. palmarum – Betasyringophiloidus seiuri
S. pensylvanica – Betasyringophiloidus seiuri
S. petechia – Syringophilopsis dendroicae
Cardellina [Wilsonia] pusilla – Neopicobia troglodytes, Syringophilopsis dendroicae
Fam. Passerellidae
Melospiza georgiana – Betasyringophiloidus seiuri
M. lincolnii – Betasyringophiloidus seiuri
M. melodia – Betasyringophiloidus seiuri
Pipilo erythrophthalmus – Aulonastus pipili
Spizella breweri – Syringophiloidus auduboni, S. sialius
Zonotrichia albicolis – Betasyringophiloidus seiuri, Syringophiloidus zonotrichia
Z. atricapilla – Aulonastus emberizicus
Fam. Passeridae
Passer domesticus – Krantziaulonastus dubinini, Picobia passeri, Syringophiloidus minor, Syringophilopsis hunanensis, S. passericus
P. hispaniolensis – Syringophiloidus minor
P. montanus – Syringophilopsis fringillae, Syringophiloidus minor, S. montanus
Petronia petronia – Syringophiloidus petronicus
Fam. Pellorneidae
Illadopsis rufipennis – Picobia illadopsae
Fam. Phylloscopidae
Phylloscopus collybita – Neosyringophilopsis phylloscopi
Ph. trochilus – Neosyringophilopsis phylloscopi
Fam. Pipridae
Corapipo altera – Syringophilopsis bochkovi
Manacus vitellinus – Syringophilopsis bochkovi
Fam. Ploceidae
Dinemellia dinemelli – Krantziaulonastus yoyomi, Picobia dinemellia
Euplectes axillaris – Neoaulonastus tanzanicus
Hyphanturgus nigricollis brachypterus – Syringophilopsis nitens
Malimbus nitens – Syringophilopsis nitens
Ploceus xanthops – Betasyringophiloidus seiuri
P. aurantius – Syringophiloidus plocei
P. cucullatus – Syringophiloidus plocei
P. nelicourvi – Neoaulonastus tanzanicus
P. nigerrimus – Syringophiloidus plocei
P. nigricollis – Syringophilopsis nitens
P. ocularis – Picobia ploceus
Ploceus baglafecht [Othyphantes stuhlmanni] – Betasyringophiloidus seiuri
Ploceus [Pachyphantes] superciliosus – Betasyringophiloidus seiuri
P. vitellinus – Syringophilopsis nitens
Pseudonigrita arnaudi – Syringophiloidus pseudonigritae
Quelea erythrops – Betasyringophiloidus seiuri, Syringophilopsis nitens
Q. quelea – Neoaulonastus quelea
Fam. Polioptilidae
Polioptila caerulea – Syringophilopsis polioptilus
Fam. Prunellidae
Prunella atrogularis – Neopicobia modularis
P. modularis – Aulonastus prunellae, Neopicobia modularis, Torotrogla modularis
Fam. Ptilogonatidae
Phainopepla nitens – Torotrogla villosa
Fam. Pycnonotidae
Chlorocichla flaviventris – Syringophilopsis veselovsky
Eurillas latirostris – Syringophilopsis veselovsky
Pycnonotus barbatus – Picobia pycnonoti, Syringophilopsis veselovsky
P. goiavier – Picobia pycnonoti
P. xanthopygos – Picobia pycnonoti, Torotrogla pycnonotus
Fam. Remizidae
Remiz pendulinus – Neoaulonastus remizus
Fam. Rhinocriptidae
Pteroptochos tarnii – Syringophiloidus tarnii
Scytalopus magellanicus – Picobia magellani
Teledromas fuscus – Syringophiloidus teledromas
Fam. Sittidae
Sitta carolinensis – Syringophilopsis sittae
S. europaea – Syringophilopsis sittae
Fam. Sturnidae
Aplonis panayensis – Picobia indonesiana
Enodes erythrophris – Picobia indonesiana
Gracula religiosa – Syringophiloidus graculae
Lamprotornis superbus – Picobia lamprotornis
Mino dumontii – Picobia indonesiana
Sturnus cineraceus – Picobia sturni
S. unicolor – Picobia sturni
S. vulgaris – Krantziaulonastus buczekae Picobia sturni, Syringophiloidus presentalis, Syringophilopsis sturni
Fam. Sylviidae
Sylvia atricapilla – Aulobia sylviae, Neoaulonastus bisetatus
S. borin – Syringophilopsis borini
S. communis – Neoaulonastus bisetatus
S. curruca – Aulobia sylviae, Neoaulonastus bisetatus, Picobia currucae, Syringophilopsis faini
S. hortensis – Picobia currucae
S. nisoria – Aulobia sylviae, Picobia currucae
Fam. Thamnophilidae
Hypocnemis cantator – Rafapicobia thamnophili
Myrmeciza ferruginea – Rafapicobia thamnophili
Myrmoborus leucophrys – Rafapicobia thamnophili
Myrmotherula longipennis – Rafapicobia thamnophili
Pygiptila stellaris – Phipicobia pygiptilae
Thamnophilus insignis – Rafapicobia thamnophili
Fam. Thraupidae
Dacnis cayana – Syringophiloidus stawarczyki
Emberizoides herbicola – Neopicobia herbicolae
Sicalis luteola – Syringophilopsis emberizae
Sporophila torqueola – Syringophiloidus sporophila
Thamnophilus rufus – Syringophiloidus stawarczyki
Fam. Timaliidae
Cyanoderma [Stachyris] pyrrhops – Syringophilopsis stachyris
Fam. Troglodytidae
Campylorhynchus brunneicapillum – Syringophiloidus sp.
Henicorhina leucosticta – Aulonastus henicorhina
H. leucophrys – Picobia troglodytidus
Microcerculus marginatus – Syringophiloidus microcerculus
Thryomanes bewickii – Neopicobia troglodytes
Thryothorus ludovicianus – Syringophiloidus thryothorus
Troglodytes aedon – Neopicobia troglodytes, Neosyringophilopsis troglodytis
T. troglodytes – Neosyringophilopsis troglodytis
Fam. Turdidae
Catharus [= Hylocichla] fuscescens – Syringophilopsis hylocichlae
Copsychus albospecularis – Rafapicobia copsychus
Hylocichla mustelina – Picobia hylocichlae
Sialia mexicana – Syringophiloidus sialius
Turdus albicollis – Syringophilopsis turdi
T. atrogularis – Syringophiloidus parapresentalis, Syringophilopsis turdi
T. iliacus – Syringophiloidus parapresentalis, Syringophilopsis turdi
T. merula – Torotrogla merulae, Syringophiloidus parapresentalis
T. migratorius – Syringophiloidus parapresenalis, S. presentalis, Syringophilopsis turdi, Torotrogla merulae
T. pilaris – Syringophiloidus parapresentalis, Syringophilopsis turdi
T. philomelos – Syringophiloidus philomelosus, Syringophilopsis turdi, Torotrogla merulae
T. torquatus – Torotrogla merulae
T. viscivorus – Torotrogla merulae
Fam. Tyrranidae
Elaenia ruficeps – Picobia myiopagi
Empidonax hammondii – Syringophilopsis empidonax
E. wrightii – Syringophilopsis empidonax
Hemitriccus margariraceiventer – Picobia schmidti
Lophotriccus pileatus – Picobia schmidti, Syringophilopsis szeffleri
Myiozetetes similis – Aulonastus similis
Sayornis nigricans – Picobia sayornis
Tolmomyias sulphurescens – Picobia ochoi
Tyrranus tyrranus – Syringophilopsis tyranni
T. dominicensis – Syringophilopsis tyranni
T. forficatus – Syringophilopsis tyranni
T. verticalis – Syringophilopsis tyranni
Myiarchus cinerascens – Syringophilopsis tyranni
M. crinitus – Syringophilopsis tyranni
Myiopagis gaimardii – Picobia myiopagi
M. subplacens – Picobia myiopagi, Syringophilopsis tyranni
Satrapa icterophrys – Syringophilopsis tyranni
Fam. Vangidae
Prionops scopifrons – Neoaulonastus prionops, Picobia meru
Fam. Vireonidae
Vireo hypochryseus – Syringophilopsis catesbyi
V. olivaceus – Syringophilopsis catesbyi
Fam. Zosteropidae
Zosterops erythropleurus – Neoaulonastus zosterops
Z. senegalensis – Neoaulonastus zosterops
Pelecaniformes
Fam. Ardeidae
Ardea alba egretta – Trypetoptila casmerodia
Ixobrychus cinnamomeus – Ixobrychiphilus wallacei
Fam. Threskiornithidae
Geronticus calurus – Stibarokris geronticus
Phimosus infuscatus – Selenonycha inspirata
Platalea leucorodia – Stibarokris platalea
Plegadis falcinellus – Stibarokris brevisetosus
Theristicus caudatus – Stibarokris theristicus
Phoenicopteriformes
Fam. Phoenicopteridae
Phoeniconaias minor – Stibarokris phoeniconaias
Phoenicopterus ruber – Stibarokris phoeniconaias
Ph. chilensis – Stibarokris phoeniconaias
Piciformes
Fam. Lybiidae
Pogoniulus bilineatus – Neosyrigophilopsis lybidus, Picineoaulonastus pogoniulus
P. pusillus – Picineoaulonastus pogoniulus
Tanopicobia trachyphoni – Trachyphonus erythrocephalus
Fam. Picidae
Campethera nubica – Picobia dryobatis
Celeus elegans – Neopicobia ea
C. flavus – Neopicobia ea
C. torquatus – Neopicobia ea
Chrysocolaptes lucidus – Neoaulonastes chrysocolaptes
Colaptes [Picumnus] rubiginosus – Neopicobia freya
Dendrocopos borealis – Picobia dryobatis
D. darjellensis – Picobia dryobatis
D. leucotos – Picobia dryobatis, Neoaulonastus picidus
D. macei – Picobia dryobatis
D. major – Picobia dryobatis, Neoaulonastus picidus
D. hortorum – Picobia dryobatis
Dendropicos fuscescens – Picobia dryobatis, Syringophiloidus picidus
D. goerthae – Neopicobia freya, Picobia dryobatis
D. griseocephalus – Neopicobia freya
D. namaquus – Picobia dryobatis
D. stierlingi – Picobia dryobatis
D. syriacus – Picobia dryobatis
Dryocopus galeatus – Neopicobia freya, Picobia dryobatis
Jynx torquilla – Picobia dryobatis
Melanerpes carolinus – Picobia dryobatis,
M. erythrocephalus – Picobia dryobatis
M. formicivorus – Syringophiloidus carolae
Picoides pubescens – Picobia dryobatis, Picisyringophilus kratos
P. mixtus – Picobia dryobatis
P. scalaris – Picobia dryobatis
P. tridactylus – Picobia dryobatis
Piculus rubiginosus – Neopicobia freya
Picumnus exilis – Neopicobia hepburni
P. olivaceus – Neopicobia hepburni
P. sclateri – Neopicobia hepburni
P. temminckii – Neopicobia hepburni
Picus awokera – Picobia heeri
P. canus – Picobia heeri, Neoaulonastus picidus
P. chlorolophus – Picobia heeri
P. erythropygius – Picobia heeri
P. flavinucha – Picobia heeri
P. mentalis – Picobia mentalis
P. puniceus – Picobia heeri
P. squamatus – Picobia heeri
P. viridis – Picobia heeri
Veneliornis cassini – Picobia dryobatis
V. dignus – Picobia dryobatis
V. fumigatus – Picobia dryobatis
V. kirkii – Picobia dryobatis
V. maculifrons – Picobia dryobatis
V. passerines – Picobia dryobatis
V. sanguineus – Picobia dryobatis
V. spilogaster – Picobia dryobatis
Fam. Rhamphastidae
Ramphastos sulfuratus – Rafaicobia ramphastos
R. vietellinus – Rafapicobia ramphastos
Fam. Semnornithidae
Semnornis ramphastinus – Rafapicobia olszanowskii
Procellariiformes
Fam. Procellariidae
Ardenna pacifica – Procellariisyringophilus bulwerius
Bulwerius bulwerius – Procellariisyringophilus bulwerius
Calonectris diomedea – Procellariisyringophilus bulwerius
Puffinus pacificus – Stibarokris mariasi
Fam. Hydrobatidae
Hydrobates pelagicus – Stibarokris hydrobates
Psittaciformes
Fam. Psittacidae
Amazona aestiva – Psittaciphilus amazonae
A. amazonica – Psittaciphilus amazonae
A. finschi – Neoaulobia mironovi
Aratinga canicularis – Neoaulobia mexicana
A. pertinax – Megasyringophilus kethleyi, Neoaulobia mexicana
A. jandaya – Megasyringophilus kethleyi, Neoaulobia aratingae
Bolborhynchus aymara – Peristerophila mucuya
Brotogeris jugularis cyanoptera – Rafapicobia brotogeris
B. versicolurus – Megasyringophilus kethleyi, Peristerophila mucuya
Forpus cyanopygius – Peristerophila forpi
Pionus senilis – Terratosyringophilus pioni
Poicephalus senegalus – Neoaulobia pulyerti, Lawrencicobia poicephali
Fam. Psittaculidae
Agapornis fischeri – Neoaulobia agapornis
A. nigrigenis – Neoaulobia agapornis
A. personatus – Neoaulobia agapornis
A. roseicollis – Neoaulobia agapornis
A. taranta – Neoaulobia agapornis
Eos bernea – Megasyringophilus eos
Geoffroyus geoffroyi – Megasyringophilus geoffroyus
Glossopsitta porphyrocephala – Neopicobia glossopsitta
Loriculus philippensis – Neoaulobia puylaerti
L. pusillus – Neoaulobia puylaerti
Lorius garrulus – Terratosyringophilus loricatus
L. lory – Terratosyringophilus reichholfi
Platycercus eximius – Megasyringophilus platycercus
Psittacula cyanocephala – Megasyringophilus cyanocephala, Neoaulobia psittaculae
P. eupatria – Megasyringophilus cyanocephala
P. krameri – Megasyringophilus cyanocephala
Trichoglossus chlorolepidotus – Megasyringophilus trichoglossus
T. euteles – Megasyringophilus trichoglossus
T. haematodus – Peristerophila mucuya, Terratosyringophilus loricinus
T. ornatus – Megasyringophilus dubinini
Fam. Cacatuidae
Cacatua galerita – Megasyringophilus cacatua
C. tenuirostris – Neoaulobia krafti
C. sulphurea – Lawrencipicobia sulphurea
Calyptorhynchus funereus – Neoaulobia cacatua
C. lathami – Lawrencipicobia calyptorhyncha
Probosciger aterrimus – Neoaulobia cacatua
Fam. Strigopidae
Nestor meridionalis – Peristerophila nestoriae
Pterocliformes
Fam. Pteroclidae
Pterocles coronatus – Columbiphilus pteroclesi
P. senegallus – Pteroclidisyringophus re, Columbiphilus pteroclesi
Strigiformes
Fam. Strigidae
Asio otus – Bubophilus aluconis, B. asiobius
Athene cunicularia – Neobubophilus atheneus
A. brama – Neobubophilus atheneus
A. noctua – Neobubophilus atheneus
Bubo ascalaphus – Bubophilus ascalaphus
B. africanus – Bubophilus ascalaphus
B. bubo – Bubophilus ascalaphus
B. virginianus – Bubophilus ascalaphus
Megascops choliba – Megasyringophilus dalmas
Strix aluco – Bubophilus aluconis
S. woodfordii – Bubophilus aluconis
Fam. Tytonidae
Tyto alba affinis – Bubophilus ascalaphus, B. tytonis
Suliformes
Fam. Phalacrocoracidae
Phalacrocorax auritus – Stibarokris phalacrus
Ph. carbo – Stibarokris dastychi
Tinamiformes
Fam. Tinamidae
Eudromia elegans – Tinamiphilopsis elegans
Nothura boraquira – Tinamiphilopsis ariconte
N. minor – Tinamiphilopsis ariconte
Rhynchotus rufescens – Tinamiphilopsis ariconte
Trogoniformes
Fam. Trogonidae
Pharomachrus antisianus – Syringophiloidus quetzali
Ph. mocinno – Syringophiloidus quetzali
Trogon citreolus – Syringophilopsis trogoni
T. melanocephalus – Syringophilopsis trogoni
Upupiformes
Fam. Phoeniculidae
Phoeniculus purpureus – Picobia phoeniculi
Fam. Upupidae
Upupa epops africana – Picobia phoeniculi
Upupa epops epops – Peristerophila upupi, Picobia phoeniculi
Cloacaroidea Bochkov & O’Connor, 2008: 335
The small family Epimyodicidae has been initially considered subfamily of Cloacaridae, but the analysis of Bochkov (2002) and Bochkov & O’Connor (2008) has shown the need to give it familial status under the new superfamily Cloacaroidea. Although small (19 sp.), the superfamily embraces mites with peculiar localisation and high value in reconstructiing the ways of formation of parasitic mites.
Cloacaridae Camin, Moss, Oliver & Singer, 1967: 263
This small (15 sp.) family has been established almost simultaneously by Camin, Moss, Oliver & Singer (1967) and by Fain (1968). The genera Cloacarus Camin et al., Emyduracarus Fain and Caminacarus Fain contain species living in the rectum of turtles and tortoises in Israel, USA, China, Africa, and Australia. They are attached to the mucose or live more or less submucose. Theodoracarus testudinis Fain live in the muscle tissue of tortoises. Chelonacarus elongatus Pence et Wright, 1998 had been discovered as parasite of sea turtles (Chelonia mydas).
Besides the typical subfamily Cloacarinae, latter has been described the only representative of Pneumophaginae (Pneumophagus bubonis Fain et Smiley, 1989) from the lungs of Bubo virginianus.
Fam. Cloacaridae Camin, Moss, Oliver et Singer, 1967 – 6 gen., 15 sp.
Subfam. Cloacarinae Camin, Moss, Oliver et Singer, 1967 – 14 sp.
Caminacarus Fain, 1968 – 9 sp.
Chelonacarus Pence et Wright, 1998 – 1 sp.
Cloacarus Camin, Moss, Oliver et Singer, 1967 – 2 sp.
Emyduracarus Fain, 1968 – 1 sp.
Theodoracarus Fain, 1968 – 1 sp.
Subfam. Pneumophaginae Fain et Smiley, 1989 – 1 sp.
Pneumophagus Fain et Smiley, 1989 – 1 sp.
Cloacaridae Camin, Moss, Oliver & Singer, 1967: 263
Type genus: Cloacarus Camin, Moss, Oliver et Singer, 1967
Cloacarinae Camin, Moss, Oliver & Singer, 1967: 263
Type genus: Cloacarus Camin, Moss, Oliver et Singer, 1967
Caminacarus Fain, 1968: 6
Type species: Caminacarus theodori Fain, 1968
Caminacarus chrysemys Pence & Casto, 1975: 133
Type locality: Laplace, Louisiana
Distribution: USA (Louisiana)
Host: Chrysemys scripta elegans (Testudines: Emydidae)
Caminacarus costai Fain, 1968: 10
Type locality: Benjamina
Distribution: Israel
Host: Mauremys caspica (Testudines: Emydidae)
Caminacarus dawsoni Bochkov & OConnor, 2008: 340
Type locality: USA
Distribution: USA
Host: Graptemys pseudogeographica (Testudines: Emydidae)
Caminacarus deirochelys Fain, 1968: 18
Type locality: Englewoord
Distribution: USA (Florida)
Host: Deirochelys reticularia (Testudines: Emydidae)
Caminacarus pelomedusae Fain, 1968: 13
Locality: Niarembe, Ituri (DR Congo), Tabora
Distribution: DR Congo, Tanzania
Hosts: Pelomedusa galeata, P. subrufa (Testudines: Pelomedusidae)
Caminacarus pelusios Fain, 1968: 13
Type locality: Lake Mohasi
Distribution: Rwanda
Host: Pelusios castaneus (Testudines: Pelomedusidae)
Caminacarus sinensis Fain, 1968: 16
Type locality: China
Distribution: China
Host: Trionyx sinensis (Testudines: Trionychidae)
Caminacarus terrapenae Pence & Casto, 1975: 13.
Type locality: Louisiana
Distribution: USA (Louisiana)
Host: Terrapene carolina (Testudines: Emydidae)
Caminacarus theodori Fain, 1968: 7
Type locality: Benjamina
Distribution: Israel
Host: Mauremys caspica (Testudines: Emydidae)
Chelonacarus Pence & Wright, 1998: 835
Type species: Chelonacarus elongatus Pence et Wright, 1998
Chelonacarus elongatus Pence & Wright, 1998: 836
Type locality: Bocas del Toro, Isla de Colon
Distribution: Panama
Host: Chelonia mydas (Testudines: Cheloniidae)
Cloacarus Camin, Moss, Oliver & Singer, 1967: 264
Type species: Cloacarus faini Camin et Singer, 1967
Cloacarus beeri Camin & Oliver, in Camin et al., 1967: 269
Type locality: Douglas Lake, Michigan
Distribution: USA (Michigan)
Host: Chrysemys picta (Chelonia: Emydidae)
Cloacarus faini Camin & Singer, in Camin et al., 1967: 265
Type locality: stream near U.S. Route 59 at county line between Douglas Co. and Franklin Co., Kansas
Distribution: USA (Kansas)
Host: Chelydra serpentina (Chelonia: Chelydridae)
Emyduracarus Fain, 1968: 20
Type species: Emydurarus australis Fain, 1968
Emydurarus australis Fain, 1968: 20; Domrow, 1991: 1347
Type locality: St. Lawrence (Queensland)
Distribution: Australia (Coastal central Queensland)
Host: Elseya [Emydura] latisternum (Testudines: Chelidae)
Theodoracarus Fain, 1968: 23
Type species: Theodoracarus testudinis Fain, 1968
Theodoracarus testudinis Fain, 1968: 23
Type locality: Jerusalem
Distribution: Israel
Host: Testudo graeca ibera (Testudines: Testudinidae)
Pneumophaginae Fain & Smiley, 1989: 112
Type genus: Pneumophagus Fain et Smiley, 1989
Pneumophagus Fain & Smiley, 1989: 112
Type species: Pneumophagus bubonis Fain et Smiley, 1989
Pneumophagus bubonis Fain & Smiley, 1989: 112
Type locality: East Lansing, Michigan
Distribution: USA (Michigan)
Host: Bubo virginianus (Strigiformes: Strigidae)
AFRICA
Congo DR (Fain, 1968) – Caminacarus pelomedusae
Rwanda (Fain, 1968) – Caminacarus pelusios
Tanzania (Fain, 1968) – Caminacarus pelomedusae
ASIA
China (Fain, 1968) – Caminacarus sinensis
Israel (Fain, 1968) – Caminacarus costai, C. theodori, Theodoracarus testudinis
NORTH AMERICA
USA (Fain & Smiley, 1989; Pence & Casto, 1975; Bochkov & OConnor, 2008) – Caminacarus chrysemys, C. deirochelys, C. dawsoni, C. terrapenae, Cloacarus beeri, C. faini, Pneumophagus bubonis
SOUTH AMERICA
Panama (Pence & Wright, 1998) – Chelonacarus elongatus
OCEANIA
Australia (Fain, 1968; Domrow, 1991) – Emyduracarus australis
Reptilia
Testudines
Fam. Chelidae
Emydura latisternum – Emydurarus australis
Fam. Cheloniidae
Chelonia mydas – Chelonacarus elongatus
Fam. Chelydridae
Chelydra serpentina – Cloacarus faini
Fam. Emydidae
Chrysemys scripta elegans – Caminacarus chrysemys
Deirochelys reticularia – Caminacarus deirochelys
Graptemys pseudogeographica – Caminacarus dawsoni
Mauremys caspica – Caminacarus costai
Terrapene carolina – Caminacarus terrapenae
Fam. Pelomedusidae
Pelomedusa galeata – Caminacarus pelomedusae
Pelomedusa subrufa – Caminacarus pelomedusae
Pelusios castaneus – Caminacarus pelusios
Fam. Testudinidae
Testudo graeca ibera – Theodoracarus testudinis
Fam. Trionychidae
Trionyx sinensis – Caminacarus sinensis
Aves
Strigiformes
Fam. Strigidae
Bubo virginianus – Pneumophagus bubonis
Epimyodicinae Fain, Lukoschus & Rosmalen, 1982: 4
Epimyodicidae: Bochkov, 2002: 651
The representatives of Epimyodicinae have been discovered in loose deep connective subcutaneous tissues of Soricomorpha and Rodentia, so far only in Europe and North America.
So far only four species are known from subcutaneous tissues of Soricomorpha (Sorex, Crocidura, Talpa) and Rodentia (Microtus, Peromyscus, Sylvaemus).
Epimyodicinae Fain, Lukoschus & Rosmalen, 1982: 4
Epimyodicidae: Bochkov, 2002: 651
Type genus: Epimyodex Fain et Orts, 1969
Epimyodex Fain & Orts, 1969: 66
Type species: Epimyodex talpae Fain et Orts, 1969
Epimyodex crocidurae Fain, Lukoschus & Rosmalen, 1982: 5
Type locality: Nijmegen
Distribution: Holland
Host: Crocidura russula (Soricomorpha: Soricidae)
Epimyodex microti Fain, Lukoschus & Rosmalen, 1982: 7; Fain & Bochkov, 2001: 223
Type locality: Nijmegen (Holland)
Distribution: Holland, Italy, USA
Hosts: Microtus arvalis, M. savii (Rodentia: Arvicolidae), Peromyscus leucopus, P. maniculatus (Rodentia: Cricetidae); Sylvaemus sylvaticus (Muridae)
Epimyodex soricis Fain & Bochkov, 2001: 221- 223.
Type locality: Lincoln Co., Oregon
Distribution: USA (Oregon)
Hosts: Sorex trowbridgii, S. vagrans (Soricomorpha: Soricidae)
Epimyodex talpae Fain & Orts, 1969: 66
Type locality: Lillois, Belgium
Distribution: Belgium
Hosts: Talpa europaea, T. romana (Soricomorpha: Talpidae, Talpinae)
EUROPE
Belgium (Fain & Orts, 1969) – Epimyodex talpae
Holland (Fain, Lukoschus & Rosmalen, 1982) – Epimyodex crocidurae, E. microti
Italy (Fain, Lukoschus & Rosmalen, 1982) – Epimyodex microti, E. talpae
NORTH AMERICA
USA (Fain, Lukoschus & Rosmalen, 1982; Fain & Bochkov, 2001) – Epimyodex microti, E. soricis
Mammalia
Soricomorpha
Fam. Soricidae
Crocidura russula – Epimyodex crocidurae
Sorex trowbridgii – Epimyodex soricis
S. vagrans – Epimyodex soricis
Fam. Talpidae
Talpa europaea – Epimyodex talpae
T. romana – Epimyodex talpae
Rodentia
Fam. Arvicolidae
Microtus arvalis – Epimyodex microti
M. savii – Epimyodex microti
Fam. Cricetidae
Peromyscus leucopus – Epimyodex microti
P. maniculatus – Epimyodex microti
Fam. Muridae
Sylvaemus sylvaticus – Epimyodex microti
Many prominent Acarologists provided literature and advice to the present author in the complicated task to put together and to update the huge information on the seven families of mites treated in this volume. Some will certainly be ommited, but cordial gratitude is first of all due to A. Fain, A. Bochkov, U. Gerson, I. Literák, F. Dusbábek, J. B. Kethley, R. F. Lawrence, B. OConnor, R. V. Southcott, J. de la Cruz, K.M. Jack, H. Rak, F. M. Summers, M. Shiba, R. Domrow, C.H.W. Flechtmann, I. M. Newell, K. Kawashima, A. Hoffmann, D. B. Pence, M. Skoracki, F.S. Lukoschus, Ch. D. Radford, V. Černy, W.W. Moss, W. B. Nutting, J.N. Izdebska, K.M.T. Giesen, M. Nadchatram, S.E. Thewke, K. Uchikawa, V. Bukva, J. O. Whitaker, A. Baker, V. Amaral, M.K.P. Meyer, E. Glowska, M. Hromada, A. Khaustov, A. Saboori, M. Klimovičová, K. Kaszewska, G. Spicer, M. Mironov, M. Zmudzinski, L. Corpuz-Raros, V.I. Volgin, J.L. Izdebska, R.L. Smiley, H. Klompen, M. Dabert, E. Ueckermann.