Labidostommatoidea Ewing, 1934: 64

Phalanx Stomatostigmata Oudemans, 1906

The superfamily contains only one family (Labidostommatidae) with five genera and 75 species.

Fam. Labidostommatidae - 4 gen., 75 recent sp.

Akrostomma Robaux, 1977 - 6 sp.

Eunicolina Berlese, 1911 - 4 sp. (plus one fossil)

Labidostomma Kramer, 1879 - 44 sp

Atyeonella Feider et Vasiliu, 1969 - 8 sp.

Labidostomma Kramer, 1879 – 16 sp.

Nicoletiella Canestrini, 1882 – 17 sp. (plus one fossil)

Cornutella Feider et Vasiliu, 1969 – 2 sp.

Pselistoma Bertrand, 1990 – 1 sp.

Pseudocornutella Sidorchuk et Bertrand, 2013 – 1 sp. (fossil)

Sellnickiella Feider et Vasiliu, 1969 – 20 sp. (plus one fossil)

Dicastriella Feider et Vasiliu, 1970 - 7 sp.

Sellnickiella Feider et Vasiliu, 1969 – 13 sp.

EUROPE

Austria (Schrank, 1781; Willmann, 1939, 1951; Franz, 1954; Vistorin, 1978; Schmölzer, 1993) –Labidostomma cornuta, L. denticulatum, L. luteum, L. lyra

Bulgaria (Beron, 1995) – Eunicolina nova, Labidostomma (Cornutella) cornutum

Croatia (Willmann, 1940, 1941; Vistorin, 1978а) – Eunicolina tuberculata, Labidostomma (Cornutella) cornutum

Czech Rep. (Willmann, 1939)

Denmark (Luxton, 1966, 1981; Hallas, 1978) – Labidostomma luteum

Finland (Lehtinen & Niemi, 1995) – Labidostomma (L.) luteum

France (contin.) (Cooreman, 1954, 1959; Coineau, 1964a, 1964b; Bertrand, 1980; Bertrand & Coineau, 1979) – Akrostomma coralloides, Labidostomma (Cornutella) cornutum, Labidostomma (Nicoletiella) repetitor (= L. (Nicoletiella) jacquemarti), L. (N.) luteum, Labidostomma (L.) integrum

Corsica (Bertrand, 1981; Bertrand et al., 2012) - Labidostomma (Nicoletiella) corsicum, L. (N.) glymma, L. (N.) vialeae, Labidostomma (L.) integrum

Georgia (Reck, 1940) – Labidostomma caucasicum (= L. integrum)

Germany (C.L.Koch, 1838; Oudemans, 1916; Willmann, 1955; Vistorin, 1978a) – Labidostomma denticulatum, L. luteum, L. lyra

Great Britain (Michael, 1880) – Labidostomma luteum

Corfu (Berlese, 1911; Sellnick, 1931) – “Labidostoma corcyraeum””(= L. denticulatum), Eunicolina nova

Hungary (Willmann, 1937; Dudich, Kolosváry & Szalay, 1940; Újváry & Kontschán, 2010; Pfliegler & Bertrand, 2011) – Labidostomma denticulata, L. lyra, L. (Cornutella) cornutum

Iceland (Sellnick, 1940) – Labidostomma denticulata (actually L. luteum, see Lehtinen & Niemi, 1995)

Ireland (Halbert, 1915) – Nicoletiella cornuta (actually L. luteum, see Lehtinen & Niemi, 1995)

Italy (contin.) (Berlese, 1887; 1911; Lombardini, 1943; Bottazzi, 1956; Vistorin, 1978a) – Akrostomma zangherii (sp. dubia), Eunicolina tuberculata, Labidostomma denticulatum, L. integrum, L. luteum

Sicily (Berlese, 1883; Bottazzi, 1956) – Labidostomma luteum

Sardinia (Bertrand et al., 2012) – Labidostomma repetitor, L. (L.) integrum

Poland (Bloszyk, 1980; Błoszyk & Czarnota, 1998; Bloszyk, Konwerski, Napierala, Goldyn & Gawronska, 2007; Bloszyk et al., 2017; Bloszyk, Adamski & Napierała, 2018) – Labidostomma (Nicoletiella) denticulatum, L. (N.) luteum, L. (Cornutella) cornutum

Romania (Cooreman, 1951; Feider & Vasiliu, 1968, 1970; Iavorschi, 1992; Kontschán, 2008) - Labidostomma motasi, L. (Cornutella) cornutum, L. (L.) integrum (= L. caucasicum), L.(Nicoletiella) denticulata, L. (Nicoletiella) romanica, Eunicolina [Grandjeanellina] nova

Slovakia (Bloszyk et al., 2022) – Labidostomma luteum

Slovenia (Willmann, 1932; Willmann, 1940, 1941; Vistorin, 1978a) – Labidostomma (Nicoletiella) lyra, Labidostomma cornuta, L. spelaeophila

Spain (cont.) (Mihelčič, 1958a,b; Bertrand, 1983 ; Bertrand et al., 2012) – Akrostomma coineaui, Labidostomma (Nicoletiella) luteum, [L. colomerense], L. franzi, L. (L.) integrum

Sweden - Labidostomma (Nicoletiella) luteum

Switzerland (Schweizer & Bader, 1963) – Labidostomma (Nicoletiella) luteum

Turkey (all) (Urhan & Özmen, 2006) – Labidostomma sp.

Baltic Area (Dunlop & Bertrand, 2011; Sidorchuk et Bertrand, 2013) - Labidostomma (Nicoletiella) paleoluteum (fossil), Labidostomma (Pseudocornutella) electri (fossil), Sellnickiella balticae (fossil)

AFRICA

Algeria (Grandjean, 1942) – Labidostomma integrum, L. (Cornutella) cornutum

Angola (Bertrand & Theron, 1990a) – Labidostomma (L.) angolensis, L. (L.) percilliata

DR Congo (Cooreman, 1954) – Labidostomma integrum schoutedeni

South Africa (Thor, 1931a; Meyer & Ryke, 1959; Bertrand & Theron, 1990a, 1992) - Labidostomma (L.) aethiopica, L. (L.) fissurata, L.(L.) mirax, L.(L.) percilliata, Sellnickiella (S.) capensis, S.(S.) hoegi, S.(S.) natalensis

ASIA

India (Bhattacharyya, 1966) – Labidostomma himalayensis

Iran (Bertrand et al., 2012; Jalaeian, Kamali & Bertrand, 2013) – Labidostomma (L.) intermedia

Israel and Palestina - Labidostomma (L.) intermedia

Japan (Shiba, 1969, 1972, 2004) – Labidostomma (Atyonella) simplex, Labidostomma (Cornutella) ezoensis, L. (Cornutella) mutisetosa

Kazakhstan (Bertrand et al., 2012) - Labidostomma (L.) integrum

Korea (N.) (Feider & Vasiliu, 1972) – Labidostomma asiatica, L. (Cornutella) mutisetosa, M. multisetosa

Nepal (Feider & Vasiliu, 1968a) - Labidostomma nepalense

Philippines (Bertrand & Corpus Raros, 1997) - Labidostomma (L.) guadalupense

NORTH AMERICA

USA (Ewing, 1913; Greenberg, 1952; Robaux, 1977; Crossley & Simmons, 2015) - Akrostomma gabriella, A. grandjeani, A. plumosum, Eunicolina porifera, Labidostomma (L.) barbae, Labidostomma (Nicoletiella) caloderum, L. (N.) fergusoni, L. (N.) pacifica,

SOUTH and CENTRAL AMERICA

Brazil (Sellnick, 1922; Feider & Vasiliu, 1970; Feider, Vasiliu & Calugar, 1974; Bertrand, 1990) – Labidostomma amasonica, Sellnickiella (S.) brasiliensis, S.(S.) decemlabeculata, S. (S.) mediumreticulata, S. (S.) schusteri, S. (S.) pulchra

Chile (contin.) (Feider & Vasiliu, 1970; Feider, Vasiliu & Calugar, 1974; Bertrand, 1990c) – Labidostomma (Nicoletiella) luteoides, L. (Pselistoma) legendrei, Sellnickiella (S.) decemlabeculata, Sellnickiella (Dicastriella) caeca, S. (D.) coineaui, S. (D.) covarubiasi, S. (D.) fungifera, S.(D.) fusiformis, S. (D.) mediohexagonata, Sellnickiella (S.) macroscopa,

Guatemala (Stoll, 1891) – Labidostomma (Labidostomma) neotropica

Peru (Bertrand, 1990c) – Labidostomma (Atyeonella) peruviana

OCEANIA

Australia (continent) (Wommersley, 1935; Atyeo & Crossley Jr., 1961a; Bertrand, 1997; Halliday, 1998) – Sellnickiella adelaideae, S. biunguiculata, S. womersleyi

New Zealand (Atyeo & Crossley Jr. 1961b) – Sellnickiella (S.) circinus, Labidostomma Atyonella fictiluteum, L. (A.) glandula, L. (A.) malleolus, L. (A.) multifarium, L. (A.) ocellatum, L. (A.) striatum

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Eupodoidea Banks, 1894

According to Zhang, Ed. (2011), the superfamily consists of nine families (four of them described recently), with 85 genera and 309 species. The four “old” families (Eupodidae, Penthaleidae, Penthalodidae and Rhagidiidae) account for 277 of these species.

Fam. Eupodidae C.L. Koch, 1842 - 14 gen., 95 sp.

Aethosolenia Baker et Lindquist, 2002 – 1 sp.

Benoinyssinae Fain, 1958 – 10 sp.

Caleupodes A. Baker, 1987 – 1 sp.

Claveupodes Strandtmann et Prasse, 1977 – 1 sp.

Cocceupodes Sigh Thor, 1934 – 8 sp.

Echinoeupodes Khaustov, 2017 – 2 sp.

Eupodes C.L. Koch, 1835 – 47 sp.

Filieupodes Jesionowska, 2010 – 9 sp.

Linopodes C.L. Koch, 1835 - 10 sp.

Neoprotereunetes Fain et Camerik, 1994 – 1 sp.

Niveupodes Barilo, 1991 – 1 sp.

Pseudeupodes Khaustov, 2014 – 1 sp.

Pseudopenthaleus Khaustov, 2015 – 2 sp.

Xerophiles Jesionowska, 2003 – 1 sp.

EUROPE

Austria (Irk, 1939; Willmann, 1951; Mihelčič, 1957, 1958; Schmölzer, 1962, 1993) – Cocceupodes mollicellus, Eupodes pseudoclavifrons, E. variegatus, Linopodes motatorius

Belgium (Cooreman, 1943) – Claveupodes sp., Linopodes motatorius

Bulgaria (Beron, 2011) – Linopodes motatorius

Croatia (Matočec, 2002) - Linopodes motatorius

Czech Rep. (Willmann, 1954) – Eupodes sp., Linopodes motatorius, [? L. subterraneus Wankel]

Denmark (Weis-Fogh, 1948; Haarløv, 1957; Hallas, 1978) – Cocceupodes mollicellus, Eupodes berlesei, E. viridis, E. voxencollinus, Filieupodes paradoxus

Greenland (Haarløv, 1942; Makarova, 2015) – Cocceupodes breweri, C. mollicellus (= Eupodes clavifrons R. Canestrini, 1886 = Cocceupodes curticlava Thor, 1934), C. paradoxus, C. cf. stellatus

Estonia (Thor & Willmann, 1941) – Protereunethes striatellus

France (contin.)(Coineau, 1976) – Eupodes strandtmanni

Germany (C.L. Koch, 1835, 1838; Willmann, 1952; Strandtmann & Prasse, 1977) – Benoinyssus ereynetoides, Claveupodes delicatus, Cocceupodes stellatus, Eupodes leucomelas, E. ocellatus, E. striola, E. signatus, E. trifasciatus, E. voxencollinus, Filieupodes trisetatus, Linopodes motatorius, L. ravus, Protereunethes striatellus

Great Britain (Hull, 1918; Evans, 1952; Turk & Turk, 1952; A. Baker, 1987) – Caleupodes reticulatus,Claveupodes sp., Eupodes clavifrons, E. unifasciatus, E. variegatus, E. voxencollinus, Filieupodes paradoxus, Linopodes antennaepes, L. motatorius

Holland (Oudemans, 1906; Thor & Willmann, 1941) - Eupodes milvinus, E. viridis, Protereunethes striatellus

Hungary (Dudich et al., 1940) – Linopodes motatorius

Iceland (Sellnick, 1940) – Eupodes variegatus, Linopodes motatorius

Surtsey (Lindroth, Andersson, Bödvarsson & Richter, 1973) – Cocceupodes clavifrons, Eupodes hiemalis, E. variegatus, Linopodes motatorius

Ireland (Halbert, 1915, 1920) – Eupodes variegatus var. halophilus

Italy (contin.)(Canestrini, 1886; Thor, 1934, Thor & Willmann, 1941; Bottazzi, 1950) - Cocceupodes mollicellus (= clavifrons), Eupodes fusifer, E. iconicus, E. milvinus, E. vallombrosae, E. variegatus, Linopodes antennaepes, L. eupodoides, L. motatorius

Sicily (Berlese, 1923) – Protereunetes agilis

Norway (Thor, 1912, 1934a, 1934c; Thor & Willmann, 1941) – Cocceupodes mollicellus (=curviclava), Eupodes pseudoclavifrons, E. decoloratus, E. hjartdaliae, E. skiaaki, E. variegatus, E. voxencollinus, Linopodes motatorius

Svalbard (Spitzbergen)(Oudemans, 1926; Thor, 1934b, 1934c) – Eupodes clavifrons, Protereunetes boerneri

Poland (Jesionowska, 2005, 2006, 2007a, 2007b, 2008, 2010; Szudarek-Trepto, Kaźmierski, Dabert et al., 2020) – Claveupodes sp., Cocceupodes longisolenidiatus, Filieupodes filiformis, F. filistellatus, Eupodes voxencollinus, Xerophiles ereynetoidalis

Romania (Ivan et al., 2016) – Eupodes sp.

Russia (cont.)

Crimea (Khaustov, 2014, 2015, 2017) – Echinoeupodes echinus, Pseudeupodes porosus, Pseudopenthaleus tauricus

Slovakia (Jesionowska, 2003) - Xerophiles ereynetoidalis

Spain (cont.)(Mihelčič, 1958b; Santamaría et al., 2012) - Cocceupodes sp., Benoinyssus ereynetoides, E. valombrosae, E. variegatus, Linopodes motatorius

Sweden (Thor & Willmann, 1941; Sellnick, 1958) - Eupodes milvinus, E. variegatus

Switzerland (Haller, 1880; Schweizer, 1922; Cooreman, 1954,1959, Schweizer & Bader, 1963) – Eupodes berlesei, E. haltica, E. vallombrosae, Linopodes eupodoides, L. motatorius

AFRICA

Angola (Meyer & Ryke, 1960; Olivier & Theron, 1997a, 1998) – Benoinyssus celatus, B. ornatus, Eupodes parafusifer

Botswana (Olivier & Theron, 1997a; 1998) - Benoinyssus serratus, Eupodes minipulus

Cabo Verde (Olivier & Theron, 1998) - Eupodes minipulus

Egypt (Trägårdh, 1904; Abou-Awad, 1984: 333; Abou-Awad & El-Bagouri, 1984, 1986; Abou-Awad, El-Sawaf & Abdel-Khalek, 2006) – Benoinyssus momeni, Cocceupodes sharkiensis, Filieupodes aegyptiacus, F. strandtmanni, Eupodes bakeri, E. momeni, E. niloticus, E. strandtmanni, E. temperatus, E. zaheri, Linopodes barnufi, L.curtipes

Ivory Coast - Eupodes sigmoidensis

Namibia (Olivier & Theron, 1997a) - Benoinyssus serratus

Rwanda (Fain, 1958, 1959) – Benoinyssus najae

South Africa (Thor, 1934c; Meyer & Ryke, 1960; Olivier & Theron, 1997a, 1997b, 1998, 2003) – Benoinyssus falcatus, B. ornatus, B. pusillus, B. serratus, C. gracilongus, Eupodes acanthus, E. fusiferellus, E. hamatus, E. indentatus, E. lutatus, E. minipilus, E. parafusifer, E. variegatus, Filieupodes fusiformis

Swaziland (Olivier & Theron, 1997a) – Benoinyssus ornatus, B. serratus

Zimbabwe (Olivier & Theron, 1998) - Eupodes minipulus

“East Africa” (Berlese, 1916) – “Linopodes agitatorius”

ASIA

China (Hagstrum et al., 2013) – Eupodes voxencollinus

India (Ghosh & Gupta, 2003; Gupta et al., 2004, Indranil et al., 2011) – Eupodes sigmoidensis

Indonesia (Corpuz-Raros, 2005) – Eupodes pusillus [Java]

Iran (Darbemamieh, Hajiqanbar & Khanjani, 2013) - Benoinyssus sp., Claveupodes sp., Cocceupodes sp., Eupodes crozetensis, E. sigmoidensis, E. voxencollinus, Linopodes antennaepes, L. cameronensis

Israel and Palestina (Lipkin & Safriel, 1971) – Eupodes variegatus var. halophila

Japan (Karsch, 1881; Morikawa, 1963; Shiba, 1969a, 1969c, 1971, 1978) – Cocceupodes communis, Echinoeupodes turgidus, Eupodes cordiformis, E. infirmus, E. longisetatus, E. okinoshimaensis, Linopodes iwatensis, L. pubescens, Protereunetes perforates, P. villosus, Pseudopenthaleus ciliatus

Malaysia (Shiba, 1976) – Eupodes berensis, E. lumbarus, E. onoi, E. pasohensis, E. pusillus, E. pseudolumbarus, Linopodes cameronensis

Philippines (Corpuz-Raros, 2005) – Eupodes callejae, E. garciai, E. onoi, E. pusillus, E. pseudolumbarus

Singapore (Corpuz-Raros, 2005) – Eupodes onoi

United Arab Emirates (Negm, 2014) – Eupodes sp.

Uzbekistan (Barilo, 1991; Khaustov, 2016) – Niveupodes nivalis

Yemen (Ueckermann et al., 2006) – Eupodes sp.

NORTH AMERICA

Canada (Miller, 1925; Behan, 1978; Moseley, 2007; Beaulieu et al., 2019 – 13 sp.) - Eupodes voxencollinus, Linopodes antennaepes

Mexico (Baker, 1945) - Benoinyssus tuberculatus

USA (Banks, 1894; 1896; 1897, 1907; 1916; Miller, 1925; Thor, 1934) – Eupodes brevipes, E. marinus, Linopodes antennaepes

Alaska (Strandtmann, 1971) – Cocceupodes breweri, Eupodes alaskanensis, E. voxencollinus, Filieupodes shepardi, Protereunetes boerneri

Hawaii (Strandtmann & Goff, 1978) - Eupodes hawaiiensis, E. sigmoidensis, E. voxencollinus

SOUTH and CENTRAL AMERICA

Argentina (Bedano, 2004) – 1 sp.

Brazil (Bizarro et al., 2020) – Benoinyssus oconnori

Costa Rica (Baker & Lindquist, 2002) – Aethosolenia laselvensis

OCEANIA

Australia (continent)(Clancy, 1981) - Linopodes sp.

New Zealand (Spain & Luxton, 1971; Martin, 1983; Luxton, 1990) – Claveupodes delicatulus, Linopodes spp., Eupodes minutus,

Campbell Island (Strandtmann, 1964) - Eupodes longisetatus

Macquarie Island (Womersley, 1937; Luxton, 1990) – Eupodes minutus

ANTARCTICA

Antarctic Continent (Strandtmann & Womersley, 1963; Strandtmann, 1967; Strandtmann & Davies, 1972; Gless, 1972; Strandtmann & Sømme, 1977; Marshall & Pugh, 1996; Llano & Gless, 2013) – Eupodes angardi, E. winsnesi, E. wisei

South Georgia (Strandtmann, 1970) – Eupodes minutus

South-Orkney-Islands (Booth, Edwards & Usher, 1985) – Eupodes exiguus, E. parvus

South Shetland Islands (Strandtmann & Tilbrook, 1968) – Cocceupodes australis

SUB-ANTARCTIC

Crozet Islands (Strandtmann & Davies, 1972) – Eupodes crozetensis, E. riedli, Possession I.

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