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ACARORUM CATALOGUS X. Trombidiformes, Prostigmata, Superfamilia Labidostommatoidea (Labidostommatidae), Superfamilia Eupodoidea, (Eupodidae, Dendrochaetidae, Rhagidiidae, Eriorhynchidae, Pentapalpidae, Penthalodidae, Penthaleidae, Proterorhagiidae, Strandtmanniidae), Superfamilia Tydeoidea, Ereynetidae, Superfamily Paratydeoidea, Paratydeidae, Superfamilia Anystoidea, (Anystidae, Erythracaridae,Teneriffiidae, Pseudocheylidae, Stigmocheylidae), Superfamilia Caeculoidea (Caeculidae), Superfamilia Adamystoidea (Adamystidae), Superfamilia Pomerantzioidea (Pomerantziidae)
expand article infoPetar Beron
‡ Unaffiliated, Sofia, Bulgaria
Open Access

Introduction

This is the last volume of the series Acarorum Catalogus. We tried to follow the system generally accepted. After the publication of the first volume of the series (Erytraeoidea, Calyptostomatoidea) in 2008 appeared a comprehensive treatment of mites (Manuel of Acarology III of Krantz & Walter, Eds. 2009), in which many changes in the system of Acari are proposed. Acariformes are already a superorder, Prostigmata – suborder, but within the order Trombidiformes.

Here is treated first the infraorder Labidostommatina with its only superfamily Labidostommatoidea and the only family Labidostommatidae with five genera and 75 species. The other infraorder, treated in the present volume (Eupodoidea) includes the superfamily Eupodoidea with nine families, including five described after 1979. Treated are also the superfamilies Tydeoidea (fam. Ereynetidae), Paratydeoidea (Paratydeidae), Anystoidea (five families), Caeculoidea (Caeculidae), Adamystoidea (Adamystidae), and Pomerantzioidea (Pomerantziidae). In this volume are analysed 230 genera and 967 species of mites.

Labidostommatidae – 75 sp. (+4 fossils)

Eupodidae and Cocceupodidae – 95 sp.

Dendrochaetidae – 1 sp.

Rhagidiidae – 150 sp.

Eriorhynchidae – 5 sp.

Pentapalpidae – 1 sp.

Penthalodidae – 44 sp.

Penthaleidae – 27 sp.

Proterorhagiidae – 1 sp.

Strandtmanniidae – 1 sp.

Ereynetidae – 180 sp.

Paratydeidae – 13 sp.

Anystidae – 102 sp.

Erythracaridae – 68 sp.

Teneriffiidae – 28 sp.

Pseudocheylidae – 23 sp.

Stigmocheylidae – 5 sp.

Caeculidae – 114 sp.

Adamystidae – 28 sp.

Pomerantziidae – 6 sp.

This last volume of the series Acarorum Catalogus ends with an update of the former nine volumes. Added to these volumes are another genera and species.

Order Trombidiformes

The recent survey of order Trombidiformes (Zhang, Ed., 2011) defined suborder Prostigmata as containing four infraorders and 40 superfamilies. This volume concerns 20 families of prostigmatid mites, mostly soil and plant inhabitants. Some (Speleognathinae, Riccardoella) are parasites of vertebrates and snails.

Infraorder Labidostommatina

The Mite Superfamily Labidostommatoidea

Labidostommatoidea Ewing, 1934: 64

Phalanx Stomatostigmata Oudemans, 1906

The superfamily contains only one family (Labidostommatidae) with five genera and 75 species.

The Mite Family Labidostommatidae Oudemans

Nicoletiellidae G. Canestrini, 1891

Labidostomidae Oudemans, 1904

Labidostommatidae Oudemans, 1906

Strongly hitinized free living mites (600 – 1500 µm long), usually yellow and with one large lense-formed organ behind the side eyes. About 75 species, living in moss, leaf litter and other organic materials. Feed on small insects and mites.

The names of the main genus have varied since their description and are used differently even at present by various authors: Labidostoma, Labidostomma, Nicoletiella (as genus or subgenus of Labidostomma), Nicoletiellidae, Labidostomidae, Labidostommatidae.

The first species has been described very early (Acarus denticulatus Schrank, 1776), but the base of the taxonomy of the family was laid with the descriptions of Labidostomma Kramer, 1879 and Nicoletiella Canestrini, 1882, than several other authors contributes with more new species and discussions: Stoll, Berlese, Lombardini, Coineau, Ewing, Feider, Greenberg, Meyer & Ryke, Oudemans, Grandjean, Robaux, Sellnick, Shiba, Štorkán, Willmann, Womersley, Iavorschi, and others. Some authors should be mentioned specially. Atyeo & Crossley (1961) described six new species from New Zealand. Revision of the family and several new descriptions are due to the Romanian authors Feider & Vasiliu, including generic/subgeneric taxa (Grandjeanellina, Atyeonella, Sellnickiella, Dicastriella, Cornutella, Mahunkiella).

The modern treatment of the family has been started by M. Bertrand since 1980 and so far he published (partly with co-authors) 24 new species and the subgenera Pseudocornutella and Pselistoma.

Composition of fam. Labidostommatidae

Fam. Labidostommatidae - 4 gen., 75 recent sp.

Akrostomma Robaux, 1977 - 6 sp.

Eunicolina Berlese, 1911 - 4 sp. (plus one fossil)

Labidostomma Kramer, 1879 - 44 sp

Atyeonella Feider et Vasiliu, 1969 - 8 sp.

Labidostomma Kramer, 1879 – 16 sp.

Nicoletiella Canestrini, 1882 – 17 sp. (plus one fossil)

Cornutella Feider et Vasiliu, 1969 – 2 sp.

Pselistoma Bertrand, 1990 – 1 sp.

Pseudocornutella Sidorchuk et Bertrand, 2013 – 1 sp. (fossil)

Sellnickiella Feider et Vasiliu, 1969 – 20 sp. (plus one fossil)

Dicastriella Feider et Vasiliu, 1970 - 7 sp.

Sellnickiella Feider et Vasiliu, 1969 – 13 sp.

Catalogue of Labidostommatidae

Family Labidostommatidae Oudemans

Nicoletiellidae G.Canestrini, 1891: (Type genus: Nicoletiella Canestrini, 1882)

Labidostomidae Oudemans, 1904; Bertrand, 1990a: 31

Labidostommatidae Oudemans, 1906

Sellnickiellinae Feider & Vasiliu, 1969: 206

Nicoletiellinae Can.: Feider & Vasiliu, 1969: 204

Eunicolinae Feider & Vasiliu, 1969: 204

Type genus: Labidostomma Kramer, 1879

Genus Akrostomma Robaux

Akrostomma Robaux, 1977: 453

Type species: Akrostomma grandjeani Robaux, 1977

Akrostomma coineaui Bertrand

Akrostomma coineaui Bertrand, 1983: 160

Type locality: Isla de Ons (Provincia Pontevedra)

Distribution: Spain

Akrostomma coralloides Bertrand et Coineau

Akrostomma coralloides Bertrand & Coineau, 1979: 101

Type locality: Banyuls (valley of Baillaurie, Pouade Road)

Distribution: France

Akrostomma gabriella Crossley et Simmons

Akrostomma gabriella Crossley & Simmons: 2015: 459

Type locality: California, Los Angeles County, San Gabriel Mts., Glendale Ridge Road, San Dimas Experimental Forest

Distribution: USA (California)

Akrostomma grandjeani Robaux

Akrostomma grandjeani Robaux, 1977: 453

Type locality: H. Cowell Redwoods State Park (Sta. Cruz Co.), California

Distribution: USA (California)

Akrostomma plumosum (Greenberg)

Labidostomma plumosum Greenberg, 1952: 201

Labidostoma (Labidostoma) plumosum Greenberg: Bertrand, 1990a: 36

Akrostomma plumosum (Greenberg): Crossley & Simmons, 2015: 459

Type locality: Gale’s Canyon, Winters, California

Distribution: USA (California)

Akrostomma zangherii Lombardini

Labidostomma zangherii Lombardini, 1943: 21

Akrostomma zangherii (Lombardini)(?): Bertrand, 1990a: 35

Type locality: Italy

Distribution: Italy

Genus Eunicolina Berlese

Eunicolina Berlese, 1911: 430

Type species: Eunicolina tuberculata Berlese, 1911. Redescription, Vistorin 1980: 376

Grandjeanellina Feider & Vasiliu, 1969: 206 - Type species: Eunicolina nova Sellnick, 1931; syn. by Bertrand, 1990a: 35

Eunicolina glaesi Bertrand, Sidorchuk et Hoffeins (fossil)

Eunicolina glaesi Bertrand, Sidorchuk & Hoffeins, 2015: 323

Type locality: Baltic amber

Distribution: Baltic amber (Eocene)

Eunicolina nova Sellnick

Eunicolina nova Sellnick, 1931: 772; Willmann,1941: 51; Beron, 1995: 5

Grandjeanellina nova (Sellnick): Feider & Vasiliu, 1969: 206

Type locality: Korfu (Greece)

Distribution: Bulgaria, Croatia, Greece, Romania

Eunicolina porifera Greenberg

Eunicolina porifera Greenberg, 1932: 201; Feider & Vasiliu, 1969: 206; Bertrand, 1990a: 35

Type locality: Cape Henry, Virginia

Distribution: USA

Eunicolina travei Coineau

Eunicolina travei Coineau, 1964: 153; Feider & Vasiliu, 1969: 206; Bertrand, 1990a: 35

Type locality: Saint-Paul d’Eygeaux (Haute-Vienne), France

Distribution: France

Eunicolina tuberculata Berlese

Eunicolina tuberculata Berlese, 1911: 430; Feider & Vasiliu, 1969: 206; Vistorin, 1978a: 279; Bertrand, 1990a: 35

Type locality: “in agro Tridentino”,”Tiarno” (Italy)

Distribution: Croatia, Italy

Genus Labidostomma Kramer

Labidostomma Kramer, 1879: 13

Labidostoma Michael, 1880:

Labidostoma Bertrand, 1990a: 36

Panoplia von Heyden, 1826

Type species: Acarus denticulatus Schrank, 1776

Ceratoacarus Ewing, 1913: 128

Type species: Ceratoacarus pacificus Ewing, 1913

Subgenus Atyeonella Feider et Vasiliu

Atyeonella Feider & Vasiliu, 1969: 205 (genus)

Labidostoma (Atyeonella): Bertrand, 1990a: 36 (subgenus)

Type species: Labidostoma fictiluteum Atyeo et Crossley, 1961

Labidostomma (Atyeonella) fictiluteum Atyeo et Crossley

Labidostoma fictiluteum Atyeo & Crossley, 1961b: 40

Atyeonella fictiluteum (Atyeo et Crossley): Feider & Vasiliu, 1969: 206

Labidostoma (Atyeonella) fictiluteum Atyeo & Crossley: Bertrand, 1990a: 36

Type locality: Ngongotaha, Rotorua, Auckland

Distribution: New Zealand

Labidostomma (Atyeonella) glandula Atyeo et Crossley

Labidostoma glandula Atyeo & Crossley, 1961b: 44

Atyeonella fictiluteum (Atyeo et Crossley): Feider & Vasiliu, 1969: 206

Labidostoma (Atyeonella) glandula Atyeo et Crossley: Bertrand, 1990a: 36

Type locality: Ngongotaha, Rotorua, Auckland

Distribution: New Zealand

Labidostomma (Atyeonella) malleolus Atyeo et Crossley

Labidostoma malleolus Atyeo & Crossley, 1961b: 37

Atyeonella maleolus (Atyeo et Crossley): Feider & Vasiliu, 1969: 206

Labidostoma (Atyeonella) malleolus Atyeo et Crossley: Bertrand, 1990a: 36

Type locality: Kapiti Island, Wellington

Distribution: New Zealand

Labidostomma (Atyeonella) multifarium Atyeo et Crossley

Labidostoma multifarium Atyeo & Crossley, 1961b: 33

Atyeonella multifarium (Atyeo et Crossley): Feider & Vasiliu, 1969: 206

Labidostoma (Atyeonella) multifarium (Atyeo et Crossley): Bertrand, 1990a: 36

Type locality: Orepuke, Southland

Distribution: New Zealand

Labidostomma (Atyeonella) ocellatum Atyeo et Crossley

Labidostoma ocellatum Atyeo & Crossley, 1961b: 42

Atyeonella ocelatum (Atyeo et Crossley): Feider & Vasiliu, 1969: 206

Labidostoma (Atyeonella) ocellatum Atyeo & Crossley: Bertrand, 1990a: 36

Type locality: Coromandel Peninsula, Auckland

Distribution: New Zealand

Labidostomma (Atyeonella) peruviana Bertrand

Labidostoma (Atyeonella) peruviana Bertrand, 1990c: 182

Type locality: Sierra Garavito, Prov. de Cusco

Distribution: Peru

Labidostomma (Atyeonella) simplex (Shiba)

Atyeonella simplex Shiba, 1972: 48

Labidostoma (Atyeonella) simplex (Shiba): Bertrand, 1990a: 36

Type locality: Hokkaido, Poroshiri-sanso at the foot of Mt. Poroshiri Hadaka

Distribution: Japan (Hokkaido)

Labidostomma (Atyeonella) striatum Atyeo et Crossley

Labidostoma striatum Atyeo & Crossley, 1961b: 46

Atyeonella striatum (Atyeo et Crossley): Feider & Vasiliu, 1969: 206

Labidostoma (Atyeonella) striatum Atyeo et Crossley: Bertrand, 1990a: 36

Type locality: Takahe Valley, Te Anau, Otago

Distribution: New Zealand

Subgenus Labidostomma Kramer

Labidostomma Kramer, 1879: 13

Labidostoma Michael, 1888

Type species: Acarus denticulatus Schrank, 1776 (= Labidostomma luteum Kramer, 1879 - Redescription, Lehtinen & Niemi 1995: 217)

Labidostomma (Labidostomma) aethiopica Meyer et Ryke

Labidostoma (Labidostoma) aethiopica Meyer & Ryke, 1959: 417; Bertrand, 1990a: 36; Bertrand & Theron, 1990: 248

Type locality: South Africa, Bathurst

Distribution: South Africa

Labidostomma (Labidostomma) amasonica Feider, Vasiliu et Calugar

Labidostomma amasonica Feider, Vasiliu & Calugar, 1974: 424

Labidostoma (Labidostoma) amasonica Feider, Vasiliu et Calugar: Bertrand, 1990a: 36

Type locality: Brazil [Amazonia]

Distribution: Brazil

Labidostomma (Labidostomma) angolensis Bertrand et Theron

Labidostoma (Labidostoma) angolensis Bertrand & Theron, 1990a: 248

Type locality: Luachine, Luangue, Tchitentga

Distribution: Angola

Labidostomma (Labidostomma) barbae Greenberg

Labidostomma barbae Greenberg, 1952: 198

Labidostoma (Labidostoma) barbae Greenberg: Bertand, 1990a: 36

Type locality: 3 mi southeast of Thomasville, Thomas County, Georgia

Distribution: USA (Georgia)

Labidostomma (Labidostomma) fissurata Bertrand et Theron

Labidostoma (Labidostoma) fissurata Bertrand & Theron, 1990a: 248

Type locality: Potchefstroom

Distribution: South Africa

Labidostomma (Labidostomma) guadalupense Bertrand et Corpuz-Raros

Labidostomma guadalupense Bertrand & Corpuz-Raros, 1997: 363

Type locality: Leyte, Guadalupe

Distribution: Philippines

Labidostomma (Labidostomma) integrum Berlese

Labidostoma integrum (Berlese): Berlese, 1911: 429; 1913: 138; Costa, 1961: 259

Labidostoma (Labidostoma) integrum (Berlese): Bertrand, 1990a: 36; Bertrand et al., 2012: 235

Labidostomma integrum : Thor, 1931: 73; Feider & Vasiliu, 1970: 371

Type locality: Umbria (Italy)

Distribution: France (incl. Corsica), Israel, Italy (incl. Sardinia), Kazakhstan, Romania, Spain

Labidostomma (Labidostomma) integrum Berlese

Labidostomma integrum : Thor, 1931: 73; Feider & Vasiliu, 1970: 371; Bloszyk et al., 2003: 173.

Labidostomma (L.) caucasicum Reck, 1940: 452; Feider & Vasiliu, 1969: 205 and Bloszyk et al., 2003: 173 consider this species as synonym of L. integrum

Type locality: D.R.Congo

Distribution: D.R.Congo

Labidostomma (Labidostomma) intermedia Bertrand, Bagheri, Akbari, Yazdanian, Irani-Nejad, Mohajer et Saboori

Labidostoma (Labidostoma) intermedia Bertrand, Bagheri, Akbari, Yazdanian, Irani-Nejad, Mohajer et Saboori, 2012: 237; Jalaeian, Kamali & Bertrand, 2013: 16

Type locality: Golestan province, Gorgan

Distribution: Iran

Labidostomma (Labidostomma) mirax Bertrand et Theron

Labidostoma (Labidostoma) mirax Bertrand & Theron, 1990a: 255

Type locality: Tsitsikama, Cape Province

Distribution: South Africa

Labidostomma (Labidostomma) motasi Iavorschi

Labidostomma motasi Iavorschi, 1992: 47

Type locality: cave Peštera de la Movile, Dobroudja

Distribution: Romania

Labidostomma (Labidostomma) himalayensis Bhattacharyya

Labidostomma himalayensis Bhattacharyya, 1966: 46

Type locality: Rohla Valley

Distribution: NW Himalaya (India)

Labidostomma (Labidostomma) neotropica Stoll

Labidostoma (Labidostoma) neotropica Stoll, 1891: 9; 1993: 27; Bertrand, 1990a: 36

Type locality: Retalhuleu

Distribution: Guatemala

Labidostomma (Labidostomma) nepalense Feider et Vasiliu

Labidostomma nepalense Feider & Vasiliu, 1968a: 630

Labidostoma (Labidostoma) nepalense Feider & Vasiliu: Bertrand, 1990a: 36

Type locality: Nepal, 27o19’ N, 87o31’E

Distribution: Nepal

Labidostomma (Labidostomma) percilliata Bertrand et Theron

Labidostoma (Labidostoma) percilliata Bertrand & Theron, 1990a: 248

Type locality: Luachine, Luangue, Tchitentga

Distribution: Angola

Subgenus Nicoletiella Canestrini

Nicoletiella Canestrini, 1882: 65

Type species: Nicoletia cornuta G. Canstrini et Fanzago, 1877

Labidostomma (Nicoletiella) asiatica (Feider et Vasiliu)

Mahunkiella asiatica Feider & Vasiliu, 1972: 382

Labidostoma (Nicoletiella) asiatica Feider & Vasiliu: Bertrand, 1990a: 36

Type locality: Prov. South Phenan, Bong ha ri

Distribution: North Korea

Labidostomma (Nicoletiella) caloderum Greenberg

Labidostomma caloderum Greenberg, 1952: 197

Labidostoma (Nicoletiella) caloderum Greenberg: Bertrand, 1990a: 36

Type locality: Univ. of Kansas Natural History Reservation, Douglas County, Kansas

Distribution: USA (Kansas)

Labidostomma (Nicoletiella) coreana (Feider et Vasiliu)

Mahunkiella coreana Feider & Vasiliu, 1972: 380

Labidostoma (Nicoletiella) coreana (Feider et Vasiliu): Bertrand, 1990a: 36

Type locality: Prov. Kanwon, Kum-gang san

Distribution: N. Korea

Labidostomma (Nicoletiella) corsicum Bertrand

Labidostoma (Nicoletiella) corsicum Bertrand, 1981: 36

Type locality: Corsica

Distribution: France (Corsica)

Labidostomma (Nicoletiella) denticulatum (Schrank)

Acarus denticulatus Schrank, 1776: 125; 1781: 520

Labidostoma denticulatum : Oudemans, 1904: 132; 1916: 250; Kontschán, 2008: 361

Labidostoma (Nicoletiella) denticula [sic](Schrank): Bertrand, 1990a: 36

Labidostoma (Nicoletiella) denticulata (Schrank): Vistorin, 1978a: 279; Bertrand et al., 2015: 331

Nicoletiella denticulata (Schrank): Sellnick, 1940: 31; Willmann, 1951: 143; 1955: 217; Franz, 1954: 363; Mihelčič, 1958b: 274

Nicoletiella (Cornutella) denticulata (Schrank): Bloszyk, 1980: 72

Nicoletiella lyra Willmann, 1932: 160; 1934: 50; 1939: 433; 1941: 47; 1955: 217; Dudich, Kolosvary & Szalay, 1940: 53; Bloszyk et al., 2003: 172. Type loc.: Postoina (Slovenia).

Nicoletiella corcyraea Berlese, 1911 (Corfu)

Labidostomma (Nicoletiella) denticulatum (Schrank): Újváry & Kontschán, 2010; Pfliegler & Bertrand, 2011: 178

Type locality: Austria

Distribution: Austria, Germany, Hungary, Island, Italy, Poland, Spain, Slovenia

Labidostomma (Nicoletiella) fergusoni (Robaux)

Nicoletiella fergusoni Robaux, 1977: 443

Labidostoma (Nicoletiella) fergusoni (Robaux): Bertrand, 1990a: 36

Type locality: Mount Madona (Ridge Road)(Sta. Clara Co.), California

Distribution: USA (California)

Labidostomma (Nicoletiella) franzi Bertrand

Labidostomma franzi Bertrand, 1983: 163

Labidostoma (Nicoletiella) franzi Bertrand: Bertrand, 1990a: 36

Type locality: Galicia, La Ramosa

Distribution: Spain

Labidostomma (Nicoletiella) glymma Grandjean

Labidostomma glymma Grandjean, 1942: 119

Labidostoma (Nicoletiella) glymma Bertrand, 1990a: 36

Type locality: France

Distribution: France (cont. and Corsica),

Labidostoma (Nicoletiella) luteoides Bertrand

Nicoletiella (Nicoletiella) luteoides Bertrand, 1990c: 181

Type locality: Osorno (40o34’S, 73o09’W)

Distribution: Chile

Labidostomma (Nicoletiella) luteum Kramer

Labidostomma luteum Kramer, 1879: 13; Thor, 1931: 72; Cooreman, 1943: 2; 1959: 17; Franz, 1954: 363; Bottazzi, 1950; Schweizer & Bader, 1963: 247; Lehtinen & Niemi, 1995: 211

Labidostoma luteum Kramer: Mihelčič, 1958a: 40; 1958b: 273; Hallas, 1978: 31; Schmölzer, 1993: 58

Nicoletiella (Nicoletiella) luteum (Kramer): Vistorin, 1978: 272; Bloszyk et al., 2003: 171

Labidostoma (Nicoletiella) luteum Kramer: Bertrand, 1990a: 36

Labidostomma (Nicoletiella) luteum elongatum Grandjean, 1942: 189 (Périgueux, France). Listed by Bertrand (1990: 37) as species.

Nicoletiella vejdovskyi Štorkán, 1938 – syn. by Lehtinen & Niemi, 1995

Type locality: Germany

Distribution: Austria, Belgium, Denmark, England, Finland, France, Germany, Holland, Ireland, Italy, Sicily, Poland, Spain, Sweden, Switzerland

Labidostomma (Nicoletiella) multisetosa Shiba

Labidostoma multisetosa Shiba, 1969: 85

Labidostoma (Nicoletiella) multisetosa Shiba: Bertrand, 1990a: 36

Type locality: Shiga Heights IBP Area

Distribution: Japan

Labidostomma (Nicoletiella) pacifica (Ewing)

Ceratoacarus pacificus Ewing, 1913: 127

Nicoletiella pacifica (Ewing): Thor, 1931: 76

Labidostoma (Nicoletiella) pacifica (Ewing): Bertrand, 1990a: 36

Type locality: Oregon

Distribution: USA

Labidostomma (Nicoletiella) paleoluteum Dunlop et Bertrand (fossil)

Labidostoma (Nicoletiella) paleoluteum Dunlop & Bertrand, 2011: 193

Type locality: Baltic camber (loc. idet.), Palaeogene (Eocene)

Distribution: Baltic area

Labidostomma (Nicoletiella) repetitor Grandjean

Labidostomma luteum repetitor Grandjean, 1942: 122

Labidostomma repetitor : Lehtinen & Niemi, 1995: 220

Labidostoma (Nicoletiella) repetitor Bertrand, 1990a: 36

Labidostoma Jacquemarti Coineau, 1964: 543 – Syn. by Lehtinen & Niemi, 1995 (Type: Banyuls-sur-Mer, France)

Labidostoma (Nicoletiella) jacquemarti Coineau: Bertrand, 1990a: 36

Type locality: France

Distribution: France

Labidostomma (Nicoletiella) romanica (Feider et Vasiliu)

Nicoletiella romanica Feider & Vasiliu, 1968: 31

Labidostoma (Nicoletiella) romanica (Feider & Vasiliu): Bertrand, 1990a: 36

Type locality: the regions of Jassy, Suceava, Ploiesti, Banat, 200 – 800 m

Distribution: Romania

Labidostomma (Nicoletiella) spelaeophila (Willmann)

Nicoletiella spelaeophila Willmann, 1940: 209; Feider & Vasiliu, 1969: 205 consider “Nicoletiella speleofila [sic]” and “N. carpaticum” to be synonymes of “N. (C.) cornutum”.

L. spelaeophila (Willmann, 1940) – Cooreman, 1951: 10

Type locality: «Carniolia, Höhle bei Radna«

Distribution: Romania, Slovenia

Labidostomma (Nicoletiella) vialeae Bertrand

Labidostomma vialeae Bertrand, 1981: 28

Labidostoma (Nicoletiella) vialeae Bertrand: Bertrand, 1990a: 36

Type locality: Galeria (Corsica)

Distribution: France (Corsica)

Labidostomma virescens (Targioni Tozzetti)

Caligonas virescens Targioni Tozzetti, 1886

Labidostomma virescens (Targioni Tozzetti)

Type locality: Italy

Distribution: Italy

Subgenus Cornutella Feider et Vasiliu

Nicoletiella (Cornutella) Feider & Vasiliu, 1969: 205

Type species: Nicoletia cornutum Canestrini et Fanzago, 1877

Mahunkiella Feider & Vasiliu, 1972: 375, synonymized by Bertrand et al., 2012: 242

Labidostomma (Cornutella) cornutum (Canestrini et Fanzago)

Nicoletia cornutum Canestrini & Fanzago, 1877: 120

Nicoletiella cornuta (Canestrini et Fanzago): R. Canestrini, 1882: 65; Vistorin, 1978a: 272

Labidostomma cornuta (Canestrini & Fanzago): Cooreman, 1954: 11; 1959: 16

Nicoletiella (Cornutella) cornuta (Canestrini et Fanzago): Feider & Vasiliu, 1970: 371

Labidostomma (Nicoletiella) cornuta (Canestrini et Fanzago): Beron, 1995: 5

Nicoletiella (Cornutella) cornutum (Canestrini et Fanzago): Pfliegler & Bertrand, 2011: 178

Type locality:

Distribution: Algeria, Austria, Bulgaria, Croatia, France, Hungary, Poland, Romania, Slovenia

Labidostomma (Cornutella) ezoensis (Shiba)

Nicoletiella (Cornutella) ezoensis Shiba, 1972: 46

Type locality: Hokkaido, Kita Cirque, Mt. Poroshiri, Hidaka

Distribution: Japan (Hokkaido)

Labidostomma (Cornutella) multisetosa Shiba

Labidostoma multisetosa Shiba, 1969: 86

Mahunkiella multisetosa (Shiba): Feider et Vasiliu, 1972: 377

Labidostomma (Cornutella) multisetosa: Bertrand et al., 2012: 234, 242

Type locality: Shiga Heights (Japan)

Distribution: Japan, North Korea

The species Labidostoma corcyraeum Berlese, 1911 from “Corcyra”(Corfu) and Nicoletiella (Cornutella) absoloni Willmann, 1932 () are listed by Feider & Vasiliu, 1969, but not in the revision of Bertrand, 1990.

Subgenus Pseudocornutella Sidorchuk et Bertrand

Pseudocornutella Sidorchuk & Bertrand, 2013: 31

Type species: Labidostomma (Pseudocornutella) electri Sidorchuk et Bertrand, 2013

Labidostomma (Pseudocornutella) electri Sidorchuk et Bertrand

Labidostoma (Pseudocornutella) electri Sidorchuk & Bertrand, 2013: 31 (fossil)

Type locality: Baltic Amber

Distribution: Baltic Amber

Subgenus Pselistoma Bertrand

Pselistoma Bertrand, 1990: 183

Type species: Labidostomma (Pselistoma) legendrei Bertrand, 1990

Labidostomma (Pselistoma) legendrei Bertrand

Labidostomma (Pselistoma) legendrei Bertrand, 1990c: 183

Type locality: Osorno region

Distribution: Chile

Genus Sellnickiella Feider et Vasiliu

Sellnickiella Feider & Vasiliu, 1969: 206

Type species: Labidostoma brasiliense Sellnick, 1922

Subgenus Dicastriella Feider et Vasiliu

Dicastriella Feider & Vasiliu, 1970a: 283 (genus)

Sellnickiella (Dicastriella): Bertrand, 1990a: 35 (subgenus)

Type species: Dicastriella covarubiasi Feider et Vasiliu, 1970

Sellnickiella (Dicastriella) caeca (Feider, Vasiliu et Calugar)

Dicastriella caeca Feider, Vasiliu & Calugar, 1974: 420

Sellnickiella (Dicastriella) caeca (Feider, Vasiliu et Calugar): Bertrand, 1990c: 35

Type locality: Misitumi (reg. Tarapaca)

Distribution: Chile

Sellnickiella (Dicastriella) coineaui (Feider et Vasiliu)

Dicastriella coineaui Feider & Vasiliu, 1970a: 303; Feider, Vasiliu & Calugar, 1974: 422

Sellnickiella (Dicastriella) coineaui (Feider et Vasiliu): Bertrand, 1990c: 35

Type locality: Polpaico (Santiago)

Distribution: Chile

Sellnickiella (Dicastriella) covarubiasi (Feider et Vasiliu)

Dicastriella covarubiasi Feider & Vasiliu, 1970a: 297; Feider, Vasiliu & Calugar, 1974: 423

Sellnickiella (Dicastriella) covarubiasi (Feider et Vasiliu): Bertrand, 1990c: 35

Type locality: Limpache (Valparaiso)

Distribution: Chile

Sellnickiella (Dicastriella) fungifera (Feider et Vasiliu)

Dicastriella fungifera Feider & Vasiliu, 1970a: 306

Sellnickiella (Dicastriella) fungifera (Feider et Vasiliu): Bertrand, 1990c: 35

Type locality: Limache (Valparaiso)

Distribution: Chile

Sellnickiella (Dicastriella) fusiformis Bertrand

Labidostomma (Dicastriella) fusiformis Bertrand, 1990c: 179

Type locality: Osorno (40o34’S, 73o09’W)

Distribution: Chile

Sellnickiella (Dicastriella) mediohexagonata (Feider, Vasiliu et Calugar)

Dicastriella mediohexagonata Feider, Vasiliu & Calugar, 1974: 419

Sellnickiella (Dicastriella) mediohexagonata (Feider, Vasiliu et Calugar): Bertrand, 1990c: 35

Type locality: Farellones (Prov. Santiago)

Distribution: Chile

Subgenus Sellnickiella Feider et Vasiliu

Sellnickiella Feider & Vasiliu, 1969: 206

Type species: Labidostoma brasiliense Sellnick, 1922

Sellnickiella (Sellnickiella) adelaideae (Womersley)

Labidostomma adelaideae Womersley, 1935: 152; Atyeo & Crossley, 1961b: 83

Sellnickiella (Sellnickiella) adelaidae (Womersley): Bertrand, 1990a: 35

Sellnickiella adelaideae (Womersley): Halliday, 1998: 66

Type locality: Adelaida

Distribution: Australia

Sellnickiella (Sellnickiella) brasiliense (Sellnick)

Labidostoma brasiliense Sellnick, 1922: 45

Sellnickiella brasiliense (Sellnick): Feider & Vasiliu, 1969: 206; 1970: 289

Sellnickiella (Sellnickiella) brasiliensis (Sellnick): Bertrand, 1990a: 35

Type locality: Passa Quatro, Sul de Minas

Distribution: Brazil

Sellnickiella (Sellnickiella) capensis Bertrand et Theron

Sellnickiella (Sellnickiella) capensis Bertrand & Theron, 1992: 46

Type locality: Hawaquas Montains nr. Paarl, Cape Province

Distribution: South Africa

Sellnickiella (Sellnickiella) circinus (Atyeo et Crossley)

Labidostoma circinus Atyeo & Crossley, 1961b: 35

Sellnickiella (Sellnickiella) circinus (Atyeo et Crossley): Bertrand, 1990a: 36

Type locality: Ngongotaha, Rotorua, Auckland

Distribution: New Zealand

Sellnickiella (Sellnickiella) decemlabeculata Feider et Vasiliu

Sellnickiella decemlabeculata Feider & Vasiliu, 1970: 297; Feider, Vasiliu & Calugar, 1974: 418

Sellnickiella (Sellnickiella) decemlabeculata: Bertrand, 1990c: 35

Type locality: Brazil, Nova Teutonia

Distribution: Brazil, Chile

Sellnickiella (Sellnickiella) hoegi (Thor)

Labidostoma höegi Thor, 1931a: 57

Labidostomma höegi Thor, 1931b: 73; Meyer & Ryke, 1959: 419

Sellnickiella (Sellnickiella) hoegi (Thor): Bertrand, 1990a: 35

Type locality: Port Elizabeth

Distribution: South Africa

Sellnickiella (Sellnickiella) macroscopa Bertrand

Sellnickiella (Sellnickiella) macroscopa Bertrand, 1990c: 179

Type locality: Cantin (39o22’ S, 72o38’W)

Distribution: Chile

Sellnickiella (Sellnickiella) mediumreticulata Feider et Vasiliu

Sellnickiella mediumreticulata Feider & Vasiliu, 1970a: 289; Feider, Vasiliu & Calugar, 1974: 417

Type locality: Nova Teutonia

Distribution: Brazil

Sellnickiella (Sellnickiella) natalensis Bertrand et Theron

Sellnickiella (Sellnickiella) natalensis Bertrand & Theron, 1992: 48

Type locality: St. Lucia (Natal)

Distribution: South Africa

Sellnickiella (Sellnickiella) pulchra Bertrand

Sellnickiella (Sellnickiella) pulchra Bertrand, 1990c: 178

Type locality: Floresta de Tijuca, reg. of Rio de Janeiro

Distribution: Brazil

Sellnickiella (Sellnickiella) schusteri Feider, Vasiliu et Calugar

Sellnickiella schusteri Feider, Vasiliu & Calugar, 1974: 414

Sellnickiella (Sellnickiella) schusteri Feider, Vasiliu et Calugar: Bertrand, 1990a: 35

Type locality: Serra de Cantareira (Sao Paulo)

Distribution: Brazil

Sellnickiella (Sellnickiella) womersleyi (Atyeo et Crossley)

Labidostomma womersleyi Atyeo & Crossley, 1961a: 84

Type locality: Remarkable Creek, Wilmington, South Australia

Distribution: Australia

Sellnickiella (Subgen.?) balticae Sidorchuk et Bertrand (fossil)

Sellnickiella balticae Sidorchuk & Bertrand, 2013: 27

Type locality: Baltic Amber

Distribution: Baltic Amber

Distribution of Labidostommatidae

EUROPE

Austria (Schrank, 1781; Willmann, 1939, 1951; Franz, 1954; Vistorin, 1978; Schmölzer, 1993) –Labidostomma cornuta, L. denticulatum, L. luteum, L. lyra

Bulgaria (Beron, 1995) – Eunicolina nova, Labidostomma (Cornutella) cornutum

Croatia (Willmann, 1940, 1941; Vistorin, 1978а) – Eunicolina tuberculata, Labidostomma (Cornutella) cornutum

Czech Rep. (Willmann, 1939)

Denmark (Luxton, 1966, 1981; Hallas, 1978) – Labidostomma luteum

Finland (Lehtinen & Niemi, 1995) – Labidostomma (L.) luteum

France (contin.) (Cooreman, 1954, 1959; Coineau, 1964a, 1964b; Bertrand, 1980; Bertrand & Coineau, 1979) – Akrostomma coralloides, Labidostomma (Cornutella) cornutum, Labidostomma (Nicoletiella) repetitor (= L. (Nicoletiella) jacquemarti), L. (N.) luteum, Labidostomma (L.) integrum

Corsica (Bertrand, 1981; Bertrand et al., 2012) - Labidostomma (Nicoletiella) corsicum, L. (N.) glymma, L. (N.) vialeae, Labidostomma (L.) integrum

Georgia (Reck, 1940) – Labidostomma caucasicum (= L. integrum)

Germany (C.L.Koch, 1838; Oudemans, 1916; Willmann, 1955; Vistorin, 1978a) – Labidostomma denticulatum, L. luteum, L. lyra

Great Britain (Michael, 1880) – Labidostomma luteum

Corfu (Berlese, 1911; Sellnick, 1931) – “Labidostoma corcyraeum””(= L. denticulatum), Eunicolina nova

Hungary (Willmann, 1937; Dudich, Kolosváry & Szalay, 1940; Újváry & Kontschán, 2010; Pfliegler & Bertrand, 2011) – Labidostomma denticulata, L. lyra, L. (Cornutella) cornutum

Iceland (Sellnick, 1940) – Labidostomma denticulata (actually L. luteum, see Lehtinen & Niemi, 1995)

Ireland (Halbert, 1915) – Nicoletiella cornuta (actually L. luteum, see Lehtinen & Niemi, 1995)

Italy (contin.) (Berlese, 1887; 1911; Lombardini, 1943; Bottazzi, 1956; Vistorin, 1978a) – Akrostomma zangherii (sp. dubia), Eunicolina tuberculata, Labidostomma denticulatum, L. integrum, L. luteum

Sicily (Berlese, 1883; Bottazzi, 1956) – Labidostomma luteum

Sardinia (Bertrand et al., 2012) – Labidostomma repetitor, L. (L.) integrum

Poland (Bloszyk, 1980; Błoszyk & Czarnota, 1998; Bloszyk, Konwerski, Napierala, Goldyn & Gawronska, 2007; Bloszyk et al., 2017; Bloszyk, Adamski & Napierała, 2018) – Labidostomma (Nicoletiella) denticulatum, L. (N.) luteum, L. (Cornutella) cornutum

Romania (Cooreman, 1951; Feider & Vasiliu, 1968, 1970; Iavorschi, 1992; Kontschán, 2008) - Labidostomma motasi, L. (Cornutella) cornutum, L. (L.) integrum (= L. caucasicum), L.(Nicoletiella) denticulata, L. (Nicoletiella) romanica, Eunicolina [Grandjeanellina] nova

Slovakia (Bloszyk et al., 2022) – Labidostomma luteum

Slovenia (Willmann, 1932; Willmann, 1940, 1941; Vistorin, 1978a) – Labidostomma (Nicoletiella) lyra, Labidostomma cornuta, L. spelaeophila

Spain (cont.) (Mihelčič, 1958a,b; Bertrand, 1983 ; Bertrand et al., 2012) – Akrostomma coineaui, Labidostomma (Nicoletiella) luteum, [L. colomerense], L. franzi, L. (L.) integrum

Sweden - Labidostomma (Nicoletiella) luteum

Switzerland (Schweizer & Bader, 1963) – Labidostomma (Nicoletiella) luteum

Turkey (all) (Urhan & Özmen, 2006) – Labidostomma sp.

Baltic Area (Dunlop & Bertrand, 2011; Sidorchuk et Bertrand, 2013) - Labidostomma (Nicoletiella) paleoluteum (fossil), Labidostomma (Pseudocornutella) electri (fossil), Sellnickiella balticae (fossil)

AFRICA

Algeria (Grandjean, 1942) – Labidostomma integrum, L. (Cornutella) cornutum

Angola (Bertrand & Theron, 1990a) – Labidostomma (L.) angolensis, L. (L.) percilliata

DR Congo (Cooreman, 1954) – Labidostomma integrum schoutedeni

South Africa (Thor, 1931a; Meyer & Ryke, 1959; Bertrand & Theron, 1990a, 1992) - Labidostomma (L.) aethiopica, L. (L.) fissurata, L.(L.) mirax, L.(L.) percilliata, Sellnickiella (S.) capensis, S.(S.) hoegi, S.(S.) natalensis

ASIA

India (Bhattacharyya, 1966) – Labidostomma himalayensis

Iran (Bertrand et al., 2012; Jalaeian, Kamali & Bertrand, 2013) – Labidostomma (L.) intermedia

Israel and Palestina - Labidostomma (L.) intermedia

Japan (Shiba, 1969, 1972, 2004) – Labidostomma (Atyonella) simplex, Labidostomma (Cornutella) ezoensis, L. (Cornutella) mutisetosa

Kazakhstan (Bertrand et al., 2012) - Labidostomma (L.) integrum

Korea (N.) (Feider & Vasiliu, 1972) – Labidostomma asiatica, L. (Cornutella) mutisetosa, M. multisetosa

Nepal (Feider & Vasiliu, 1968a) - Labidostomma nepalense

Philippines (Bertrand & Corpus Raros, 1997) - Labidostomma (L.) guadalupense

NORTH AMERICA

USA (Ewing, 1913; Greenberg, 1952; Robaux, 1977; Crossley & Simmons, 2015) - Akrostomma gabriella, A. grandjeani, A. plumosum, Eunicolina porifera, Labidostomma (L.) barbae, Labidostomma (Nicoletiella) caloderum, L. (N.) fergusoni, L. (N.) pacifica,

SOUTH and CENTRAL AMERICA

Brazil (Sellnick, 1922; Feider & Vasiliu, 1970; Feider, Vasiliu & Calugar, 1974; Bertrand, 1990) – Labidostomma amasonica, Sellnickiella (S.) brasiliensis, S.(S.) decemlabeculata, S. (S.) mediumreticulata, S. (S.) schusteri, S. (S.) pulchra

Chile (contin.) (Feider & Vasiliu, 1970; Feider, Vasiliu & Calugar, 1974; Bertrand, 1990c) – Labidostomma (Nicoletiella) luteoides, L. (Pselistoma) legendrei, Sellnickiella (S.) decemlabeculata, Sellnickiella (Dicastriella) caeca, S. (D.) coineaui, S. (D.) covarubiasi, S. (D.) fungifera, S.(D.) fusiformis, S. (D.) mediohexagonata, Sellnickiella (S.) macroscopa,

Guatemala (Stoll, 1891) – Labidostomma (Labidostomma) neotropica

Peru (Bertrand, 1990c) – Labidostomma (Atyeonella) peruviana

OCEANIA

Australia (continent) (Wommersley, 1935; Atyeo & Crossley Jr., 1961a; Bertrand, 1997; Halliday, 1998) – Sellnickiella adelaideae, S. biunguiculata, S. womersleyi

New Zealand (Atyeo & Crossley Jr. 1961b) – Sellnickiella (S.) circinus, Labidostomma Atyonella fictiluteum, L. (A.) glandula, L. (A.) malleolus, L. (A.) multifarium, L. (A.) ocellatum, L. (A.) striatum

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Infraorder Eupodina

The Mite Superfamily Eupodoidea

Eupodoidea Banks, 1894

According to Zhang, Ed. (2011), the superfamily consists of nine families (four of them described recently), with 85 genera and 309 species. The four “old” families (Eupodidae, Penthaleidae, Penthalodidae and Rhagidiidae) account for 277 of these species.

The Mite Family Eupodidae Koch

Eupodides C.L. Koch, 1842: 60

Eupodidae: Murray, 1877: 117

Small or middle-sized (160 – 1000 µm) mites, living in moss, leaf litter or on plants. Predators or plant parasites. Members of Benoinyssus Fain have been found in the nasal mucus of Naja melanoleuca (Serpentes: Elapidae).

Longtime after the descriptions of genus Eupodes Koch, 1835 and the family Eupodidae again by C.L. Koch (1842), the family was left with this genus plus Protereunetes Berlese, 1923, Linopodes C.L. Koch, 1835 and later Cocceupodes Sig Thor, 1934 (the last two being considered after 2010 to belong to a separate family Cocceupodidae Jesionowska, later synonymized with Eupodidae and restaured). Followed the description of five further genera by Fain (1958), A. Baker (1987), Barilo (1991), Fain & Camerik (1994) and Baker & Lindquist (2002). The important revision of Thor & Willmann (1941) was followed by modern contributions by Meyer & Ryke (1960, South Africa), Strandtmann (1971, Alasca), Strandtmann & Goff (1978, Hawaii), Shiba (1978, Japan), Qin Ting-Kui (1998, Australia and New Zealand), Olivier & Theron (1997, 1998, Southern Africa), and others. Several other authors contributed considerably to the study of these mites: Thor in Norway, Strandtmann in Antarctica, Abou-Awad in Egypt, Corpuz-Raros in the Philippines. Most of the modern research is being done in countries outside Europe and the identity of the 26 “Eupodes” species, described by C.L. Koch from Germany and enumerated by Thor & Willmann (1941), remains unclear. Only eight of them have been discussed by modern authors.

Cocceupodidae is a small family of mites, detached from Eupodidae and consisting of three genera and 23 valid species (Jesionowska, 2010). The study of the family starts with Acarus [Linopodes] motatorius Linnaeus, 1758. Linopodes C.L. Koch, 1835 is the oldest genus, includes mites with very long front legs. They live in wet places, rotten wood and under stones, often in caves. Koch (1838) describes from Germany 12 “Linopodes” species, their identity is still unclear. Some Cocceupodes are known from the far North or from Antarctica.

Khaustov (2014) synonymized Cocceupodidae under Eupodidae and Filieupodes under Cocceupodes and described from Crimea a new genus and species Pseudeupodes porosus. Later Khaustov (2015) added another two new genera and species Pseudopenthaleus tauricus and Echinoeupodes echinus, thus the families Eupodidae and Cocceupodidae (combined) seem to include 14 genera and 95 species. Other Polish specialists, based on molecular phylogeny, restaured the validity of the family Cocceupodidae and of the genus Filieupodes Jesionowska. They concluded that “(1) the genera Linopodes, Filieupodes and Cocceupodes are a separate monophyletic lineage of familial rank and the sister group of Eupodidae s.s., and (2) the genus Filieupodes should be considered a separate genus being a sister group of the Linopodes-Cocceupodes clade” (Szudarek-Trepto, Kaźmierski, Dabert et al. 2020).

Composition of the families Eupodidae and Cocceupodidae

Fam. Eupodidae C.L. Koch, 1842 - 14 gen., 95 sp.

Aethosolenia Baker et Lindquist, 2002 – 1 sp.

Benoinyssinae Fain, 1958 – 10 sp.

Caleupodes A. Baker, 1987 – 1 sp.

Claveupodes Strandtmann et Prasse, 1977 – 1 sp.

Cocceupodes Sigh Thor, 1934 – 8 sp.

Echinoeupodes Khaustov, 2017 – 2 sp.

Eupodes C.L. Koch, 1835 – 47 sp.

Filieupodes Jesionowska, 2010 – 9 sp.

Linopodes C.L. Koch, 1835 - 10 sp.

Neoprotereunetes Fain et Camerik, 1994 – 1 sp.

Niveupodes Barilo, 1991 – 1 sp.

Pseudeupodes Khaustov, 2014 – 1 sp.

Pseudopenthaleus Khaustov, 2015 – 2 sp.

Xerophiles Jesionowska, 2003 – 1 sp.

Catalogue of Eupodidae and Cocceupodidae

Family Eupodidae C.L. Koch

Eupodides C.L. Koch, 1842: 60

Eupodini Canestrini, 1886: 693

Eupodidae Murray, 1877: 117

Type genus: Eupodes C.L. Koch, 1835

Benoinyssinae Fain, 195. Type genus: Benoinyssus Fain, 1958: 178.

Cocceupodidae Jesionowska, 2010: 639. Synonymized by Khaustov, 2014, but restored by Szudarek-Trepto, Kaźmierski, Dabert et al. (2020)

Type genus: Cocceupodes Sig Thor, 1934 (sensu Jesionowska, 2005)

Genus Aethosolenia Baker et Lindquist

Aethosolenia Baker & Lindquist, 2002: 2

Type species: Aethosolenia laselvensis Baker et Lindquist, 2002

Aethosolenia laselvensis Baker et Lindquist

Aethosolenia laselvensis Baker & Lindquist, 2002: 4

Type locality: Heredia Province, La Selva Biological Station, 50-150 m

Distribution: Costa Rica

Genus Benoinyssus Fain

Benoinyssus Fain, 1958: 178

Type species: Benoinyssus najae Fain, 1958

Benoinyssus celatus Olivier et Theron

Benoinyssus celatus Olivier & Theron, 1997a: 317

Type locality: Cafunfe river

Distribution: Angola

Benoinyssus ereynetoides (Strandtmann et Prasse)

Eupodes ereynetoides Strandtmann & Prasse, 1976: 2; Santamaría et al., 2012: 1164

Benoinyssus ereynetoides (Strandtmann et Prasse): Olivier & Theron, 1997a: 302

Type locality: Etzsdorf/ Saalkreis (East Germany)

Distribution: Germany, Spain

Benoinyssus falcatus Olivier et Theron

Benoinyssus falcatus Olivier & Theron, 1997a: 311

Type locality: Northern Province, Louis Trichardt

Distribution: South Africa

Benoinyssus momeni (Abou-Awad)

Eupodes momeni Abou-Awad, 1984: 333

Benoinyssus momeni (Abou-Awad): Olivier & Theron, 1997a: 302

Type locality: Ain Shams Univ., El-qaliubiya Region

Distribution: Egypt

Benoinyssus najae Fain

Benoinyssus najae Fain, 1958: 138; 1959: 358

Type locality: Kaninya

Distribution: Rwanda

Host: Naja melanoleuca (Serpentes: Elapidae), nasal mucus

Benoinyssus oconnori Bizarro Wurlitzer et Da Silva

Benoinyssus oconnori Bizarro Wurlitzer & Da Silva, 2020: 3

Type locality: municipality of Novo Mundo, Mato Grosso state

Distribution: Brazil

Host: from soil and litter

Benoinyssus ornatus Olivier et Theron

Benoinyssus ornatus Olivier & Theron, 1997a: 317

Type locality: Northern Province, Giyani (South Africa)

Distribution: Angola, Swaziland, South Africa

Benoinyssus pusillus Olivier et Theron

Benoinyssus pusillus Olivier & Theron, 1997a: 314

Type locality: Northern Province, Warmbaths

Distribution: South Africa

Benoinyssus serratus Olivier et Theron

Benoinyssus serratus Olivier & Theron, 1997a: 302

Type locality: Northern Province, Turfloop (South Africa)

Distribution: Botswana, Namibia, South Africa, Swaziland

Benoinyssus sp. – Iran (Darbemamieh, Hajiqanbar & Khanjani, 2013: 454)

Benoinyssus tuberculatus (Baker)

Opsereynetes tuberculatus Baker, 1945: 18

Benoinyssus tuberculatus (Baker): Hunter, 1964: 181

Ereynetes (Opsereynetes) tuberculatus Baker: Fain, 1964b: 106

Type locality: Desierto de los Leones

Distribution: Mexico

Genus Caleupodes A. Baker

Caleupodes A. Baker, 1987: 103

Type species: Caleupodes reticulatus A. Baker, 1987

Caleupodes reticulatus A. Baker

Caleupodes reticulatus A. Baker, 1987: 104

Type locality: Suffolk, Hollesley Common

Distribution: England

Genus Claveupodes Strandtmann et Prasse

Claveupodes Strandtmann & Prasse, 1977: 10

Type species: Claveupodes delicatus Strandtmann et Prasse, 1977

Claveupodes delicatus Strandtmann et Prasse

Claveupodes delicatus Strandtmann & Prasse, 1977: 11; Martin, 1983: 15

Type locality: Etzdorf/Saalkreis, East Germany

Distribution: Germany, New Zealand

Claveupodes sp. - Iran (Darbemamieh, Hajiqanbar & Khanjani, 2013: 454)

Genus Cocceupodes Sig Thor

Cocceupodes Sig Thor, 1934: 128 (sensu Jesionowska, 2005)

Type species: Eupodes clavifrons R. Canestrini, 1886

Cocceupodes australis Strandtmann et Tilbrook

Cocceupodes australis Strandtmann & Tilbrook, 1968: 51; Shiba, 1971: 222; Jesionowska, 2010: 643. Strandtmann & Prasse (1977: 9) synonymize it with C. mollicellus.

Type locality: Deception Island

Distribution: Deception Island (South Shetland Islands), Japan

Cocceupodes breweri Strandtmann

Cocceupodes breweri Strandtmann, 1971: 87; Jesionowska, 2010: 641

Type locality: Anaktuvik Pass, Alaska

Distribution: USA (Alaska), Greenland

Cocceupodes communis Shiba

Cocceupodes communis Shiba, 1969c: 68; Jesionowska, 2010: 643

Type locality: Shiga Heights IBP Area, Nagano Pref.

Distribution: Japan

Cocceupodes gracilongus Olivier et Theron

Cocceupodes gracilongus Olivier & Theron, 2003: 186; Jesionowska, 2010: 641

Type locality: S. Afr., Northern Province, Ga-Selati River, near Tzaneen

Distribution: South Africa

Cocceupodes longisolenidiatus Jesionowska

Cocceupodes longisolenidiatus Jesionowska, 2007: 352; 2010: 641

Type locality: Poland

Distribution: Poland

Cocceupodes mollicellus (C.L. Koch)

Eupodes mollicellus C.L. Koch, 1838: fa. 19, t. 23

Eupodes clavifrons R. Canestrini, 1886: 716 (type loc. Trentino); Haarløv, 1942 : 28

Cocceupodes clavifrons (R. Canestrini): Sig Thor, 1934: 128; Schweizer & Bader, 1963: 218; Strandtmann, 1971: 85

Cocceupodes mollicellus : Oudemans, 1937: 891; Thor & Willmann, 1941: 36; Mihelčič, 1957;

Macfadyen, 1954: 283; Haarløv, 1957: 26; Strandtmann & Prasse, 1977: 8; Strandtmann &

Goff, 1978: 133; Jesionowska, 2010: 641

Cocceupodes curticlava Thor, 1934: 178; Thor & Willmann, 1941: 36; Macfadyen, 1954: 283;

Haarløv, 1957: 27 (syn. by Strandtmann & Prasse, 1977: 8)

Type locality: Germany

Distribution: Austria, Denmark, Germany, Italy, Norway, Hawaii, Greenland, Switzerland, Yan Mayen I.

Cocceupodes planiticus Shiba

Cocceupodes planiticus Shiba, 1978: 141; Jesionowska, 2010: 643

Type locality: Japan

Distribution: Japan

Cocceupodes stellatus Strandtmann et Prasse

Cocceupodes stellatus Strandtmann & Prasse, 1977: 6; Jesionowska, 2010: 641

Type locality: Etzsdorf/ Saalkreis (East Germany)

Distribution: Germany, Greenland (cf. stellatus)

Genus Echinoeupodes Khaustov

Echinoeupodes Khaustov, 2017: 29

Type species: Echinoeupodes Khaustov, 2017: 29

Echinoeupodes echinus Khaustov

Echinoeupodes echinus Khaustov, 2017: 31

Type locality: nr. Yalta

Distribution: Crimea

Echinoeupodes turgidus (Shiba)

Protereunetes turgidus Shiba, 1978: 145

Echinoeupodes turgidus (Shiba): Khaustov, 2017: 31

Type locality: Gobo, Wakayama

Distribution: Japan

Genus Eupodes C.L. Koch

Eupodes C.L. Koch, 1835: fa. 1, t. 18 - 19

Megamerus Dugès, 1834: 50 (non Mac Leay, 1827). Enumerated are: longipes, inflatus, ovalis, celer, roseus, castaneus, fallax

Egypteupodes B.A. Abou-Awad, 1984: 329 (syn. by Olivier & Theron,1997: 58). Type: E. strandtmanni Abou-Awad, 1984

Type species: Eupodes striola C.L. Koch, 1835

Eupodes acanthus Olivier et Theron

Eupodes acanthus Olivier & Theron, 1998: 284

Type locality: Northern Province, Debegeni Falls

Distribution: South Africa

Eupodes alaskanensis Strandtmann

Eupodes alaskanensis Strandtmann, 1971: 79

Type locality: Alaska, Cape Beaufort

Distribution: USA (Alaska)

Eupodes angardi Strandtmann et Sømme

Eupodes angardi Strandtmann & Sømme, 1977: 138

Type locality: Queen Maud Land, Brattskarvet

Distribution: Antarctica

Eupodes bakeri Abou-Awad, El-Sawaf et Abdel-Khalek

Eupodes bakeri Abou-Awad, El-Sawaf & Abdel-Khalek, 2006: 46

Type locality: Egypt

Distribution: Egypt

Eupodes berensis Shiba

Eupodes berensis Shiba, 1976: 85

Type locality: Malaysia

Distribution: Malaysia

Eupodes berlesei Thor

Eupodes berlesei Thor, 1912: 236; Schweizer & Bader, 1963: 217

Type locality: Italy

Distribution: Denmark, Italy, Switzerland

Eupodes brevipes Banks

Eupodes brevipes Banks, 1916: 14

Type locality: Laguna Beach, California

Distribution: USA (California)

Eupodes callejae Corpuz- Raros

Eupodes callejae Corpuz- Raros, 2005: 135

Type locality: Baybay, Leyte Is.

Distribution: Philippines (Leyte Is., Luzon Is.)

Eupodes cordiformis Shiba

Eupodes cordiformis Shiba, 1978: 135

Type locality: Kuwabara, Matsuyama, Ehime

Distribution: Japan

Eupodes crozetensis Strandtmann et Davies

Eupodes crozetensis Strandtmann & Davies, 1972: 50; Darbemamieh, Hajiqanbar & Khanjani, 2013: 454

Type locality: French Base Buildings, Possession Island, Crozet Islands

Distribution: Iran, Possession Island, Crozet Islands (sub-antarctic)

Eupodes exiguus Booth, Edwards et Usher

Eupodes exiguus Booth, Edwards & Usher, 1985: 392

Type locality: South-Orkney-Islands

Distribution: Antarctica (South-Orkney-Islands)

Eupodes fusifer R. Canestrini

Eupodes fusifer R. Canestrini, 1886: 714

Type locality: San Remo, Veneto

Distribution: Italy

Eupodes fusiferellus Meyer et Ryke

Eupodes fusiferellus Meyer & Ryke, 1960: 483

Type locality: Bathurst

Distribution: South Africa

Eupodes garciai Corpuz- Raros

Eupodes garciai Corpuz- Raros, 2005: 137

Type locality: Luzon (UPLB campus, College, Laguna)

Distribution: Philippines (Luzon, Leyte Isls)

Eupodes ghardagae Schuster

Eupodes ghardagae Schuster

Type locality: Egypt

Distribution: Egypt

Eupodes gilvus C.L. Koch

Eupodes gilvus C.L. Koch, 1838: fa. 20, t. 1

Type locality: Regensburg

Distribution: Germany

Eupodes haltica (Haller)

Megamerus haltica Haller, 1880: 364

Eupodes haltica (Haller): Thor & Willmann, 1941: 25

Type locality: “Moos am Thunersee”

Distribution: Switzerland

Eupodes hamatus Olivier et Theron

Eupodes hamatus Olivier & Theron, 1997: 72

Type locality: Western Cape, Lamberts Bay

Distribution: South Africa

Eupodes hawaiiensis Strandtmann et Goff

Eupodes hawaiiensis Strandtmann & Goff, 1978: 129; Goff, 1987: 31

Type locality: Hawaii Isl., Kilauea Forest Reserve, 1620 m

Distribution: Hawaii

Eupodes hiemalis C.L. Koch

Eupodes hiemalis C.L. Koch, 1838: fa. 19, t. 4; Thor & Willmann, 1941: 20; Lindroth et al., 1973: 125

Type locality: Regensburg (Germany)

Distribution: Germany, Iceland

Eupodes hjartdaliae Thor

Eupodes hjartdaliae Thor, 1934a: 213

Type locality: Hjartdal, Thelemarken

Distribution: Norway

Eupodes iconicus C.L. Koch

Eupodes iconicus C.L. Koch, 1838: fa.19, t. 7; 1842: fa. 3, p.69; Walckenaer, 1847: 537; Oudemans, 1937: 884; Thor & Willmann, 1941: 12

Type locality: Regensburg (Germany)

Distribution: Germany, Italy

Eupodes indentatus Olivier et Theron

Eupodes indentatus Olivier & Theron, 1997: 69

Type locality: Mt. Inn Hotel, Soutpansberg

Distribution: South Africa

Eupodes infirmus Shiba

Eupodes infirmus Shiba, 1978: 133

Type locality: Mt. Saragamine, Shigenobu-cho, Ehime

Distribution: Japan

Eupodes leucomelas C.L. Koch

Eupodes leucomelas C.L. Koch, 1838: fa.19, t.14

Type locality: Regensburg

Distribution: Germany

Eupodes littoralis Shiba

Eupodes littoralis Shiba, 1978: 141

Type locality: Wake, Matsuyama, Ehime

Distribution: Japan

Eupodes longipilus Thor

Eupodes longipilus Thor, 1934: 178; Meyer & Ryke, 1960: 487

Type locality: Cape Town

Distribution: South Africa

Eupodes longisetatus Strandtmann

Eupodes longisetatus Strandtmann, 1964: 148; Shiba, 1969c: 66; 1969a: 149

Type locality: Campbell Island

Distribution: Japan, New Zealand (Campbell Island)

Eupodes lumbarus Shiba

Eupodes lumbarus Shiba, 1976: 88

Type locality: Malaysia

Distribution: Malaysia

Eupodes lutatus Olivier et Theron

Eupodes lutatus Olivier & Theron, 1997: 72

Type locality: Zululand, St Lucia

Distribution: South Africa

Eupodes marinus Banks

Eupodes marinus Banks, 1896: 75; 1907: 596; Thor, 1934: 206; Thor & Willmann, 1941: 27

Type locality: Sea Cliff, New York

Distribution: USA

Eupodes milvinus C.L. Koch

Eupodes milvinus C.L. Koch, 1838: fa. 19, t.1; 1842: fa. 3, p.70; Walckenaer, 1847: 537; Oudemans, 1937: 890; Thor & Willmann, 1941: 22

Type locality: Oberpfalz

Distribution: Germany, Italy, Holland, Sweden

Eupodes minipilus Olivier et Theron

Eupodes minipilus Olivier & Theron, 1998: 281

Type locality: Northern Province, Giyani (South Africa)

Distribution: Botawana, Cabo Verde Islands, South Africa, Zimbabwe

Eupodes minutus (Strandtmann)

Protereunetes minutus Strandtmann, 1967: 55

Eupodes minutus (Strandtmann): Strandtmann, 1970: 89; Luxton, 1990: 400

Tydeus antipodus : Womersley (1937)

Eupodes antipodus : Baker (1970)

Type locality: South Georgia, Bay of Isles

Distribution: South Georgia, Macquarie I., New Zealand (South Island)

Eupodes niloticus Abou-Awad et El-Bagouri

Eupodes niloticus Abou-Awad & El-Bagouri, 1985: 5; Abdelgayed, 2019: 32

Type locality: Egypt

Distribution: Egypt

Eupodes ocellatus Willmann

Eupodes ocellatus Willmann, 1952: 159

Type locality: Wanderooge I.

Distribution: Germany

Eupodes oedipus Oudemans, 1937: 888 (nom. nov. pro Eupodes macropus Koch, 1838)

Eupodes okinoshimaensis Shiba

Eupodes okinoshimaensis Shiba, 1978: 138

Type locality: Nakamura, Okinoshima, Shimane

Distribution: Japan

Eupodes onoi Shiba

Eupodes onoi Shiba, 1976: 92

Type locality: Malaysia

Distribution: Singapore, Malaysia

Eupodes parafusifer Meyer et Ryke

Eupodes parafusifer Meyer & Ryke, 1960: 483; Olivier & Theron, 1998: 276

Type locality: Mooi River, Potchefstroom (South Africa)

Distribution: Angola, South Africa

Eupodes parvus Booth, Edwards et Usher

Eupodes parvus Booth, Edwards & Usher, 1985: 394

Type locality: South-Orkney-Islands

Distribution: Antarctica (South-Orkney-Islands)

Eupodes parvus grahamensis Booth, Edwards et Usher

Eupodes parvus grahamensis Booth, Edwards & Usher, 1985: 395

Type locality: Antarctica, Takaki Promontory, Brabant Island

Distribution: Antarctica

Eupodes pasohensis Shiba

Еupodes pasohensis Shiba, 1976: 86

Type locality: Pasoh Forest Reserve, Negeri Sembilan, W. Malaysia

Distribution: Malaysia

Eupodes pseudoclavifrons R. Canestrini

Eupodes pseudoclavifrons R. Canestrini, 1886: 717; Thor, 1934: 205; Schmölzer, 1993: 59

Cocceupodes pseudoclavifrons (R. Canestrini): Schweizer & Bader, 1963: 218

Type locality: Italy (Trentino)

Distribution: Austria, Italy, Switzerland

Eupodes pseudolumbarus Shiba

Eupodes pseudolumbarus Shiba, 1976: 90; Corpuz-Raros, 2005: 140

Type locality: Pasoh Forest Reserve, Negeri Sembilan, W. Malaysia

Distribution: W.Malaysia, Philippines (Luzon, Leyte)

Eupodes pusillus Shiba

Eupodes pusillus Shiba, 1976: 93; Corpuz-Raros, 2005: 143c

Type locality: Pasoh Forest Reserve, Negeri Sembilan, W. Malaysia

Distribution: W.Malaysia, Philippines (Luzon, Leyte)

Eupodes riedli Schuster

Eupodes riedli Schuster, 1965

Type locality: Possession I.

Distribution: Possession I., (Crozet Isls, Antarctica)

Eupodes sigmoidensis Strandtmann et Goff

Eupodes sigmoidensis Strandtmann & Goff, 1978: 123; Goff, 1987: 31; Gupta et al., 2004: 44 Darbemamieh, Hajiqanbar & Khanjani, 2013: 454

Type locality: Hawaii I., east slope of Mauna Loa

Distribution: Hawaii, India, Iran, Ivory Coast

Eupodes signatus C.L. Koch

Eupodes signatus C.L. Koch, 1835: fasc. 1, t.19

Type locality: Pleistein

Distribution: Deutschland

Eupodes skiaaki Thor

Eupodes skiaaki Thor, 1934: 213

Type locality: Linseim, Grjotli, Videsaeter in Skiaak (Gudbrandsdalen)

Distribution: Norway

Eupodes strandtmanni Coineau

Eupodes strandtmanni Coineau, 1976: 56

Type locality: Banyuls-sur-Mer

Distribution: France

Eupodes striola C.L. Koch

Eupodes striola C.L. Koch, 1836: fa.1, t.18

Type locality: Germany

Distribution: Germany

Eupodes temperatus Shiba

Eupodes temperatus Shiba, 1978: 135; Abou-Awad, 1984: 335

Type locality: Yuyama, Matsuyama

Distribution: Egypt, Japan

Eupodes tottanfjella Strandtmann

E. tottanfjella Strandtmann, 1967: 54

Type locality: Queen Maud Land

Distribution: Antarctica

Eupodes trifasciatus C.L. Koch

Eupodes trifasciatus C.L. Koch, 1838: fa.19, t.15

Type locality: Regensburg

Distribution: Germany

Eupodes unifasciatus C.L. Koch

Eupodes unifasciatus C.L. Koch, 1838: fa.19, t. 11; 1841: 436; 1842: fa.3, p.69; Walckenaer, 1847: 537; Oudemans, 1937: 897; Thor & Willmann, 1941: 15; Turk & Turk, 1952: 497

Type locality: Regensburg (Germany)

Distribution: England, Germany,

Eupodes vallombrosae Thor

Eupodes vallombrosae Thor, 1934: 212; Mihelčič, 1958b: 269; Schweizer & Bader, 1963: 217

Type locality: “Vallombrosa, Firenze”, Italy

Distribution: Italy, Schwitzerland, Spain

Eupodes variegatus C.L. Koch

Eupodes variegatus C.L. Koch, 1838: fa.19, t.2; 1841: v.3, 435; 1842: fa.3, p. 69; Walckenaer, 1847: 537; Thor, 1912: 236; Halbert, 1915: 110; 1920: 139 (var. halophilus); Hull, 1918: 34; Sellnick, 1940: 32; Haarløv, 1942; Thor & Willmann, 1941: 20; Macfadyen, 1954: 283; Bottazzi, 1950: 37; Mihelčič, 1958a: 104; 1958b: 269; Sellnick, 1958: 38; Meyer & Ryke, 1960: 487; Lipkin & Safriel, 1971: 7 (var. halophila)

Type locality: Regensburg (Germany)

Distribution: Austria, England, Germany, Greenland, Ireland, Island, Israel, Italy, Norway, South Africa, Spain, Sweden, Yan Mayen I.

Eupodes versicolor C.L. Koch

Eupodes versicolor C.L. Koch, 1838: fa.19, t.13

Type locality: Regensburg

Distribution: Germany

Eupodes viridis Oudemans

Eupodes viridis Oudemans, 1906: 40

Type locality: Greenland

Distribution: Greenland

Eupodes voxencollinus Thor

Eupodes voxencollinus Thor, 1934: 208; Weis-Fogh, 1948; Evans, 1952: 666; Hallas, 1978: 31; Strandtmann & Goff, 1978: 126; Abou-Awad, 1984: 333; Goff, 1987: 31; Darbemamieh, Hajiqanbar & Khanjani, 2013: 454; Hagstrum D. et al. 2013: 546

Eupodes alaskanensis Strandtmann, 1971: 79; Strandtmann & Prasse, 1977: 4

Eupodes acuminatus Willmann, 1952 – Wangerooge Is.

Type locality: “Voksenkollen bei Oslo” (Norway)

Distribution: Alaska, Canada, China, Denmark, Egypt, England, Germany Hawaii, Iran, Norway

Eupodes winsnesi Strandtmann et Sømme

Eupodes winsnesi Strandtmann & Sømme, 1977: 140

Type locality: Queen Maud Land, Brattskarvet, Sverdrupfjella

Distribution: Antarctica

Eupodes wisei Womersley et Strandtmann

Eupodes wisei Womersley & Strandtmann, 1963: 452

Type locality: Hallett Station

Distribution: Antarctica

Eupodes zaheri Abou-Awad, El-Sawaf et Abdel-Khalek

Eupodes zaheri Abou-Awad, El-Sawaf & Abdel-Khalek, 2006: 44

Type locality: Egypt

Distribution: Egypt

Eupodes sp. – Yemen (Ueckermann et al., 2006: 262)

Genus Filieupodes Jesionowska

Filieupodes Jesionowska, 2010: 641

Type species: Filieupodes filiformis Jesionowska, 2010

Filieupodes aegyptiacus (Abou-Awad et El-Bagouri)

Cocceupodes aegyptiacus Abou-Awad & El-Bagouri, 1984: 68

Filieupodes aegyptiacus (Abou-Awad et El-Bagouri): Jesionowska, 2010: 642

Type locality: Egypt (on Vitis vinifera)

Distribution: Egypt

Filieupodes filiformis Jesionowska

Filieupodes filiformis Jesionowska, 2010: 645

Type locality: The Bialowieza Forest. Hajnówka (Podlaskie District)

Distribution: Poland

Filieupodes filistellatus Jesionowska

Filieupodes filistellatus Jesionowska, 2010: 651

Type locality: Ciechocinek near Toruń (Kujawsko-Pomorskie Distr.)

Distribution: Poland

Filieupodes fusiformis (Olivier et Theron)

Cocceupodes fusiformis Olivier & Theron, 2003: 165

Filieupodes fusiformis (Olivier et Theron): Jesionowska, 2010: 652

Type locality: S.Afr., Northern Province, near Nylstroom

Distribution: South Africa

Filieupodes paradoxus (Weis-Fogh)

Cocceupodes paradoxus Weis-Fogh, 1948: 263; Evans, 1952: 666; Haarløv, 1957: 27

Filieupodes paradoxus (Weis-Fogh): Jesionowska, 2010: 652

Type locality: Denmark

Distribution: England, Denmark, Greenland

Filieupodes sharkiensis (Abou-Awad, El-Sawaf et Abdel-Khalek)

Cocceupodes sharkiensis Abou-Awad, El-Sawaf & Abdel-Khalek, 2006: 48

Filieupodes sharkiensis : Jesionowska, 2010: 642

Type locality: Sharki

Distribution: Egypt

Filieupodes shepardi (Strandtmann)

Cocceupodes shepardi Strandtmann, 1971: 88

Filieupodes shepardi (Strandtmann): Jesionowska, 2010: 642

Type locality: Cape Beaufort (Alaska)

Distribution: USA (Alaska)

Filieupodes strandtmanni (Abou-Awad et El-Bagouri)

Cocceupodes strandtmanni Abou-Awad & El-Bagouri, 1986: 68

Filieupodes strandtmanni (Abou-Awad et El-Bagouri): Jesionowska, 2010: 642

Type locality: Egypt

Distribution: Egypt

Filieupodes trisetatus (Strandtmann et Prasse)

Cocceupodes trisetatus Strandtmann & Prasse, 1977: 7; Strandtmann & Goff, 1978: 130; Goff, 1987: 31

Filieupodes trisetatus (Strandtmann et Prasse): Jesionowska, 2010: 641

Type locality: Etzsdorf/ Saalkreis (East Germany)

Distribution: Germany, Hawaii

Genus Linopodes C.L. Koch

Linopodes C.L. Koch, 1836: fa. 1, t. 16

Type species: Linopodes ravus C.L. Koch, 1836

Linopodes antennaepes Banks

Linopodes antennaepes Banks, 1894: 221; Miller, 1925: 93; Darbemamieh, Hajiqanbar & Khanjani, 2013: 454

Type locality: USA

Distribution: Australia, Canada, Great Britain, Iran, Italy, USA

Linopodes barnufi Abou - Awad Badawi, El-Sawaf et Abdel-Khalek

Linopodes barnufi Abou - Awad Badawi, El-Sawaf & Abdel-Khalek, 2006: 50

Type locality: Egypt

Distribution: Egypt

Linopodes cameronensis Shiba

Linopodes cameronensis Shiba, 1976: 96; Darbemamieh, Hajiqanbar & Khanjani, 2013: 454

Type locality: Malaysia

Distribution: Iran, Malaysia

Linopodes eupodoides R. Canestrini

Linopodes eupodoides R. Canestrini, 1886: 701; Schweizer & Bader, 1963: 219

Type locality: Italy, Trentino

Distribution: Italy, Switzerland

Linopodes iwatensis Morikawa

Linopodes pubescens iwatensis Morikawa, 1963: 15

Linopodes iwatensis : Shiba, 1969c: 69

Type locality: Uchimagi, Yamagata-mura, Iwate Pref.

Distribution: Japan

Linopodes kochi Sig Thor (in Thor & Willmann, 1941: 43, nomen nov. for Linopodes longipes C.L. Koch, 1838: 15 non Trombidium longipes Hermann, 1804)

Type locality: Germany

Distribution: Germany

Linopodes motatorius (Linnaeus)

Acarus motatorius Linnaeus, 1758: 616

Gamasus motatorius : Fabricius, 1805: 360

Linopodes motatorius (Linnaeus): Canestrini, 1886: 699; Oudemans, 1897: 112; 1937: 916; Willmann, 1951a: 146; 1954: 234; Bottazzi, 1956: 38; Haarløv, 1957: 30; Cooreman, 1959: 17; Schmölzer, 1962: 283; Schweizer & Bader, 1963: 219 (full synonymy until 1938 in Thor & Willmann, 1941).

Type locality: Europe

Distribution: Austria, Bulgaria, Belgium, Croatia, Czech Rep., Denmark, Germany, Great Britain, Hungary, Iceland, Italy, Norway, Spain, Switzerland

Linopodes motatorius africanus Meyer et Ryke

Linopodes motatorius africanus Meyer & Ryke, 1960: 489

Type locality: Bathurst

Distribution: South Africa

Linopodes obsoletus C.L. Koch

Linopodes obsoletus C.L. Koch, 1838: fasc. 18, t.18

Type locality: Regensburg

Distribution: Germany

Linopodes pubescens Morikawa

Linopodes pubescens Morikawa, 1963: 13

Type locality: Shiroyama, Matsuyama City, Ehime Pref.

Distribution: Japan

Linopodes ravus C.L. Koch

Linopodes ravus C.L. Koch, 1836: fa. 1, t. 17

Type locality: Regensburg

Distribution: Germany

Genus Neoprotereunetes Fain et Camerik

Neoprotereunetes Fain & Camerik, 1994: 153

Type species: Ereynetes lapidarius Oudemans, 1906

Neoprotereunetes lapidarius (Oudemans)

Ereynetes lapidarius Oudemans, 1906: 40

Protereunetes lapidarius : Bujtendijk, 1945: 319; Thor & Willmann, 1941: 58

Neoprotereunetes lapidarius (Oudemans): Fain & Camerik, 1994: 153

Type locality: Arnhem

Distribution: Holland

Genus Niveupodes Barilo

Niveupodes Barilo, 1991: 140

Type species: Niveupodes niveus Barilo, 1991

Niveupodes niveus Barilo

Niveupodes niveus Barilo, 1991: 140; Khaustov, 2016: 258

Type locality: Zarafshan Range, 20 km south of Samarkand

Distribution: Uzbekistan

Genus Pseudeupodes Khaustov

Pseudeupodes Khaustov, 2014: 13

Type species: Pseudeupodes porosus Khaustov, 2014

Pseudeupodes porosus Khaustov

Pseudeupodes porosus Khaustov, 2014: 13

Type locality: Yalta mountain-forest Nature Reserve

Distribution: Crimea

Genus Pseudopenthaleus Khaustov

Pseudopenthaleus Khaustov, 2015: 931

Type species: Pseudopenthaleus tauricus Khaustov, 2015

Pseudopenthaleus ciliatus (Shiba)

Penthaleus ciliatus Shiba, 1978: 147

Pseudopenthaleus ciliatus (Shiba): Khaustov, 2015: 927

Type locality: Mt. Sagaramine, Shigenobu-sho, Ehime

Distribution: Japan

Pseudopenthaleus tauricus Khaustov

Pseudopenthaleus tauricus Khaustov, 2015: 931

Type locality: Crimea

Distribution: Crimea

Genus Xerophiles Jesionowska

Xerophiles Jesionowska, 2003: 12

Type species: Xerophiles ereynetoidalis Jesionowska, 2003

Xerophiles ereynetoidalis Jesionowska

Xerophiles ereynetoidalis Jesionowska, 2003: 12

Type locality: Szczecin (Poland)

Distribution: Poland, Slovakia

Among the species described by Müller, 1776, Schrank, 1776, Hermann, 1804, Dugès, 1834, C.L. Koch, 1838, Wankel, 1861, Karsch, 1881, Banks, 1894, 1897, Berlese, 1917, but with unclear identity, are Eupodes cerinus Koch, E. celerrimus Koch, E. variabilis Banks, E. formosulus Koch, E. dugesi Sig Thor (as nom. nov.), E. dilectus Koch, E. cornifer (Müller) (Acarus c.), E. cinctus Koch, E. chloromelas Koch, E. celer (Dugès) (Megamerus c.), E. lineatus Koch, E. lineola Koch, E. fasciola Koch, Е. ochlochlorus Koch, E. saltatorius Schrank (Acarus s.), E. macropus Koch, E. melanurus Koch, E. pallescens Koch, Linopodes ambustus Koch, L. decoloratus Koch, L. decoloreus Koch L. longipes , L. macropus Herm. (Trombidium m.), L. gracilipes Karsch, L. flexuosus Koch, L. flavipes Koch, L. riparius Koch, L. subterraneus Wankel, L. mammuthia Banks (Eupodes m.), L. maculatus Koch, L. melaleucus Koch, L. rubiginosus Koch, L. agitatorius Berlese.

Distribution of Eupodidae and Cocceupodidae

EUROPE

Austria (Irk, 1939; Willmann, 1951; Mihelčič, 1957, 1958; Schmölzer, 1962, 1993) – Cocceupodes mollicellus, Eupodes pseudoclavifrons, E. variegatus, Linopodes motatorius

Belgium (Cooreman, 1943) – Claveupodes sp., Linopodes motatorius

Bulgaria (Beron, 2011) – Linopodes motatorius

Croatia (Matočec, 2002) - Linopodes motatorius

Czech Rep. (Willmann, 1954) – Eupodes sp., Linopodes motatorius, [? L. subterraneus Wankel]

Denmark (Weis-Fogh, 1948; Haarløv, 1957; Hallas, 1978) – Cocceupodes mollicellus, Eupodes berlesei, E. viridis, E. voxencollinus, Filieupodes paradoxus

Greenland (Haarløv, 1942; Makarova, 2015) – Cocceupodes breweri, C. mollicellus (= Eupodes clavifrons R. Canestrini, 1886 = Cocceupodes curticlava Thor, 1934), C. paradoxus, C. cf. stellatus

Estonia (Thor & Willmann, 1941) – Protereunethes striatellus

France (contin.)(Coineau, 1976) – Eupodes strandtmanni

Germany (C.L. Koch, 1835, 1838; Willmann, 1952; Strandtmann & Prasse, 1977) – Benoinyssus ereynetoides, Claveupodes delicatus, Cocceupodes stellatus, Eupodes leucomelas, E. ocellatus, E. striola, E. signatus, E. trifasciatus, E. voxencollinus, Filieupodes trisetatus, Linopodes motatorius, L. ravus, Protereunethes striatellus

Great Britain (Hull, 1918; Evans, 1952; Turk & Turk, 1952; A. Baker, 1987) – Caleupodes reticulatus,Claveupodes sp., Eupodes clavifrons, E. unifasciatus, E. variegatus, E. voxencollinus, Filieupodes paradoxus, Linopodes antennaepes, L. motatorius

Holland (Oudemans, 1906; Thor & Willmann, 1941) - Eupodes milvinus, E. viridis, Protereunethes striatellus

Hungary (Dudich et al., 1940) – Linopodes motatorius

Iceland (Sellnick, 1940) – Eupodes variegatus, Linopodes motatorius

Surtsey (Lindroth, Andersson, Bödvarsson & Richter, 1973) – Cocceupodes clavifrons, Eupodes hiemalis, E. variegatus, Linopodes motatorius

Ireland (Halbert, 1915, 1920) – Eupodes variegatus var. halophilus

Italy (contin.)(Canestrini, 1886; Thor, 1934, Thor & Willmann, 1941; Bottazzi, 1950) - Cocceupodes mollicellus (= clavifrons), Eupodes fusifer, E. iconicus, E. milvinus, E. vallombrosae, E. variegatus, Linopodes antennaepes, L. eupodoides, L. motatorius

Sicily (Berlese, 1923) – Protereunetes agilis

Norway (Thor, 1912, 1934a, 1934c; Thor & Willmann, 1941) – Cocceupodes mollicellus (=curviclava), Eupodes pseudoclavifrons, E. decoloratus, E. hjartdaliae, E. skiaaki, E. variegatus, E. voxencollinus, Linopodes motatorius

Svalbard (Spitzbergen)(Oudemans, 1926; Thor, 1934b, 1934c) – Eupodes clavifrons, Protereunetes boerneri

Poland (Jesionowska, 2005, 2006, 2007a, 2007b, 2008, 2010; Szudarek-Trepto, Kaźmierski, Dabert et al., 2020) – Claveupodes sp., Cocceupodes longisolenidiatus, Filieupodes filiformis, F. filistellatus, Eupodes voxencollinus, Xerophiles ereynetoidalis

Romania (Ivan et al., 2016) – Eupodes sp.

Russia (cont.)

Crimea (Khaustov, 2014, 2015, 2017) – Echinoeupodes echinus, Pseudeupodes porosus, Pseudopenthaleus tauricus

Slovakia (Jesionowska, 2003) - Xerophiles ereynetoidalis

Spain (cont.)(Mihelčič, 1958b; Santamaría et al., 2012) - Cocceupodes sp., Benoinyssus ereynetoides, E. valombrosae, E. variegatus, Linopodes motatorius

Sweden (Thor & Willmann, 1941; Sellnick, 1958) - Eupodes milvinus, E. variegatus

Switzerland (Haller, 1880; Schweizer, 1922; Cooreman, 1954,1959, Schweizer & Bader, 1963) – Eupodes berlesei, E. haltica, E. vallombrosae, Linopodes eupodoides, L. motatorius

AFRICA

Angola (Meyer & Ryke, 1960; Olivier & Theron, 1997a, 1998) – Benoinyssus celatus, B. ornatus, Eupodes parafusifer

Botswana (Olivier & Theron, 1997a; 1998) - Benoinyssus serratus, Eupodes minipulus

Cabo Verde (Olivier & Theron, 1998) - Eupodes minipulus

Egypt (Trägårdh, 1904; Abou-Awad, 1984: 333; Abou-Awad & El-Bagouri, 1984, 1986; Abou-Awad, El-Sawaf & Abdel-Khalek, 2006) – Benoinyssus momeni, Cocceupodes sharkiensis, Filieupodes aegyptiacus, F. strandtmanni, Eupodes bakeri, E. momeni, E. niloticus, E. strandtmanni, E. temperatus, E. zaheri, Linopodes barnufi, L.curtipes

Ivory Coast - Eupodes sigmoidensis

Namibia (Olivier & Theron, 1997a) - Benoinyssus serratus

Rwanda (Fain, 1958, 1959) – Benoinyssus najae

South Africa (Thor, 1934c; Meyer & Ryke, 1960; Olivier & Theron, 1997a, 1997b, 1998, 2003) – Benoinyssus falcatus, B. ornatus, B. pusillus, B. serratus, C. gracilongus, Eupodes acanthus, E. fusiferellus, E. hamatus, E. indentatus, E. lutatus, E. minipilus, E. parafusifer, E. variegatus, Filieupodes fusiformis

Swaziland (Olivier & Theron, 1997a) – Benoinyssus ornatus, B. serratus

Zimbabwe (Olivier & Theron, 1998) - Eupodes minipulus

East Africa” (Berlese, 1916) – “Linopodes agitatorius

ASIA

China (Hagstrum et al., 2013) – Eupodes voxencollinus

India (Ghosh & Gupta, 2003; Gupta et al., 2004, Indranil et al., 2011) – Eupodes sigmoidensis

Indonesia (Corpuz-Raros, 2005) – Eupodes pusillus [Java]

Iran (Darbemamieh, Hajiqanbar & Khanjani, 2013) - Benoinyssus sp., Claveupodes sp., Cocceupodes sp., Eupodes crozetensis, E. sigmoidensis, E. voxencollinus, Linopodes antennaepes, L. cameronensis

Israel and Palestina (Lipkin & Safriel, 1971) – Eupodes variegatus var. halophila

Japan (Karsch, 1881; Morikawa, 1963; Shiba, 1969a, 1969c, 1971, 1978) – Cocceupodes communis, Echinoeupodes turgidus, Eupodes cordiformis, E. infirmus, E. longisetatus, E. okinoshimaensis, Linopodes iwatensis, L. pubescens, Protereunetes perforates, P. villosus, Pseudopenthaleus ciliatus

Malaysia (Shiba, 1976) – Eupodes berensis, E. lumbarus, E. onoi, E. pasohensis, E. pusillus, E. pseudolumbarus, Linopodes cameronensis

Philippines (Corpuz-Raros, 2005) – Eupodes callejae, E. garciai, E. onoi, E. pusillus, E. pseudolumbarus

Singapore (Corpuz-Raros, 2005) – Eupodes onoi

United Arab Emirates (Negm, 2014) – Eupodes sp.

Uzbekistan (Barilo, 1991; Khaustov, 2016) – Niveupodes nivalis

Yemen (Ueckermann et al., 2006) – Eupodes sp.

NORTH AMERICA

Canada (Miller, 1925; Behan, 1978; Moseley, 2007; Beaulieu et al., 2019 – 13 sp.) - Eupodes voxencollinus, Linopodes antennaepes

Mexico (Baker, 1945) - Benoinyssus tuberculatus

USA (Banks, 1894; 1896; 1897, 1907; 1916; Miller, 1925; Thor, 1934) – Eupodes brevipes, E. marinus, Linopodes antennaepes

Alaska (Strandtmann, 1971) – Cocceupodes breweri, Eupodes alaskanensis, E. voxencollinus, Filieupodes shepardi, Protereunetes boerneri

Hawaii (Strandtmann & Goff, 1978) - Eupodes hawaiiensis, E. sigmoidensis, E. voxencollinus

SOUTH and CENTRAL AMERICA

Argentina (Bedano, 2004) – 1 sp.

Brazil (Bizarro et al., 2020) – Benoinyssus oconnori

Costa Rica (Baker & Lindquist, 2002) – Aethosolenia laselvensis

OCEANIA

Australia (continent)(Clancy, 1981) - Linopodes sp.

New Zealand (Spain & Luxton, 1971; Martin, 1983; Luxton, 1990) – Claveupodes delicatulus, Linopodes spp., Eupodes minutus,

Campbell Island (Strandtmann, 1964) - Eupodes longisetatus

Macquarie Island (Womersley, 1937; Luxton, 1990) – Eupodes minutus

ANTARCTICA

Antarctic Continent (Strandtmann & Womersley, 1963; Strandtmann, 1967; Strandtmann & Davies, 1972; Gless, 1972; Strandtmann & Sømme, 1977; Marshall & Pugh, 1996; Llano & Gless, 2013) – Eupodes angardi, E. winsnesi, E. wisei

South Georgia (Strandtmann, 1970) – Eupodes minutus

South-Orkney-Islands (Booth, Edwards & Usher, 1985) – Eupodes exiguus, E. parvus

South Shetland Islands (Strandtmann & Tilbrook, 1968) – Cocceupodes australis

SUB-ANTARCTIC

Crozet Islands (Strandtmann & Davies, 1972) – Eupodes crozetensis, E. riedli, Possession I.

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  • Strandtmann R. W., J. Prasse. 1977. Prostigmatic mites from the experimental farm in Etzsdorf/ Saalkreis, G. D. R. – Abhandl. Ber. Naturk. Görlitz, 50(2): 1 – 36.
  • Strandtmann R. W., L. Sømme. 1977. Prostigmate mites (Acari, Prostigmata) from Sverdrupfjella, Dronning Maud Land, with description of four new species. – Norw. J. Ent., 24: 137 – 146.
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The Mite Family Dendrochaetidae Olivier

Dendrochaetidae Olivier, 2008: 16

The family contains so far only one genus and species (Dendrochaetus acarus Olivier) from South Africa. These mites live in the soil.

Catalogue of Dendrochaetidae

Fam. Dendrochaetidae Olivier

Dendrochaetidae Olivier, 2008: 16

Type genus: Dendrochaetus Olivier, 2009 [repl. name for Dendrodus Olivier, 2008, praeocc.]

Genus Dendrochaetus Olivier

Dendrodus Olivier, 2008: 19 (praeocc.)

Dendrochaetus Olivier, 2009: 68

Type species: Dendrochaetus acarus Olivier, 2009 [for Dendrodus acarus Olivier, 2008]

Dendrochaetus acarus (Olivier)

Dendrodus acarus Olivier, 2008: 19

Dendrochaetus acarus (Olivier): Olivier, 2009: 68

Type locality: South Africa

Distribution: South Africa

Distribution of Dendrochaetidae

AFRICA

South Africa (Olivier, 2008, 2009) – Dendrochaetus [Dendrodus] acarus

Bibliography of Dendrochaetidae

  • Olivier P.A.S. 2008. Dendrochaetidae, a new family of mites (Acari, Prostigmata), with description of a new genus and species from South Africa. – African Zoology, 43(1): 16 – 24.
  • Olivier P.A.S. 2009. A replacement name for Dendrodus Olivier, 2008, and validation of Dendrochaetidae Olivier, 2008 nomen nudum (Acari, Prostigmata). – Zootaxa, 2293: 68.

The Mite Family Rhagidiidae Oudemans

Rhagidiidae Oudemans, 1922: 83

The Rhagidiids are soft bodied, predaceous soil mites. Thorell (1872) described the genus Rhagidia and Cambridge (1876) described from the far away Kerguelen Island another genus – Poecilophysis. Described by Oudemans (1922), the family Rhagidiidae has been studied until 1980 by several researchers (C.L. Koch, Canestrini, Thorell, Berlese, Banks, Hull, Sig Thor, Trägårdh, Willmann, Elliot, Strandtmann, Ehrnstberger, Abou-Awad and El-Bagoury, Womersley, Morikawa, Shiba, Baltac, Halliday, Khaustov, and others). The first review of the family (Thor & Willmann, 1941) contained three genera, 37 certain and 8 uncertain species and five “varieties”. R.W. Strandtmann tried to distinguish some more reliable diagnostic characters, but only the world revision of the family (Zacharda, 1980) achieved the modern system of the family. From the 20 genera present in the monograph Zacharda described 16 as new and from 74 species 35 were described as new.

In the 40 years elapsed since this monograph have been described the new genera Krantzia Zaharda, 1983, Neothoria Abou-Awad et El-Bagoury, 1986 Lindquistula Zacharda, 1986, Kouchibouguacia Zacharda, 1986, Arhagidia Lindquist et Zacharda, 1987, Dongnaia Zacharda, 1994, Janes Jesionowska, 2002, Paracrassocheles Khaustov, 2015 and several new subgenera and species. Now (September 2021) the family Rhagidiidae comprises 28 genera and 150 species of whitish mites, living in the soil and often in caves.

Composition of fam. Rhagidiidae

Family Rhagidiidae Oudemans, 1922 - 28 gen., 150 sp.

Arctorhagidia Zacharda, 1980 – 2 sp.

Arhagidia Lindquist et Zacharda, 1987 – 1 sp.

Brevipalpia Zacharda, 1980 - 1 sp.

Coccorhagidia Sig Thor, 1934 – 8 sp.

Crassocheles Zacharda, 1980 - 2 sp.

Elliotta Zacharda, 1980 - 1 sp.

Eskimaia Zacharda, 1980 – 1 sp.

Evadorhagidia Zacharda, 1980 - 4 sp.

Flabellorhagidia Elliott, 1976 - 1 sp.

Foveacheles Zacharda, 1980 - 33 sp.

Foveacheles Zacharda, 1980 - 11 sp.

Hirschmannetta Zacharda, 1980 - 1 sp.

Mediostella Zacharda, 1980 - 5 sp.

Propriorhagidia Zacharda, 1980 - 2 sp.

Proxistella Zacharda, 1980 - 2 sp.

Spelaeocheles Zacharda, 1988 – 1 sp.

Ternirhagidia Zacharda, 1980 - 1 sp.

Trofocheles Zacharda, 1982 - 3 sp.

Usitorhagidia Zacharda, 1980 – 7 sp.

Hammenia Zacharda, 1980 - 1 sp.

Janes Jesionowska, 2002 – 1 sp.

Kouchibouguacia Zacharda, 1986 – 2 sp.

Latoempodia Zacharda, 1980 - 2 sp.

Lindquistula Zacharda, 1986 – 1 sp.

Neothoria Abou-Awad et El-Bagoury, 1986 – 2 sp.

Paracrassocheles Khaustov, 2015 – 1 sp.

Parallelorhagidia Zacharda, 1980 - 3 sp.

Poecilophysis Cambridge, 1876 - 16 sp.

Dentocheles Zacharda, 1980 - 5 sp.

Poecilophysis Cambridge, 1876 - 3 sp.

Procerocheles Zacharda, 1980 - 5 sp.

Soprocheles Zacharda, 1980 - 2 sp.

Wankelia Zacharda, 1980 - 1 sp.

Rhagidia Thorell, 1872 - 28 sp.

Austrorhagidia Zacharda, 1980 – 3 sp.

Deharvengiella Zacharda, 1987 – 3 sp.

Zachardaia Özdikmen, 2008 (= Krantzia Zacharda, 1983) - 1 sp.

Dongnaia Zacharda, 1994 – 1 sp.

Noerneria Canestrini, 1886 – 14 sp.

Rhagidia Thorell, 1872 - 7 sp.

Robustocheles Zacharda, 1980 - 19 sp.

Amoveocheles Zacharda, 1980 - 3 sp.

Lewia Zacharda, 1980 - 5 sp.

Mexia Zacharda, 1997 - 3 sp.

Robustocheles Zacharda, 1980 - 8 sp.

Shibaia Zacharda, 1980 - 6 sp.

Thoria Zacharda, 1980 - 2 sp.

Traegaardhia Zacharda, 1980 - 10 sp.

Troglocheles Zacharda, 1980 - 10 sp.

Tuberostoma Zacharda, 1980 – 3 sp.

Catalogue of Rhagidiidae

Family Rhagidiidae Oudemans

Rhagidiidae Oudemans, 1922: 83

Type genus: Rhagidia Thorell, 1872

Genus Arctorhagidia Zacharda

Arctorhagidia Zacharda, 1980: 719

Type species: Rhagidia mariehammerae Strandtmann, 1971

Arctorhagidia mariehammerae (Strandtmann)

Rhagidia mariehammerae Strandtmann, 1971: 113

Arctorhagidia mariehammerae (Strandtmann): Zacharda, 1980: 719

Type locality: Alaska, Cape Beaufort

Distribution: USA (Alaska)

Arctorhagidia sateri (Strandtmann)

Coccorhagidia sateri Strandtmann, 1971: 95

Arctorhagidia sateri (Strandtmann): Zacharda, 1980: 721

Type locality: Alaska, Barrow

Distribution: USA (Alaska)

Genus Arhagidia Lindquist et Zacharda

Arhagidia Lindquist et Zacharda, 1987: 2149

Type species: Arhagidia monothrix Zacharda, 1987

Arhagidia monothrix Lindquist et Zacharda

Arhagidia monothrix Lindquist & Zacharda, 1987: 2156

Type locality: Jornada ecological research site of the Chihuahuan Desert in southern New Mexico

Distribution: USA (New Mexico)

Genus Brevipalpia Zacharda

Brevipalpia Zacharda, 1980: 712

Type species: Brevipalpia minima Zacharda, 1980

Brevipalpia minima Zacharda

Brevipalpia minima Zacharda, 1980: 712; 2012; Zacharda & Daniel, 1988: 58; Darbemamieh, Hajiqanbar & Khanjani, 2013: 455

Type locality: Bohemia centr., Závist-Knoll near Zbraslav

Distribution: Czech Republic, Iran, Nepal

Genus Coccorhagidia Sig Thor

Coccorhagidia Sig Thor, 1934: 318

Type species: Nörneria clavifrous Canestrini, 1886

Coccorhagidia clavifrons (Canestrini)

Nörneria clavifrons Canestrini, 1886: 710

Rhagidia clavifrons : Berlese, 1921: 90; Bedano, 2004

?Rhagidia mordax: Willmann, 1936: 296

Coccorhagidia clavifrons : Sig Thor, 1934: 318; Dudich et al., 1940: 54; Thor & Willmann, 1941: 128; Willmann, 1951b: 143; Turk & Turk, 1952: 496; Macfadyen, 1954: 284; Haarløv, 1957: 27; Meyer & Ryke, 1960: 493; Shiba, 1969: 81; Strandtmann & Prasse, 1976: 13; Strandtmann & Goff, 1978: 143; Zacharda, 1980: 704; 2013; Abou-Awad, 1985: 258; Goff, 1987: 34; Qin, 1998: 300; Darbemamieh, Hajiqanbar & Khanjani, 2013: 455

Type locality: Doss (Tavon, Trento), Italy

Distribution: Argentina, Austria, Czech Republic, Denmark, Egypt, England, Germany, Hungary, Iran, Italy, Japan, Slovakia; Canada, Hawaii, South Africa, New Zealand, Yan Mayen I.

Coccorhagidia cubana Baltac

Coccorhagidia cubana Baltac, 1977a: 197

Type locality: Cueva de México, Province de Camagüey

Distribution: Cuba

Coccorhagidia montana Bannova, Sarmanova et Zhuravleva

Coccorhagidia montana Bannova, Sarmanova & Zhuravleva, 1977: 59

Type locality: Romania

Distribution: Romania

Coccorhagidia divergens Mihelčič (sp. inquirend.)

Coccorhagidia divergens Mihelčič, 1957: 103

Type locality: Ötztaler Alpen: Hochfirst, Nivale Polsterpflanzen aus der O-Wand, ca. 3400 m.

Distribution: Austria

Coccorhagidia pittardi Strandtmann

Coccorhagidia pittardi Strandtmann, 1971: 98; Zacharda, 1980: 708; 2012: ; Zacharda & Daniel, 1988: 58; Darbemamieh, Hajiqanbar & Khanjani, 2013: 455

Type locality: Alaska, Wainwright

Distribution: Slovakia, Tatra (2200-2367 m); Alaska, Iran, Nepal (3900 m),

Coccorhagidia semiclavifrons Shiba (sp. inquir.)

Coccorhagidia semiclavifrons Shiba, 1969: 83

Type locality: Shiga Heigths, IBP Area

Distribution: Japan

Coccorhagidia subterranea (Berlese)

Rhagidia subterranea Berlese, 1905: 232

Coccorhagidia berlesei Thor & Willmann, 1941: 129 (as nom. nov. pro Scyphius subterraneus nom. nud. Absolon, 1899)

Coccorhagidia subterranea (Berlese): Evans, 1952: 666; Haarløv, 1957: 27

Type locality: Florence (Italy)

Distribution: Denmark, England, Italy

Genus Crassocheles Zacharda

Crassocheles Zacharda, 1980: 550

Type species: Crassoheles muralis Zacharda, 1980

Crassocheles muralis Zacharda

Crassocheles muralis Zacharda, 1980: 551

Type locality: Velký Berdĕz-Hill, Czech Republic

Distribution: Czech Republic

Crassocheles virgo Zacharda

Crassocheles virgo Zacharda, 1980: 554

Type locality: Provodin, Provodingké kameny, Czech Republic

Distribution: Czech Republic

Genus Elliotta Zacharda

Elliotta Zacharda, 1980: 597

Type species: Flabellorhagidia howarthi Elliott, 1976

Elliotta howarthi (Elliott)

Flabellorhagidia howarthi Elliott, 1976: 5-10

Elliotta howarthi : Zacharda, 1980: 597

Type locality: Klickitat Co., Washington, Cheese Cave, 2 km SW Trout Lake, USA

Distribution: USA (Washington, Hawaii, Idaho)

Genus Eskimaia Zacharda

Eskimaia Zacharda, 1980: 723

Type species: Coccorhagidia capitata Strandtmann, 1971

Eskimaia capitata (Strandtmann)

Coccorhagidia capitata Strandtmann, 1971: 98

Eskimaia capitata (Strandtmann): Zacharda, 1980: 723

Type locality: Alaska, Anaktuvik Pass, Brooks Range

Distribution: USA (Alaska)

Genus Evadorhagidia Zacharda

Evadorhagidia Zacharda, 1980: 725

Type species: Evadorhagidia oblikensis Zacharda, 1980

Evadorhagidia bezdezensis Zacharda

Evadorhagidia bezdezensis Zacharda, 1980: 734; Ružička, Hajer & Zacharda, 1995: 47

Type locality: Bohemia bor., Malý Bezdĕz-Hill

Distribution: Czech Republic

Evadorhagidia corcontica Zacharda

Evadorhagidia oblikensis corcontica Zacharda, 1993: 57

Evadorhagidia corcontica Zacharda:

Type locality: Krkonose Mts.

Distribution: Czech Republic

Evadorhagidia janetscheki (Willmann)

Rhagidia janetscheki Willmann, 1953: 218; Strandtmann, 1971: 103

Evadorhagidia janetscheki : Zacharda, 1980: 733

Type locality: Tyrol, the Zillertall Alps, Rossrücken, 3000 m

Distribution: Austria

Evadorhagidia oblikensis Zacharda

Evadorhagidia oblikensis Zacharda, 1980: 726

Type locality: Bohemia bor., Oblik-Hill near Louny

Distribution: Czech Republic

Evadorhagidia quinqueseta Zacharda

Evadorhagidia quinqueseta Zacharda, 1980: 731

Type locality: Canada, N.W. Territory, Atkinson Point

Distribution: Canada (N.W. Territory)

Evadorhagidia sp. – Sweden (Hippa et al., 1989: 61), Finland (Zacharda & Kučera, 2010: 5)

Genus Flabellorhagidia Elliott

Flabellorhagidia Elliott, 1976: 1

Type species: Flabellorhagidia pecki Elliott, 1976

Flabellorhagidia pecki Elliott

Flabellorhagidia pecki Elliott, 1976: 2; Zacharda, 1980: 657

Type locality: Butte Co., Idaho, Boy Scout Cave (lava)

Distribution:USA (Idaho)

Genus Foveacheles Zacharda

Foveacheles Zacharda, 1980: 661

Type species: Rhagidia osloensis Sig Thor, 1934

Subgenus Achaetorhagidia Zacharda

Achaetorhagidia Zacharda, 1994: 36

Type species: Foveacheles (Achaetorhagidia) tropica Zacharda, 1994

Foveacheles (Achaetorhagidia) namcatiensis Zacharda

Foveacheles (Achaetorhagidia) namcatiensis Zacharda, 1994: 38

Type locality: Nam Cat Tien National Park, South Vietnam

Distribution: Vietnam

Foveacheles (Achaetorhagidia) tropica Zacharda

Foveacheles (Achaetorhagidia) tropica Zacharda, 1994: 44

Type locality: Nam Cat Tien National Park, South Vietnam

Distribution: Vietnam

Subgenus Foveacheles Zacharda

Foveacheles Zacharda, 1980: 661

Type species: Rhagidia osloensis Sig Thor, 1934

Foveacheles (Foveacheles) auricularia Zacharda et Elliott

Foveacheles (Foveacheles) auricularia Zacharda & Elliott, 1985: 468

Type locality: California

Distribution: USA

Foveacheles (Foveacheles) brevichelae Zacharda

Foveacheles (Foveacheles) brevichelae Zacharda, 1980: 666; 2012: ; Darbemamieh, Hajiqanbar & Khanjani, 2013: 455; Zacharda & Kučera, 2010: 5

Type locality: Bohemia centr., Karlštejn

Distribution: Austria, Canada, Czech Republic, Finland, Iran, Russia

Foveacheles (Foveacheles) cannadasi Barranco et Amate

Foveacheles (Foveacheles) cannadasi Barranco & Amate, 2000: 15

Type locality: Cueva Larga, Sierra de los Filabres, Almeria

Distribution: Spain

Foveacheles (Foveacheles) caucasica Zacharda

Foveacheles (Foveacheles) caucasica Zacharda, 1983b: 305; 1993: 60

Type locality: Kabardino-Balkaria, Poljana Azau, 2 km W of Terskol, N-slope of Ceget Mt., 2600 m

Distribution: Czech Republic, Russia (Caucasus)

Foveacheles (Foveacheles) cegetensis Zacharda

Foveacheles (Foveacheles) cegetensis Zacharda, 1983b: 307; 1993: 60; Zacharda & Kučera, 2010: 5; Darbemamieh, Hajiqanbar & Khanjani, 2013: 455

Type locality: Kabardino-Balkaria, Poljana Azau, 2 km W of Terskol, N-slope of Ceget Mt., 2600 – 2800 m

Distribution: Czech Republic, Finland, Iran, Russia (Caucasus)

Foveacheles (Foveacheles) gigantea Zacharda

Foveacheles (Foveacheles) gigantea Zacharda, 2000: 248

Type locality: Ceske Stredohori Mts,

Distribution: Czech Rep.

Foveacheles (Foveacheles) halltalensis Zacharda

Foveacheles (Foveacheles) halltalensis Zacharda, 2000: 253

Type locality: Northern Alps,

Distribution: Austria

Foveacheles (Foveacheles) incognita Zacharda

Foveacheles (Foveacheles) incognita Zacharda, 1980: 670

Type locality: Patria incognita

Distribution: Patria incognita

Foveacheles (Foveacheles) lemoni Abou-Awad, El-Sawaf et Abdel-Khalek

Foveacheles (Foveacheles) lemoni Abou-Awad, El-Sawaf & Abdel-Khalek, 2009: 147

Type locality: El - Kassasin, El – Sharkiya

Distribution: Egypt

Foveacheles (Foveacheles) osloensis (Sig Thor)

Rhagidia osloensis Sig Thor, 1934: 314; Thor & Willmann, 1941: 120

Foveacheles (Foveacheles) osloensis: Zacharda, 1980: 662; Hippa et al., 1989: 61; Darbemamieh, Hajiqanbar & Khanjani, 2013: 455

Type locality: Norway

Distribution: Austria, Czech Republic, Germany, Iran, Norway, Sweden

Foveacheles (Foveacheles) proxima Zacharda

Foveacheles (Foveacheles) proxima Zacharda, 2000: 258

Type locality: Northern Alps

Distribution: Austria

Foveacheles (Foveacheles) thaleri Zacharda

Foveacheles (Foveacheles) thaleri Zacharda, 2010: 58

Type locality: Cedar Valley, Troodos

Distribution: Cyprus

Subgenus Hirschmannetta Zacharda

Hirschmannetta Zacharda, 1980: 694

Type species: Foveacheles (Hirschmannetta) magna Zacharda, 1980

Foveacheles (Hirschmannetta) magna Zacharda

Foveacheles (Hirschmannetta) magna Zacharda, 1980: 695

Type locality: Harz, Bielshöhle Cave

Distribution: Germany

Subgenus Mediostella Zacharda

Mediostella Zacharda, 1980: 672

Type species: Foveacheles (Mediostella) rupestris Zacharda, 1980

Foveacheles (Mediostella) canestrini (Berlese et Trouessart)

Noerneria canestrini Berlese & Trouessart, 1889: 142

Rhagidia gigas var. canestrini: Thor & Willmann, 1941: 112

?Rhagidia fragosa Schuster, 1958: 95

Foveacheles (Mediostella) canestrini: Zacharda, 1980: 679

Type locality: Plage du Croisic (Loire Tufe)

Distribution: France

Foveacheles (Mediostella) pribilovi Zacharda

Foveacheles (Mediostella) pribilovi Zacharda, 1996: 344

Type locality: Alaska, Pribilof Islands, St. Paul’s Island

Distribution: USA (Alaska)

Foveacheles (Mediostella) rupestris Zacharda

Foveacheles (Mediostella) rupestris Zacharda, 1980: 672

Type locality: the High Tatras, Koprovský štit, 2376 m

Distribution: Slovakia

Foveacheles (Mediostella) vancouverensis Zacharda

Foveacheles (Mediostella) vancouverensis Zacharda, 1986: 2542

Type locality: British Columbia, Vancouver Island, SE end of Long Beach

Distribution: Canada

Foveacheles (Mediostella) willmanni Zacharda

Foveacheles (Mediostella) willmanni Zacharda, 1980: 676

Type locality: Bösensteingipfel

Distribution: Austria

Subgenus Propriorhagidia Zacharda

Propriorhagidia Zacharda, 1980: 690

Type species: Foveacheles (Propriorhagidia) mira Zacharda, 1980

Foveacheles (Propriorhagidia) mira Zacharda

Foveacheles (Propriorhagidia) mira Zacharda, 1980: 690

Type locality: Wien

Distribution: Austria

Foveacheles (Propriorhagidia) tuktoyaktukensis Zacharda

Foveacheles (Propriorhagidia) tuktoyaktukensis Zacharda, 1980: 692; 1985: 78

Type locality: Canada, N.W. Territory, Tuktoyaktut, Beaufort Sea

Distribution: Canada (N.W. Territory), USA

Subgenus Proxistella Zacharda

Proxistella Zacharda, 1980: 700

Type species: Scyphius terricola C. L. Koch, 1835

Foveacheles (Proxistella) simulata Zacharda

Foveacheles (Proxistella) simulata Zacharda, 2000: 106

Type locality: Berounka River Valley

Distribution: Czech Rep.

Foveacheles (Proxistella) terricola (C. L. Koch)

Scyphius terricola C. L. Koch, 1835: Fasc. 1, 15

Scyphius terricolus C. L. Koch, 1836: Heft 1, Nr 15

Rhagidia terricola : Willmann, 1936: 294; 1954: 234; Thor, 1912: 235; Halbert, 1915: 110; Vitzthum, 1925: 153; Irk, 1939: 153; Thor & Willmann, 1941: 102; 1951: 146; Macfadyen, 1854: 284; Mihelčič, 1958: 102; Cooreman, 1959: 26; Schmölzer, 1962: 282; 1993: 59; Rack, 1974: 133;

Foveacheles (Proxistella) terricola Zacharda, 1980: 701

Type locality: Germany

Distribution: Austria, Czech Rep., France, Germany, Norway, Switzerland, Yan Mayen I.

Subgenus Spelaeocheles Zacharda

Foveacheles (Spelaeocheles) Zacharda, 1988: 75

Type species: Foveacheles (Spelaeocheles) troglodyta Zacharda, 1988

Foveacheles (Spelaeocheles) troglodyta Zacharda

Foveacheles (Spelaeocheles) troglodyta Zacharda, 1988: 76

Type locality: East Slovakia, Stratenska vysočina

Distribution: Slovakia

Subgenus Ternirhagidia Zacharda

Ternirhagidia Zacharda, 1980: 696

Type species: Foveacheles (Ternirhagidia) alpina Zacharda, 1980

Foveacheles (Ternirhagidia) alpina Zacharda

Foveacheles (Ternirhagidia) alpina Zacharda, 1980: 697

Type locality: the Zillertall Alps, Tetterjoch, 3000 m, Schnoarzensteingipfel, 3320 m

Distribution: Austria

Foveacheles (Ternirhagidia) emendata Zacharda

Foveacheles (Ternirhagidia) emendata Zacharda, 1980: 699

Type locality: Patria incognita

Distribution: Patria incognita

Subgenus Trofocheles Zacharda

Trofocheles Zacharda, 1982: 275

Type species: Foveacheles (Trofocheles) tenorioae Zacharda, 1982

Foveacheles (Trofocheles) goffi Zacharda

Foveacheles (Trofocheles) goffi Zacharda, 1982: 277

Type locality: Hawaii I., Kau Forest Es., Manu Cave, 2100 m

Distribution: USA (Hawaii)

Foveacheles (Trofocheles) paralleloseta Zacharda

Foveacheles (Trofocheles) paralleloseta Zacharda, 1985: 77

Type locality: Sam Six Cave, 4 mi. W Wytheville, Wythe Co., Virginia

Distribution: USA (Virginia)

Foveacheles (Trofocheles) tenorioae Zacharda

Foveacheles (Trofocheles) tenorioae Zacharda, 1982: 278

Type locality: Hawaii I., Hawaii Volcanoes Nat. Park, Kipuka Puaulu, Cave No 3

Distribution: USA (Hawaii)

Subgenus Usitorhagidia Zacharda

Usitorhagidia Zacharda, 1980: 682

Type species: Rhagidia arenaria Willmann, 1952

Foveacheles (Usitorhagidia) arenaria (Willmann)

Rhagidia arenaria Willmann, 1952: 155; Ehrnsberger, 1977: 67

Foveacheles (Usitorhagidia) arenaria Zacharda, 1980: 682

Type locality: Germany, Wangerooge Islands

Distribution: Germany (Wangerooge Islands)

Foveacheles (Usitorhagidia) bentonensis Zacharda

Foveacheles (Usitorhagidia) bentonensis Zacharda, 1983: 1000

Type locality: Oregon

Distribution: USA (Oregon)

Foveacheles (Usitorhagidia) clavacrinita (Ehrnsberger)

Rhagidia clavacrinita Ehrnsberger, 1977a: 95; 1977b: 67

Foveacheles (Usitorhagidia) clavacrinita Zacharda, 1980: 685

Type locality: Kiel, beach forest litter

Distribution: Germany

Foveacheles (Usitorhagidia) harzensis Zacharda

Foveacheles (Usitorhagidia) harzensis Zacharda, 1980: 688

Type locality: Harz, Einhornhöhle-Cave

Distribution: Germany

Foveacheles (Usitorhagidia) mexicana Zacharda

Foveacheles (Usitorhagidia) mexicana Zacharda, 1996: 350

Type locality: Mexico, 15 mi SW El Salto, Durango

Distribution: Mexico

Foveacheles (Usitorhagidia) titanica Zacharda et Elliott

Foveacheles (Usitorhagidia) titanica Zacharda & Elliott, 1985: 471

Type locality: California,

Distribution: USA (California)

Foveacheles (Usitorhagidia) unguiculata Zacharda

Foveacheles (Usitorhagidia) unguiculata Zacharda, 1994: 86

Type locality: North Tyrol, the Ötztal Alps, Pitztal, Wildgrat 2974 m

Distribution: Austria

Genus Hammenia Zacharda

Hammenia Zacharda, 1980: 760

Type species: Hammenia macrostella Zacharda, 1980

Hammenia macrostella Zacharda

Hammenia macrostella Zacharda, 1980: 760

Type locality: Bohemia bor., Velký Bezdĕz-Hill

Distribution: Czech Republic

Genus Janes Jesionowska

Janes Jesionowska, 2002: 174

Type species: Janes rafalskii Jesionowska, 2002

Janes rafalskii Jesionowska

Janes rafalskii Jesionowska, 2002: 174

Type locality: Poland

Distribution: Poland

Genus Kouchibouguacia Zacharda

Kouchibouguacia Zacharda, 1986: 2532

Type species: Kouchibouguacia cryptosolenidiata Zacharda, 1986

Kouchibouguacia cryptosolenidiata Zacharda

Kouchibouguacia cryptosolenidiata Zacharda, 1986: 2533

Type locality: New Brunswick, Kouchibouguac Nat. Park, Kollock Creek trail

Distribution: Canada

Kouchibouguacia transcendata Zacharda

Kouchibouguacia transcendata Zacharda, 1995: 589

Type locality: Canada, Ontario, St. Lawrence Islands Nat. Park, Grenadier Island

Distribution: Canada (Alberta, Ontario, New Brunswick), USA (New Mexico)

Genus Latoempodia Zacharda

Latoempodia Zacharda, 1980: 585

Type species: Latoempodia macroempodiata Zacharda, 1980

Latoempodia macroempodiata Zacharda

Latoempodia macroempodiata Zacharda, 1980: 585

Type locality: Oblik-Hill near Louny

Distribution: Czech Republic

Latoempodia similis Zacharda

Latoempodia similis Zacharda, 1980: 589

Type locality: the High Tatra, Mengusovská dolina Valley, 1730 m

Distribution: Slovakia

Genus Lindquistula Zacharda

Lindquistula Zacharda, 1986: 2531

Type species: Lindquistula multisolenidiata Zacharda, 1980

Lindquistula multisolenidiata Zacharda

Lindquistula multisolenidiata Zacharda, 1986: 2531

Type locality: Ontario, Pt. Pelee Nat. Park

Distribution: Canada

Genus Neothoria Abou-Awad et El-Bagoury

Neothoria Abou-Awad & El-Bagoury, 1986: 59

Type species: Neothoria niloticus Abou-Awad et El-Bagoury, 1986

Neothoria niloticus Abou-Awad et El-Bagoury

Neothoria niloticus Abou-Awad & El-Bagoury, 1986: 59

Type locality: Lower Egypt

Distribution: Egypt

Neothoria zacharda Abu-Awad

Neothoria zacharda Abu-Awad, 2010: 141

Type locality: Egypt

Distribution: Egypt

Genus Paracrassocheles Khaustov

Paracrassocheles Khaustov, 2015: 619

Type species: Paracrassocheles clavisetosus Khaustov, 2015

Paracrassocheles clavisetosus Khaustov

Paracrassocheles clavisetosus Khaustov, 2015: 619

Type locality: Crimea

Distribution: Crimea

Genus Parallelorhagidia Zacharda

Parallelorhagidia Zacharda, 1980: 590

Type species: Coccorhagidia evansi Strandtmann et Prasse, 1976

Parallelorhagidia evansi (Strandtmann et Prasse)

Coccorhagidia evansi Strandtmann & Prasse, 1976: 12

Parallelorhagidia evansi (Strandtmann et Prasse): Zacharda, 1980: 591

Type locality: Etzsdorf/ Saalkreis (East Germany)

Distribution: Czech Republic, Germany

Parallelorhagidia hawaiiensis Zacharda

Parallelorhagidia hawaiiensis Zacharda, 1980: 594; Goff, 1987: 35

Type locality: Hawaii, Big Island

Distribution: USA (Hawaii)

Parallelorhagidia parva Zacharda

Parallelorhagidia parva Zacharda, 1986: 2538

Type locality: New Brunswick, Kouchibouguac Nat. Park, Kouchibouguac Campground

Distribution: Canada

Genus Poecilophysis Cambridge

Poecilophysis Cambridge, 1876: 261

Type species: Poecilophysis kerguelenensis Cambridge, 1876

Subgenus Dentocheles Zacharda

Dentocheles Zacharda, 1980: 621

Type species: Scyphius pratensis C. L. Koch, 1835

Poecilophysis (Dentocheles) affinis Zacharda

Poecilophysis (Dentocheles) affinis Zacharda, 1983: 996

Type locality: Oregon, Coffin Butte

Distribution: USA (Oregon)

Poecilophysis (Dentocheles) extraneostella Zacharda

Poecilophysis (Dentocheles) extraneostella Zacharda, 1985: 73

Type locality: Bowling Cave near Ben Hur, Lee Co., Virginia

Distribution: USA (Virginia)

Poecilophysis (Dentocheles) melanoseta Zacharda et Elliot

Poecilophysis (Dentocheles) melanoseta Zacharda & Elliot, 1985: 646

Type locality: California

Distribution: USA (California)

Poecilophysis (Dentocheles) pratensis (C. L. Koch)

Scyphius pratensis C. L. Koch, 1835: fasc. 1, 14

Poecilophysis pratensis : Oudemans, 1905: 202; Goff, 1987: 35; Zacharda & Kučera, 2010: 5

Rhagidia pratensis : Willmann, 1932: 5; 1936: 229; 1951b: 143; 1954: 234; Sellnick, 1940: 31; Haarløv, 1957: 31; Cooreman, 1959: 20

Rhagidia japonica Morikawa, 1963: 15; Shiba, 1969: 76

Rhagidia whartoni Strandtmann, 1971: 107

Poecilophysis (Dentocheles) pratensis Zacharda, 1980: 621; 1983b: 312; 1985: 74

Type locality: Germany

Distribution: Alaska, Austria, Azerbaidzhan, Belgium, Czech Republic, Denmark, Finland, France, Germany, Hawaii, Iceland, Iran, Japan, Russia, Slovakia, Switzerland, USA

Poecilophysis (Dentocheles) wankeli (Zacharda)

Rhagidia reflexa (part.?): Wankel, 1936: 301; 1953: 218

Rhagidia wankeli Zacharda, 1978: 226

Poecilophysis (D.) wankeli Zacharda, 1980: 630

Type locality: Moravia

Distribution: Czech Rep., Slovakia, Austria

Poecilophysis (Dentocheles) weyerensis (Packard)

Bryobia ? (or Penthaleus ?) weyerensis Packard, 1888: 42

Poecilophysis (D.) weyerensis Zacharda, 1980: 627; 1985: 74; Hippa et al., 1989: 61

Rhyncholophus cavernarum Packard, 1888: 42

Type locality: Augusta County, Virginia, Weyer’s Cave, USA

Distribution: Sweden, USA

Subgenus Poecilophysis Cambridge

Poecilophysis Cambridge, 1876: 261; Zacharda, 1980: 601

Type species: Poecilophysis kerguelenensis Cambridge, 1876

Poecilophysis (P.) kerguelenensis Cambridge

Poecilophysis kerguelenensis Cambridge, 1876: 261

Rhagidia megalochela Trägårdh, 1907: 20

Rhagidia kerguelensis : André, 1947: 78

Rhagidia kerguelensis (part.): Strandtmann & Davies, 1972: 39

Poecilophysis (P.) kerguelenensis: Zacharda, 1980: 602

Type locality: Subantarctic region

Distribution: Possession I., Crozet I., Saint-Paul I., Kerguelen Islands,

Poecilophysis (P.) macquariensis (Womersley et Strandtmann)

Rhagidia macquariensis Womersley & Strandtmann, 1963: 465

Poecilophysis macquariensis Zacharda: Qin, 1998: 300

Poecilophysis (P.) macquariensis Zacharda, 1980: 606

Type locality: Macquarie Island

Distribution: Macquarie Island, Antarctica

Subgenus Procerocheles Zacharda

Procerocheles Zacharda, 1980: 609

Type species: Rhagidia gelida faeroensis Trägårdh, 1931

Poecilophysis (Procerocheles) faeroensis (Trägårdh)

Rhagidia gelida faeroensis Trägårdh, 1931: 42

Rhagidia reflexa Willmann, 1936: 300; Thor & Willmann, 1941: 104 (part.?); Cooreman, 1959: 22

Poecilophysis (Procerocheles) faeroensis Zacharda, 1980: 609; Hippa et al., 1989: 61

Type locality: Faroe Isl., Thorshawn (Denmark)

Distribution: Denmark (Faroe Is.), Germany, Czech Republic, Iran, Slovakia, Switzerland

Poecilophysis (Procerocheles) oregonensis Zacharda

Poecilophysis (Procerocheles) oregonensis Zacharda, 1983: 995

Type locality: Oregon, Washington Co.

Distribution: USA (Oregon)

Poecilophysis (Procerocheles) pseudoreflexa Zacharda

Poecilophysis (Procerocheles) pseudoreflexa Zacharda, 1980: 614; 1983: 311; Hippa et al., 1989: 61; Zacharda & Kučera, 2010: 5

Type locality: Oblik-Hill near Louny (Czech Rep.)

Distribution: Canada, Czech Republic, Finland, Kirghizstan, Russia, Slovakia, Sweden

Poecilophysis (Procerocheles) recussa (Sig Thor)

Rhagidia recussa Sig Thor, 1909: 14; Leruth, 1939: 412

Poecilophysis (Procerocheles) recussa (Sig Thor): Zacharda, 1993: 52

Type locality: Shara-Ullah Mts., North Siberia

Distribution: Belgium, Czech Republic, Russia

Poecilophysis (Procerocheles) spelaea (Wankel)

Scyphius spelaeus Wankel, 1861: 258

Rhagidia recussa Sig Thor, 1909: 14 ?

Rhagidia spelaea (Wankel): Willmann, in Thor & Willmann, 1941: 108; Willmann, 1954: 232

Poecilophysis (Procerocheles) spelaea: Zacharda, 1980: 617; Zacharda & Kučera, 2010: 5

Type locality of Scyphius spelaeus: Moravia

Type locality of Rhagidia recussa: Shara-Ullah-Mountains, North Siberia

Distribution: Czech Republic, England, Finland, France, Germany, Belgium, Poland, Romania, Switzerland; Siberia ?

Subgenus Soprocheles Zacharda

Soprocheles Zacharda, 1980: 634

Type species: Rhagidia saxonica Willmann, 1934

Poecilophysis (Soprocheles) arena Zacharda

Scyphius elongatus : Valle, 1955: 17, nec Scyphius elongatus C. L. Koch, 1838

Poecilophysis (Soprocheles) arena Zacharda, 1980: 639

Type locality: Moravia Mer., Valtice

Distribution: Czech Republic, Iran, Italy, Hawaii

Poecilophysis (Soprocheles) saxonica (Willmann)

Rhagidia saxonica Willmann, 1934: 306; 1936: 297

? Coccorhagidia semiclavifrous Shiba, 1969: 83

Rhagidia shibai Strandtmann, 1971: 111

Poecilophysis (Soprocheles) saxonica: Zacharda, 1980: 635; 1983b: 312

Poecilophysis saxonica : Zacharda & Daniel, 1988: 58; Zacharda & Kučera, 2010: 5

Type locality: Leipzig, Anwald b. Dölitz (Germany)

Distribution: Czech Republic, Finland, Germany, Nepal (3900 m), Slovakia, Kirghizstan

Subgenus Wankelia Zacharda

Wankelia Zacharda, 1980: 642

Type species: Rhagidia reflexa wolmsdorfensis Willmann, 1936

Poecilophysis (Wankelia) wolmsdorfensis (Willmann)

Rhagidia terricola longipes : Pax, Maschke, 1935: 13

Rhagidia wolmsdorfensis : Willmann, 1941: 53; Willmann, 1954: 232; Baltac, 1973: 167

Rhagidia reflexa wolmsdorfensis Willmann, 1936: 302

Poecilophysis (Wankelia) wolmsdorfensis: Zacharda, 1980: 642; 1985: 74

Type locality: Wolmsdorf, Germany

Distribution: Czech Republic, Germany, Poland, Romania, USA, “Yugoslavia”

Genus Rhagidia Thorell

Megamerus Dugès, 1834: 50, 53 (nec Megamerus W.S. Mac Leay, 1827)

Scyphius C. L. Koch, 1835: fasc. 1, 14

Rhagidia Thorell, 1872: 700; Zacharda, 1980: 558

Nörneria Canestrini, 1886: 697

Scyphioides Berlese, 1886: 16, nom. nov. for Scyphius C. L. Koch nec Risso

Scyphoides Banks, 1900: 31 (lapsus calami?)

Type species: Rhagidia gelida Thorell, 1872

Subgenus Austrorhagidia Zacharda

Austrorhagidia Zacharda, 1980: 560

Type species: Rhagidia mildredae Strandtmann, 1964

Rhagidia (Austrorhagidia) campbellensis Zacharda

Rhagidia mildredae (part.): Strandmann, 1964: 157-160

Rhagidia campbellensis Zacharda: Qin, 1998: 300

Rhagidia (Austrorhagidia) campbellensis Zacharda, 1980: 563

Type locality: Auckland I.

Distribution: Auckland I.

Rhagidia (Austrorhagidia) gerlachei Trouessart

Rhagidia gigas gerlachei Trouessart, 1903: 4-5

Rhagidia gerlachei Trouessart: Womersley & Strandtmann, 1963: 460; Strandtmann, 1967: 65; 1970: 98

Rhagidia (Austrorhagidia) gerlachei Zacharda, 1980: 563

Type locality: Antarctica, Détroit de Gerlache, South Georgia

Distribution: Antarctica (South Georgia)

Rhagidia (Austrorhagidia) mildredae Strandtmann

Rhagidia mildredae Strandtmann, 1964: 157-160; Qin, 1998: 300

Rhagidia (Austrorhagidia) mildredae Zacharda, 1980: 561

Type locality: New Zealand, Beeman Hill, Campbell Island

Distribution: Subantarctic region (Auckland Is., Campbell I.)

Subgenus Deharvengiella Zacharda

Deharvengiella Zacharda, 1987: 2051

Type species: Rhagidia (Deharvengiella) troglomorphica Zacharda, 1987

Rhagidia (Deharvengiella) paraleloseta Zacharda

Rhagidia (Deharvengiella) paraleloseta Zacharda, 2011: 669

Type locality: France

Distribution: France

Rhagidia (Deharvengiella) serpentiforma Zacharda

Rhagidia (Deharvengella) serpentiforma Zacharda, 2011: 672

Type locality:

Distribution:

Rhagidia (Deharvengiella) troglomorphica Zacharda

Rhagidia (Deharvengiella) troglomorphica Zacharda, 1987: 2051

Type locality: the Pyrenees, Haute-Garonne, Couledoux, Grotte de la Buhadére

Distribution: France

Subgenus Dongnaia Zacharda

Dongnaia Zacharda, 1994: 49

Type species: Dongnaia edaphica Zacharda, 1994

Rhagidia (Dongnaia) edaphica Zacharda

Rhagidia (Dongnaia) edaphica Zacharda, 1994: 51

Type locality: Nam Cat Tien National Park, South Vietnam

Distribution: Vietnam

Subgenus Noerneria Canestrini

Nörneria Canestrini, 1886: 708; Zacharda, 1980: 565

Type species: Nörneria gigas Canestrini, 1886

Rhagidia (Noerneria) arizonensis Zacharda

Rhagidia (Noerneria) arizonensis Zacharda, 1995: 1259

Type locality: Arizona, Madera Canyon above Roundup Camp

Distribution: USA

Rhagidia (Noerneria) breviseta Zacharda

Rhagidia breviseta Zacharda, 1995: 1248; Zacharda & Kučera, 2010: 5

Type locality: Quebec

Distribution: Canada, Finland

Rhagidia (Noerneria) compressa Zacharda

Rhagidia (Noerneria) compressa Zacharda, 1986: 2538

Type locality: British Columbia, Domonion Lake

Distribution: Canada

Rhagidia (Noerneria) distisolenidiata Zacharda

Rhagidia (Noerneria) distisolenidiata Zacharda, 1995: 1258

Type locality: New-Brunswick

Distribution: Canada

Rhagidia (Noerneria) gigas (Canestrini)

Nörneria gigas Canestrini, 1886: 708 -709

Rhagidia gigas : Trägårdh, 1904b: 3; Hull, 1918: 34; Thor & Willmann, 1941: 111-112; Ružička, Hajer & Zacharda, 1995: 47; Zacharda & Kučera, 2010: 5

Rhagidia intermedia Willmann, 1936: 293-294; 1951: 146; Haarløv, 1957: 27; Schmölzer, 1993: 59

Rhagidia danica Ehrnberger, 1977b: 95-100

Rhagidia carpatica Baltac, 1973: 168-171

Rhagidia (Nörneria) gigas Zacharda, 1980: 566

Type locality: Italy, Trentino

Distribution: Austria, Czech Republic, Denmark, Finland, Germany, Italy, Slovakia,

Rhagidia (Noerneria) longiseta Zacharda

Rhagidia (Noerneria) longiseta Zacharda, 2000: 113; Zacharda & Kučera, 2010: 5

Type locality: Austria

Distribution: Austria, Finland

Rhagidia (Noerneria) parvilobata Zacharda

Rhagidia (Noerneria) parvilobata Zacharda, 1995: 1252; Zacharda & Kučera, 2010: 5

Type locality: Quebec

Distribution: Canada, Finland

Rhagidia (Noerneria) pasadenensis Zacharda

Rhagidia (Noerneria) pasadenensis Zacharda, 1986: 2535

Type locality: Newfoundland, Pasadena

Distribution: Canada

Rhagidia (Noerneria) ponderosae Zacharda

Rhagidia (Noerneria) ponderosae Zacharda, 1983: 1000

Type locality: Baker Co., S Fork Forest Camp, 9 mi SW Unity

Distribution: USA (Oregon)

Rhagidia (Noerneria) punkva Zacharda

Rhagidia (Noerneria) punkva Zacharda, 1980: 575; 1983b: 314

Type locality: Moravia, Skalni mlýn, Czech Republic

Distribution: Czech Republic, Russia (Siberia)

Rhagidia (Noerneria) quadrisolenidiata Zacharda

Rhagidia (Noerneria) quadrisolenidiata Zacharda, 1995: 1262

Type locality: 53 mi north of Oaxaca, km 77, Oaxaca Highway 175, 3200 m

Distribution: Mexico

Rhagidia (Noerneria) rackae Zacharda

Rhagidia (Noerneria) rackae Zacharda, 1980: 582; Zacharda & Daniel, 1988: 58

Type locality: Velký-Bezdĕz-Hill, Czech Republic

Distribution: Czech Republic, Nepal (3900 m)

Rhagidia (Noerneria) ruseki Zacharda

Rhagidia (Noerneria) ruseki Zacharda, 1980: 578; Hippa et al., 1989: 61

Type locality: Oblik-Hill near Louny, Czech Republic

Distribution: Czech Republic, Sweden

Rhagidia (Noerneria) varia Zacharda

Rhagidia (Noerneria) varia Zacharda, 1985: 68

Type locality: Butler Cave, N of Burnsville, Both Co., Virginia

Distribution: USA

Subgenus Rhagidia Thorell

Rhagidia Thorell, 1872: 700; Zacharda, 1980: 558

Type species: Rhagidia gelida Thorell, 1872

Rhagidia (Rhagidia) comelicensis Lombardini

Rhagidia comelicensis Lombardini, 1962: 191

Type locality: Comelico (Alpi orientali)

Distribution: Italy

Rhagidia crenata Shiba

Rhagidia crenata Shiba, 1969a: 77; Shiba, 1971a: 158; 1971b: 226

Type locality: Shiga Heights IBP Area

Distribution: Japan

Rhagidia (Rhagidia) diversicolor (C. L. Koch)

Scyphius diversicolor C. L. Koch, 1838: fasc. 17, 22

Rhagidia diversicolor : Willmann, 1936: 297-298; 1954: 234; Cooreman, 1954: 10; 1959: 17; Zacharda & Kučera, 2010: 5

Rhagidia (Nörneria) diversicolor: Zacharda, 1980: 570

Rhagidia (Noerneria) diversicolor: Zacharda, 1985: 71

Type locality: France, Germany

Distribution: Czech Republic, Finland, Germany, Poland, Switzerland, USA

Rhagidia (Rhagidia) gelida Thorell

Rhagidia gelida : Thorell, 1872: 701; Trägårdh, 1904: 44; Thor, 1912: 235; Banks, 1919: 11; Oudemans, 1926: 106; Sellnick, 1940: 31; Thor & Willmann, 1941: 109-111; Macfadyen, 1954: 284; Cooreman, 1959: 27 (var.); Strandtmann, 1971: 106-107; Seniczak et al., 2020: 1

Rhagidia (R.) gelida Zacharda, 1980: 559

Type locality: Norway, Beeren Eiland, Svalbard

Distribution: Canada, Czech Rep., Greenland, Yan Mayen I., Iceland, Norway, Russia, Spitzbergen, Switzerland, USA (Alaska)

Rhagidia (Rhagidia) orghidani Bannova, Sarmanova et Zhuravleva

Rhagidia orghidani Bannova, Sarmanova & Zhuravleva, 1977: 57

Type locality: Romania

Distribution: Romania

Rhagidia (Rhagidia) qaliubiensis Abou-Awad, El-Sawaf et Abdel-Khalek

Rhagidia (Rhagidia) qaliubiensis Abou-Awad, El-Sawaf & Abdel-Khalek, 2009: 150

Type locality: El-Kaliubiya region

Distribution: Egypt

Rhagidia buettneri Willmann

Rhagidia buettneri Willmann, 1955: 209; Zacharda, 1980 (sp. inc. sed., “The holotype fragmentary, a tritonymph”)

Type locality: SW Saxony, Brummeisenmoor

Distribution: Germany

Rhagidia meyerae Halliday

Rhagidia meyerae Halliday, 2005: 42

Type locality: South Africa

Distribution: South Africa

Rhagidia uenoi Shiba

Rhagidia uenoi Morikawa, 1963: 18; Shiba, 1971: 224

Type locality: Japan

Distribution: Japan

Genus Robustocheles Zacharda

Robustocheles Zacharda, 1980: 527

Type species: Robustocheles mountana Zacharda, 1980

Subgenus Amoveocheles Zacharda

Amoveocheles Zacharda, 1980: 540

Type species: Robustocheles (Amoveocheles) tricuspidata Zacharda, 1980

Robustocheles (Amoveocheles) californiensis Zacharda

Robustocheles (Amoveocheles) californiensis Zacharda, 1986: 2535

Type locality: California, Albany

Distribution: USA (California)

Robustocheles (A.) dentata Zacharda

Robustocheles (A.) dentata Zacharda, 1980: 543

Type locality: Prague, Czech Republic

Distribution: Czech Republic

Robustocheles (A.) tricuspidata Zacharda

Robustocheles (A.) tricuspidata: Zacharda, 1980: 540

Type locality: Závist-Knoll near Zbraslav, Czech Republic

Distribution: Czech Republic

Subgenus Lewia Zacharda

Lewia Zacharda, 1980: 547

Type species: Rhagidia hilli Strandtmann, 1971

Robustocheles (Lewia) hilli (Strandtmann)

Rhagidia hilli Strandtmann, 1971: 113

Rhagidia grahami Elliott, 1976: 11-14

Robustocheles (Lewia) hilli Zacharda, 1980: 547; 1985: 79

Type locality of Rhagidia hilli: Alaska, Fairbanks

Type locality of Rhagidia grahami: Unita Co., Little Brush Creek Cave, Utah

Distribution: Canada, N-W Territorium, Bathurst I.; USA

Robustocheles (Lewia) lacustris Zacharda

Robustocheles (Lewia) lacustris Zacharda, 1997: 1090

Type locality: Cypress Hills Provincial Park (Alberta – Saskatchewan)

Distribution: Canada

Robustocheles (Lewia) obscuritata Zacharda

Robustocheles (Lewia) obscuritata Zacharda, 1997: 1094; Zacharda & Kučera, 2010: 5

Type locality: Gaspésie Provincial Park (Quebec)

Distribution: Canada, Finland

Robustocheles (Lewia) spinaea Zacharda

Robustocheles (Lewia) spinaea Zacharda, 1997: 1094

Type locality: 20 m SE Gallup, New Mexico

Distribution: USA

Subgenus Mexia Zacharda

Mexia Zacharda, 1997: 1084

Type species: Robustocheles (Mexia) longichelae Zacharda, 1996

Robustocheles (Mexia) longichelae Zacharda

Robustocheles (Mexia) longichelae Zacharda, 1997: 1085

Type locality: Chiapas, 7 m S El Bosque

Distribution: Mexico

Robustocheles (Mexia) occulta Zacharda et Pugsley

Robustocheles (Mexia) occulta Zacharda & Pugsley, 1988: 646

Type locality: Alberta (Banff Nat. Park, Castleguard Cave); Iowa, Washington

Distribution: Canada, USA

Subgenus Robustocheles Zacharda

Robustocheles Zacharda, 1980: 528

Type species: Robustocheles montana Zacharda, 1980

Robustocheles (R.) deltacus Abou-Awad, El-Sawat et Abdel-Khalek

Robustocheles (R.) deltacus Abou-Awad, El-Sawat & Abdel-Khalek, 2009: 145

Type locality: Ain Shams Univ., El Qaluibiya

Distribution: Egypt

Robustocheles (R.) hamedanensis Zacharda

Robustocheles hamedanensis Zacharda, 2012: 31

Type locality: Hamedan-Shokriyeh

Distribution: Iran

Robustocheles (R.) montana Zacharda

Robustocheles (R.) montana Zacharda, 1980: 529

Type locality: Velká Fatra-Mountains, Černy Kameň, Slovakia

Distribution: Germany, Slovakia

Robustocheles (R.) mucronata (Willmann)

Rhagidia mucronata Willmann, 1936: 297; 1951b: 143; Sellnick, 1940: 31; Evans, 1952: 666; Cooreman, 1954: 10; Haarløv, 1957: 31; Strandtmann & Prasse, 1977: 14

Robustocheles (R.) mucronata (Willmann): Zacharda, 1980: 532; Abou-Awad, 1985: 258; Zacharda & Kučera, 2010: 5; Darbemamieh, Hajiqanbar & Khanjani, 2013: 456

Rhagidia neoshibai Shiba, 1976: 132; syn.: Zacharda, 1984: 870

Robustocheles mucronata (Willmann): Zacharda, 1994: 53

Type locality: Nuremberg, Germany

Distribution: Brazil, Corsica, Czech Republic, Danemark, Egypt, England, Finland, Germany, Iceland, Iran, Poland, Russia (Caucasus), Slovakia, Thailand, USA (Alaska), Vietnam

Robustocheles (R.) pascuaria Zacharda

Robustocheles (R.) pascuaria Zacharda, 1997: 1076

Type locality: Cypress Hills Provincial Park

Distribution: Canada

Robustocheles (R.) robusta Zacharda

Rhagidia mordax grandis Willmann, 1935: 28; 1936: 296; Cooreman, 1959: 19; Leruth, 1939: 411

Robustocheles (R.) robusta Zacharda, 1980: 537

Type locality: Czech Republic, Oblik Hill

Distribution: Belgium, Germany, Czech Republic, Switzerland

Robustocheles (R.) similis Zacharda

Robustocheles (R.) similis Zacharda, 1997: 1080

Type locality: Nova Scotia, Cape Breton Highlands

National Park, Paquettes Lake Trail

Distribution: Canada

Genus Shibaia Zacharda

Shibaia Zacharda, 1980: 737

Type species: Rhagidia longisensilla Shiba, 1969

Shibaia coreaensis Zacharda

Rhagidia coreaensis Shiba, 1969: 608

Shibaia coreaensis Zacharda, 1980: 742; 1985: 598

Type locality: Samsheong-gun, Kangweon-do, Yongyeon-gul Cave near Yongyeon-gog, Changseong-eub

Distribution: South Korea

Shibaia heteropoda (Berlese)

Rhagidia heteropoda Berlese, 1910: 243

Rhagidia longisensilla Shiba, 1969a: 79; Baltac, 1974: 37

Type locality: “in muscis Etruriae”, Toiano, Piza

Distribution: Italy, Japan, Romania

Shibaia longisensilla (Shiba)

Rhagidia longisensilla Shiba, 1969: 79; 1969b: 608; Strandtmann, 1971: 104; ? Elliott & Strandtmann, 1971: 475; Baltac, 1974: 41; Strandtmann & Goff, 1978: 135; Goff, 1987: 35; Darbemamieh, Hajiqanbar & Khanjani, 2013: 456

Shibaia longisensilla Zacharda, 1980: 738; Zacharda & Daniel, 1988: 58

Shibaia longicornis (Shiba, 1959) - Iran

Type locality: Japan

Distribution: Austria (3133 m), Czech Republic; Germany, Iran, Japan, Alaska, Hawaii, Korea, Nepal (3900 m)

Shibaia shereefi Abou-Awad

Shibaia shereefi Abou-Awad, 1985: 253

Type locality: Giza region

Distribution: Egypt

Shibaia spiraesolenidiata Zacharda

Shibaia spiraesolenidiata Zacharda, 1995: 594

Type locality: Nova Scotia, Kejim Kujuk Nat. Park

Distribution: Canada

Shibaia tatrica Zacharda

Shibaia tatrica Zacharda, 1980: 744

Type locality: High-Tatras, Mlynická dolina, 1450 m

Distribution: Slovakia

Genus Thoria Zacharda

Thoria Zacharda, 1980: 747

Type species: Thoria brevisensilla Zacharda, 1980

Thoria brevisensilla Zacharda

Rhagidia uniseta : Strandtmann, 1971: 105 (not R. uniseta Sig Thor, 1909)

Thoria brevisensilla Zacharda, 1980: 748; Goff, 1987: 35

Type locality: Bohemia bor., Oblik-Hill near Luny

Distribution: Czech Republic, Hawaii

Thoria uniseta s. I. (Sig Thor)

Rhagidia uniseta Sig Thor, 1909: 10; 1912: 236; Sellnick, 1940: 31; Thor & Willmann, 1941: 117; Mihelčič, 1958a: 103; Strandtmann, 1971: 105; Schmölzer, 1993: 59;

?Rhagidia crassipes Schweizer & Bader, 1963: 246

Thoria uniseta Zacharda, 1980: 752; Zacharda & Kučera, 2010: 5

Type locality: Norway

Distribution: Alaska, Austria, Czech Republic, Finland, Iceland, Norway, Switzerland

Genus Traegaardhia Zacharda

Traegaardhia Zacharda, 1980: 715

Type species: Rhagidia dalmatina Willmann, 1939

Traegaardhia cavernarum Zacharda

Traegaardhia cavernarum Zacharda, 2010: 26

Type locality: Buso del Marmo Cave (1202 V/BL), Quero (Belluno), Massiccio del Monte Grappa, Veneto, Italy (340 m)

Distribution: Italy

Traegaardhia cavernicola Zacharda

Traegaardhia cavernicola Zacharda, 2010: 13

Type locality: Grotte Vecchia Diga Cave, Barcis (Pordenone), Monte Cavallo

mountain range, the Stubai Alps, Friuli

Distribution: Italy

Traegaardhia dalmatina (Willmann)

Rhagidia dalmatina Willmann, 1939: 244; 1951: 146; Mihelčič, 1958a: 103; 1958c: 38; Strandtmann, 1971: 103

? Rhagidia gigas var. longipes Trägårdh, 1912: 608

? Rhagidia longipes: Cooreman, 1959: 18

Rhagidia terricola var. longipes (Trägårdh): Verhoeff, 1912: 155

Traegaardhia dalmatina Zacharda, 1980: 715

Type locality: Croatia, Dalmatia, cave near Krivošije

Distribution: Austria, Croatia, ?Germany, France, Spain

Traegaardhia distosolenidia Zacharda

Traegaardhia distosolenidia Zacharda, 2010: 36; Çetin et al., 2021: 232

Type locality: Arcugnano (Vicenza), the Monti Berici Mts, Veneto

Distribution: Italy, Turkey

Traegaardhia gracilis Zacharda

Traegaardhia gracilis Zacharda, 2010: 31

Type locality: the Boia valley, Valdagno(Vicenza), the Lessinian Mts, Venetia

Distribution: Italy

Traegaardhia holsingeri (Zacharda)

Foveacheles (Mediostella) holsingeri Zacharda, 1980: 677

Foveacheles cf. holsingeri: Zacharda, 1985: 78

Traegaardhia holsingeri (Zacharda): Zacharda et al., 2010: 45

Type locality: Kentucky, Mammoth Cave

Distribution: USA

Traegaardhia nasuta Zacharda

Traegaardhia nasuta Zacharda, 2010: 21

Type locality: Buso dei Zambon Cave, San Pietro di Feletto (Treviso), the Hills of

Conegliano, Veneto, northeastern Italy

Distribution: Italy

Traegaardhia similis Zacharda

Traegaardhia similis Zacharda, 2010: 40

Type locality: Toney Bend Mine No. 2, Marion Co., Arkansas

Distribution: Italy

Traegaardhia subterranea Zacharda

Traegaardhia subterranea Zacharda, 2010: 52

Type locality: Amarcord Cave (6391 V/VI), Valdagno (Vicenza), Lessinian Mts

Distribution: Italy

Traegaardhia vicenzaensis Zacharda

Traegaardhia vicenzaensis Zacharda, 1994b: 93

Type locality: Veneto, M. Lessini, Torrebelvicino (Vicenza); quota 386 in a cave, Buso del Monte Naro

Distribution: Italy

Genus Troglocheles Zacharda

Troglocheles Zacharda, 1980: 646

Type species: Rhagidia strasseri Willmann, 1932

Troglocheles aggerata Zacharda

Troglocheles aggerata Zacharda, 2000: 472

Type locality: Tyrol, Oetztal Alps, Obergurgl

Distribution: Austria

Troglocheles archetypica Zacharda

Troglocheles archetypica Zacharda, 2000: 464

Type locality: Tyrol, Oetztal Alps, Obergurgl

Distribution: Austria

Troglocheles christiani Zacharda, Isaia et Piva

Troglocheles christiani Zacharda, Isaia & Piva, 2011: 656

Type locality: Türkenloch Cave near Kleinzell, 721 m

Distribution: Austria

Troglocheles gineti (Cooreman)

Rhagidia gineti Cooreman, 1959: 23; Strandtmann, 1971: 103; Dobat, 1976

?Rhagidia conciana Lombardini, 1951: 247

Troglocheles gineti : Zacharda, 1980: 655

Type locality: Ain, Grotte de la Buire

Distribution: France, Germany, Switzerland

Troglocheles lanai Zacharda, Isaia et Piva

Troglocheles lanai Zacharda, Isaia & Piva, 2011: 649

Type locality: Vallone dell’Infernotto

Distribution: Italy

Troglocheles odontochela (Turk)(species dubia)

Rhagidia odontochela Turk, 1972: 190

Troglocheles odontochela (Turk): Zacharda, 1984: 869

Type locality: England

Distribution: England

Troglocheles quinquesolenidiata Zacharda, Isaia et Piva

Troglocheles quinquesolenidiata Zacharda, Isaia & Piva, 2011: 643

Type locality: Buso Brutto Cave, 918 m

Distribution: Italy

Troglocheles strasseri (Willmann)

Rhagidia strasseri Willmann, 1932: 3; Thor & Willmann, 1941: 119; Mihelčič, 1958; Cooreman, 1954: 11; 1959: 22

Rhagidia gineti : Rack, 1974: 129, not gineti Cooreman

Troglocheles strasseri Zacharda, 1980: 650; 2000:

Type locality: shaft near Na Lazi, Tarnowaner Wald -Trnovski Gozd-Forest, Slovenia (lectotype)

Distribution: Austria, France, Slovenia, Switzerland

Troglocheles strasseri tirolensis Willmann

Rhagidia strasseri tirolensis Willmann (unpubl.); Zacharda, 1980: 650

Type locality: Austria, the Kaisergebirge-Mts., Estendorfer Schacht-shaft

Distribution: Austria

Troglocheles vandeli Zacharda

Troglocheles vandeli Zacharda, 1987: 2052

Type locality: The Pyrenees, Haute-Garonne, Herran, Trou Mille

Distribution: France

Troglocheles vornatscheri (Willmann)

Rhagidia vornatscheri Willmann, 1953: 218

Troglocheles vornatscheri Zacharda, 1980: 653; 2000:

Type locality: Austria, Herdengelhöhle Cave

Distribution: Austria

Genus Tuberostoma Zacharda

Tuberostoma Zacharda, 1980: 763

Type species: Coccorhagidia gressitti Womersley et Strandtmann, 1963

Tuberostoma gressitti (Womersley et Strandtmann)

Coccorhagidia gressitti Womersley & Strandtmann, 1963: 467; Strandtmann, 1967: 66

Tuberostoma gressitti (Womersley et Strandtmann): Zacharda, 1980: 764

Type locality: Antarctica, W.S.W. from Hallet Station

Distribution: Antarctica

Tuberostoma leechi (Strandtmann)

Rhagidia leechi Strandtmann, 1963: 463; Strandtmann, 1970: 100

Tuberostoma leechi (Strandtmann): Zacharda, 1980: 768

Type locality: Livingstone L, South Shetland Is., 62°42‘S, by 60°26/W

Distribution: Antarctica, South Shetland, South Georgia

Tuberostoma keithi (Strandtmann)

Coccorhagidia keithi Strandtmann, 1967: 67

Tuberostoma keithi (Strandtmann) Zacharda, 1980: 766

Type locality: Antarctica, Nunatak, 3 mi. NE, Mt. Mirray, Mowson Ge. area

Distribution: Antarctica

Genus Zachardaia Özdikmen

Krantzia Zacharda, 1983: 993 (nomen praeoccupatum)

Zachardaia Özdikmen, 2008: 220

Type species: Krantzia quadriseta Zacharda, 1983

Zachardaia quadriseta (Zacharda)

Krantzia quadriseta Zacharda, 1983: 993

Type locality: Oregon, Jefferson Co., Camp Sherman

Distribution: USA

In the monograph of Zacharda (1980) are listed 48 species as «Species incertae sedis et inquirenda«. Here is this list (shortly):

1. Scyphius subterraneus Absolon, 1899 - The type unknown.

2. Rhagidia margaretae Baltac, 1974 – Type not seen by Zacharda.

3. Rhagidia strandtmanni Baltac, 1976 - Type not seen by Zacharda.

4. Rhagidia grandjeani Baltac, 1976 - Type not seen by Zacharda.

5. Rhagidia pallida Banks, 1894 – The type unknown. Published also by Miller, 1925.

6. Rhagidia cavicola Banks, 1897 - The type unknown.

7. Nörneria ? maxillata Berlese, 1893 - Type not seen by Zacharda.

8. Rhagidia subterranea Berlese, 1905 - The type unknown.

9. Rhagidia heteropoda Berlese, 1910 - The type unknown.

10. Megamerus fallax Dugès, 1834 - The type unknown.

11. Megamerus roseus Dugès, 1834 - The type unknown. France.

12. Rhagidia trisetata Elliot et Strandtmann, 1971 - Type not seen by Zacharda.

13. Rhagidia muscicola Hull, 1918 - The type unknown. England.

14. Scyphoides maxillatus Karpeles, 1891 - The type unknown.

15. Scyphius pyrrholeucus Koch, 1838 – The type unknown.

16. Scyphius cerinus Koch, 1838 – The type unknown.

17. Scyphius coarctatus Koch, 1838 – The type unknown.

18. Scyphius cylindricus Koch, 1838 – The type unknown.

19. Scyphius diaphanus Koch, 1838 – The type unknown.

20. Scyphius albellus Koch, 1838 – The type unknown.

21. Scyphius oblitteratus Koch, 1838 – The type unknown.

22. Scyphius elongatus Koch, 1838 – The type unknown.

23. Scyphius reflexus Koch, 1838 – The type unknown.

24. Scyphius hamatus Kramer et Neuman, 1883 - The type unknown.

25. Scyphius japonicus Kramer et Neuman, 1883 - The type unknown.

26. Nörneria cylindrica Kramer, 1898 (nec Scyphius cylindricus Koch, 1838) - The type unknown.

27. Gamasus halophilus Laboulbene, 1851 - The type unknown.

28. Rhagidia conciana Lombardini, 1951 - The type unknown.

29. Rhagidia comelicensus Lombardini, 1962 - The type unknown.

30. Rhagidia japonica Morikawa, 1963 - Type not seen by Zacharda.

31. Rhagidia mordax Oudemans, 1906 - Type not seen by Zacharda.

32. Rhagidia fragosa Schuster, 1958 – The holotype unknown.

33. Rhagidia raetica Schweizer et Bader, 1963 - Type not seen by Zacharda.

34. Rhagidia crassipes Schweizer et Bader, 1963 - Type not seen by Zacharda.

35. Coccorhagidia clavifrons Shiba, 1969 - Type not seen by Zacharda.

36. Coccorhagidia semiclavifrons Shiba, 1969 - Type not seen by Zacharda.

37. Rhagidia crenata Shiba, 1969 - Type not seen by Zacharda.

38. Scyphius maniacus Stoll, 1890 - The type unknown.

39. Rhagidia bidens Sig Thor, 1909 - The type unknown.

40. Rhagidia birulai Sig Thor, 1909 - The type unknown. Yan Mayen I. (Macfadyen, 1954)

41. Rhagidia megalochela Trägårdh, 1907 - The type unknown.

42. Rhagidia gigas var. longipes Trägårdh, 1907 - The type unknown.

43. Rhagidia mordax var. grandis Willmann, 1936 - The type unknown.

44. Rhagidia unidentata Willmann, 1939 - The type unknown.

45. Rhagidia dalmatina gigantea Willmann, 1941 - The type unknown.

46. Rhagidia paxi Willmann, 1942 - The holotype fragmentary, probably a nymph.

47. Rhagidia strouhali Willmann, 1953 - The type unknown.

48. Rhagidia pseudocrenata Shiba, 1976 Malaysia Pasoh Forest Reserve

Distribution of Rhagidiidae

EUROPE

Austria (Irk, 1939; Franz, 1943, 1954; Willmann, 1951a, 1951b, 1953; Janetschek, 1959, 1993; Mihelčič, 1957, 1958; Schmölzer, 1962, 1993; Zacharda, 1980, 1994, 2000d, Zacharda, Isaia & Piva, 2011; Zacharda & Kučera, 2006) – Coccorhagidia clavifrons, C. berlesei, C. divergens (inquir.), Evadorhagidia janetscheki, Foveacheles brevichelae, F. mira, F. osloensis, F. proxima, F. unguiculata, F. willmanni, Foveacheles (Proxistella) terricola, Foveacheles (Ternirhagidia) alpina, Poecilophysis (D.) pratensis, P. (D.) wankeli, Rhagidia gigas (= intermedia), R. longiseta, R. reflexa, Robustocheles mucronata, Shibaia longisensilla, Thoria uniseta, Traegaardhia dalmatina, Troglocheles aggerata, T. archetypica, T. christiani, T. strasseri, T. strasseri tirolensis, T. vornatscheri,

Azerbaidzhan (Zacharda, 1983b) - Poecilophysis (Dentocheles) pratensis

Belgium (Leruth, 1939; Cooreman, 1943) – Foveacheles terricola, Poecilophysis spelaea, Poecilophysis (Dentocheles) pratensis, Robustocheles (R.) robusta,

Bulgaria (Turk, 1970) – Poecilophysis spelaea, P. wolmsdorfensis; Inc. sedis: Rhagidia strouhali

Croatia (Willmann, 1939; Zacharda, 1980) – Traegaardhia dalmatina

Czech Rep. (Wankel, 1861; Willmann, 1954; Zacharda, 1980, 1993, 2000d; Zacharda et al., 2005) – Brevipalpia minima, Coccorhagidia clavifrons, Crassocheles muralis, C. virgo, Evadorhagidia bezdezensis, E. oblikensis, E. corcontica, Foveacheles osloensis, F. brevichelae, Foveacheles (Proxistella) terricola, F. (P.) simulate, Hammenia macrostella, Latoempodia macroempodiata, Poecilophysis arena, P. spelaea, P. pratensis, P. recussa, P. saxonica, P. wolmsdorfensis, P. (D.) wankeli, Rhagidia diversicolor, R. gelida, R. gigas, R. punkva, R. rackae, R. ruseki, [R. reflexa], Robustocheles mucronata, R. robusta, R. dentata, R. tricuspidata, Shibaia longisensilla, Thoria brevisensilla, Th. uniseta, [R. berlesei]

Denmark (Haarløv, 1957) – Coccorhagidia clavifrons, C. subterranea, Poecilophysis (Dentocheles) pratensis, Robustocheles mucronata, Rhagidia intermedia (?)

Greenland (Trägårdh, 1904) – Rhagidia gelida

Faeroe Is. (Trägårdh, 1931) – Poecilophysis faeroensis

Yan Mayen I. (Macfadyen A. 1954) – Coccorhagidia clavifrons, Rhagidia birulai, Rh. gelida, Rh. terricola

Finland (Zacharda & Kučera, 2010) – Rhagidia breviseta, Rh. diversicolor, Rh. gigas, Rh. longiseta, Rh. parvilobata, Poecilophysis pratensis, P. pseudoreflexa, P. spelaea, P. saxonica, Foveacheles brevichela, F. cegetensis, Robustocheles mucronata, R. obscuritata, Thoria uniseta, Evadorhagidia sp.

France (contin.) (Berlese & Trouessart, 1889; Cooreman, 1954, 1959; Zacharda, 1980; 1987, Zacharda, Grafitti & Piva, 2011) – Foveacheles canestrini, Foveacheles (Proxistella) terricola, Poecilophysis (Dentocheles) pratensis, P. spelaea, Rhagidia (Deharvengiella) troglomorphica, Rhagidia (Noerneria) diversicolor, Rh. longipes, Traegaardhia dalmatina, Troglocheles gineti, T. strasseri, T. vandeli, Rh. halophila

Corsica (Cooreman, 1954) - Robustocheles mucronata

Germany (C.L. Koch, 1835; Strandtmann & Prasse, 1977; Ehrnsberger, 1977a, 1977b; Willmann, 1934, 1955; Dobat, 1976; Zacharda, 1980) – Coccorhagidia clavifrons, Foveacheles osloensis, F. magna, F. harzensis, F. clavacrinita, Poecilophysis faeroensis, P. (Dentocheles) pratensis, Rhagidia Buettneri, Rh. gigas (= danica), R. diversicolor, R. clavicrinita, R. longipes, Robustocheles montana, R. mucronata, R. robusta, Shibaia longisensilla, Parallelorhagidia evansi, Troglocheles gineti

Great Britain (Hull, 1918; Evans, 1952; Turk & Turk, 1952; Turk, 1972) – Coccorhagidia clavifrons, C. subterranea, Poecilophysis spelaea, Rhagidia gigas, Robustocheles mucronata, Troglocheles odontochela, “Rhagidia vitzthumi”, “Rhagidia muscicola

Hungary (Dudich et al., 1940) - Coccorhagidia clavifrons

Iceland (Sellnick, 1940) – Poecilophysis faeroensis = reflexa, P. mucronata, P. pratensis, P. recussa, Rhagidia birulai, Rh. gelida, Rh. terricola, Thoria uniseta,

Surtsey (Lindroth, Andersson, Bödvarsson & Richter, 1973) - Rhagidiidae

Ireland (Halbert, 1915) – Rhagidia halophila, Rh. terricola

Italy (contin.) (Canestrini, 1886; Berlese, 1905, 1910; Bottazzi, 1950; Lombardini, 1962; Zacharda, 2010, 2011a, Zacharda, Grafitti & Piva, 2011; Zacharda, Isaia & Piva, 2011,) – Rhagidia comelicensis, Rh. gigas, Coccorhagidia clavifrons, C. subterranea, Foveacheles terricola, Traegaardhia cavernarum, T. cavernicola, T. distosolenidia, T. gracilis, T. nasuta, T. subterranea, Troglocheles lanai, T. quinquesolenidiata

Sardinia (Zacharda, Grafitti & Piva, 2011) -

Norway (Sig Thor, 1909, 1912, 1934; Hippa & Koponen, 1989; Hippa et al., 1989; Zacharda & Kučera, 2010) – Foveacheles (F.) osloensis, Foveacheles (Proxistella) terricola, Poecilophysis spelaea, Thoria uniseta

Beeren Eiland (Thorell, 1872) – Rhagidia gelida

Spitzbergen (Svalbard)(Thorell, 1971; Oudemans, 1926; Szeniczak et al., 2020) – Rhagidia gelida

Poland (Jesionowska, 2002) – Janes rafalskii, Rhagidia diversicolor, Robustocheles mucronata

Romania (Cooreman, 1951; Baltac, 1973, 1974, 1976, 1977b; Bannova, Sarmanova, Zhuravleva, 1977) – Coccorhagidia montana, Poecilophysis spelaea, P. wolmsdorfensis, Rhagidia longipes, Rh. orghidani, Rh. margaretae (sp.inquir.), Rh. strandtmanni (sp.inquir.), Rh. grandjeani (sp.inquir.), Shibaia heteropoda (sub «Rhagidia longisensilla)

Russia (all, continent.)(Thor, 1909; Mac Lean, Behan & Fjellbeg, 1978; Zacharda, 1983b) – [Rh. birulae Thor] Coccorhagidia clavifrons, C. pittardi, Evadorhagidia bezdezensis, Foveacheles (F.) brevichelae, F. (F.) caucasica, F. (F.) cegetensis, Poecilophysis (Procerocheles) pseudoreflexa, Poecilophysis (Procerocheles) recussa, P. (Sophrocheles) saxonica, P. (Dentocheles) pratensis, Rhagidia (Noerneria) punkva, Thoria brevisensilla, Th. uniseta

Crimea (Khaustov, 2015) – Paracrassocheles clavisetosus

Slovakia (Zacharda, 1980; 1988; Zacharda & Kučera, 2006; Kalúz, 2017) – Coccorhagidia clavifrons, C. pittardi, Foveacheles rupestris, Foveacheles (Spelaeocheles) troglodyta, Latoempodia similis, Poecilophysis pratensis, P. saxonica, Poecilophysis (D.) wankeli, Robustrocheles montana, R. mucronata, Rhagidia gigas, Shibaia tatrica

Slovenia (Willmann, 1932) – Troglocheles strasseri

Spain (cont.)(Mihelčič, 1958c; Barranco & Amate, 2000) - Foveacheles cannadasi, F. terricola, Traegaardhia dalmatina

Sweden (Hippa et al., 1989; Zacharda & Kučera, 2010) – Evadorhagidia sp., Foveacheles osloensis, Poecilophysis faeroensis, P. pseudoreflexa, P. spelaea, P. weyerensis, Rhagidia gigas, Rh. ruseki, Robustocheles robusta

Switzerland (Cooreman, 1954, 1959) – Foveacheles (Proxistella) terricola, Poecilophysis (Procerocheles) faeroensis (= Rh. reflexa), Poecilophysis spelaea, Rhagidia diversicolor, Rh. gelida, Robustocheles (R.) robusta,Troglocheles gineti, T. strasseri

Turkey (Çetin et al., 2021) – Traegaardhia distosolenidia

AFRICA

Egypt (Trägårdh, 1904b; Abou-Awad, 1985; Abou-Awad & El-Bagouri, 1986; Abou-Awad, El-Sawaf & Abdel-Khalek, 2009) - Coccorhagidia clavifrons, Foveacheles (F.) lemoni, Neothoria niloticus, Robustocheles (R.) deltacus, R. (R.) mucronata, Shibaia shereefi, Thoria sp., Neothoria zacharda, Robustocheles (R.) deltacus, Rhagidia (R.) qualiubeinsis

NigeriaRhagidiidae gen.sp.

South Africa (Meyer & Ryke, 1960) - Coccorhagidia clavifrons

ASIA

Cyprus (Zacharda, 2010) – Foveacheles thaleri

Iran (Zacharda et al., 2012; Darbemamieh, Hajiqanbar & Khanjani, 2013; Yazanpanah, Kazemi & Zacharda, 2014) – Brevipalpia minima, Coccorhagidia clavifrons; C. pittardi; Foveacheles brevichelae, F. cegetensis, F. osloensis, Poecilophysis arena, P. faeroensis, P. pratensis, Rhagidia cf. breviseta, Robustocheles hamedanensis, R. mucronata

Japan (Ehara, 1960; Morikawa, 1963; Shiba, 1969, 1971a, 1971b) – Poecilophysis pratensis (= Rhagidia japonica Morikawa), [Rhagidia crenata, Rh. uenoi], Shibaia heteropoda (= longisensilla)

Kirghizstan (Zacharda, 1983b) - Poecilophysis (Procerocheles) pseudoreflexa, P. (Dentocheles) pratensis, P. (Sophrocheles) saxonica, Robustocheles mucronata, Thoria brevisensilla

Korea (Shiba, 1969b; Zacharda, 1980) – Shibaia coreanensis,

Malaysia (Shiba, 1976) - Rhagidia whartoni = Poecilophysis pratensis, Robustocheles (R.) mucronata (sub Rhagidia neoshibai), Rhagidia strandtmanni, Rh. pseudocrenata (species inquirendae)

Nepal (Zacharda & Daniel, 1987) – Brevipalpia minima, Coccorhagidia pittardi, Poecilophysis saxonica, Rhagidia rackae, Shibaia longisensilla

Thailand (Zacharda, 1994c) – Robustocheles mucronata

Vietnam (Zacharda, 1994c) – Foveacheles (Achaetorhagidia) namcatiensis, F. (A.) tropica, Rhagidia (Dongnaia) edaphica, Robustocheles mucronata

NORTH AMERICA

Canada (Banks, 1919; Zacharda, 1980; 1985, 1986, 1995, 1997; Zacharda & Pugsley, 1988; Beaulieu et al., 2019) – Coccorhagidia clavifrons, Evadorhagidia quinqueseta, Foveacheles arenaria, F. brevichelae, F. tuktoaktukensis, Foveacheles (Mediostella) vancouverensis, Kouchibouguacia cryptosolenidiata, K. transcendata, Lindquistula multisolenidiata, Parallelorhagidia parva, Poecilophysis pseudoreflexa, Rhagidia gelida, Rhagidia (Noerneria) compressa, Rh. (N.) distisolenidiata, Rh. (N.) parvilobata, Rh. (N.) pasadenensis, Robustocheles (Lewia) lacustris, R. (L.) obscuritata, R. (R.) similis, Robustocheles (Mexia) occulta, R. pasquaria, Shibaia spiraesolenidiata

Mexico (Elliott & Strandtmann, 1971; Zacharda & Elliott, 1986; Zacharda, 1995, 1997) - Rhagidia (N.) quadrisolenidiata, ?Robustocheles infernalis, R. longichelae, ?R. trisetatus

USA (Banks, 1894, 1897, 1904; Call, 1897; Holsinger, 1965; Strandtmann, 1971; Elliott, 1976; Zacharda, 1980; 1983; 1985, 1986, 1995, 2010, 2011; Zacharda & Elliott, 1983; Lindquist & Zacharda, 1987; Zacharda et al., 2010) – Arhagidia monothrix, Coccorhagidia clavifrons, Elliotta howarthi, Foveacheles bentonensis, Foveacheles (Trofocheles) paralleloseta, Foveacheles (Propriorhagidia) tuktoyaktukensis, Zachardaia quadriseta, Poecilophysis (Dentocheles) affinis, P. (D.) extraneostella, P. (D.) pratensis, P. (D.) weyerensis, Poecilophysis (Procerocheles) oregonensis, P. melanoseta, P. saxonica, P. (Wankelia) wolmsdorfensis, Rhagidia cavicola, Rh. diversicolor, Rh. (Noerneria) arizonensis, Rh. (Noerneria) ponderosae, R. varia, Robustocheles (R.) pascuaria, Robustocheles (Amoveocheles) californiensis, Robustocheles (Lewia) hilli, R. mucronata, Traegaardhia holsingeri, T. similis

Alaska (Strandtmann, 1971; Zacharda, 1980, 1996; Zacharda & Elliott, 1985, Zacharda & Pugsley, 1988) – Arctorhagidia mariehammerae, A. sateri, Coccorhagidia pittardi, Eskimaia capitata, Foveacheles pribilovi, Kouchibouguacia transcendata, Robustocheles mucronata, R. hilli, R. occulta, Rhagidia gelida, Thoria uniseta [Rh. whartoni, Rh. shibai]

Hawaii (Strandtmann & Goff, 1978; Zacharda, 1980; 1982; Goff, 1987) – Coccorhagidia clavifrons, Elliotta howarthi, Foveacheles goffi, F. tenorioae, Parallelorhagidia hawaiiensis

SOUTH and CENTRAL AMERICA

Argentina (Bedano, 2004) - Coccorhagidia clavifrons

BrazilRobustocheles mucronata

Chile (Kramer, 1898) – Rhagidia krameri Sig Thor, 1941= Nörneria cylindrica Kramer, 1898

Cuba (Baltac, 1977a) – Coccorhagidia cubana (sp. inquir.)

Nicaragua (Maes et al., 1989) - Robustocheles (R.) mucronata

OCEANIA

Australia (continent)(Wood, 1970, Qin, 1998) – Rhagidia sp.

New Zealand (Qin, 1998) – Coccorhagidia clavifrons

Macquarie Island (Womersley & Strandtmann, 1963; Luxton, 1990) – Poecilophysis macquariensis

Campbell Island (Strandtmann, 1964) – Rhagidia mildredae

ANTARCTICA

Antarctida (Trouessart, 1903; Zacharda, 1980) – Rhagidia (Austrorhagidia) gerlachei

Possession I. (Strandtmann & Davies, 1972) – Poecilophysis kerguelenensis

South Georgia (Strandtmann, 1970) - Rhagidia (Austrorhagidia) gerlachei, Tuberostoma leechi

Patria incognita (Zacharda, 1980) – Foveacheles emendata

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  • Willmann C. 1935. Exploration biologique des Cavernes de la Belgique et du Limbourg hollandais. XXV. Contr.: Acari. - Bull. Mus. r. Hist. nat. Belg., 11: 1 -41.
  • Willmann C 1936a. Die Gattung Rhagidia (Rhagidiidae, Acari). - Zool. Anz., 116: 289-303.
  • Willmann C. 1936b. Neue Acari aus schlesischen Wiesenboden. - Zool. Anz., 113: 273 -290.
  • Willmann C. 1951a. Die hochalpine Milbenfauna der mittlern Hohen Tauern ins besondere des Grossglockner-Gebietes (Acari). – Bonn. Zool. Beitr., 2(1-2): 141 – 176.
  • Willmann C. 1951b. Untersuchungen über die terrestrische Milbenfauna im pannonischen Klimagebiet Österreichs. – Sitzungsber. der Österr. Akademie der Wissenschaften, Mathem. – naturwiss. Kl., Abt. I, 160 Bd.: 91 - 176.
  • Willmann C. 1952. Die Milbenfauna der Nordseeinsel Wangerooge. - Veröff. Inst. Meeresforsch. Bremerh., 1: 139-186.
  • Willmann C. 1953a. Tarsale Sinnesorgane bei der Gattung Rhagidia und anderen prostigmatischen Milben. - Zool. Anz., 150: 215-223.
  • Willmann C. 1953b. Neue Milben aus den östlichen Alpen. – Sitz.ber. Österr. Akad. d. Wiss., math.-nat. Kl., 162: 449 – 519.
  • Willmann C. 1954. Mährische Acari hauptsächlich aus dem Gebiete des mährischen Karstes. - Čsl. Parasit., 1: 213-272.
  • Willmann C. 1955. Milben aus dem südwestlichen Sachsen – Abhandlungen und Berichte aus dem Staatlichen Museum für Tierkunde in Dresden. 22: 207 - 225.
  • Wolf B. 1934 -1938. Animalium cavernarum catalogus. 3 vols. W. Junk, Gravenhage.
  • Womersley H., R. W. Strandtmann. 1963. On some free-living prostigmatic mites of Antarctica. - Pacif. Insects, 5: 451-472.
  • Yazanpanah Sh., Sh. Kazemi, M. Zacharda. 2014. A review of Iranian Rhagidiidae (Acari: Prostigmata) and new reports of one genus and three subgenera from Iran. - Iranian Journal of Plant Protection Science (Iranian Journal of Agricultural Sciences), fall 2013-winter 2014, 44(2): 203 - 210.
  • Zacharda M. 1978. Terrestrial Prostigmatic Mites from the Amateurs’ Cave, the Moravian kars [sic], Czechoslovakia. – Vĕstník Českoslov. Spol. Zoologické, 42(3): 215 – 240.
  • Zacharda M. 1979a. The evaluation of the morphological characters in Rhagidiidae. – In: J.G.Rodriguez (Ed.) Recent Advances in Acarology, Vol. II: 509 – 514.
  • Zaharda M. 1979b. The ecology of Rhagidiidae. – Recent Advances in Acarology, 1: 537 – 542.
  • Zacharda M. 1980. Soil mites of the family Rhagidiidae (Actinedida: Eupodoidea). Morphology, systematics, ecology. – Acta Univ. Carol. Biol., 1978(5 – 6): 489 – 785.
  • Zacharda M. 1982. The cavernicolous fauna of Hawaiian lava tubes. 13. A new subgenus and two new species of Rhagidiidae (Acari: Eupodoidea). – Pacific Insects, 24: 275 – 280.
  • Zacharda M. 1983a. The Rhagidiidae (Acarina: Prostigmata) from the Central Caucasus, Siberia, and some other parts of the USSR. – Vĕstnik Českoslov. Spol. Zool., 47: 304 – 319.
  • Zacharda M. 1983b. New Rhagidiidae (Acarina: Prostigmata) from Oregon. – Can. J. Zool., 61: 993 – 1001.
  • Zacharda M. 1984. Further remarks on the synonymy of the Rhagidiidae (Acarina: Prostigmata). – J. of Natural History, 18: 869 - 872.
  • Zacharda M. 1985. New Rhagidiidae (Acarina: Prostigmata) from caves of the U.S.A. - Vĕstnik Českoslov. Spol. Zool., 49: 67 – 80.
  • Zacharda M. 1986. New taxa of Rhagidiidae from North America. Part I. - Can. J. Zool., 64: 2531 – 2542.
  • Zacharda M. 1987. New taxa of Rhagidiidae (Acari: Prostigmata) from Pyrenean caves. – Can. J. Zool., 65: 2051 - 2056.
  • Zacharda M. 1988. Foveacheles (Spelaeocheles) troglodyta sp.n., subgen. n. (Acari: Prostigmata: Rhagidiidae) from Stratenská Cave, the Western Carpathians, Slovakia. - Vĕstnik Českoslov. Spol. Zool., 52: 75 – 78.
  • Zacharda M. 1993. Glacial relict Rhagidiidae (Acari: Prostigmata) from superficial underground enclosures in the Krkonoše Mountains, Czechoslovakia. – J. Nat. Hist., 27: 47 – 61.
  • Zacharda M. 1994a. Foveacheles unguiculata n.sp., a New Rhagidiid Mite from the Central Alps (Tyrol, Austria)(Acari: Prostigmata: Rhagidiidae).– Ber. Nat. - med. Verein Innsbruck, 81: 85 – 91.
  • Zacharda M. 1994b. A new troglobitic Rhagidiid Mite, Traegaardhia vicenzaensis n.sp. from Northern Italy (Acari: Prostigmata). – Ber. Nat.-med. Verein Innsbruck, 81: 93 – 98.
  • Zacharda M. 1994c. New taxa of Rhagidiidae (Acari Prostigmata) from tropical rain forest in Nam Cat Tien National Park, South Vietnam. – Tropical Zoology, 7: 35 – 56.
  • Zacharda M. 1995a. New taxa of Rhagidiidae from North America. Part II. The genera Kouchibouguacia Zacharda and Shibaia Zacharda. – Can. J. Zool., 73: 589 – 599.
  • Zacharda M. 1995b. New taxa of Rhagidiidae (Acari: Prostigmata) from North America. Part 3A. Genus Rhagidia Thorell: the gigas species-group - Canadian Journal of Zoology, 73: 1247-1258.
  • Zacharda M. 1995c. New taxa of Rhagidiidae (Acari: Prostigmata) from North America. Part IIIB. Genus Rhagidia Thorell: the quadrisolenidiata species-group – Canadian Journal of Zoology, 73: 1259-1267.
  • Zacharda M. 1996. New taxa of Rhagidiidae (Acari: Prostigmata) from North America. Part 4. Genus Foveacheles Zacharda, with a key to world species. – Canadian Journal of Zoology, 74: 343-356.
  • Zacharda M. 1997. New taxa of Rhagidiidae (Acari: Prostigmata) from North America. Part 5. Genus Robustocheles Zacharda, with a key to world species of the genus. - Journal of Natural History, 31: 1075 -1103.
  • Zacharda M. 2000a. A new species of the genus Evadorhagidia (Acari: Prostigmata: Rhagidiidae) from talus ecosystems, with a key to the world species of the genus. – Acarologia (Paris), 40: 373-383.
  • Zacharda M. 2000b. New species of the rhagidiid genus Foveacheles (Acari: Prostigmata: Eupodoidea) with a lyrifissure-like structure on the chelicerae. – Journal of Natural History, 34: 247-265.
  • Zacharda M. 2000c. New species of the genus Troglocheles (Acari: Prostigmata: Rhagidiidae) from Oetztal Alps, Tyrol, with a key to adult species of the genus. – Journal of Natural History, 34: 463-478.
  • Zacharda M. 2000d. New and little-known species of Rhagidiidae from talus ecosystems, in the Czech Republic and Austria. - Journal of Zoology, 251: 105 - 118.
  • Zacharda M. 2010. A review of the genus Traegaardhia (Acari, Prostigmata, Rhagidiidae) with descriptions of new species and a key to species – Zootaxa, 2474: 1-64.
  • Zacharda M., G. Grafitti, E. Piva. 2011. New taxa of Rhagidia and Foveacheles (Acari: Prostigmata: Rhagidiidae) from Italian and French caves, with keys to adults of subgenera Deharvengiella and Mediostella. - Journal of Natural History, 45: 667-683.
  • Zacharda M., M. Daniel. 1987. The first record of the family Rhagidiidae (Acari: Prostigmata) from the Himalayan Region. - Vĕstnik Českoslov. Spol. Zool., 51: 58 – 59.
  • Zacharda M., W.R. Elliot. 1981a. Holarctic cave mites of the family Rhagidiidae (Actinedida: Eupodoidea). – Proc. 8th Intern. Congress on Speleology, Bowling Green, KY, p. 604 – 607.
  • Zacharda M., W.R. Elliot. 1983. New species of the family Rhagidiidae (Acarina: Actinedida: Eupodoidea) from California caves. – Acta Univ. Carol. - Biologica, 1981: 463 – 475.
  • Zacharda, M., W.R. Elliott. 1986. Robustocheles infernalis sp. n. (Acarina: Actinedida: Rhagidiidae) from Cueva del Diablo, Veracruz, Mexico. - Acta Universitatis Carolinae Biologica, 1981: 477- 481.
  • Zacharda M., D. Fong, H. Horton, I. Hobbs, E. Piva, E. Slay, J. Taylor. 2010. A review of the genus Traegaardhia (Acari, Prostigmata, Rhagidiidae) with descriptions of new species and a key to species. - Zootaxa, 2474: 1–64.
  • Zacharda M., M. Gude, S. Krause, Ch. Hauck, R. Molenda, V. Ružička, 2005. The relict Mite Rhagidia gelida (Acari: Rhagidiidae) as a Biological Cryoindicator of Periglacial Microclimate in European Highland Screes. – Arctic, Antarctic, and Alpine Research, 37(3): 402 – 408.
  • Zacharda M., M. Isaia, E. Piva. 2011. New troglobitic species of the genus Troglocheles (Acari: Prostigmata: Rhagidiidae) from caves in northern Italy and Austria, with a key to adult species of the genus. - Journal of Natural History, 45(11-12): 641-666.
  • Zacharda M., T.Kučera. 2006. Diversity of Predatory Rhagidiid Mites (Acari: Rhagidiidae) Inhabiting Montane Stony Debris in the Ötztal Alps, North Tyrol, Austria. – Arctic, Antarctic and Alpine Research, 38(2): 292 – 300.
  • Zacharda M., T. Kučera. 2010. The Rhagidiidae (Acari: Prostigmata) in NW Lapland: Could their assemblages be climate warming monitors related to environmental and habitat patterns? - Pedobiologia, 54(1): 1-8.
  • Zacharda M., C.W. Pugsley. 1988. Robustocheles occulta sp.n., a new troglobitic mite (Acari: Prostigmata: Rhagidiidae) from North American caves. – Can. J. Zool., 66: 646 – 650.
  • Zacharda M., E. A. Ueckermann, E. Rostami, H. Abbasipour, M. Khanjani. 2012. A survey of Rhagidiidae (Acari: Prostigmata) in fruit orchards in Iran, with description of a new species of Robustocheles. - International Journal of Acarology, 38: 30-34.

The Mite Family Eriorhynchidae Qin et Halliday

Eriorhynchidae Qin & Halliday, 1997: 152

Small family of five Australian species, described by Womersley (1941) and Qin & Halliday (1997) and recently detached from the Penthalodidae.

Catalogue of fam. Eriorhynchidae

Family Eriorhynchidae Qin et Halliday

Eriorhynchidae Qin & Halliday, 1997: 152

Type genus: Eriorhynchus Qin et Halliday, 1997

Genus Eriorhynchus Qin et Halliday

Eriorhynchus Qin & Halliday, 1997: 153

Type species: Eriorhynchus womersleyi Qin et Halliday, 1997

Eriorhynchus australicus (Womersley)

Penthalodes australicus Womersley, 1941: 294

Eriorhynchus australicus (Womersley): Qin & Halliday, 1997: 103

Type locality: Victoria, Acheron Way

Distribution: Australia (Victoria)

Eriorhynchus hades Qin et Halliday

Eriorhynchus hades Qin & Halliday, 1997: 156

Type locality: Victoria, Rokewood Cemetery

Distribution: Australia (Victoria)

Eriorhynchus ramosus Qin et Halliday

Eriorhynchus ramosus Qin & Halliday, 1997: 157

Type locality: Tasmania, Big Sassy Creek

Distribution: Australia (Tasmania)

Eriorhynchus walteri Qin et Halliday

Eriorhynchus walteri Qin & Halliday, 1997: 161

Type locality: Miala Nat. Park, Brisbane

Distribution: Australia (Queensland)

Eriorhynchus womersleyi Qin et Halliday

Eriorhynchus womersleyi Qin & Halliday, 1997: 161

Type locality: New South Wales, 34.6 km south of Bombala

Distribution: Australia (N.S.Wales)

Distribution of Eriorhynchidae

OCEANIA

Australia (continent)(Womersley, 1941; Qin Ting-Kui & Halliday, 1997) – Eriorhynchus australicus, E. hades, E. waltheri, E. womersleyi

Tasmania (Qin Ting-Kui & Halliday, 1997) – Eriorhynchus ramosus

Bibliography of Eriorhynchidae

  • Qin Ting-Kui, R.B. Halliday. 1997. Eriorhynchidae, a new family of Prostigmata (Acarina), with a cladistic analysis of eupodoid species of Australia and New Zealand. – Systematic Entomology, 22(2): 151 – 171.
  • Womersley H. 1941. The red-legged earth-mite (Acarina, Penthaleidae) of Australia. – Transactions of the Royal Society of South Australia, 65: 292 – 294.

The Mite Family Pentapalpidae Olivier et Theron

Pentapalpidae Olivier & Theron, 2000: 386

Small family of one known genus and species from South Africa.

Composition of fam. Pentapalpidae

Fam. Pentapalpidae Olivier et Theron, 2000 - 1 gen., 1 sp.

Pentapalpus Olivier et Theron, 2000 - 1 sp.

Catalogue of Pentapalpidae

Family Pentapalpidae Olivier et Theron

Pentapalpidae Olivier & Theron, 2000: 386

Type genus: Pentapalpus Olivier et Theron, 2000

Genus Pentapalpus Olivier et Theron

Pentapalpus Olivier & Theron , 2000: 386

Type species: Pentapalpus unguempodius Olivier et Theron, 2000

Pentapalpus unguempodius Olivier et Theron

Pentapalpus unguempodius Olivier & Theron, 2000: 386

Type locality: Eastern Cape Province, soil, Summerstrand, Porth Elizabeth, 33.58S, 25.38E

Distribution: South Africa

Distribution of Pentapalpidae

AFRICA

South Africa (Olivier & Theron, 2000) - Pentapalpus unguempodius

Bibliography of Pentapalpidae

  • Olivier P.A.S., P.D. Theron. 2000. Pentapalpidae, a new family of eupodoid mites (Prostigmata: Eupodoidea) from South Africa. – Acarologia, 40(4), 1999: 385 – 392.

The Mite Family Penthalodidae Thor

Penthalodidae Sig Thor, 1933: 274

Small or medium-sized (330 – 1500 µm) strongly sclerotisezed mites living in soil or moss. Most (39) species are within the two “old” genera - Penthalodes Murray, 1877 and Stereotydeus Berlese, 1901. Further contributions are due to Baker, Trägårdh, Khaustov, Mihelčič, Strandtmann, Goff, Qin, Thor, Jesionowska, Shiba, Womersley, Olivier, Trouessart, starting with Dugès (1834).

Composition of fam. Penthalodidae

Family Pentalodidae Thor, 1933 - 5 gen., 44 sp.

Callipenthalodes Qin, 1998 – 1 sp.

Hawaiieupodes Strandtmann et Goff, 1978 – 1 sp.

Penthalodes Murray, 1877 - 10 sp.

Protopenthalodes Jesionowska, 1989 – 2 sp.

Stereotydeus Berlese, 1901 - 30 sp.

Catalogue of Penthalodidae

Family Penthalodidae Sig Thor

Penthalodidae Sig Thor, 1933: 272

Type genus: Megamerus Dugès, 1834 (nec McLeay, 1827), renamed Penthalodes Murray, 1877

Genus Callipenthalodes Qin

Callipenthalodes Qin, 1998: 221

Type species: Callipenthalodes pennyae Qin, 1998

Callipenthalodes pennyae Qin

Callipenthalodes pennyae Qin, 1998: 224

Type locality: New Zealand

Distribution: New Zealand

Genus Hawaiieupodes Strandtmann et Goff

Hawaiieupodes Strandtmann & Goff, 1978: 133

Type species: Hawaiieupodes thermophilus Strandtmann et Goff, 1978

Hawaiieupodes thermophilus Strandtmann et Goff

Hawaiieupodes thermophilus Strandtmann & Goff, 1978: 133; Jesionovska, 2010: 337

Type locality: Hawaii I., HVNP, Steaming Bluff Trail

Distribution: Hawaii

Genus Penthalodes Murray

Megamerus Dugès, 1834: 50 (nec McLeay, 1827)

Penthalodes Murray, 1877: 123

Type species: Megamerus ovalis Dugès, 1834

Penthalodes alaskaensis Jesionowska

Penthalodes alaskaensis Jesionowska, 2010: 31

Type locality: Alaska

Distribution: USA (Alaska)

Penthalodes boneti Baker

Penthalodes boneti Baker, 1946: 424

Type locality: Desierto de los Leones, Vera Cruz

Distribution: Mexico

Penthalodes carinatus Shiba

Penthalodes carinatus Shiba, 1978: 150

Type locality: Kuroki, Oozu, Ehime

Distribution: Japan

Penthalodes columbiae (Berlese)

Chromotydaeus columbiae Berlese, 1921: 193

Penthalodes columbiae : Thor & Willmann, 1941: 193; Strandtnann, 1979: 48

Type locality: “Columbia”

Distribution: North America

Penthalodes hawaiiensis Jesionowska

Penthalodes ovalis sensu Strandtmann & Goff, 1978: 121

Penthalodes hawaiiensis Jesionowska, 2010: 32

Type locality: Hawaii

Distribution: Hawaii

Penthalodes oregonensis Baker

Penthalodes oregonensis Baker, 1946: 424

Type locality: Corvallis

Distribution: USA (Oregon)

Penthalodes ovalis (Dugès)

Megamerus ovalis Dugès, 1834: 50; 1836: 518; Lucas, 1840: 472; 1850: 472

Trombidium ovale Gervais, 1844: 170

Penthaleus ovatus : C.L. Koch, fa. 18, t.14; Halbert, 1915: 111; Starkoff & Starkoff, 1950: 98

Penthalodes ovalis (Dugès): Oudemans, 1905: 241; 1937: 861; Irk, 1939; Sellnick, 1940: 32; Thor & Willmann, 1941: 60; Willmann, 1951a: 147; 1951b: 142; 1954: 235; Macfadyen, 1954: 283; Bottazzi, 1950: 38; Haarløv, 1957: 30; Mihelčič, 1958a: 104; 1958b: 270; Strandtmann, 1971: 92; Strandtmann & Goff, 1978: 135; Schmölzer, 1962: 283; 1993: 59; Kaluz, 2000: 477

Chromotydaeus ovalis : Berlese, 1903: 252

Chromotydaeus ovatus (Koch): Sig Thor, 1912: 237; Halbert, 1920: 139

Penthaleus arcticus Trägårdh, 1907: 18

Chromotydaeus arcticus (Trägårdh): Summerhayes & Elton, 1928: 249

Type locality: Europe

Distribution: Alaska, Austria, Canada, Czech Rep., Denmark, Hawaii, Iceland, Ireland (ovatus), Italy, Slovakia, Spain, Svalbard, Turkey, Yan Mayen I.

Penthalodes ovasimilis Schweizer

Penthalodes ovasimilis Schweizer, 1951: 62

Type locality: Swiss Alps, many loc. at 2200 – 2850 m

Distribution: Switzerland

Penthalodes polonicus Jesionowska

Penthalodes polonicus Jesionowska, 2010: 32; Hajizadeh & Noei, 2012; Darbemamieh, Hajiqanbar & Khanjani, 2013: 455

Type locality: Zakopane, The Tatra Mountains Nat. Park (Poland)

Distribution: Iran, Poland

Penthalodes tristiculus (Koch et Berendt) - fossil

Penthaleus tristiculus Koch & Berendt, 1854: 108

Penthalodes tristiculus (Koch et Berendt): Dunlop et al., 2013

Type locality: amber

Distribution: amber

Penthalodes turneri Baker

Penthalodes turneri Baker, 1946: 425

Type locality: Stoddard County, Mo.

Distribution: USA (Missouri, South Carolina, Texas)

Genus Protopenthalodes Jesionowska

Protopenthalodes Jesionowska, 1989: 57

Type species: Protopenthalodes Jesionowska, 1989

Protopenthalodes coniunctus Jesionowska

Protopenthalodes coniunctus Jesionowska, 1989: 59

Type locality: Poland

Distribution: Poland

Protopenthalodes reticulatus Khaustov

Protopenthalodes reticulatus Khaustov, 2015: 139

Type locality: Yalta mountain – forest Nature Reserve

Distribution: Crimea

Genus Stereotydeus Berlese, in Berlese & Leonardi

Stereotydeus Berlese, in Berlese & Leonardi, 1901: 14

Type species: Stereotydeus notophalloides Leonardi, in Berlese & Leonardi 1901

Tectopenthalodes Trägårdh, 1907: 11 (syn. by Womersley, 1941)

Type species: Penthaleus villosus Trouessart, 1902

Stereotydeus areolatus Womersley

Stereotydeus areolatus Womersley, 1935: 79; Thor & Willmann, 1941: 70; Strandtmann, 1979: 49; Qin, 1994: 1308; 1998: 299; Fredes, 2015: 33

Type locality: South Australia (Adelaide)

Distribution: Australia, Argentina

Stereotydeus australicus Thor

Stereotydeus australicus Thor, 1934: 65; Thor & Willmann, 1941: 69; Strandtmann, 1979: 49; Qin, 1994: 1310; 1998: 299

Type locality: Western Australia, Crawley

Distribution: Australia

Stereotydeus belli (Trouessart)

Penthaleus belli Trouessart, 1902: 225

Chromotydaeus belli : Thor & Willmann, 1941: 66

Stereotydeus belli (Trouessart): Womersley & Strandtmann, 1863: 458; Strandtmann, 1979: 49

Type locality: North Victoria Land

Distribution: Antarctica

Stereotydeus brevipalpus Qin

Stereotydeus brevipalpus Qin, 1994: 1313; 1998: 299

Type locality: South Australia

Distribution: Australia

Stereotydeus capensis Womersley

Stereotydeus capensis Womersley, 1935: 81

Type locality: Cape Province

Distribution: South Africa

Stereotydeus delicatus Strandtmann

Stereotydeus delicatus Strandtmann, 1967: 63

Type locality: Balleny Is: Sabrina I.

Distribution: Antarctica

Stereotydeus gamasoides Leonardi

Stereotydeus gamasoides Leonardi, 1901: 15

Type locality: Temuco

Distribution: Chile

Stereotydeus ineffabilis Brunetti et Siepel

Stereotydeus ineffabilis Brunetti & Siepel, 2021: 118

Type locality: Inexpressible Island

Distribution: Central Victoria Land, Continental Antarctica

Stereotydeus intermedius (Trouessart)

Penthalodes intermedius Trouessart, 1907: 35

Chromotydaeus intermedius (Trt.): Thor & Willmann, 1941: 66

Stereotydeus intermedius (Trouessart): Strandtmann, 1979: 50

Type locality: South Orkney Islands, Laurie Island

Distribution: Antarctica

Stereotydeus lattimorei Baker

Stereotydeus lattimorei Baker, 1946: 423

Type locality: Brownsville, Tx, from orchid roots from the Canal Zone

Distribution: Panama

Stereotydeus longipes Strandtmann

Stereotydeus longipes Strandtmann, 1970: 94

Type locality: South Georgia, Bird I., Wanderer Valley

Distribution: South Georgia (Subantarctic)

Stereotydeus mexicanus Baker

Stereotydeus mexicanus Baker, 1946: 422

Type locality: Patzcuaro, Michoacán, Mt. Popocatepetl (at 3000 m)

Distribution: Mexico

Stereotydeus meyeri Strandtmann

Stereotydeus meyeri Strandtmann, 1967: 63

Type locality: Enderby Land: Molodozhnaya

Distribution: Antarctica

Stereotydeus mollis Womersley et Strandtmann

Stereotydeus mollis Womersley & Strandtmann, 1963: 453; Wise, 1964: 522; Pittard, Robers & Strandtmann, 1971: 88

Type locality: Observation Hill, Ross I.

Distribution: Antarctica, southern Victoria Land from about 74° to 79° S lat., Balleny Islands

Stereotydeus notophalloides Leonardi

Stereotydeus notophalloides Leonardi, in Berlese & Leonardi, 1901: 15; Thor & Willmann, 1941: 69

Type locality: Temuco

Distribution: Chile

Stereotydeus nudisetatus Strandtmann

Stereotydeus nudisetatus Strandtmann, 1964: 152; Qin, 1998: 299

Type locality: Campbell Island

Distribution: New Zealand (Campbell Island)

Stereotydeus nunatakensis Brunetti

Stereotydeus nunatakensis Brunetti, 2021: 124

Type locality: Starr Nunatak

Distribution: South Victoria Land, Continental Antarctica

Stereotydeus occidentalis Womersley

Stereotydeus occidentale Womersley, 1935: 80; Strandtmann, 1964: 49; 1979: 49

Stereotydeus occidentalis Womersley: Thor & Willmann, 1941: 71; Qin, 1994: 1315; 1998: 299

Type locality: South Australia

Distribution: Australia

Stereotydeus oregonensis Baker

Stereotydeus oregonensis Baker, 1946: 424

Type locality: Oregon, Corvallis

Distribution: USA

Stereotydeus pallotus Olivier

Stereotydeus pallotus Olivier, 2006: 55

Type locality: Eastern Cape Province: soil, Hogsback near Alice

Distribution: South Africa

Stereotydeus pseudopulcher Qin

Stereotydeus pseudopulcher Qin, 1994: 1318; 1998: 299

Type locality: Tasmania

Distribution: Australia (Tasmania)

Stereotydeus pulcher Strandtmann

Stereotydeus pulcher Strandtmann, 1964: 154; Qin, 1998: 299

Type locality: Campbell Island

Distribution: New Zealand (Campbell Island)

Stereotydeus punctatus Strandtmann

Stereotydeus punctatus Strandtmann, 1967: 62

Type locality: North Victoria Land: W side of Islands Point

Distribution: Antarctica

Stereotydeus reticulatus Strandtmann

Stereotydeus reticulatus Strandtmann, 1970: 90

Type locality: South Georgia, Bird I., Macaroni Coulm, 200-230 m

Distribution: South Georgia

Stereotydeus rupicolamartimus Shiba

Stereotydeus rupicolamartimus Shiba, 1976: 97

Type locality: Malaysia

Distribution: Malaysia

Stereotydeus shoupi Strandtmann

Stereotydeus shoupi Strandtmann, 1967: 60

Type locality: Queen Maud Mts: Shackleton Glacier area: E side of Massam Glacier

Distribution: Antarctica

Stereotydeus tasmanicus Qin

Stereotydeus tasmanicus Qin, 1994: 1320; 1998: 299

Type locality: Tasmania

Distribution: Australia (Tasmania)

Stereotydeus undulatus Strandtmann

Stereotydeus undulatus Strandtmann, 1964: 150; Qin, 1998: 299

Type locality: Campbell Island

Distribution: New Zealand (Campbell Island)

Stereotydeus vaginatus Olivier

Stereotydeus vaginatus Olivier, 2006: 55

Type locality: KwaZulu-Natal: soil, mouth of Tugela River

Distribution: South Africa

Stereotydeus villosus (Trouessart)

Penthaleus villosus Trouessart, 1902: 227

Penthalodes villosus , Trouessart, 1912: 86

Tectopenthalodes villosus (Trouessart): Trägårdh, 1907: 18; Thor & Willmann, 1941: 71

Stereotydeus (Tectopenthalodes) villosus: Berlese, 1917: 8

Stereotydeus villosus : Womersley & Strandtmann, 1963: 457; Strandtmann, 1979: 50

Type locality: South Shetland Islands, South Orkney Islands, Antarctic Peninsula

Distribution: Antarctica

Distribution of Penthalodidae

EUROPE

Austria (Irk, 1939; Franz Beier, 1948; Franz, 1954; Willmann, 1951a; Mihelčić, 1958; Schmölzer, 1962, 1993) – Penthalodes ovalis

Bulgaria (Beron, 2011) - Penthalodes ovalis

Czech Rep. (Willmann, 1954) – Penthalodes ovalis

Denmark (Haarløv, 1957) - Penthalodes ovalis

Yan Mayen I. (Macfadyen, 1954) – Penthalodes ovalis

France (contin.) (Dugès, 1834; Kontschan et al., 2018) – Penthalodes ovalis, Inc. sedis: Penthalodes inflatus

Great Britain (Hull, 1918) – Penthalodes ovalis

Iceland (Sellnick, 1940) – Penthalodes ovalis

Ireland (Halbert, 1915) – Penthalodes ovalis

Italy (contin.) (Thor & Willmann, 1941; Bottazzi, 1950) – Penthalodes ovalis

Norway (Mehl, 1979) – Penthalodes ovalis

Svalbard (Spitzbergen)(Thor & Willmann, 1941) – Penthalodes ovalis

Poland (Jesionowska, 1989, 2010) – Penthalodes polonicus, Protopenthalodes coniunctus

Russia (all, continent.)

Crimea (Khaustov, 1915) – Protopenthalodes reticulatus

Slovakia (Kalúz, 2000) – Penthalodes ovalis

Spain (cont.)(Mihelčič, 1958b) - Penthalodes ovalis

Sweden (Strandtnann, 1979) – Penthalodes ovalis

Switzerland (Schweizer, 1951; Schweizer & Bader, 1963) - Penthalodes ovalis

Turkey (all)(Kalúz, 2000) - Penthalodes ovalis, P. ovasimilis

AFRICA

South Africa (Womersley, 1935; Olivier, 2006) – Stereotydeus pallotus, S. vaginatus

ASIA

Iran (Hajizadeh & Noei, 2012; Hajizadeh & Noei, 2012; Darbemamieh, Hajiqanbar & Khanjani, 2013) – Penthalodes polonicus

Japan (Shiba, 1978) – Penthalodes carinatus

Malaysia (Shiba, 1976) - Stereotydeus rupicolamartimus

NORTH AMERICA

Canada (Strandtmann, 1979; Behan, 1978; Beaulieu et al., 2019) – Penthalodes ovalis

Mexico (Baker, 1946) - Penthalodes boneti

USA (Baker, 1946) – Penthalodes oregonensis, P. turneri

Alaska (Strandtmann, 1971; Jesionowska, 2010) – Penthalodes alascaensis, P. ovalis

Hawaii (Strandtmann & Goff, 1978; Goff, 1983, 1987; Jesionowska, 2008, 2010) – Hawaiieupodes termophilus, Penthalodes hawaiiensis, P. ovalis

SOUTH and CENTRAL AMERICA

Argentina (Fredes, 2015) – Stereotydeus areolatus

Chile (contin.) (Leonardi, 1901, 1903) – Stereotydeus notophalloides, S. gamasoides

Panama (Baker, 1946) - Stereotydeus lattimorei

OCEANIA

Australia (continent)(Womersley, 1935; Qin, 1994) – Stereotydeus areolatus, S. australicus, S. brevipalpus, S. occidentalis

Tasmania (Qin, 1994) – Stereotydeus pseudopulcher, S. tasmanicus

New Zealand (Qin, 1998) - Callipenthalodes pennyae

Campbell Island (Strandtmann, 1964) – Stereotydeus nudisetatus, S. pulcher, S. undulatus

ANTARCTICA

Antarctida (Trouessart, 1902; Womersley & Strandtmann, 1963; Strandmann, 1967; Wise, 1964; Pittard, 1971; Pittard, Roberts & Strandtmann, 1971; Brunetti et al., 2021a, b) – Stereotydeus belli, S. delicates, S. meyeri, S. mollis, S. nunatakensis, S. punctatus, S. shoupi, S. villosus

South Georgia (Strandtmann, 1970) – Stereotydeus longipes, S. reticulatus

South Shetland IslandsStereotydeus villosus

South Orkney Islands (Trouessart, 1912) - Stereotydeus villosus

Inexpressible Island (Brunetti et al., 2021a) – Stereotydeus ineffabilis

Bibliography of Penthalodidae

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The Mite Family Penthaleidae Oudemans

Penthaleidae Oudemans, 1931: 252; Sig Thor, 1933: 2

Middle sized (800 – 1000 µm) mites, black with red legs, living on plants and sometimes pests on some agricultural crops.

Composition of fam. Penthaleidae

Family Penthaleidae - 7 gen., 27 sp.

Chromotydaeus Berlese, 1903 - 3 sp.

Halotydeus Berlese, 1891 - 8 sp.

Linopenthaleus Willmann, 1951 - 1 sp.

Linopenthaloides Strandtmann, 1981- 1 sp.

Penthaleus C.L. Koch, 1836 - 12 sp.

Propenthalodes Jesionowska, 1989 – 1 sp.

Turanopenthalodes Barilo, 1988 – 1 sp.

Catalogue of fam. Penthaleidae

Family Penthaleidae Oudemans

Penthaleidae Oudemans, 1931: 252

Type genus: Penthaleus C.L. Koch, 1836

Genus Chromotydaeus Berlese

Chromotydaeus Berlese, 1903: 252

Type species: Penthaleus ovalis (Dugès, 1834)

Chromotydaeus anauniensis (R. Canestrini)

Penthaleus anauniensis R. Canestrini, 1886: 720

Chromotydaeus anauniensis (R. Canestrini): Thor & Willmann, 1941: 65

Type locality: Trient, Dosso di Tavon

Distribution: Italy

Chromotydaeus egregius (Berlese)

Penthaleus egregius Berlese, 1891(1888 ?): 3; Starkoff & Starkoff, 1950: 98

Chromotydaeus egregius (Berlese): Berlese, 1903: 252

Type locality: Italy

Distribution: Italy

Chromotydaeus illustris (Berlese)

Penthaleus illustris Berlese, 1916: 294; Bernini et al., 1994-96: 34

Chromotydaeus illustris (Berlese): Thor & Willmann, 1941: 67

Type locality: Garden “Boboli”, Florence

Distribution: Italy

Chromotydaeus quartus Qin et Halliday

Chromotydaeus quartus Qin & Halliday, 1996: 1834

Type locality: Western Australia

Distribution: Australia

Chromotydaeus similis Mihelčič

Chromotydaeus similis Mihelčič, 1958a: 39

Type locality: Sierra Nevada

Distribution: Spain

Genus Halotydeus Berlese

NB. Linopodes globosus Canestrini, 1884 was listed under Chromotydeus by Thor & Willmann, 1941: 65, but under line was noted that it could belong also to the [still undescribed] genus Linopenthaleus.

Halotydeus Berlese, 1891: 9

Type species: Notophallus hydrodromus Berlese et Trouessart, 1889

Halotydeus anthropus Qin et Halliday

Halotydeus anthropus Qin & Halliday, 1996: 442

Type locality: South Africa

Distribution: South Africa

Halotydeus bakerae Qin et Halliday

Halotydeus egregius (Berlese, 1891) [syn. Baker 1996:373]

Halotydeus bakerae Qin & Halliday, 1996: 445

Type locality: South Australia

Distribution: Australia (South Australia)

Halotydeus castellus Qin et Halliday

Halotydeus castellus Qin & Halliday, 1996: 445

Type locality: New-South-Wales

Distribution: Australia (New-South-Wales)

Halotydeus destructor (R.W. Jack)

Penthaleus destructor R.W. Jack, 1908; Tucker, 1925: 21

Halotydeus destructor (Tucker): Womersley, 1933: 108; Thor & Willmann, 1941: 91; Macfadyen, 1954: 283; Meyer & Ryke, 1960: 489. Redescription, Qin & Halliday 1996: 447; Robinson & Hoffmann, 2001: 699; Maino, Binns & Umina, 2018: 216

Type locality: South Africa

Distribution: Australia, South Africa, Yan Mayen I. (? introduced)

Halotydeus hydrodromus (Berlese et Trouessart)

Notophallus hydrodromus Berlese & Trouessart, 1889: 141

Halotydeus hydrodromus (Berlese & Trouessart): Berlese, 1891: Fasc. 60, No10; Halbert, 1915: 111 (var. albolineatus), 1920: 139; Hull, 1918: 33; Thor & Willmann, 1941: 90

Type locality: France

Distribution: England, France, Ireland, Italy

Halotydeus mollis Luxton

Halotydeus mollis Luxton, 1990: 668

Type locality: Hong Kong, marine

Distribution: China (Hong Kong)

Halotydeus signiensis Strandtmann et Tilbrook

Halotydeus signiensis Strandtmann & Tilbrook, 1968: 54

Type locality: Robin Peak, Signy Island

Distribution: South Orkney Islands (Signy I.)

Halotydeus spectatus Qin et Halliday

Halotydeus spectatus Qin & Halliday, 1996: 449

Type locality: New-South-Wales

Distribution: Australia (New-South-Wales)

Genus Linopenthaleus Willmann

Genus Linopenthaleus Willmann, 1951a: 159

Type species: Linopenthaleus irki Willmann, 1951

Linopenthaleus irki Willmann

Linopenthaleus irki Willmann, 1951a: 160; Schweizer & Bader, 1963: 220

Type locality: Grossglockner area (Austria)

Distribution: Austria, Switzerland

Genus Linopenthaloides Strandtmann

Linopenthaloides Strandtmann, 1981: 392

Type species: Linopenthaloides novazealandicus Strandtmann, 1981

Linopenthaloides novazealandicus Strandtmann

Linopenthaloides novazealandicus Strandtmann, 1981: 392

Type locality: N.Z., South I.: W coast, Charleton, ex Metro Cave, Ananui catchement

Distribution: New Zealand

Genus Penthaleus C.L. Koch

Penthaleus C.L. Koch, 1836: fa.1, t.12 and 13

Type species: Penthaleus haematopus C.L. Koch, 1836 = P. maior (Dugès)

Notophallus Canestrini, 1886: 697

Penthaleus arcticus Trägårdh

Penthaleus arcticus Trägårdh, 1901: 19

Type locality: Bären Island

Distribution: Norway

Penthaleus bipustulatus (Hermann)

Trombidium bipustulatum Hermann, 1804: 40

Penthaleus bipustulatus (Hermann): C.L. Koch, 1838: fa.18, t.7; Macfadyen, 1954: 283

Penthaleus bipustulatum (Hermann): Oudemans, 1937: 852

Type locality: France

Distribution: France, Germany, Yan Mayen I.

Penthaleus crinitus Narayan

Penthaleus crinitus Narayan, 1957: 111

Type locality: Mexico

Distribution: Mexico

Penthaleus dorsalis (Banks)

Notophallus dorsalis Banks, 1902: 172; Considered synonym of Penthaleus major by Baker (1946: 421).

Penthaleus dorsalis : Bruce, 2021

Type locality: USA

Distribution: USA

Penthaleus erythrocephalus C.L. Koch

Penthaleus erythrocephalus C.L. Koch, 1838: fa.18, t.8; Mihelčič, 1958a: 40; 1958b: 269

Type locality: Germany

Distribution: Germany, Spain

Penthaleus falcatus Qin et Halliday

Penthaleus falcatus Qin & Halliday, 1996: 1837; Halliday, 1998: 77; Robinson & Hoffmann, 2001: 699

Type locality: New South Wales

Distribution: Australia

Penthaleus fallax (Dugès)

Megamerus fallax Dugès, 1834: 50, 53

Penthaleus fallax (Dugès): Oudemans, 1937: 854

Type locality: France

Distribution: France

Penthaleus guttatus C.L. Koch

Penthaleus guttatus C.L. Koch: fasc. 18, nr. 13; Oudemans, 1937: 855; Thor & Willmann, 1941: 83

Type locality: Regensburg

Distribution: Germany

Penthaleus longipilis (Canestrini)

Notophallus longipilis Canestrini, 1886: 706

Penthaleus longipilis (Canestrini): Shiba, 1969: 71

Type locality: Italy

Distribution: England, Italy, Japan

Penthaleus major (Dugès)

Tétranyque majeur Dugès, 1834: 53 (nom. nud.)

Rhyncholophus major : Murray, 1877: 126

Penthaleus haematopus C.L. Koch, 1836: fasc. 1, nr.12; Marchal, 1908: 13; Thor, 1912: 237; Hull, 1918: 33

Penthaleus erythropus C.L. Koch, 1838: fasc. 18, nr.6

Penthaleus insulanus Thorell, 1872: 702

Tetranychus borealis L. Koch, 1879: 129

Notophallus insulanus : Trägårdh, 1900: 16

Notophallus haematopus : R. Canestrini, 1886: 704

Notophallus bicolor Froggatt, 1921: 33

Penthaleus bicolor Womersley, 1933: 110

Penthaleus major (Dugès): Meyer & Ryke, 1960: 491; Balevski & Kontev, 1969; Rekk, 1976: 49; Zhang & Hong, 2010: 6

Penthaleus maior (Dugès): Sellnick, 1940: 32; Thor & Willmann, 1941: 80; Schmölzer, 1993: 59; Halliday, 1998: 77; Darbemamieh, Hajiqanbar & Khanjani, 2013: 455

Type locality: Europe

Distribution: Argentina, Australia, Austria, Brazil, Bulgaria, Canada, China, France, Georgia, Germany, Great Britain, Greece, Holland, Hungary (cf. P. major), Iran, Iceland, Italy, Mexico, New Zealand, Norway, Poland, Portugal, Russia, Slovakia, Slovenia, South Africa, Spain, USA

Penthaleus minor (Canestrini)

Notophallus minor Canestrini, 1886: 705; Berlese, 1892: fasc. 61, No5

Penthaleus minor (Canestrini): Oudemans, 1905: 241; Thor & Willmann, 1941: 85; Qin & Halliday, 1996; Halliday, 1998: 77; Zhang & Hong, 2010: 6; Darbemamieh, Hajiqanbar & Khanjani, 2013: 455

Type locality: Padua (Italy)

Distribution: Australia, China, Great Britain, Germany, Iran, Italy, Switzerland

Penthaleus saxatilis C.L. Koch

Penthaleus saxatilis C.L. Koch, 1836: fa.1, t.13; Thor & Willmann, 1941: 81

Type locality: Regensburg

Distribution: Germany

Penthaleus tectus Halloway

Penthaleus tectus Halloway, 2005b: 144

Type locality: East Australia

Distribution: Australia

Genus Turanopenthalodes Barilo

Turanopenthalodes Barilo, 1988: 619

Type species: Turanopenthalodes polytrichus Barilo, 1988

Turanopenthalodes polytrichus Barilo

Turanopenthalodes polytrichus Barilo, 1988: 619; Khaustov, 2016: 674

Type locality: Samarkand, settl. Agalyk

Distribution: Uzbekistan

As species with uncertain affinity are listed by Thor & Willmann, 1941 or described before or after this monograph: Penthaleus nemoralis (Müller) (Acarus n.), P. agilis Berlese, 1905, P. hermanni Sig Thor, Notophallus viridis Banks, 1917 (synonymized with Penthaleus major by Baker (1946: 421), Penthaleus guayavicola (Stoll)(Trombidium g.), P. trilineatus (Stoll)(Trombidium g.), P. albicollis (Stoll)(Trombidium a.). Here could be added also some species, described by C.L. Koch (1838): Penthaleus amictus, P. militaris, P. virellus, P. rhodomelas and the described by Ludvig Koch from Novaja Zemlja Penthaleus borealis, P. crassipes.

Distribution of Penthaleidae

EUROPA

Austria (Irk, 1942; Willmann, 1951a, 1951b; Mihelčič, 1958c; Schmölzer, 1993) – Linopenthaleus irki, Penthaleus bipustulatus, P. major

Bulgaria (Balevski & Kontev, 1969; Stoeva, 2006; Atanasov, Marinova & Atanasova, 2008; Maneva & Atanasova, 2016) - Penthaleus major

Denmark (Müller O.F. 1776) - ? Penthaleus nemoralis

Yan Mayen I. (Macfadyen, 1954) - Penthaleus bipustulatus

France (contin.)(Hermann, 1804; Dugès, 1834; Qin & Halliday, 1996) – Penthaleus bipustulatus, P. fallax, P. major

Georgia (Rekk, 1976) - Penthaleus major

Germany (C.L. Koch, 1835, 1836, 1838; Qin & Halliday, 1996) – Penthaleus bipustulatus, P. erythrocephalus, P. major, P. saxatilis

Great Britain (Hull, 1918; Wallace & Mahon, 1971) - Halotydeus hydrodromus, Penthaleus longipilis, P. major

Greece (Hadzinicolis, 1986) - Penthaleus major

Holland (Wallace & Mahon, 1971) - Penthaleus major

Hungary (Kontschán et al., 2018) - Penthaleus cf. major

Iceland (Sellnick, 1940; Gudleifsson et al., 2002; Hallas & Gudleifsson, 2005) – Penthaleus major

Ireland (Halbert, 1915, 1920) - Halotydeus hydrodromus, Penthaleus major

Italy (contin.)(Canestrini, 1886, Berlese, 1916; Bernini F., M. Castagnoli, R. Nannelli. 1994-1996. Checklist delle specie de la fauna italiana. Fascicolo 24 Arachnidi Acari. – Bologna, 132 pp. Wallace & Mahon, 1971) – Chromotydaeus anauniensis, Ch. illustris, Penthaleus longipilis, P. major, P. minor

Norway (Thor, 1912; Johansen & Haug, 2002) – Penthaleus major (= haematopus)

Poland (Kontschán et al., 2018) - Penthaleus major

Portugal (cont.)(Wallace & Mahon, 1971; Bernini et al., 1994–96) - Penthaleus major

Russia (all, continent.) (L. Koch, 1879; Stuxberg, 1886; Kornilov & Kositsyna, 1981) – [Penthaleus borealis, P. crassipes], Penthaleus major

Slovenia (Devetak et al., 2019) - Penthaleus major

Spain (cont.)(Mihelčič, 1958a, 1958b; Wallace & Mahon, 1971) – Chromotydaeus similis, Penthaleus major

Switzerland (Schweizer & Bader, 1963) – Linopenthaleus irki, Penthaleus longipilis, P. major, P. minor

AFRICA

South Africa (Jack, 1908; Tucker, 1925; Meyer & Ryke, 1960; Qin & Halliday, 1996; Halliday, 2005) – Halotydeus anthropus, H. destructor, Penthaleus major

ASIA

China (Narayan, 1962; Zhang & Hong, 2010: 6) - Penthaleus major, P. minor

Iran (Darbemamieh, Hajiqanbar & Khanjani, 2013) - Penthaleus major, P. minor

Japan (Shiba, 1969; Kanda & Hirai, 1990) – Penthaleus longipilis, P. major

Uzbekistan (Barilo, 1988; Khaustov, 2016) - Turanopenthalodes polytrichus

NORTH AMERICA

Canada (Behan, 1978; Qin & Halliday, 1996; Beaulieu, 2019) – Penthaleus major,

Mexico (Baker, 1946; Narayan, 1957) – Penthaleus major, Stereotydeus mexicanus

USA (Banks, 1902; Ewing, 1910; Brust, 2018, 2021a, b) – Penthaleus dorsalis, P. major

SOUTH and CENTRAL AMERICA

Argentina (Qin & Halliday, 1996) - Penthaleus major

Brazil (Valle da Silva Pereira et al., 2017) - Penthaleus major

Guatemala (Stoll, 1886 - 1893) - ? Penthaleus trilineatus, ? Penthaleus albicollis

OCEANIA

Australia (continent)(Newman, 1923, 1924, 1925a, 1925b, 1936; Swan, 1934; Womersley, 1941; Wallace, 1970; Wallace & Mahon, 1971; Halliday, 1991, 1998, 2005b; Qin & Halliday, 1995, 1996a, 1996b; Weeks, Fripp & Hoffmann, 1995; Qin, 1998; Weeks & Hoffmann, 1999; James, 1987, 2000; Robinson & Hoffmann, 2001; Umina & Hoffmann, 2003, 2004; Umina, Hoffmann & Weeks, 2004) – Chromotydaeus quartus, Halotydeus bakerae, H. castellus, H. destructor, H. spectatus, Penthaleus major, P. minor, P. tectus

New Zealand (Strandtmann, 1981; Wallace & Mahon, 1971) – Linopenthaloides novazealandicus, Penthaleus major

ANTARCTICA

South Orkney Islands (Strandtmann & Tilbrook, 1968) – Halotydeus signiensis

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  • Willmann C. 1951b. Untersuchungen über die terrestrische Milbenfauna im pannonischen Klimagebiet Österreichs. – Sitzungsber. der Österr. Akademie der Wissenschaften, Mathem. – naturwiss. Kl., Abt. I, 160 Bd.: 91 - 176.
  • Womersley H. 1941. The red-legged earth-mite (Acarina, Penthaleidae) of Australia. - Transactions of the Royal Society of South Australia, 65(2): 292–294.
  • Xuan Cheng, A. A Hoffmann, J. L Maino, P. A. Umina. 2018. A cryptic diapause strategy in Halotydeus destructor (Tucker) (Trombidiformes: Penthaleidae) induced by multiple cues. - Pest Management Science, https://doi.org/10.1002/ps.5053
  • Xuan Cheng, P. Umina, A. Hoffmann. 2018. Influence of previous host plants on the reproductive success of a polyphagous mite pest, Halotydeus destructor (Trombidiformes: Penthaleidae). - Journal of Economic Entomology, 111(2): 680 – 688.
  • Zhang Zhi-Qiang. 2010. Terrestrial Parasitengona (except chiggers) of China: a review of progress in systematics and biology, with a checklist of species. – Zoosymposia, 4: 94 - 105.
  • Zhang Zhi-Qiang, Xiao-Yue Hong. 2010. Xin Centenary and Progress in Chinese Acarology: an Introduction. – Zoosymposia 4: 5 – 10.

The Mite Family Proterorhagiidae Lindquist et Palacios-Vargas

Proterorhagiidae Lindquist & Palacios-Vargas, 1991: 341

Family containing only one species from Mexico.

Catalogue of Proterorhagiidae

Family Proterorhagiidae Lindquist et Palacios-Vargas

Proterorhagiidae Lindquist & Palacios-Vargas, 1991: 341

Type genus: Proterorhagia Lindquist et Palacios-Vargas, 1991

Genus Proterorhagia Lindquist et Palacios-Vargas

Proterorhagia Lindquist & Palacios-Vargas, 1991: 341

Type species: Proterorhagia oztotloica Lindquist et Palacios-Vargas, 1991

Proterorhagia oztotloica Lindquist et Palacios-Vargas

Proterorhagia oztotloica Lindquist & Palacios-Vargas, 1991: 344

Type locality: border area between the states of Colima and Jalisco, Pozo La Escondida near Lagunitas Village, 2200 m

Distribution: Mexico

Distribution of Proterorhagiidae

NORTH AMERICA

Mexico (Lindquist & Palacios-Vargas, 1991) – Proterorhagia oztotloica

Bibliography of Proterorhagiidae

  • Lindquist E.E., J.G. Palacios-Vargas. 1991. Proterorhagiidae (Acari, Endeostigmata), a new family of Rhagidiid-like mites from Mexico. – Acarologia, 32(4): 341-363.

The Mite Family Strandtmanniidae Zacharda

Strandtmanniidae Zacharda, 1979: 77

Monotypical family known from the Chech Republic.

Catalogue of Strandtmanniidae

Family Strandtmanniidae Zacharda

Strandtmanniidae Zacharda, 1979: 77

Type genus: Strandtmannia Zacharda, 1979

Genus Pilorhagidia Strandtmann et Goff, 1978

Pilorhagidia Strandtmann & Goff, 1978: 141

Strandtmannia Zacharda, 1979: 78

Type species: Pilorhagidia hirsuta Strandtmann et Goff, 1978

Pilorhagidia celtarum (Zacharda)

Strandtmannia celtarum Zacharda, 1979: 78

Pilorhagidia celtarum (Zacharda):

Type locality: Bohemia centr., Zavist-knoll near Zbraslav

Distribution: Czech Rep.

Pilorhagidia hirsuta Strandtmann et Goff

Pilorhagidia hirsuta Strandtmann & Goff, 1978: 141

Type locality: Hawaii I: HVNP, Mauna Loa Trail, 2040 rn

Distribution: Hawaii Islands

Distribution of Strandtmanniidae

EUROPE

Czech Rep. (Zacharda, 1979) - Pilorhagidia celtarum

OCEANIA

Hawaii Islands (Strandtmann & Goff, 1978) – Pilorhagidia hirsuta

Bibliography of Strandtmanniidae

  • Strandtmann R.W., M.L.Goff. 1978. The Eupodoidea of Hawaii (Acarina: Prostigmata). - Pacific insects, 19(3/4): 121-143.
  • Zacharda M. 1979. Strandtmanniidae, a new family of Eupodoidea Acarina Prostigmata. - Vestnik Ceskoslovenske společnosti zoologicke, 43(1): 76-81.

Mite Superfamily Tydeoidea

Tydeoidea Cunliffe, 1955

The Superfamily contains the families Ereynetidae, Iolinidae, Triophtydeidae and Tydaeidae.

The Mite Family Ereynetidae Oudemans

The family Ereynetidae contains three subfamilies: Ereynetinae (free living in the soil), Speleognathinae (parasites in the respiratory passages of molluscs, birds and mammals) and Lawrencarinae (parasites in the respiratory passages of frogs and toads). Until 1936 only the Ereynetinae were known, described by several early acarologists, starting with Schrank (1776) and C.L. Koch (1836). In 1883 A. Berlese described the genus Ereynetes and in 1931 Oudemans created the family Ereynetidae. Some other specialists who contributed to the study of Ereynetinae are Vitzthum (1931), André & Lamy (1930, 1931), Thor (1932a, 1932b, 1936), E.W. Baker (1945), Turk & Phillips (1945), R.A. Baker (1970a, b), Cooreman (1947), Willmann (1936, 1949, 1953), Mihelčic (1958), Hunter (1964), Corpuz-Raros (1975), Shiba (1976), Zabludovskaya (1994) and again Berlese (1910, 1923). Many publications were devoted to the peculiar mites parasitizing the snails and slugs (Riccardoella). The modern arrangement of this interesting group is due to A. Fain and his co-authors J. L. van Goethem, A.M. Camerik and G.M. Barker.

The mighty presence of A. Fain in the study of Ereynetidae began in 1955. Firstly he submitted part of his collection from Rwanda to J. Cooreman, who described the first African member of the family as new genus and species Speleognathopsis galli Cooreman, 1955. This was the end of the pre-Fain era in the study of Ereynetidae. Only from the present day species have been known by this time from the parasitic subfamilies Speleognathinae and Lawrencarinae, and they were wrongly positioned in the system. In 1936 H. Womersley described the peculiar mite Speleognathus australis, for which he created the new family (now subfamily) Speleognathidae. Followed E. Boyd with “Speleognathus” sturni Boyd, 1948, the first among the many nasal parasites of birds and mammals we know now. Womersley (1953) named this mite Boydaia sturni. Meanwhile Lawrence (1952) described the first mite endoparasite of Anura, but named it Riccardoella eweri. Followed several other parasites of frogs and toads, now 12 species in the genera Lawrencarus Fain, 1957, Batracarus Fain, 1961 and Xenopacarus Fain, Baker & Tinsley, 1969. Fain (1962) summarized the data on the mites parasitizing Anura. Kethley (1971) described one new genus (Hydranetes) with two new species of Ereynetid mites, living under elytrae of Hydrophilid beetles in Florida.

The rich collection of endoparasitic mites, brought by Fain from Rwanda and Congo, was the basis for the many descriptions of new taxa in the subfamily Speleognathinae. The mites of this subfamily in the Afrotropical Region have been reviewed by Skoracki et al. (2018).

Very few people interfered in the stream of papers by Fain and co-authors (Aitken, Bafort, Tinsley, Baker, Barker, Cameric, Goff, Herin, Puylaert, Hyland, Klompen, Lukoschus, Mortelmans, Nadchatram, Prasse, van Goethem, Vercammen-Grandjean, Wagner, Flechtmann, Leal). Dubinin (1957) was the author of the new genus Ophthalmognathus. Other specialists describing Speleognathinae were Crossley (1952), Womersley (1936, 1953), Van Eyndhoven (1955), Ford (1959, 1962), Clark (1958, 1960, 1964, 1967a, 1967b), Domrow (1965, 1969, 1991), Pence (1973), Pence & Casto (1976), OConnor (1978).

The family Ereynetidae contains now (end 2021) 180 species of 35 genera and three subfamilies. The progress since the first general paper on the family (Thor, 1933) is obvious: by this time only 15 species of 3 (actually two – Ereynetes and Riccardoella) genera have been recognized in Ereynetidae. Most species are endoparasites.

Fam. Ereynetidae35 genera, 180 sp.

Subfam. Ereynetinae78 sp.

Tribe Austrereynetini

Austrereynetes Fain et Barker, 2003 – 1 sp.

Tribe Ereynetini

Ereynetes Berlese, 1883 – 55 sp.

Anereynetes Fain, 1964 - 24

Ereynetes Berlese, 1883 - 21

Gymnereynetes Fain, 1964 - 4

Huntereynetes Fain, 1964 - 3

Opsereynetes Thor, 1932 - 3

Hanriccardoella André, Ducarme et Lebrun, 2004 - 1

Hydranetes Kethley, 1971 - 2

Protereunetes Berlese, 1923 - 9

Pseudotydeus Baker et Delfinado, 1974 - 2

Riccardoella Berlese, 1923 - 8

Proriccardoella Fain et van Goethem, 1986 - 6

Riccardoella Berlese, 1923 – 2

Subfam. Speleognathinae - 90 sp.

Tribe Boydaiaini

Boydaia Womersley, 1953 - 41

Coboydaia Fain, 1971 - 5

Coboydaia (Apodiboydaia) Fain, 1985 - 1

Coboydaia (Coboydaia) Fain, 1971 - 4

Tribe Trispeleognathini

Astrida Fain, 1955 - 4

Astrida Fain, 1955 - 2

Cerylonyssus Fain et Aitken, 1970 – 1

Aureliania Fain, 1958 – 1

Meropiboydaia Fain, 1985 - 1

Metaboydaia Fain, 1962 - 1

Neastrida Fain, 1962 - 1

Neoboydaia Fain, 1958 - 3

Ophthalmognathus Dubinin, 1957 - 2

Phoenicopteriella Fain, 1970 - 2

Picinyssus Fain, 1969 - 1

Psittaboydaia Fain, 1985 - 2

Pterniboydaia Fain, 1985 - 1

Trispeleognathus Fain, 1958 -

Tribe Paraspeleognathopsini

Paraspeleognathopsis Fain, 1958 - 4

Tribe Speleochirini

Hipposideroptes Fain, 1970 - 1

Neospeleognathopsis Fain, 1958 – 3

Neospeleognathopsis (Neospeleognathopsis) Fain, 1958 – 1

Neospeleognathopsis (Speleomyotis) - 1

Neospeleognathopsis (Pteropignathus) - 1

Ralliboydaia Fain, 1962 – 3

Speleochir Fain, 1966 - 5

Neospeleochir Fain, 19 - 1

Speleognathopsis Cooreman, 1954 - 6

Speleomys Fain, 1970 – 1

Speleorodens Fain, 1962 - 6

Tribe Speleognathini

Domrownetes OConnor, 1978 - 1

Speleognathus Womersley, 1936 – 4

Subfamily Lawrencarinae Fain, 1957 - 12 sp.

Tribe Lawrencarini Fain

Batracarus Fain, 1961 - 1

Lawrencarus Fain, 1957 - 8

Tribe Xenopacarini Fain, Baker et Tinsley, 1969

Xenopacarus Fain, Baker et Tinsley, 1969 - 3

Catalogue of Ereynetidae

Family Ereynetidae Oudemans

Ereynetidae Oudemans, 1931: 253

Type genus: Ereynetes Berlese, 1883

Speleognathidae Womersley, 1936 (syn. by Fain, 1957: 250)

Type genus: Speleognathus Fain, 1936: 313

Microereunetidae Bottazzi, 1950: 38; syn. by Fain, 1954a: 110

Type genus: Microereunetes Berlese, 1924: syn. by Fain, 1954a: 110

Subfamily Ereynetinae Fain

Ereynetinae Fain, 1957: 250

Type genus: Ereynetes Berlese, 1883

Pseudotydeinae Baker & Delfinado, 1974: 444 – syn. by André & Ducarme, 2003: 373

Type genus: Pseudotydeus Baker et Delfinado, 1974

Tribe Austrereynetini Fain et Barker

Austrereynetini Fain & Barker, 2003: 233

Type genus: Austrereynetes Fain et Barker, 2003

Genus Austrereynetes Fain et Barker

Austrereynetes Fain & Barker, 2003: 234

Type species: Austrereynetes maudensis Fain et Barker, 2003

Austrereynetes maudensis Fain et Barker

Austrereynetes maudensis Fain & Barker, 2003: 235

Type locality: Maud Island, Marlborough Sounds, New Zealand

Distribution: New Zealand

Host: Pseudaneitea schauinslandi (Gastropoda: Pulmonata: Athoracophoridae)

Tribe Ereynetini Fain

Type genus: Ereynetes Berlese, 1883

Genus Ereynetes Berlese

Ereynetes Berlese, 1883: 5.1

Ereunetes Berlese, 1883: fasc.V, 1, tav.22 (non Ereunetes Illiger, 1911: Aves)

Microereunetes Berlese, 1921: 24; 1923: 243

Type species: Tydeus polymitus C.L. Koch, 1836 [Ereynetes galeatus Berlese, 1883], see Fain, 1985b: 257

Neoereunetes Bottazzi, 1950: 44 (n.nov.), type sp. polymytis. Syn. by Fain, 1964a: 111

Ereynetoides Fain et Nadchatram, 1962a: 69: type sp. Ereynetoides malayi Fain et Nadchatram, 1962a: 72. Syn. by Fain, 1964a: 110

Subgenus Ereynetes Berlese

Ereynetes Berlese, 1883: 5.1; Fain, 1985b: 257

Type species: Tydeus polymitus C.L. Koch, 1836 [Ereynetes galeatus Berlese, 1883]

Ereynetes (Ereynetes) agilis (Berlese)(sp. inquirenda)

Micrereunetes (Protereunetes) agilis Berlese, 1923: 245; Fain, 1964a: 99

Ereynetes (Ereynetes) agilis Berlese: Fain, 1964a: 106; Fain & Camerik, 1994: 150

Protereunetes agilis (Berlese): Thor & Willmann, 1941: 54; Strandtmann, 1971: 82

Type locality: Palermo

Distribution: Sicily

Host: moss

Ereynetes (Ereynetes) amplectorus (Hunter)

Ereynetoides amplectorus Hunter, 1964: 86; Siahkamari et al., 2020:

Ereynetes (Ereynetes) amplectorus Hunter: Fain, 1964a: 110; Fain & Camerik, 1994: 156

Type locality: American Fork, Utah

Distribution: Mexico, USA (Utah), South Africa, Iran

Host: on celery

Ereynetes (Ereynetes) arcuatus (Berlese)

Micrereunetes arcuatus Berlese, 1923: 244

Ereynetes arcuatus (Berlese): Thor, 1933: 72

Ereynetes (Ereynetes) arcuatus Berlese: Fain, 1964a: 106

Type locality: Portici

Distribution: Italy

Ereynetes (Ereynetes) boharti Hunter et Cross

Ereynetes (Ereynetes) boharti Hunter & Cross, 1968: 156; Fain & Camerik, 1994: 151

Type locality: Shoshoni, Wyoming

Distribution: USA (Wyoming)

Host: from artificial nest of bee, Nomia melanderi (Hymenoptera, Halictidae)

Ereynetes (Ereynetes) bolitophilus Fain et Camerik

Ereynetes (Ereynetes) bolitophilus Fain & Camerik, 1994: 156; Abdelgayed et al., 2019: 32

Type locality: Sandton (South Africa)

Distribution: South Africa, Egypt

Host: cow dung

Ereynetes (Ereynetes) cavernarum Shiba

Ereynetes cavernarum Shiba, 1971: 226

Type locality: Kaminari-ana Pit, Kansu-yama, Fujiwara, Susono-shi, Shizuoka Pref.

Distribution: Japan

Ereynetes (Ereynetes) centralis (Berlese)

Micrereunetes galeatus var. centralis Berlese, 1923: 243

Ereynetes centralis (Berlese): Thor, 1933: 74

Ereynetes (Ereynetes) galeatus centralis (Berlese): Fain, 1964a: 106

Neoereunetes centralis (Berlese): Bottazzi, 1950: 40

Type locality: Italy (“in agro Florentino”)

Distribution: Italy

Host: manure, humus, chestnut wood

Ereynetes (Ereynetes) excellens (Berlese)

Micrereunetes excellens Berlese, 1923: 243

Ereynetes excellens Berlese: Thor, 1933: 73

Ereynetes (Ereynetes) excellens Berlese: Fain, 1964a: 106

Type locality: Japan

Distribution: Japan

Ereynetes (Ereynetes) faini (Hunter)

Ereynetoides faini Hunter, 1964: 188

Ereynetes (Ereynetes) faini Hunter: Fain, 1985b: 257; Fain & Camerik, 1994: 151

Type locality: Boston

Distribution: USA (on beet root from Denmark and lily bulbs from England), Mexico

Ereynetes (Ereynetes) galeatus (Berlese)

Micrereunetes galeatus Berlese, 1923: 243

Acarus limacum , Berlese, 1883 (non Schrank, 1776)

Ereynetes polymitus , R. Canestrini, 1886 (non Koch, 1838)

Ereynetes ministralis , Oudemans, 1928: 382 (non Koch, 1838)

Micrereunetes polymitus , Berlese, 1920 (non Koch, 1838)

Ereynetes galeatus Berlese: Thor, 1933: 74

Ereynetes (Ereynetes) berlesei Oudemans, 1928: 382

Ereynetes (Ereynetes) galeatus (Berlese): Fain, 1964a: 106; Zabludovskaya, 1994: 35

Type locality: Palermo

Distribution: Sicily, Crimea

Host: in soil

Ereynetes (Ereynetes) gandensis Fain et Bafort

Ereynetes (Ereynetes) gandensis Fain & Bafort, 1973: 409; Fain, 1985b: 257; Fain & Camerik, 1994: 151; Zabludovskaya, 1994: 35

Type locality: Gand (Belgium)

Distribution: Belgium, Ukraine

Host: in humus

Ereynetes (Ereynetes) husatoi Shiba

Ereynetes (Ereynetes) husatoi Shiba, 1976: 139

Type locality: Malaya, Pasoh Forest Reserve

Distribution: Malaysia

Ereynetes (Ereynetes) luriei Fain et Camerik

Ereynetes (Ereynetes) luriei Fain & Camerik, 1994: 157

Type locality: Sandton

Distribution: South Africa

Host: cow dung

Ereynetes (Ereynetes) malayi (Fain et Nadchatram)

Ereynetoides malayi Fain & Nadchatram, 1962a: 72

Ereynetes (Ereynetes) malayi Fain & Nadchatram: Fain, 1964a: 106; 1985b: 257; Fain & Camerik, 1994: 150

Type locality: Fort Betis, Ulu Kelantan, Malaya

Distribution: Malaysia

Host: nasal cavity of Arachnothera l. longirostra (Passeriformes: Nectariniidae)

Ereynetes (Ereynetes) pegazzanoae Fain

Ereynetes (Ereynetes) pegazzanoae Fain, 1964a: 95; Zabludovskaya, 1994: 35

Type locality: Tiarno (Italy)

Distribution: Italy, Ukraine

Host: in dung

Ereynetes (Ereynetes) philippinensis Corpuz-Raros

Ereynetes (Ereynetes) philippinensis Corpuz-Raros, 1975: 326; Fain & Camerik, 1994: 151

Type locality: College, Laguna

Distribution: Philippines

Host: Scarabaeid beetle (Coleoptera)

Ereynetes (Ereynetes) punctatus Zabludovskaya

Ereynetes (Ereynetes) punctatus Zabludovskaya, 1994: 37

Type locality: Migya Village, Pervomayskiy Rayon, Nikolaevskaya oblast

Distribution: Ukraine

Host: in dung

Ereynetes (Ereynetes) septentrionalis (Berlese)

Micrereunetes galeatus var. septentrionalis Berlese, 1923: 244

Ereynetes septentrionalis (Berlese): Thor, 1933: 73; Çobanoglu et al., 2020:

Ereynetes (Ereynetes) galeatus septentrionalis (Berlese): Fain, 1964a: 106

Ereynetes (Ereynetes) septentrionalis (Berlese): Fain & Camerik, 1994: 150

Type locality: Nord Italy

Distribution: Italy, Turkey

Host: manure

Ereynetes (Ereynetes) tadakii Shiba

Ereynetes (Ereynetes) tadakii Shiba, 1976: 136

Type locality: Malacca

Distribution: Malaysia

Ereynetes (Ereynetes) tshernogoriensis Zabludovskaya

Ereynetes (Ereynetes) tshernogoriensis Zabludovskaya, 1994: 36

Type locality: Karpatskiy Zapovednik [Reserve], Tshernogorskiy Rayon, Transkarpathia; paratypes from Karadagskiy Zapovednik, Crimea

Distribution: Crimea, Ukraine

Ereynetes (Ereynetes) ueckermanni Fain et Camerik

Ereynetes (Ereynetes) ueckermanni Fain & Camerik, 1994: 154

Type locality: Cluny Farm, Transvaal

Distribution: South Africa

Host: cow and horse dung

Ereynetes (Ereynetes) watsoni Fain

Ereynetoides watsoni Fain, 1962a: 925

Ereynetes (Ereynetes) watsoni (Fain): Fain, 1964a: 106; 1985b: 257; Fain & Camerik, 1994: 150

Type locality: Macquarie Id., Antarctica

Distribution: Macquarie I., Antarctica

Host: Stilbocarpa litter

Subgenus Anereynetes Fain

Anereynetes Fain, 1964a: 104; 1985b: 257

Type species: Ereynetes hydrophilus Cooreman, 1947

Ereynetes (Anereynetes) beauchampi Hunter

Ereynetes beauchampi Hunter, 1964: 189

Ereynetes (Anereynetes) beauchampi Hunter: Fain & Camerik, 1994: 151

Type locality: on ginger root, from China at Boston

Distribution: ? China

Host: on ginger roots

Ereynetes (Anereynetes) bipilosus Willmann

Ereynetes bipilosus Willmann, 1953: 482; Franz, 1954: 365

Ereynetes (Anereynetes) bipilosus Willmann: Fain, 1964a: 107; Fain & Camerik, 1994: 153 (with ?)

Type locality: Germany

Distribution: Austria, Germany

Host: soil

Ereynetes (Anereynetes) biscutatus (Berlese)

Micrereunetes biscutatus Berlese, 1923: 243

Ereynetes biscutatus (Berlese): Thor, 1933: 75; Franz, 1954: 365

Ereynetes (Anereynetes) biscutatus (Berlese): Fain, 1964a: 107; Fain & Camerik, 1994: 152 (with ?)

Type locality: “in Agro Veneto”

Distribution: Austria, Italy

Host: dung

Ereynetes (Anereynetes) botswanensis Fain et Camerik

Ereynetes (Anereynetes) botswanensis Fain & Camerik, 1994: 158

Type locality: Mahalapye

Distribution: Botswana

Host: Metacatharsius sp. (Coleoptera: Scarabaeidae)

Ereynetes (Anereynetes) coenobitus Hunter et Poe

Ereynetes (Anereynetes) coenobitus Hunter & Poe, 1971: 135; Fain, 1985b: 257; Fain & Camerik, 1994: 151

Type locality: Puerto Rico

Distribution: Puerto Rico

Host: Coenobita clypeatus (Crustacea, Decapoda, Brachiura: Coenobitidae)

Ereynetes (Anereynetes) copridis Fain et Camerik

Ereynetes (Anereynetes) copridis Fain & Camerik, 1994: 158

Type locality: Mahalapye

Distribution: Botswana

Host: Copris subsidens (Coleoptera: Scarabaeidae)

Ereynetes (Anereynetes) corticalis (Vitzthum)

Micrereunetes corticalis Vitzthum, 1923: 98

Ereynetes corticalis (Vitzthum): Vitzthum, 1930: 425; Mihelčič, 1958: 274

Ereynetes (Anereynetes) copridis (Vitzthum): Fain & Camerik, 1994: 152 (with ?)

Type locality: Waidhofen an der Thaya (Austria)

Distribution: Austria, Spain

Host: “im Mittelgang von Hylesinus und Myelophilus”, under Alisma plantago

Ereynetes (Anereynetes) cruciatus (Berlese)

Ereynetes cruciatus Berlese, 1910: 243

Micrereunetes cruciatus (Berlese): Fain, 1964a: 96

Ereynetes (Anereynetes) cruciatus (Berlese): Fain, 1964a: 107; Fain & Camerik, 1994: 151

Type locality: Java

Distribution: Indonesia (Java)

Host: Copris molossus (Coleoptera: Scarabaeidae)

Ereynetes (Anereynetes) davisi Hunter

Ereynetes davisi Hunter, 1964: 190

Ereynetes (Anereynetes) davisi Hunter: Fain & Camerik, 1994: 151

Type locality: Gadsden, Arizona

Distribution: USA (Arizona)

Host: “Under bark, cottonwood”

Ereynetes (Anereynetes) hydrophilus Cooreman

Ereynetes hydrophilus Cooreman, 1947: 6

Ereynetes (Anereynetes) hydrophilus Cooreman: Fain, 1964a: 107, 1985b: 257; Zabludovskaya, 1994: 36; Fain & Camerik, 1994: 151

Type locality: Kalmhoute (Belgium)

Distribution: Belgium, Ukraine

Host: in dung; described from the nest of Larus ridubundus

Ereynetes (Anereynetes) inermis Fain

Ereynetes (Gymnereynetes) inermis Fain, 1964a: 96; Fain & Camerik, 1994: 151

Type locality: Florence

Distribution: Italy

Host: humus

Ereynetes (Anereynetes) insularis Willmann

Ereynetes insularis Willmann, 1952: 158

Ereynetes (Anereynetes) insularis Willmann: Fain, 1964a: 107; Fain & Camerik, 1994: 153 (with ?)

Type locality: Wangerooge Is.

Distribution: Germany

Host: soil near the coast

Ereynetes (Anereynetes) lyratus Fain et Prasse

Ereynetes (Anereynetes) lyratus Fain & Prasse, 1973b: 233; 1985b: 257; Fain & Camerik, 1994: 151

Type locality: Etzdorf/Saalkreis

Distribution: Germany

Host: soil

Ereynetes (Anereynetes) meliponae Fletchtmann, Fain et Leal

Ereynetes (Gymnereynetes) meliponae Fletchtmann, Fain & Leal, 1985c: 95

Ereynetes (Aneynetes) meliponae: Fain & Camerik, 1994: 151

Type locality: Recife, Pernambuco

Distribution: Brazil

Host: Melipona scutellaris (Hymenoptera: Apidae, Meliponinae)

Ereynetes (? Anereynetes) ministralis (C. L. Koch)

Tydeus ministralis C. L. Koch, 1838: fasc. 20, No 12

Ereynetes ministralis (C. L. Koch): Oudemans, 1928: 382; Thor, 1932: 28; Schweizer & Bader, 1963: 248

Ereynetes (? Anereynetes) ministralis (C. L. Koch): Fain & Camerik, 1994: 152

Type locality: Germany

Distribution: Germany, Switzerland

Ereynetes (Anereynetes) minutus Willmann

Ereynetes minutus Willmann, 1949: 127

Ereynetes (Anereynetes) minutus Willmann: Fain, 1964a: 107; Fain & Camerik, 1994: 153 (with ?)

Type locality: Ciechocinek (Poland, not Germany!)

Distribution: Poland

Host: soil

Ereynetes (Anereynetes) ornatus Mihelčič

Ereynetes ornatus Mihelčič, 1958: 274

Ereynetes (Anereynetes) ornatus Mihelčič: Fain, 1964a: 107; Fain & Camerik, 1994: 153 (with ?)

Type locality: Spain

Distribution: Spain

Host: under Alisma plantation

Ereynetes (Anereynetes) papuanus Fain et Van Goethem

Ereynetes (Anereynetes) papuanus Fain & Van Goethem, 1978: 125; 1985b: 257; Fain & Camerik, 1994: 151

Type locality: Laing Island, Morobe Prov.

Distribution: Papua New Guinea

Host: Coenobita rugosa (Crustacea, Decapoda, Brachiura: Coenobitidae)

Ereynetes (Anereynetes) potator Vitzthum

Ereynetes potator Vitzthum, 1932: 78; Thor, 1933: 75

Ereynetes (Anereynetes) potator Vitzthum: Fain, 1964a: 107; Fain & Camerik, 1994: 152 (with ?)

Type locality: S. Sumatra (Ranau)

Distribution: Indonesia (Sumatra)

Host: in bamboo stumps

Ereynetes (Anereynetes) proximus Mihelčič

Ereynetes proximus Mihelčič, 1958: 275

Ereynetes (Anereynetes) proximus Mihelčič: Fain, 1964a: 107; Fain & Camerik, 1994: 153 (with ?)

Type locality: Spain

Distribution: Spain

Host: under Alisma plantation

Ereynetes (Anereynetes) sabinensis Baker

Ereynetes sabinensis Baker, 1945: 19

Ereynetes (Anereynetes) sabinensis Baker: Fain, 1964a: 107; Fain & Camerik, 1994: 153 (with ?)

Type locality: Mexico

Distribution: Mexico

Host: bat guano

Ereynetes (Anereynetes) simplex Willmann

Ereynetes simplex Willmann, 1936: 196; Turk, 1970: 40

Ereynetes (Anereynetes) simplex Willmann: Fain, 1964a: 107; Fain & Camerik, 1994: 153 (with ?)

Type locality: graphite mine nr. Mährisch-Altstadt (Stare Mesto), Glatzer Schneeberg

Distribution: Bulgaria, Czech Rep. (on the border with Poland)

Host: unknown

Ereynetes (? Anereynetes) sittardiensis Oudemans

Ereynetes sittardiensis Oudemans, 1912: 247; 1914: 120; Vitzthum, 1931: 81; Thor, 1933: 73

Ereynetes (? Anereynetes) sittardiensis Oudemans: Fain, 1964a: 107; Fain & Camerik, 1994: 152; Zabludovskaya, 1994: 36

Type locality: Sittard (Holland)

Distribution: Holland, Ukraine

Host: in the nests of rodents and insectivorans

Ereynetes (Anereynetes) upelbensis Corpuz-Raros

Ereynetes (Anereynetes) upelbensis Corpuz-Raros, 1975: 329; Fain & Camerik, 1994: 151

Type locality: Mt. Makiling, Laguna

Distribution: Philippines

Subgenus Gymnereynetes Fain in,

Gymnereynetes Fain, 1964a: 104; 1985b: 257

Type species: Ereynetes macquariensis Fain, 1962

Ereynetes (Gymnereynetes) aurantipes Fain et Bafort

Ereynetes (Gymnereynetes) aurantipes Fain & Bafort, 1973a: 411; 1985b: 257; Zabludovskaya, 1994: 36; Fain & Camerik, 1994: 152

Type locality: De Pinte nr. Gand (Belgium)

Distribution: Belgium, Ukraine, Crimea

Host: in humus and moss

Ereynetes (Gymnereynetes) brevipes (Berlese)

Micrereunetes (Protereunetes) brevipes Berlese, 1923: 245

Protereunetes brevipes (Berlese): Macfadyen, 1954: 283

Ereynetes (Gymnereynetes) brevipes (Berlese): Fain, 1964a: 107; 1985a: 275; Fain & Camerik, 1994: 152

Type locality: Chianti (in the description: “habitat in Etruria”)

Distribution: Italy, Yan Mayen I.

Host: unknown

Ereynetes (Gymnereynetes) exilis Fain et Prasse

Ereynetes (Gymnereynetes) exilis Fain & Prasse, 1973b: 230; 1985b: 257; Zabludovskaya, 1994: 36; Fain & Camerik, 1994: 152

Type locality: Etzdorf/Saalkreis, Germany

Distribution: Germany, Ukraine, Crimea

Host: in soil

Ereynetes (Gymnereynetes) macquariensis Fain

Ereynetes macquariensis Fain, 1962a: 921; Strandtmann, 1970: ; Crooker, 1977: 489; Luxton, 1990: 397

Ereynetes (Gymnereynetes) macquariensis: Fain, 1964a: 107; 1976: 319; 1985b: 257; Strandtmann & Tilbrook, 1968: 55; Fain & Camerik, 1994: 152

Type locality: Macquarie I.

Distribution: Macquarie I., Signy I., Kerguelen I., Marion I., Antarctic Peninsula

Subgenus Huntereynetes Fain

Huntereynetes Fain, 1964a: 110

Type species: Ereynetoides scutulis Hunter, 1964

Ereynetes (Huntereynetes) propescutulis Hunter et Rosario

Ereynetes (Huntereynetes) propescutulis Hunter et Rosario in Hunter, Rosario & Moser 1989: 16; Penttinen, Viiri & Moser, 2013: 3

Type locality: Georgia (USA)

Distribution: Finland, USA

Host: pine beetles (Coleoptera: Curculionidae, Ips) and their tunnels

Ereynetes (Huntereynetes) scutulis (Hunter)

Ereynetoides scutulis Hunter, 1964: 181

Ereynetes (Huntereynetes) scutulis (Hunter): Fain, 1964a: 110; 1985b: 258; Fain & Camerik, 1994: 152; Pernek et al., 2008:

Ereynetes scutulis (Hunter): Fernandes et al., 2013: 359

Type locality: Athens, Georgia

Distribution: Croatia, USA (Georgia)

Host: “from Ips tunnel in pine log”; phoretic on Pityokteines spinidens and P. vorontzowi (Col., Scolytidae)

Ereynetes (Huntereynetes) sinescutulis Hunter, Rosario et Moser

Ereynetes (Huntereynetes) sinescutulis Hunter, Rosario & Moser, 1989: 18; Fain & Camerik, 1994: 152 (with ?)

Type locality: Idaho

Distribution: USA

Host: on pine beetles (Ips pini)

Subgenus Opsereynetes Thor

Opsereynetes Thor, 1932: 27(as genus)

Ereynetes (Opsereynetes): Fain, 1964a: 106

Type species: Opsereynetes norvegicus Thor, 1932

Ereynetes (Opsereynetes) norvegicus (Thor)

Opsereynetes norvegicus Thor, 1932: 27 – 28; 1933: 77; Shiba, 1969b: 87

Ereynetes (Opsereynetes) norvegicus (Thor): Fain, 1964a: 106; 1985b: 258; Fain & Camerik, 1994: 152; Donczyk, 2006: 165

Type locality: South Norway

Distribution: Japan, Norway, Poland

Ereynetes (Opsereynetes) robustus (Baker)

Opsereynetes robustus Baker, 1945: 18

Ereynetes (Opsereynetes) robustus Baker: Fain, 1964a: 106; Fain & Camerik, 1994: 152

Type locality: Desierto de los Leones

Distribution: Mexico

Ereynetes (Opsereynetes) simplexus Baker

Ereynetes (Opsereynetes) simplexus Baker, 1945: 17

Ereynetes (Opsereynetes) simplexus Baker: Fain, 1964a: 106

Type locality: Desierto de los Leones

Distribution: Mexico

Fain & Cemeric (1994) indicate that: “Several other species have been included in the genus Ereynetes as species of Incertae sedis by Thor (1933): i.e. Acarus padi Schrank, 1803, Acarus junci Hermann, 1804, Acarus mammilaris Canestrini & Fanzago, 1876 and Acarus armatus Canestrini & Fanzago, 1876

Genus Hanriccardoella André, Ducarme et Lebrun

Hanriccardoella André, Ducarme & Lebrun, 2004: 81

Type species: Hanriccardoella faini André, Ducarme et Lebrun, 2004

Hanriccardoella faini André, Ducarme et Lebrun

Hanriccardoella faini André, Ducarme & Lebrun, 2004: 83

Type locality: Han Cave (Rochefort)

Distribution: Belgium

Host: free living

Genus Hydranetes Kethley

Hydranetes Kethley, 1971: 176

Type species: Hydranetes tropisternus Kethley, 1971

Hydranetes citrinensis Kethley

Hydranetes citrinensis Kethley, 1971: 181

Type locality: Citra, Marion County, Florida

Distribution: USA (Florida

Host: Tropisternus lateralis nimbatus (Col.: Hydrophilidae)

Hydranetes tropisternus Kethley

Hydranetes tropisternus Kethley, 1971: 181

Type locality: Citra, Marion County, Florida

Distribution: USA (Florida

Hosts: Tropisternus lateralis nimbatus, T. quadristriatus, T. striolatus (Col.: Hydrophilidae)

Genus Protereunetes Berlese

Micrereunetes (Protereunetes) Berlese, 1923: 245 (subgenus)

Protereunetes Berlese: Oudemans, 1931: 252 (genus)

Type species: Micrereunetes (Protereunetes) agilis Berlese, 1923

Protereunetes boerneri Thor

Protereunetes boerneri Thor, 1934: 129; Macfadyen, 1954: 283; Strandtmann, 1971: 83

Type locality: Svalbard

Distribution: Svalbard, USA (Alaska), Yan Mayen I.

Protereunetes clavifrons (R. Canestrini)

Eupodes clavifrons R. Canestrini, 1886: 716

Protereunetes clavifrons : Strandtmann, 1971: 85

Eupodes mollicellus C.L. Koch, 1838: fasc. 19, nr. 23

Cocceupodes mollicellus : Sig Thor, 1934: 178; Haarløv, 1957: 26

Type locality: Italy

Distribution: Denmark, Germany, Italy, Norway, Swalbard, Sweden

Protereunetes crozeti Strandtmann

Protereunetes crozeti Strandtmann, 1972: 48

Type locality: Crozet Island

Distribution: Crozet Island

Protereunetes maudae Strandtmann

Protereunetes maudae Strandtmann, 1967: 56

Type locality: Victoria Land, Antarctica

Distribution: Antarctica

Protereunetes minutus Strandtmann

Protereunetes minutus Strandtmann, 1967: 55

Type locality: Antarctic Peninsula

Distribution: Antarctica

Protereunetus paulinae Llano et Gless

Protereunetus paulinae Llano & Gless, 2013: 299

Type locality: Hallett Station

Distribution: Antarctica

Protereunetes perforatus Shiba

Protereunetes perforatus Shiba, 1978: 147

Type locality: Mt. Sagaramine, Shigenobu-sho, Ehime

Distribution: Japan

Protereunetes striatellus (C.L. Koch)

Eupodes striatellus C.L. Koch, 1838: fa. 19, t.16

Protereunetes striatellus (C.L. Koch): Oudemans, 1938: 870; Thor & Willmann, 1941: 55

Type locality: Regensburg (Germany)

Distribution: Germany, Estonia, Holland

Protereunetes villosus Shiba

Protereunetes villosus Shiba, 1978: 145

Type locality: Tomisato, Betshimura, Ehime

Distribution: Japan

Genus Pseudotydeus Baker et Delfinado

Pseudotydeus Baker & Delfinado, 1974: 444

Type species: Pseudotydeus perplexus Baker et Delfinado, 1974

Pseudotydeus lebruni André et Ducarme

Pseudotydeus lebruni André & Ducarme, 2003: 375

Type locality: Nou Moulin Cave, Wallonia

Distribution: Belgium

Pseudotydeus perplexus Baker et Delfinado

Pseudotydeus perplexus Baker & Delfinado, 1974: 447

Type locality: Columbus, Ohio

Distribution: USA (Ohio)

Genus Riccardoella Berlese

Micrereunetes (Riccardoella) Berlese, 1923: 244

Type species: Acarus limacum Schrank, 1776

Subgenus Proricardoella Fain et van Goethem

Proricardoella Fain & van Goethem, 1986: 129

Type species: Riccardoella oudemansi Thor, 1932

Riccardoella (Proriccardoella) canadensis Fain et Van Goethem

Riccardoella (Proriccardoella) canadensis Fain & Van Goethem, 1986: 134; Zabludovskaya, 1995: 90; André, Ducarme & Lebrun, 2004: 81

Type locality: Morgan Arboretum (Canada)

Distribution: Canada, Ukraine

Host: humus

Riccardoella (Proriccardoella) novaezealandiae Fain et Barker

Riccardoella (Proriccardoella) novaezealandiae Fain & Barker, 2004: 44

Type locality: Mt. Hikurangi, Poverty Bay

Distribution: New Zealand

Host: undescribed genus and species of Athoracophoridae (Gastropoda: Pulmonata)

Riccardoella (Proriccardoella) oudemansi Thor

Riccardoella oudemansi Thor, 1932a: 249; Sellnick, 1940: 33; Willmann, 1941: 58; 1951: 142; Franz, 1954: 365; Varga & Kontschan, 2004: 95

Riccardoella jenynsi Thor, 1933: 66

Riccardoella (Proriccardoella) oudemansi Thor: Fain & Van Goethem, 1986: 129; Fain & Barker, 2004: 48

Type locality: Holland

Distribution: Austria, Bulgaria, Croatia, England, Holland, Hungary, Island, Italy, Mexico, ?Slovakia, South Africa

Hosts: Arion ater, A. lusitanicus, A. rufus (Gastropoda: Arionidae), Deroceras reticulatum, Limax agrestis, L. cinereoniger, L. cinereus, L. flavus, L. maximus, (Gastropoda: Limacidae), Tandonia budapestensis (Gastropoda: Milacidae), Oxychilus draparnaudi, Meledella werneri (Gastropoda: Zonitidae), Cepaea nemoralis (Gastropoda: Helicidae)

Riccardoella (Proriccardoella) reaumuri Fain et Van Goethem

Riccardoella (Proriccardoella) reaumuri Fain & Van Goethem, 1986: 134; Fain & Barker, 2004: 48; Waki & Shimano, 2019: 97

Type locality: Jura, nr. Ponarlier

Distribution: France, Japan

Host: Arianta arbustorum (Gastropoda: Helicidae)

Riccardoella (Proriccardoella) triodopsis Fain et Klompen

Riccardoella (Proriccardoella) triodopsis Fain & Klompen, 1990: 187

Type locality: Bankhead Nat. Forest, Lawrence Co., Alabama

Distribution: USA (Alabama), Japan

Hosts: Xolotrema obstructa [Triodopsis obstricta] (Gastropoda: Polygyridae),

Acusta despecta sieboldtiana (Gastropoda: Bradybaenidae)

Riccardoella (Proriccardoella) zadielensis André, Ducarme et Lebrun

Riccardoella zadielensis André, Ducarme & Lebrun, 2004: 85

Type locality: Zádiel Gorge, Zádiel and Silica (Slovakia), Amateur’s Cave (Czech Rep.)

Distribution: Czech Rep., Slovakia

Host: free living

Subgenus Riccardoella Berlese

Riccardoella Berlese, 1923: 244 (subgenus); Fain & van Goethem, 1986: 129

Type species: Acarus limacum Schrank, 1776

Riccardoella (R.) limacum (Schrank)

Acarus limacum Schrank, 1776: 13; 1781: 521

?Acarus helicis Schrank, 1803: 209

Ereynetes limacum (Schrank): Berlese, 1883 (in part); Mégnin, 1889

Micrereunetes limacum : Fain, 1964a: 100

Micrereunetes (Ricardoella) limacum: Berlese, 1923: 244

Riccardoella limacum (Schrank): Thor, 1929: 145; Oudemans, 1929: 254; 1931: 253; Turk & Phillips, 1945: 448 (in part); Schweizer & Bader, 1963: 249; Arutyunyan, 1972: 108; Haralabidis et al., 1983: 144

Riccardoella (R.) limacum (Schrank): Fain & van Goethem, 1986: 130; Fain & Klompen, 1990: 190

Type locality: France

Distribution: Armenia, Belgium, Bulgaria, England, France, Greece, Italy, N. Macedonia (“S. Yugoslavia”), Poland, New Zealand, Serbia, South Africa, Switzerland, USA (recorded also from other countries, but, according to Fain & van Goethem, 1986, these data may concern Riccardoella oudemansi)

Hosts: Arianta arbustorum, Helix aspersa, H. lucorum, H. pomatia (Gastropoda: Helicidae), Cernuella aginnica (Gastropoda: Hygromiidae), Athoracophorus bitentaculatus (Gastropoda: Athoracophoridae) and other snails

Riccardoella (R.) tokyoensis Waki, Hiruta et Shimano

Riccardoella tokyoensis Waki, Hiruta & Shimano, 2018: 163

Type locality: Tokyo

Distribution: Japan

Hosts: Tauphaedusa tau (Gastropoda: Clausiliidae)

Species inquirenda:

Riccardoella concolor (Haldeman)

Hypopus concolor Haldeman, 1851

Ereynetes concolor : Banks, 1915: 22

Riccardoella concolor : Thor, 1933

Distribution: ?North America

Subfamily Speleognathinae Womersley

Spelaeognathidae Womersley, 1936: 312 (see the comments of Domrow, 1969: 380)

Speleognathinae: Fain, 1957: 251 and auct.

Type genus: Speleognathus Womersley, 1936: 313

Tribe Boydaiini Fain

Boydaiini Fain, 1985a: 144

Type genus: Boydaia Womersley, 1953

Genus Boydaia Womersley

Boydaia Womersley, 1953: 82; Fain, 1962 : 7; 1963: 17

Type species: Speleognathus sturni Boyd, 1948

Boydaia agelaii Fain et Aitken

Boydaia (Boydaia) agelaii Fain & Aitken, 1968: 82; Fain & Lukoschus, 1972a: 90

Boydaia agelaii Fain & Aitken: Bernardon et al., 2015, 2018

Type locality: La Paille Village, Trinidad

Distribution: Brazil, Suriname, Trinidad, USA

Hosts: Agelaius icterocephalus, A. phoeniceus, A. ruficapillus, Chrysomus ruficapillus (Passeriformes: Icteridae)

Boydaia americana Fain

Boydaia spatulata americana Fain, 1963c: 46

Boydaia americana : Fain & Aitken, 1968: 80; 1970: 330

Boydaia (Boydaia) americana Fain: Fain, 1971a: 108

Type locality: South America, Brazil (Zoo birds)

Distribution: Brazil, South Africa, Trinidad

Hosts: Guiraca coerulea coeruela, Sporophila americana, Cyanocompsa cyanoides rotschildii, Volatinia jacarina, Piranga rubra rubra, Tangara gyrola, Rhamphocelus carbo, Coereba flaveola

Boydaia aratingae Fain

Boydaia aratingae Fain, 1963: 42; Fain & Aitken, 1969: 34

Boydaia (Boydaia) aratingae Fain: Fain, 1971a: 111

Type locality: Anvers (coming from Brazil)

Distribution: Brazil

Host: Aratinga jandaya (Psittaciformes: Psittacidae, Psittacinae)

Boydaia bradornis Fain

Boydaia bradornis Fain, 1956c: 654; Zabludovskaya, 1998, 2012; Skoracki et al., 2018: 421

Boydaia (Boydaia) bradornis Fain, 1958a: 176

Type locality: Akanyaru (Rwanda)

Distribution: Rwanda, Ukraine

Hosts: Agricola [Bradornis] pallidus griseus, Muscinia muscinia (Passeriformes: Muscicapidae)

Boydaia buphagi Fain

Boydaia (Boydaia) buphagi Fain: Fain, 1961: 58; 1963d: 56, 1971a: 110; Skoracki et al., 2018: 422

Type locality: Rwanda

Distribution: Rwanda

Hosts: Buphagus africanus (Passeriformes: Buphagidae), Lamprocolius chloropterus (Passeriformes: Sturnidae)

Boydaia cecropis Fain

Boydaia (Boydaia) cecropis Fain, 1969c: 374; Skoracki et al., 2018: 423

Type locality: Akanyaru

Distribution: Rwanda

Host: Cecropis abyssinica unitatis (Passeriformes: Hirundinidae)

Boydaia cinnyris Fain

Boydaia cinnyris Fain, 1969c: 373, 1971: 98; Skoracki et al., 2018: 423

Type locality: Akanyaru

Distribution: Rwanda

Hosts: Cinnyris cupreus cupreus (= septentrionalis), Chalcomitra senegalensis lamperti (= aequatorialis) (Passeriformes: Nectariniidae)

Boydaia clarki Fain

Boydaia clarki Fain, 1963: 41

Boydaia (Boydaia) clarki Fain: Fain, 1971a: 110

Type locality: Zoo d’Anvers (from the USA)

Distribution: USA

Host: Callipepla squamata (Galliformes: Odontophoridae)

Boydaia colini Clark

Boydaia colini Clark, 1958: 78; Pence, 1973a: 365

Boydaia (Boydaia) colini Clark: Fain, 1971a: 110

Type locality: Maryland

Distribution: USA (Louisiana, Maryland)

Host: Colinus virginianus (Galliformes: Odontophoridae)

Boydaia cyanerpes Fain

Boydaia cyanerpes Fain, 1963c: 45; Fain & Aitken,1969: 42

Type locality: Recife (dead in Anvers)

Distribution: Brazil

Host: Cyanerpes cyaneus (Passeriformes: Thraupidae)

Boydaia faini Dusbábek et Černý

Boydaia faini Dusbábek & Černý, 1970: 276

Type locality: Havana

Distribution: Cuba

Host: Dendroica cerulea (Passeriformes: Parulidae)

Boydaia falconis Fain

Boydaia falconis Fain, 1956c: 657, 1963c: 60, 1972: 610; Domrow, 1969: 381; Skoracki et al., 2018: 423

Type locality: Akanyaru (Rwanda)

Distribution: Australia, Rwanda, Chad (Tibesti)

Hosts: Falco berigora, F. cuvieri, F. t. tinnunculus (Falconiformes: Falconidae)

Boydaia formicarii Fain et Aitken

Boydaia formicarii Fain & Aitken, 1970: 331

Type locality: Moscueiro Ferry, Benavides

Distribution: Brazil

Hosts: Formicarius analis, F. colma (Passeriformes: Formicariidae)

Boydaia hirundoae Fain

Boydaia hirundoae Fain, 1956c: 661; 1960: 114; 1963d: 179; Domrow, 1965: 43; 1969: 381; Pence, 1973a: 365; 1973 : 887; 1975: 18; Fain & Aitken, 1968: 81; Fain & Hyland, 1970: 40; Skoracki et al., 2018: 423

Boydaia (Boydaia) hirundoae Fain: Fain, 1958a: 176; 1960 : 114;1963 : 56, 1971: 100; Fain, Herin & Puylaert, 1977: 393

Type locality: Akanyaru (Rwanda)

Distribution: Australia, Belgium, Brazil, Mexico, Rwanda, South Africa, Trinidad & Tobago, USA

Hosts: Hirundo abyssinica, H. neoxena, H. rustica, Petrochelidon [Hylochelidon] ariel, Progne chalybea, P. subis, Stelgidopteryx ruficollis (Passeriformes: Hirundinidae)

Boydaia indica Fain

Boydaia indica Fain: 1969c: 373; Skoracki et al., 2018: 424

Boydaia (Boydaia) indica Fain: Fain, 1971 : 101

Type locality: Anvers Zoo

Distribution: DR Congo, India, East Africa

Hosts: Acridotheres tristis, Gracupita [Sturnopastor] contra, Sturnus malabaricus, S. pagodarum, Lamprocolius chloropterus, L. chalybaeus, Spreo superbus (Passeriformes: Sturnidae)

Boydaia jordani Van Eyndhoven

Boydaia jordani Van Eyndhoven, 1955: 203

Boydaia (Boydaia) jordani Van Eyndhoven: Fain, 1958a: 176

Boydaia turdi Clark, 1967: 294 – Type: ex Turdus migratorius, Bitterroot Valley, Montana, USA

Type locality: Kampen, Holland

Distribution: Holland, USA

Hosts: Turdus migratorius, T. pilaris (Passeriformes: Turdidae)

Boydaia laticoxa Fain et Aitken

Boydaia laticoxa Fain & Aitken, 1969: 34; 1970: 328

Type locality: APEG Forest nr. Belem

Distribution: Brazil

Host: Glyphorynchus spirurus (Passeriformes: Dendrocolaptidae)

Boydaia laticoxa philydori Fain et Aitken

Boydaia (Boydaia) laticoxa philydori Fain & Aitken, 1970: 328

Type locality: APEG Forest (Station A), IPEAN, Belém

Distribution: Brazil

Host: Philydor pyrrhodes (Passeriformes: Furnariidae)

Boydaia locustellae Fain

Boydaia locustellae Fain, 1970b: 320

Type locality: Hoboken nr. Anvers

Distribution: Belgium

Host: Locustella luscinioides (Passeriformes: Locustellidae)

Boydaia loxiae Fain

Boydaia loxiae Fain, 1963d: 180

Type locality: Belgium

Distribution: Belgium

Host: Loxia curvirostra (Passeriformes: Fringillidae)

Boydaia maluri Domrow

Boydaia maluri Domrow, 1969: 386

Type locality: Cowley Beach, Kowanyama

Distribution: Australia

Hosts: Malurus amabilis, M. melanocephalus (Passeriformes: Sylviidae)

Boydaia mimi Fain et Hyland

Boydaia mimi Fain & Hyland, 1970: 40; Pence, 1973: 365

Type locality: Vera Cruz

Distribution: Mexico

Hosts: Mimus gilvus, M. polyglottos (Passeriformes: Mimidae)

Boydaia morenoi Dusbábek et Černý

Boydaia morenoi Dusbábek & Černý, 1970: 274

Type locality: El Dorado, Prov. of Las Villas

Distribution: Cuba

Host: Mimus polyglottos orpheus (Passeriformes: Mimidae)

Boydaia myzomelae Domrow

Boydaia myzomelae Domrow, 1969: 383

Type locality: Esk

Distribution: Australia

Host: Myzomela sanguinolenta (Passeriformes: Meliphagidae)

Boydaia nectarinia Fain

Boydaia (Boydaia) nectarinia Fain, 1958a: 180, 1969

Type locality: Akanyaru

Distribution: Rwanda

Hosts: Cyanomitra verticalis viridisplendens, Chalcomitra senegalensis lamperti (= aequatorialis), Cinnyris cupreus cupreus (= septentrionalis) (Passeriformes: Nectariniidae)

Boydaia pheucticola Pence et Casto

Boydaia pheucticola Pence & Casto, 1976: 468; Spicer, 1977b: 498

Type locality: Texas, Jeff Davis Co., Fort Davis

Distribution: Mexico, USA

Hosts: Pheucticus melanocephalus, Spiza americana (Passeriformes: Cardinalidae)

Boydaia podargi Fain et Lukoschus

Boydaia podargi Fain & Lukoschus, 1979a: 21; Koch, 1986: 61

Type locality: W. Australia

Distribution: Australia

Host: Podargus strigoides (Caprimulgiformes: Podargidae)

Boydaia psalidoprocnei Fain

Boydaia psalidoprocnei Fain, 1956c: 660, 1963 : 57, 1963c: 56; Pence, 1973a: 365

Boydaia (Boydaia) psalidoprocnei Fain, 1958a: 176, 1971: 100

Type locality: Akanyaru (Rwanda)

Distribution: Canada, Russia, Rwanda, USA (Louisiana)

Hosts: Iridoprocne bicolor, Psalidoprocne albiceps, Riparia riparia, Tachycineta bicolor (Passeriformes: Hirundinidae)

Boydaia psittaci Fain et Mortelmans

Boydaia (Boydaia) psittaci Fain & Mortelmans, 1959: 16

Boydaia psittaci : Fain & Aitken, 1969: 42

Type locality: Antwerp Zoo [Peru]

Distribution: Brazil, Mexico, Peru

Hosts: Pionites melanocephala, P. leucogaster, Bolborhynchus lineola, Forpus passerinus, Brotogeris versicolorus, Amoropsittaca aymara (Psittaciformes: Psittacidae)

Boydaia pycnonoti Fain

Boydaia pycnonoti Fain, 1956c: 655; Skoracki et al., 2018: 425

Boydaia (Boydaia) pycnonoti Fain: Fain, 1958a: 176, 1963c: 57; 1971: 98; Fain, Herin & Puylaert, 1977: 393

Type locality: Rwanda

Distribution: India, Kenya, Malaysia, Rwanda, South Africa, Zimbabwe

Hosts: Pycnonotus barbatus tricolor, P. goiavier personatus, P. jocosus, P. leucogenys leucotis, P. leucotis, P. xanthopygus (Passeriformes: Pycnonotidae)

Boydaia quiscali Clark

Boydaia quiscali Clark, 1960: 108; Fain & Aitken, 1970: 330; Pence, 1973a: 365

Boydaia (Boydaia) quiscali Clark, 1960: Fain, 1971 : 109

Type locality: Louisiana

Distribution: Brazil, Mexico, USA

Hosts: Quiscalus quiscula (Passeriformes: Icteridae), Saltator maximus, S. coerulescens (Passeriformes: Thraupidae)

Boydaia rosickyi Dusbábek et Černý

Boydaia rosickyi Dusbábek & Černý, 1970: 273

Type locality: Havana

Distribution: Cuba

Host: Passerina cyanea (Passeriformes: Fringillidae)

Boydaia saxicolae Fain

Boydaia saxicolae Fain, 1969: 370, 1971: 98; Skoracki et al., 2018: 425

Boydaia (Boydaia) saxicolae Fain: Fain, 1971a: 100

Type locality: Astrida (= Bucavu), Rugege Forest (Rwanda)

Distribution: Holland, Rwanda

Hosts: Saxicola torquatus axillaris, Monticola angolensis, Myrmecocichla nigra (Passeriformes: Muscicapidae), Turdus pilaris (Passeriformes: Turdidae)

Boydaia sinensis Fain et Bafort

Boydaia (Boydaia) sinensis Fain & Bafort, 1963: 80

Type locality: China

Distribution: China

Host: Leothrix lutea (Passeriformes: Timaliidae)

Boydaia spatulata Fain

Boydaia sturni var. spatulata Fain, 1955b: 695

Boydaia spatulata Fain, 1956a: 47; 1956c: 651; 1956a: 33; 1958a: 10; Domrow 1969: 381; Skoracki et al., 2018: 426

Boydaia (Boydaia) spatulata Fain, 1958a: 176; 1960 : 114, 1963c: 56; 1971: 100

Type locality: Akanyaru, Butare (= Astrida)(Rwanda)

Distribution: Australia, Botswana, Guinea, Rwanda, South Africa, USA

Hosts: Melaniparus leucomelas insignis (Passeriformes: Paridae), Erythropygia [Cercotrichos] hartlaubi, Pogonocichla stellata ruwenzorii, Sheppardia polioptera, Cercomela tractrac (Passeriformes: Muscicapidae), Sialia sialis (Passeriformes: Turdidae), Tiaris canorus, Tangara sp. (Passeriformes: Thraupidae), Cissomela pectoralis, Philemon citreogularis, P. corniculatus (Passeriformes: Melliphagidae), Iduna similis (Passeriformes: Acrocephalidae), Schistolais leucopogon reichenowi (Passeriformes: Cisticolidae), Macronyx croceus (Passeriformes: Motacillidae),

Boydaia sturni (Boyd)

Speleognathus sturni Boyd, 1948: 9; 1951: 79

Boydaia sturni : Womersley, 1953: 82; Cooreman, 1954: 426; Fain, 1956a: 42; 1956c: 651; 1961: 57; 1963c: 56; Domrow, 1969: 381; Pence, 1973a: 365

Boydaia (Boydaia) sturni: Fain, 1958a: 176

Type locality: Ithaca, N.Y.

Distribution: [Australia], Belgium, USA

Hosts: Acridotheres tristis, Ampeliceps coronatus, Sturnus vulgaris, (Passeriformes: Sturnidae), Quiscalus [Cassidix] mexicanus (Passeriformes: Icteridae)

Boydaia synallaxis Fain, Hyland et Aitken

Boydaia (Boydaia) synallaxis Fain, Hyland & Aitken, in Fain, 1969c: 376; Fain & Aitken, 1970: 332

Type locality: Musqueiro Ferry, Para, Belém (Brazil), Vera Cruz Cruz (Mexico)

Distribution: Brazil, Mexico

Hosts: Synallaxis rutilans, S. gujanensis, S. erythrothorax (Passeriformes: Furnariidae), Empidonax sp. (Passeriformes: Tyrannidae)

Boydaia tyrannis Ford

Boydaia tyrannis Ford, 1959: 380; Fain & Aitken, 1970: 330

Boydaia (Boydaia) tyrannis Ford: Fain, 1971a: 109

Type locality: Hickory Corners, Michigan

Distribution: Brazil, Mexico, Trinidad, USA (Michigan)

Hosts: Tyrannus tyrannus, Elaenia flavogaster, Myiophobus fasciatus, Pyrocephalus rubinus (Passeriformes: Tyrannidae), Manacus manacus, Pipra pipra, P. fasciicauda, P. iris (Passeriformes: Pipridae), Pachyramphus [=Platypsaris] aglaiae (Passeriformes: Tytiridae)

Boydaia willmanni Fritsch

Boydaia willmanni Fritsch, 1962: 237

Type locality: Еrlangen

Distribution: Germany

Host: Gallinago [Capella] gallinago (Charadriiformes: Scolopacidae)

Boydaia zosteropis Fain

Boydaia zosteropis Fain, 1963c: 43; 1970b: 320; Domrow, 1969: 381; Skoracki et al., 2018: 426

Boydaia (Boydaia) zosteropis Fain: Fain, 1971a: 110

Type locality: Rugege (Rwanda)

Distribution: Australia, Rwanda

Hosts: Zosterops lateralis, Z. senegalensis stuhlmanni (= scotti) (Passeriformes: Zosteropidae)

Boydaia zumpti Fain

Boydaia zumpti Fain, 1955b: 695; 1956a: 45; 1956c: 654; Skoracki et al., 2018: 427

Boydaia (Boydaia) zumpti Fain, 1958a: 176

Type locality: Astida (= Butare)

Distribution: Rwanda

Hosts: Euryllus [Andropadus] latirostris latirostris [= eugenius], Bleda syndactyla woosnami, (Passeriformes: Pycnonotidae), Schoenicola brevirostris alexinae (Passeriformes: Locustellidae)

Genus Coboydaia Fain

Boydaia (Coboydaia) Fain, 1971: 98 (subgenus)

Coboydaia Fain: Fain, 1985a: 146 (genus)

Type species: Boydaia clavata Fain, 1955

Subgenus Apodiboydaia Fain

Coboydaia (Apodiboydaia) Fain, 1985a: 146

Type species: Boydaia trochila Fain, 1958

Coboydaia (Apodiboydaia) trochila (Fain)

Boydaia trochila Fain, 1958: 11

Boydaia (Coboydaia) trochila Fain: Fain, 1971a: 112

Boydaia (Apodiboydaia) trochila Fain: Fain, 1985a: 146

Type locality: Antwerp Zoo

Distribution: Brazil

Host: Chlorestes notatus (Apodiformes: Trochilidae)

Subgenus Coboydaia Fain

Boydaia (Coboydaia) Fain, 1971a: 98 (subgenus)

Coboydaia Fain: Fain, 1985a: 146 (genus)

Type species: Boydaia clavata Fain, 1955

Coboydaia (Coboydaia) amandavae (Fain)

Boydaia amandavae Fain, 1962b: 363, 1963 : 58, 1970a: 319

Coboydaia amandavae : Fain, 1970b: 319; 1971: 101; Skoracki et al., 2018: 427

Type locality: Antwerp Zoo

Distribution: Central Africa, Indonesia (Java)

Hosts: Amandava subflava, A. amandava, Erythrura prasina, Sporaeginthus subflavus (Passeriformes: Estrildidae)

Coboydaia (Coboydaia) clavata (Fain)

Boydaia clavata Fain, 1955b: 694; 1956c: 655, 1956a: 44, 1960b: 101, 1963 : 371, 1963c: 59; Skoracki et al., 2018: 428

Boydaia (Apodiboydaia) clavata Fain: Zabludovskaya, 2002: 93

Boydaia (Coboydaia) clavata Fain: Fain, Herin & Puylaert, 1977: 392

Type locality: Rwanda

Distribution: Burundi, Cameroon, Ghana, Guinea, Rwanda, Senegal, South Africa, Zambia, Zimbabwe

Hosts: Euplectes hordeaceus, E. ardens, E. macroura, Othyphantes baglafecht stuhlmanni, Ploceus baglafecht stuhlmanni, P. [Textor] cucullatus, P. [Textor] cucullatus graueri, P. pelzelni, P. velatus, P. vitellinus, P. xanthops (= P. x. camburni), Quelea quelea, Q. quelea centralis, Q. quelea aethiopica, Q. erythrops (Passeriformes: Ploceidae), Saxicola rubetra (Passeriformes: Turdidae) , Vidua chalybeata, V. chalybeata amauropteryx (Passeriformes: Viduidae), Streptopelia vinacea (Columbiformes: Columbidae)

Coboydaia (Coboydaia) nigra (Fain)

Boydaia nigra Fain, 1955b: 695; 1956a: 44; 1956c: 658; 1959: 33; 1962: 266; 1963a: 168, 1963c: 58, 1963d: 179; Fain & Aitken, 1968: 81; Pence 1973a: 365, 1973c: 888

Boydaia (Coboydaia) nigra Fain, 1955b: 695; 1971: 101; Fain & Goff, 1980: 506; Fain, Herin & Puylaert, 1977: 393

Boydaia (Coboydaia) nigra nigra: Fain & Hyland 1975: 207

Boydaia nigra icteri Fain & Hyland, 1970: 38

Ereynetes crassipes Berlese & Trouessart, 1889: 141; Fain, 1957: 253

Boydaia crassipes (Berlese et Trouessart, 1889): Domrow 1969: 386 (Australia): Riccardoella crassipes: Thor, 1933: 66

Boydaia sturni , sensu Porter & Strandtmann, 1952 not Speleognathus sturni Boyd, 1948; Fain, 1958a: 179

Coboydaia nigra (Fain): Skoracki, 2018: 430

Type locality: Rwanda

Distribution: Belgium, Botswana, Burundi, Cameroon, Guinea, Kazakhstan, Mexico, Russia, Rwanda, South Africa, Trinidad & Tobago, Turkmenistan, Ukraine, USA (Hawaii)

Hosts: Icterus galbula, I. spurius (Passeriformes: Icteridae), Motacilla capensis, M. flava (Passeriformes: Motacillidae), Carduelis carduelis, Carpodacus erythrinus, Euphonia violacea, Fringilla coelebs, F. montifringilla, Haemorhous mexicanus, Serinus sulphuratus shelleyi, S. citrinelloides frontalis, S. mozambicus barbatus, S. [Poliospiza] burtoni tanganyicae, Serinus [Poliospiza] gularis (Passeriformes: Fringillidae), Emberiza aureola, E. citrinella, E. pusilla, E. rustica, E. rutila, E. spodocephala, E. tristrami, Spizella passerina (Passeriformes: Emberizidae), Passer ammodendri, P. domesticus, P. griseus, P. hispaniolensis, P. montanus (Passeriformes: Passeridae), Tanagra violacea (Passeriformes: Thraupidae), Oenanthe isabellina (Passeriformes: Turdidae), Lalage tricolor (Passeriformes: Campophagidae)

Coboydaia (Coboydaia) nigra motacillae (Fain)

Boydaia nigra motacillae Fain, 1969c: 375

Coboydaia nigra motacillae Fain: Zabludovskaya, 2002: 93

Type locality: Nyakibanda, Butare (= Astrida)(Rwanda), Busumbura (Burundi)

Distribution: Belgium, Burundi, Rwanda, Russia, South Africa

Hosts: Anthus trivialis, Motacilla aguimp vidua, M. alba, M. capensis, M. flava (Passeriformes: Motacillidae), Acrocephalus arundinaceus, A. s. scirpaceus (Passeriformes: Acrocephalidae)

Coboydaia (Coboydaia) sturnellae (Clark)

Boydaia sturnellae Clark, 1960: 104

Boydaia (Coboydaia) sturnellae: Fain, 1971a: 112

Type locality: Laurel, Maryland

Distribution: USA

Host: Sturnella magna (Passeriformes: Icteridae)

Tribe Trispeleognathini Fain

Trispeleognathini Fain, 1985a: 144

Type genus: Trispeleognathus Fain, 1958

Genus Astrida Fain

Astrida Fain, 1955b: 696

Type species: Astrida caprimulgi Fain, 1955

Subgenus Astrida Fain

Astrida Fain, 1955b: 696

Type species: Astrida caprimulgi Fain, 1955

Astrida (Astrida) caprimulgi Fain

Astrida caprimulgi Fain, 1955b: 696; 1956a: 33; 1956c: 646, 1962a: 9, 1963c: 62; Pence, 1973b: 879, 1973c: 886; Skoracki et al., 2018: 403

Type locality: Muheri nr. Astrida (= Butare)(Rwanda)

Distribution: Malaysia, Rwanda, USA

Hosts: Antrostomus carolinensis, Scotornis fossii welwitschi (Caprimulgiformes: Caprimulgidae), Bubo africanus, Glaucidium perlatum, Otus senegalensis, O. spilocephalus vulpes (Strigiformes: Strigidae)

Astrida (Astrida) onychognathi Fain

Astrida onychognathi Fain, 1964c: 37

Type locality: Butare (= Astrida)

Distribution: Rwanda

Host: Onychognathus walleri elgonensis (Passeriformes: Sturnidae)

Subgenus Cerylonyssus Fain et Aitken

Cerylonyssus Fain & Aitken, 1970: 326

Type species: Astrida (Cerylonyssus) chlorocerylei Fain et Aitken, 1970

Astrida (Cerylonyssus) chlorocerylei Fain et Aitken

Astrida (Cerylonyssus) chlorocerylei Fain & Aitken, 1970: 327

Type locality: Sapucajuba, IPEAN, Belém

Distribution: Brazil

Host: Chloroceryle aenea (Coraciiformes: Alcedinidae, Cerylinae)

Genus Aureliania Fain

Aureliania Fain, 1958a: 177 (subgenus)

Aureliania Fain, 1970b: 321 (genus)

Type species: Boydaia aureliani Fain, 1955

Aureliania aureliani (Fain)

Boydaia aureliani Fain, 1955b: 694; 1956a: 36

Boydaia (Aureliania) aureliani Fain: Fain, 1958a: 177

Neoboydaia (Aureliania) aureliani (Fain): Fain, 1963 : 61, 1963d: 181, 1963c: 52; Domrow, 1969: 389.

Aureliania aureliani (Fain): Fain 1962a: 9, 1985a: 146; Domrow 1975: 18, 1991: 1289; Skoracki et al., 2018: 404

Type locality: Butare (= Astrida)(Rwanda)

Distribution: Australia, Belgium, Rwanda

Hosts: Tyto alba alba, T. alba affinis (Strigiformes: Tytonidae)

Aureliania longipilis (Rosas Costa)

Speleognathus longipilis Rosas Costa, 1955: 79

Boydaia (Aureliania) longipilis (Rosas Costa): Fain, 1958a: 179

Neoboydaia (Aureliania) longipilis (Rosas Costa): Fain, 1963c: 61

Aureliania longipilis (Rosas Costa):

Type locality: Peia. de Buenos-Aires

Distribution: Argentina

Host: Guira guira (Cuculiformes: Cuculidae)

Genus Meropiboydaia Fain

Meropiboydaia Fain, 1985a: 145

Type species: Boydaia merops Fain, 1955

Meropiboydaia merops (Fain)

Boydaia merops Fain, 1955b: 694; 1956a: 41, 1956d: 30

Aureliania merops , Fain 1958b: 177

Neoboydaia (Aureliania) merops: Fain, 1963c: 61

Neoboydaia merops (Fain): Domrow, 1966: 210, 1969: 389

Meropiboydaia merops (Fain): Fain, 1985a: 146; Domrow, 1991: 1294

Type locality: Astrida (= Butare), Rwanda

Distribution: Australia, Rwanda

Hosts: Merops apiaster, M. ornatus (Coraciiformes: Meropidae)

Genus Metaboydaia Fain

Metaboydaia Fain, 1962 : 7; 1985a: 149

Type species: Speleognathus poffei Fain, 1955

Metaboydaia poffei (Fain)

Speleognathus poffei Fain, 1955b: 695; 1956a: 23

Speleognathus (Neospeleognathus) poffei: Fain, 1958a: 178

Neoboydaia (Metaboydaia) poffei: Fain 1962a: 7

Trispeleognathus (Neospeleognathus) poffei: Fain, 1963c: 24

Metaboydaia poffei : Fain, 1985a: 149

Type locality: Akanyaru, Astrida (= Butare)

Distribution: Rwanda

Hosts: Scopus umbretta (Pelecaniformes: Scopidae), Coracias caudatus caudatus (Coraciiformes: Coraciidae)

Genus Neastrida Fain

Neastrida Fain, 1962a: 9

Type species: Astrida parrae Fain, 1956

Neastrida parrae (Fain)

Astrida parrae Fain, 1956e: 113

Astrida (Neastrida) parrae: Fain, 1962a: 9, 1963c: 26

Neastrida parrae : Fain 1985a: 148.

Type locality: Butare (Rwanda)

Distribution: Rwanda, South Africa

Host: Actophilornis africanus (Charadriiformes: Jacanidae)

Genus Neoboydaia Fain

Boydaia (Neoboydaia) Fain, 1958a: 177 (subgenus)

Neoboydaia Fain, 1961b: 162, 1962a: 7, 1963c: 52

Type species: Boydaia philomachi Fain, 1956

Neoboydaia colymbiformi Clark

Neoboydaia colymbiformi Clark, 1964: 158; Domrow, 1969: 189; Pence, 1973a: 364, 1975: 18; Fain, 1985a: 148

Type locality: California

Distribution: USA (California, Louisiana)

Hosts: Podiceps [Colymbus] nigricollis, P. ruficollis, Podilymbus podiceps (Podicipediformes: Podicipedidae)

Neoboydia galachrysiae Fain

Neoboydia galachrysiae Fain, 1961b: 129; 1985a: 148; Skoracki et al., 2018: 409

Neoboydaia (Neoboydaia) galachrysiae: Fain, 1963c: 19; 1964 : 137

Type locality: Kasongo (Maniema Prov.)

Distribution: Democratic Republic of the Congo

Hosts: Glareola [Galachrysia] cinerea, G. nuchalis (Aves, Glareolidae)

Neoboydaia philomachi (Fain)

Boydaia philomachi Fain, 1956a: 41; 1956d: 27; 1962e: 300

Boydaia (Neoboydaia) philomachi Fain, 1958a: 177, 1962a: 7, 1964b: 141

Neoboydaia (Neoboydaia) philomachi: Fain, 1962a: 7, 1963c: 18, 1964b: 141; Domrow, 1965: 43

Neoboydaia philomachi eroliae Fain et Hyland, 1970: 42, 1975: 204; Zabludovskaya, 1998: 48

Neoboydaia philomachi : Clark, 1964: 16, fig. 2; Domrow, 1965: 43, 1969: 388; Fain & Aitken 1968: 84; Pence 1973a: 364, 1973c: 884, 885, 891; Fain, 1985a: 148; 1985b: 254

Neoboydaia philomachi thalasseus Fain & Hyland 1975: 204

Type locality: Butare (= Astrida), Rwanda

Distribution: Australia, Russia, Rwanda, USA

Hosts: Actitis macularius, Arenaria interpres, Gallinago gallinago, Limnodromus griseus, L. scolopaceus, Limosa lapponica, Calidris [Philomachus] pugnax, C. melanotos, C. minutilla, Tringa flavipes, T. melanoleuca, T. glareola, T. solitaria (Charadriiformes: Scolopacidae), Sterna hirundo, Thalasseus maximus (Charadriiformes: Laridae, Sterninae)

Genus Ophthalmognathus Dubinin

Ophthalmognathus Dubinin, 1957: 66

Neospeleognathus Fain, 1958a: 178 (subgenus of Speleognathus); 1985a: 147 (syn.)

Type species of Ophthalmognathus: Ophthalmognathus dogieli Dubinin, 1957

Type species of Neospeleognathus: Speleognathus schoutedeni Fain, 1955

Ophthalmognathus schoutedeni (Fain)

Speleognathus schoutedeni Fain, 1955b: 695; 1956b: 18; 1962e: 301

Speleognathus (Neospeleognathus) schoutedeni: Fain, 1958a: 178

Ophthalmognathus dogieli Dubinin, 1957: 68 – Russia, syn. by Domrow, 1969: 392

Neoboydaia (Neospeleognathus) schoutedeni: Fain, 1962 : 7; 1963c: 62, 1963d: 23

Trispeleognathus (Neospeleognathus) schoutedeni: Fain, 1963c: 23

Ophthalmognathus schoutedeni (Fain): Domrow, 1969: 392; 1991: 1295; Fain & Goff, 1980: 506; Fain, 1985a: 147; Halliday, 1998: 50; Skoracki et al., 2018: 418

Type locality: Butare (= Astrida)

Distribution: Australia, Rwanda, Russia

Hosts: Ardea cinerea, Ardeola idae, Nycticorax caledonicus, N. nycticorax (Pelecaniformes: Ardeidae)

Ophthalmognathus tenorioae Fain et Goff

Ophthalmognathus tenorioae Fain & Goff, 1980: 506; Goff, 1983: 167; Fain, 1985a: 147

Type locality: Oahu I., Honolulu, Campbell Estate Road

Distribution: Rwanda, USA (Hawai, Oahu I.)

Hosts: Nycticorax nycticorax, N. n. hoactli (Pelecaniformes: Ardeidae)

Genus Phoenicopteriella Fain

Phoenicopteriella Fain, 1970b: 323

Type species: Phoenicopteriella mirabilis Fain, 1970

Phoenicopteriella mirabilis Fain

Phoenicopteriella mirabilis Fain, 1970b: 323; 1985a: 152

Type locality: North Africa

Distribution: North Africa

Host: Phoenicopterus ruber (Phoenicopteriformes: Phoenicopteridae)

Phoenicopteriella womersleyi (Fain)

Speleognathus womersleyi Fain, 1955b: 695; 1956a: 21; Clark, 1958: 84

Trispeleognathus womersleyi : Fain 1958b: 178, 1962a: 9; Fain & Hyland 1975: 205; Kaneko, Matsudaira & Prince Masahito, 1978: 147

Trispeleognathus (Trispeleognathus) womersleyi: Fain, 1963c: 23

Ophthalmognathus womersleyi : Domrow, 1969: 390; Spicer, 1977: 498

Phoenicopteriella womersleyi (Fain): Fain, 1985a: 152, 1985b: 254; Skoracki et al., 2018: 412

Type locality: Butare (= Astrida)(Rwanda)

Distribution: Japan, Rwanda, USA

Hosts: Authya affinis, Anas acuta, A. clypeata, A. discors, A. strepera, Dendrocygna viduata, Sarkidiornis melanotus (Anseriformes: Anatidae)

Genus Picinyssus Fain

Picinyssus Fain, 1969c: 369

Type species: Picinyssus buccanodon Fain, 1969

Picinyssus buccanodon Fain

Picinyssus buccanodon Fain, 1969c: 370; 1985a: 151

Type locality: Butahiro and Kamahia (Nord Kivu)

Distribution: DR Congo

Host: Buccanodon duchaillui (Piciformes: Lybiidae)

Genus Psittaboydaia Fain

Psittaboydaia Fain, 1985a: 145

Trichoglossiella Fain, 1985a: 145

Type species of Psittaboydaia: Neoboydaia psittaculae Fain, 1962

Type species of Trichoglossiella: Aureliania trichoglossi Fain, 1970

Psittaboydaia amazona (Fain)

Trispeleognathus (Neospeleognathus) amazona Fain, 1972: 90 (in Fain & Lukoschus, 1972)

Psittaboydaia (P.) amazona (Fain): Fain, 1985a: 145

Type locality: Meerzorg

Distribution: Suriname

Host: Amazona amazonica (Psittaciformes: Psittacidae)

Psittaboydaia psittaculae (Fain)

Neoboydaia psittaculae Fain, 1962: 7; 1962 : 318; Domrow, 1969: 388

Neoboydaia (Neoboydaia) psittaculae Fain, 1963c: 19

Psittaboydaia (Psittaboydaia) psittaculae (Fain): Fain, 1985a: 147; Skoracki et al., 2018: 415

Type locality: Central Africa

Distribution: Australia, Central Africa

Hosts: Agapornis roseicollis, Trichoglossus moluccanus, Psitteuteles versicolor, Psittacula krammeri (Psittaciformes: Psittaculidae)

Psittaboydaia trichoglossi (Fain)

Aureliania trichoglossi Fain, 1970b: 321

Psittaboydaia (Trichoglossiella) trichoglossi (Fain): Fain, 1985a: 145

Type locality: Anvers Zoo

Distribution: New Guinea

Host: Trichoglossus haematod forsteni (Psittaciformes: Psittaculidae)

Genus Pterniboydaia Fain

Pterniboydaia Fain, 1985a: 146

Type species: Boydaia pternistis Fain, 1955

Pterniboydaia pternistis (Fain)

Boydaia pternistis Fain, 1955a: 145, 1956c: 646

Aureliania pternistis (Fain): Fain, 1963 : 373

Neoboydaia pternistis (Fain): Fain, 1963c: 21; Kaneko, Kadosaka & Toshioka, 1977: 163

Neoboydaia (Aureliania) pternistis (Fain): Fain, 1963c: 61

Pterniboydaia pternistis (Fain): Fain, 1985a: 146; Skoracki et al., 2018: 416

Type locality: Astrida (= Butare) (Rwanda)

Distribution: Belgium, Japan, Rwanda, Zambia

Hosts: Bambusicola thoracica, Pternistis afer, P. swainsoni, Perdix p. perdix (Galliformes: Phasianidae)

Genus Ralliboydaia Fain

Neoboydaia (Ralliboydaia) Fain, 1962b: 365 (subgenus)

Ralliboydaia Fain: Fain, 1985a: 151 (genus); Fain, 1963: 52 is synonymizing Ralliboydaia with Neoboydaia.

Type species: Neoboydaia (Ralliboydaia) latiralli Fain, 1962

Ralliboydaia coccyzae (Pence)

Astrida coccyzae Pence, 1973a: 366

Neoboydaia (Ralliboydaia)

Ralliboydaia coccyzae (Pence): Fain, 1985a: 151

Type locality: Mandeville, La.

Distribution: USA (Louisiana)

Host: Coccyzus americanus (Cuculiformes: Cuculidae)

Ralliboydaia latiralli (Fain)

Neoboydaia (Ralliboydaia) latiralli Fain, 1962b: 365

Neoboydaia (Neoboydaia) latiralli Fain, 1963c: 20

Ralliboydaia latiralli (Fain): Fain, 1985a: 151

Type locality: Brazil

Distribution: Brazil

Host: Latirallus melanophaius oenops (Ralliformes: Rallidae)

Ralliboydaia porphyrionis (Domrow)

Speleognathopsis porphyrionis Domrow, 1965: 44; 1969: 393

Ralliboydaia porphyrionis (Domrow): Fain, 1985a: 151

Type locality: Samford, S. E. Queensland

Distribution: Australia (Queensland)

Host: Porphyrio melanotus (Gruiformes: Rallidae)

Genus Speleognathopsis Cooreman

Speleognathopsis Cooreman, 1954: 428

Type species: Speleognathopsis galli Cooreman, 1954

Subgenus Speleognathopsis Cooreman

Speleognathopsis Cooreman, 1954: 428

Type species: Speleognathopsis galli Cooreman, 1954

Speleognathopsis (S.) accipitris (Domrow)

Ophthalmognathus accipitris Domrow, 1969: 390; Fain, 1985a: 146

Speleognathopsis accipitris (Domrow): Domrow, 1991: 1301

Type locality: Kowanyama

Distribution: Australia

Host: Accipiter fasciatus (Falconiformes: Accipitridae)

Speleognathopsis (S.) benoiti Fain [see syn. of wai and charadricola in Domrow, 1969: 393)]

Speleognathopsis benoiti Fain, 1955b: 696; 1956a: 29; 1962 : 9; 1963c: 25; 1985a: 150; Domrow, 1966: 210; 1969: 393; 1991: 1301; McDaniel, 1967: 94

Speleognathopsis wai Fain, Vercammen-Grandjean & Wagner 1966: 115. Type locality: Bloemhof, Transvaal, South Africa (syn. by Domrow, 1969: 393)

Type locality: Butare (= Astrida), Rwanda

Distribution: Australia, Rwanda, USA (Texas)

Hosts: Charadrius melanops, Erythrogonys cinctus, Lobibyx miles, Pluvialis squatarola, Vanellus [Hemiparra] crassirostris, V. senegallus (Charadriiformes: Charadriidae)

Speleognathopsis (S.) charadricola Fain

Speleognathopsis charadricola Fain, 1964c: 35; 1985a: 150

Type locality: Zoo Anvers

Distribution: Central Africa

Host: Charadrius (Leucopolius) pecuarius (Charadriiformes: Charadriidae)

Speleognathopsis (S.) galli Cooreman

Speleognathopsis galli Cooreman, 1954: 426; Fain, 1963 : 53; 1963c: 62; 1985a: 149; Domrow, 1969: 392; Talbot, 1969: 582; Faccini, 1981: 135; Panda et al., 1987: 507

Type locality: Astrida (= Butare), Rwanda

Distribution: Brazil, India, Papua New Guinea, Rwanda

Hosts: Gallus gallus domesticus (Galliformes: Phasianidae), Numida meleagris intermedia (Galliformes: Numididae)

Subgenus Speleomyotis Fain

Speleomyotis Fain, 1970: 513.

Type species: Speleognathopsis bastini Fain, 1958

Speleognathopsis (Speleomyotis) bastini Fain

Speleognathopsis bastini Fain, 1958b: 342

Speleognathopsis (Speleomyotis) bastini Fain: Fain, 1966: 108; 1970c: 513

Speleognathopsis (Speleomyotis) bastini bastini Fain: Fain & Lukoschus, 1971c: 285

Type locality: Belgium

Distribution: Belgium, Japan, Suriname

Hosts: Eptesicus fuscus, E. melanopterus, Myotis myotis, M. nigricans (Chiroptera: Vespertilionidae)

Speleognathopsis (Speleomyotis) bastini didelphis Fain et Aitken

Speleognathopsis bastini didelphis Fain & Aitken, 1969: 41

Type locality: Jandiai, Caratateua, Bragança (Pará)

Distribution: Brazil

Host: Didelphis marsupialis (Didelphimorphia: Didelphidae)

Speleognathopsis (Speleomyotis) molossus Fain et Lukoschus

Speleognathopsis (Speleomyotis) molossus Fain & Lukoschus, 1971c: 285

Type locality: Paramaribo

Distribution: Suriname

Hosts: Cynomops planirostris, Molossus ater, M. molossus (Chiroptera: Molossidae)

Genus Trispeleognathus Fain

Speleognathus (Trispeleognathus) Fain, 1958a: 178

Trispeleognathus Fain: 1962: 7

Type species: Speleognathus striatus Crossley, 1952

Trispeleognathus euricoi Amaral

Trispeleognathus euricoi Amaral, 1965: 31

Ophthalmognathus euricoi (Amaral): Domrow, 1969: 391

Trispeleognathus (Trispeleognathus) euricoi: Amaral, 1968: 124; Fain & Aitken, 1969: 43

Type locality: Itaporanga (Estado de São Paulo)

Distribution: Brazil

Hosts: Leptotila verreauxi, Nothura m. maculosa (Columbiformes: Columbidae)

Trispeleognathus striatus (Crossley)

Speleognathus striatus Crossley, 1952: 385; Fain, 1955b: 695, 1956b: 18,

Boydaia striatus : Womersley, 1953: 82; Cooreman, 1954: 426; Fain, 1955 : 695; 1962 : 300

Speleognathus (Trispeleognathus) striatus: Fain 1958a: 178

Ophthalmognathus striatus : Domrow 1969: 389; Pence & Casto, 1976: 466

Trispeleognathus (Trispeleognathus) striatus: Fain, 1963c: 22; Amaral 1963: 91; Fain & Aitken, 1969: 43, 1970: 328

Trispeleognathus striatus : Fain 1962a: 9, 1985a: 149; Amaral 1963: 91; Fain & Hyland, 1970: 42; Skoracki et al., 2018: 420

Type locality: Garza Co., Lubbock Co., Texas

Distribution: Brazil, Mexico, Rwanda, Trinidad & Tobago, USA,

Hosts: Columba livia domestica, Columbigallina passerina, Geotrygon montana, Leptotila verreauxi, Streptopelia s. semitorquata (Columbiformes: Columbidae)

Tribe Speleochirini Fain

Speleochirini Fain, 1985a: 144

Type genus: Speleochir Fain, 1966

Genus Hipposideroptes Fain

Hipposideroptes Fain, 1970c: 514

Type species: Hipposideroptes kimuenzae Fain, 1970

Hipposideroptes kimuenzae Fain

Hipposideroptes kimuenzae Fain, 1970c: 514

Type locality: Kimuenza

Distribution: DR Congo

Host: Hipposideros commersoni gigas (Chiroptera: Hipposideridae)

Hipposideroptes saccopteryx Fain et Lukoschus

Hipposideroptes saccopteryx Fain & Lukoschus, 1971c: 289

Type locality: Lelydorp

Distribution: Suriname

Hosts: Saccopteryx bilineata, S. leptura (Chiroptera: Emballonuridae)

Genus Neospeleochir Fain

Speleochir (Neospeleochir) Fain, 1970c: 514

Neospeleochir Fain:

Type species: Boydaia duboisi Fain, 1955

Neospeleochir duboisi (Fain)

Boydaia Duboisi Fain, 1955b: 691

Boydaia (Aureliania) duboisi Fain: Fain, 1958a: 176

Speleochir duboisi Fain: Fain, 1966: 108

Speleochir (Neospeleochir) duboisi Fain: Fain, 1970c : 514

Neospeleochir duboisi Fain: Fain, 1970c: 520

Type locality: Astrida (= Butare)

Distribution: Rwanda

Host: Nycteris sp. (Chiroptera: Nycteridae)

Genus Neospeleognathopsis Fain

Speleognathopsis (Neospeleognathopsis): Fain, 1958a: 177

Neospeleognathopsis Fain: Fain, 1962 : 10

Type species: Speleognathopsis chiropteri Fain, 1955

Subgenus Neospeleognathopsis Fain

Neospeleognathopsis Fain, 1958a: 177

Type species: Speleognathopsis chiropteri Fain, 1955

Neospeleognathopsis (N.) chiropteri (Fain)

Speleognathopsis chiropteri Fain, 1955b: 692; 1956a: 31

Neospeleognathopsis (N.) chiropteri (Fain): Fain, 1966: 108; 1970c: 513

Type locality: Astrida (Butare)(Rwanda)

Distribution: D.R.Congo, Egypt, Rwanda

Hosts: Eidolon helvum, Epomophorus labiatus minor, Rousettus aegyptiacus, R. aegyptiacus leachi (Chiroptera: Pteropodidae)

Subgenus Pteropignathus Fain et Lukoschus

Pteropignathus Fain & Lukoschus, 1979: 58

Type species: Neospeleognathopsis (Pteropignathus) pteropus Fain et Lukoschus, 1979

Neospeleognathopsis (Pteropignathus) pteropus (Fain et Lukoschus)

Neospeleognathopsis (Pteropignathus) pteropus Fain & Lukoschus, 1979: 58

Pteropignathus pteropus (Fain et Lukoschus):

Type locality: Brooking Springs

Distribution: Australia

Host: Pteropus scapulatus (Chiroptera: Pteropodidae)

Genus Speleochir Fain

Speleochir Fain, 1966: 103

Type species: Speleochir aitkeni Fain, 1966

Speleochir aitkeni Fain

Speleochir aitkeni Fain, 1966: 107

Speleochir (S.) aitkeni Fain: Fain, 1970c: 517

Type locality: Aripo Cave, Northern Range

Distribution: Trinidad (W.I.)

Host: Anoura g. geoffroyi (Chiroptera: Phyllostomatidae)

Speleochir barbulata Fain et Aitken

Speleochir (Speleochir) barbulata Fain & Aitken, 1970: 334

Type locality: APEG Forest, IPEAN, Belém

Distribution: Brazil

Host: Chiroptera indet.

Speleochir brasiliensis Fain et Aitken

Speleochir brasiliensis Fain & Aitken, 1969: 39

Type locality: Mucambo Forest, IPEAN, Brazil

Distribution: Brazil

Hosts: Artibeus jamaicensis, Vampyrodes caraccioloi (Chiroptera: Phyllostomatidae)

Speleochir phyllostomi (Clark)

Neospeleognathopsis phyllostomi Clark, 1967: 241

Speleochir phyllostomi Clark: Fain & Aitken, 1969: 39

Type locality: Colombia

Distribution: Colombia

Host: Phyllostomus hastatus (Chiroptera: Phyllostomatidae)

Genus Speleochiroides Fain

Speleochiroides Fain, 1985a: 146

Type species: Speleochir carollia Fain et Liukoschus, 1971

Speleochiroides carollia (Fain et Lukoschus)

Speleochir carollia Fain & Lukoschus, 1971: 293

Speleochiroides carollia (Fain et Lukoschus): Fain, 1985a: 146

Type locality: Lelydorp

Distribution: Suriname

Host: Carollia perspicillata (Chiroptera: Phylostommatidae)

Tribe Paraspeleognathopsini Fain

Paraspeleognathopsini Fain, 1985a: 145

Type genus: Paraspeleognathopsis Fain, 1958

Genus Paraspeleognathopsis Fain

Speleognathopsis (Paraspeleognathopsis) Fain, 1958a: 177

Paraspeleognathopsis Fain: Fain, 1970c: 510 (genus). According to Domrow (2997), Paraspeleognathopsis Fain, 1958 = Speleorodens Fain, 1962 = Speleomys Fain, 1971

Type species: Speleognathopsis bakeri Fain, 1955

Paraspeleognathopsis bakeri (Fain)

Speleognathopsis bakeri Fain, 1955b: 148; 1955a: 695; 1956a: 26; 1956d: 159

Speleognathopsis (Paraspeleognathopsis) bakeri Fain: Fain, 1958a: 177

Paraspeleognathopsis bakeri (Fain): Domrow, 1967: 775; 1991: 1296

Paraspeleognathopsis (P.) bakeri (Fain): Fain & Lukoschus, 1968: 85; Beron, 1973: 189

Type locality: Butare (= Astrida), Rwanda

Distribution: Australia, Belgium, Bulgaria, Holland, Japan, Korea, Rwanda, Spain

Hosts: Apodemus agrarius coreae, A. speciosus, Rattus natalensis (sub Mastomys coucha ugandae), Melomys cervinipes, Sylvaemus flavicollis, S. sylvaticus, S. sylvaticus callipides (Rodentia: Muridae)

Paraspeleognathopsis exaeta Domrow

Paraspeleognathopsis exaeta Domrow, 1991: 1298

Type locality: S.E.New South Wales, Bondo State Forest

Distribution: Australia

Host: Petaurus breviceps (Diprotodontia: Petauridae)

Paraspeleognathopsis galagoensis (Fain)

Speleognathopsis galagoensis Fain, 1956a: 32; 1956: 167

Paraspeleognathopsis (P.) galagoensis Fain, 1963c: 64

Type locality: Akanyaru

Distribution: Rwanda

Host: Galago senegalensis moholi (Primates: Galagonidae)

Paraspeleognathopsis graphiuri Fain

Paraspeleognathopsis graphiuri Fain, 1970c: 510

Type locality: Butare (= Astrida)

Distribution: Rwanda

Host: Graphiurus (Claviglis) microtis (Rodentia: Gliridae)

Genus Speleomys Fain [acc. to Domrow (1975, 2007) syn. of Paraspeleognathopsis Fain, 1958]

Speleomys Fain, 1970c: 513

Type species: Speleognathopsis galliardi Fain, 1955

Speleomys galliardi (Fain)

Speleognathopsis galliardi Fain, 1955b: 149; 1956a: 26; 1956: 163

Speleognathus galliardi Fain, 1956: 163

Speleomys galliardi (Fain): 1970c: 513

Type locality: Kansi and Butare (= Astrida)

Distribution: Rwanda

Hosts: Arvicanthis abyssinicus, Gerbilliscus boehmi (Rodentia: Muridae)

Speleomys neotropicalis (Fain et Aitken)

Paraspeleognathopsis (Speleorodens) neotropicalis Fain & Aitken, 1969: 37

Speleomys neotropicalis (Fain et Aitken): Fain, 1970c: 515

Type locality: Jandiai, Caratateua, Bragança (Pará)

Distribution: Brazil

Host: Zygodontomys lasiurus fuscinus (Rodentia: Cricetidae, Sigmodontinae)

Genus Speleorodens Fain [acc. to Domrow (1975, 2007) syn. of Paraspeleognathopsis Fain, 1958]

Speleorodens Fain, 1962: 10

Paraspeleognathopsis (Speleorodens) Fain, 1963c: 55

Type species: Speleognathopsis strandtmanni Fain, 1956

Speleorodens cricetidarum (Clark)

Paraspeleognathopsis cricetidarum Clark, 1967: 240

Paraspeleognathopsis (Speleorodens) cricetidarum: Fain & Aitken, 1969: 36

Type locality: Panama

Distribution: Brazil, Panama, Trinidad

Hosts: Oryzomys talamancae, O. laticeps velutinus, Zygodontomys cherriei, Heteromys desmarestianus, Neracomys guianae (Rodentia: Cricetidae)

Speleorodens derricki (Womersley)

Boydaia derricki Womersley, 1954: 65; Domrow & Smith, 1956: 203

Astrida derricki (Womersley): Fain, 1956: 167

Speleognathopsis derricki (Womersley): Domrow, 1961: 376

Paraspeleognathopsis (Speleorodens) derricki: Fain, 1963: 64

Speleomys derricki (Womersley):

Paraspeleognathopsis derricki (Womersley): Domrow, 1967: 775

Speleorodens derricki (Womersley): Fain, 1971c: 516

Type locality: Mt. Glorious, S.E. Queensland

Distribution: Australia

Hosts: Hydromys chrysogaster, Mus musculus, Rattus assimilis, R. conatus, R. fuscipes (Rodentia: Muridae)

Speleorodens michigensis (Ford)

Speleognathopsis michigensis Ford, 1962: 104

Paraspeleognathopsis (Speleorodens) clethrionomys Fain & Lukoschus, 1968: 86 – Holland, ex Myodes [Clethrionomys] glareolus

Speleorodens michigensis (Ford): OConnor, 1978: 124

Type locality: Ingham County, Michigan

Distribution: Holland, USA

Hosts: Microtus agrestis, M. pennsylvanicus, Myodes [Clethrionomys] gapperi, Myodes [C.] glareolus (Rodentia: Arvicolidae)

Speleorodens neotropicalis (Fain et Aitken)

Paraspeleognathopsis (Speleorodens) neotropicalis Fain & Aitken, 1969: 37

Type locality: Jandiai, Caratateua, Bragança (Pará), Brazil

Distribution: Brazil

Host: Zygodontomys lasiurus fuscinus (Rodentia: Cricetidae)

Speleorodens ruandae Fain

Speleorodens ruandae Fain, 1962b: 367

Type locality: Butare (= Astrida)

Distribution: Rwanda

Host: «dans une termitière«, accidental

Speleorodens strandtmanni (Fain)

Speleognathopsis strandtmanni Fain, 1955b: 696; 1956c: 645

Speleognathopsis sciuri Clark, 1960: syn. by Fain, 1961a: 160

Speleognathopsis (Neospeleognathopsis) strandtmanni (Fain): Fain, 1961a: 160

Paraspeleognathopsis (Speleorodens) strandtmanni (Fain): Fain, 1963c: 64

Speleorodens strandtmanni (Fain): Fain, 1970c: 518

Type locality: Astrida (= Butare), Rwanda

Distribution: Belgium, DR Congo, Panama, Rwanda, USA

Hosts: Sciurus granatensis, S. vulgaris, S. carolinensis, Funisciurus carruthersi, Tamiscus emini (Rodentia: Sciuridae)

Tribe Speleognathini Fain

Speleognathini Fain, 1985a: 144

Type genus: Speleognathus Womersley, 1936

Genus Domrownetes OConnor

Domrownetes OConnor, 1978: 128

Type species: Paraspeleognathopsis exul Domrow, 1975

Domrownetes exul (Domrow)

Paraspeleognathopsis exul Domrow, 1975: 97; 1991: 1298

Domrownetes exul (Domrow): OConnor, 1978: 97

Type locality: Mount Sabine, Otway Ranges, Victoria

Distribution: Australia (Victoria)

Host: Antechinus stuartii (Dasyuromorphia: Dasyuridae)

Genus Speleognathus Womersley

Speleognathus Womersley, 1936: 313

Type species: Speleognathus australis Womersley, 1936

Speleognathus australis Womersley

Speleognathus australis Womersley, 1936: 313; 1954; Domrow, 1961: 374; 1991: 1302; Fain, 1955b: 695; 1956c: 662; 1961; 1962 : 9; 1963c: 54; 1971 : 515; Drummond & Medley, 1964; Southcott, 1986: 201; Halliday, 1998: 51

Speleognathus (Speleognathus) australis Womersley: Fain, 1958a: 178

Speleognathus bovis Fain, 1955; 1956d: 156; 1956b: 18; 1956c: 662 (syn. de S. australis) - Type locality: Astrida (= Butare, Rwanda), ex Artiodactyla: Bovidae

Type locality: on water surface (not “in moss”, see Fain, 1956c: 662, at Glen Osmond, Adelaide, South Australia

Distribution: Australia, USA

Hosts: Bos taurus, Bison bison (Artiodactyla: Bovidae)

Subfamilia Lawrencarinae Fain

Lawrencarinae Fain, 1957: 251

Type genus: Lawrencarus Fain, 1957

Tribe Lawrencarini Fain

Lawrencarini Fain

Type genus: Lawrencarus Fain, 1957

Genus Batracarus Fain

Batracarus Fain, 1961a: 254

Type species: Batracarus hylaranae Fain, 1961

Batracarus hylaranae Fain

Batracarus hylaranae Fain, 1961a: 254; 1962b: 58

Type locality: Bogor (= Buitenzorg), Java

Distribution: Indonesia (Java)

Host: Hylarana erythraea (Anura: Ranidae)

Genus Lawrencarus Fain

Lawrencarus Fain, 1957: 250

Type species: Riccardoella eweri Lawrence, 1952

Subgenus Cernycarus de la Cruz

Cernycarus de la Cruz, 1971: 2

Type species: Lawrencarus (Cernycarus) hollandsae de la Cruz, 1971

Lawrencarus (Cernycarus) hollandsae de la Cruz

Lawrencarus (Cernycarus) hollandsae de la Cruz, 1971: 2

Type locality: Culandrillo

Distribution: Cuba

Host: Eleutherodactylus sp. (Anura: Ceratophryidae)

Subgenus Lawrencarus Fain

Lawrencarus Fain, 1957: 250

Type species: Riccardoella eweri Lawrence, 1952

Lawrencarus (L.) angelae (Womersley)

Boydaia angelae Womersley, 1953: 83; Fain, 1955b: 694; 1956c: 647

Lawrencarus angelae (Womersley): Fain, 1961a: 246; Domrow, 1961: 379; 1991: 1293; Fain, 1962 :

Type locality: South Australia

Distribution: Australia

Hosts: Limnodynastes tasmaniensis, L. peronii (Anura: Ceratophryidae, Cycloraninae)

Lawrencarus (L.) americanus Fain

Lawrencarus americanus Fain, 1961a: 250

Type locality: Florida

Distribution: USA (Florida)

Host: Hyla septentrionalis (Anura: Hylidae)

Lawrencarus (L.) brasiliensis Fain

Lawrencarus brasiliensis Fain, 1961a: 252; 1962: 52

Type locality: Alto da Serra, nr. Cubatan, State of São Paulo

Distribution: Brazil

Hosts: Cyclorhamphus asper, Eleutherodactylus guntheri (Anura: Ceratophryidae,

Ceratophryinae)

Lawrencarus (L.) brasiliensis desantisi Mauri et Bischoff

Lawrencarus brasiliensis desantisi Mauri & Bischoff, 1984: 338

Type locality: Averias, Prov. De Santiago del Estero

Distribution: Argentina

Hosts: Bufo arenarum, B. paracnemis (Anura: Bufonidae)

Lawrencarus (L.) ceratobatrachi Fain

Lawrencarus ceratobatrachi Fain, 1961a: 251; 1962 : 47

Type locality: Buin

Distribution: Bougainville I. (Papua New Guinea)

Host: Ceratobatrachus guntheri (Anura: Ranidae)

Lawrencarus (L.) domrowi Fain

Lawrencarus domrowi Fain, 1961a: 253; 1962 : 54; Domrow, 1991: 1293

Type locality: Dinner Ck, Nord Queensland

Distribution: Australia (Queensland)

Host: frog (Anura)

Lawrencarus (L.) eweri (Lawrence)

Riccardoella eweri Lawrence, 1952: 747; Lufty, 1960: 183

Boydaia angelae : Fain, 1956: 647

Lawrencarus eweri : Fain, 1957: 250; 1958a: 183; 1961: 253; 1962: 20

Lawrencarus (L.) eweri cubanus de la Cruz

Lawrencarus eweri cubanus de la Cruz, 1971: 3

Type locality: Soledad

Distribution: Cuba

Host: Bufo peltacephalus (Anura: Bufonidae)

Lawrencarus (L.) eweri dahuricus Zabludovskaya

Lawrencarus eweri dahuricus Zabludovskaya, 1992: 66

Type locality: Chitinskaya oblast, Borzyanskiy Rayon, Borzya River, loc. Arshan

Distribution: Russia

Host: Bufo raddei (Anura: Bufonidae)

Lawrencarus (L.) eweri eweri (Lawrence)

Lawrencarus eweri eweri (Lawrence): Fain, 1962 : 21

Type locality: Natal Museum, Pietermaritzburg, South Africa

Distribution: DR Congo, Egypt, Italy, Morocco, Rwanda, South Africa

Hosts: Bufo regularis, B. mauretanicus, B. carens, B. kisoloensis, B. superciliaris, Bufotes v. viridis (Anura: Bufonidae)

Lawrencarus (L.) eweri thailandiae Fain

Lawrencarus eweri thailandiae Fain, 1961a: 252; 1962: 24

Type locality: Pak Shong

Distribution: Thailand

Host: Bufo melanostictus (Anura: Bufonidae)

Lawrencarus (L.) hylae Fain

Lawrencarus hylae Fain, 1961a: 247; Domrow, 1974: 18; 1991: 1293

Lawrencarus (L.) hylae hylae Fain

Lawrencarus hylae hylae : Fain, 1962: 33

Type locality: Queensland

Distribution: Australia, North America, Brazil

Hosts: Hyla coerulea, H. hayi, H. nasuta (Anura: Hylidae)

Lawrencarus (L.) hylae afrixali Fain

Lawrencarus afrixali Fain, 1961a: 251

Lawrencarus hylae afrixali : Fain, 1962: 37

Type locality: Virunga Nat. Park (formerly Albert N.P.)

Distribution: DR Congo

Hosts: Afrixalus fulvovittatus leptosomus, Hyperolius castaneus, Phlyctimanus verrucosus (Anura: Hyperoliidae)

Lawrencarus hylae intermedius Fain

Lawrencarus hylae intermedius Fain, 1962: 39; Mauri & Bischoff, 1984: 338

Type locality: Alto da Serra, Cubatan, Sao Paolo Prov. (Brazil)

Distribution: Argentina, Brazil

Hosts: Hyla c. cinerea, H. hayii (Anura: Hylidae), Pleurodema sp. (Anura: Leptodactylidae)

Lawrencarus hylae brevicipitis Fain

Lawrencarus hylae brevicipitis Fain, 1962: 41

Type locality: Amari, Usumbara (DR Congo)

Distribution: DR Congo, India

Hosts: Probreviceps macrodactylus, Ramanella triangularis (Anura: Microhylidae, Microhylinae, Brevicipitinae)

Lawrencarus lechriodi Fain

Lawrencarus lechriodi Fain, 1962: 31

Type locality: New Guinea

Distribution: New Guinea

Host: Lechriodus platyceps (Anura: Ceratophryidae, Cycloraninae)

Tribe Xenopacarini Fain, Baker et Tinsley

Xenopacarini Fain, Baker & Tinsley, 1969: 341

Type genus: Xenopacarus Fain, Baker et Tinsley, 1969

Genus Xenopacarus Fain, Baker et Tinsley

Xenopacarus Fain, Baker & Tinsley, 1969: 341

Type species: Xenopacarus africanus Fain, Baker et Tinsley, 1969.

Xenopacarus africanus Fain, Baker et Tinsley

Xenopacarus africanus Fain, Baker & Tinsley, 1969: 341

Type locality: South Africa (imported into Leeds, England)

Distribution: South Africa

Host: Xenopus laevis (Amphibia: Pipidae, Xenopodinae)

Xenopacarus kenyensis Fain et Tinsley

Xenopacarus kenyensis Fain & Tinsley, 1975: 854

Type locality: Kiambu, Nairobi; Marsabit

Distribution: Kenya

Host: Xenopus borealis (Amphibia: Pipidae, Xenopodinae)

Xenopacarus kivuensis Fain et Tinsley

Xenopacarus kivuensis Fain & Tinsley, 1993: 513

Type locality: Ebisha, about 8 km from R.St. Irangi, Kivu Prov.

Distribution: D.R. Congo

Host: Xenopus sp. group fraseri (Amphibia: Pipidae, Xenopodinae)

Distribution of Ereynetidae

EUROPE

Armenia (Arutyunyan, 1972, 1981) – Boydaia sturni, Riccardoella limacum,

Austria (Willmann, 1951; Franz, 1954; Sixl, 1972) – Ereynetes papilosus, E. biscutatus, Paraspeleognathopsis bakeri, Riccardoella oudemansi

Belgium (Cooreman, 1947; Fain, 1958b, 1962 , 1963c; Fain & Bafort, 1973; Fain & Lukoschus, 1968: 85; Fain & van Goethem, 1986; André & Ducarme, 2003; André et al., 2004) – Ereynetes aurantipes, E. gandensis, E. hydrophilus, Aureliania aureliani, Boydaia loxiae, B. nigra motacillae, B. n. nigra, B. sturni, Hanriccardoella faini, Paraspeleognathopsis bakeri, P. strandtmanni, Pseudotydeus lebruni, Pterniboydaia pternistis, Riccardoella oudemansi, R. limacum

Bulgaria (Turk, 1970; Beron, 1973) – Ereynetes (Anereynetes) simplex, Paraspeleognathopsis bakeri, Riccardoella limacum, R. oudemansi

Croatia (Willmann, 1941; Pernek et al., 2008) – Ereynetes scutulis, Riccardoella oudemansi

Czech Rep. (Willmann, 1936; Zacharda, 1978; André et al., 2004;) – Ereynetes (Anereynetes) simplex, Riccardoella oudemansi, R. zadielensis

Yan Mayen I. (Macfadyen, 1954) – Ereynetes [Protereunetes] brevipes

Finland (Penttinen, Viiri & Moser, 2013) – Ereynetes propescutulis

France (contin.) (Megnin, 1889; Fain & van Goethem, 1986; Morand & Deguzan, 1986) – Riccardoella reaumuri, R. limacum

Germany (Willmann, 1952, 1953; Fritsch, 1962; Schmidt, 1987) – Boydaia willmanni, Ereynetes bipilosus, E. insularis, E. lyratus

Great Britain (Turk & Phillips, 1945; White, 1959; Baker, 1970a, b; Fain & van Goethem, 1986; Graham et al., 1993) – Riccardoella limacum, R. oudemansi,

Greece (Haralabidis et al., 1983) - Riccardoella limacum

Holland (Oudemans, 1912, 1928; Thor, 1932; Van Eyndhoven, 1955; Fain & Lukoschus, 1968: 85; OConnor, 1978; Mienis, 1984; Fain & van Goethem, 1986) – Boydaia saxicolae, Ereynetes sittardiensis, Paraspeleognathopsis bakeri, Ricardoella oudemansi, Speleorodens michigensis (= clethrionomys)

Hungary (András & Jenö, 2004; Varga & Kontschan, 2004) - Riccardoella

oudemansi

Iceland (Sellnick, 1940) – Riccardoella oudemansi

Ireland (Fauna Europaea) – 1 sp.

Italy (contin.)(Berlese, 1923; Bottazzi, 1950; Fain, 1964b, 2004) – Ereynetes arcuatus, E. biscutatus, E. brevipes, E. centralis, E. inermis, E. ministralis, E. pegazzanoae, E. septentrionalis, Protereunetes brevipes, Riccardoella limacum, R. oudemansi

Sicily (Berlese, 1923) – Ereynetes agilis, E. galeatus

N. Macedonia (Haralabidis et al., 1983) – Riccardoella limacum

Norway (Thor, 1932) – Ereynetes norvegicus

Poland (Karbarz-Wiktorowicz, 1973; Dończyk, 2006) – Ereynetes (Opsereynetes) norvegicus, Ereynetes (Gymnereynetes), Riccardoella limacum

Russia (all, continent.)(Dubinin, 1957; Zabludovskaya, 1985, 1986, 1990a, 1990b, 1991, 1992, 1998, 2012) – Boydaia sturni, B. bradornis, B. jordani, B. psalidoprocnei, Coboydaia nigra nigra, C. nigra motacillae, Neoboydaia phylomachi eroliae, Ophthalmognathus schoutedeni (= dogieli), Phoenicopteriella womersleyi, Lawrencarus eweri dahuricus

Crimea (Zabludovskaya, 1994) - Ereynetes (E.) g. galeatus, E. (G.) exilis, E. (Gymnereynetes) aurantipes, E. (E.) tshernogoriensis

Serbia (Stojnić et al., 2016) – Riccardoella limacum, R. oudemansi

Slovakia (André et al., 2004; Shoaib & Cagáň, 2004) – Riccardoella zadielensis

Spain (cont.) (Mihelčič, 1958; Fain & Lukoschus, 1968: 85) – Ereynetes (Anereynetes) corticalis, E. (A.) ornatus, E. (A.) proximus, Paraspeleognathopsis bakeri

Sweden (Moser et al., 1989) – Ereynetes propescutulis

Switzerland (Schweizer, 1922, 1963) – Ereynetes ministralis, Riccardoella limacum

Turkey (all) (Çobanoğlu, Cilbircioğlu & Zabludovskaya, 2020) – Ereynetes septentrionalis

Ukraine (Zabludovskaya, 1994, 1995, 2002, 2005, 2012) – Boydaia psalidoprocnei, Coboydaia nigra, Ereynetes (E.) gandensis, E. (E.) tshernogoriensis, E. (E.) pegazzanoe, E. (E.) punctatus, E.(Anereynetes) hydrophilus, E. (A.) sittardiensis, E. (Gymnereynetes) aurantipes, E. (G.) exilis, Paraspeleognathopsis bakeri, Riccardoella canadensis

AFRICA

Botswana (Fain, 1959, 1960, 1969; Fain & Camerik, 1994; Skoracki et al., 2018) – Boydaia spatulata, Coboydaia nigra, Ereynetes copridis, E. botswanensis

Burundi (Fain, 1963 ; Skoracki et al., 2018) - Coboydaia clavata, C. nigra

Cameroon (Fain, Herin & Puylaert, 1977; Zabludovskaya, 2002) – Boydaia hirundoae, B. pycnonoti, Coboydaia clavata, C. nigra

Chad (Fain, 1963) – Boydaia falconis

Congo DR (Fain, 1961a, 1964 ; 1966: 108; 1970b; Fain & Tinsley, 1993; Skoracki et al., 2018) – Hipposideroptes kimuenzae, Neoboydaia galachrysiae, Neospeleognathopsis chiropteri, Xenopacarus kivuensis

Egypt (Fain, 1963c, 1966: 108; Lufty, 1960; Abdelgayed et al., 2019) – Ereynetes bolitophilus, Lawrencarus eweri eweri, Neospeleognathopsis chiropteri

Ghana (Skoracki et al., 2018) - Coboydaia clavata

Guinea (Zabludovskaya, 2002) – Coboydaia clavata, C. nigra, Pycnonotus barbatus

Kenya (Fain & Tinsley, 1975; Skoracki et al., 2018) – Boydaia pycnonoti, Xenopacarus kenyensis

Rwanda (Cooreman, 1954; Fain, 1955b, 1956a, 1956b, 1956d, 1962, 1963, 1969, 1971, 1972, 1985a, 1985b; Pence & Casto, 1976, Fain & Goff, 1980; Skoracki et al., 2018) – Astrida caprimulgi, Boydaia bradornis, B. buphagi, B. cecropis, B. cinnyris, B. duboisi, B. falconis, B. psalidoprocnei, B. saxicolae, B. spatulata, B. zosteropis, B. zumpti, Coboydaia clavata, C. nigra, Meropiboydaia merops, Neastrida poffei, Neoboydaia philomachi, Ophthalmognathus tenorioae, Phoenicopteriella womersleyi, Pterniboydaia pternistis, Speleorodens strandtmanni

Senegal (Fain, 1963) – Coboydaia clavata

South Africa (Fain, 1956, 1958, 1959, 1963c, 1971; Fain, Baker & Tinsley, 1969; Fain, Herin & Puylaert, 1977; Fain & Camerik, 1994; Ueckermann & Tiedt, 2003; Skoracki et al., 2018) – Boydaia pycnonoti, Ereynetes aplectorus, E. bolitophilus, E. luriei, E. ueckermanni, Lawrencarus eweri eweri, Coboydaia nigra, Neastrida parrae, Neoboydaia psittaculae, Riccardoella limacum, R. oudemansi, Xenopacarus africanus

Zambia (Fain, 1960 , 1963) – Coboydaia clavata, Pterniboydaia pternistis

Zimbabwe (Skoracki et al., 2018) - Boydaia pycnonoti, Coboydaia clavata

ASIA

China (Li, Xuan & Fan, 1992;) – Ereynetes sp.

India (Fain, 1969; Bhat, Kulkarni & Mishra, 1983; Panda, Mishra & Panda, 1987) – Boydaia indica, Speleognathopsis galli

Indonesia (Berlese, 1910; Vitzthum, 1932, Fain, 1961 , 1963) - Batracarus hylaranae, Coboydaia amandavae [Java], Ereynetes cruciatus [Java], E. potator [Sumatra]

Iran (Siahkamari et al., 2020) – Ereynetes [Ereynetoides] amplectorus

Israel and Palestina (Mienis, 1990) – Riccardoella sp.

Japan (Berlese, 1923; Kaneko, 1968; Shiba, 1969а, 1971; Kaneko, Kadosaka & Toshioka, 1977; Kaneko, Matsudaira & Prince Masahito, 1978; Uchikawa & Suzuki, 1984; Kadosaka, Kaneko & Asanuma, 1983, 1987; Waki, Hiruta & Shimano, 2018; Waki et al., 2018; Waki & Shimano, 2019; Waki, Shimano & Asami, 2019; Waki, Shimano & Shiba, 2019; Waki et al., 2020) – Ereynetes cavernarum, E. excellens, E. (Opsereynetes) norvegicus, Speleognathopsis (Speleomyotis) bastini, Paraspeleognathopsis bakeri, Pterniboydaia pternistis, Riccardoella (Proriccardoella) triodopsis, R. tokyoensis, Trispeleognathus womersleyi

Okinawa (Waki et al., 2018) - Riccardoella reaumuri

Kazakhstan (Zabludovskaya, 1998) - Coboydaia nigra

Kyrghyzstan (Zabludovskaya, 2002) - Coboydaia nigra motacillae

Korea (Fain & Lukoschus, 1968; Shiba, 1969b) – Paraspeleognathopsis bakeri

Malaysia (Fain, 1963c, Fain & Nadchatram, 1962; Shiba, 1976; McClure, Ratanaworadhan et al., 1973) – Astrida caprimulgi, Boydaia pycnonoti, B. spatulata, Ereynetes husatoi, E. malayi, E. tadakii

New Guinea (Fain, 1962) - Lawrencarus lechriodi

Philipppines (Corpuz-Raros, 1975) – Ereynetes philippinensis, Ereynetes (Anereynetes) upelbensis

Turkmenistan (Zabludovskaya, 1998) – Coboydaia nigra

Thailand (Fain, 1961a) – Lawrencarus eweri thailandiae

NORTH AMERICA

Canada (Fain & van Goethem, 1986; Knee, Proctor & Galloway, 2008) – Boydaia aratingae, B. faini, B. psalidoprocnei, B. sturni, Neoboydaia colymbiformi, Riccardoella canadensis,

Mexico (Baker, 1945, Clark, 1960; Fain, 1963c; Fain & Hyland, 1970; Polaco & Mendl, 1988; Vásquez-Rojas & Estrada-Venegas, 2010) – Boydaia mimi, Ereynetes (Opsereynetes) robustus, E. (O.) simplexus, E. (Opsereynetes) tuberculatus, E. (Anereynetes) sabinensis, E. (Ereynetes) amplectorus, E. (E.) faini, Riccardoella oudemansi, Trispeleognathus striatus

USA (Boyd, 1948, 1951; Clark, 1958, 1960, 1964, 1967a, 1967b; Hunter, 1964; McDaniel, 1967; Hunter & Cross, 1968; Kethley, 1971; Pence, 1973a, 1973b, 1973c; Baker & Delfinado,1974; Pence & Casto, 1976, Spicer, 1977; Fain & Klompen, 1990; Hilario-Pérez & Danel, 2016; Hilario-Pérez & Dowling, 2018) – Astrida caprimulgi, A. coccyzae, Boydaia agelaii, B. colini, B. hirundoae, B. mimi, B. pheucticola, B. psalidoprocnei, B. quiscali, B. spatulata, B. sturnellae, B. sturni, B. turdi, Coboydaia nigra, Ereynetes amplectorus, E. boharti, E. davisi, E. scutulis; Hydranetes citrinensis, H. tropisternus; Neoboydaia philomachi, Ophthalmognathus womersleyi, Pseudotydeus perplexus, Riccardoella (Proriccardoella) triodopsis, Ricardoella (R.) limacum, Trispeleognathus striatus, Speleognathopsis benoiti

Hawai Is. (Fain & Goff, 1980) – Coboydaia nigra, Ophthalmognathus tenorioae,

SOUTH and CENTRAL AMERICA

Argentina (Rosas Costa, 1955; Mauri & Bischoff, 1984) – Aureliania longipilis,

Lawrencarus brasiliensis desantisi, L. hylae intermedius

Brazil (Amaral, 1963, 1965, 1967; Fain, 1962, 1963 ; Fain & Aitken, 1969, 1970; Faccini et al., 1981; Flechtmann & Baggio, 1985; Mascarenhas & Brum, 2006; Bernardon et al., 2015; Bernardon, Mascarenhas et al., 2018; Mendoza – Roldan, 2019) – Boydaia agelaii, B. americana, B. aratingae, B. cyanerpes, B. laticoxa, B. psittaci, B. quiscali, B. synallaxis, B. tyrannis, Coboydaia trochila, Lawrencarus brasiliensis, Neoboydaia lateralli, Speleochir barbulata, S. brasiliensis, Speleomys neotropicalis, Neospeleognathopsis (Speleomyotis) bastini didelphis, Riccardoella limacum, Speleognathopsis galli, Trispeleognathus euricoi, T. striatus

Cuba (Dusbábek & Černy, 1970; de la Cruz, 1971) – Boydaia faini, B. morenoi, B. rosickyi, Lawrencarus (L.) eweri cubanus, L. (Cernycarus) hollandsae

Panama (Clark, 1967a) - Speleorodens strandtmanni

Puerto Rico (Hunter & Poe, 1971) – Ereynetes coenobitus

Suriname (Fain & Lukoschus, 1971, 1972) – Boydaia agelaii, Hipposideroptes saccopteryx, Neospeleognathopsis (Speleomyotis) bastini bastini, N. (S.) molossus, Speleochiroides carollia, Psittaboydaia amazona

Trinidad and Tobago (Fain & Aitken, 1968) - Boydaia agelaii, B. americana, B. hirundoae, B. tyrannis, Trispeleognathus striatus

OCEANIA

Australia (continent) (Rainbow, 1906; Domrow, 1960, 1962, 1965, 1966, 1967, 1969, 1975, 1991; OConnor, 1978; Fain & Lukoschus, 1968, 1979; Womersley, 1936, 1953, 1954) – Aureliania aureliani, Boydaia spatulata, Domrownetes exul, Meropiboydaia merops, Neoboydaia philomachi, N. psittaculae, Paraspeleognathopsis bakeri, P. exaeta, Speleorodens derricki, Pteropignathus pteropi, Speleognathopsis benoiti, Riccardoella limacum (probably R. oudemansi, see Fain, 2004)

New Zealand (Barker & Ramsay, 1978; Fain & Barker, 2003, 2004) – Austreynetes maudensis, Riccardoella (Riccardoella) limacum, Riccardoella (Proriccardoella) novaezealandiae

Macquarie I. (Fain, 1962d; Luxton, 1990) – Ereynetes (Gymnereynetes) macquariensis

Papua New Guinea (Domrow, 1969; Talbot, 1969; Fain & Van Goethem, 1978) – Ereynetes (Anereynetes) papuanus, Lawrencarus ceratobatrachi, Speleognathopsis galli

ANTARCTICA

Kerguelen I. (Fain, 1976) - Ereynetes macquariensis

Marion I. (Fain, 1976) - Ereynetes macquariensis

Signy I. (Strandmann & Tilbrook, 1968) - Ereynetes macquariensis

Antarctic Peninsula (Strandtmann, 1967; Crooker, 1977; Llano & Gless, 2013) - Ereynetes macquariensis, Protereunetes minutus, P. paulinae

Hosts of Ereynetidae

Mollusca

Gastropoda

Pulmonata

Fam. Arionidae

Arion ater - Riccardoella oudemansi

A. lusitanicus - Riccardoella oudemansi

A. rufus - Riccardoella oudemansi

Fam. Athoracophoridae

Pseudaneitea schauinslandi - Austreynetes maudensis

Athoracophorus bitentaculatusRiccardoella limacum

Athoracophoridae gen. sp. – Riccadoella (Proriccardoella) novaezealandiae

Fam. Clausiliidae

Tauphaedusa tauRiccardoella tokyoensis

Fam. Bradybaenidae

Acusta despecta sieboldtianaRiccardoella (Proriccardoella) triodopsis

Fam. Helicidae

Cepaea nemoralis - Riccardoella oudemansi

Helix lucorum - Riccadoella limacum

H. pomatia - Riccadoella limacum

H. aspersa - Riccadoella limacum

Fam. Limacidae

Deroceras reticulatum - Riccardoella oudemansi

Limax agrestis - Riccardoella oudemansi

L. cinereoniger - Riccardoella oudemansi

L. cinereus - Riccardoella oudemansi

L. flavus - Riccardoella oudemansi

L. maximus - Riccardoella oudemansi

L. cinereoniger - Riccardoella oudemansi

Monochroma brunneum – Riccardoella limacum

Fam. Milacidae

Tandonia budapestensis - Riccardoella oudemansi

Fam. Polygiridae

Xolotrema obstructa [Triodopsis obstrictus] – Riccardoella (Proriccardoella) triodopsis

Fam. Zonitidae

Meledella werneri – Riccardoella oudemansi

Oxychilus draparnaudi - Riccardoella oudemansi

Phylum Arthropoda

Subphylum Crustacea

Order Decapoda

Infraorder Brachiura

Fam. Coenobitidae

Coenobita clypeatus – Ereynetus coenobitus

Insecta

Order Coleoptera

Fam. Hydrophilidae

Tropisternus lateralis nimbatus - Hydranetes citrinensis

Tropisternus lateralis nimbatus - Hydranetes tropisternus

T. striolatus - Hydranetes tropisternus

T. quadristriatus – Hydranetes tropisternus

Fam. Scarabaeidae

Copris subsidens - Ereynetes (Anereynetes) copridis

C. molossus – Ereynetes cruciatus

Metacatharsius sp. - Ereynetes (Anereynetes) botswanensis

Scarabaeidae indet. – Ereynetes philippinensis

Order Hymenoptera

Fam. Apidae

Melipona scutellaris - Ereynetes (Gymnereynetes) meliponae

Subphylum Vertebrata

Class Amphibia

Order Anura

Fam. Bufonidae

Bufo arenarum - Lawrencarus brasiliensis desantisi

B. melanostictus - Lawrencarus eweri thailandiae

B. regularis - Lawrencarus eweri eweri

B. mauretanicus - Lawrencarus eweri eweri

B. carens - Lawrencarus eweri eweri

B. kisoloensis - Lawrencarus eweri eweri

B. paracnemis – Lawrencarus brasiliensis desantisi

B. peltacephalus - Lawrencarus eweri cubanus

B. raddei - Lawrencarus eweri dahuricus

B. superciliaris - Lawrencarus eweri eweri

Bufotes v. viridis - Lawrencarus eweri eweri

Fam. Ceratophryidae

Cyclorhamphus asper - Lawrencarus brasiliensis

Eleutherodactylus guntheri - Lawrencarus brasiliensis

Eleutherodactylus sp. – Lawrencarus (Cernycarus) hollandsae

Lechriodus platyceps - Lawrencarus lechriodi

Limnodynastes tasmaniensis - Lawrencarus angelae

L. peronii - Lawrencarus angelae

Fam. Hyperoliidae

Afrixalus fulvovittatus leptosomus - Lawrencarus hylae afrixali

Hyperolius castaneus - Lawrencarus hylae afrixali

Phlyctimanus verrucosus - Lawrencarus hylae afrixali

Fam. Hylidae

Hyla cinerea cinerea - Lawrencarus hylae intermedius

H. coerulea - Lawrencarus hylae hylae

H. hayii - Lawrencarus hylae intermedius

H. nasuta - Lawrencarus hylae hylae

H. septentrionalis – Lawrencarus americanus

Fam. Leptodactylidae

Pleurodema sp. - Lawrencarus hylae intermedius

Fam. Microhylidae

Probreviceps macrodactylus – Lawrencarus hylae brevicipitis

Ramanella triangularis - Lawrencarus hylae brevicipitis

Fam. Pipidae

Xenopus borealisXenopacarus kenyensis

X. laevis – Xenopacarus africanus

Xenopus sp. group fraseri – Xenopacarus kivuensis

Fam. Ranidae

Hylarana erythraea - Batracarus hylaranae

Ceratobatrachus guntheri - Lawrencarus ceratobatrachi

Class Aves

Order Anseriformes

Fam. Anatidae

Aythia affinis - Phoenicopteriella womersleyi

Anas acuta – Phoenicopteriella womersleyi

A. clypeata - Phoenicopteriella womersleyi

A. discors – Phoenicopteriella womersleyi

A. strepera – Phoenicopteriella womersleyi

Dendrocygna viduata – Phoenicopteriella womersleyi

Sarkidiornis melanotos – Phoenicopteriella womersleyi

Order Caprimulgiformes

Fam. Caprimulgidae

Antrostomus carolinensis - Astrida caprimulgi

Scotornis fossii welwitschi - Astrida caprimulgi

Fam. Podargidae

Podargus strigoidesBoydaia podargi

Order Charadriiformes

Fam. Charadriidae

Afribyx senegallus - Speleognathopsis benoiti

Charadrius melanops - Speleognathopsis benoiti

Ch. pecuarius – Speleognathopsis charadricola

Erythrogonys cinctus - Speleognathopsis benoiti

Hemiparra crassirostris – Speleognathopsis benoiti

Lobibyx miles - Speleognathopsis benoiti

Philomachus pugnax – Neoboydaia philomachi

Pluvialis squatarola - Speleognathopsis benoiti

Tringa glareola - Neoboydaia philomachi

Fam. Glareolidae

Glareola [Galachrysis] cinerea – Neoboydaia galachrysiae

G. nuchalis - Neoboydaia galachrysiae

Rhinoptilus africanus – Speleognathopsis benoiti (= wai)

Fam. Jacanidae

Actophilornis africanusNeastrida parrae

Fam. Scolopacidae

Actitis macularius - Neoboydaia philomachi

Arenaria interpres - Neoboydaia philomachi

Calidris melanotos - Neoboydaia philomachi

C. minutilla - Neoboydaia philomachi

Gallinago gallinago - Neoboydaia philomachi, Boydaia willmanni

Limnodromus griseus - Neoboydaia philomachi

L. scolopaceus - Neoboydaia philomachi

Limosa lapponica – Neoboydaia philomachi

Neoboydaia philomachi - Neoboydaia philomachi

Philomachus pugnax - Neoboydaia philomachi

Tringa flavipes - Neoboydaia philomachi

T. melanoleuca - Neoboydaia philomachi

T. glareola - Neoboydaia philomachi

T. solitaria - Neoboydaia philomachi

Fam. Laridae

Sterna hirundo - Neoboydaia philomachi

Thalasseus maximus - Neoboydaia philomachi

Order Columbiformes

Fam. Columbidae

Columba livia domesticaTrispeleognathus striatus

Columbigallina passerina Trispeleognathus striatus

Geotrygon montana – Trispeleognathus striatus

Leptotila verreauxi - Trispeleognathus striatus

Streptopelia s. semitorquata – Trispeleognathus striatus

S. vinacea - Coboydaia clavata

Order Coraciiformes

Fam. Coraciidae

Coracias caudatus caudatusMetaboydaia poffei

Fam. Meropidae

Merops apiaster – Meropiboydaia merops

M. ornatus - Meropiboydaia merops

Order Cuculiformes

Fam. Cuculidae

Guira guira - Carcinopodacarus polymorphus

Order Falconiformes

Fam. Accipitridae

Accipiter fasciatusSpeleognathopsis accipitris

Fam. Falconidae

Falco berigora - Boydaia falconis

F. cuvieri – Boydaia falconis

F. t. tinnunculus – Boydaia falconis

Order Galliformes

Fam. Odontophoridae

Callipepla squamata - Boydaia clarki

Colinus virginianusBoydaia colini

Fam. Phasianidae

Bambusicola thoracica – Pterniboydaia pternistis

Gallus gallus domesticusSpeleognathopsis galli

Pternistis afer – Pterniboydaia pternistis

P. swainsoni – Pterniboydaia pternistis

Perdix p. perdix – Pterniboydaia pternistis

Fam. Numididae

Numida meleagris intermedia - Speleognathopsis galli

Order Gruiformes

Fam. Rallidae

Laterallus melanophaius oenops - Neoboydaia lateralli

Order Passeriformes

Fam. Acrocephalidae

Acrocephalus a. arundinaceus - Coboydaia nigra

A. s. scirpaceus - Coboydaia nigra

Iduna similis - Boydaia spatulata

Fam. Buphagidae

Buphagus africanus - Boydaia buphagi

Fam. Campophagidae

Lalage tricolor – Coboydaya nigra (sub Boydaia crassipes)

Fam. Cardinalidae

Spiza americana – Boydaia pheucticola

Fam. Cisticolidae

Schistolais leucopogon reichenowi - Boydaia spatulata

Fam. Dendrocolaptidae

Glyphorynchus spirurus - Boydaia laticoxa

Fam. Emberizidae

Emberiza aureola – Coboydaia nigra

E. citrinella – Coboydaia nigra

E. pusilla – Coboydaia nigra

E. rustica – Coboydaia nigra

E. rutila – Coboydaia nigra

E. spodocephala – Coboydaia nigra

E. tristrami – Coboydaia nigra

Spizella passerina – Coboydaia nigra

Fam. Estrildidae

Amandava amandava - Coboydaia amandavae

A. subflava - Coboydaia amandavae

Erythrura prasina - Coboydaia amandavae

Sporaeginthus subflavus - Coboydaia amandavae

Fam. Formicariidae

Formicarius analis - Boydaia formicarii

F. colma - Boydaia formicarii

Fam. Fringillidae

Carduelis c. carduelis - Coboydaia nigra

Carpodacus erythrinus – Coboydaia nigra

Euphonia violacea - Coboydaia nigra

Fringilla coelebs – Coboydaia nigra

F. montifringilla - Coboydaia nigra

Haemorhous mexicanus - Coboydaia nigra

Loxia curvirostraBoydaia loxiae

Passerina cyaneaBoydaia rosickyi

Serinus burtoni tanganjicae - Coboydaia nigra

S. frontalis - Coboydaia nigra

S. [Poliospiza] gularis – Coboydaia nigra

S. mozambicus - Coboydaia nigra

S. sulphuratus shelleyi - Coboydaia nigra

Fam. Furnariidae

Philydor pyrrhodes - Boydaia laticoxa philydori

Synallaxis rutilans - Boydaia synallaxis

S. gujanensis - Boydaia synallaxis

S. erythrothorax - Boydaia synallaxis

Fam. Hirundinidae

Cecropis abyssinica unitatisBoydaia cecropis

Hirundo abyssinica – Boydaia hirundoae

H. neoxena - Boydaia hirundoae

H. rustica - Boydaia hirundoae

Hylochelidon ariel - Boydaia hirundoae

Progne chalybea - Boydaia hirundoae

P. subis - Boydaia hirundoae

Psalidoprocne albiceps - Boydaia psalidoprocnei

Riparia riparia - Boydaia psalidoprocnei

Stelgidopteryx ruficollis - Boydaia hirundoae

Tachycineta bicolor - Boydaia psalidoprocnei

Fam. Icteridae

Agelaius icterocephalus – Boydaia agelaii

A. phoeniceus - Boydaia agelaii

A. ruficapillus - Boydaia agelaii

Chrysomus ruficapillus - Boydaia agelaii

Icterus galbula - Coboydaia nigra

I. spurius – Coboydaia nigra icteri

Quiscalus quiscula - Boydaia buphagi, B. quiscali

Q. mexicanus – Boydaia sturni

Sturnella magnaBoydaia buphagi, Coboydaia sturnellae

Fam. Laniidae

Corvinella corvina - Coboydaia nigra

Fam. Locustellidae

Locustella luscinioides - Boydaia locustellae

Schoenicola brevirostris – Boydaia zumpti

Fam. Meliphagidae

Cissomela pectoralis - Boydaia spatulata

Myzomela sanguinolentaBoydaia myzomelae

Philemon citreogularis Boydaia spatulata

P. corniculatus - Boydaia spatulata

Fam. Mimidae

Mimus gilvus – Boydaia mimi

M. polyglottos - Boydaia mimi

M. polyglottos orpheus – Boydaia morenoi

Fam. Motacillidae

Anthus spinoletta - Coboydaia nigra

A. trivialis – Coboydaia nigra motacillae

Macronyx croceus - Boydaia spatulata

Motacilla aguimp vidua - Coboydaia nigra motacillae

M. alba - Coboydaia nigra motacillae

M. capensis - Coboydaia nigra motacillae

M. flava - Coboydaia nigra motacillae

Fam. Muscicapidae

Agricola pallidus griseus - Boydaia bradornis

Cercomela tractrac - Boydaia spatulata

Erythropygia [Cercotrichos] hartlaubi – Boydaia spatulata

Luscinia luscinia - Boydaia bradornis

Muscicapa rufigastra - Boydaia spatulata

Pogonocichla stellate ruwenzorii - Boydaia spatulata

Sheppardia polioptera - Boydaia spatulata

Fam. Nectariniidae

Arachnothera l. longirostris - Ereynetes malayi

Chalcomitra senegalensis lamperti (=aequatorialis) - Boydaia cinnyris, B. nectarinia

Cinnyris cupreus cupreus (= septentrionalis) - Boydaia cinnyris, B. nectarinia

Cyanomitra verticalis viridisplendens - Boydaia nectarinia

Fam. Paridae

Melaniparus leucomelas insignis [as Parus niger insignis] – Boydaia spatulata

Fam. Parulidae

Dendroica cerulea - Boydaia faini

Fam. Passeridae

Passer ammodendri - Coboydaia nigra

P. domesticus – Coboydaia nigra

P. griseus - Coboydaia nigra nigra

P. hispaniolensis - Coboydaia nigra

P. montanus - Coboydaia nigra

Fam. Pipridae

Manacus manacus - Boydaia tyrannis

Pipra fasciicauda – Boydaia tyrannis

P. iris - Boydaia tyrannis

P. pipra - Boydaia tyrannis

Fam. Ploceidae

Euplectes hordeaceus – Coboydaia clavata

E. ardens - Coboydaia clavata

E. macroura - Coboydaia clavata

Hypochera amauropteryx - Coboydaia clavata

H. chalybeata - Coboydaia clavata

Othyphantes baglafecht stuhlmanni - Coboydaia clavata

Ploceus baglafecht stuhlmanni - Coboydaia clavata

P. [Textor] cucullatus - Coboydaia clavata

P. [Textor] cucullatus graueri - Coboydaia clavata

P. pelzelni - Coboydaia clavata

P. velatus - Coboydaia clavata

P. vitellinus - Coboydaia clavata

P. xanthops (= P. x. camburni) - Coboydaia clavata

Quelea quelea - Coboydaia clavata

Q. quelea centralis - Coboydaia clavata

Q. quelea aethiopica - Coboydaia clavata

Q. erythrops - Coboydaia clavata

Fam. Pycnonotidae

Bleda syndactyla woosnami – Boydaia zumpti

Euryllas latirostris latirostris - Boydaia zumpti

Pycnonotus barbatus tricolor - Boydaia pycnonoti

P. erythrophthalmus - Boydaia pycnonoti

P. jocosus - Boydaia pycnonoti

P. goiavier personatus - Boydaia pycnonoti

P. leucogenys leucotis - Boydaia pycnonoti

P. leucotis - Boydaia pycnonoti

P. simplex - Boydaia pycnonoti

P. xanthopygus - Boydaia pycnonoti

Fam. Sturnidae

Ampeliceps coronatus – Boydaia sturni

Acridotheres tristis – Boydaia indica, B. sturni

Lamprocolius chloropterusBoydaia buphagi, B. indica

L. chalybaeus - Boydaia indica

Onychognathus walleri elgonensis - Astrida onychognathi

Spreo superbus - Boydaia indica

Sturnopastor contra - Boydaia indica

Sturnus malabaricus – Boydaia indica

S. pagodarum - Boydaia indica

S. vulgaris - Boydaia indica

Fam. Sylviidae

Malurus amabilis – Boydaia maluri

M. melanocephalus - Boydaia maluri

Fam. Thraupidae

Cyanerpes cyanea – Boydaia cyanerpes

Tangara sp. - Boydaia spatulata

Tiaris canorus - Boydaia spatulata

Fam. Turdidae

Oenanthe isabellina - Coboydaia nigra

Saxicola rubetra - Coboydaia clavata

Sialia sialisBoydaia spatulata

Turdus migratoriusBoydaia jordani

T. pilaris - Boydaia jordani (= turdi), B. saxicolae

Fam. Tyrannidae

Empidonax sp. – Boydaia synallaxis

Pyrocephalus rubinua - Boydaia tyrannis

Tyrannus tyrannusBoydaia tyrannis

Fam. Tytiridae

Pachyramphus [=Platypsaris] aglaiae - Boydaia tyrannis

Fam. Viduidae

Vidua chalybeata - Coboydaia clavata

V. chalybeata amauropteryx - Coboydaia clavata

Fam. Zosteropidae

Zosterops lateralis – Boydaia zosteropis

Z. senegalensis stuhlmanni (= scotti) - Boydaia zosteropis

Order Pelecaniformes

Fam. Ardeidae

Ardea cinerea - Ophthalmognathus schoutedeni (= O. dogieli)

Ardeola idae – Ophthalmognathus schoutedeni

Nycticorax caledonicus - Ophthalmognathus schoutedeni

N. nycticorax - Ophthalmognathus schoutedeni, O. tenorioae

N. n. hoactli - Ophthalmognathus tenorioae

Fam. Scopidae

Scopus umbrettaMetaboydaia poffei

Order Piciformes

Fam. Lybiidae

Buccanodon duchaillui - Picinyssus buccanodon

Order Podicipidiformes

Fam. Podicipedidae

Podiceps nigricollis – Neoboydaia colymbiformi

P. ruficollis - Neoboydaia colymbiformi

Podilymbus podiceps - Neoboydaia colymbiformi

Order Psittaciformes

Fam. Psittacidae

Amazona amazonica - Psittaboydaia amazona

Aratinga jandayaBoydaia aratingae

Fam. Psittaculidae

Agapornis roseicollis - Psittaboydaia psittaculae

Psittacula krameri – Psittaboydaia psittaculae

Agapornis roseicollis - Psittaboydaia psittaculae

Trichoglossus haematod forsteniPsittaboydaia trichoglossi

Trichoglossus moluccanus - Psittaboydaia psittaculae

Order Ralliformes

Fam. Rallidae

Latirallus melanophaius oenopsRalliboydaia latiralli

Porphyrio melanotusRalliboydaia porphirionis

Order Strigiformes

Fam. Strigidae

Bubo africanus – Astrida caprimulgi

Glaucidium perlatum – Astrida caprimulgi

Otus senegalensis – Astrida caprimulgi

O. spilocephalus vulpes - Astrida caprimulgi

Fam. Tytonidae

Tyto alba affinis – Aureliana aureliani

T. alba alba - Aureliana aureliani

Class Mammalia

Order Dasiuromorphia

Fam. Dasyuridae

Antechinus stuartii - Domrownetes exul

Order Diprotodontia

Fam. Petauridae

Petaurus breviceps - Paraspeleognathopsis exaeta

Order Didelphimorphia

Fam. Didelphidae

Didelphis marsupialisSpeleomyotis bastini

Order Chiroptera

Fam. Pteropodidae

Eidolon helvum – Neospeleognathopsis chiropteri

Epomophorus labiatus minor – Neospeleognathopsis chiropteri

Pteropus scapulatusPteropignathus pteropus

Rousettus aegyptiacus - Neospeleognathopsis chiropteri

R. aegyptiacus leachi - Neospeleognathopsis chiropteri

Fam. Hipposideridae

Hipposideros commersoni gigas - Hipposideroptes kimuenzae

Fam. Nycteridae

Nycteris sp. – Neospeleochir duboisi

Fam. Phyllostomatidae

Anoura g. geoffroy – Speleochir aitkeni

Artibeus jamaicensis – Speleochir brasiliensis

Carollia perspicillataSpeleochiroides carollia

Phyllostomus hastatusSpeleochir phyllostomi

Vampyrodes carracioloi - Speleochir brasiliensis

Fam. Rhinolophidae

Rhinolophus cornutus – Neospeleognathopsis (Speleomyotis) bastini

Fam. Vespertilionidae

Eptesicus fuscus - Neospeleognathopsis (Speleomyotis) bastini

E. melanopterus - Neospeleognathopsis (Speleomyotis) bastini

Myotis myotisNeospeleognathopsis (Speleomyotis) bastini

M. nigricans - Neospeleognathopsis (Speleomyotis) bastini

Fam. Molossidae

Cynomops planirostris – Neospeleognathopsis (Speleomyotis) molossi

Molossus ater - Neospeleognathopsis (Speleomyotis) molossi

M. molossus - Neospeleognathopsis (Speleomyotis) molossi

Fam. Emballonuridae

Saccopteryx bilineata – Hipposideroptes saccopteryx

S. leptura - Hipposideroptes saccopteryx

Order Rodentia

Fam. Arvicolidae

Microtus agrestis – Speleorodens michigensis (= clethrionomys)

M. pennsilvanicus - Speleorodens michigensis

Myodes [Clethrionomys] gapperi - Speleorodens michigensis

M. glareolus - Speleorodens michigensis (= clethrionomys)

Fam. Cricetidae

Neacomys guianae - Speleorodens cricetidarum

Zygodontomys lasiurus fuscinus - Speleorodens neotropicalis

Fam. Gliridae

Claviglis murinus microtis – Paraspeleognathopsis graphiuri

Fam. Muridae

Murinae

Apodemus agrarius coreae - Paraspeleognathopsis bakeri

A. flavicollis - Paraspeleognathopsis bakeri

A. speciosus – Paraspeleognathopsis bakeri

Arvicanthis abysinicus – Speleomys galliardi

Dasymys incomtus medius – Speleomys galliardi

Hydromys chrysogaster – Speleorodens derricki

Mastomys coucha ugandae - Paraspeleognathopsis bakeri

Melomys cervinipes - Paraspeleognathopsis bakeri

Mus musculus - Speleorodens derricki

Pelomys fallax concolor – Speleomys galliardi

Rattus assimilis - Speleorodens derricki

R. conatus – Speleorodens derricki

R. rattus – Speleorodens derricki

Sylvaemus sylvaticus – Paraspeleognathopsis bakeri

S. sylvaticus callipides – Paraspeleognathopsis bakeri

Gerbillinae

Gerbilliscus boehmi – Speleomys galliardi

Fam. Sciuridae

Funisciurus carruthersi – Speleorodens strandtmanni

Sciurus carolinensis – Speleorodens strandtmanni (= P. sciuri)

S. granatensis - Speleorodens strandtmanni

S. vulgaris - Speleorodens strandtmanni

Tamiscus emini – Speleorodens strandtmanni

Order Primates

Fam. Galagonidae

Galago senegalensis moholi - Paraspeleognathopsis galagoensis

Order Artiodactyla

Fam. Bovidae

Bos taurus – Speleognathus australis (= bovis)

Bison bison - Speleognathus australis

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The Mite Superfamily Paratydeoidea

Paratydeoidea Baker, 1949

The superfamily contains only the family Paratydeidae.

The Mite Family Paratydeidae Baker

Paratydeidae Baker, 1949: 119

Small family of 13 known species of six genera, described by Berlese (1910), Baker (1949, 1950), Theron, Meyer & Ryke (1969), Kuznetzov (1973), Soliman (1974), Kandeel (1992), Kandeel & Hoda (1992), Flechtmann (1992), Seeman & Walter (2000).

Catalogue of Paratydeidae

Family Paratydeidae Baker

Paratydeidae Baker, 1949: 119

Type genus: Paratydeus Baker, 1949

Genus Hexatydeus Soliman

Hexatydeus Soliman, 1974: 198

Type species: Hexatydeus aegyptiacus Soliman, 1974

Hexatydeus aegyptiacus Soliman

Hexatydeus aegyptiacus Soliman, 1974: 198

Type locality: Dakahlia, El Manzala

Distribution: Egypt

Hexatydeus amabilis Kandeel

Hexatydeus amabilis Kandeel, 1992: 6

Type locality: Egypt

Distribution: Egypt

Genus Paratydeus Baker

Paratydeus Baker, 1949: 119

Type species: Paratydeus alexanderi Baker, 1949

Paratydeus alexanderi Baker

Paratydeus alexanderi Baker, 1949: 120

Type locality: intercepted at Brownsville, Texas on twig with bananas from Mexico

Distribution: Mexico

Genus Sacotydeus Theron, Meyer et Ryke

Sacotydeus Theron, Meyer & Ryke, 1969: 698

Type species: Sacotydeus lootsi Theron, Meyer et Ryke, 1969

Sacotydeus lootsi Theron, Meyer et Ryke

Sacotydeus lootsi Theron, Meyer & Ryke, 1969: 699

Type locality: Potchefstroom, Transvaal

Distribution: South Africa

Genus Scolotydaeus Berlese

Scolotydaeus Berlese, 1910: 214

Type species: Scolotydaeus bacillus Berlese, 1910

Neotydeus Baker, 1950: 289. Syn. by Delfinado & Baker, 1974: 207

Type species: Neotydeus ardisanneae Baker, 1950

Scolotydeus ardisannae (Baker)

Neotydeus ardisannae Baker, 1950: 291

Scolotydeus ardisannae (Baker): Syn. by Delfinado & Baker, 1974: 207

Type locality: Sanburn, Johnson Co., Illinois

Distribution: USA

Scolotydaeus bacillus Berlese

Scolotydaeus bacillus Berlese, 1910: 214; Thor, 1932: 90; 1933: 44; Theron, Meyer & Ryke, 1969: 708

Type locality: Palermo (“in agri Panormitani”)

Distribution: Norway, Italy (incl. Sicily)

Scolotydaeus corticicola Flechtmann

Scolotydaeus corticicola Flechtmann, 1992: 299

Type locality: Brazil

Distribution: Brazil

Scolotydaeus simplex Delfinado et Baker

Scolotydaeus simplex Delfinado & Baker, 1974: 207

Type locality: New York, Long Island, Adirondack Oark

Distribution: U.S.A. (New York)

Genus Tanytydeus Theron, Meyer et Ryke

Tanytydeus Theron, Meyer & Ryke, 1969: 703

Type species: Tanytydeus cristatus Theron, Meyer et Ryke

Tanytydeus cristatus Theron, Meyer et Ryke

Tanytydeus cristatus Theron, Meyer & Ryke, 1969: 703

Type locality: Potchefstroom, Transvaal

Distribution: South Africa, Crimea

Tanytydeus kakadu Seeman et Walter

Tanytydeus kakadu Seeman & Walter, 2000: 399

Type locality: NT, Kakadu Nat. Park, Baroalba Gorge, Gubara Walk

Distribution: Australia (Northern Territory)

Tanytydeus lamington Seeman et Walter

Tanytydeus lamington , Seeman & Walter, 2000: 395

Type locality: Lamington Nat. Park, Mt Bithongabel

Distribution: Australia (Queensland)

Tanytydeus neocristatus Kandeel et Hoda

Tanytydeus neocristatus Kandeel & Hoda, 1992: 311

Type locality: Egypt

Distribution: Egypt

Genus Walytydeus Kuznetzov

Walytydeus Kuznetzov, 1973: 12

Type species: Walytydeus tauricus Kuznetzov, 1973

Walytydeus tauricus Kuznetzov

Walytydeus tauricus Kuznetzov, 1973: 13

Type locality: Crimea

Distribution: Crimea

Distribution of Paratydeidae

EUROPE

Crimea (Kuznetzov, 1973) – Tanytydeus cristatus, Walytydeus tauricus

Italy (contin.)(Berlese, 1882)

Sicily (Berlese, 1910) - Scolotydaeus bacillus

Norway (Thor, 1932) - Scolotydaeus bacillus

AFRICA

Egypt (Kandeel, 1992; Kandeel & Hoda, 1992; Soliman, 1974) – Hexatydeus

aegypticus, H. amabilis, Tanytydeus neocristatus

South Africa (Theron, Meyer & Ryke, 1969) – Sacotydeus lootsi, Tanytydeus

cristatus

NORTH AMERICA

Mexico (Baker, 1949) - Paratydeus alexanderi

USA (Baker, 1950; Delfinado & Baker, 1974) - Scolotydaeus (= Neotydeus) ardisannae, S. simplex

SOUTH and CENTRAL AMERICA

Brazil (Flechtmann, 1992) - Scolotydaeus corticicola

OCEANIA

Australia (continent.)(Seeman & Walter, 2000) – Tanytydeus lamington, T. kakadu

Bibliography of Paratydeidae

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  • Kandeel M.M.H. 1992. Revision of family Paratydeidae with the description of Hexatydeus amabilis n. sp. from Egypt (Acari: Actinedida). – Bull. Ent. Soc. Egypt (A.R.E.), 70: 1 - 9.
  • Kandeel M.M.H., F.M. Hoda. 1992. First record of Paratydeidae from Egypt with the description of a new species (Acari: Actinedida). – Agric. Res. Review Cairo, 62: 311 - 316.
  • Kuznetzov N.N. 1973. Mites of the family Paratydeidae (Acariformes, Prostigmata); description of a new genus and species from Crimean material. – Nauchnye doklady vyssh. Shk. Biol. Nauki, 1973(11): 11 - 16.
  • Seeman O.D., D.E. Walter. 2000. A review of the Paratydeidae (Acari: Prostigmata) with descriptions of the first Australian representatives, Tanytydeus lamington sp. nov. and T. kakadu sp.nov. – Acarologia, 40(4), 1999: 393 - 400.
  • Soliman Z.R. 1974. New genus of family Paratydeidae from Egypt. – Bull. Soc. Ent. Egypt, 58: 197- 200.
  • Theron P. D., M. K. P. Meyer, P. A. J. Ryke. 1969. Two new genera of the family Paratydeidae (Acari: Prostigmata) from South African soils. – Acarologia, 11(4): 697 - 710.
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Supercohort Anystides

Cohort Anystina

The cohort includes (Walter et al, in Krantz & Walter, 2009) the superfamilies Anystoidea, Caeculoidea, Adamystoidea, Paratydeoidea, and Pomerantzioidea, all treated in the present volume.

The Mite Superfamily Anystoidea

Anystoidea Cunliffe, 1955: 213; Southcott, 1957: 173

Phalanx Hamatostomata Wainstein, 1965: 77

In this superfamily are included the families Anystidae, Erythracaridae, Teneriffiidae, Pseudocheylidae, and Stigmocheylidae. They are predators.

The Mite Families Anystidae Oudemans and Erythracaridae Oudemans

The Anystids (s.lat.) are fast moving predators, moderately large (500 – 1500 µm), preying upon small mites and insects. Most often they are found on the soil surface or on plants. As early as in 1927 it was shown by Hirst that they can reach the highest mountains (Tibet, Mount Everest, 4950 m).

Acarus baccarum Linnaeus, 1758 and A. salicinus (now Anystis) were the first members of Anystidae to be described. Since the description of the genus Anystis von Heyden, 1826 on the basis of Trombidium cornigerum Hermann, 1804, many people contributed with new descriptions of Anystids: Haller (1882), Berlese (1882, 1886, 1903, 1905, 1923, described the genera Tarsotomus, Erythracarus), Thor (1912) described Tarsolarkus, Banks (1916), Schweizer (1922), Hirst (1927, 1931 - described Anandia), Trägårdh (1905, 1931). Among the early students of Anystidae are also worth mentioning Dugès (1834), C.L. Koch (1836, 1847), Stoll (1886, 1887).

It was Oudemans, who revised the family in 1936, describing nine new genera (Autenriethia, Barellea, Walzia, Scharfenbergia, Snartia, Tencateia, Chaussieria, Bechsteinia, Siblyia) and new species and dividing the family into two subfamilies Anystinae and Erythracarinae, the only recognized ever since.

After 1936 the most important contributions were the partial revision of Meyer & Ueckermann (1987) in South Africa (with the descriptions of the new genera Namandia and Paranandia and of 7 new species) and the revision of Erythracarinae of Otto (1999b), of Erythracarus of Otto (1999a), of Chaussieria (Otto, 1999c) and of Tarsotomus and Paratarsotomus (Otto, 1999d), in Australia. Other acarologists who contributed to the system or the biology of Anystidae after 1936 were Evans & Browning, Barilo, Naudo, Willmann (described the genus Absoloniana), Reda, Kuznetsov (described the genus Paratarsotomus), Womersley and others.

Finally. Pepato & Klimov (2015) upgraded the subfamily Erythracarinae on family level.

Composition of Anystidae and Erythracaridae

Fam. Anystidae Oudemans, 1902 - 16 gen., 102 sp. and Fam. Erythracaridae Oudemans – 8 gen., 68 sp.

Subfam. Anystinae Oudemans, 1936 - 8 gen., 34 sp.

Anystis von Heyden, 1826 - 21 sp.

(= Actineda C.L. Koch, 1836)

Autenriethia Oudemans, 1936 - 1 sp.

Barellea Oudemans, 1936 - 1 sp.

Mesoanystis Zacharda, 1985 - 1 sp. (fossil)

Scharfenbergia Oudemans, 1936 - 2 sp.

Snartia Oudemans, 1936 - 1 sp.

Tencateia Oudemans, 1936 - 3 sp.

Walzia Oudemans, 1936 - 5 sp.

Fam. Erythracaridae Oudemans: Pepato & Klimov, 2015 - 8 gen., 68 sp.

Chaussieria Oudemans, 1937 - 10 sp.

Erythracarus Berlese, 1903 - 19 sp.

Erythrocheylus Berlese, 1903 - 2 sp.

Namadia Meyer & Ueckermann, 1987 - 1 sp.

Paratarsotomus Kuznetsov, 1983 – 5 sp.

Siblyia Oudemans, 1936 - 1 sp.

Tarsolarcus Thor, 1912 - 3 sp.

Tarsotomus Berlese, 1882 - 27 sp.

(= Absoloniana Willmann, 1940)

(= Paranandia Meyer еt Ueckermann, 1987)

(= Anandia Hirst, 1927)

(= Parabsolonia Barilo, 1984)

(= Bechsteinia Oudemans, 1936)

(= Chabrieria Oudemans, 1936)

Catalogue of the Families Anystidae and Erythracaridae

Family Anystidae Oudemans

Anystinae Oudemans, 1902:

Anystidae Thor, 1903; Oudemans, 1909; 1936: 364

Erythracaridae Miller, 1925: 97

Type genus: Anystis von Heyden, 1826

Subfamily Anystinae Oudemans

Anystinae: Oudemans, 1936: 383

Type genus: Anystis von Heyden, 1826

Genus Anystis von Heyden

Anystis von Heyden, 1826: 609

Type species: Trombidium cornigerum Hermann, 1804

Actineda C.L. Koch, 1836: 6

Type species: Actineda hilaris C.L. Koch, 1836

Anystis agilis (Banks)

Actineda agilis Banks, 1894: 211; 1895: 432; 1903: 275

Anystis agilis (Banks): Banks, 1907: 598; Ewing, 1909: 432; Miller, 1925: 98; Cuthbertson & Murchie, 2007: 276

Type locality: USA

Distribution: Bermuda Is., Great Britain, Italy, USA

Anystis andrei Oudemans

Anystis andrei Oudemans, 1936: 408

Type locality: Beausoleil, France, 350 m

Distribution: France

Anystis baccarum (Linnaeus)

Acarus baccarum Linnaeus, 1758: 106

Actineda baccarum : Stoll, 1886

Anystis baccarum (L.): Trägårdh, 1905: 63; 1931: 624; Thor, 1912 : 388; Hull, 1918: 29; Vitzthum, 1926: 121; André, 1927: 168; Oudemans, 1906: 85; 1936: 387; Irk, 1939: 154; Willmann, 1939: 13; 1951b: 144; 1954: 236; Irk,1939: 154; Duduch et al., 1940: 54; Evans, 1953: 280; Franz, 1954: 367; Mihelčič, 1958a: 40; 1958b: 278; Shiba, 1969: 101; 2006: 87; Hardman et al., 1885; Smith-Meyer & Ueckermann, 1987: 2 (redescription), Gupta, 2002: 12. Detailed synonymy in Vitzthum, 1926

Anystes baccarum (L.): Irk, 1939; Schweizer, 1951: 66

Actineda coccinea Targioni-Tozzetti, 1878: 258.

Type locality: Sweden

Distribution: Algeria, Australia, Austria, Canada, Chile (Juan Fernandes Is.), China, Egypt, Faroes Is., Germany, Great Britain, Hungary, Iceland, India, Iran, Israel, Italy, Japan, Korea, Madeira, Malaysia, Mexico, New Guinea, Norway, St-Helena, South Africa, Spain, Sweden, Switzerland, Tanzania, Tunisia, Turkey, U.S.A.

Anystis berlesei Oudemans

Anystis berlesei : Oudemans, 1936: 395

Type locality: Italy

Distribution: Italy, St. Helena

Anystis borrusica Oudemans

Anystis borrusica : Oudemans, 1936: 405

Type locality: Königsberg, now Kaliningrad, Russia

Distribution: Russia (Kaliningrad District)

Anystis citreola Oudemans

Anystis flaveola Stoll, 1887 (nec flaveola C.L. Koch, 1836)

Anystis citreola : Oudemans, 1936: 410

Type locality: Guatemala

Distribution: Guatemala

Anystis cornigerum (Hermann)

Trombidium cornigerum Hermann, 1804: 47

Actineda cornigera (Hermann): C.L. Koch, 1836: Heft 17. Considered by Vitzthum (1926) who listed some names of C.L. Koch as synonyms of A. baccarum: Actineda flaveola, A. pini, A. triangularis, A. robuscula, A. rubis, and others. In his revision of Anystidae Oudemans (1936) indicates that “Kochschen “Arten” wahrscheinlich Nymphen sind”(p.365, 368). as synonym of Anystis baccarum.

Erythraeus cornigerus: Dugès, 1834: 16, 44.

Anystis cornigerum (Hermann): von Heyden, 1826. Type of genus Anystis von Heyden, 1826.

Type locality: France

Distribution: France, Great Britain, Italy

Anystis cornutum (Hermann)

Trombidium cornutum Hermann, 1804: 47

Anystis sp.: Thor & Willmann, 1947: 513

Anystis cornutum (Hermann): Otto, 1999a: 895 (with “?”)

Type locality: France

Distribution: France

Anystis cursorium (Gervais)

Trombidium cursorium Gervais, 1844: 187

Anystis cursorium (Gervais):

Type locality: France

Distribution: France

Anystis germanica Oudemans

Anystis germanica : Oudemans, 1936: 403

Type locality: Königsberg, now Kaliningrad, Russia

Distribution: Russia (Kaliningrad District)

Anystis indica Gupta

Anystis indica Gupta, 1992: 144; 2002:

Type locality: India

Distribution: India

Anystis jabanica (Berlese)

Actineda jabanica Berlese, 1905: 156

Anystis jabanica (Berlese): Holm & Wallace, 1989: 77

Type locality: Buitenzorg (Java)

Distribution: Australia, Indonesia (Java)

Anystis lahorense Khan

Anystis lahorense Khan, 1967: 400

Type locality: Lahor

Distribution: Pakistan

Anystis langei Barilo

Anystis langei Barilo, 1984c: 452

Type locality: Zeravshan Range, 20 km south of Samarkand

Distribution: Uzbekistan

Anystis nagalandensis Gupta

Anystis nagalandensis Gupta, 1991: 235; 1992:

Type locality: Botsa, Nagaland

Distribution: India

Anystis pallescens (C.L. Koch)

Actineda pallescens C.L. Koch, 1838: 6

Type locality: Germany

Distribution: Germany, Holland, Italy, Sainte-Helene

Anystis salicinus (Linnaeus)

Acarus salicinus : Linnaeus, 1758: 618

Anystis salicinus (L.): Oudemans, 1936: 393; Smith – Meyer & Ueckermann, 1987: 8; Otto, 1992: 33; Shiba, 2006: 87; Zhang & Hong, 2010: 8

Anystis rosae Oudemans, 1936: 397- Type locality: Arnhem, Holland

Anystis kochi Oudemans, 1936: 399 - Type locality: Prov. Gelderland, Holland

Type locality: (?) Sweden

Distribution: China, France, Germany, Great Britain, Holland, Japan, Sweden; Sainte-Helene, South Africa (introduced)

Anystis sellnicki Oudemans

Anystis sellnicki : Oudemans, 1936: 401

Type locality: Königsberg, now Kaliningrad, Russia

Distribution: Russia (Kaliningrad District)

Anystis venustula (Koch et Berendt)(fossil)

Actineda venustula Koch & Berendt, 1854: Plate XIII, fig. 106

Anystis venustula (Koch et Berendt): Oudemans, 1936: 411

Type locality: Baltic Amber (Paleogene)

Distribution: Baltic Amber (Paleogene)

Anystis vitis (Schrank)

Acarus vitis Schrank, 1781: 519

Actineda vitis (Schrank): Halbert, 1915: 114

Anystis vitis (Schrank): Oudemans, 1936: 464; Bottazzi, 1950: 29; Starkoff & Starkoff, 1950: 95

Type locality: Austria

Distribution: Austria, Ireland, Italy

Anystis voigtsi Oudemans

Anystis voigtsi : Oudemans, 1936: 405

Type locality: Bremen, Germany

Distribution: Germany

Anystis wallacei Otto

Anystis salicinus : Meyer & Ueckermann, 1987: 8, in part; Holm & Wallace, 1989: 79 (nec Acarus salicinus Linnaeus)

Anystis wallacei Otto, 1992: 25; Halliday, 2005: 7; Barbar, 2018

Type locality: France

Distribution: China, France, Portugal, Spain, Morocco, Syria; South Africa, Australia (introduced)

Genus Autenriethia Oudemans

Autenriethia Oudemans, 1936: 425

Type species: Actineda velox Berlese, 1905

Autenriethia velox (Berlese)

Actineda velox : Berlese, 1905: 156

Autenriethia velox (Berlese): Oudemans, 1936: 427

Type locality: Johore, Malaysia

Distribution: Malaysia

Genus Barellea Oudemans

Barellea Oudemans, 1936: 427

Type species: Anystis sinensis Berlese, 1923

Barellea sinensis (Berlese)

Anystis sinensis : Berlese, 1923: 237

Barellea sinensis (Berlese): Oudemans, 1936: 427; Zhang & Hong, 2010: 8

Type locality: Peking

Distribution: China

Genus Mesoanystis Zacharda (fossil)

Mesoanystis Zacharda, 1985: 150

Type species: Mesoanystis taymirensis Zacharda, 1985

Mesoanystis taymirensis Zacharda (fossil)

Mesoanystis taymirensis Zacharda, 1985: 150

Type locality: Taymir Peninsula, Jantardach (Upper Cretaceous amber)

Distribution: Russia (Taymir)

Genus Scharfenbergia Oudemans

Scharfenbergia Oudemans, 1936: 412

Type species: Actineda hilaris C.L.Koch, 1836

Scharfenbergia hilaris (C.L. Koch)

Actineda hilaris : C.L. Koch, 1836: 6

Scharfenberhia hilaris (C.L.Koch): Oudemans, 1936: 412

Type locality: Esslarn, Germany

Distribution: Germany, France

Scharfenbergia gauthieri Oudemans

Scharfenbergia gauthieri Oudemans, 1936: 414

Type locality: Reghaia, Algeria

Distribution: Algeria

Genus Snartia Oudemans

Snartia Oudemans, 1936: 417

Type species: Snartia nepenthus Oudemans, 1936

Snartia nepenthus Oudemans

Snartia nepenthus Oudemans, 1936: 417

Type locality: Haranggaol, Lake Toba, Sumatra

Distribution: Sumatra (Indonesia)

Genus Tencateia Oudemans

Tencateia Oudemans, 1936: 422

Type species: Tencateia besselingi Oudemans, 1936

Tencateia besselingi Oudemans

Tencateia besselingi : Oudemans, 1936: 424; Willmann, 1954: 236; Meyer & Ueckermann, 1987: 14

Type locality: Roosendaal, Noord-Brabant

Distribution: Czech Rep., Holland

Tencateia kanthiensis Gupta

Tencateia kanthiensis Gupta, 1992: 146

Type locality: W. Bengal, Distr. Medinpur, Kanthi

Distribution: India

Tencateia toxopei Oudemans

Tencateia toxopei : Oudemans, 1936: 425; Willmann, 1951a: 147; Shiba, 2006: 87

Type locality: Oisterwijk, Noord-Brabant (Holland)

Distribution: Austria, Holland, Japan

Tencateia villosa Meyer et Ueckermann

Tencateia villosa : Meyer & Ueckermann, 1987: 10

Type locality: South Africa (10 km from Postmasburg, Cape Prov., and many other places)

Distribution: South Africa

Genus Walzia Oudemans

Walzia Oudemans, 1936: 419

Type species: Actineda antiguensis Stoll, 1886

Walzia antiguensis (Stoll)

Actineda antiguensis : Stoll, 1886: 7

Walzia antiguensis (Stoll): Oudemans, 1936: 419

Type locality: Antigua, Guatemala

Distribution: Guatemala, Venezuela

Walzia australica Womersley

Walzia australica : Womersley, 1942: 18; Holm & Wallace, 1989; Su et al., 2006

Type locality: South Australia, Queensland

Distribution: Australia (South Australia, Queensland), China

Walzia darjeelingensis Gupta

Walzia darjeelingensis Gupta, 1992: 148; 2002:

Type locality: West Bengal, Darjeeling Botanical Garden

Distribution: India

Walzia hammeni Meyer et Ueckermann

Walzia hammeni : Meyer & Ueckermann, 1987: 16

Type locality: South Africa, Blyde River Canyon, Transvaal, and many other places

Distribution: South Africa

Walzia indiana Meyer et Ueckermann

Walzia indiana : Meyer & Ueckermann, 1987: 14; Gupta, 2002:

Type locality: Midnapur, West Bengal, India

Distribution: India

Fam. Erythracaridae Oudemans

Subfamilia Erythracarinae Oudemans

Erythracarinae Oudemans, 1936: 427

Erythracaridae: Pepato & Klimov, 2015

Type genus: Erythracarus Oudemans, 1936

Genus Chaussieria Oudemans

Schellenbergia Oudemans, 1936: 433 (praeocc.)

Chaussieria Oudemans, 1937: 662 (replacement name); Meyer & Ueckermann, 1987: 18; Otto, 1999b: 254

Chausseria (misspelling): Willmann, 1951b: 147; Hutterer & Rack, 1977: 421

Type species: Erythraeus domesticus C.L. Koch, 1847

Chaussieria aprica Otto

Chaussieria aprica Otto, 1999b: 258

Type locality: 10 miles south of Larrimah, Northern Territory

Distribution: Australia

Chaussieria berlesei (Oudemans)

Erythraeus parietinus (Hermann): Berlese, 1882: (not Hermann, 1804)

Tarsotomus berlesei Oudemans, 1906: 87

Schellenbergia berlesei (Oudemans): Oudemans, 1936: 434

Chausseria berlesei (Oudemans): Willmann, 1951a: 147

Chaussieria berlesei (Oudemans): Meyer & Ueckermann, 1987: 18

Type locality: Italy

Distribution: Austria, Italy

Chaussieria capensis Meyer et Ryke

Chaussieria capensis Meyer & Ryke, 1960: 185; Ueckermann & Smith Meyer, 1988: 41 (synonymized it with Ch. venustissima, but Otto, 1999b: 264 restored it as full species); López-Campos & Vázquez-Rojas, 2010: 157

Type locality: South Africa, Bathurst

Distribution: South Africa

Chaussieria dissimilis Naudo

Chaussieria dissimilis : Naudo, 1977: 298; Otto, 1999b: 265

Type locality: St. Helena, Prosperous Bay Plain

Distribution: St. Helena

Chaussieria domestica (C.L.Koch)

Erythraeus domesticus : C.L.Koch, 1847: 257

Tarsotomus domesticus : Oudemans, 1921: 358

Schellenbergia domesticus (C.L.Koch): Oudemans, 1936: 433

Schellenbergia berlesei (Oudemans): Oudemans, 1936: 434

Chaussieria domestica (C.L.Koch): Meyer & Ueckermann, 1987: 22; Otto, 1999b: 266; Shiba, 2006: 87

Erythraeus venustissimus : Berlese, 1882: fasc. 2(3); 1888: 3; Canestrini, 1885: 1652; 1886: 164; Schweizer, 1922: 81

Erythracarus venustissimus (Berlese): Type: Sicily. Vitzthum, 1928: 47; Schweizer, 1951: 66; Mihelčič, 1958a: 40; Schweizer & Bader, 1963: 266

Schellenbergia venustissima (Berlese): Oudemans, 1936: 436

Chaussieria venustissima (Berlese): Naudo, 1977: 300; Meyer & Ueckermann, 1987: 20 (in part, European material only); Ueckermann & Smith Meyer, 1988: 41; Otto & Olomski, 1994: 287

Chausseria venustissima (Berlese)(misspelling): Hutterer & Rack, 1976: 434

Tarsotomus berlesei Oudemans, 1905: 87 (syn. by Otto, 1999b: 266)

Chaussieria berlesei (Oudemans): Meyer & Ueckermann, 1987: 18

Chaussieria maritima : Evans & Browning, 1953: 419. Type: Babbacombe, South Devon, England. Meyer & Ueckermann, 1987: 22. Syn. by Otto, 1999b: 266

Type locality: Germany

Distribution: Austria, Germany, Great Britain, Greece (Crete), Holland, Iran, Italy (incl. Sicily), Japan, Russia, Spain, Switzerland, Uzbekistan

Chaussieria pilifera Otto

Chaussieria pilifera Otto, 1999b: 270

Type locality: South Australia, 31 km NW of Renmark

Distribution: Australia (South Australia, Victoria, Western Australia)

Chaussieria pulleni Otto

Chaussieria pulleni Otto, 1999b: 274

Type locality: 79 km NNW of Renmark, South Australia

Distribution: Australia

Chaussieria pumila Otto

Chaussieria pumila Otto, 1999b: 275

Type locality: NSW, 43 km E of Balranald

Distribution: Australia (NSW, Northern Territory, South Australia, Victoria, Western Australia)

Chaussieria sanctaehelenae Naudo

Chaussieria sanctaehelenae : Naudo, 1977: 292; Meyer & Ueckermann, 1987: 22; Otto, 1999b: 282

Chaussieria brevis Naudo, 1977: 298; Meyer & Ueckermann, 1987: 22. Syn. by Otto, 1999b: 283

Chaussieria benoiti Naudo, 1977: 297; Meyer & Ueckermann, 1987: 22.

Type locality: Prosperous Bay Plain

Distribution: St. Helena

Chaussieria warregense (Hirst)

Tarsotomus warregense : Hirst, 1931: 562; Oudemans, 1936: 442

Schellenbergia warregense (Hirst): Oudemans, 1936: 442; Womersley, 1942: 20

Chaussieria warregense (Hirst): Naudo, 1977: 300; Meyer & Ueckermann, 1987: 18; Holm & Wallace, 1989; Otto, 1999b: 286

Type locality: Barrigun, near Warrego River, New South Wales

Distribution: Australia (New South Wales, ACT, Northern Territory, South Australia, Western Australia, Tasmania, Queensland)

Genus Erythracarus Berlese

Erythracarus Berlese, 1903: 26

Type species: Trombidium ruricola (not parietinum Hermann, 1804, fide Otto, 1999a)

Palaeoerythracarus Zacharda, 1985: 150 – syn. by Otto, 2000: 457

Type species: Palaeoerythracarus sachalinensis Zacharda, 1985

Bechsteinia Oudemans, 1936: 436 – syn. by Otto, 1999a: 828

Chabrieria Oudemans, 1936: 446 – syn. by Otto, 1999a: 828

Type species of Bechsteinia Oudemans: Bechsteinia schneideri Oudemans, 1936

Type species of Chabrieria Oudemans: Tarsotomus terminalis Banks, 1916

Erythracarus amnicolus Otto

Erythracarus amnicolus Otto, 1999a: 860

Type locality: 67 km S of Menindee, N.S.W.

Distribution: Australia (N.S.W., Western Australia)

Erythracarus avius Otto

Erythracarus avius Otto, 1999a: 864

Type locality: Tasmania, Peters Link Road

Distribution: Australia (Tasmania)

Erythracarus barbarus Otto

Erythracarus barbarus Otto, 1999a: 865

Type locality: McNary (Arizona)

Distribution: USA (Arizona)

Erythracarus ciliatus Otto

Erythracarus ciliatus Otto, 1999a: 868

Type locality: 5 km W of Cowangie, Victoria

Distribution: Australia (Vtoria, Western Australia)

Erythracarus decoris Otto

Erythracarus decoris Otto, 1999a: 831

Type locality: Bethungra, N.S.W.

Distribution: Australia (N.S.W., A.C.T., Victoria, Northern Territory, Western Australia)

Erythracarus elegans Otto

Erythracarus elegans Otto, 1999a: 835

Type locality: 13 km NW of Goulbourn, N.S.W., road to Crookwell

Distribution: Australia (N.S.W., A.C.T., Victoria)

Erythracarus epigeus (C.L. Koch)

Erythraeus epigeus C.L. Koch, 1837: 16.24; 1842: 53

Erythracarus ? epigeus (C.L. Koch): Oudemans, 1936: 431

Erythracarus epigeus (C.L. Koch): Otto, 1999a: 876

Type locality: along the river Danube near Regensburg

Distribution: Germany

Erythracarus festinus (Meyer et Ueckermann)

Bechsteinia festina : Meyer & Ueckermann, 1987: 28

Erythracarus festinus (Meyer et Ueckermann): Otto, 1999a: 840

Type locality: Transvaal, near Golela, Pongola River

Distribution: South Africa

Erythracarus flavipes Otto

Erythracarus flavipes Otto, 1999a: 878

Type locality: A.C.T., Canberra, Mount Ainslie

Distribution: Australia

Erythracarus grahami (Meyer et Ryke)

Bechsteinia grahami : Meyer & Ryke, 1960: 184; Meyer & Ueckermann, 1987: 28

Erythracarus grahami (Meyer et Ryke): Otto, 1999a: 883

Type locality: Bathurst, near Albany, Cape Province, South Africa

Distribution: Angola, South Africa

Erythracarus ? hirsutus (Menge) (fossil)

Erythraeus hirsutus Menge, 1854: 107

Erythracarus ? hirsutus (Menge): Oudemans, 1936: 432; Otto, 1999a: 884

Type locality: amber

Distribution: amber

Erythracarus ? lagopus (Menge) (fossil)

Erythraeus lagopus Menge, 1854: 107

Erythracarus ? lagopus (Menge): Oudemans, 1936: 432; Otto, 1999a: 885

Type locality: amber

Distribution: amber

Erythracarus katherinae Otto

Erythracarus katherinae Otto, 1999a: 841

Type locality: Burrewarra Point, N.S.W.

Distribution: Australia, incl. Tasmania

Erythracarus longisetae Reda

Bechsteinia longisetae (Reda), 1991: 728

Erythracarus longisetae (Reda): Otto, 1999a:

Type locality: Egypt

Distribution: Egypt

Erythracarus nasutus Otto

Bechsteinia festina : Meyer & Ueckermann, 1987: 28 (in part, misidentification)

Erythracarus nasutus Otto, 1999a: 844; Lopez-Campos & Vazquez-Rojas, 2010: 157; de Oliveira Bernardi et al., 2010: 119

Type locality: Transvaal, Loskop Dam

Distribution: Brazil, Mexico, South Africa

Erythracarus nobarii (Reda)

Bechsteinia nobarii Reda, 1991: 729

Type locality: Egypt

Distribution: Egypt

Erythracarus parietinus (Hermann)

Trombidium parietinum : Hermann, 1804: 37

Erythraeus parietinus Latr.: Haller, 1882: 314

Erythraeus parietinus (Hermann): C.L. Koch, 1837: 16.23; 1842: 53 + figure 25

Tarsotomus parietinus (Herm.): Oudemans, 1906: 85

Erythracarus parietinus (Herm.): Thor, 1912 : 388; Schweizer, 1951: 66; Franz, 1954: 367; Bottazzi, 1950: 29; Mihelčič, 1958a: 40; 1958b: 278

Erythraeus parietinus Herm.: Hull, 1918: 29

Erythracarus parietinum (Hermann): Oudemans, 1936: 429

Erythraeus comes Berlese, 1886: 6; syn. by Oudemans, 1906: 87 (as Tarsotomus comes)

Tarsotomus comes (Berlese): André, 1927: 169

Erythracarus comes (Berlese): Berlese, 1903: 26; 1920: 8; Starkoff & Starkoff, 1950: 96

Anystis parietinum : Oudemans, 1936:

Bechsteinia solarii Ueckermann, 1987: 56. Type: Hamburg, Germany, Syn. by Otto, 1999a: 886

Erythraeus spinatus Banks, 1894: 210. Type: New York State, Long Island, Sea Cliff Village. Oregon. Syn. by Otto, 1999a: 886

Type locality of E. parietinus: France

Distribution: Algeria, Austria, France, Germany, Great Britain, Holland, Italy, Norway, Spain, Switzerland, Ukraine, USA

Erythracarus proavus (Menge)(fossil)

Erythraeus proavus Menge, 1854: 107

Type locality: amber

Distribution: amber

Erythracarus pyrrholeucus (Hermann)

Trombidium pyrrholeucum Hermann, 1804: 58

Erythraeus ruricola Dugès, 1834: 40; C.L. Koch, 1837: 1.5, 1842: 53; Canestrini, 1885: 1650; Berlese, 1887: No2,3. Synonymy by Oudemans, 1936: 432

Erythracarus ruricola (Dugès): Berlese, 1903: 26; Oudemans, 1936: 432; Schweizer, 1951: 66; Mihelčič, 1958a: 40; Schweizer & Bader, 1963: 266 (in part, specimen from the locality ‘Scanf’ only, fide Otto, 1999a: 846)

Erythracarus pyrrholeucus : Oudemans, 1936: 432; Otto, 1999a: 850 (redescription); Barbar, 2018

Tarsotomus terminalis : Banks, 1916: 13. Type: Claremont, California. Syn. by Otto, 1999a: 847

Chabrieria terminalis (Banks): Oudemans, 1936: 446

Bechsteinia schneideri : Type: San Remo, Italy; Oudemans, 1936: 436; Cooreman, 1946: 4000 (Italy, Cyprus); Ueckermann, 1987: 57. Syn. by Otto, 1999a: 847

Bechsteinia californica McGregor, 1956: 23; Ueckermann, 1987: 56. Type: Whittier, California. Syn. by Otto, 1999a: 847

Type locality of Erythracarus pyrrholeucus: France

Distribution: Algeria, Australia, Cyprus, Iran, Italy, France, Spain, Ukraine, Crimea, U.S.A. (California)

Erythracarus raripilus (Menge)(fossil)

Erythraeus raripilus Menge. 1854: 107

Erythracus [sic] raripilus (Menge): Oudemans, 1936: 433

Erythracarus raripilus (Menge): Otto, 1999a: 894

Type locality: amber

Distribution: amber

Erythracarus sachalinensis (Zacharda) (fossil)

Palaeoerythracarus sachalinensis Zacharda, in Zacharda & Krivoluckiy, 1985: 150

Erythracarus sachalinensis (Zacharda)

Type locality: South Sachalin I., Starodubskoe (Paleogene amber)

Distribution: Russia (Sachalin)

Erythracarus tricolor (Lucas)

Erythraeus tricolor Lucas,1846: 311

Erythracarus tricolor (Lucas): Oudemans, 1936: 433; Otto, 1999b: 895 (with “?”)

Type locality: Algeria

Distribution: Algeria

Genus Erythrocheylus Berlese

Erythrocheylus Berlese, 1903: 27

Type species: Pseudocheylus erythraeoides Leonardi, 1901

Erythrocheylus erythraeoides (Leonardi)

Pseudocheylus erythraeoides Leonardi in Berlese & Leonardi, 1901: 110

Erythrocheylus erythraeoides (Leonardi, 1901): Berlese, 1903:

Type locality: Funchal

Distribution: Chile

Erythrocheylus littoralis Berlese

Erythrocheylus littoralis Berlese, 1903: 27

Type locality: Italy

Distribution: Italy

Genus Lacteoscythis Pogrebnyak

Lacteoscythis Pogrebnyak, 1995: 11

Type species: Lacteoscythis arenaria Pogrebnyak, 1995

Lacteoscythis arenaria Pogrebnyak

Lacteoscythis arenaria Pogrebnyak, 1995: 11

Type locality: Black Sea Natural Reserve (south part of Ukraine)

Distribution: Ukraine

Genus Namadia Meyer et Ueckermann

Namadia Meyer & Ueckermann, 1987: 32

Type species: Namadia floreata Meyer et Ueckermann, 1987

Namadia floreata Meyer et Ueckermann

Namadia floreata Meyer & Ueckermann, 1987: 32

Type locality: Gamoep

Distribution: South Africa

Genus Paratarsotomus Kuznetsov

Paratarsotomus Kuznetsov, 1983: 91

Type species: Paratarsotomus scutellatus Kuznetsov, 1983

Paratarsotomus behningii (Thor)

Tarsotomus behningii Thor, 1912a: 471 (the species is named after A. Behning); Oudemans, 1936: 441

Paratarsotomus behningii (Thor): Otto, 1999c: 797

Type locality: “in einem Nebenteich des Salzsees Baskuntschak”, Russia

Distribution: Russia

Paratarsotomus callunae (Oudemans)

Tarsotomus callunae Oudemans, 1936: 438; Meyer & Ueckermann, 1987: 26

Paratarsotomus callunae (Oudemans): Otto, 1999c: 797

Type locality: Bergen-op-Zoom, Noord - Brabant

Distribution: Holland

Paratarsotomus macropalpis (Banks)

Tarsotomus macropalpis : Banks, 1916: 13

Paratarsotomus macropalpis (Banks): Otto, 1999c: 798; Abdelgayed et al., 2019: 31

Type locality: Claremont, California

Distribution: California (USA), Egypt

Paratarsotomus sabulosus (Berlese)

Erythraeus sabulosus Berlese, 1885: No 7; 1886: fa. 29; Schweizer, 1922: 81

Tarsotomus sabulosus (Berlese): Oudemans, 1936: 411

Erythracarus sabulosus (Berl.): Berlese, 1903: 26; Mihelčic, 1958b: 278

Erythracarus sabulosus var. oelandicus Trägårdh, 1915: 126. Oeland Insel. Syn. by Otto, 1999c: 800

Tarsotomus rackae : Meyer & Ueckermann, 1987: 26 -Type locality: Oberweiden, Austria. Syn. by Otto, 1999c: 800

Tarsotomus oelandicus (Trägårdh): Oudemans, 1936: 442. Syn. by Otto, 1999c: 800

Paratarsotomus sabulosus (Berlese): Otto, 1999c: 800; Ueckermann et al., 2006: 255

Type locality: Italy

Distribution: Austria, Germany, Italy, Spain, Sweden, Ukraine, Yemen

Paratarsotomus scutellatus Kuznetsov

Paratarsotomus scutellatus Kuznetsov, 1983: 91; Otto, 1999c: 802; Lopez-Campos & Vazquez-Rojas, 2010: 157

Type locality: Uzbekistan

Distribution: Mexico, Uzbekistan

Genus Pedidromus Otto

Pedidromus Otto, 2000: 458

Type species: Pedidromus peliculus Otto, 2000

Pedidromus agitatus Otto

Pedidromus agitatus Otto, 2000: 459

Type locality: N.S. Wales, 80 km East of Hay

Distribution: Australia (NSW, Northern Territory, Western Australia)

Pedidromus curiosus Otto

Pedidromus curiosus Otto, 2000: 460

Type locality: 12 km SW of Calperum, South Australia

Distribution: Australia

Pedidromus durongensis Otto

Pedidromus durongensis Otto, 2000: 461

Type locality: 8 km NW of Durong, Queensland

Distribution: Australia

Pedidromus peliculus Otto

Pedidromus peliculus Otto, 2000: 462

Type locality: N.S. Wales, 43 km East of Balranald

Distribution: Australia (NSW, South Australia)

Pedidromus pilotrix Otto

Pedidromus pilotrix Otto, 2000: 462

Type locality: 10 km SSW of Warren, NSW

Distribution: Australia

Pedidromus velox Otto

Pedidromus velox Otto, 2000: 464

Type locality: Taplan, South Australia

Distribution: Australia (South Australia, Western Australia, Queensland, Victoria)

Genus Siblyia Oudemans

Siblyia Oudemans, 1936: 445

Type species: Erythraeus ignipes Dugès, 1834

Siblyia ignipes (Dugès)

Erythraeus ignipes : Dugès, 1834: 43

Siblyia ignipes (Dugès): Oudemans, 1936: 445

Type locality: ? Montpellier, France

Distribution: France

Genus Tarsolarkus Thor

Tarsolarkus Thor, 1912a: 466

Tarsolarcus Thor, 1912a: 465

Neotarsolarcus Kuznetsov, 1983: 87. Syn. by Oto, 2000: 470. Type: Neotarsolarcus clypeatus Kuznetsov, 1983, Tadjikistan and Russia.

Type species: Tarsolarkus articulosus Thor, 1912

Tarsolarkus articulosus Thor

Tarsolarkus articulosus : Thor, 1912a: 466; Irk, 1939: 154; Sellnick, 1949: 128; Willmann, 1951b: 147; Hull, 1918: 30; Irk, 1939: 154; Oudemans, 1936: 444; Sellnick, 1949: 128; Franz, 1954: 367; Schmölzer, 1956: 548; Mehl, 1979: 36; Otto, 2000: 470

Tarsolarcus articulosus : Willmann, 1951b: 147

Erythracarus cluozzai Schweizer & Bader, 1963: 265. Type: Switzerland, Val Cluozza, 2100 m; syn. by Otto, 1999a: 896

Tarsolarkus oudemansi Barilo, 1984a: 1006 from Uzbekistan. Syn. by Otto, 2000: 470

Type locality: Norway (many places)

Distribution: Austria, France, Great Britain, Norway, Sweden, Switzerland, Uzbekistan

Tarsolarkus clypeatus (Kuznetsov)

Neotarsolarcus clypeatus Kuznetsov, 1983: 88

Tarsolarkus clypeatus (Kuznetsov): Otto, 2000: 470

Type locality: Russia

Distribution: Russia, Tajikistan

Tarsolarkus longisetus Barilo

Tarsolarkus longisetus Barilo, 1984a: 1004; Otto, 2000: 472

Type locality: 8 km East of Samarkand

Distribution: Uzbekistan

Tarsolarkus sp. – Yemen (Ueckermann et al., 2006)

Genus Tarsotomus Berlese

Tarsotomus Berlese, 1882: 2

Type species: Erythraeus hercules Berlese, 1882

Anandia Hirst, 1927: 445 – syn. by Otto, 1999c: 751

Type species: Anandia alticola Hirst, 1927

Absoloniana Willmann, 1940: 215 – syn. by Otto, 1999c: 751

Type species: Absoloniana diversipes Willmann, 1940

Paranandia Meyer & Ueckermann, 1987: 34 – syn. by Otto, 1999c: 751

Type species: Anandia bathurstensis Meyer et Ryke, 1960

Parabsolonia Barilo, 1984: 1736 – syn. by Otto, 1999c: 751

Type species: Parabsolonia ignicola Barilo, 1984

Tarsotomus abruptus Otto

Tarsotomus abruptus Otto, 1999c: 756

Type locality: 10 km SSW of Warren, NSW

Distribution: Australia (NSW, ACT, Victoria)

Tarsotomus aiolos Otto

Tarsotomus aiolos Otto, 1999c: 758

Type locality: 64 km SE of Cooya Pooya Homestead, Western-Australia

Distribution: Australia (WA)

Tarsotomus aleantis Otto

Tarsotomus aleantis Otto, 1999c: 759

Type locality: Millstream Chichester National Park; Western-Australia

Distribution: Australia (WA)

Tarsotomus alticolus (Hirst)

Anandia alticola : Hirst, 1927: 445; Oudemans, 1936: 445; Meyer & Ryke, 1960: 180; Meyer & Ueckermann, 1987: 22

Tarsotomus alticolus (Hirst): Otto, 1999c: 785

Type locality: “Upper Rombok, Mount Everest, 4950 m”, Tibet

Distribution: Tibet (China)

Tarsotomus ambitus Otto

Tarsotomus ambitus Otto, 1999c: 762

Type locality: Millstream Chichester National Park; Western-Australia

Distribution: Australia (WA)

Tarsotomus amentatus (Barilo)

Absoloniana amentata : Barilo, 1984a: 1009; Meyer & Ueckermann, 1987: 24

Tarsotomus amentatus (Barilo): Otto, 1999c: 786; Ueckermann et al., 2006: 255

Type locality: Dzhambayski Region, Samarkandskaya oblast (Uzbekistan)

Distribution: Uzbekistan, Yemen

Tarsotomus anubis Otto

Tarsotomus anubis Otto, 1999c: 763

Type locality: 8 km W of Underbool, road to Murrayville; Victoria

Distribution: Australia (Victoria, Northern Territory, South Australia)

Tarsotomus argillus Otto

Tarsotomus argillus Otto, 1999c: 767

Type locality: 3 km east of Swan Hill; Victoria

Distribution: Australia (Victoria, NSW, South Australia)

Tarsotomus bathurstensis (Meyer et Ryke)

Anandia bathurstensis : Meyer & Ryke, 1960: 182

Paranandia bathurstensis (Meyer et Ryke): Meyer & Ueckermann, 1987: 35

Tarsotomus bathurstensis (Meyer et Ryke): Otto, 1999c: 751

Type locality: Albany District, Bathurst, Cape Province

Distribution: South Africa

Tarsotomus bestiolus (Barilo)

Absoloniana bestiola Barilo, 1984b: 1734; Meyer & Ueckerman, 1987: 24

Tarsotomus bestiolus (Barilo): Otto, 1999c: 786

Type locality: Uzbekistan

Distribution: Uzbekistan

Tarsotomus colossus Otto

Tarsotomus colossus Otto, 1999c: 769

Type locality: Mount Newman, Balfour Downs; Western-Australia

Distribution: Australia (WA, Western Australia)

Tarsotomus comosus Otto

Tarsotomus comosus Otto, 1999c: 771

Type locality: 10 km SSW of Warren, NSW

Distribution: Australia (NSW)

Tarsotomus desertorus Otto

Tarsotomus desertorus Otto, 1999c: 772

Type locality: 32 km south of Murrayville, road to Nhill; Victoria

Distribution: Australia (Victoria, NSW, Northern Territory, South Australia)

Tarsotomus dianellus Otto

Tarsotomus dianellus Otto, 1999c: 775

Type locality: south of Walpeup, road to Patchewollock; Victoria

Distribution: Australia (Victoria, NSW, South Australia)

Tarsotomus dioxis Otto

Tarsotomus dioxis Otto, 1999c: 777

Type locality: 80 km east of Hay (NSW)

Distribution: Australia (NSW)

Tarsotomus erraticus Banks

Tarsotomus erraticus : Banks, 1912: 441; Oudemans, 1936: 441; Otto, 1999c: 787

Type locality: Springer, New Mexico

Distribution: USA (New Mexico)

Tarsotomus heliophilus Otto

Tarsotomus heliophilus Otto, 1999c: 778

Type locality: 14 km north of Balranald, road to Ivanhoe; New-South-Wales

Distribution: Australia (NSW, Victoria, South Australia, Western Australia)

Tarsotomus hercules (Berlese)

Erythraeus hercules : Berlese, 1882: fa. 2; 1888: 3

Tarsotomus hercules (Berlese): Trägårdh, 1905: 64; Oudemans, 1936: 441; Starkoff & Starkoff, 1950: 96; Cooreman, 1955: 123; Bottazzi, 1950: 30; Mihelčič, 1958a: 40; Ueckermann et al., 2006: 255

Absoloniana diversipes : Willmann, 1940: 215; 1941: 56; Barilo, 1984a: 1007; Meyer & Ueckermann, 1987: 24 -Type locality: Cave Čančarica, Bjelashnica planina, South Bosna. Syn. by Otto, 1999c: 787

Type locality of Erythraeus hercules: Sicily, Acireale

Distribution: Bosnia, Egypt, France, Iran, Italy, Salamin Is. (Greece), Sicily, Spain, Sudan, Switzerland, Ukraine, Uzbekistan, Yemen

Tarsotomus ignicola (Barilo)

Parabsolonia ignicola Barilo, 1984: 1738

Tarsotomus ignicolus (Barilo): Otto, 1999c: 789

Type locality: Uzbekistan

Distribution: Ukraine, Uzbekistan

Tarsotomus nivalis (Schweizer)

Erythraeus hercules var. nivalis Schweizer, 1922: 80

Tarsotomus nivalis : Oudemans, 1936: 441; Bottazzi, 1951: 222

Type locality: Alps (2760 - 3000 m)

Distribution: Italy, Switzerland (Cima del Pezzacul, 2300 m)

Tarsotomus pentagoneus (Meyer et Ueckermann)

Absoloniana pentagonea : Meyer & Ueckermann, 1987: 24

Tarsotomus pentagoneus Otto, 1999c: 789

Type locality: Malelane and Pongola, Transvaal, South Africa

Distribution: South Africa

Tarsotomus primitivus Otto

Tarsotomus primitivus Otto, 1999c: 790

Type locality: 32 km south of Murrayville, road to Nhill; Victoria

Distribution: Australia (Victoria)

Tarsotomus pusillus Otto

Tarsotomus pusillus Otto, 1999c: 779

Type locality: 64 km SE of Cooya Pooya Homestead, Western-Australia

Distribution: Australia (WA)

Tarsotomus pygmaeus Otto

Tarsotomus pygmaeus Otto, 1999c: 781

Type locality: Kurunjini (Hamersley Range) National Park; Western Australia

Distribution: Australia (WA)

Tarsotomus solatus Otto

Tarsotomus solatus Otto, 1999c: 781

Type locality: Uluru National Park, Mount Olga, Northern Territory

Distribution: Australia (NT)

Tarsotomus undulatus (Mahunka et Mahunka-Papp)

Diversipes undulatus Mahunka & Mahunka-Papp, 1993: 326

Tarsotomus undulatus (Mahunka et Mahunka-Papp):

Type locality: Madagascar

Distribution: Madagascar

Tarsotomus velopes Otto

Tarsotomus velopes Otto, 1999d: 783

Type locality: north of Roebourne; Western-Australia

Distribution: Australia (WA)

Tarsotomus sp. – Mexico (López-Campos & Vázquez-Rojas, 2010)

Distribution of Anystidae and Erythracaridae

EUROPE

Austria (Irk, 1939; Willmann, 1951a, 1951b; Hutterer & Rack, 1976; Meyer & Ueckermann, 1987; Otto, 1999b) – Anystis baccarum, Chaussieria berlesei, Ch. domestica (= venustissima), Tarsolarkus articulosus, Paratarsotomus sabulosus (= Tarsotomus rackae), Tencateia toxopei

Belgium (Cooreman, 1943) – Anystis baccarum

Bosna and Herzegovina (Willmann, 1940, 1941) – Tarsotomus hercules (= Absoloniana diversipes)

Bulgaria (Beron, in prep.)

Czech Rep. (Willmann, 1954) – Anystis baccarum, Tencateia besselingi

Denmark (Hallas, 1978) -

Feroes Is. (Trägårdh, 1931) – Anystis baccarum

France (contin.)(Hermann, 1804; Dugès, 1834; Gervais, 1844; Oudemans, 1936; Schmölzer, 1956; Otto, 1992, 1999b) – Anystis andrei, A. cornigerum, A. cornutum, A. cursorium, A. salicinus, A. wallacei, Erythracarus parietinus, Scharfenbergia hilaris, Siblyia ignipes, Tarsolarkus articulosus, Tarsotomus hercules

Germany (C.L. Koch, 1836, 1842, 1847; Haller, 1882; Oudemans, 1936; Ueckermann, 1987; Otto, 1999b) - Anystis baccarum, A. salicinus, A. voigtsi, Chaussieria domestica (= venustissima), Erythracarus epigeus, E. parietinus (= Bechsteinia solarii), Paratarsotomus sabulosus, Scharfenbergia hilaris

Great Britain (Thor, 1912a; Hull, 1918; Evans & Browning, 1953; Otto, 1999b; Cuthbertson, Bell & Murchie, 2003; Cuthbertson & Murchie, 2004a, b, 2007, 2010) – Anystis baccarum, A. salicinus, A. cornigerum, A. cursorium, Chaussieria domestica (= maritima), Erythracarus parietinus, Tarsolarkus articulosus

Greece (Sellnick, 1931) - Anystis baccarum

Crete (Hutterer, 1978) - Chaussieria domestica (= venustissima)

Salamine (Cooreman, 1955) – Tarsotomus hercules

Holland (Oudemans, 1906, 1936) – Anystis salicinus, Chaussieria domestica, Erythracarus parietinus, Tencateia besselingi, Paratarsotomus callunae

Hungary (Dudich et al., 1940; Ripka et al., 2002) – Anystis baccarum

Iceland (Sellnick, 1940) – Anystis baccarum

Ireland (Halbert, 1915) – Anystis vitis

Italy (contin.) (Berlese, 1886; Oudemans, 1936; Starkoff & Starkoff, 1950; Bottazzi, 1950, 1951; Castagnoli & Nannelli, 1995; Otto, 1999b) – Anystis agilis, A. baccarum, A. conigerum, A.. pallescens, A. vitis, Erythracarus parietinus [= comes], E. pyrrholeucus (= E. [Bechsteinia] schneideri), Chaussieria domestica, Tarsotomus hercules, T. nivalis

Sicily (Berlese, 1882; Bottazzi, 1950; Otto, 1999b) – Chaussieria domestica, Erythracarus pyrrholeucus,Tarsotomus hercules

Latvia (Kuznetsov & Petrov, 1984; Petrova, Salmane & Čudare, 2004) - Anystis baccarum

Norway (Thor, 1912a, 1912b, 1912c) – Anystis baccarum, Erythracarus parietinus (= comes), Tarsolarkus articulosus

Portugal (cont.) (Halliday, 2005) – Anystis wallacei

Madeira (Willmann, 1939) – Anystis baccarum

Romania (Călugăr & Ivan, 2016) - Anystidae

Russia (all, continent.)(Thor, 1912; Oudemans, 1936; Lange, Drozdovskij & Bushkovskaya. 1974a, b; Kuznetsov, 1983; Meyer & Ueckermann, 1987) – Anystis borrusica, A. germanica, A. sellnicki (from Kaliningrad District), Chaussieria domestica (= venustissima), Paratarsotomus behningii, Tarsolarkus clypeatus

Crimea (Otto, 1999a) - Erythracarus pyrrholeucus

Sachalin (Zacharda, 1985; Otto, 2000) – Erythracarus [Palaeoerythracarus] sachalinensis

Serbia (Mladenović et al., 2013) – Anystis sp.

Spain (cont.) (Mihelčič, 1958a, 1958b; Otto, 1992) – Anystis baccarum, A. wallacei, Chaussieria domestica (= venustissima), Erythracarus parietinus, E. pyrrholeucus (= ruricola), Tarsotomus hercules

Sweden (Linnaeus, 1758; Trägårdh, 1915; Sellnick, 1949) – Anystis baccarum, A. salicinus, Paratarsotomus sabulosus [Öland, as “var. oelandicus Träg.], Tarsolarkus articulosus

Switzerland (Schweizer, 1922; Schweizer & Bader, 1963) – Anystis baccarum, Chaussieria domestica (= venustissima), Erythracarus parietinus, E. pyrrholeucus (= ruricola), Tarsolarkus articulosus (= Erythracarus cluozzai Schweizer et Bader), Tarsotomus hercules (= T. nivalis)

Turkey (Ülgentürk, Szentkirályi et al. 2013) – Anystis baccarum

Ukraine (Pogrebnyak, 1995; Otto, 1999a, 1999b) – Erythracarus parietinus, E. pyrrholeucus, Lacteoscythis arenaria, Paratarsotomus sabulosus, Tarsotomus hercules, T. ignicolus

AFRICA

Algeria (Lucas, 1946; Berlese, 1886; André, 1927; Oudemans, 1936) – Anystis baccarum, Scharfenbergia gauthieri, Erythracarus parietinus [Tarsotomus comes], E. tricolor

AngolaErythracarus [Bechsteinia] grahami

Egypt (Trägårdh, 1905; Reda, 1991; Reda & Gomma, 1991; Abdelgayed, Abd El-Wahed, Ali & Eraky, 2019) – Anystis baccarum, Paratarsotomus macropalpis, Erythracarus nobarii, Tarsotomus hercules, Erythracarus nobarii, Neoaegyptocus chelicifurcata

Malawi (Meyer & Ueckermann, 1987) – Chaussieria domestica (= venustissima)

Morocco (Otto, 1992) – Anystis wallacei

Reunion (Quilici, Kreiter, Ueckermann & Vincenot, 1997) – 2 sp.

Sainte-Helene (Naudo, 1977; Meyer & Ueckermann, 1987; Otto, 1999b: 182) – Anystis baccarum, A. berlesei, A. kochi, Chaussieria dissimilis, Ch. sanctaehelenae (= brevis, = benoiti)

South Africa (Oudemans, 1936; Meyer, 1973; Meyer & Ueckermann, 1987; Ueckermann, 1987; Ueckermann & Meyer Smith, 1988; Otto, 1999a, 1999b; Halliday, 2005) – Anystis baccarum, A. wallacei (introduced), Erythracarus [Bechsteinia] grahami, Erythracarus festinus (= Bechsteinia festina), Chaussieria domestica (= venustissima), Ch. capensis, Namadia floreata, Tarsotomus bathurstensis, Tarsotomus [Absoloniana] pentagonea, Tencateia villosa, Walzia hammeri

Sudan (Trägårdh, 1905) – Tarsotomus hercules

Tanzania (Evans, 1953) – Anystis baccarum

Tunisia (Kreiter et al., 2002) - Anystis baccarum

ASIA

China (Berlese, 1923; Hirst, 1927; Su et al., 2006; Zeng et al., 2007; Zhang & Hong, 2010) – Anystis baccarum, A. salicinus, A. wallacei,Tarsotomus [Anandia] alticola (Tibet), Barellea sinensis, Walzia australica

Cyprus (Cooreman, 1946) – Erythracarus [Bechsteinia] pyrrholeucus (= schneideri)

India (Meyer & Ueckermann, 1987; Gupta, 1991, 1992, 2002) – Anystis baccarum, A. indica, A. nagalandensis, Tencateia kanthiensis, Walzia derjeelingensis, W. indiana

Indonesia (Berlese, 1905; Oudemans, 1936; Holm & Wallace, 1989;) – Anystis [Actineda] jabanica (Jawa), Snartia nepenthus (Sumatra)

Iran (Otto, 1999a, 1999b) – Erythracarus pyrrholeucus, Chaussieria domestica, Tarsotomus hercules

Israel and PalestinaAnystis baccarum

Japan (Shiba, 1969, 1976, 2006) - Anystis baccarum, A. salicinus, Tencateia toxopei, Chaussieria domestica, Erythracarus [Bechsteinia] sp.

Okinawa (Kishida, 1959) – Erythracarus simku (nomen nudum)

Korea (Lee Kim & Kim, 1995) - Anystis baccarum

Malaysia (Berlese, 1905; Vitzthum, 1926) – Anystis baccarum, Autenriethia velox

Syria (Barbar, 2018) – Anystis wallacei, Erythracarus pyrrholeucus

Tajikistan (Kuznetsov, 1983) - Tarsolarkus clypeatus

Uzbekistan (Barilo, 1984a, 1984b; Otto, 1999b) – Paratarsotomus scutellatus, Chaussieria domestica, Tarsotomus [Absoloniana] amentata, T. [A.] bestiola, T. hercules (= Absoloniana diversipes), T. [Parabsolonia] ignicola, Tarsolarkus longisetus

Yemen (Ueckermann et al., 2006) – Anystis baccarum, Erythracarus sp. near nasutus, E. sp. near parietinus, Paratarsotomus sabulosum Tarsolarkus sp., Tarsotomus amentatus, T. hercules

NORTH AMERICA

Canada (Hardman et al., 1885) - Anystis baccarum

Mexico (Vitzthum, 1933; López- Campos & Vázquez-Rojas, 2010) – Anystis baccarum, Chaussieria capensis, Erythracarus nasutus, Paratarsotomus sculellatus, Tarsotomus sp., new genera 1 and 2

USA (Banks, 1894, 1895, 1907, 1912, 1916; Ewing, 1909; Miller, 1925; McGregor, 1956; Sorensen, Kinn, Doutt & Cate, 1976; Otto, 1999a) – Anystis agilis, A. baccarum, Erythracarus barbarus, E. parietinus, Erythracarus pyrrholeucus [= Tarsotomus [Bechsteinia] terminalis, = Bechsteinia californica], Tarsotomus erraticus, Paratarsotomus macropalpis

SOUTH and CENTRAL AMERICA

Bermuda Isls (Banks, 1903) – Anystis agilis

Brazil (de Oliveira Bernardi et al., 2010) – Erythracarus nasutus

Chile (contin.)(Leonardi, in Berlese & Leonardi, 1901) - Anystidae

Juan Fernandez Is. (Trägårdh, 1931) – Anystis baccarum

Costa Rica (Banks, 1916) - Anystidae

Guatemala (Stoll, 1886, 1887) – Anystis citreola, Walzia antiguensis

Venezuela (Oudemans, 1936) – Walzia antiguensis

OCEANIA

Australia (continent)(Hirst, 1931; Womersley, 1933, 1942; Holm & Wallace, 1989; Otto & Halliday, 1991; Otto, 1992, 1999b,c; Halliday, 1998) – Anystis baccarum, A. jabanica, A. wallacei [intr.], Chaussieria pilicola, Ch. puleni, Ch. pumila, Ch. warregense, Erythracarus amnicolus, E. ciliatus, E. decoris, E. elegans, E. flavipes, E. katherinae, E. pyrrholeucus, Pedidromus agitatus, P. curiosus, P. durongensis, P. peliculus, P. pilotrix, P. velox, Tarsotomus anubis, T. argillus, T. colossus, T. comosus, T. desertorus, T. dianellus, T. dioxis, T. heliophilus, T. primitivus, T. pusillus, T. solatus, Walzia australica

Tasmania (Otto, 1999a,b) – Erythracarus avium, Chaussieria warregense

Papua New Guinea (Canestrini, 1898) – “Actineda vitis

Bibliography on Anystidae and Erythracaridae

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The Mite Family Teneriffiidae Thor

Teneriffiidae Thor, 1911: 179

Recorded first by Thor (1911) from Canary Is. and Paraguay, this small family contains now 26 species of predaceous free-living mites, living from the dunes of Namibia and Somalia and the intertidal zone of Japan to the high altitude in Alps (3207 m) and Himalaya (5050 m). Their generic taxonomy remains subject of discussions, especially after the revision of McDaniel, Morihara & Lewis (1976), in which only the two “initial” genera were recognized for the family Teneriffiidae: Teneriffia Thor, 1911 and Parateneriffia Thor, 1911. All genera, established between 1911 and 1976 by various authors: Neoteneriffiola Hirst, 1924, Heteroteneriffia Hirst, 1925, Austroteneriffia Womersley, 1935, Mesoteneriffia Irk, 1939, as well as the name Teneriffiola Strand, 1911 and the family name Teneriffiolidae, based on it, are considered by the McDaniel et al. (1976) as synonyms. Mesoteneriffiolla Schmölzer, 1956 and “Neoteneriffiolatadjikistanica Wainstein, 1969 were not considered. Other acarologists added to the taxonomy (and sometimes to the confusion) of Teneriffiidae: Vitzthum (1942), Irk (1939a, b), Radford (1950), Schmölzer (1952, 1956), Tibbetts (1958), Besch (1962), Eller & Strandtmann (1963), Ehara (1965), Wainstein (1969), Shiba & Furukawa (1975), and others.

New genera and species described after 1976 by Schmölzer (2002 – Himalteneriffia from Ladakh), Luxton (1993), Yin, Bei & Li (1995 - Sinoteneriffia from China), Luo et al. (1996 – Sinoteneriffia from China), Luo, Yin et al. (2007, Neoteneriffiola from China), Bernardi et al. (2012, Brazil), Ueckermann & Khanjani (2002), Khanjani, Asali Fayaz & Ueckermann (2011), Khanjani, Yazdanpanah & Asali Fayaz (2013), Khanjani, Hoseini & Asali Fayaz (2014), Ueckermann & Durucan (2020, Teneriffia from Turkey), Zmudzinski, Skoracki & Friedrich (2021, Teneriffia from Ethiopia).

Judson (1994, 1995) not only described new and redescribed the older species of Teneriffiidae, but questioned the lumping of several genera by McDaniel et al. (1976) into two original units. Based on very detailed morphological studies, he reviewed and removed from synonymy the genera Neoteneriffiola and Austroteneriffia and described new species from Namibia and Australia.

In our collection in Sofia we have many specimens of Teneriffiidae from Greece, Afghanistan, Chinese Karakorum, Somalia, Nigeria, Tanzania, Zambia, Peru, and Mexico, which represent new species and are being described.

Composition of Teneriffiidae

Fam. Teneriffiidae – 28 sp.

Austroteneriffia Womersley, 1935 - 10

Heteroteneriffia Hirst, 1925 - 2

Mesoteneriffia Irk, 1939 - 1

Mesoteneriffiola Schmölzer, 1956 - 1

Himalteneriffia Schmölzer, 2002 - 1

Neoteneriffiola Hirst, 1924 - 5

Parateneriffia Thor, 1911 - 1

Sinoteneriffia Yin, Bei et Li, 1995 – 2

Teneriffia Thor, 1911 - 5

Catalogue of Teneriffiiidae

Family Teneriffiidae Thor

Teneriffiidae Thor, 1911: 179

Teneriffiolidae Hirst, 1924: 1078

Teneriffiinae Womersley, 1935: 334

Type genus: Teneriffia Thor, 1911

Genus Austroteneriffia Womersley

Austroteneriffia Womersley, 1935: 334

Type species: Austroteneriffia hirsti Womersley, 1935

Austroteneriffia hirsti Womersley

Austroteneriffia hirsti : Womersley, 1935: 335; Vitzthum, 1942: 808; Radford, 1950: 76; Judson, 1995: 830

Parateneriffia hirsti : McDaniel, Morihara & Lewis, 1976: 532

Type locality: Menindie (S. Australia)

Distribution: Australia

Austroteneriffia hojoensis (Shiba et Furukawa)

Neoteneriffiola hojoensis : Shiba & Furukawa, 1975: 118

Austroteneriffia hojoensis : Judson, 1995: 833

Type locality: Kashima, Hojo, Ehime Pref., near the sea shore, Japan

Distribution: Japan

Austroteneriffia japonica (Ehara)

Neoteneriffiola japonica : Ehara, 1965: 226; Shiba & Furukawa, 1975: 111

Austroteneriffia japonica (Ehara): Judson, 1995: 833

Type locality: Sapporo, Japan

Distribution: Japan

Austroteneriffia kamalii Ueckermann et Khanjani

Austroteneriffia kamalii Ueckermann & Khanjani, 2002: 168

Type locality: Famenin, Hamadan prov., litter of Medicago sativa

Distribution: Iran

Austroteneriffia khorramabadiensis Khanjani, Hoseini et Asali Fayaz

Austroteneriffia khorramabadiensis Khanjani, Hoseini & Asali Fayaz, 2014: 70

Type locality: Kaka Reza region, Khorramabad vic., Lorestan Province

Distribution: Iran

Austroteneriffia leei Judson

Austroteneriffia leei : Judson, 1995: 835

Type locality: Coomandook, S. Australia

Distribution: South Australia (Australia)

Austroteneriffia littorina Shiba et Furukawa

Austroteneriffia littorina : Shiba & Furukawa, 1975: 123; Judson, 1995: 837

Type locality: Wake, Matsuyama, Ehime Pref., Japan

Distribution: Japan

Austroteneriffia shiraziensis Khanjani, Yazdanpanah et Asali Fayaz

Austroteneriffia shiraziensis Khanjani, Yazdanpanah & Asali Fayaz, 2013: 36

Type locality: Koohmare Sorkhi region, Fars province

Distribution: Iran

Austroteneriffia tadjikistanica (Wainstein)

Neoteneriffiola tadjikistanica Wainstein, 1969: 1250; 1978: 202

Austroteneriffia tadjikistanica (Wainstein): Judson, 1995: 838; Ueckermann, van Harten & Smith Meyer, 2006: 256

Type locality: Tadjikistan

Distribution: Tadjikistan, Yemen (incl. Socotra Island)

Austroteneriffia zamaniani Khanjani, Asali Fayaz et Ueckermann

Austroteneriffia zamaniani Khanjani, Asali Fayaz & Ueckermann, 2011: 551

Type locality: Sorkh Abad Village, Hamedan vic., Hamedan Prov.

Distribution: Iran

Austroteneriffiaandrei Besch (species inquirenda)

Austroteneriffia Andréi: Besch, 1962: 236

Austroteneriffia” andrei Besch: Judson, 1995: 827 (another genus!)

Type locality: “Arroyo Lopez, Gletscherbach bei Bariloche”, Prov. Rio Negro, Distribution: Argentina

Genus Heteroteneriffia Hirst

Heteroteneriffia Hirst, 1925: 1278; synonymized by McDaniel et al., 1976: 529, but restaured by Ueckermann & Durucan, 2020

Type species: Heteroteneriffia marina Hirst, 1925

Heteroeneriffia marina Hirst

Heteroteneriffia marina : Hirst, 1925: 1278

Heteroteneriffiolla marina : Shiba, 1976: 134

Teneriffia marina (Hirst): McDaniel et al., 1976: 529

Type locality: Teluk Kemang, near Port Dickson, Negri Sembilan, Malaysia

Distribution: Japan, Malaysia

Heteroteneriffia mortoni Luxton

Heteroteneriffia mortoni Luxton, 1993: 104

Teneriffia mortoni (Luxton): Judson, 1994: 115

Type locality: Hong Kong marine littoral

Distribution: China (Hong Kong)

Genus Himalteneriffia Schmölzer

Himalteneriffia Schmölzer, 2002: 133

Type species: Himalteneriffia riccabonai Schmölzer, 2002

Himalteneriffia riccabonai Schmölzer

Himalteneriffia riccabonai Schmölzer, 2002: 133

Type locality: “Basislager am Markha – Fluss ca. 50 km nördlich von Leh, ca. 77o öL., 34o nBr., nur wenige km vom Kongmaru-La entfernt; ca. 5050 m Höhe”

Distribution: India (Ladakh)

Genus Mesoteneriffia Irk

Mesoteneriffia Irk, 1939: 220

Type species: Mesoteneriffia steinbocki Irk, 1939

Mesoteneriffia steinbocki Irk

Mesoteneriffia steinbocki : Irk, 1939: 222; Strandtmann, 1965: 261

Mesoteneriffia steinboecki : Schmölzer, 1952: 22; 2002: 124; Schweizer & Bader, 1963: 263

Type locality: “Ötztaler und Stubaier Alpen”, 2719 - 3050 m, Austria

Distribution: Austria, Switzerland

Genus Mesoteneriffiola Schmölzer

Mesoteneriffiola Schmölzer, 1956: 549

Type species: Mesoteneriffiola alpina Schmölzer, 1956

Mesoteneriffiola alpina Schmölzer

Mesoteneriffiola alpina Schmölzer, 1956: 549; 2002: 124

Type locality: “Unterhalb d. Roche d’Alvau, 2900 m”, Daufiné

Distribution: France

Genus Neoteneriffiola Hirst

Neoteneriffiola Hirst, 1924: 1078

Parateneriffia Thor: McDaniel et al., 1976: 532 (in part)

Type species: Neoteneriffiola luxoriensis Hirst, 1924

Neoteneriffiola coineaui Judson

Neoteneriffiola coineaui : Judson, 1994: 117

Type locality: Hamilton Range, Namibia

Distribution: Namibia

Neoteneriffiola luxoriensis Hirst

Neoteneriffiola luxoriensis : Hirst, 1924: 1078; Judson, 1994: 115

Parateneriffia luxoriensis :

Type locality: Assuan and Luxor, Egypt

Distribution: Egypt

Neoteneriffiola uta Tibbetts

Neoteneriffiola uta : Tibbetts, 1958: 44; López-Campos & Vázquez-Rojas, 2010: 157

Parateneriffia uta (Tibbetts): McDaniel et al., 1976: 532

Type locality: Utah

Distribution: Mexico, U.S.A. (Utah)

Neoteneriffiola xerophila De Oliveira Bernardi, Pellegrini et Ferreira

Neoteneriffiola xerophila De Oliveira Bernardi, Pellegrini & Ferreira, 2012: 411

Type locality: Brazil

Distribution: Brazil

Neoteneriffiola yunnanensis Luo Youzhen, Yin Suigong, Chen Bin et Zhang Yidong

Neoteneriffiola yunnanensis Luo Youzhen, Yin Suigong, Chen Bin & Zhang Yidong, 1997: 87

Type locality: Yunnan

Distribution: China (Yunnan)

Neoteneriffiola sp

Distribution: South Africa (Meyer et al., 1973)

Genus Parateneriffia Thor

Parateneriffia Thor, 1911: 176

Type species: Parateneriffia bipectinata Thor, 1911

Parateneriffia bipectinata Thor

Parateneriffia bipectinata Thor, 1911: 177

Type locality: Paraguay

Distribution: Paraguay

Genus Sinoteneriffia Yin, Bei et Li

Sinoteneriffia Yin, Bei & Li, 1995: 433

Type species: Sinoteneriffia nuda Yin, Bei et Li, 1995

Sinoteneriffia kunmingensis Luo, Yin, Zhang et Bin

Sinoteneriffia kunmingensis Luo, Yin, Zhang & Bin, 1997: 243

Type locality: Kunming

Distribution: China

Sinoteneriffia nuda Yin, Bei et Li

Sinoteneriffia nuda Yin, Bei & Li, 1995: 434 (Engl. 437)

Type locality: Liaoning Prov., from Sabina chinensis

Distribution: China

Genus Teneriffia Thor

Teneriffia Thor, 1911: 172

Teneriffiola Strand, 1911: 287

Type species: T. quadripapillata Thor, 1911

Teneriffia aethiopica Zmudzinski, Skoracki et Friedrich

Teneriffia aethiopica Zmudzinski, Skoracki & Friedrich, 2021: 318

Type locality:

Distribution: Ethiopia

Teneriffia mexicana McDaniel, Morihara et Lewis

Teneriffia mexicana : McDaniel et al., 1976: 529

Type locality: Sonora, Mexico

Distribution: Mexico

Teneriffia quadripapillata Thor

Teneriffia quadripapillata Thor, 1911: 173; Schmölzer, 2002: 131

Teneriffiola quadripapillata (Thor): Hirst, 1924: 1080

Type locality: Orotava, Teneriffa

Distribution: Teneriffa (Canary Is., Spain)

Teneriffia sebahatae Ueckermann et Durucan

Teneriffia sebahatae Ueckermann & Durucan, 2020: 1141

Type locality: sea shore nr. Antalya (Kundu)

Distribution: Turkey

Teneriffia hajiqanbari Paktinat-Saeij et Kazemi

Teneriffia hajiqanbari Paktinat-Saeij & Kazemi, 2022: 263

Type locality: Queshm Island, southern Hormozgan Province

Distribution: Iran

Distribution of Teneriffiidae

EUROPE

Austria (Irk, 1939; Schmölzer, 1952) - Mesoteneriffia steinbocki

France (Schmölzer, 1956) - Mesoteneriffiola alpina

Greece (Beron, in prep.) - Teneriffiidae

Switzerland (Schweizer & Bader, 1963) - Mesoteneriffia steinbocki

Turkey (Ueckermann & Durucan, 2020) – Teneriffia sebahatae

AFRICA

Canary Is.(Thor, 1911) - Teneriffia quadripapillata

Egypt (Hirst, 1924) - Neoteneriffiola luxoriensis

Ethiopia (Zmudzinsk, Skoracki & Friedrich, 2021) – Teneriffia aethiopica

Namibia (Judson, 1994) - Neoteneriffiola coineaui

Nigeria (Beron, in prep.) - Teneriffiidae

Somalia (Beron, in prep.) - Teneriffiidae

South Africa (Meyer et al., 1973) - Neoteneriffiola sp.

Tanzania (Beron, in prep.) - Teneriffiidae

Zambia (Beron, in prep.) - Teneriffiidae

ASIA

Afghanistan (Beron, in prep.) - Teneriffiidae

China (Yin Suigong, Bei Naxir, Li Xuejin, 1995; Luo Youzhen, Yin Suigong, Zhang Yidong, Chen Bin, 1996; Luo Youzhen, Yin Suigong, Chen Bin, Zhang Yidong, 1997; Beron, in prep.) – Neoteneriffiola yunnanica, Sinoteneriffia kunmingensis, S. nuda

India (Schmölzer, 2002; Paktinat-Saeij & Kazemi, 2022 Zmudzinski et al., 2021) – Himalteneriffia riccabonai

Iran (Ueckermann & Khanjani, 2002; Khanjani, Asali Fayaz & Ueckermann, 2011; Khanjani, Yazdanpanah & Asali Fayaz, 2013; Khanjani, Hoseini & Asali Fayaz, 2014) – Austroteneriffia kamalii, A. khorramabadiensis, A. shiraziensis, A. zamaniensis

Japan (Ehara, 1965; Shiba, 1976; Shiba & Furukawa, 1975; Judson, 1995) – Austroteneriffia hojoensis, A. japonica, A. littorina, Teneriffia marina

Malaysia (Hirst, 1925) - Teneriffia marina

Tadjikistan (Wainstein, 1969) - Austroteneriffia tadjikistanica

Yemen (incl. Socotra)(Ueckermann, van Harten & Smith Meyer, 2006) – Austroteneriffia tadjikistanica, Parateneriffia sp.

NORTH and CENTRAL AMERICA

Mexico (McDaniel, Morihara et Lewis, 1976; López-Campos & Vázquez-Rojas, 2010; Beron, in prep.) - Neoteneriffiola uta, Teneriffia mexicana

USA (Tibbetts, 1958) - Neoteneriffiola uta

SOUTH AMERICA

Argentina (Besch, 1962) - “Austroteneriffia” andrei

Brazil (Bernardi et al., 2012) – Neoteneriffiola

Paraguay (Thor, 1911) - Parateneriffia bipectinata

Peru (Beron, in prep.) - Teneriffiidae

OCEANIA

Australia (Womersley, 1935; Judson, 1995) - Austroteneriffia hirsti, A. leei

Bibliography on Teneriffiidae

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  • Tibbetts T. 1958. A new species in the genus Neoteneriffiola from Utah (Acarina: Anystoidea: Teneriffiidae). - Great Basin Naturalist, 18: 43 - 45.
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The Mite Family Pseudocheylidae Oudemans

Pseudocheylidae Oudemans, 1909

Small free - living mites, found under bark, in moss, soil or in bird’s nest. Three genera with 24 known species. Widely distributed, but rarely found.

Catalogue of the Family Pseudocheylidae

Family Pseudocheylidae Oudemans

Pseudocheylidae Oudemans, 1909:

Type genus: Pseudocheylus Berlese, 1888

Genus Anoplocheylus Berlese

Anoplocheylus Berlese, 1910: 210 (subgenus)

Anoplocheylus (Berlese): Baker & Atyeo, 1964: 262 (genus)

Type species: Pseudocheylus (Anoplocheylus) europaeus Berlese, 1910

Rhagina Womersley, 1935: 336 (Syn. by Baker & Wharton, 1952)

Type species: Rhagina protea Womersley, 1935

Anoplocheylus aegypticus Baker et Atyeo

Anoplocheylus aegypticus Baker & Atyeo, 1964: 267; Gerson, 1967: 367; van Dis & Ueckermann, 1991: 113

Type locality: Ismaelia - Port Said Road, 50 kilometers

Distributon: Egypt

Anoplocheylus bonabjadidiensis Navaei-Bonab, Bagheri, Ueckermann et Zarei

Anoplocheylus bonabjadidiensis Navaei-Bonab et al., 2011: 421; Navaei-Bonab & Kazazi, 2013: 28

Type locality: Bonabjadid, Marand, East Azerbaijan Province, Iran

Distributon: Iran

Anoplocheylus brasiliensis Khaustov et Tolstikov

Anoplocheylus brasiliensis Khaustov & Tolstikov, 2015: 60

Type locality: Rio de Janeiro, Leme, Atlantic rain forest

Distribution: Brazil

Anoplocheylus brevisetosus Ueckermann et Khanjani

Anoplocheylus brevisetosus Ueckermann & Khanjani, 2004: 59

Type locality: Rustenburg, North West Province

Distributon: South Africa

Anoplocheylus clavatus Baker et Atyeo

Anoplocheylus clavatus Baker & Atyeo, 1964: 265; van Dis & Ueckermann, 1991: 106

Type locality: Gnangara (16 miles north of Perth), Western Australia

Distributon: Australia

Anoplocheylus corticicola Fungarworn et Butcher

Anoplocheylus corticicola Fungarworn & Butcher, 2015: 2

Type locality: Bangkok, Pathumwan Dist., Chulalongkorn Univ. Campus

Distribution: Thailand

Anoplocheylus europaeus (Berlese)

Pseudocheylus (Anoplocheylus) europaeus Berlese, 1910: 210

Anoplocheylus europaeus (Berlese): Baker & Atyeo, 1964: 263; van Dis & Ueckermann, 1991: 106; Livshitz & Mitrofanov, 1973: 771; Ueckermann & Khanjani, 2004: 55

Type locality: Palermo

Distributon: Italy (Sicily)

Anoplocheylus hapsiscutus Flechtmann

Anoplocheylus hapsiscutus Flechtmann, 1971: 49

Type locality: Piracicaba, São Paulo

Distributon: Brazil

Anoplocheylus kazemii Bagheri

Anoplocheylus kazemii Bagheri, in Bagheri et al., 2013: 292

Type locality: Iran

Distributon: Iran

Anoplocheylus malayeriensis Ueckermann et Khanjani

Anoplocheylus malayeriensis Ueckermann & Khanjani, 2004: 55; Navaei-Bonab & Kazazi, 2013: 28

Type locality: Mey, 23 km from Malãyer to Arãk

Distributon: Iran

Anoplocheylus marivaniensis Khanjani, Hoseini et Amini

Anoplocheylus marivaniensis Khanjani, Hoseini & Amini, 2014: 186

Type locality: Merivan, Kurdistan Province

Distributon: Iran

Anoplocheylus paraclavatus van Dis et Ueckermann

Anoplocheylus paraclavatus van Dis & Ueckermann, 1991: 106; Ueckermann & Khanjani, 2004: 55

Type locality: Edenville, Pongola

Distributon: South Africa

Anoplocheylus protea (Womersley)

Rhagina protea Womersley, 1935: 336

Anoplocheylus protea (Womersley): Baker & Atyeo, 1964: 264; van Dis & Ueckermann, 1991: 111; Ueckermann & Khanjani, 2004: 57

Type locality: Glen Osmond, South Australia

Distributon: Australia

Anoplocheylus qorvehiensis Khanjani, Hoseini et Amini

Anoplocheylus qorvehiensis Khanjani, Hoseini & Amini, 2014: 189

Type locality: Qorveh, Kurdistan Province

Distributon: Iran

Anoplocheylus sinai Bagheri

Anoplocheylus sinai Bagheri, in Bagheri et al., 2013: 290; Navaei-Bonab & Kazazi, 2013: 28

Type locality: Iran

Distributon: Iran

Anoplocheylus tauricus Livshitz et Mitrofanov

Anoplocheylus tauricus Livshitz & Mitrofanov, 1973: 770; van Dis & Ueckermann, 1991: 113; Ueckermann & Khanjani, 2004: 65; Bayram & Cobanoğlu, 2006:

Type locality: cape Kazantip, Crimea

Distributon: Turkey, Crimea, Iran

Anoplocheylus tellustrus van Dis et Ueckermann

Anoplocheylus tellustrus van Dis & Ueckermann, 1991: 109

Anoplocheylus reticulatus van Dis & Ueckermann, 1991: 112. Type: Pongola, Edenville, South Africa. Syn. by Ueckermann & Khanjani, 2004: 61

Type locality: Pongola, Edenville

Distributon: South Africa

Anoplocheylus transiens Delfinado et Baker

Anoplocheylus transiens Delfinado & Baker, 1974: 209; van Dis & Ueckermann, 1991: 113; Ueckermann & Khanjani, 2004: 61

Type locality: Heckscher Park, Long Islang, New York

Distributon: USA

Genus Neocheylus Trägårdh

Neocheylus Trägårdh, 1906: 870

Type species: Neocheylus natalensis Trägårdh, 1906

Neocheylus collis Baker et Atyeo

Neocheylus collis Baker & Atyeo, 1964: 270

Type locality: Mount Toolbrunup (Stirling Ranges, 300 miles south from Perth), W. Australia

Distributon: Australia

Neocheylus natalensis Trägårdh

Neocheylus natalensis Trägårdh, 1906: 870

Type locality: Natal

Distributon: South Africa

Neocheylus nidicolus Lawrence

Neocheylus nidicolus Lawrence, 1954: 74

Type locality: Karamoja (Uganda)

Distributon: Uganda

Genus Pseudocheylus Berlese

Pseudocheylus Berlese, 1888: 189

Type species: Pseudocheylus biscalatus Berlese, 1888

Pseudocheylus americanus (Ewing)

Cheyletiella americana Ewing, 1909: 429

Pseudocheylus americanus (Ewing): Baker & Wharton, 1952: 226; Skvarla et al., 2013: 401

Type locality: Urbana, Illinois

Distributon: USA (Illinois)

Pseudocheylus biscalatus Berlese

Pseudocheylus biscalatus Berlese, 1888: 189; Flechtmann, 1971: 33

Type locality: Rio-Apa (Paraguay), Mato Grosso (Brazil)

Distribution: Brazil, Paraguay

Distribution of Pseudocheylidae

EUROPE

Italy (contin.)

Sicily (Berlese, 1910) – Anoplocheylus europaeus

Russia

Crimea (Livshitz & Mitrofanov, 1973) – Anoplocheylus tauricus

Turkey (all) (Bayram & Cobanoğlu, 2006) - Anoplocheylus tauricus

AFRICA

Egypt (Baker & Atyeo, 1964) – Anoplocheylus aegyptiacus

South Africa (Trägårdh, 1906; van Dis & Ueckermann, 1991; Ueckermann & Khanjani, 2004) – Anoplocheylus brevisetosus, A. paraclavatus, A. tellustrus (= reticulatus), Neocheylus natalensis

Uganda (Lawrence, 1954) – Neocheylus nidicolus

ASIA

Iran (Ueckermann & Khanjani, 2004; Navaei-Bonab et al., 2011; Bagheri et al., 2013; Navaei-Bonab & Kazazi, 2013: 28; Khanjani, Hoseini & Amini, 2014) – Anoplocheylus bonabjadidiensis, A. kazemii, A. malayeriensis, A. marivaniensis, A. qorvehiensis, A. sinai, A. tauricus

Israel (Gerson, 1967) - Anoplocheylus aegyptiacus

Thailand (Fuangarworn & Butcher, 2015) – Anoplocheylus corticicola

NORTH AMERICA

Canada (Smith & Lindquist,1979) – one species

USA (Ewing, 1909; Delfinado & Baker, 1974; Skvarla et al., 2013) – Anoplocheylus transiens, Pseudocheylus americanus

SOUTH and CENTRAL AMERICA

Brazil (Berlese, 1888; Flechtmann, 1971; Khaustov & Tolstikov, 2015) – Anoplocheylus brasiliensis, A. hapsiscutus, Pseudocheylus biscalatus

Paraguay (Berlese, 1888) - Pseudocheylus biscalatus

OCEANIA

Australia (continent) (Womersley, 1935; Baker & Atyeo, 1964) – Anoplocheylus clavatus, A. protea, Neocheylus collis

Bibliography of Pseudocheylidae

  • Baker E., W. Atyeo. 1964. A Review of the Mites of the Family Pseudocheylidae Oudemans, 1909 (Acarina, Prostigmata). – Bull. Univ. of Nebrasca State Museum, 4(12): 257 - 272.
  • Baker E.W., G.W. Wharton. 1952. An Introduction to Acarology. - Macmillan Co., New York.
  • Bagheri M., E. Zarei, M. Ahaniazad, Gh. Gharekhany, R. Navaei-Bonab. 2013. Two new species of the genus Anoplocheylus Berlese, 1910 (Acari: Trombidiformes: Pseudocheylidae) from Iran. – Zootaxa, 3599(3): 291 – 297.
  • Bayram S., S. Cobanoglu. 2006. Astigmata and Prostigmata (Acari) of Bulbaceous Ornamental Plants in Ankara, Turkey. – Acta Phytopathologica et Entomologica Hungarica, 41(3-4): 367 - 381
  • Berlese A. 1888. Acari Austro-Americani quos collegit Aloysius Balzan. Manipulus primus. – Bull. Soc. Ent. Ital., 20: 171 - 222.
  • Berlese A. 1910. Acari nuovi. Manipulus V. – Redia, 6: 199 - 234. Delfinado M., E.W. Baker. 1974. Terrestrial Mites of New York (Acarina: Prostigmata), I – Tarsocheylidae, Paratydeidae, and Pseudocheylidae. – New York Ent. Soc., 82: 202 - 211.
  • Ewing H.E. 1909. A systematic and biological study of the Acarina of Illinois. – Univ. Illinois Bull., 7(14): 77 - 78.
  • Flechtmann C.H.W. 1971. Alguns Trombidiformes do Brasil e do Paraguai (Acari). Piracicaba, Estado de Sao Paulo: 63+16 pp.
  • Fuangarworn M., B.A. Butcher. 2015. Contribution to the family Pseudocheylidae (Acari, Trombidiformes) from Thailand: one new species and one new record of Anoplocheylus Berlese, with observations on their onthogeny. - Intern. J. of Acarology, 41(8): 625 – 641.
  • Gerson U. 1967. Some cheyletid and pseudocheyletid mites from Israel. – Acarologia, 9: 359 – 369.
  • Khanjani M., M.A. Hoseini, F. Amini. 2014. Two new Anoplocheylus species (Acari: Trombidiformes: Pseudocheylidae) from Kurdistan province of Iran. – Zootaxa, 3861(2): 185 – 192.
  • Khaustov A.A., A.V. Tolstikov. 2015. First record of mites of the genus Anoplocheylus (Acari: Pseudocheylidae) in South America with description of a new species from Brazil. – Neotropical Entomology, 44(1): 59 – 67.
  • Lawrence R.F. 1954. The known African species of Cheyletidae and Pseudocheylidae (Acarina, Prostigmata). – Ann. Natal Museum, 13(1): 65 - 77.
  • Livshitz I.S., V.I. Mitrofanov. 1973. A new species of the genus Anoplocheylus (Trombidiformes, Pseudocheylidae) from Crimea. – Zool. Zhurnal, 52(5): 70 - 771 (in Russian, summ. Engl.).
  • Navaei-Bonab R., M. Bagheri, E.A. Ueckermann, E. Zarei. 2011. Description of a new species of Anoplocheylus Berlese, 1910 (Acari: Trombidiformes: Pseudocheylidae) from Iran. – Acarologia, 51(4): 419 – 423.
  • Navaei-Bonab R., M. Kazazi. 2013. Some Pseudocheylidae (Acari: Trombidiformes) of Marand (Northwest of Iran). In: Program and Abstract book of the Second Intern. Persian Congress of Acarology, Karaj, Iran, p.28.
  • Oudemans A.C. 1909. Acarologische Aantek. 26. – Ent. Berichte, II(47): 317 – 320. Skvarla M.J., J. R. Fisher, A.P.G. Dowling. 2013. On some mites (Acari: Prostigmata)from the Interior Highlands: descriptions of the male, immature stages, and female reproductive system of Pseudocheylus americanus (Ewing, 1909) and some new state records for Arkansas. – Zootaxa, 3641(4): 401 – 419.
  • Smith I.M., E.E. Lindquist. 1979. Prostigmata. In: H.V. Danks (ed.), Canada and its Insect fauna. - Memoirs of the Entomology Society of Canada, 108: 267–277.
  • Trägårdh I. 1906. Neue Acariden aus Natal und Zululand. – Zool. Anz., 30: 870 - 877.
  • Trägårdh I. 1950. Description of a new species of Heterocheylus Lombardini from Africa, with notes on the classification of the Pseudocheylidae. - Entomologisk Tidskrift, 71(2): 104–110.
  • Ueckermann E.A., M. Khanjani. 2004. A revision of the genus Anoplocheylus Berlese (Acari: Pseudocheylidae), with the description of two new and re-description of four known species. – Syst. & Applied Acarology, 9: 53 - 68.
  • Van Dis J.C.S., E.A. Ueckermann. 1991. A review of the Pseudocheylidae Oudemans (Acari: Prostigmata). – Phytoprophylactica, 23(2): 105 - 113.
  • Womersley H. 1935. On the occurrence in Australia of Acarina of the Family Teneriffiidae (Trombidoidea). - Rec. South Australian Mus., 5: 333 - 338.

The Mite Family Stigmocheylidae Berlese

Stigmocheylidae Berlese, 1910: 209

Small family (five known species) of soil mites.

Catalogue of Stigmocheylidae

Family Stigmocheylidae Berlese

Stigmocheylidae Berlese, 1910b: 209; Kethley, 1990: 698

Type genus: Stigmocheylus Berlese, 1910

Genus Stigmocheylus Berlese

Stigmocheylus Berlese, 1910a: 209

Type species: Stigmocheylus brevisetus Berlese, 1910

Stigmocheylus americanus Bochkov

Stigmocheylus americanus Bochkov, 2008: 315

Type locality: Las Cruces, Jornada, New Mexico

Distribution: USA (New Mexico)

Stigmocheylus brevisetus Berlese

Stigmocheylus brevisetus Berlese, 1910a: 209

Type locality: Pisa

Distribution: Italy

Stigmocheylus oberon Bochkov

Stigmocheylus oberon Bochkov, 2008: 314

Type locality: New Mexico

Distribution: USA (New Mexico)

Stigmocheylus persicus Damavandian et Paktinat-Saeij

Stigmocheylus persicus Damavandian & Paktinat-Saeij, 2018: 2160

Type locality: Noor, Mazandaran province, northern Iran

Distribution: Iran

Stigmocheylus pilosus (Soliman et Zaher)

Hemitarsocheylus pilosus Soliman & Zaher, 1975: 105

Stigmocheylus pilosus (Soliman et Zaher): Bochkov, 2008: 314

Type locality: Giza

Distribution: Egypt

Distribution of Stigmocheylidae

EUROPE

Italy (Berlese, 1910) - Stigmocheylus brevisetus

AFRICA

Egypt (Soliman & Zaher, 1975) – Stigmocheylus pilosus

ASIA

Iran (Naghibinejad, Ahadiyat, & Ostovan, 2009; Damavandian & Paktinat-Saeij, 2018) – Stigmocheylus persicus

NORTH AMERICA

USA (Bochkov, 2008) – Stigmocheylus americanus, S. oberon

Bibliography of Stigmocheylidae

  • Baker E., W. Atyeo. 1964. A Review of the Mites of the Family Pseudocheylidae Oudemans, 1909 (Acarina, Prostigmata). – Bull. Univ. of Nebrasca State Museum, 4(12): 257 - 272.
  • Berlese A. 1910a. Acari nuovi. Manipulus V. – Redia, 6: 199 - 234.
  • Bochkov A. V. 2008. A review of the mite family Stigmocheylidae Berlese (Acari: Prostigmata). – Annales Zoologici (Warsaw), 58(2): 311 - 325. Damavandian M.R., S. Paktinat-Saeij. 2018. First new species of the family Stigmocheylidae (Acari: Prostigmata: Paratydeoidea) from Asia. - Syst. & Applied Acarology, 23(11): 2159 – 2164.
  • Damavandian M. R., S Paktinat - Saey. 2018. First new species of the family Stigmocheylidae (Acari: Prostigmata: Paratydeoidea) from Asia. Syst. and applied acarology 23 (11).
  • Kethley J.B. 1990. Acarina: Prostigmata (Actinedida), pp. 667–756. In: D. L. Dindal (ed). Soil Biology Guide. Wiley, New York.
  • Lindquist E.E. 1976. Transfer of the Tarsocheylidae to the Heterostigmata, and reassignement of the Tarsonemina and Heterostigmata to lower hierarchic status in the Prostigmata (Acari). – The Canadian Entomologist, 108: 23 - 48.
  • Naghibinejad M., A. Ahadiyat, H. Ostovan. 2009. First report of the family Stigmocheylidae (Trombidiformes: Paratydeoidea) from Iran. - Plant Protection Journal, 3(1): 341–343 (In Persian).
  • Soliman Z.R., M.A. Zaher. 1975. Hemitarsocheylus a new genus from the family Tarsocheylidae with a description of a new species. - Acarologia, 17: 103–105.

The mite Superfamily Caeculoidea

Superfamily Caeculoidea Ewing, 1934

Phalanx Caeculata Wainstein, 1965: 77

The superfamily contains one family – Caeculidae.

The mite family Caeculidae

Family Caeculidae Berlese, 1883

Dufour (1832) started the study of these peculiar mites, called after the mythical son of Vulcan. They prefer the open rocky habitats and are known already from all continents. The second genus in the family (Procaeculus) was described by Jacot (1936), more than a century after the first. The low intensity of the studies of Caeculidae of many countries is shown by the multitude of species, found in the USA by several authors, mostly by Mulaik (1945), and (after 1952) in Spain, Morocco, N. Macedonia and Chad by H. Franz, who revised the family (Franz, 1952), describing the new genera Allocaeculus and Microcaeculus, and compiling a list of all known Caeculidae. Very profound research on the family was enterprised by Y. Coineau since 1963. He described several new species, mainly from Southern Africa, and the new genera Calocaeculus, Andocaeculus, and Neocaeculus. Three new species have been described from Australia (Taylor, Gunawardene & Kinnear, 2013, Taylor, 2014) and one from Thailand (Fuangarworn & Butcher, 2015) and Canada (Bernard et al., 2020). One new Allocaeculus was described from Slovakia (Mangová et al., 2014), another one from Turkey (Per et al., 2017). New checklist of the family was published by Taylor et al. (2013).

Fossil species have been described from amber from Poland, Myanmar, Mexico, and Dominican Republic (Coineau & Magowski, 1994, Coineau & Poinar, 2000, Rivas et al., 2016, Porta et al., 2019).

Now the family is considered to contain at least 114 sp. in seven genera.

Fam. Caeculidae – 114 sp.

Allocaeculus Franz, 1952 – 36 sp.

Andocaeculus Coineau, 1974 - 5

Caeculus Dufour, 1832 - 21

Calocaeculus Coineau, 1974 - 1

Microcaeculus Franz, 1952 - 25

Neocaeculus Coineau, 1967 – 12

Procaeculus Jacot, 1936 - 13

Incertae sedis - 1

Catalogue of Caeculidae

Family Caeculidae Berlese

Hoplopini Canestrini & Fanzago, 1877: 481

Caeculidae Berlese, 1883

Type genus: Caeculus Dufour, 1832

Genus Allocaeculus Franz

Allocaeculus Franz, 1952: 108

Type species: Allocaeculus relictus Franz, 1952

Allocaeculus andalusiacus Franz

Allocaeculus andalusiacus Franz, 1952: 113

Type locality: Sevilla, Cinca de Pino, Andalusia

Distribution: Spain

Allocaeculus catalanus Franz

Allocaeculus catalanus Franz, 1952: 2; 1954: 210; 1955: 12

Type locality: Gerona (Spain), Eastern Pyrenees (France)

Distribution: France, Kazachstan, Russia, Spain

Allocaeculus circinatus Coineau

Allocaeculus circinatus Coineau, 1974b: 278

Type locality: Ghaub Farm in Ovidi highlands, 25 km S Tsumeb

Distribution: Namibia

Allocaeculus dubius (Kulczynski)

Caeculus dubius Kulczynski, 1901: 367

Allocaeculus dubius (Kulczynski): Franz, 1952: 122

Type locality: Algeria

Distribution: Algeria

Allocaeculus echinatus Franz

Allocaeculus echinatus Franz, 1952: 118

Type locality: Sevilla, Andalusia

Distribution: Spain

Allocaeculus erinaceus Coineau

Allocaeculus erinaceus Coineau, 1974b: 277

Type locality: Ingwavuma

Distribution: South Africa

Allocaeculus grandjeani Franz

Allocaeculus grandjeani Franz, 1957: 10

Type locality: Khenifra

Distribution: Morocco

Allocaeculus hirsutus Coineau

Allocaeculus hirsutus Coineau, 1974b: 277

Type locality: Glencoe

Distribution: South Africa

Allocaeculus hoggarensis (André)

Caeculus hoggarensis André, 1936: 111

Allocaeculus hoggarensis (André): Franz, 1952: 122; Piffl, 1959: 54

Type locality: Hoggar, Mt Ilaman

Distribution: Algeria

Allocaeculus hystriciformis Franz

Allocaeculus hystriciformis Franz, 1952: 120

Type locality: Mte Arastepa in Sierra Ronda

Distribution: Spain

Allocaeculus hystrix (Lawrence)

Caeculus hystrix Lawrence, 1939: 542

Allocaeculus hystrix (Lawrence): Franz, 1952: 122

Type locality: Zululand, Hluhluve Game Reserve

Distribution: South Africa

Allocaeculus indicus Piffl

Allocaeculus indicus Piffl, 1959: 52

Type locality: Saidu Sharif, Swat Area, NW Pakistan

Distribution: Pakistan

Allocaeculus kalahariensis Coineau

Allocaeculus kalahariensis Coineau, 1974b: 277

Type locality: Kalahari Gemsbok National Park

Distribution: South Africa

Allocaeculus kenyae Franz

Allocaeculus kenyae Franz, 1964: 82

Type locality: 8 km N Nairobi

Distribution: Kenya

Allocaeculus kocheri Franz

Allocaeculus kocheri Franz, 1964: 84

Type locality: Oukaimeden, 2300 m

Distribution: Morocco

Allocaeculus logonensis Franz

Allocaeculus logonensis Franz, 1957: 6

Type locality: Logon, Deressia nr. Lai

Distribution: Chad

Allocaeculus meseticola Franz

Allocaeculus meseticola Franz, 1952: 114; 1955: 14

Type locality: Alcala de Henares

Distribution: Spain

Allocaeculus mosambicensis (André)

Caeculus mosambicensis André, 1936: 115

Allocaeculus mosambicensis (André): Franz, 1952: 122; 1957: 12; Piffl, 1959: 54

Type locality: Vila Pery

Distribution: Mozambique

Allocaeculus multispinosus Franz

Allocaeculus multispinosus Franz, 1955: 11; 1960: 86

Type locality: Puerto de Perales, Sierra de Gata

Distribution: Spain, Turkey

Allocaeculus nigeriensis Coineau

Allocaeculus nigeriensis Coineau, 1974b: 278

Type locality: Ibadan

Distribution: Nigeria

Allocaeculus ouadaiensis Franz

Allocaeculus ouadaiensis Franz, 1957: 7

Type locality: Mont Kelinguen nr. Abecher

Distribution: Chad

Allocaeculus pilosus Lawrence

Caeculus pilosus Lawrence, 1938: 211

Allocaeculus pilosus (Lawrence): Franz, 1952: 122; 1957: 12; Piffl, 1959: 54

Type locality: N. Damaraland

Distribution: Namibia

Allocaeculus relictus Franz

Allocaeculus relictus Franz, 1952: 111; 1954b: 369; 1955: 15; Winkler, 1953: 55

Type locality: “Kalvarienberg b. Gumpoldskirchen”, Austria

Distribution: Austria, Czech Rep.

Allocaeculus sandbergensis Mangová, Krumpál et Ľuptáčik

Allocaeculus sandbergensis Mangová, Krumpál & Ľuptáčik, 2014: 214

Type locality: National nature reserve Devínska Kobyla near Sandberg

Distribution: Slovakia

Allocaeculus sarhroi Franz

Allocaeculus sarhroi Franz, 1964: 86

Type locality: Tizi-n-Tazazerte, Jbel Sarhro, 2100 m

Distribution: Morocco

Allocaeculus schusteri Franz

Allocaeculus schusteri Franz, 1960: 83

Type locality: Ohrid

Distribution: N. Macedonia

Allocaeculus sclerodermatus André

Caeculus sclerodermatus André, 1936: 106

Allocaeculus sclerodermatus (André): Franz, 1952: 122; 1957: 12; Piffl, 1959: 54

Type locality: South Tunisia, Nefta

Distribution: Tunisia

Allocaeculus sculptus (Karpelles)

Hoplopus sculptus Karpelles, 1893: 39

Caeculus sculptus Karpelles, 1893: 39

Allocaeculus sculptus (Karpelles): Franz, 1952: 122; Piffl, 1959: 54

Type locality: Hungary

Distribution: Hungary

Allocaeculus senegalensis (André)

Caeculus senegalensis André, 1938: 264

Allocaeculus senegalensis (André): Taylor et al., 2013: 448

Type locality: Dakar

Distribution: Senegal

Allocaeculus sirwani Paktinat-Saeij, Ghobari et Samani

Allocaeculus sirwani Paktinat-Saeij, Ghobari & Samani, 2020: 834

Type locality: Marivan city, Kurdistan Province

Distribution: Iran

Allocaeculus spatulifer (Michael)

Caeculus spatulifer Michael, 1890: 432; Vitzthum, 1933: 87

Allocaeculus spatulifer (Michael): Franz, 1952: 122

Type locality: Algeria

Distribution: Algeria, Croatia, Spain

Allocaeculus spinosissimus Franz

Allocaeculus spinosissimus Franz, 1952: 116

Allocaeculus spinosissimus rondae Franz, 1952: 118. Type locality: Mte Arastepa in Sierra Ronda

Type locality: Sierra Guadarrama above Escorial

Distribution: Spain

Allocaeculus sudanensis (Trägårdh)

Caeculus sudanensis Trägårdh, 1904: 70

Allocaeculus sudanensis : Franz, 1952: 122

Type locality: “Gebelein am weissen Nil”

Distribution: Sudan

Allocaeculus tenerifae Franz

Allocaeculus tenerifae Franz, 1965: 67

Type locality: Las Madrigueras (S. Tenerife)

Distribution: Spain (Canary Is.)

Allocaeculus tschadensis Franz

Allocaeculus tschadensis Franz, 1957: 8; 1964: 86

Type locality: Mont Kelinguen nr. Abecher (Chad)

Distribution: Chad, Morocco, Senegal

Allocaeculus tschadensis rifensis Franz

Allocaeculus tschadensis rifensis Franz, 1960: 84

Type locality: Zoco el Arba, road Tetuan - Xauen

Distribution: Morocco

Allocaeculus turcicus Per, Doğan, Zeytun et Ayyıldız

Allocaeculus turcicus Per, Doğan, Zeytun & Ayyıldız, 2017: 371

Type locality: Karamlıkdere Valley, Yozgat, 844 m

Distribution: Turkey

Genus Andocaeculus Coineau

Andocaeculus Coineau, 1974b: 279

Type species: Microcaeculus brundini Franz, 1962

Andocaeculus beatrizrosso Porta, Pizarro-Araya et Ramírez

Andocaeculus beatrizrosso Porta, Pizarro-Araya & Ramírez, 2021: 42

Type locality: Argentina

Distribution: Argentina

Andocaeculus brundini (Franz)

Microcaeculus Brundini Franz, 1962: 99

Andocaeculus brundini (Franz): Coineau, 1974b: 279

Type locality: Arroyo Lopez, W from Bariloche (Prov. Rio Negro)

Distribution: Argentina

Andocaeculus burmeisteri Porta, Pizarro-Araya et Ramírez

Andocaeculus burmeisteri Porta, Pizarro-Araya & Ramírez, 2021: 28

Type locality: Argentina

Distribution: Argentina

Andocaeculus caioi Ott et Ott

Andocaeculus caioi Ott & Ott, 2014: 356

Type locality: Fazenda Rincão dos Moraes, Área de Proteção Ambiental do Rio Ibirapuitã, Sant’Ana do Livramento, Rio Grande do Sul

Distribution: Brazil

Andocaeculus weinrauchi (Franz)

Microcaeculus weinrauchi Franz, 1964: 89

Andocaeculus weinrauchi :

Type locality: Puna “El Infernillo”, 3000 m

Distribution: Argentina

Genus Caeculus Dufour

Caeculus Dufour, 1832: 289

Hoplopus Canestrini & Fanzago, 1877: 480 (replacement name)

Type species: Caeculus echinipes Dufour, 1832

Pseudocaeculus Coineau, 1974 (unavailable name)

Pseudocaeculus americanus (Banks): Taylor et al., 2013: 448.

Caeculus americanus Banks

Caeculus americanus Banks, 1899: 221; McDaniel & Boe, 1990: 722

Type locality: Palm Springs, California

Distribution: USA (California)

Caeculus archeri Mulaik

Caeculus archeri Mulaik, 1945: 13; Franz, 1952: 121; McDaniel & Boe, 1990: 723

Type locality: Monte Sano, Madison Co., Alabama

Distribution: USA (Alabama, Tennessee)

Caeculus calechius Mulaik

Caeculus calechius Mulaik, 1945: 5; Franz, 1952: 121; McDaniel & Boe, 1990: 723

Type locality: Edinbu.rg, Hidalgo Co., Texas

Distribution: USA (Montana, Texas, Utah)

Caeculus cassiopeiae Bernard et Lumley

Caeculus cassiopeiae Bernard & Lumley, in: Bernard, Lumley, Buck & Cobb. 2020: 2

Type locality: Alberta, Medicine Hat, near Gas City Campground; 50°2.23’N, 110°43.56’W

Distribution: Canada

Caeculus clavatus Banks

Caeculus clavatus Banks, 1906: 135; Franz, 1952: 121; McDaniel & Boe, 1990: 722

Type locality: USA (California)

Distribution: USA (California)

Caeculus cremnicolus Enns

Caeculus cremnicolus Enns, 1958: 107; Skvarla, Fisher & Dowling, 2013: 401; Bernard et al., 2020:

Type locality: Baskett Wildlife Research and Education Area, Devil’s Backbone; Easley; Wilton, Missouri

Distribution: Canada, USA (Arkansas, Missouri)

Caeculus crossleyi Coineau

Caeculus crosleyi Coineau, 1974b: 276

Type locality: DeKalb & Rockdale Counties

Distribution: USA (Georgia)

Caeculus dorotheae Mulaik

Caeculus dorotheae Mulaik, 1945: 9; Franz, 1952: 121; McDaniel & Boe, 1990: 722 (dortheae)

Type locality: Lockes, Nye Co., Nevada

Distribution: USA (Arizona, Nevada)

Caeculus echinipes Dufour

Caeculus echinipes Dufour, 1832: 289; André, 1927: 168; 1932: 485; Irk, 1939: 154; Willmann, 1951: 148; Franz, 1952: 98; 1954a: 212; 1954b: 369; Kratochvil, 1962: 174; Schmölzer, 1962: 286; Coineau & Haupt, 1976

Hoplopus echinipes : Canestrini & Fanzago, 1877: 477

Type locality: Moxente (Valencia)

Distribution: Algeria, Austria, Bulgaria, France, Germany, Greece, Italy, Spain, Switzerland

Caeculus echinipes crosbyi Jacot

Caeculus echinipes crosbyi Jacot, 1936: 18; Franz, 1952: 121

Caeculus echinipes : Franz, 1955: 14

Type locality: Pic du Midi de Bigore, Pyrenees

Distribution: France

Caeculus gertschi Mulaik

Caeculus gertschi Mulaik, 1945: 8; Franz, 1952: 121

Type locality: Rio Grande City, Starr Co., Texas

Distribution: USA (Texas)

Caeculus hardyi Mulaik et Allred

Caeculus hardyi Mulaik & Allred, 1954: 36; McDaniel & Boe, 1990: 723

Type locality: Laredo, Harlingen (New Mexico, Texas), San Pedro (Mexico)

Distribution: Mexico, USA

Caeculus hypopachus Mulaik

Caeculus hypopachus Mulaik, 1945: 10; Franz, 1952: 122; McDaniel & Boe, 1990: 723

Type locality: Edinburg, Hidalgo Co., Texas, in stomach of toad (Hypopachus cuneus)

Distribution: USA (Texas)

Caeculus janetae Higgins et Mulaik

Caeculus janetae Higgins & Mulaik, 1957b: 267

Type locality: SW USA

Distribution: USA

Caeculus kerrulius Mulaik

Caeculus kerrulius Mulaik, 1945: 8; Franz, 1952: 122; McDaniel & Boe, 1990: 722

Type locality: Kerrville, KerrCo., Texas

Distribution: USA (Texas)

Caeculus krantzi Coineau

Caeculus krantzi Coineau, 1974b: 276

Type locality: Manzanita, 32 mi E of Sweet Home, Oregon

Distribution: USA (Oregon)

Caeculus lewisi McDaniel et Boe

Caeculus lewisi McDaniel & Boe, 1990: 716

Type locality: South Dakota

Distribution: USA (South Dakota)

Caeculus mariae Higgins et Mulaik

Caeculus mariae Higgins & Mulaik, 1957a: 27

Type locality: Oregon

Distribution: USA (Oregon)

Caeculus pettiti Nevin

Caeculus pettiti Nevin, 1943: 389; 1945: 195; Franz, 1952: 122; McDaniel & Boe, 1990: 723

Type locality: Virginia

Distribution: USA

Caeculus tipus Mulaik

Caeculus tipus Mulaik, 1945: 6; Mulaik & Allred, 1954: 38; Franz, 1952: 122; McDaniel & Boe, 1990: 723

Type locality: Edinburg, Hidalgo Co., Texas

Distribution: USA (Utah, Texas)

Caeculus valverdius Mulaik

Caeculus valverdius Mulaik, 1945: 6; Franz, 1952: 122; McDaniel & Boe, 1990: 72

Type locality: Edinburg, Hidalgo Co., Texas

Distribution: USA (Arizona, Texas)

Caeculus veracruzensis Porta et Vázquez Rojas

Caeculus veracruzensis Porta & Vázquez Rojas, 2020: 473

Type locality: La Mancha, Veracruz

Distribution: Mexico

Genus Calocaeculus Coineau

Calocaeculus Coineau, 1974b: 279

Type species: Calocaeculus lawrencei Coineau, 1974

Calocaeculus lawrencei Coineau

Calocaeculus lawrencei Coineau, 1974b: 279

Type locality: Pearston in Somerset East

Distribution: South Africa

Genus Microcaeculus Franz

Microcaeculus Franz, 1952: 101

Type species: Microcaeculus austriacus Franz, 1952

Sclerocaeculus Coineau, 1974

Type species: Sclerocaeculus namibensis (Piffl, 1965)

Microcaeculus andrei (Lawrence)

Caeculus Andrei Lawrence, 1939: 540

Microcaeculus andrei (Lawrence): Franz, 1952: 122; Halliday, 2005: 28

Type locality: South Africa

Distribution: South Africa

Microcaeculus austriacus Franz

Microcaeculus austriacus Franz, 1952: 103; 1954b: 369; Pieper, 1970: 106; Thaler et al., 1993: 311

Type locality: “Kalvarienberg b. Gumpoldskirchen a. O-Hang des Anningers südl. v. Wien” (Austria)

Distribution: Austria, Czech Rep., Germany

Microcaeculus canariensis Franz

Microcaeculus canariensis Franz, 1965: 68

Type locality: Ortes Los Cristianos (Tenerife)

Distribution: Spain (Canary Is.)

Microcaeculus castrii Franz

Microcaeculus castrii Franz, 1964: 87

Type locality: Cordillera de la Costa, Prov. De Coquimbo, Alcones (and several other localities)

Distribution: Chile

Microcaeculus deserticolus (Lawrence)

Caeculus deserticolus Lawrence, 1939: 537

Microcaeculus deserticolus (Lawrence): Franz, 1952: 122

Sclerocaeculus deserticolus (Lawrence) – Coineau, 1974: 59

Type locality: Kakamas, Namaqualand

Distribution: South Africa

Microcaeculus franzi Coineau

Microcaeculus franzi Coineau, 1968: 143

Type locality: Ghisonaccia, Corsica

Distribution: Corsica

Microcaeculus graecus Franz

Microcaeculus graecus Franz, 1964: 91

Type locality: Arachova, 2000 m

Distribution: Greece

Microcaeculus hispanicus Franz

Microcaeculus hispanicus Franz, 1952: 105; Coineau, 1966: 23

Type locality: Alcalá de Henares (Spain)

Distribution: France, Spain

Microcaeculus insolitus (André)

Caeculus insolitus André, 1935: 113

Microcaeculus insolitus (André): Franz, 1952: 122

Type locality: Grande Salvage

Distribution: Salvages Islands (Portugal)

Microcaeculus insularis Franz

Microcaeculus insularis Franz, 1964: 90

Type locality: The Big Island against Essaouira (Mogador)

Distribution: Morocco

Microcaeculus laoshanensis (Jacot)

Caeculus laoshanensis Jacot, 1936: 20

Microcaeculus laoshanensis (Jacot): Franz, 1952: 122

Type locality: Laoshan

Distribution: China

Microcaeculus liguricus (Vitzthum)

Caeculus liguricus Vitzthum, 1933: 87; Willmann, 1954: 237

Microcaeculus liguricus (Vitzthum): Franz, 1952: 122

Type locality: Coast of Liguria

Distribution: Czech Rep., Italy, Morocco

Microcaeculus monticolus (Lawrence)

Caeculus monticolus Lawrence, 1939: 539

Microcaeculus monticolus (Lawrence): Franz, 1952: 122; Halliday, 2005: 28

Type locality: South Africa

Distribution: South Africa

Microcaeculus muscorum (Lucas)

Caeculus muscorum Lucas, 1849: 307

Microcaeculus muscorum (Lucas): Franz, 1952: 122

Type locality: Algeria

Distribution: Algeria

Microcaeculus namibensis Piffl

Microcaeculus namibensis Piffl, 1965: 1

Sclerocaeculus namibensis (Pifl): Taylor et al., 2013: 449

Type locality: Namibia

Distribution: Namibia

Microcaeculus nicoleae Coineau

Microcaeculus nicoleae Coineau, 1974b: 276

Type locality: Bloeddrif

Distribution: South Africa

Microcaeculus nudus Franz

Microcaeculus nudus Franz, 1964: 88

Type locality: Serro El Roble, Cordillera de la Costa, Provincias de Valparaiso y .Santiago, Gipfel del Cerro

Distribution: Chile

Microcaeculus pica Otto

Microcaeculus pica Otto, 2009: 6

Type locality: New South Wales, Guerilla Bay

Distribution: Australia (NSW)

Microcaeculus pisanus (André)

Caeculus pisanus André, 1936: 123; Franz, 1952: 122 (Inc. sed.)

Microcaeculus pisanus (André): Taylor et al., 2013: 449

Type locality: Italy

Distribution: Italy

Microcaeculus sabulicola Franz

Microcaeculus sabulicola Franz, 1952: 107; Coineau, 1969b: 207

Type locality: “Andalusien, am Sandstrand westl. Algeciras”(Spain)

Distribution: France, Morocco, Portugal, Spain

Microcaeculus sinensis (Jacot)

Caeculus sinensis Jacot, 1936: 22

Microcaeculus sinensis (Jacot): Franz, 1952: 122

Type locality: China

Distribution: China

Microcaeculus sinensis taishanicus (Jacot)

Caeculus sinensis taishanicus Jacot, 1936: 22

Microcaeculus sinensis taishanicus (Jacot): Franz, 1952: 122

Type locality: Taishan

Distribution: China

Microcaeculus steineri Franz

Microcaeculus hispanicus steineri Franz, 1955: 9

Microcaeculus steineri Franz:

Type locality: Sierra Nevada, 2700 m

Distribution: Spain

Microcaeculus steineri delamarei Coineau

Microcaeculus steineri delamarei Coineau, 1974b: 276

Type locality: Font Estramer, 12 km N from Perpignan

Distribution: France

Microcaeculus stephani Coineau et Haupt

Microcaeculus stephani Coineau & Haupt, 1978: 622

Type locality: Formentera, Playa Mitjora

Distribution: Spain (Formentera, Pityusas)

Microcaeculus tibbettsi (Higgins et Mulaik)

Caeculus tibbettsi Higgins & Mulaik, 1958: 17

Microcaeculus tibbettsi (Higgins et Mulaik): Franz, 1962: 100

Type locality: Seoul

Distribution: Korea

Microcaeculus uchidai (Asanuma)

Caeculus uchidai Asanuma, 1940: 271

Microcaeculus uchidai (Asanuma): Taylor et al., 2013: 449

Type locality: Japan

Distribution: Japan

Genus Neocaeculus Coineau

Neocaeculus Coineau, 1967: 58

Type species: Neocaeculus luxtoni Coineau, 1967

Neocaeculus bornemisszai Coineau et Enns

Neocaeculus bornemisszai Coineau & Enns, 1969: 690

Type locality: Kimberley, 100 km S Wyndham

Distribution: Australia

Neocaeculus bruchi (Berlese)

Caeculus bruchi Berlese, 1916: 289; Franz, 1952: 122 (Inc. sed., Microcaeculus ?)

Neocaeculus bruchi : Coineau, 1974: Taylor et al., 2013: 449

Type locality: “La Plata”

Distribution: “La Plata”(Argentina)

Neocaeculus imperfectus Taylor, Gunawardene et Kinnear

Neocaeculus imperfectus Taylor, Gunawardene & Kinnear, 2013: 440

Type locality: Barrow Island, Western Australia

Distribution: Australia

Neocaeculus johnstoni Coineau

Neocaeculus johnstoni Coineau, 1974: 68

Type locality: Lord Howe Island

Distribution: Lord Howe Island

Neocaeculus kinnearae Taylor

Neocaeculus kinnearae Taylor, 2014: 348

Type locality: Barrow Island, Western Australia

Distribution: Australia

Neocaeculus knoepffleri Coineau et Enns

Neocaeculus knoepffleri Coineau & Enns, 1969: 679

Type locality: Kimberley, 100 km S Wyndham

Distribution: Australia

Neocaeculus lamorali Coineau

Neocaeculus lamorali Coineau, 1974b: 278

Type locality: De Waal farm, 10 km N of Leonardville

Distribution: Namibia

Neocaeculus luxtoni Coineau

Neocaeculus luxtoni Coineau, 1967a: 58

Type locality: Alexandra (Otago)

Distribution: New Zealand

Neocaeculus nudonates Taylor

Neocaeculus nudonates Taylor, 2014: 343

Type locality: Barrow Island, Western Australia

Distribution: Australia

Neocaeculus orientalis Fuangarworn et Butcher

Neocaeculus orientalis Fuangarworn & Butcher, 2015: 251

Type locality: Thailand

Distribution: Thailand

Neocaeculus setecidades Ott et Ott

Neocaeculus setecidades Ott & Ott, 2018: 2

Type locality: Parque Nacional Sete Cidades (04°06’44.2”S - 41°41’47.7”W), sample site PNC7C 0323, Piracuruca, Piauí

Distribution: Brazil

Neocaeculus womersleyi Coineau

Neocaeculus womersleyi Coineau, 1974: 68

Type locality: Risdon

Distribution: Tasmania

Genus Procaeculus Jacot

Procaeculus Jacot, 1936: 25

Type species: Procaeculus bryani Jacot, 1936

Procaeculus aitkeni Coineau

Procaeculus aitkeni Coineau, 1967b: 719

Type locality: Trinidad

Distribution: Trinidad

Procaeculus brevis (Mulaik)

Caeculus brevis Mulaik, 1945: 11; Franz, 1952: 121; McDaniel & Boe, 1990: 723

Procaeculus brevis (Mulaik): Taylor et al., 2013: 449.

Type locality: Houston, Harris Co., Texas

Distribution: USA (Texas)

Procaeculus bryani Jacot

Procaeculus bryani Jacot, 1936: 25; Franz, 1952: 121; Coineau, 1967b: 713

Type locality: Peale Island, Wake Island, Western end of the Hawaiian Island group

Distribution: Hawaiian Islands

Procaeculus coineaui Porta et al. (fossil)

Procaeculus coineaui Porta et al., 2016: 38

Type locality: Myanmar

Distribution: Myanmar (Burma)

Procaeculus dominicensis Coineau et Poinar (fossil)

Procaeculus dominicensis Coineau & Poinar, 2000: 143

Type locality: Mine in Northern Cordillera, El Mamey Formation (Upper Eocene)

Distribution: Dominican Republic

Procaeculus eridanosae Coineau et Magowski (fossil)

Procaeculus eridanosae Coineau & Magowski, 1994: 243

Type locality: Baltic amber (Eocene)

Distribution: Poland

Procaeculus magnus Coineau

Procaeculus magnus Coineau, 1974b: 279

Type locality: Mazatlan, Sin.

Distribution: Mexico

Procaeculus mexicanus (Mulaik et Allred)

Caeculus mexicanus Mulaik & Allred, 1954: 34; McDaniel & Boe, 1990: 723

Procaeculus mexicanus (Mulaik et Allred): Coineau, 1967b: 718

Type locality: San Luis Potosi (also intercepted in Texas)

Distribution: Mexico

Procaeculus orchidicolis (Mulaik et Allred)

Caeculus orchidicolis Mulaik & Allred, 1954: 30

Procaeculus orchidicolis (Mulaik et Allred): Coineau, 1967b: 718

Type locality: Chilpancingo (also intercepted in Texas)

Distribution: Mexico

Procaeculus oregonus (Mulaik et Allred)

Caeculus oregonus Mulaik & Allred, 1954: 28; McDaniel & Boe, 1990: 723

Procaeculus oregonus (Mulaik et Allred): Coineau, 1967b: 718

Type locality: Oak Ridge, Oregon

Distribution: USA (Oregon)

Procaeculus potosi (Mulaik et Allred)

Caeculus potosi Mulaik & Allred, 1954: 32

Procaeculus potosi (Mulaik et Allred): Coineau, 1967: 718

Type locality: Tamazunchale (also intercepted in Texas)

Distribution: Mexico

Procaeculus puertoricus Mulaik

Caeculus puertoricus Mulaik, 1945: 12; Franz, 1952: 122

Procaeculus puertoricus (Mulaik): Taylor et al., 2013: 450

Type locality: Como Springs

Distribution: Puerto Rico

Procaeculus willmanni (Vitzthum)

Caeculus willmannii Vitzthum, 1933: 86

Procaeculus willmanni (Vitzthum): Franz, 1952: 121

Procaeculus aitkeni Coineau, 1967a: 719; syn. by Taylor et al., 2013: 450

Type of P. aitkeni: Bush-Bush Forest, Trinidad

Type locality: Guatemala

Distribution: Guatemala

Incertae sedes

Caeculus hirtipes Berlese

Caeculus hirtipes Berlese, 1910: 369; Franz, 1952: 122 (Inc. sed.)

Type locality: “Mechieria”

Distribution: Algeria

Caeculus hirtipes graecus Berlese

Caeculus hirtipes var. graecus Berlese, 1910: 369; Franz, 1952: 122 (Inc. sed.)

Type locality: Naxos

Distribution: Greece

Distribution of Caeculidae

EUROPE

Austria (Irk, 1939; Willmann, 1951; Franz, 1952, 1955; Thaler, 1979; Thaler, Knoflach & Meyer, 1993) – Allocaeculus relictus, Caeculus echinipes, Microcaeculus austriacus

Bulgaria (Kratochvil, 1962) – Caeculus echinipes

Croatia (Vitzthum, 1933) – Allocaeculus spatulifer

Czech Rep. (Franz, 1952; Winkler, 1953; Willmann, 1954) – Allocaeculus relictus, Microcaeculus austriacus, M. liguricus

France (Moniez, 1893; André, 1932; Jacot, 1936; Franz, 1954; Coineau, 1966, 1969b, 1974) – Allocaeculus catalanus, Caeculus echinipes, C. echinipes crosbyi, Microcaeculus hispanicus, M. sabulicola, M. steineri delamarei

Corsica (Coineau, 1968) - Microcaeculus franzi

Germany (Pieper, 1970; Coineau & Haupt, 1976) – Caeculus echinipes, Microcaeculus austriacus

Greece (Berlese, 1910; Sellnick, 1931; Franz, 1964) – Caeculus echinipes, Microcaeculus graecus; inc. sed.: Caeculus hirtipes var. graecus

Italy (Vitzthum, 1933; Franz, 1954) – Allocaeculus sculptus, Caeculus echinipes, Microcaeculus liguricus, M. pisanus

Sicily (Berlese, 1883) – Caeculus echinipes

Hungary (Karpelles, 1893) - Caeculus

N. Macedonia (Franz, 1960) – Allocaeculus schusteri

Poland (Coineau & Magowski, 1994) - Procaeculus eridanosae (fossil)

Portugal (Coineau, 1969b) – Microcaeculus sabulicola

Salvages Islands (André, 1935) – Microcaeculus insolitus

Slovakia (Mangová, Krumpál & Ľuptáčik, 2014) – Allocaeculus sandbergensis

Spain (Dufour, 1832; Vitzthum, 1933; Franz, 1952, 1954, 1955, 1960; Mihelčič, 1958) – Allocaeculus andalusiacus, A. catalanus, A. echinatus, A. hystriciformis, A. meseticola, A. multispinosus, A. spatulifer, A. spinosissimus (incl. A. spinosissimus rondae), Caeculus echinipes, Microcaeculus hispanicus, M. sabulicola, M. steineri

Canary Is. (Franz, 1965) – Allocaeculus tenerifae, Microcaeculus canariensis

Balear and Pityusas Islands (Coineau & Haupt, 1977) – Microcaeculus stephani [Formentera]

Switzerland (Schweizer, 1951; Schweizer & Bader, 1963; Pieper, 1970) – Caeculus echinipes

Turkey (Karaca et al., 2012; Per, Doğan, Zeytun & Ayyıldız, 2017; Per, Doğan & Ayyıldız, 2019) – Allocaeculus multispinosus, A. turcicus

AFRICA

Algeria (Lucas, 1849; Michael, 1890; Kulczynski, 1901; André, 1927; 1935) – Allocaeculus dubius, A. hoggarensis, A. muscorum, A. spatulifer, Caeculus echinipes

Chad (Franz, 1957, 1964) – Allocaeculus logonensis, A. ouadaiensis, A. tschadensis

Kenya (Franz, 1964) - Allocaeculus kenyae

Morocco (Coineau, 1969b; Franz, 1960, 1964) – Allocaeculus grandjeani, A. kocheri, A. sarhroi, A. tschadensis rifensis, Microcaeculus insularis, M. liguricus, M. sabulicola

Mozambique (André, 1936) - Allocaeculus mosambicensis

Namibia (Lawrence, 1938; Piffl, 1965; Coineau, 1974b) – Allocaeculus circinatus, A. pilosus, Microcaeculus namibensis, Neocaeculus lamorali

Nigeria (Coineau, 1974b) – Allocaeculus nigeriensis

Senegal (André, 1938; Franz, 1964) – Allocaeculus sp. cf. andalusiacus, A. tschadensis, A. senegalensis

Somalia (Berlese, 1916) – “Caeculus pr. spathuliger Michael”

South Africa (Lawrence, 1939; Coineau, 1974b; Halliday, 2005) – Allocaeculus erinaceus, A. hirsutus, A. hystrix, A. kalahariensis, Calocaeculus lawrencei, Microcaeculus andrei, M. deserticolus, M. monticolus, M. nicoleae

Sudan (Trägårdh, 1904) - Allocaeculus sudanensis

Tunisia (André, 1936) – Allocaeculus sclerodermatus

ASIA

China (Jacot, 1936) - Microcaeculus laoshanensis, M. sinensis

Iran (Paktinat-Saeij, Ghobari & Samani, 2020) – Allocaeculus sirwani

Japan (Asanuma, 1940) – Microcaeculus uchidai

KazachstanAllocaeculus catalanus

Korea (Higgins & Mulaik, 1958) – Caeculus tibbettsi

Myanmar (Burma)(Porta et al., 2019) – Procaeculus coineaui

Pakistan (Piffl, 1959) – Allocaeculus indicus

Thailand (Fuangarworn & Butcher, 2015) – Neocaeculus orientalis

NORTH AMERICA

Canada (Bernard, Lumley, Buck & Cobb, 2020) - Caeculus cremnicolus, C. cassiopeiae

Mexico (Mulaik & Allred, 1954; Coineau, 1974b; Rivas et al., 2016; Porta & Vázquez Rojas, 2020) – Caeculus veracruzanus, Procaeculus magnus, P. mexicanus, P. orchidicolis, P. potosi

United States (Banks, 1899, 1906; Nevin, 1943, 1945; Mulaik, 1945; Mulaik & Allred, 1954; Higgins & Mulaik, 1957a, 1957b; Enns, 1958; Coineau, 1974b; Hagan, 1985; McDaniel & Boe, 1990; Skvarla, Fisher & Dowling, 2013) – Caeculus americanus, C. archeri, C. calechius, C. clavatus, C. cremnicolus, C. dorotheae, C. janetae, C. gertschi, C. krantzi, C. lewisi, C. mariae, Procaeculus brevis, P. oregonus

Hawaii (Jacot, 1936) – Procaeculus bryani

CENTRAL and SOUTH AMERICA

Argentina (Berlese, 1916; Franz, 1962, 1964; Porta, Pizarro-Araya & Ramírez, 2021) – Andocaeculus beatrizrosso, A. brundini, A. burmeisteri, A. weinrauchi, Neocaeculus bruchi

Brazil (Ott & Ott, 2014, 2018) – Andocaeculus caioi, Neocaeculus setecidades

Chile (Franz, 1964) – Microcaeculus castrii, M. nudus

Dominican Republic (Coineau & Poinar, 2000) – Procaeculus dominicensis (fossil)

Guatemala (Vitzthum, 1933) – Procaeculus willmanni

Puerto Rico (Mulaik, 1945) – Procaeculus puertoricus

Trinidad (Coineau, 1967b) – Procaeculus aitkeni

OCEANIA

Australia (Coineau & Enns, 1970; Otto, 1993; Taylor, Gunawardene & Kinnear, 2013; Taylor, 2014) – Microcaeculus pica, Neocaeculus bornemisszai, N. imperfectus, N. kinnearae, N. knoepffleri, N. nudonates

Tasmania (Coineau, 1974) - Neocaeculus womersleyi

New Zealand (Coineau, 1967a) – Neocaeculus luxtoni

Lord Howe Island (Coineau, 1974) - Neocaeculus johnstoni

Bibliography of Caeculidae

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The mite Superfamily Adamystoidea

Saxidromoidea Coineau, 1974a: 1059

Adamystoidea Cunliffe, 1957

The mite Superfamily Adamystoidea contains only one family – Adamystidae.

The mite family Adamystidae

Rare or little studied free-living mites, first annouced by Cunliffe (1957) and Coineau (1974a, 1974b, 1976). They are known so far only from 14 countries, but are certainly much more widespread. Ueckermann (1989) revised the family and described six new species, five of them sympatric in the area of Edenville, Orange Free State, South Africa. Another two new genera have been described fom South Africa by Coineau, Theron & Fernandez (2006). Rafalski (1982) described one very high living species in Hindu Kush (Adamystis coineaui, 5100 m). Other researchers described species from Middle Asia (Uzbekistan, Tajikistan, Barilo, 1986), USA (McDaniel & Bolen, 1983), Ukraine (Pogrebnyak, 2007), Cuba (Palacios-Vargas & Prieto, 1995), Mexico (López-Campos, 1996), Iran (Beyzavi, Ueckermann & Ostovan 2012; Fuangarworn & Beyzavi, 2012; Khanjani, Alvandy, Asalifayaz & Ueckermann, 2012), and Thailand (Fuangarworn & Lekprayoon, 2010). So far only 28 species are known in this family, from the genera Adamystis Cunliffe, 1957 (21 sp.), Nannodromus (1 sp.), Saxidromus Coineau, 1974 (4 sp.), Bovidromus Coineau, Theron et Fernandez, 2006 (1 sp.) and Rhinodromus Coineau, Theron et Fernandez, 2006 (1 sp.)

Catalogue of Adamystidae

Family Adamystidae Cunliffe

Adamystinae Cunliffe, 1957: 172

Saxidromidae Coineau, 1974a: 1059

Adamystidae: Coineau, 1974b: 431

Type genus: Adamystis Cunliffe, 1957

Subfamily Adamystinae Cunliffe

Adamystinae Cunliffe, 1957: 172

Type genus: Adamystis donnae Cunliffe, 1957

Genus Adamystis Cunliffe

Adamystis Cunliffe, 1957: 172

Type species: Adamystis donnae Cunliffe, 1957

Adamystis alvandicus Khanjani, Alvandy, Asali Fayaz et Ueckermann

Adamystis alvandicus Khanjani, Alvandy, Asali Fayaz & Ueckermann, 2010: 32

Type locality: Spiny Hawthorn trees, Crataegus aronia (L.) (Rosaceae), highland of Alvand Mountain, Hamedan, Hamedan province (34°39.8’27”N, 48°23’0.35”E).

Distribution: Iran

Adamystis beckyanneae McDaniel et Bolen

Adamystis beckyanneae McDaniel & Bolen, 1983: 177; Ueckermann, 1989: 236

Type locality: Olton Town, Lamb County, Texas

Distribution: USA (Texas)

Adamystis buchelis Ueckermann

Adamystis buchelis Ueckermann, 1989: 229

Type locality: Edenville, Orange Free State

Distribution: South Africa

Adamystis coineaui Rafalski

Adamystis coineaui Rafalski, 1982: 135; Ueckermann, 1989: 236

Type locality: Valley of Mandaras, Valley of Darum, High Hindu Kush, 5100 m

Distribution: Pakistan

Adamystis donnae Cunliffe

Adamystis donnae Cunliffe, 1957: 172; Ueckermann, 1989: 236

Type locality: Tahoe City, California

Distribution: USA (California)

Adamystis doumengei Coineau

Adamystis doumengei Coineau, 1974: 1061; Ueckermann, 1989: 239

Type locality: Ermelo Road, Transvaal

Distribution: South Africa

Adamystis ekaterinae Ghasemi-Moghadam et Saboori

Adamystis ekaterinae Ghasemi-Moghadam & Saboori, 2019: 14

Type locality: Chitgar Forest Park

Distribution: Iran

Adamystis fonsi Coineau

Adamystis fonsi Coineau, 1974: 1061; Ueckermann, 1989: 239; Pogrebnyak, 2007: 13

Type locality: Banyuls-sur-mer, Eastern Pyrenees

Distribution: France, Ukraine

Adamystis iranoturanianensis Beyzavi, Ueckermann et Ostovan

Adamystis iranoturanianensis Beyzavi, Ueckermann & Ostovan, 2012: 28

Type locality: from soil and humus under oak trees (Quercus sp., Fagaceae), Kamfiruze, Fars province (30°7.15’55”N, 52°15’56.17”E).

Distribution: Iran

Adamystis monomaculata Ueckermann

Adamystis monomaculata Ueckermann, 1989: 232

Type locality: Edenville, Orange Free State; Potchefstroom, Transvaal

Distribution: South Africa

Adamystis nudis Ueckermann

Adamystis nudis Ueckermann, 1989: 231

Type locality: Edenville, Orange Free State

Distribution: South Africa

Adamystis oculeusis Ueckermann

Adamystis oculeusis Ueckermann, 1989: 226

Type locality: Edenville, Orange Free State

Distribution: South Africa

Adamystis persiangulfensis Paktinat-Saeij et Kazemi

Adamystis persiangulfensis Paktinat-Saeij & Kazemi, 2021: 260

Type locality: Qeshm Island

Distribution: Iran

Adamystis saboorii Paktinat-Saeij et Ahaniazad

Adamystis saboorii Paktinat-Saeij & Ahaniazad, 2019: 133

Type locality: Yarshahr village, Maragheh city, East Azerbaijan Province

Distribution: Iran

Adamystis sarae Hunter et Crossley

Adamystis sarae Hunter & Crossley, 1968: 181; Ueckermann, 1989: 229

Type locality: Savannah River Plant Reservation near Aiken, South Carolina

Distribution: USA

Adamystis sartorum Barilo

Adamystis sartorum Barilo, 1986: 1414

Type locality: Samarkand, Buhara (Uzbekistan)

Distribution: Tadjikistan, Uzbekistan

Adamystis thailandensis Fuangarworn et Lekprayoon

Adamystis thailandensis Fuangarworn & Lekprayoon, 2010: 61

Type locality: Pang-Nga Province, Thaimuang District, Hard-Thaimuang Khao-Champee National Park, 8°29’38”N, 98°13’39”E

Distribution: Thailand

Adamystis theroni Fuangarworn et Beyzavi

Adamystis theroni Fuangarworn & Beyzavi, 2012: 449

Type locality: Zagros mountain, south of Kamfiruz region, Fars province (30°8’N, 52°17’E).

Distribution: Iran

Adamystis trimaculata Ueckermann

Adamystis trimaculata Ueckermann, 1989: 239

Type locality: Edenville, Orange Free State

Distribution: South Africa

Adamystis ueckermanni Fuangarworn et Beyzavi

Adamystis ueckermanni Fuangarworn & Beyzavi, 2012: 452

Type locality: Zagros mountain, south of Kamfiruz region, Fars province (30°8’N, 52°17’E).

Distribution: Iran

Adamystis yvesi Ueckermann

Adamystis yvesi Ueckermann, 1989: 236

Type locality: Pongola, Transvaal

Distribution: South Africa

Subfamily Saxidrominae Coineau

Saxidrominae Coineau, 1974a: 1059

Saxidromidae: Coineau, 1974a: 1059; 1974b: 431

Type genus: Saxidromus Coineau, 1974

Genus Bovidromus Coineau, Theron et Fernandez

Bovidromus Coineau, Theron & Fernandez, 2006: 67

Type species: Bovidromus roussouwi Coineau, Theron et Fernandez, 2006

Bovidromus roussouwi Coineau, Theron et Fernandez

Bovidromus roussouwi Coineau, Theron et Fernandez, 2006: 67

Type locality: South Africa

Distribution: South Africa

Genus Nannodromus Fernandez, Coineau, Theron et Tiedt

Nannodromus Fernandez, Coineau, Theron & Tiedt, 2014: 19

Type secies: Nannodromus reveilleti Fernandez, Coineau, Theron et Tiedt, 2014

Nannodromus reveilleti Fernandez, Coineau, Theron et Tiedt

Nannodromus reveilleti Fernandez, Coineau, Theron & Tiedt, 2014: 19

Type locality: Kruger National Park, N’Wanetsi: S24°27’30.56”, E31°58’35.30”

Distribution: South Africa

Genus Rhinodromus Coineau, Theron et Fernandez

Rhinodromus Coineau, Theron & Fernandez, 2006: 73

Type species: Rhinodromus lootsi Coineau, Theron et Fernandez, 2006

Rhinodromus lootsi Coineau, Theron et Fernandez

Rhinodromus lootsi Coineau, Theron & Fernandez, 2006: 73

Type locality: South Africa

Distribution: South Africa

Genus Saxidromus Coineau

Saxidromus Coineau, 1974: 1060

Type species: Saxidromus delamarei Coineau, 1974

Saxidromus annahoffmannae Lopes-Campos

Saxidromus annahoffmannae Lopes-Campos, 1996: 215

Type locality: Veracruz

Distribution: Mexico

Saxidromus caribeus Palacios-Vargas et Prieto

Saxidromus caribeus Palacios-Vargas & Prieto, 1995: 213

Type locality: Santiago de las Vegas, Habana region

Distribution: Cuba

Saxidromus delamarei Coineau

Saxidromus delamarei Coineau, 1974: 1060; Coineau & Naudo, 1986: 303; Ueckermann, 1989: 227

Type locality: Banyuls-sur-Mer, Eastern Pyrenees

Distribution: France

Saxidromus knoepffleri Coineau

Saxidromus knoepffleri Coineau, 1974: 1061; Ueckermann, 1989: 227

Type locality: Ermelo Road, Melspruit, Transvaal

Distribution: South Africa

Distribution of Adamystidae

EUROPE

France (Coinneau, 1974) – Adamystis fonsi, Saxidromus delamarei

Ukraine (Pogrebnyak, 2007) - Adamystis fonsi

AFRICA

South Africa (Coineau, 1974; Ueckermann, 1989; Coineau, Theron & Fernandez, 2006; Fernandez, Coineau, Theron & Tiedt, 2014) – Adamystis buchelis, A. doumengei, A. monomaculata, A. nudis, A. oculensis, A. trimaculata, A. yvesi, Bovidromus roussouwi, Nannodromus reveilleti, Rhinodromus lootsi, Saxidromus knoepffleri

ASIA

Iran (Beyzavi, Ueckermann & Ostovan 2012; Fuangarworn, Beyzavi & Ostovan, 2012; Khanjani, Alvandy, Asalifayaz & Ueckermann, 2012; Paktinat-Saeij et al., 2019; Paktinat-Saeij & Kazemi, 2021) – Adamystis alvandicus, A. ekaterinae, A. iranoturanianensis, A. persiangulfensis, A. saboorii, A. theroni, A. ueckermanni

Japan (Shiba, 1999) – Adamystidae gen. sp.

Pakistan (Rafalski, 1982) – Adamystis coineaui

Tadjikistan (Barilo, 1986) – Adamystis sartorum

Thailand (Fuangarworn & Lekprayoon, 2010) - Adamystis thailandensis

Uzbekistan (Barilo, 1986) - Adamystis sartorum

NORTH AMERICA

Mexico (López-Campos, 1996) – Saxidromus annahoffmannae

United States (Cunliffe, 1957; Hunter & Crossley, 1968; McDaniel & Bolen, 1983) – Adamystis beckyanneae, A. donnae, A. sarae

Hawaii (Swift, 2000) – Adamystidae gen. sp.

CENTRAL and SOUTH AMERICA

Cuba (Palacios-Vargas & Prieto, 1995) – Saxidromus caribeus

French Antilles (Saint-Martin)(Coineau, 1979) – Adamystidae gen. sp.

OCEANIA

Australia (Walter & O’Dowd, 1995) – Adamystidae gen. sp.

Bibliography of Adamystidae

  • Alberti G., Y. Coineau, N. Fernandez, P.D. Theron. 2010. On the female genital system of Saxidromidae (Prostigmata Actinotrichida). – XIII International Congress of Acarology, Recife, Brazil, 23 – 27 August.
  • Barilo A.B. 1987. On the mites of the family Adamystidae (Acariformes, Prostigmata) found in Middle Asia. – Zool. Zhurnal, 66(9): 1414 -1416.
  • Beyzavi G., E.A.Ueckrmann, H. Ostovan. 2012. A new Adamystis Cunliffe, 1957 species from Iran (Acari: Trombidiformes: Adamystidae). - Systematic & Applied Acarology, 17: 27–30.
  • Coineau Y. 1974a. Un nouveau type d’Acariens Prostigmates libres: les Saxidromoidea nouvelle super-famille. – C.R. Acad. Sci., 278: 1059 - 1062.
  • Coineau Y. 1976. Les pariades sexuelles des Saxidrominae Coineau 1974 (Acariens Prostigmates, Adamystidae). – Acarologia, 18(2): 234 - 240.
  • Coineau Y. 1979. Les Adamystidae, une étonnante famille d’Acariens Prostigmates primitifs. – Proc. 4th Intern. Congress of Acarology, Saalfelden, 1974: 431 - 435.
  • Coineau Y., P.D. Theron, N. Fernandez. 2006. Parades et dimorphismes sexuels comparés chez deux nouveaux genres de Saxidromidae (Acari: Alycina) d’Afrique du Sud. – Acarologia, 46(1-2): 65 – 87.
  • Cunliffe F. 1957. Notes on the Anystidae with a description of a new genus and species, Adamystis donnae, and a new subfamily, Adamystinae (Acarina). – Proc. Entomol. Soc. Wash., 59(4): 172 - 175.
  • Fernandez N., Y. Coineau, P.D. Theron, L. Tiedt. 2014. Nannodromus reveilleti (Acari, Anystida, Saxidromidae) a new genus and species from South Africa. – ZooKeys, 378: 17–39.
  • Fuangarworn M., G. Beyzavi, H. Ostovan. 2012. Adamystis Cunliffe, 1957 (Acari: Prostigmata: Adamystidae) in Iran: two new species and a key to the Iranian species. – Systematic & Applied Acarology, 17: 448–457.
  • Fuangarworn M., Ch. Lekprayoon. 2010. Adamystis thailandensis sp. nov. (Acari: Prostigmata: Adamystidae), a new species of soil mites from Thailand with a key to world species of Adamystidae. – Zootaxa, 2649: 61 - 68.
  • Ghasemi-Moghadam, A. Ahadiyat, A. Saboori, Y. Fathipour. 2019. A new species of Adamystis (Acari: Trombidiformes: Adamystidae) from Iran. – Zootaxa, 4647(1): 14 - 22.
  • Hunter P.E., D.A. Crossley. 1968. Adamystis sarae, a new species of soil mite from cornfield litter in South Carolina (Acarina: Anystidae). – J. Georgia Entomol. Soc., 3(4): 181 - 183.
  • Khanjani M., S. Alvandy, B. Asalifayaz, E.A. Ueckermann. 2012. A new species of the genus Adamystis (Acari: Anystina: Adamystidae) from western Iran. – Systematic & Applied Acarology, 17: 31–35.
  • López-Campos M.G. 1996. Especie nueva de Saxidromus (Prostigmata: Adamystidae) del Estado de Veracruz, México. – Anales Inst. Biol. Univ. Nac. Autón. México, Ser. Zool., 67(2): 213 - 223.
  • McDaniel B., E.G. Bolen. 1983. A new species of Adamystis (Acari: Adamystidae) from the southern high plains of Texas. – Proc. Entomol. Soc. Wash., 85(1): 177 – 179.
  • Paktinat-Saeij S., M. Ahaniazad, M. Bagheri, M. R. Damavandian. 2019. A new species of the genus Adamystis Cunliffe (Acari: Trombidiformes: Adamystidae) from Iran, with a key to world species of the genus. - Systematic & Applied Acarology, 24(1): 132–138.
  • Paktinat-Saeij S., Sh. Kazemi. 2021. A new species of Adamystis Cunliffe (Acari: Trombidiformes: Adamystidae) from mangrove in the Persian Gulf, Iran, with a key to world species of the genus. - Systematic & Applied Acarology, 26(1): 260 – 271.
  • Palacios-Vargas J., T.D. Prieto. 1995. A new species of Saxidromus (Endeostigmata: Adamystidae) from Cuba. – Acarologia, 36(3): 213 - 217.
  • Pogrebnyak S. 2007. Adamystis fonsi (Prostigmata, Adamystidae) external morphology peculiarities – new for Ukraine family of Prostigmatid mites. – Zbirnik praz Zoologichnogo muzeyu, 39: 13 – 20.
  • Rafalski J. 1982. Adamystis coineaui sp. n. (Acari, Adamystidae) from Hindu-Kush. – Acarologia, 23(2): 135 - 143.
  • Shiba M. 1999. Prostigmata. In: Aoki J. (Ed.), Pictorial Key to Soil Animals of Japan. Tokay Univ. Press, Tokyo, pp. 211 – 311.
  • Swift S.F. 2000. New records of Hawaiian prostigmatid mites (Acari: Prostigmata). – Bishop Museum Occasional Papers, 64: 50 – 51.
  • Ueckermann E.A. 1989. A revision of the family Adamystidae Cunliffe (Acari: Prostigmata). – Phytoprophylactica, 21: 227 - 240.
  • Walter D.E., D.J. O’Dowd. 1995. Life on the forest phylloplane: hairs, little houses, and myriad mites. In: Lowman M.E. & N. Nadkarni (Eds.), Forest Canopies. Academic Press, New York, pp. 325 – 351.
  • Walter D.E., E.E. Lindquist, I.M. Smith, D.R. Cook, G.W. Krantz. 2009. Order Trombidiformes. In: Krantz, G.W. & Walter, D.E. (Eds.) A manual of acarology. Third edition. Lubbock, Texas Tech University Press, pp. 233–420.

The mite Superfamily Pomerantzioidea

The subfamily contains only one family – Pomerantziidae.

The mite Family Pomerantziidae

Small family (six known species), established by Baker (1949) for one species, found In Georgia (USA). Price (1971, 1974, 1975) added new species from the USA.

Quing-Hai Fan & Yan Chen (2005) revised the family and separated two of the species of Price in a new genus Apomerantzia. One new species (Pomerantzia subterranea) was found in deep humus in Fuzhou, China, another one (Pomerantzia philippina Bochkov et Walter, 2007) – in the Philippines.

Catalogue of Pomerantziidae

Family Pomerantziidae

Pomerantziidae Baker, 1949: 269

Type genus: Pomerantzia Baker, 1949

Genus Apomerantzia Quing-Hai Fan et Yan Chen

Apomerantzia Quing-Hai Fan & Yan Chen, 2005: 8

Type species: Pomerantzia prolata Price, 1971

Apomerantzia kethleyi (Price)

Pomerantzia kethleyi Price, 1975: 487; Kethley, 1989: 101

Apomerantzia kethleyi (Price): Quing-Hai Fan & Yan Chen, 2005: 7

Type locality: Indiana, Lake Co., St. John

Distribution: USA (Indiana, Illinois, Minnesota)

Apomerantzia prolata (Price)

Pomerantzia prolata Price, 1971: 395

Apomerantzia prolata (Price): Quing-Hai Fan & Yan Chen, 2005: 5

Type locality: California, Nevada Co.

Distribution: USA (California)

Genus Pomerantzia Baker

Pomerantzia Baker, 1949: 269

Type species: Pomerantzia charlesi Baker, 1949

Pomerantzia benhami Price

Pomerantzia benhami Price, 1974: 425

Type locality: California, Fresno Co.

Distribution: USA (California)

Pomerantzia charlesi Baker

Pomerantzia charlesi Baker, 1949: 269; Price, 1974: 420

Type locality: Upson County, Georgia

Distribution: USA (Georgia)

Pomerantzia philippina Bochkov et Walter

Pomerantzia philippina Bochkov & Walter, 2007: 159

Type locality: Romblon Prov., Sibuyan Isl, Magdiwang, beach near of Puto

Point, 12°26’N, 122°41’E, soil

Distribution: the Philippines

Pomerantzia subterranea Quing-Hai Fan et Yan Chen

Pomerantzia subterranea Quing-Hai Fan & Yan Chen, 2005: 8

Type locality: Fuzhian

Distribution: China

Distribution of Pomerantziidae

ASIA

China (Quing-Hai Fan & Yan Chen, 2005) – Pomerantzia subterranea

Philippines (Bochkov & Walter, 2007) – Pomerantzia philippina

NORTH AMERICA

United States (Baker, 1949; Price, 1971, 1974, 1975; Price & Benham, 1976) – Apomerantzia kethleyi, A. prolata, Pomerantzia benhami, P. charlesi

Bibliography of Pomerantziidae

  • Baker 1949. Pomerantziidae, a new family of prostigmatic mites. – J. Wash. Acad. Sci., 39(8): 269 - 271.
  • Bochkov A.V., D.E. Walter. 2007. The life-cycle of Pomerantzia philippina sp.n. (Prostigmata: Pomerantziidae) described from the Philippines. – Acarina, 15(1): 159 – 170.
  • Fan Quing-Hai, Yan Chen. 2005. A review of the Pomerantziidae (Acari: Prostigmata: Pomerantzioidea), with the description of a new genus. – Zootaxa, 1037: 1 - 22.
  • Kethley J. 1989. Occurrence of Pomerantzia kethleyi (Acari: Prostigmata: Pomerantziidae) in Illinois and Minnesota. - The Great Lakes entomologist, 22(2): 101.
  • Kethley J. 1990. Acarina: Prostigmata (Actinedida). In: Dindal D.L. (Ed.). Soil Biology Guide. New York: John Wiley and Sons. p. 667-756.
  • Price D. W. 1971. A new species of Pomerantzia Baker from California (Acarina: Pomerantziidae). – Proc. ent. Soc. Wash., 73(4): 394 - 398.
  • Price D. W. 1974. Notes on the genus Pomerantzia Baker, with a description of second species from California. – Proc. ent. Soc. Wash., 76(4): 419 - 427.
  • Price D. W. 1975. Notes on the genus Pomerantzia Baker with a description of a new species from Indiana. - Proc. ent. Soc. Wash., 77(4): 487 - 490.
  • Price D.W., G.S.Jr. Benham. 1976. Vertical distribution of pomerantziid mites (Acarina: Pomerantziidae).- Proceedings of the Entomological Society of Washington, 78: 487–490.

The series Acarorum Catalogus – supplementa and corrigenda in the former volumes

In the period of 14 years (2008 - 2021) of publication of my 9 volumes of the series Acarorum Catalogus, the fast development of the Acarology brought to light many new taxa of Acari and many other mites have been re-arranged or synonymized. This task requires a supplement to the published volumes (one, concerning Vol.I, has been already published). These supplements will be sertainly followed by many other changes, but it would be proper to fix by 2021 the situation within the 97 families of mites considered in the series.

First update of Volume I:

Beron P. 2017. Acarorum Catalogus I Acariformes: Calyptostomatoidea (Calyptostomatidae), Erythraeoidea (Smarididae, Erythraeidae). Pensoft & Nat. Mus. Natur. Hist. Sofia: supplement 1. - Historia naturalis bulgarica, 24: 131 – 154.

Second update (2017 – 2021)

Beron P. 2008. Acarorum Catalogus I Acariformes: Calyptostomatoidea (Calyptostomatidae), Erythraeoidea (Smarididae, Erythraeidae). Pensoft & Nat. Mus. Natur. Hist. Sofia: 271 pp.

Introduction

Many specialists from Poland, Montenegro, Iran, Turkey, China, Brazil, and other countries added new information to the knowledge on these groups. As a result of the recent contributions, the number of genera and species in Calyptostomatoidea and Erythraeoidea (some of them of uncertain status) is updated as follows:

Calyptostomatoidea

Calyptostomatidae – one genus, ten species

Erythraeoidea

Smarididae – 10 genera, 59 species

Erythraeidae – 57 genera, ca. 890 species

New contributions to the Catalogue

Fam. Calyptostomatidae

Calyptostoma seemani Masoumi et Saboori

Calyptostoma seemani Masoumi & Saboori, 2020: 1823

Type locality: Kandovan region, Alborz province

Distribution: Iran

Erythraeoidea

Fam. Smarididae

Unlike most terrestrial parasitengone mites (chiggers, erythraeids), known mostly from parasitic and easy-to-collect larvae, smaridid systematics are mainly based on the post larval instar (Dos Santos, Welbourn, Klimov & Pepato, 2021).

Clavismaris maquine Ott et Ott

Clavismaris maquine Ott & Ott, 2018: 2

Type locality: Atlantic forest area at FEPAGRO Maquiné, Rio Grande do Sul

Distribution: Brazil

Hirstiosoma baenai (Mayoral & Barranco, 2017) in place of Sphaerotarsus baenai Mayoral & Barranco, 2017 (Dos Santos et al., 2021)

Hirstiosoma quercus (Yazdanpanah, Saboori & Hakimitabar, 2016) in place of Sphaerotarsus quercus Yazdanpanah, Saboori & Hakimitabar, 2016 (Dos Santos et al., 2021)

Sphaerotarsus sadafae Amin, Khanjani et Nadri

Sphaerotarsus sadafae Amin, Khanjani & Nadri, 2020: 1965

Type locality:

Distribution: Iran

Genus Southcottiana Dos Santos, Welbourn, Klimov et Pepato

Southcottiana Dos Santos, Welbourn, Klimov & Pepato, 2021: 104

Type species: Sphaerotarsus monticolus Southcott, 1997

Southcottiana monticola (Southcott)

Sphaerotarsus monticolus Southcott, 1997: 6

Southcottiana monticola (Southcott, 1997): Dos Santos, Welbourn, Klimov & Pepato, 2021: 104

Type locality: Mt. Lofty, S. Australia

Disribution: Australia (S. Australia)

Trichosmaris calcarensis Dos Santos, Welbourn, Klimov et Pepato

Trichosmaris calcarensis Dos Santos, Welbourn, Klimov & Pepato, 2021: 108

Type locality: cave, 5 km E Pain Municipality, Minas Gerais

Distribution: Brazil

Trichosmaris paulensis Dos Santos, Welbourn, Klimov et Pepato

Trichosmaris paulensis Dos Santos, Welbourn, Klimov & Pepato, 2021: 112

Type locality: Americana Municipality, São Paulo State

Distrinution: Brazil

According to Dos Santos, Welbourn, Klimov & Pepato (2021), Surasmaris Southcott, 1995 is a junior synonym of Trichosmaris Southcott, 1963 and Surasmaris longirostris Southcott, 1995 becomes Trichotarsus longirostris

Fam. Erythraeidae

Erythraeinae

Collemboerythraeus izadpanahi Noei, Sabbori, Sedghi et Hakimitabar

Collemboerythraeus izadpanahi Noei, Sabbori, Sedghi & Hakimitabar, 2017: 2

Type locality: Simkan region, Jahrom city, Fars province

Distribution: Iran

Host: Collembola: Sminthuridae

Curteria duzgunesae (Saboori, Çobanoğlu et Bayram, 2007) – recorded on Uroleucon sp. (Homoptera: Aphidae) by Oner et al. (2021)

Erythraeus (E.) ankaraicus Saboori, Çobanoğlu et Bayram, 2007 - recorded also on Uroleucon aeneum, U. sonchi, Aphis craccivora, A. nerii, Aulacorthum solani,

Brachycaudus sp., Hyperomyzus lactucae, Macrosiphoniella sanborni (Homoptera: Aphidae) by Oner et al. (2021).

Erythraeus (E.) cinereus (Dugès) – Montenegro (Haitlinger & Šundiċ, 2018a)

Erythraeus (E.) hubeiensis Xu, Yi, Guo et Jin

Erythraeus (E.) hubeiensis Xu, Yi, Guo & Jin, 2019: 55

Type locality: Qizimeishan National Natural Reserve, Hubei Province

Distribution: China

Host: Typhlocybini (Cicadellidae: Typhlocybinae)

Erythraeus (E.) moeritzensis Haitlinger, 2007 – Albania (Haitlinger & Šundiċ, 2018a)

Erythraeus (E.) southcotti Goldarazena et Zhang, 1998 - Albania (Haitlinger & Šundiċ, 2018a)

Erythraeus (E.) regalis (C. L. Koch, 1837) – Montenegro, Greece (Lesbos) (Haitlinger & Šundiċ, 2018a)

Erythraeus (E.) phalangoides (De Geer, 1778) - new for Turkey, recorded on Myzus sp., Sitobion sp. and Urocyon sonchi (Homoptera: Aphidae) by Oner et al. (2021)

E.(E.) uhadi Kamran et Alatawi, 2014 – new for Turkey, recorded on Uroleucon sonchi (Homoptera: Aphidae) by Oner et al. (2021)

Erythraeus (Parerythraeus) thomasi Roland et Gabryś

Erythraeus (Parerythraeus) thomasi Roland & Gabryś, 2020: 326

Type locality: Biogeographic province: Varsinais-Suomi, loc.:670:25 V, Kaarina Kuusiston Kartano, Kuiva keto

Distribution: Finland

Note: the subgenus Parerythraeus is reinstalled as subgenus, out of synonymy with Erythraeus, by Roland & Gabryś (2020).

Erythraeus (Zaracarus) coleopterus Mortazavi, Hajikanbar et Saboori, 2012 - new for Turkey, recorded on Uroleucon sonchi (Homoptera: Aphidae) by Oner et al. (2021)

E. (Z.) iranicus Saboori et Akrami, 2001 - new for Turkey, recorded on Uroleucon sonchi and Aphis nerii (Homoptera: Aphidae) by Oner et al., (2021)

E. (Z.) kurdistaniensis Khanjani et Ueckermann, 2005 - new for Turkey, recorded on Brachycaudus tragopogonis, Hyperomyzus lactucae and Uroleucon sonchi (Oner et al., 2021)

E. (Zaracarus) hainanensis Xu, Yi, Guo et Jin

E. (Zaracarus) hainanensis Xu, Yi, Guo & Jin, 2019: 64

Type locality: Bawangling National Natural Reserve, Hainan Province

Distribution: China (Hainan)

Host: planthoppers (Hemiptera: Delphacidae)

Erythraeus (Zaracarus) lancifer Southcott, 1995 – Albania (Haitlinger & Šundiċ, 2018a: 1199)

Erythraeus (Zaracarus) ueckermanni is considered a valid species and not a synonyme of E. budapestensis by Saboori & Shirvani (2021)

Lasioerythraeus jessicae Dos Santos Costa

Lasioerythraeus jessicae Dos Santos Costa 2019: 184

Type locality: 19°05’46.0”S, 43°21’25.0”W, in an iron ore terrain cave, Conceição do Mato Dentro municipality, Minas Gerais State

Distribution: Brazil

Leptinae

Leptus (Leptus) andongensis Hakimitabar, Joharchi et Jung

Leptus (L.) andongensis Hakimitabar, Joharchi & Jung, 2020: 1

Type locality: Andong

Distribution: S. Korea

Host: Meimuna sp. (Insecta: Homoptera: Cicadidae)

Leptus (Leptus) brasilicus Haitlinger, Šundiċ et Pompermaier

Leptus (Leptus) brasilicus Haitlinger, Šundiċ & Pompermaier. 2017: 874

Type locality: 30 km northeast of Brasilia

Distribution: Brazil

Leptus (Leptus) cameroonicus Haitlinger, Šundiċ et Nkwala

Leptus (L.) cameroonicus Haitlinger, Šundiċ & Nkwala, 2020: 608

Type locality: forest reserve of the town Mbalmayo (3º28’34.218” N; 11º29’37.416E)

Distribution: Cameroon

Host: Eyprepocinemis plorans ibandana (Orthoptera: Acrididae)

Leptus (Leptus) candangus Šundiċ, Haitlinger et Pompermaier

Leptus candangus Šundiċ, Haitlinger & Pompermaier, 2017: 90

Type locality: Serrado-Pianaltinade (pitfall traps)

Distribution: Brazil

Leptus (Leptus) chingombensis Bernard, Felska et Makol

Leptus (Leptus) chingombensis Bernard, Felska & Makol, 2019: 794

Type locality: Mikwa stream, Michinga Escarpment, 1160 m

Distribution: Zambia

Hosts: Heliaeschna fuliginosa (Odonata: Aeschnidae), Orthetrum julia (Libellulidae), Paragomphus cognatus (Gomphidae), Pseudagrion spernatum (Coenagrionidae)

Leptus (Leptus) darvishi Saboori, Hakimitabar et Khademi

Leptus (Leptus) darvishi Saboori, Hakimitabar & Khademi, 2018: 95

Type locality: Komaijan and Mahallat cities, Markazi Province

Distribution: Iran

Host: undetermined grasshoppers (Orthoptera: Acrididae) and undetermined beetles (Coleoptera: Buprestidae and Tenebrionidae)

Leptus (Leptus) flechtmanni Bassini-Silva, Jacinavicius et Barros-Battesti

Leptus (L.) flechtmanni Bassini-Silva, Jacinavicius & Barros-Battesti, 2020: 2

Type locality: Serra dos Carajás,Parauapebas, Pará, Brazil, (6° 6’29’’S, 50° 18’16’’W)

Distribution: Brazil

Host: Unidentified opilion (Opiliones: Cosmetidae)

Leptus (Leptus) grancanaricus Haitlinger et Šundiċ

Leptus (Leptus) grancanaricus Haitlinger & Šundiċ, 2020: 836

Type locality: Gran Canaria, Tejada: Mirador de Timagada, altitude 1,258 m

Distribution: Canary Islands

Host: Herpisticus guanarteme (Coleoptera: Curculionidae)

Leptus (L.) guarani Haitlinger, Šundiċ, Ázara et Bernardi

Leptus (L.) guarani Haitlinger, Šundiċ, Ázara & Bernardi, 2020: 1923

Type locality: State Parana, Morretes

Distribution: Brazil

Hosts: Acanthogonyleptes editus (Opiliones: Gonyleptidae)

Leptus (L.) haitlingeri Jacinavicius, Bassini-Silva et Barros-Battesti

Leptus (L.) haitlingeri Jacinavicius, Bassini-Silva & Barros-Battesti, 2018: 975; Bassini-Silva et al., 2020: 8

Type locality: Serra da Cantareira, Săo Paulo municipality, State of Săo Paulo

Distribution: Brazil

Hosts: Fidena (Fidena) sp. (Diptera: Tabanidae); Panstrongylus megistus (Hemiptera: Reduviidae), Dysdercus sp. (Hemiptera: Pyrrhocoridae), Obiricodesmus rosascens (Diplopoda, Polydesmida: Chelodesmidae)

Leptus (Leptus) machadoi Haitlinger et Šundiċ

Leptus (Leptus) machadoi Haitlinger & Šundiċ, 2020: 839

Type locality: Fuerteventura, Jandia, Pico de la Zarza, 807 m

Distribution: Canary Islands

Host: Laparocerus maxorata (Coleoptera: Curculionidae)

Leptus (Leptus) mariae – Drosophila subobscura (Diptera: Drosophilidae) (Haitlinger & Šundiċ, 2018a: 1199)

Leptus (Leptus) molochinus – new for Slovakia (Haitlinger & Šundiċ, 2021)

Leptus (Leptus) planaltensis Haitlinger, Šundiċ et Pompermaier

Leptus planaltensis Haitlinger, Šundiċ & Pompermaier, 2017: 879

Type locality: Brazil

Distribution: Brazil

Leptus (Leptus) pouryayevalii Hakimitabar, Saboori et Fadaei

Leptus pouryayevalii Hakimitabar, Saboori & Fadaei, 2021: 138

Type locality: Koushk Bala Village, Alborz Prov., Chalous road, Karaj, 1824 m

Distribution: Iran

Host: shaking the leaves of weeds

Leptus (L.) sidorchukae dos Santos Costa

Leptus (L.) sidorchukae dos Santos Costa, 2019: 159

Type locality: 19°34’36.0”S, 44°01’13.0”W, in an unnamed limestone cave, Pedro Leopoldo municipality, Minas Gerais State

Distribution: Brazil

Leptus sp. on Chrysops brunneus and Hybomitra lasiophthalma (Diptera: Tabanidae, Canada)(Mullen et al., 1989)

Abrolophinae

Abrolophus bochkovi Hakimitabar, Saboori et Fadaei

Abrolophus bochkovi Hakimitabar, Saboori & Fadaei, 2020: 108

Type locality: Qazvin province

Distribution: Iran

Abrolophus diaoluoensis Xu et Jin

Abrolophus diaoluoensis Xu & Jin, 2021: 24

Type locality: Hainan, Diaoluoshan National Natural Reserve

Distribution: China (Hainan)

Abrolophus hajikanbar Noei

Abrolophus hajikanbar Noei, 2022: 226

Type locality: Chahardeh and Chahkand villages, Birjand, South Khorasan province

Distribution: Iran

Abrolophus karamani Haitlinger et Šundiċ (syn.)

Abrolophus karamani Haitlinger & Šundiċ, 2019: 108; the species is considered as a junior synonym of A. aitapensis (Southcott 1948)(Hakimitabar, Saboori & Fadaei, 2020b).

Type locality: Port Louis

Distribution: Guadeloupe

Abrolophus marianopolicus Haitlinger et Šundiċ

Abrolophus marianopolicus Haitlinger & Šundiċ, 2018b: 354.

Type locality: 5 km north of Marianopoli (prov. Caltainisetta)

Distribution: Sicily

Abrolophus poljankensis Haitlinger et Šundiċ

Abrolophus poljankensis Haitlinger & Šundiċ, 2021: 39

Type locality: Nižna Poljanka n. Bardejov (Slovakia)

Distribution: Poland, Slovakia

Abrolophus quisquiliarus (Hermann, 1804) – Albania (Haitlinger & Šundiċ, 2018a: 1202); Canary Islands (La Gomera)(Haitlinger, 2019, first finding in Canaries and Africa)

Abrolophus wratislaviensis (Haitlinger, 1986) – Calabria, Sicily (Haitlinger & Šundiċ, 2018a)

Abrolophus zelandicus Luxton

Abrolophus zelandicus Luxton, 1989: 416; 1990: 398

Type locality: Auckland, Nelson (on a sea shore)

Distribution: New Zealand

Marantelophus iranicus (Haitlinger et Saboori, 1996) – Albania ((Haitlinger & Šundiċ, 2018a: 1199)

Charletonia elbasani Šundić, Haitlinger et Milosević

Charletonia elbasani Šundić, Haitlinger & Milosević, 2017: 564

Type locality: Elbasan

Distribution: Albania

Host: on grasses

Charletonia kalithensis Haitlinger, 2006 – Albania, Montenegro (Haitlinger & Šundiċ, 2018a: 1199)

Charletonia braunsi (Oudemans, 1910) – Cameroon (Haitlinger, Šundiċ & Nkwala, 2020)

Callidosomatinae

Caeculisoma allopenlineatus Xu et Jin

Caeculisoma allopenlineata Xu & Jin, 2020: 286

Type locality: Diaoluoshan National Natural Reserve, Hainan Province

Distribution: China

Hosts: Hemiptera: Cicadellidae, Delphacidae

Caeculisoma penlineatus Xu et Jin

Caeculisoma penlineatus Xu & Jin, 2019c: 511; Xu et al., 2020: 295

Type locality: Chongqing Municipality

Distribution: China (Chongqing Municipality, Guizhou Prov., Hainan Prov.)

Hosts: Mileewa margheritae (Hemiptera: Cicadellidae), Gen. sp. (Hem.: Delphacidae)

Caeculisoma semispinus Xu et Jin

Caeculisoma semispinus Xu & Jin, 2019d: 561

Type locality: Qizimeishan National Reserve, Hubei Province

Distribution: China

Host: Shaddai sp. (Hemiptera: Cicadellidae)

Dambullaeus adonis Makol et Moniuszko

Dambullaeus adonis Makol & Moniuszko, 2014: 5

Type locality: Western Madagascar

Distribution: Madagascar

Host: Latois nigrolineata (Hemiptera: Fulgoromorpha: Flatidae)

Dambullaeus hainanensis Xu et Jin

Dambullaeus hainanensis Xu & Jin, 2019: 456

Type locality: Hainan

Distribution: China

Dambullaeus jianfengensis Xu et Jin

Dambullaeus jianfengensis Xu & Jin, 2020: 297

Type locality: Jianfengling National Natural Reserve, Hainan Province

Distribution: China (Hainan)

Iguatonia seemani Xu et Jin

Iguatonia seemani Xu & Jin, 2020: 306

Type locality: Jianfengling National Natural Reserve, Hainan Province

Distribution: China (Hainan)

Iguatonia xinfengi Xu et Jin

Iguatonia xinfengi Xu & Jin, 2020: 315

Type locality: Jianfengling National Reserve, Hainan Province

Distribution: China (Hainan)

Balaustiinae

Balaustium akramii Noei

Balaustium akramii Noei, 2017: 2218; Oner et al., 2021: 2

Type locality: Sari city, Mazandaran province, Iran

Distribution: Iran, Turkey

Host: Uroleucon sanchi (Homoptera: Aphidae)

Balaustium izmirensis Noei et Ersin

Balaustium izmirensis Noei & Ersin, 2019: 31

Type locality: Bornova, Izmir

Distribution: Turkey

Balaustium medicagoense Meyer et Ryke, 1959 – new for New Zealand (Xu & Zhang, 2019)

Balaustium ryszardi Šundiċ et Noei

Balaustium ryszardi Šundiċ & Noei, 2021: 2609

Type locality: Peloponese

Distribution: Greece

Distribution of Calyptostomatoidea and Erythraeoidea (suppl.)

EUROPE

Albania (Šundiċ, Haitlinger & Milošević, 2017; Haitlinger & Šundiċ, 2018a) - Abrolophus quisquiliarus, Charletonia elbasani, Erythraeus (Zaracarus) lancifer, E. (E.) moeritzensis, E. (E.) southcotti, Marantelophus iranicus, Charletonia kalithensis

Finland (Roland & Gabryś, 2020) – Erythraeus (Parerythraeus) thomasi

Greece (Haitlinger & Šundiċ, 2018a; Šundiċ & Noei, 2021) – Balaustium ryszardi, Abrolophus anzelmi, A. petanovicae, A. wratislaviensis, Erythraeus (E.) regalis

Hungary (Haitlinger & Šundiċ, 2018) - Charletonia cardinalis, Moldoustium baltiensis

Italy (Haitlinger & Šundiċ, 2018a) - Abrolophus wratislaviensis, A. anzelmi, A. dagmarae

Sicily (Haitlinger & Šundiċ, 2018b) - Abrolophus marianopolicus, A. wratislaviensis

Montenegro (Haitlinger & Šundiċ, 2018a) - Erythraeus (E.) cinereus, E. (E.) regalis, Charletonia kalithensis

Poland (Haitlinger & Šundiċ, 2021) – Abrolophus poljankus

Slovakia (Haitlinger & Šundiċ, 2021) - Abrolophus poljankus, Leptus molochinus

Turkey (Gencsoylu, 2007; Noei et al., 2017; Noei, Ersin & Çakmak, 2019; Oner et al., 2021) – Balaustium akrami, B. izmirensis, E.(E.) phalangoides, E.(E.) uhadi, E. (Zaracarus) coleopterus, E.(Z.) iranicus, E. (Z.) kurdistaniensis. Charletonia krendowskyi

AFRICA

Canary Islands (Haitlinger, 2019, Haitlinger & Šundiċ, 2020) - Leptus (Leptus) grancanaricus, L.(L.) machadoi, Abrolophus quisquiliarus

Cameroon (Haitlinger, Šundiċ & Nkwala, 2020) – Leptus cameroonicus

Madagascar (Makol et al., 2014) – Dambullaeus adonis

Zambia (Bernard, Felska & Makol, 2019) – Leptus (L.) chingombensis

ASIA

China (Xu, Yi, Guo & Jin, 2019a, b, c, d, 2020, 2021) – Abrolophus diaoluoensis, Caeculisoma allopenlineatus, C. penlineatus, C. semispinus, Dambullaeus jianfengensis, Erythraeus (E.) hubeiensis

Hainan (Xu, Yi, Guo & Jin, 2020) - Caeculissoma penlineatus, Erythraeus (Z.) hainanensis, Dambullaeus hainanensis, Iguatonia seemani, I. xinfengi

India (Gupta et al., 2004; Gupta A. et al., 2006) - Erythraeidae

Iran (Noei et al., 2017; Noei, Maroufpoor, Faizi & Ostovan, 2017; Noei & Asadollahzadeh et al., 2017; Saboori et al., 2018; Hakimitabar, Saboori & Saaveh, 2018; Noei, Saboori, Sedghi & Hakimitabar, 2018; Noei & Rabieh, 2019; Noei & Honarmand. 2020; Hakimitabar, Saboori & Fadaei, 2020, 2021; Masoumi, Saboori & Seiedy, 2020; Amin, Khanjani & Nadri, 2020; Saboori & Shirvani, 2021, Noei, 2022) – Calyptostoma seemani; Smarididae: Sphaerotarsus sadafae; Erythraeidae: Abrolophus bochkovi, Balaustium akrami, Collemboerythraeus izadpanahi, Leptus darwishi, L. pouryayevalii

South Korea (Hakimitabar, Johardii & Jong, 2020) – Leptus andongensis

NORTH AMERICA

Canada (Mullen et al. 1989) – Leptus sp.

Mexico (Montiel-Parra & Pérez-Ortiz, 2007; Chaires-Grijalva et al., 2021) – Leptus

indianensis, L. lomani, L. onnae, L. ruginus

SOUTH and CENTRAL AMERICA

Argentina (Cédola, Nolasco & Minardi, 2018) – Balaustium sp.

Bolivia (Torrico-Bazoberry et al., 2020) – Leptus hringuri

Brazil (Pinto et al., 2014; Šundić et al., 2017; Ott & Ott, 2018; Jacinavicius, 2019; Haitlinger, Šundiċ & Pompermaier, 2017; Šundiċ, Haitlinger & Pompermaier, 2017; Guimarraes et al., 2019; Bassini-Silva et al., 2019, 2020; Haitlinger et al., 2020; Dos Santos et al., 2021) – Smarididae: Clavismaris maquine, Trichosmaris calcarensis, T. paulensis; Erythraeidae: Leptus brasilicus, L. candangus, L. guarani, L. flechtmanni, L. haitlingeri, L. planaltensis, L. simonettae,

Costa Rica (Dos Santos et al., 2021) – Trichosmaris longirostris

OCEANIA

Australia (Dos Santos et al., 2021) - Southcottiana monticola

New Zealand (Luxton, 1989, 1990; Xu & Zhang, 2020) – Abrolophus zelandicus, Balaustium medicagoense

Hosts

Arachnida

Opiliones

Fam. Cosmetidae

Acromares vittatum - Leptus ca. indianensis

Cynorta sp. - Leptus ruginus

Holovonones compressus - Leptus ruginus

Heterovonones incrassatus – Leptus indianensis

Vonones pelaezi - Leptus ruginus

Unidentified opilion – Leptus flechtmanni

Fam. Gonyleptidae

Acanthogonyleptes editusLeptus (Leptus) guarani

Caldassius nigripes – Leptus sp.

Gonyleptes gonyleptoides - Leptus sp.

Moreiranula mamillata - Leptus sp.

Fam. Stygnopsidae

Paramitraceras granulatus - Leptus ruginus

Fam. Sclerosomatidae

Metopilio sp. - Leptus ruginus

Leiobunum nycticorpum - Leptus ruginus

Paranelima sp. - Leptus ruginus

Prionostemma sp. - Leptus ruginus, L. nr. indianensis

Krusa sp. - Leptus ruginus

Scorpiones

Fam. Buthidae

Centruroides limpidus limpidus – Leptus ruginus

Fam. Caraboctonidae

Hadrurus gerstchi - Leptus ruginus

Fam. Vejovidae

Vejovis mexicanus – Leptus sp.

Vejovis spinigerus – Leptus sp.

Vejovis variegatus – Leptus ruginus

Amblypygi

Fam. Phrynidae

Phrynus garridoi – Leptus onnae

Paraphrynus pococki – Leptus onnae

Araneae

Fam. Filistatidae

Kukulcania sp. - Leptus ca. indianensis

Ixodida

Fam. Argasidae

Ornithodoros sp. - Leptus (Leptus) simonettae

Myriapoda

Diplopoda - Polydesmida

Fam. Chelodesmidae

Obiricodesmus rosascens - Leptus haitlingerii

Insecta s.l.

Collembola

Fam. Sminthuridae - Collemboerythraeus izadpanahi

Odonata

Fam. Aeschnidae

Heliaeschna fuliginosa - Leptus (Leptus) chingombensis

Fam. Gomphidae

Paragomphus cognatus - Leptus (Leptus) chingombensis

Fam. Libellulidae

Orthetrum julia - Leptus (Leptus) chingombensis

Fam. Coenagrionidae

Pseudagrion spernatum - Leptus (Leptus) chingombensis

Orthoptera

Fam. Acrididae

Eyprepocinemis plorans ibandana Charletonia braunsi, C. cameroonensis, C. justinae, Leptus cameroonicus

Zonocerus variegatus – Charletonia cameroonensis

Hemiptera Heteroptera

Fam. Reduviidae

Panstrongylus megistusLeptus haitlingerii

Fam. Pyrrhocoridae

Dysdercus sp. - Leptus haitlingerii

Hemiptera

Fam. Cicadellidae

Mileewa margheritaeCaeculisoma penlineatus

Shaddai sp. – Caeculisoma semispinus

Fam. Delphacidae

Gen. sp. - Caeculisoma penlineatus

Fam. Cicadidae

Meimuna sp. - Leptus (L.) andongensis

Fam. Membracidae

Alchisme grossa - Leptus hringuri

Fam. Flatidae

Latois nigrolineata - Dambullaeus adonis

Homoptera

Fam. Aphididae

Uroleucon (U.) sonchi - Balaustium akrami, Erythraeus ankaraicus, E. uhadi, E. (Z.) coleopterus, E.(Z.) iranicus, E.(Z.) kurdistaniensis

Uroleucon (U.) aeneum - Erythraeus ankaraicus

Uroleucon sp. – Curteria duzgunesae

Aphis craccivora - Erythraeus ankaraicus

A. nerii - Erythraeus ankaraicus, E.(Z.) iranicus

Aulacorthum solani - Erythraeus ankaraicus

Brachycaudus tragopogonis – Erythraeus (Z.) kurdistaniensis

Brachycaudus sp. - Erythraeus ankaraicus

Hyperomyzus lactucae - Erythraeus ankaraicus, Erythraeus (Z.) kurdistaniensis

Hyalopterus pruni - Erythraeus ankaraicus

Macrosiphoniella sanborni – Erythraeus ankaraicus

Myzus sp. – Erythraeus phalangoides

Sitobion sp. - Erythraeus phalangoides

Coleoptera

Fam. Curculionidae

Herpisticus guanartemeLeptus grancanaricus

Laparocerus maxorata - Leptus machadoi

Fam. Elateridae

Monocrepidius flavangulus - Leptus sp.

Glyphonyx flohri – Leptus sp.

Diptera

Fam. Tabanidae

Fidena (Fidena) sp. - Leptus haitlingeri

Chrysops brunneusLeptus sp.

Hybomitra lasiophthalmaLeptus sp.

Fam. Drosophilidae

Drosophila subobscura – Leptus mariae

Publications on Calyptostomatoidea and Erythraeoidea

(suppl. 2017 – 2021 and ommited)

  • Bassini-Silva R., F. de Castro Jacinavicius, S. Muñoz-Leal, F. A. Terassini, G.M. Barboza de Oliveira, C. Welbourn, M. Šundić, M. B. Labruna, D. M. Barros-Battesti. 2019. Leptus (Leptus) simonettae Haitlinger, 2000 (Trombidiformes: Erythraeidae) parasitizing a soft tick (Ixodida: Argasidae) in Brazil. - International Journal of Acarology, DOI: 10.1080/01647954.2019.1667433
  • Bassini-Silva R. et al. (9 other authors). 2020. A new species of Leptus (Leptus) (Trombidiformes: Erythraeidae) and new records of Leptus (Leptus) haitlingeri Jaciniavicius, Bassini-Silva & Welbourn, 2019 for Brazil. – Intern. Journal of Acarology, 46(4): 1 - 9.
  • Berbard M., M. Felska, J. Makol. 2019. Erythraeid larvae parasitizing dragonflies in Zambia – description of Leptus (Leptus) chingombensis sp. nov. with data on biology and ecology of host-parasite interactions. – Systematic & Applied Acarology, 24(5): 790 – 813.
  • Cédola C.V., V. Nolasco, G.M. Minardi. 2018. Trophic breadth niche, prey preference and developmental time of a Balaustium sp. (Acari: Erythraeidae) from Argentina. Persian Journal of Acarology, 7(3), 255–264.
  • Chaires-Grijalva M.P., I.M. Vázquez-Rojas, E.O. Martínez-Luque. 2021. New Host Data and Distribution for an Ectoparasitic Larva of Leptus at Tamaulipas, Mexico. - Southwestern Entomologist, 46(1): 265-270.
  • Dos Santos Costa S. G. 2019. Multi-instar descriptions of cave dwelling Erythraeidae (Trombidiformes: Parasitengona) employing an integrative approach. – Zootaxa, 4717(1): 137 – 184.
  • Dos Santos S.G., C. Welbourn, P. Klimov, A. R. Pepato. 2021. Integrating phylogeny, ontogeny and systematics of the mite family Smarididae (Prostigmata, Parasitengona): Classification, identification key, and description of new taxa. – Systematic and Applied Acarology, 26(1): 85 – 123.
  • Elverici M., E. Buga, S. Sevsay. 2022. A contribution to the knowledge of the genus Charletonia [курсив] Oudemans from Turkey. - Persian J. Acarol., 11(3): 471 - 482.
  • Gabryś G., W. Roland. 2018. A Redescription of Erythraeus italicus Berlese, 1920(Acari: Actinotrichida: Erythraeidae). - Annales Zoologici, 68(3): 421-429.
  • Gencsoylu I. 2007. A new species of ectoparasite, Erythraeus ankaraicus Saboori, Çobanoğlu and Bayram, on Asymmetrasca (= Empoasca) decedens incotton. - Phytoparasitica, 35: 433–435.
  • Gupta A., A. Chalterjee, A. K. Sanyal, S. K. Gupta. 2006. On a collection of soil prostigmatid mites (Acari) from southern parts of West Bengal, India. - Rec. Zool.Surv. India: l06(4): 61-71.
  • Gupta S.K., S. Ghoshal, A. Choudhury, B. Mukherjee. 2004. Phytophagous and predatory mite fauna of Sundarbans Biosphere Reserve: II. Some predatory mites occurring on mangrove vegetation and agri-horticultural crops. - Rec. zool. Surv.India, 103(3-4) : 33-45.
  • Guimarraes R.R., R.R. Guimarrães-Junior, H.R. Rodrigues, R.R. Guimarrães, R.W.Carvalho. 2019. First record of Leptus sp. (Latreille) (Acari: Erythraeidae), parasitizing horse fly (Diptera: Tabanidae) from Marambaia Island, Mangaratiba, Rio de Janeiro. Brazil. – Entomol. Bras., 12(1): 31–34.https://doi.org/10.12741/ebrasilis.v12i1.731
  • Haitlinger R. 2019. New records of mites from La Gomera, Canary Islands, Spain (Trombidiformes: Erythraeidae, Trombidiidae). - Ecologica Montenegrina, 20: 110–113.
  • Haitlinger R. 2020. New larval mites (Acari: Prostigmata: Erythraeidae,Microtrombidiidae, Trombidiidae) from Turkey, Peru and Poland. - Wiadomosci Parazytologiczne, 46(3): 379-396.
  • Haitlinger R., M. Šundiċ. 2018a. New records of mites from Albania, Greece(Lesbos), Italy and Montenegro, with notes on some species (Acari: Prostigmata: Erythraeidae, Microtrombidiidae, Neotrombidiidae, Trombellidae, Trombidiidae). –Linzer biol. Beitr., 50(1): 1195 -1212.
  • Haitlinger R., M. Šundiċ. 2018b. Abrolophus marianopolicus sp. nov.(Trombidiformes: Erythraeidae) from Sicily. Italy with notes on other Abrolophusspecies. – Syst. & Applied Acarology, 23(2): 353 – 359.
  • Haitlinger R., M. Šundiċ. 2019a. A new species of larval Abrolophus Berlese 1891 with notes on A. aitapensis (Southcott, 1948) (Acari, Prostigmata, Erythraeidae) from Guadeloupe (Lesser Antilles, France). - The Montenegrin Academy of Sciences and Arts Proceedings of the Section of Natural Sciences, 23: 107 – 117.
  • Haitlinger R., M. Šundiċ. 2019b. New records of mites (Trombidiformes:Erythraeidae,Microtrombidiidae, Trombidiidae) from Greece and Hungary and the list of terrestrial parasitengona found in both countries. - Agriculture & Forestry, 65(3): 51-63.
  • Haitlinger R., M. Šundiċ. 2020. Two new species of Leptus Latreille, 1796(Trombidiformes: Erythraeidae) from the Canary Islands, parasitising Curculionidae (Insecta: Coleoptera), with new metrical data for some Leptus spp. –Systematic Parasitology, 97: 835–846.
  • Haitlinger R., M. Šundiċ. 2021. A new larval Abrolophus Berlese, 1891 from Polandand Slovakia (Acari, Trombidiformes, Erythraeidae) – Spixiana, 44(1): 39-42.
  • Haitlinger R., M. Šundiċ, L. Ázara, L. F. O. Bernardi. 2020. A new species of larval Leptus (Leptus) (Trombidiformes: Erythraeidae) from Brazil with list of host-parasite associations between Leptus and arthropods in America. - Biologia(2020). https://doi.org/10.2478/s11756-020-00453-7
  • Haitlinger R., M. Šundiċ, A.L.D. Nkwala. 2020. Description of Leptus (Leptus) cameroonicus sp. nov. and first record of Charletonia braunsi (Oudemans, 1910) from Cameroon (Trombidiformes: Erythraeidae), with new metric and meristic datafor some African Leptus. - Systematic & Applied Acarology, 25(4): 607–621.
  • Haitlinger R., M. Šundiċ, V.T. Pompermaier. 2017. Two New Larval Leptus Latreille(Trombidiformes: Erythraeidae) From Brazil. - Systematic and Applied Acarology, 22(6): 874.
  • Hakimitabar M., O. Joharchi, C. Jung. 2020. A new species of. Leptus, the first erythraeid mite (Acari: Trombidiformes) from South Korea. - International Journal of Acarology, 47(4): 1–12.
  • Hakimitabar M., A. Saboori, H.M. Saaveh. 2018. New morphological data and new host record of Lasioerythraeus saboorii (Trombidiformes: Erythraeidae) from Iran.- Persian J. Acarol., 7(1): 41- 49.
  • Hakimitabar M., A. Saboori, E. Fadaei. 2020. New species of Abrolophus (Acari:Erythraeidae) from Iran with a key to species with a comb-like seta on the palpaltarsus. – Syst. & Applied Acarology, 25(12): 2300–2314.
  • Hakimitabar M., A. Saboori, E. Fadaei. 2021. A new species of Leptus (Acari:Erythraeidae) from Iran. – Persian Journal of Acarology, 10(2): 137 – 143.
  • Jacinavicius F. de Castro, R. Bassini-Silva, D. M. Barros-Battesti. 2019. Descriptionof Leptus (Leptus) haitlingeri n.sp. (Trombidiformes: Erythraeidae), parasitising horse flies (Diptera: Tabanidae), and a key to the larvae of Leptus spp. in Brazil. –Systematic Parasitology, 96(8): 723–734.
  • Luxton M. 1989. New taxa of intertidal mites. - Journal of Natural History, 23: 407-428.
  • Luxton M. 1990. The marine littoral mites of the New Zealand region. - Journal of theRoyal Society of New Zealand, 20(4): 367 - 418.
  • Mąkol J., H. Moniuszko, D. Świerczewski, A. Stroiński. 2014. Planthopper (Hemiptera: Flatidae) Parasitized by Larval Erythraeid Mite (Trombidiformes:Erythraeidae) - A Description of Two New Species From Western Madagascar. –Journal of insect science, 14(1): 1 – 12.
  • Masoumi H. R., A. Saboori, M. Seiedy. 2020. Descriptions of all active instars of anew species of Calyptostoma (Trombidiformes: Calyptostomatidae) from Iran. –Systematic and Applied Acarology, 25(10): 1821-1839.
  • Montiel-Parra G., T. Ma. Pérez-Ortiz. 2007. Nuevos registros de acaros de las familias Trombidiidae y Erythraeidae parasitos de Arácnidos en Mexico. – Entomología Mexicana 6. Editors: E. G. Estrada-Venegas, A. Equihua Martínez,C. Luna-León & J. L. Rosas-Acevedo.
  • Mullen G.L., D.J. Leprince, B.M. OConnor. 1989. Acarine associates of horse flies and deer flies (Diptera: Tabanidae) in North America. In: Channa Basavanna, G.P. & Viraktamath, C.A. (Eds) Progress in Acarology, 2,291- 296.
  • Noei J., S. Asadollahzadeh, I. Çakmak, A. Hadizadeh. 2017. A new larval species of Balaustium (Acari: Erythraeidae) from northern Iran and Turkey with a key to the genera of larval Balaustiinae and species of Balaustium. - Syst. & AppliedAcarology, 22: 2218-2232.
  • Noei J., F. Ersin, J. Çakmak. 2019. A new larval species of Balaustium (Acari:Actinotrichida: Erythraeidae) from Turkey. - Turkish Journal of Zoology, 43(1): 30 –42.
  • Noei J., A. Honarmand. 2020. Records of terrestrial parasitengone mites (Acari:Erythraeidae, Microtrombidiidae) ectoparasitic on grasshoppers from Khorasan Razavi province with new morphological data. – Pland Pest Res., 10(4): 15 – 26.
  • Noei J., M. Maroufpoor, F. Faizi, H. Ostovan. 2017. Second record of Erythraeus (Zaracarus) coleopterus (Acari: Erythraeidae) from Iran with new morphological data. – Persian Journal Acarology, 6(2): 71 – 80.
  • Noei J., M.M. Rabieh. 2019. New data on Nothrombidium, Southcottella and Eatoniana larvae (Acari: Trombellidae, Neothrombiidae, Erythraeidae) from Iran. –Persian J. Acarol., 8(3): 179 – 187.
  • Noei J., A. Saboori, A. Sedghi, M. Hakimitabar. 2018. A new larval species of the genus Collemboerythraeus (Acariformes: Erythraeidae) ectoparasitic on Collembola from Iran. - Syst. Appl. Acarol., 23(1): 1-12.
  • Noei J. 2022. A new species of larval Abrolophus () from Iran with a key to species without a comb-like seta on the palptarsus. – Persian Journal of Acarology,11(2), serial No41: 225 – 236. Oner D., Ş. Kök, A. Saboori, I. Cakmak. 2021. Mites parasitizing aphids in the parksand gardens of Aydin, with eight newly recorded mite species for Turkey and re-description of Allothrombium clavatum (Acari: Trombidiidae). - International Journalof Acarology, DOI: 10.1080/01647954.2021.1919758
  • Erythraeus (Erythraeus) phalangoides, E. (E.) uhadi, Erythraeus (Zaracarus)coleopterus, E. (Z.) iranicus, E. (Z.) kurdistaniensis (Erythraeidae) are new records to the Turkish fauna.
  • Ott A.P., R. Ott. 2018. A new species of velvet mite from meridional South America ofthe genus Clavismaris (Acari, Erythraeoidea, Smarididae, Hirstiosomatinae). – Iheringia, Série Zoologia, 108:
  • Pinto Z.T., C. Carriçao, R.L. Caetano, R.R. Barbosa, R.M. Mendonça, M.M. Queiroz.2014. First record of Leptus sp. Latreille, 1796 (Acari: Erythraeidae) fromItaipuaçu beach, Maricá, RJ, Brazil. Check List, 10(4): 954–956.
  • Roland W., G. Gabryś. 2020. Erythraeus (Parerythraeus) thomasi sp. nov. (Acari:Actinotrichida: Erythraeidae) from Finland, with a redefinition of Parerythraeus Southcott, 1946 stat. nov. as a subgenus and a key to species. – Zootaxa,4718(3): 324 – 336.
  • Saboori A., M. Hartmann, M. Hakimitabar, N. Khademi, A.-R. Katouzian. 2018. A newspecies of larval Leptus (Acari: Erythraeidae) from Markazi Province of Iran. –International Journal of Acarology, 44: 90 – 95.
  • Saboori A., M. Hakimitabar, N. Khademi, H. Masoumi, A.-R. Katouzian. 2020.Corrections and additions to Leptus Latreille (Trombidiformes: Erythraeidae) of the world: revised classification and keys. – Persian J. Acarol., 9(2): 209 – 212.
  • Saboori A., Z. Shirvani. 2021. A checklist of Acari type specimens deposited in the Jalal Afshar Zoological Museum, Karaj, Iran. - Zootaxa, 4949(2): 289–311.
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Update 2011 - 2021 to Vol. II

Beron P. 2011. Acarorum Catalogus. II. Acariformes Acaridida Listrophoroidea (Listrophoridae, Dromiciocoptidae, Myocoptidae, Chirodiscidae, Atopomelidae). Pensoft & Nat. Mus. Natur. Hist. Sofia: 263 pp.

Introduction

As a result of the recent contributions (mostly by A. Bochkov and co-authors), the number of genera and species in Listrophoroidea (some of them uncertain status) is updated as follows:

Listrophoridae – 24 genera, 177 species

Dromiciocoptidae – 1 genus, 3 species

Myocoptidae – 5 genera, 86 species

Chirodiscidae - 26 genera, 241 species

Atopomelidae - 44 genera, 413 species

Total 100 genera, 925 sp.

New contributions to the Catalogue

Fam. Listrophoridae

Afrolistrophorus (Afrolistrophorus) laonastes Bochkov, Abramov et Hugot

Afrolistrophorus (Afrolistrophorus) laonastes Bochkov, Abramov & Hugot, 2011: 11

Type locality: Kammouane Province, 18 km north of Thakhek, near Ban Doy village, 17º33’26’’N, 104º49’28’’E, 140 m a. s. l.

Distribution: Laos

Host: Laonastes aenigmamus (Rodentia: Diatomyidae)

Afrolistrophorus (Afrolistrophorus) hapalomys Bochkov et Abramov

Afrolistrophorus (Afrolistrophorus) hapalomys Bochkov & Abramov, 2014: 139

Type locality: Binh Phuoc Province, Bu Gia Map National Park, 13

km NE Bu Gia Map Village, 540 m a.l.s., 12°11’37”N, 107°12’21”E

Distribution: Vietnam

Host: Hapalomys delacouri Thomas (Rodentia, Muridae)

Afrolistrophorus (Afrolistrophorus) saxatilomys Bochkov, Abramov et Hugot

Afrolistrophorus (Afrolistrophorus) saxatilomys Bochkov, Abramov & Hugot, 2011: 14

Type locality: Kammouane Province

Distribution: Laos

Host: Saxatilomys paulinae (Rodentia: Muridae)

Amlistrophorus geoxus Sikora et Bochkov

Amlistrophorus geoxus Sikora & Bochkov, 2012: 390

Type locality: Los Lagos, Lago de Todoslos Santos, 41°06’S 72°12’W

Distribution: Chile

Host: Geoxus valdivianus (Rodentia: Cricetidae, Sigmodontinae)

Dubininetta abramovi Bochkov

Dubininetta abramovi Bochkov, 2017: 440

Type locality: 5 km east of Na Hang, Tuyen Quang Province, Vietnam; 22°20’59”N, 105°25’36”E

Distribution: Vietnam

Host: Euroscaptor subanura (Soricomorpha: Talpidae)

Dubininetta euroscaptor Bochkov

Dubininetta euroscaptor Bochkov, 2017: 438

Type locality: 14 km north of Kon Plong Town, Kon Plong District, Kon Tum Province, Vietnam; 14°43’20”N, 108°18’58”E

Distribution: Vietnam

Host: Euroscaptor parvidens (Soricomorpha: Talpidae)

Dubininetta taiwanensis (Fain & Lukoschus, 1978) – is recorded on Mogera robusta from the Far East of Russia by Bochkov (2017)

Leporacarus sylvilagi Fain, Whitaker et Lukoschus, 1981 on Sylvilagus brasiliensis (Linnaeus, 1758) (Lagomorpha: Leporidae) - on Minas Gerais and Rio Grande do Sul (Brazil)(Bochkov & Valim, 2016)

Lynxacarus (Lynxacarus) serrafreirei Bochkov et Valim

Lynxacarus (Lynxacarus) serrafreirei Bochkov & Valim, 2016: 30

Type locality: Rio de Janeiro

Distribution: Brazil

Host: Galictis cuja (Carnivora: Mustelidae)

Prolistrophorus akodon Fain and Lukoschus, 1982 from Akodon montensis from Argentina (Sikora & Bochkov, 2012)

Prolistrophorus amazonicus amazonicus Fain, 1971 – on Calomys callosus from Argentina and Bolivia, C. laucha and C. musculinus from Argentina (Sikora & Bochkov, 2012)

Prolistrophorus argentinus (Hirst, 1921) – on Holochilus brasiliensis and H. chacarius from Argentina (Sikora & Bochkov, 2012)

Prolistrophorus (Amlistrophorus) musculinus Fain, 1973 (formerly a subspecies of P. amazonicus) from Mus musculus (Rodentia: Muridae) from Suriname is raised to species status. The full generic status for the subgenus Amlistrophorus of the genus Prolistrophorus proposed by Fain et al.,1996 is not supported (Sikora & Bochkov, 2012)

P. nectomys Fain, 1971 – on Nectomys palmipes from Peru and Melanomys caliginosus from Panama (Sikora & Bochkov, 2012)

P. bidentatus Fain et Lukoschus, 1984 from Akodon cursor (Rodentia: Cricetidae) (new host-from Minas Gerais and Rio Grande do Sul, for the first time in Brazil) (Bochkov & Valim, 2016)

Prolistrophorus (Aprolistrophorus) parabidentatus Bochkov, Lareschi et Barreto

Prolistrophorus (Aprolistrophorus) parabidentatus Bochkov, Lareschi & Barreto, 2014: 590

Type locality: Argentina

Distribution: Argentina

Host: Akodon azarae (Rodentia: Cricetidae, Sigmodontinae)

Prolistrophorus (Aprolistrophorus) tylomys Bochkov, Lareschi et Barreto

Prolistrophorus (Aprolistrophorus) tylomys Bochkov, Lareschi & Barreto, 2014: 592

Type locality: Guatemala

Distribution: Guatemala

Host: Tylomys nudicaudus (Rodentia: Cricetidae)

Prolistrophorus (Beprolistrophorus) hirstianus Fain, 1973 from Scapteromys aquaticus from Argentina (Bochkov, Lareschi & Barreto, 2014)

Prolistrophorus (P.) argentinus (Hirst, 1921) – on Holochilus brasiliensis and H. chacarius from Argentina (Sikora & Bochkov, 2012); on Melanomys caliginosus, Akodon affinis from Colombia and Scapteromys aquaticus from Argentina (Bochkov, Lareschi & Barreto, 2014)

Prolistrophorus ctenomys Fain, 1970 on Ctenomys torquatus (Rodentia: Ctenomyidae) from Brazil (Bochkov & Valim, 2016)

Prolistrophorus (P.) grassii (Radford, 1954) on Zygodontomys brevicauda from Colombia (Bochkov, Lareschi & Barreto, 2014)

Prolistrophorus (Prolistrophorus) frontalis (Hirst, 1921) on Oligoryzomys sp. from Argentina (Bochkov, Lareschi & Barreto, 2014)

Prolistrophorus hylaeamys Bochkov et Valim

Prolistrophorus hylaeamys Bochkov & Valim, 2016: 27

Type locality: Minas Gerais

Distribution: Brazil

Host: Hylaeamys laticeps (Rodentia: Cricetidae, Sigmodontinae)

Prolistrophorus musculinus Fain, 1973 (formerly a subspecies of P. amazonicus) from Mus musculus (Rodentia: Muridae) from Suriname is raised to species status (Sikora & Bochkov, 2012)

Pteromychirus menetes Bochkov, Abramov et Hugot

Pteromychirus menetes Bochkov, Abramov & Hugot, 2011: 17

Type locality: Kammouane Province, 18 km north of Thakhek, near Ban Doy village, 17º33’26’’N, 104º49’28’’E, 140 m a. s. l.

Distribution: Laos

Host: Menetes berdmorei (Rodentia: Sciuridae)

Sclerolistrophorus oxymycteris Fain, 1976 – on Oryzomys laticeps from Brazil (Sikora & Bochkov, 2012)

Fam. Myocoptidae

Genus Apocalyptoides Bochkov

Apocalypsis Bochkov, 2010: 148 (praeoccupied name, not Butler, 1876, Lepidoptera).

Apocalyptoides Bochkov, 2016b: (replacement name)

Type species Criniscansor [not Gliricoptes !] allactaga Fain et Lukoschus, 1979

Apocalyptoides allactaga (Fain et Lukoschus)

Criniscansor [not Gliricoptes] allactaga Fain et Lukoschus, 1979: 236

Apocalypsis allactaga (Fain et Lukoschus): Bochkov, 2010: 148

Apocalyptoides allactaga (Fain et Lukoschus): Bochkov, 2016b:

Type locality: Char-nur

Distribution: Mongolia

Host: Allactaga sibirica (Rodentia: Dipodidae)

Criniscansor congolensis Fain, 1970 – on Grammomys macmillani from Tanzania (Bochkov, OConnor & Skoracki, 2016)

Gliricoptes graphiuri Fain, 1970 is synonymised with G. vulcanorum Fain, 1970 by Bochkov, OConnor & Skoracki, 2016

Gliricoptes vulcanorum Fain, 1970 in Tanzania (Bochkov, OConnor & Skoracki, 2016)

Myocoptes (M.) grammomys Fain, 1970 – on Grammomys dryas from Burundi and Uganda (Bochkov, OConnor & Skoracki, 2016)

Myocoptes (M.) lophuromys Bochkov, OConnor et Skoracki

Myocoptes (M.) lophuromys Bochkov, OConnor & Skoracki, 2016: 588

Type locality: Uganda

Distribution: Uganda

Host: Lophuromys voosnami (Rodentia: Muridae)

Myocoptes meriones Bochkov

Myocoptes (M.) meriones Bochkov, 2016b: 477

Type locality: Kirghizia

Distribution: Kirghizia, Tajikistan

Hosts: Meriones tamariscinus, M. libycus (Rodentia: Muridae)

Myocoptes (M.) spinulatus Fain, 1970 – on Dendromus mysticalis from Tanzania (Bochkov, OConnor & Skoracki, 2016)

Myocoptes (Comyoptes) nesokia Bochkov et OConnor

Myocoptes (Comyoptes) nesokia Bochkov & OConnor, 2017: 47

Type locality: Pakistan

Distribution: Pakistan

Host: Nesokia indica (Rodentia: Muridae)

Trichoecius angolensis Fain, 1972 is described for the first time by Bochkov, OConnor & Skoracki, 2016. Found in South Africa

Trichoecius dubininae Bochkov

Trichoecius dubininae Bochkov, 2016b: 462

Type locality: Chelyabinsk Province, Russia

Distribution: Kazakhstan, Kirghizia, Russia

Host: Lasiopodomys gregalis (Rodentia: Arvicolidae)

Trichoecius faini Bochkov, OConnor et Skoracki

Trichoecius faini Bochkov, OConnor & Skoracki, 2016: 595

Type locality: Tanzania

Distribution: Tanzania

Host: Hylomyscus arcimontanus (Rodentia: Muridae)

Trichoecius hylomyscus Bochkov, OConnor et Skoracki

Trichoecius hylomyscus Bochkov, OConnor & Skoracki, 2016: 593

Type locality: Tanzania

Distribution: Gabon, Tanzania

Hosts: Hylomyscus anselli, H. stella (Rodentia: Muridae)

Trichoecius lemmus Bochkov

Trichoecius lemmus Bochkov, 2016b: 466

Type locality: Taymyr peninsula

Distribution: Russia

Host: Lemmus sibiricus (Rodetia: Arvicolidae)

Trichoecius lootensis Fain, 1970 in Tanzania (Bochkov, OConnor & Skoracki, 2016)

Trichoecius meriones Bochkov

Trichoecius meriones Bochkov, 2016b: 468

Type locality: Turkmenia

Distribution: Turkmenia

Host: Meriones meridianus (Rodentia: Muridae)

Trichoecius otomys Fain, 1970 – on Otomys typus from Uganda (Bochkov, OConnor & Skoracki, 2016)

Fam. Chirodiscidae

Lutrilichus javanicus Fain, 1970 – on Melogale moschata from Vietnam (Bochkov, 2018)

Schizocarpus azasicus Bochkov et Saveljev

Schizocarpus azasicus Bochkov & Saveljev, 2012: 12

Type locality: Tuva, Todzhinskii District

Distribution: Russia (Tuva)

Host: Castor fiber tuvinicus (Rodentia: Castoridae)

Schizocarpus brachyurus (Dubinina, 1964)(= Sch intercalatus Fain et Lukoschus, 1985 = Sch. parabrachyurus Fain et Lukoschus, 1985) - on Castor fiber tuvinicus from Russia (Tuva)(Bochkov & Saveljev, 2012)

Schizocarpus faini Bochkov, Labrzycka, Skoracki et Saveljev

Schizocarpus faini Bochkov, Labrzycka, Skoracki & Saveljev, 2012: 46

Type locality: Suwalki

Distribution: Poland

Host: Castor fiber belorussicus (Rodentia: Castoridae)

Schizocarpus fedjushini (Dubinina, 1964) – on Castor fiber tuvinicus from Russia (Tuva)(Bochkov & Saveljev, 2012); on Castor fiber belorussicus from Poland (Bochkov et al., 2012)

Schizocarpus gozdziewskii Bochkov, Labrzycka, Skoracki et Saveljev

Schizocarpus gozdziewskii Bochkov, Labrzycka, Skoracki & Saveljev, 2012: 51; Bochkov & Saveljev, 2012: 16

Type locality: Suwalki

Distribution: Poland, Russia (Tuva)

Hosts: Castor fiber belorussicus, C.f. orientoeuropaeus, C.f. tuvinicus (Rodentia: Castoridae)

Schizocarpus heatherae Bochkov, Labrzycka, Skoracki et Saveljev

Schizocarpus heatherae Bochkov, Labrzycka, Skoracki & Saveljev, 2012: 49

Type locality: Suwalki

Distribution: Poland

Host: Castor fiber belorussicus (Rodentia: Castoridae)

Schizocarpus heideckei Bochkov et Saveljev

Schizocarpus heideckei Bochkov & Saveljev, 2012: 9

Type locality: Tuva, Todzhinskii District

Distribution: Russia (Tuva)

Host: Castor fiber tuvinicus (Rodentia: Castoridae)

Schizocarpus insignis Fain et Lukoschus, 1985 – on Castor fiber tuvinicus from Russia (Tuva)(Bochkov & Saveljev, 2012); on Castor fiber belorussicus from Poland (Bochkov et al., 2012)

Schizocarpus klompeni Bochkov, Labrzycka, Skoracki et Saveljev

Schizocarpus klompeni Bochkov, Labrzycka, Skoracki & Saveljev, 2012: 50

Type locality: Suwalki

Distribution: Poland

Host: Castor fiber belorussicus (Rodentia: Castoridae)

Schizocarpus lavrovi Bochkov et Saveljev

Schizocarpus lavrovi Bochkov & Saveljev, 2012: 7

Type locality: Tuva, Todzhinskii District

Distribution: Russia (Tuva)

Host: Castor fiber tuvinicus (Rodentia: Castoridae)

Schizocarpus modestus Fain et Lukoschus, 1985 – on Castor fiber tuvinicus from Russia (Tuva)(Bochkov & Saveljev, 2012)

Schizocarpus parahumilis Bochkov, Labrzycka, Skoracki et Saveljev

Schizocarpus parahumilis Bochkov, Labrzycka, Skoracki & Saveljev, 2012: 53

Type locality: Suwalki

Distribution: Poland

Host: Castor fiber belorussicus (Rodentia: Castoridae)

Schizocarpus pseudonumerosus Bochkov, Labrzycka, Skoracki et Saveljev

Schizocarpus pseudonumerosus Bochkov, Labrzycka, Skoracki & Saveljev, 2012: 47

Type locality: Suwalki

Distribution: Poland

Host: Castor fiber belorussicus (Rodentia: Castoridae)

Schizocarpus pusillus Fain et Lukoschus, 1985 – on Castor fiber tuvinicus from Russia (Tuva)(Bochkov & Saveljev, 2012)

Schizocarpus radiatus Fain et Lukoschus, 1985 – on Castor fiber tuvinicus from Russia (Tuva)(Bochkov & Saveljev, 2012); on Castor fiber belorussicus from Poland (Bochkov et al., 2012)

Schizocarpus saveljevi Bochkov

Schizocarpus saveljevi Bochkov, 2012: 168

Type locality: Leningrad Province

Distribution: Russia

Host: Castor fiber (Rodentia: Castoridae)

Schizocarpus subornatus Fain et Lukoschus, 1985 – on Castor fiber tuvinicus from Russia (Tuva)(Bochkov & Saveljev, 2012)

Schizocarpus testiculatus Bochkov, Labrzycka, Skoracki et Saveljev

Schizocarpus testiculatus Bochkov, Labrzycka, Skoracki & Saveljev, 2012: 44

Type locality: Suwalki

Distribution:. Poland

Host: Castor fiber belorussicus (Rodentia: Castoridae)

Schizocarpus tuvinicus Bochkov et Saveljev

Schizocarpus tuvinicus Bochkov & Saveljev, 2012: 10

Type locality: Tuva, Todzhinskii District

Distribution: Russia

Host: Castor fiber tuvinicus (Rodentia: Castoridae)

Schizocarpus unzhakovi Bochkov et Saveljev

Schizocarpus unzhakovi Bochkov et Saveljev, 2012: 8

Type locality: Tuva, Todzhinskii District

Distribution: Russia (Tuva)

Host: Castor fiber tuvinicus (Rodentia: Castoridae)

Schizocarpus zurowskii Bochkov, Labrzycka, Skoracki et Saveljev

Schizocarpus zurowskii Bochkov, Labrzycka, Skoracki & Saveljev, 2012: 52; Bochkov & Saveljev, 2012: 16

Type locality: Suwalki

Distribution: Poland, Russia

Host: Castor fiber belorussicus (Rodentia: Castoridae)

Schizocoptes daberti Bochkov

Schizocoptes daberti Bochkov, 2016: 005

Type locality: South Kivu, Plantation Lemera

Distribution: DR Congo

Host: Chrysochloris stuhlmanni (Afrosoricida: Chrysochloridae)

Soricilichus sylvisorex Bochkov, Mbaltini et Verheyen

Soricilichus sylvisorex Bochkov, Mbaltini & Verheyen, 2016: 249

Type locality: DR Congo

Distribution:. DR Congo

Hosts: Sylvisorex granti, S. lunaris, S. vulcanorum (Soricomorpha: Soricidae)

Fam. Atopomelidae

Atopomelus megalotis Bochkov, Abramov et Hugot

Atopomelus megalotis Bochkov, Abramov & Hugot, 2011: 6

Type locality: Kammouane Province, 18 km north of Thakhek, near Ban Doy village, 17º33’26’’N, 104º49’28’’E, 140 m a. s. l.

Distribution: Laos

Host: Hylomys megalotis (Erinaceomorpha: Erinaceidae)

Didelphoecius micoureus Bochkov et Valim

Didelphoecius micoureus Bochkov & Valim, 2016: 32

Type locality: Minas Gerais

Distribution: Brazil

Host: Micoureus paraguayanus (Didelphimorphia: Didelphidae)

Listrophoroides oconnori Bochkov, Abramov et Hugot

Listrophoroides oconnori Bochkov, Abramov & Hugot, 2011: 3

Type locality: Kammouane Province, 18 km north of Thakhek, near Ban Doy village, 17º33’26’’N, 104º49’28’’E, 140 m a. s. l.

Distribution: Laos

Host: Saxatilomys paulinae (Rodentia: Muridae)

Distribution of Listrophoroidea (suppl.)

EUROPE

Poland (Bochkov, Labrzycka, Skoracki & Saveljev, 2012) – Schizocarpus brachyurus, Sch. faini, Sch. fedjushini, Sch. gozdziewskii, Sch. heatherae, Sch. insignis, Sch. klompeni, Sch. parahumilis, Sch. pseudonumerosus, Sch. radiatus, Sch. testiculatus, Sch. zurowskii

Russia (entire)(Bochkov & Saveljev, 2012; Bochkov, 2012, 2016b) – Schizocarpus brachyurus (= Sch. intercalatus, = Sch. parabrachyurus), Sch. gozdziewskii, Sch. saveljevi, Schi. azasicus, Sch. daberti, Sch. heidekei, Sch. insignis, Sch. lavrovi, Sch. modestus, Sch. pusillus, Sch. radiatus, Sch. subornatus, Sch. tuvinicus, Sch. unzhakovi, Trichoecius dubininae, T. lemmus

AFRICA

Burundi (Bochkov, OConnor & Skoracki, 2016) - Myocoptes grammomys

DR Congo (Bochkov, Mbaltini & Verheyen, 2016; Bochkov, 2016) - Soricilichus sylvisorex, Schizocoptes daberti

Gabon (Bochkov, OConnor & Skoracki, 2016) - Trichoecius hylomyscus

South Africa (Bochkov, OConnor & Skoracki, 2016) – Trichoecius angolensis

Tanzania (Bochkov, OConnor & Skoracki, 2016) – Criniscansor congolensis, Gliricoptes vulcanorum, Myocoptes spinulatus, Trichoecius faini, T. hylomyscus, T. lootensis

Uganda (Bochkov, OConnor & Skoracki, 2016) - Lophuromys voosnami, Myocoptes grammomys, Trichoecius otomys

ASIA

Kazakhstan (Bochkov, 2016) - Trichoecius dubininae

Kirghizia (Bochkov, 2016) - Trichoecius dubininae

Laos (Bochkov, Abramov & Hugot, 2011) – Afrolistrophorus laonastes, A. saxatilomys, Pteromychirus menetes, Atopomelus megalotis, Listrophoroides oconnori

Vietnam (Bochkov, 2017, 2018) – Dubininetta abramovi, D. euroscaptor, Lutrilichus javanicus

SOUTH and CENTRAL AMERICA

Argentina (Sikora & Bochkov, 2012) - Prolistrophorus (Aprolistrophorus) parabidentatus, Prolistrophorus akodon, P. amazonicus, P. argentines, P. (P.) frontalis, P. (Beprolistrophorus) hirstianus

Bolivia (Sikora & Bochkov, 2012) - Prolistrophorus amazonicus

Brazil (Sikora & Bochkov, 2012, Bochkov & Valim, 2016; Gorza et al., 2018) – Didelphoecius micoureus, Leporacarus gibbus, L. sylvilagi, Lynxacarus serrafreirei, Parakosa tadarida, Prolistrophorus bidentatus, P. ctenomys, P. hylaeamys, Sclerolistrophorus oxymycteris

Chile (Sikora & Bochkov, 2012) - Amlistrophorus geoxus

Colombia (Bochkov, Lareschi & Barreto, 2014) - Prolistrophorus (Prolistrophorus) grassii, P. (P.) argentinus

Far East of Russia (Bochkov, 2017) - Dubininetta taiwanensis

Grenada (Springer et al., 2018) – Afrolistrophorus sp., Atopomelidae

Guatemala (Bochkov, Lareschi & Barreto, 2014) - Prolistrophorus (Aprolistrophorus) tylomys

Panama (Sikora & Bochkov, 2012) - Prolistrophorus nectomys

Peru (Sikora & Bochkov, 2012) - Prolistrophorus nectomys

Suriname (Sikora & Bochkov, 2012) - Prolistrophorus musculinus

Hosts

Didelphimorphia

Fam. Didelphidae

Metachirus nudicaudatusDidelphoecius validus

Micoureus paraguayanus - Didelphoecius micoureus

Erinaceomorpha

Fam. Erinaceidae

Hylomys megalotis - Atopomelus megalotis

Afrosoricida

Fam. Chrysochloridae

Chrysochloris stuhlmanni Schizocoptes daberti

Soricomorpha

Fam. Soricidae

Sylvisorex granti - Soricilichus sylvisorex

S. lunaris - Soricilichus sylvisorex

S. vulcanorum - Soricilichus sylvisorex

Fam. Talpidae

Euroscaptor parvidensDubininetta euroscaptor

E. subanura – Dubininetta abramovi

Mogera robusta - Dubininetta taiwanensis

Chiroptera

Fam. Molossidae

Molossus molossus – Parakosa tadarida

Rodentia

Fam. Arvicolidae

Lasiopodomys gregalisTrichoecius dubininae

Lemmus sibiricus - Trichoecius lemmus

Fam. Cricetidae

Akodon affinis - Prolistrophorus (Prolistrophorus) argentinus

A. azarae - Prolistrophorus (Aprolistrophorus) parabidentatus

A. cursor – Prolistrophorus bidentatus

A. montensis – Prolistrophorus akodon

Calomys callosus - Prolistrophorus amazonicus

C. laucha - Prolistrophorus amazonicus

C. musculinus - Prolistrophorus amazonicus

Geoxus valdivianusAmlistrophorus geoxus

Hylaeamys laticeps - Prolistrophorus hylaeamys

Melanomys caliginosus - Prolistrophorus argentinus, P. nectomys

Nectomys palmipesProlistrophorus nectomys

Oligoryzomys sp. – Prolistrophorus (P.) frontalis

Oryzomys laticeps - Sclerolistrophorus oxymycteris

Scapteromys aquaticus - Prolistrophorus argentines, P. (Beprolistrophorus) hirstianus

Tylomys nudicaudus - Prolistrophorus (Aprolistrophorus) tylomys

Zygodontomys brevicauda - Prolistrophorus (Prolistrophorus) grassii

Fam. Ctenomyidae

Ctenomys torquatusProlistrophorus ctenomys

Fam. Muridae

Dendromus mysticalisMyocoptes spinulatus

Grammomys dryasMyocoptes grammomys

G. macmillani – Criniscansor congolensis

Hapalomys delacouriAfrolistrophorus hapalomys

Hylomyscus anselli – Trichoecius hylomyscus

H. arcimontanus – Trichoecius faini

H. stella - Trichoecius hylomyscus

Meriones tamariscinus – Myocoptes meriones

M. libycus - Myocoptes meriones

M. meridianus – Trichoecius meriones

Otomys typusTrichoecius otomys

Saxatilomys paulinae - Afrolistrophorus saxatilomys, Listrophoroides oconnori

Lagomorpha

Fam. Leporidae

Sylvilagus brasiliensis Leporacarus sylvilagi

Fam. Castoridae

Castor fiberSchizocarpus saveljevi

Castor fiber belorussicusSchizocarpus brachyurus, Sch. heatherae, Sch. faini,

Sch. fedjushini, Sch. gozdziewskii, Sch. insignis, Sch. klompeni, Sch. parahumilis, Sch. pseudonumerosus, Sch. radiatus, Sch. testiculatus, Sch. zurowskii

Castor fiber tuvinicus - Schizocarpus azasicus, Sch. daberti, Sch. gozdziewskii, Sch. grandis, Sch. heidekei, Sch. insignis, Sch. lavrovi, Sch. modestus, Sch. pusillus, Sch. radiatus, Sch. subornatus, Sch. tuvinicus, Sch. unzhakovi, Sch. zurowskii

Carnivora

Fam. Mustelidae

Melogale moschata - Lutrilichus javanicus

Galictis cuja - Lynxacarus (Lynxacarus) serrafreirei

Publications on Listrophoroidea

(suppl. 2011 - 2021)

  • Bochkov A.V. 2012. Schizocarpus saveljevi sp. nov. (Acariformes: Chirodiscidae) parasitizing the Eurasian beaver–Castor fiber Linnaeus, 1758 (Rodentia: Castoridae) from Leningrad Province (Russia). - Proc. Zool. Inst. RAS, 316(2): 166 – 171.
  • Bochkov A.V. 2016a. A new mite species Schizocoptes daberti sp. n. (Acariformes: Chirodiscidae) from Chrysochloris stuhlmanni Matsche (Afrosoricida: Chrysochloridae) in the Democratic Republic of the Congo. - Folia Parasitologica, 63: 005.
  • Bochkov A.V. 2016b. Myocoptid mites (Acariformes: Myocoptidae) of the fauna of the former USSR. - Zootaxa, 4193(3): 451–485. DOI: 10.1164
  • Bochkov A.V. 2017. A review of fur-mites of the genus Dubininetta Fain & Lukoschus, 1978 (Acariformes: Listrophoridae), ectoparasites of moles (Soricomorpha: Talpidae). - Systematic Parasitology, 94(6): 635–647. https://doi.org/10.1007/s11230-017-9732-6
  • Bochkov A.V. 2018. External morphology of postembrionic stages of Lutrilichus javanicus (Acariformes: Chirodiscidae) from Melogale moschata (Carnivora: Mustelidae) from Vietnam. - Acarologia, 58(4): 763 – 772.
  • Bochkov, A.V., A.V. Abramov. 2014. Acariform mites (Acariformes)–permanent symbionts of Hapalomys delacouri Thomas (Rodentia, Muridae) in Vietnam. - ZooKeys, 459: 137–145.
  • Bochkov A.V., A. Labrzycka, M. Skoracki, A.P. Saveljev. 2012. Fur mites of the genus Schizocarpus Trouessart (Acari: Chirodiscidae) parasitizing the Eurasian beaver Castor fiber belorussicus Lavrov (Rodentia: Castoridae) in NE Poland (Suwałki). - Zootaxa, 3152: 39 – 59.
  • Bochkov A.V., M. Lareschi, M. Baretto. 2014. New species and records of the mite genus Prolistrophorus (Acariformes: Listrophoridae) from rodents of the subfamily Sigmodontinae (Rodentia: Cricetidae). - Acta Parasitologica, 59(4): 586 – 595.
  • Bochkov A.V., S.G. Mbalitini, E. Verheyden. 2016. Myocoptid mites (Acariformes: Review of the fur-mite genus Soricilichus Fain, 1970 (Acariformes: Chirodiscidae) – symbionts of the African shrews of the subfamily Crocidurinae (Soricomorpha: Soricidae). - Zootaxa, 4072(2): 235 - 253. DOI: 10.11646/zootaxa.4072.2.6
  • Bochkov A.V., B.M.OConnor, M. Skoracki. 2017. New species and new records of mites of the family Myocoptidae (Acariformes: Sarcoptoidea) from African rodents. - Journal of Medical Entomology, 54(3): 587 – 596.
  • Bochkov A.V., B.M.OConnor. 2017. A new species of myocoptid mites Myocoptes (Comyocoptes) nesokia sp. nov. (Acariformes: Myocoptidae) parasitizing Nesokia indica (Rodentia: Muridae) in Pakistan. - Acta Parasitologica, 62(1): 46 – 49.
  • Bochkov A.V., A.P. Saveljev. 2014. Mites of the genus Schizocarpus Trouessart, 1896 (Acariformes: Chirodiscidae) from the North American beavers (Castor canadensis) in Russia. - Parazitologiya, 48(6): 430 – 436.
  • Bochkov A.V., M.P. Valim. 2016. New species and records of mites of the superfamily Sarcoptoidea (Acariformes: Psoroptidia) from mammals in Brazil. - Acta Parasitologica, 61: 22 – 44. All data on host-parasite associations of sarcoptoids in Brazil are summarized. Totally, 61 sarcoptoid species of 8 families are recorded in Brazil.
  • Gorza L.L., C. M. Ferraz, D. M.Araujo, A. D. C. G. Lopes, L. B. Carvalho, F. L. G. Leite, P. C. G. Farias, A. R. Oliveira, F. R. Braga. 2018. Leporacarus gibbus (Acari: Listrophoridae) in a naturally-infested domestic rabbit: first report of its occurrence in the state of Espírito Santo, Brazil. - Bioscience Journal, 34(6): 1693 – 1696.
  • Sikora B., A.V. Bochkov. 2012. Fur mites of the family Listrophoridae (Acariformes: Sarcoptoidea) associated with South American sigmodontine rodents (Cricetidae: Sigmodontinae). - Acta Parasitologica, 57(4): 388 – 396.
  • Springer C. C.; K. Tiwari, A.M. Acuna, R. Sharma. 2018. Observation of Fur-mites of the Family Atopomelidae and Listrophoridae (Acari: Astigmata) from Brown Rats (Rattus norvegicus) in Grenada, West Indies: A Preliminary Report. - Journal of Animal Research; New Delhi, 8(1): 163-164.

Update 2014 - 2021 to Vol. III

Beron P. 2014. Acarorum Catalogus. III. Opilioacarida, Holothyrida, Mesostigmata (Dermanyssoidea). Pensoft & Nat. Mus. Natur. Hist. Sofia: 286 pp.

Introduction

As a result of the recent contributions, the number of genera and species in Beron, 2014 (some of them of uncertain status) is updated as follows:

Opilioacarida – 13 genera, 55 recent and 3 fossil species

Holothyrida – 15 genera, 30 species

Macronyssidae – 34 genera, 236 species

Dermanyssidae – 3 genera, 39 species

Halarachnidae – 6 genera, 40 species

Raillietiidae – 1 genus, 7 species

Entonyssidae - 8 genera, 24 species

Manitherionyssidae – 1 genus, 1 species

Hystrichonyssidae - 1 genus, 1 species

Dasyponyssidae - 2 genera, 2 species

Spelaeorhynchidae - 1 genus, 7 species

New contributions to the Catalogue

Opilioacarida

Fam. Opilioacaridae

Genus Amazonacarus Vázquez, de Araújo et Feres

Amazonacarus Vázquez, de Araújo & Feres, 2014: 152

Type species: Amazonacarus setosus Vázquez, de Araújo et Feres, 2014

Amazonacarus paraensis Vázquez, de Araújo et Feres

Amazonacarus paraensis Vázquez, de Araújo & Feres, 2014: 162

Type locality: Para

Distribution: Brazil

Amazonacarus setosus Vázquez, de Araújo et Feres

Amazonacarus setosus Vázquez, de Araújo & Feres, 2014: 155

Type locality: Pará, Alter do Chão

Distribution: Brazil

Genus Brasilacarus Vázquez, de Araújo et Feres

Brasilacarus Vázquez, de Araújo & Feres, 2015: 377

Type species: Brasilacarus cocaris Vázquez, de Araújo et Feres, 2015

Brasilacarus cocaris Vázquez, de Araújo et Feres

Brasilacarus cocaris Vázquez, de Araújo & Feres, 2015: 375

Type locality: Manaus, near Rio Negro and Taruma-Mirim stream

Distribution: Brazil

Caribeacarus ojasti (Lehtinen, 1980) – moved from Neocarus o. by Araújo et al. (2020)

Neocarus belizensis Vázquez, May, Alamilla et Klompen

Neocarus belizensis Vázquez, May, Alamilla & Klompen, 2018: 132

Type locality: Maya Mountains

Distribution: Belize

Neocarus caipora Bernardi, Klompen et Ferreira

Neocarus caipora Bernardi, Klompen & Ferreira, 2014: 47

Type locality: Canãa dos Carajas, Gruta Cav11-S11 (06°24’38.3”S 50°19’38.7”W)

Distribution: Brazil

Neocarus chactemalensis Vázquez et Klompen

Neocarus chactemalensis Vázquez & Klompen, 2015: 537

Type locality: Quintana Roo, Chetumal, campus Quintana Roo State University, 8 m

Distribution: Mexico

Neocarus comalensis Vázquez et Klompen

Neocarus comalensis Vázquez & Klompen, 2015: 538

Type locality: Guerrero, El Comal, elevation 1749 m

Distribution: Mexico

Neocarus coronatus Araújo, Bichuette, Bauchan, Ochoa et Feres

Neocarus coronatus : Araújo, Bichuette, Bauchan, Ochoa & Feres, 2018: 303

Type locality: São Desidério county, Bahia state

Distribution: Brazil

Neocarus entrerios Vázquez, Bernardi et Klompen

Neocarus entrerios Vázquez, Bernardi & Klompen, 2020: 512

Type locality: Misiones, P.N. Iguazu, Area de la Garganta del Diablo, 185 m

Distribution: Argentina

Neocarus misiones Vázquez, Bernardi et Klompen

Neocarus misiones Vázquez, Bernardi & Klompen, 2020: 508

Type locality: Misiones, Iguazu, Puerto Libertad, 173 m, 25.9189°S 54.5818°W

Distribution: Argentina

Neocarus simmonsi Bernardi, Zampaulo et Oliveira

Neocarus simmonsi Bernardi, Zampaulo & Oliveira, 2020: 14

Type locality: Minas Gerais State, São Gonçalo do Rio Abaixo city, Simmons’ cave, 19°53’55”S, 43°28’22”W

Distribution: Brazil

Neocarus spelaion Vázquez, Bernardi et Klompen

Neocarus spelaion Vázquez, Bernardi & Klompen, 2020: 3

Type locality: Minas Gerais State, Matozinhos, Cave 013, 19°31’57.21”S, 44°5’30.30”W

Distribution: Brazil

Opilioacarus baeticus Moraza Zorrilla, Prieto et Balanzateguic

Opilioacarus baeticus Moraza Zorrilla, Prieto & Balanzateguic, 2021: 130

Type locality: Isleta del Moro: Barranco del Paraíso (Níjar, Almería)

Distribution: Spain

Opilioacarus brignolii Araújo et Di Palma

Opilioacarus brignolii Araújo & Di Palma, 2018: 138

Type locality: Apulia, Bari, Monopoli

Distribution: Italy, incl. Sardinia

Opilioacarus groehni Dunlop et Bernardi (fossil)

Opilioacarus groehni Dunlop & Bernardi, 2014: 760

Type locality: from the Cretaceous Burmese amber

Distribution: Burma

Opilioacarus thaleri Vázquez, Ávila et Jus

Opilioacarus thaleri Vázquez, Ávila & Jus, 2021: 552

Type locality: Creta, Georgioupolis

Distribution: Greece (Creta)

Holothyrida

Fam. Neothyridae

Diplothyrus lehtineni Vázquez, Araújo et Feres

Diplothyrus lehtineni Vázquez, Araújo & Feres, 2016: 271

Type locality: Pará, São Geraldo do Araguaia, Serra das Andorinhas

Distribution: Brazil

Mesostigmata

Fam. Macronyssidae

Cryptonyssus flexus from Pipistrellus nathusii (Northern Caucasus, Orlova, Smirnov et al., 2020)

Macronyssus diversipilis from Myotis tschuliensis (Northern Caucasus, Orlova, Smirnov et al., 2020)

Macronyssus leislerianus from Nyctalus leisleri (Northern Caucasus, Orlova, Smirnov et al., 2020)

Ornithonyssus bacoti – dermatitis in Italy (Cafiero et al., 2015)

Ornithonyssus bursa from Sturnus vulgaris in Argentina (Lareschia et al., 2017)

Ornithonyssus sylviarum (Canestrini et Fanzago) – from Columba livia domestica in Egypt (Halawa et al., 2019)

Parichoronyssus alexanderfaini Morales-Malacara et Guerrero

Parichoronyssus alexanderfaini Morales-Malacara & Guerrero, 2020: 405

Type locality: Loc. near Bartica, Brit. Guiana, Forest Reserve 24 mi. along Potaro Rd.

Distribution: Guiana

Host: Rhinophylla pumilio (Chiroptera: Phyllostomatidae)

Parichoronyssus gettingeri Morales-Malacara et Guerrero

Parichoronyssus gettingeri Morales-Malacara & Guerrero, 2020: 410

Type locality:

Distribution:

Hosts: Rhynchonycteris naso (Chiroptera: Embalonuridae), Noctilio leporinus (Chiroptera: Noctilionidae)

Pellonyssus [as Steatonyssus] flabellifer (Gupta et Paul, 1985)(suppl.) - West Bengal: Midnapur Dist., Patharkumkumi, from Ploceus philippinensis (p.20)

Steatonyssus afer from Namibia on Cistugo seabrae (Orlova, Laverty et al., 2020)

Steatonyssus noctulus from Miniopterus schreibersii (Orlova, Stanyukovich & Orlov, 2015), Nyctalus noctula (Northern Caucasus, Orlova, Smirnov et al., 2020)

Steatonyssus longipes from Passer domesticus niloticus, Columba livia nilotica and Streptopelia senegalensis aegyptiaca in Egypt (Halawa et al., 2019. Note: the Steatonyssus are parasites of Mammals, mostly on bats, so these records on birds should be confirmed)

Steatonyssus olesovi Orlova, Stanyukovich et Orlov

Steatonyssus olesovi Orlova, Stanyukovich & Orlov, 2015: 97

Type locality: Russia

Distribution: Russia

Hosts: Myotis dasycneme (probably the main host), M. blythii, M. daubentoni, M. brandtii, M. mystacinus, and Ognev’s long-eared bat.

Distribution of Opilioacarida, Holothyrida, and some Mesostigmata (suppl.)

Opilioacarida

EUROPE

Italy (Araújo, Di Palma & Feres, 2018) – Opilioacarus brignolii

Sardinia (Araújo, Di Palma & Feres, 2018) - Opilioacarus brignolii

Greece

Crete (Vázquez et al., 2021) – Opilioacarus thaleri

Spain (Moraza Zorrilla, Prieto & Balanzateguic, 2021) – Opilioacarus baeticus

NORTH AMERICA

Mexico (Vázquez & Klompen, 2015) - Neocarus chactemalensis, N. comalensis,

SOUTH and CENTRAL AMERICA

Argentina (Vázquez, Bernardi & Klompen, 2020) – Neocarus entrerios, N. misiones

Belize (Vázquez, May, Alamilla & Klompen, 2018) – Neocarus belizensis

Brazil (Araújo, Bichuette, Bauchan, Ochoa & Feres, 2018; Bernardi & Borges-Filho, 2018; Bernardi, Klompen & Ferreira, 2014; Vázquez, de Araújo & Feres, 2014, 2015; Bernardi, Zampaulo & Oliveira, 2020) – Amazonacarus paraensis, A. setosus, Brasilacarus cocaris, Neocarus caipora, N. coronatus, N. simmonsi, N. spelaion

Holothyrida

SOUTH and CENTRAL AMERICA

Brazil (Bernardi, Azara & Ferreira, 2011; Vázquez, Araújo & Feres, 2016) – Diplothyrus lehtineni, D. schubarti According to Bernardi et al., 2011 (fide Lehtinen), Holothyrida are known also from the Philippines, Costa Rica and Panama.

Some Mesostigmata

EUROPE

Italy (Cafiero et al., 2015) - Ornithonyssus bacoti

Russia (Orlova, Smirnov et al., 2017; Orlova, Krilov et al., 2021) – Cryptonyssus flexus, Macronyssus diversipilis, M. leislerianus, Steatonyssus noctulus

Turkey (Ciloglu et al., 2020) – Dermanyssus gallinae

AFRICA

Egypt (Negm et al., 2018; Halawa et al., 2019) – Ornithonyssus sylviarum, Steatonyssus longipes, Dermanyssus

Namibia (Orlova, Laverty et al., 2020) - Steatonyssus (S.) afer

NORTH AMERICA

Mexico (Morales-Malacara & Guerrero, 2020) - Parichoronyssus gettingeri

SOUTH and CENTRAL AMERICA

Argentina (Lareschia et al., 2017) - Ornithonyssus bursa

Guiana (Morales-Malacara & Guerero, 2020) - Parichoronyssus alexanderfaini

Peru (Morales-Malacara & Guerrero, 2020; Angoma et al., 2021) – acari on bats

Venezuela (Morales-Malacara & Guerrero, 2020) – Parichoronyssus gettingeri

Hosts

Aves

Columbiformes

Fam. Columbidae

Columba livia domestica - Ornithonyssus bursa, O. sylviarum, Steatonyssus longipes

Streptopelia senegalensis aegyptiaca - Ornithonyssus bursa, Steatonyssus longipes

Passeriformes

Fam. Ploceidae

Ploceus philippinus – Pellonysus flabellifer

Fam. Passeridae

Passer domesticus niloticus - Ornithonyssus bursa, Steatonyssus longipes

Fam. Pycnonotidae

Pycnonotus barbatus – Ornithonyssus bacoti

Fam. Sturnidae

Sturnus vulgaris - Ornithonyssus bursa

Mammalia

Chiroptera

Fam. Cistugidae

Cistugo seabrae - Steatonyssus (S.) afer

Fam. Vespertilionidae

Myotis tschuliensisMacronyssus diversipilis

Nyctalus leisleriMacronyssus leislerianus

N. noctula – Steatonyssus noctulus

Pipistrellus nathusiiCryptonyssus flexus

Bibliography of Opilioacarida, Holothyrida, and some Mesostigmata

(suppl. 2014 - 2021)

Opilioacarida

  • Araújo M. S., A. Di Palma, R. J. Fazzio Feres. 2018. A new species of Opilioacarus With, 1902 (Acari: Opilioacaridae) from Italy, and a new diagnosis of the genus. - Zootaxa, 4500(1): 135 – 145.
  • Araújo M.S., A. Di Palma, R.J.F. Feres. 2020. Catalog of the Opilioacarida (Acari: Parasitiformes). - Zootaxa, 4895: 332–356.
  • Araújo M.S.d., M.E. Bichuette, G.R. Bauchan, R. Ochoa, R.J.F. Feres. 2018. A new species of cave dwelling Neocarus (Acari: Opilioacaridae) from Bahia state, Brazil, with remarks on taxonomic characters. - Zootaxa, 4402: 303-322. doi: 10.11646/zootaxa.4402.2.4
  • Bernardi L.F.d.O., E.L. Borges-Filho. 2018. Neocarus spelaion sp. n. (Parasitiformes, Opilioacaridae), a new species of cave dwelling Neocarus from Minas Gerais state, Brazil. - Subterranean Biology, 27: 1-16. doi: 10.3897/subtbiol.27.25777
  • Bernardi L.F.d.O., H. Klompen, M.S. Zacarias, R.L. Ferreira. 2013. A new species of Neocarus Chamberlin & Mulaik, 1942 (Opilioacarida: Opilioacaridae) from Brazil, with remarks on postembryonic development. - ZooKeys, 358: 69-89. doi: 10.3897/zookeys.358.6384
  • Bernardi L.F.d.O., H. Klompen, R.L. Ferreira. 2014. Neocarus caipora, a new mite species (Parasitiformes: Opilioacarida: Opilioacaridae) from Brazilian Amazon caves. – Acarologia, 54(1): 47–56.
  • Bernardi L.F.d.O., R.A. Zampaulo, M.P.A. Oliveira. 2020. A new species of Neocarus (Opilioacaridae) from a Brazilian ferruginous geosystem and notes on natural History. – Subterranean Biology, 36: 11 – 13.
  • Ciloglu A., A. Yildirim, Z. Onder, G. Yetismis, O. Duzlu, E. Simsek, A. Inci. 2020. Molecular characterization of poultry red mite, Dermanyssus gallinae lineages in Turkey and first report of Plasmodium species in the mite populations. - International Journal of Acarology, 46(4): 241
  • Dunlop J.A., L.F.O. Bernardi. 2014. An opilioacarid mite in Cretaceous Burmese amber. - Naturwissenschaften, 101: 759–763. doi: 10.1007/s00114-014-1212-0
  • Halawa A.M., S.A.A. Osman, H.A.A. Ahmed. 2019. Diversity and infestation rate of mites associated with some birds in upper and lower Egypt. – Annals of Agric. Sci., Moshtohor, 57(2): 517 – 524.
  • Moraza Zorrilla M. L., C. E. Prieto, I. Balanzateguic. 2021. A new species of the genus Opilioacarus With, 1902 (Acari: Opilioacarida) for the Iberian Peninsula. - Acarologia (Paris), 61(1): 128 - 147.
  • Vázquez M.M., M.S. de Araújo, R. J. F. Feres. 2014. A new genus and two new species of Opilioacaridae (Acari: Parasitiformes) from Amazonia, Brazil with a key to world genera. - Zootaxa, 3814(2): 151-176.
  • Vázquez M.M., M.S. de Araújo, R. J. F. Feres. 2015. Brasilacarus cocaris (Acari: Opilioacaridae), a new genus and species from Amazonia, Brazil. – Zootaxa, 3915(3): 375-389.
  • Vázquez M., L.F.O. Bernardi, H. Klompen. 2020. The family Opilioacaridae (Acari: Parasitiformes) in Argentina, with description of two new species. – Acarologia, 60(3): 505-519.
  • Vázquez M.M., H. Klompen. 2015. The family Opilioacaridae (Parasitiformes: Opilioacarida) in Mexico, description of two new species and notes on biology and geographical distribution. – Zootaxa, 3957(5): 535-552.
  • Vázquez M.M., D. May, E. Alamilla, H. Klompen. 2018. A new species of Opilioacaridae (Parasitiformes: Opilioacarida) from Belize with some observations on life history and behaviour. - Systematic & Applied Acarology, 23(1): 132–144.
  • Vázquez M.M., I. M. Á. Herrera, P. Just,,A..C. R. Lerma, M. Chatzaki, T. L. Heller, J. Král. 2021. A new opilioacarid species(Parasitiformes: Opilioacarida) from Crete (Greece) with notes on its karyotype. - Acarologia, 61(3): 548-563; DOI 10.24349/acarologia/20214449
  • Holothyrida

  • Bernardi L.F.d.O.; L.N. d. Azara, R.L. Ferreira. 2011. Description of the female of Diplothyrus schubarti Lehtinen, 1999 (Holothyrida: Neothyridae) and new species occurrences in Brazil. – Acarologia, 51(3): 311 – 319.
  • Vázquez M.M., M.S. Araújo, R.J.F. Feres. 2016. A new species of Diplothyrus (Parasitiformes: Neothyridae) from Brazil. - Acarologia, 56(3): 269–278.
  • Some Mesostigmata

  • Angoma D. M., J. L. Mendoza, S.Jorge, L. M. Silva, J.Iannacone. 2021. Ectoparasitic fauna on the common vampire bat Desmodus rotundus (Geoffroy, 1810) (Chiroptera: Phyllostomidae) from Huarochiri, Lima, and a checklist of ectoparasites in bats of Peru. – Graellsia, 77(1): e135
  • Cafiero M.A., D.A. Raele, G. Mancini, D. Galante. 2015. Dermatitis by Tropical Rat Mite, Ornithonyssus bacoti (Mesostigmata, Macronyssidae) in Italian city‐dwellers: a diagnostic challenge. - The Journal of the European Academy of Dermatology and Venereology. https://doi.org/10.1111/jdv.13162
  • Canto-Osorio J.M., A. Cuxim-Koyoc, H. A. Ruiz-Piña, J. B. Morales-Malacara, E. Reyes-Novelo. 2020. Ectoparasites of Didelphis virginiana from Yucatan, Mexico. - Journal of Medical Entomology, 57(6): 1821-1829.
  • Gupta S.K., K. Paul. 1985. Some mites associated with birds’ nests in West Bengal, with descriptions of eleven new species. - Bull.Zool. Surv. of India, 7(1) 1-23.
  • Halawa A.M., S. A. A. Osman, H. A. A. Ahmed. 2019. Diversity and infestation rate of mites associated with some birds in upper and lower Egypt. – Annals of Agric. Sci., Moshtohor, 57(2): 517 – 524.
  • Lareschia M., G.L. Cicuttinc, M. N. De Salvoc, L. Ibañez, D. Montalti. 2017. The tropical fowl mite Ornithonyssus bursa (Acari: Mesostigmata: Macronyssidae) parasitizing the European starling Sturnus vulgaris (Aves: Passeriformes: Sturnidae), an invasive bird in central Argentina. An approach to the bacterial fauna of this mite. - Revista mexicana de biodiversidad, 88(2): 455.
  • Lima-Barbero J.F., M. S. Sánchez, A. Cabezas-Cruz, L. Mateos-Hernández, M. Contreras, I. G. Fernández de Mera, M. Villar, J. de la Fuente. 2019. Clinical gamasoidosis and antibody response in two patients infested with Ornithonyssus bursa (Acari: Gamasida: Macronyssidae). - Experimental and Applied Acarology, 78: 555–564.
  • McCulloch J.B., J. P. Owen, N.C. Hinkle, B.A. Mullens, J. W. Busch. 2020. Genetic Structure of Northern Fowl Mite (Mesostigmata: Macronyssidae) Populations Among Layer Chicken Flocks and Local House Sparrows (Passeriformes: Passeridae). - Journal of Medical Entomology, 57(1): 122–130.
  • Morales-Malacara J.B., R. Guerrero. 2020. Two New Species and New Records of Mites of the Genus Parichoronyssus (Acari: Macronyssidae) from South American Bats (Chiroptera), with a Key to the known species of the Genus. - Journal of Medical Entomology, 57(2): 404–417. https://doi.org/10.1093/jme/tjz196
  • Negm M.W., A.A. Mohamed, H.M.Kh. El-Gepaly, S.M. Abdelaziz. 2018. Mesostigmata mites (Acari, Parasitiformes) associated with birds and their nests from Egypt. –Turkish Journal of Zoology, 42: 722 – 731.
  • Nieri-Bastos F.A., B. L. Marcelo, A. Marcili, L. A. Durden, L. Mendoza-Uribe, D. M. Barros-Battesti. 2011. Morphological and molecular analysis of Ornithonyssus spp. (Acari: Macronyssidae) from small terrestrial mammals in Brazil. – Exp. Appl. Acarology, 55(4): 305 – 327.
  • Nordenfors H., J. Höglund. 2000. Long term dynamics of Dermanyssus gallinae in relation to mite control measures in aviary systems for layers. - Br. Poult. Sci., 41: 533-540.
  • Orlova M.V., P.B. Klimov, O.L. Orlov, D. G. Smirnov, A.V. Zhigalin, I.V. Budaeva, A.A. Emelyanova, N.V. Anisimov. 2021. A checklist of bat-associated macronyssid mites (Acari: Gamasina: Macronyssidae) of Russia, with new host and geographical records. – Zootaxa, 4974(3): 537–564.
  • Orlova M. V., Th.M. Laverty, W.K. Reeves, E.M. Gratton, M.L. Davies, N.V. Anisimov. 2020. New geographical and host record of bat ectoparasite Steatonyssus (Steatonyssus) afer Radovsky and Yunker, 1963 (Mesostigmata: Gamasina: Macronyssidae). – Acarologia, 60(4): 951-958.
  • Orlova M.V., D. G. Smirnov, N.V. Anisimov, O. L. Orlov, P. B. Klimov, V. P. Vekhnik, E. S. Murashko, A. M. Lukyanenko. 2020. Parasitic macronyssid mites (Mesostigmata: Macronyssidae) from bats of Northern Caucasus with key for females of the genus Macronyssus Kolenati, 1858 of Russia and adjacent countries. – International J. of Acarology, 46(5): 364-372. DOI: 10.1080/01647954.2020.1801839
  • Orlova M.V., M.K. Stanyukovich, O.L. Orlov. 2015. Gamasid mites (Mesostigmata: Gamasina) associated with bats (Chiroptera: Vespertilionidae, Rhinolophidae, Molossidae) of boreal Palaearctic zone (Russia and adjacent countries). Tomsk: Publishing House of Tomsk State University, 150 р.
  • Salinas-Ramos V.B., J. B. Morales-Malacara. 2019. Seasons influence acarine infestation in bats from tropical dry forests of Western Mexico. - Mammalian Biology, 96(1): 37-44.

Update 2015 – 2021 to Vol. IV

Beron P. 2015. Acarorum Catalogus. IV. Myobioidea, Pterygosomatoidea. Pensoft & Nat. Mus. Natur. Hist. Sofia: 251 pp.

Introduction

As a result of the recent contributions, the number of genera and species in Myobiidae and Pterygosomatoidea (some of them with uncertain status) is updated as follows:

Myobiidae – 55 genera, 511 species

Pterygosomatidae – 13 genera, 196 species

New contributions to the Catalogue

Fam. Myobiidae

Acanthophthirius lasiurus (Fain & Whitaker, 1976) recorded in Mexico on the bat Lasiurus cinereus (Palisot de Beauvois) by Herrera-Mares et al. (2017)

Crocidurobia faini Bochkov, Mbalitini et Verheyen

Crocidurobia faini Bochkov, Mbalitini & Verheyen, 2016: 496

Type locality: DR Congo

Distribution: DR Congo

Hosts: Crocidura denti, Crocidura cf. niobe, Crocidura cf. littoralis (Soricomorpha: Soricidae)

Crocidurobia lukoschusi Bochkov

Crocidurobia lukoschusi Bochkov, 2017: 212

Type locality: Vietnam

Distribution: Vietnam

Host: Crocidura tanakae (Soricomorpha: Soricidae)

Eudusbabekia saguei - recorded in Mexico on Pteronotus davyi by Herrera-Mares et al. (2017)

Eudusbabekia provirilia Morales-Malacara, Guzmán-Cornejo & López-Ortega, 2002 recorded in Mexico on Leptonycteris yerbabuenae by Herrera-Mares et al. (2017)

Ioanella mimon (Fain, 1973) - on Mimon cozumelae (Herrera-Mares et al., 2017)

Ioannela leptonycteris (Fain, 1973) - on Leptonycteris yerbabuenae by Herrera-Mares et al. (2017)

Eudusbabekia mimon (Fain, 1973) - on Mimon cozumelae (Herrera-Mares et al., 2017)

Radfordia (Hesperomyobia) akodon Bochkov

Radfordia (Hesperomyobia) akodon Bochkov, 2017b

Type locality: Bolivia

Distribution: Bolivia

Host: Akodon varius (Rodentia: Cricetidae, Sigmodontinae)

Radfordia (Hesperomyobia) vandenberghi Fain and Lukoschus on Oligoryzomys flavescens (Waterhouse) and Oligoryzomys sp. (Rodentia: Cricetidae: Sigmodontinae) from Bolivia (Bochkov, 2017)

Radfordia (Microtimyobia) stekolnikovi Bochkov et Mironov, 1998, syn. of R. lemnina (Koch, 1841)

Radfordia (Microtimyobia) stenocrani Bochkov et Mironov, 1998, syn. of R. lemnina (Koch, 1841)

Radfordia (Radfordia) mirabilis Bochkov et Abramov

Radfordia (Radfordia) mirabilis Bochkov & Abramov, 2014: 142

Type locality: Binh Phuoc Province, Bu Gia Map National Park, 13

km NE Bu Gia Map Village, 540 m a.l.s., 12°11’37”N, 107°12’21”E

Distribution: Vietnam

Host: Hapalomys delacouri (Rodentia: Muridae)

Protohylomysobia erinaceophilus Sidorchuk et Bochkov, 2018 (2019)

Fam. Pterygosomatidae

Genus Callopistiella Silva-de la Fuente, Casanueva et Moreno Salas

Callopistiella Silva-de la Fuente, Casanueva & Moreno Salas, 2015: 66

Type species: Callopistiella atacamensis Silva-de la Fuente, Casanueva et Moreno Salas, 2015

Callopistiella atacamensis Silva-de la Fuente, Casanueva et Moreno Salas

Callopistiella atacamensis Silva-de la Fuente, Casanueva & Moreno Salas, 2015: 67

Type locality: Caldera: Copiapó Province, Atacama Region, 26°56’S, 70°44’W

Distribution: Chile

Host: Callopistes maculatus (Sauria: Teiidae)

Geckobia africana Fajfer

Geckobia africana Fajfer, 2018: 2

Type locality: Irimga Region, Mufindi District, Uzungwa Scarp Forest Reserve

Distribution: Tanzania

Host: Cnemaspis africana (Sauria: Gekkonidae)

Geckobia andina Machado, Gazêta, Pérez, Cunha et Giupponi

Geckobia andina Machado et al., 2019: 335

Type locality: Cerro Verde, Andean highlands

Distribution: Peru

Host: Phyllodactylus gerrhopygus (Sauria: Gekkonidae)

Geckobia circumdata Machado, Gazêta, Pérez, Cunha et Giupponi

Geckobia circumdata Machado et al., 2019: 342

Type locality: Arequipa, Cerro Verde, 2345 m

Distribution: Peru

Host: Phyllodactylus gerrhopygus (Sauria: Gekkonidae)

Geckobia milii Fajfer

Geckobia milii Fajfer, 2019: 6

Type locality: Far North Queensland, Cape York Peninsula

Distribution: Australia (Queensland)

Host: Underwoodisaurus milii (Sauria: Carphodactylidae)

Fajfer (2019) revised the genus Pterygosoma and left as such only the species of Asia and Africa. Gerrhosaurobia Lawrence was raised in a new status of full genus, the three species in it became Gerrhosaurobia bicolor (Lawrence), G. gerrhosauri (Lawrence) and G. hystrix (Lawrence). A new genus Neopterygozoma Fajfer has been described for the parasites on the South American lizards of the genus Liolaemus (Liolaemidae). Fajfer (2020b) revised the entire genus Pterygosoma

Geckobiella stamii (Jack) – on Amblyomma dissimile (Acari: Ixodidae) associated with Iguana iguana from Panama (Domínguez et al., 2020)

Genus Neopterygozoma Fajfer

Neopterygozoma Fajfer, 2019: 422

Type species: Pterygosoma chilensis Fajfer et González-Acuña, 2013

Neopterygosoma schroederi Fajfer

Neopterygosoma schroederi Fajfer, 2020: 544

Type locality: Maule region, Curicó Prov., Curicó

Distribution: Chile

The species Pterygosoma chilensis, P. cyanopteri, P. formosus, P. levissima, P. ligare, P. ovata, described by Fajfer & Gonzalez – Acuña in 2013, as well as Pterygosoma patagonica Ditmar de la Cruz, Morando et Avila, 2004, have been transferred by Fajfer (2020a) in the genus Neopterygosoma.

Pterygosoma aculeatum Lawrence, 1936 – restaured as species from P. agamae aculeatum by Fajfer, 2020b: 93

Pterygosoma agamae Peters, 1849 – published again from Tanzania and Mozambique (Fajfer, 2020b: 99)

Pterygosoma expansum Bertrand, Finkelman et Paperna, 2000 is synonymised with Pterygosoma adramitana Jack, 1961 by Fajfer, 2020b

Pterygosoma angolae Jack, 1962 – raised as species from P. melanum angolae by Fajfer, 2020b: 51

Pterygosoma balochistani Fajfer

Pterygosoma balochistani Fajfer, 2016a: 348; 2020b: 26

Type locality: Balochistan, Darzi Chan

Distribution: Pakistan

Host: Landakia nupta nupta (Sauria: Agamidae)

Pterygosoma blanfordi Fajfer

Pterygosoma blanfordi Fajfer, 2016a: 344

Type locality: South India, Kerala State, Palakkad district, Parambiculam Tiger Reserve

Distribution: India

Host: Psammophilus blanfordianus (Sauria: Agamidae)

Pterygosoma capensis Jack, 1962 - raised as species from P. melanum capensis by Fajfer, 2020b: 55

Pterygosoma gladiator Bertrand, Finkelman et Paperna, 2000 is synonymised with P. neumanni (Berlese, 1910) by Fajfer, 2020b: 111

Pterygosoma livingstonei Bertrand et Modry, 2004 is synonymised with P. circularis Jack, 1962 by Fajfer, 2020b

Pterygosoma longipalpae Lawrence, 1936 - restaured as species from Pterygosoma melanum longipalpae by Fajfer, 2020b: 59

Pterygosoma orbicularis Jack, 1962 - raised as species from P. spinosa orbicularis by Fajfer, 2020b: 119

Pterygosoma pallidum Fajfer, Melnikov et Dabert

Pterygosoma pallidum Fajfer, Melnikov & Dabert, 2016: 792; Fajfer, 2020b: 11

Type locality: Aquaba Province, 11 km north-east of Ar- Risha

Distribution: Jordan

Host: Trapelus pallidum (Sauria: Agamidae)

Pterygosoma parasiniatum Fajfer, Melnikov et Dabert

Pterygosoma parasiniatum Fajfer, Melnikov & Dabert, 2016: 801

Type locality: NE of Ar-Risha

Distribution: Jordan

Host: Pseudotrapelus cf. sinaiticus (Sauria: Agamidae)

Pterygosoma problematica Jack, 1962 - raised as species from P. fimbriata problematica by Fajfer, 2020a: 83

Pterygosoma pseudorbicularis Jack, 1962 - raised as species from P. bibronii pseudorbicularis by Fajfer, 2020b: 121

Pterygosoma rhipidostichata Bertrand, Finkelman and Paperna, 2000 is synonymised with P. mutabilis Jack, 1961 by Fajfer, 2020b

Pterygosoma theobaldi Fajfer, Melnikov et Dabert

Pterygosoma theobaldi Fajfer, Melnikov & Dabert, 2016: 806

Type locality: North India

Distribution: India

Host: Phrynocephalus teobaldi (Sauria: Agamidae)

Pimeliaphylus hemidactyli Fajfer et Karanth

Pimeliaphylus hemidactyli Fajfer & Karanth, 2021: 442

Type locality: Karnataka State, Bangalore, IISc campus

Distribution: India

Hosts: Hemidactylus murrayi, H. leschenaultia, H. parvimaculatus (Squamata: Gekkonidae)

Zonurobia circularis limpopoensis was proposed by OConnor (2020) as a replacement name for Zonurobia circularis transvaalensis Lawrence, not Zonurobia transvaalensis Lawrence

New hosts and distribution records are given for Pterygosoma neumanni (Berlese, 1910), P. bibronii Jack, 1962, P. longipalpae Lawrence, 1936, P. gracilipalpis and P. sinaita Jack, 1961.

Distribution of Myobioidea and Pterygosomatoidea

(suppl. 2015 - 2021)

Myobioidea

ASIA

Vietnam (Bochkov, 2017) – Crocidurobia lukoschusi

NORTH AMERICA

Mexico (Herrera-Mares et al., 2017; Herrera-Mares, Guzmán-Cornejo & Morales-Malacara, 2021) - Acanthophthirius lasiurus, Ioanella mimon, Eudusbabekia jimenezi, E. mimon, E. saguei

SOUTH and CENTRAL AMERICA

Bolivia (Bochkov, 2017) - Radfordia (Hesperomyobia) akodon, R. (H.) vandenberghi

Pterygosomatoidea

AFRICA

Tanzania (Fajfer, 2018) – Geckobia africana

South Africa (OConnor, 2020) - Zonurobia circularis limpopoensis

ASIA

India (Fajfer, 2016a) - Pterygosoma blanfordi, P. theobaldi

Jordan (Fajfer, 2016b) - Pterygosoma pallidum, P. parasiniatum

Pakistan (Fajfer, 2016a) - Pterygosoma balochistani

SOUTH and CENTRAL AMERICA

Argentina (Ditmar de la Cruz, Morando & Avila, 2004, Fajfer, 2020) – Neopterygosoma patagonica

Chile (Fajfer, 2019, 2020) – Neopterygosoma spp.(formerly in Pterygosoma), Neopterygosoma schroederi, Callopistiella atacamensis

Cuba (Borroto Paez et al., 2020) – Geckobia hemidactyli

Panama (Domínguez et al., 2020) - Geckobiella stamii

Peru (Machado et al., 2019) – Geckobia andina, G. circumdata

OCEANIA

Australia (Fajfer, 2018) – Geckobia milii

Hosts

Myobioidea

Soricomorpha

Fam. Soricidae

Crocidura denti - Crocidurobia faini

Crocidura cf. niobe - Crocidurobia faini

Crocidura cf. littoralis - Crocidurobia faini

Crocidura tanakae - Crocidurobia lukoschusi

Chiroptera

Fam. Phyllostomatidae

Mimon cozumelae - Eudusbabekia mimon, Ioanella mimon

Rodentia

Fam. Cricetidae

Akodon varius - Radfordia (Hesperomyobia) akodon

Oligoryzomys flavescens - Radfordia (Hesperomyobia) vandenberghi

Fam. Muridae

Hapalomys delacouri - Radfordia (Radfordia) mirabilis

Pterygosomatoidea

Reptilia

Sauria

Fam. Agamidae

Landakia nupta nupta - Pterygosoma balochistani

Phrynocephalus teobaldi - Pterygosoma theobaldi

Psammophilus blanfordianus - Pterygosoma theobaldi

Pseudotrapelus cf. sinaiticus - Pterygosoma parasiniatum

Trapelus pallidum - Pterygosoma pallidum

Fam. Carphodactylidae

Underwoodisaurus miliiGeckobia milii

Fam. Gekkonidae

Cnemaspis africanaGeckobia africana

Hemidactylus mabouiaGeckobia hemidactyli

H. murrayi – Pimeliaphilus hemidactyli

H. parvimaculatus - Pimeliaphilus hemidactyli

H. leschenaultia - Pimeliaphilus hemidactyli

Phyllodactylus gerrhopygusGeckobia andina, G. circumdata

Fam. Teiidae

Callopistes maculatus - Callopistiella atacamensis

Publications on Myobioidea and Pterygosomatoidea

(suppl. 2015 - 2021)

Myobioidea

  • Bochkov A.V. 2015. Comparison of external morphology of tritonymphs in myobiid mites (Myobiidae) associated with marsupials (Marsupialia). - International Journal of Acarology, 41: 590–599. DOI: 10.1080/01647954.2015.1084046
  • Bochkov A.V. 2017. To the systematics of the subgenus Hesperomyobia Bochkov, 1997 (Acariformes: Myobiidae: Radfordia). - Systematic and Applied Acarology, 22(6): 785 – 788.
  • Bochkov A.V., A.V. Abramov. 2014. Acariform mites (Acariformes)–permanent symbionts of Hapalomys delacouri Thomas (Rodentia, Muridae) in Vietnam. - ZooKeys, 459: 137–145.
  • Bochkov A.V., S.G. Mbalitini, E. Verheyen. 2016. Crocidurobia faini sp. n. (Acariformes: Myobiidae) - a new mite species parasitizing shrews of the genus Crocidura (Soricimorpha: Soricidae) in DR Congo. - Systematic Parasitology, 93: 493–499. DOI: 10.1007/s11230-016-9623-2
  • Bochkov A.V., A.A. Mirolubov. 2015. Description of postembrionic immature stages of Protomyobia onoi Jameson & Dusbábek, 1971 (Acariformes: Myobiidae) and comparative analysis of the juvenile external morphology in the tribe Protomyobiini. - International Journal of Acarology, 41: 115–127. DOI: 10.1080/01647954.2014.1003966
  • Bochkov A.V. 2017a. Crocidurobia lukoschusi sp.nov. (Acariformes: Myobiidae) parasitizing Crocidura tanakae Kuroda, 1938 (Soricimorpha: Soricidae) in Vietnam, the first record of the genus from oriental white-toothed shrews. - International Journal of Acarology, 43: 212–216. DOI: 10.1080/01647954.2016.1268204213.
  • Bochkov A.V. 2017b. To the systematics of the subgenus Hesperomyobia Bochkov, 1997 (Acariformes: Myobiidae: Radfordia). - Systematic and Applied Acarology, 22 (6): 785–788. DOI: 10.11158/saa.22.6.4.
  • Herrera-Mares A., C. Guzmán-Cornejo, L. León-Paniagua, G. Rivas. 2017. Myobiid mites (Trombidiformes, Myobiidae) of the golden bat Mimon cozumelae from Mexico. Description of the male and tritonymph of Ioanella mimon and new records of Eudusbabekia mimon. – ZooKeys, 658: 1 – 8.
  • Herrera-Mares A., C. Guzmán-Cornejo, J.-B. Morales - Malacara. 2021. The myobiid mites (Acariformes, Eleutherengona, Myobiidae) from Mexico: hosts, distribution and identification key for the genera and species. - Systematic and Applied Acarology, 26(4): 724 – 748.
  • Sidorchuk E. A., A.V. Bochkov, T. Weiterschan, O.F. Chernova. 2019. A case of mite-on-mammal ectoparasitism from Eocene Baltic amber (Acari: Prostigmata: Myobiidae and Mammalia: Erinaceomorpha). - Journal of Systematic Paleontology, 17(4): 331–347. DOI: 10.1080/14772019.2017.1414889
  • Pterygosomatoidea

  • Borroto-Páez R., C. C. Martínez-Rivera, D. R. Pérez. 2020. Mites (Geckobia hemidactyli, Actinedida: Pterygosomatidae) in Tropical House Geckos (Hemidactylus mabouia) in Cuba: A Review with New Distribution Records. - Reptiles & Amphibians, 27(3): 382–389.
  • Díaz J.A., R. A. Torres, L. Paternina, D. Santana, R. Miranda. 2020. Traveling with an invader: ectoparasitic mites of Hemidactylus frenatus (Squamata: Gekkonidae) in Colombia. – Cuadernos de Herpetología, 34
  • Dittmar de la Cruz K., M. Morando, L. Avila. 2004. Description of a New pterygosomatid mite (Acari, Actinedida: Pterygosomatidae) parasitic on Liolaemus spp. (Iguania: Liolaemini) from Argentina. - Zootaxa, 521: 1–6.
  • Domínguez L. G., J.J. Lezcano, I. Rodríguez, R.J. Miranda, J. Roberto, S. C. Bermúdez, C. Sergio. 2020. Is Geckobiella stamii (Acari: Pterygosomatidae) a hyperparasite or phoretic on Amblyomma dissimile (Acari: Ixodidae) associated with Iguana iguana from Panama? – Acarologia, 60(1): 40 – 44.
  • Fajfer M. 2016a. Two new species of the genus Pterygosoma (Acariformes: Pterygosomatidae) parasitizing agamid lizards (Sauria: Agamidae) from the Indian subcontinent. - Acta Parasitologica, 61(2): 343–354.
  • Fajfer M. 2018. New species and records of scale mites (Acariformes: Pterygosomatidae) from geckos (Squamata: Gekkonidae and Carphodactylidae). - BioMed Research International, 2018.
  • Fajfer M. 2019. Systematics of reptile-associated scale mites of the genus Pterygosoma (Acariformes: Pterygosomatidae) derived from external morphology. - Zootaxa 4603: 401 – 440.
  • Fajfer M. 2020a. A systematic revision of the genus Neopterygosoma Fajfer, 2019 (Acariformes: Pterygosomatidae) with the description of a new species. – Systematic Parasitology, 97: 535 – 551.
  • Fajfer M. 2020b. A systematic revision of the scale mite genus Pterygosoma Peters, 1849 (Acariformes: Pterygosomatidae). – Zootaxa, 4805(1): 1 – 147.
  • Fajfer M., P. Karanth. 2021. Integrating a morphological description with DNA barcode data of a new species of the genus Pimeliaphilus (Acariformes: Pterygosomatidae) with an analysis of its host specifity and a key to the genus. – Systematic & Applied Acarology, 26(2): 438 – 454.
  • Fajfer M., D. Melnikov, M. Dabert. 2016b. Three new species of the genus Pterygosoma Peters, 1849 (Acariformes: Pterygosomatidae) from agamid lizards (Sauria: Agaminae) with DNA barcode data. - Systematic Parasitology, 93(8): 791 – 814.
  • Machado I.B., G. S. Gazêta, J. Pérez, R. Cunha, A. P. de l. Giupponi. 2019. Two new species of the genus Geckobia Mégnin, 1878 (Acariformes, Prostigmata, Pterygosomatidae) from Peru. – Zootaxa, 4657(2): 333 – 351.
  • OConnor B.M. 2020. Correction of a case of homonymy in the genus Zonurobia (Acari: Pterygosomatidae). - Acarologia (Paris), 60(3): 557 – 558.
  • Paredes-León R. 2013. Sistemática de la familia Pterygosomatidae (Acari: Prostigmata). Tesis de Doctorado. Universidad Nacional Autónoma de México, Mexico City, 127 pp.
  • Sidorchuk E.A., A. A. Khaustov. 2018. A parasite without host: The first fossil pterygosomatid mite (Acari: Prostigmata: Pterygosomatidae) from French Lower Cretaceous amber. - Cretaceous Research, 91: 131-139.
  • Silva-de la Fuente M.C., R. Paredes-León, M. E. Casanueva, G. Escobar-Huerta, L. Moreno Salas. 2015. A new genus and species of pterygosomatid mite (Acari: Pterygosomatidae) parasitizing Callopistes maculatus (Squamata: Teiidae) from Chile. – Zootaxa, 3972(1): 65 – 74.

Update 2015 - 2021 to Vol. V

Beron P. 2015. Acarorum Catalogus. V. Sarcoptoidea. Pensoft & Nat. Mus. Natur. Hist. Sofia: 380 pp.

Introduction

As a result of the recent contributions, the number of genera and species in Sarcoptoidea (some of them uncertain status) is updated as follows:

Lemurnyssidae – 2 genera, 4 species

Lobalgidae – 2 genera, 9 species

Psoroptidae – 31 genera, 58 species

Listropsoralgidae - 4 genera, 19 species

Bochkovocoptidae – 1 genus, 1 species

Rhyncoptidae - 5 genera, 13 species

Sarcoptidae - 17 genera, 117 species

Pneumocoptidae - 1 genus, 4 species

Gastronyssidae - 9 genera, 42 species

Chirorhynchobiidae - 1 genus, 3 species

Pyroglyphidae - 22 genera, 51 species

Dermationidae - 12 genera, 54 species

Epidermoptidae - 16 genera, 51 species

Gaudoglyphidae – 1 genus, 1 species

Ptyssalgidae – 1 genus, 1 species

Turbinoptidae - 9 genera, 36 species

Cytoditidae - 2 genera, 14 species

Laminosioptidae – 6 genera, 27 species

Guanolichidae - 3 genera, 4 species

New (or ommited) contributions to the Catalogue

Fam. Lobalgidae

Echimytricalges brasiliensis Fain, 1970 is described actually not from Echimys brasiliensis, but (according to Bochkov & OConnor, 2010) from Phyllomys pattoni. This species was recorded also from Trinomys albosignatus and T. iheringi (Brazil, Bochkov & OConnor, 2010)

Echimytricalges guianensis Fain, 1970 - on Proechimys simonsi from Peru (Bochkov & OConnor, 2010)

Echimytricalges hoplomys Fain, 1970 – recorded also on Proechimys semispinosus and Hoplomys gymnurus from Ecuador and Costa Rica (Bochkov & OConnor, 2010)

Echimytricalges lonchothrix Bochkov et OConnor

Echimytricalges lonchothrix Bochkov & OConnor, 2010: 218

Type locality: Para, Rio Tapajoz, Boim, 03°05’S, 55°18’W

Distribution: Brazil

Host: Lonchothrix emiliae (Rodentia: Echimyidae)

Echimytricalges mesomys Fain, Lukoschus et Mendez, 1982 - on Mesomys hispidus from Brazil (Bochkov & OConnor, 2010)

Echimytricalges proechimys Bochkov et OConnor

Echimytricalges proechimys Bochkov & OConnor, 2010: 214

Type locality: Oriente, Rio Jatun Yacu

Distribution: Ecuador

Host: Proechimys brevicauda (Rodentia: Echimyidae)

Echimytricalges surinamensis Fain and Lukoschus, 1970 – also on Proechimys oconneli from Colombia (Bochkov & OConnor, 2010)

Echimytricalges whitakeri Fain et Ritzi

Echimytricalges whitakeri Fain & Ritzi, 2001 – validated by Bochkov & OConnor, 2010: 221 from nomen nudum

Fam. Psoroptidae

Caparinia ictonyctis Lawrence

Caparinia ictonyctis Lawrence, 1955: 54

Type locality: South Africa

Distribution: South Africa

Host: Atelerix albiventris (Erinaceomorpha: Erinaceidae)

Gaudalges averianovi Bochkov

Gaudalges averianovi Bochkov, 2018: 593

Type locality: Madagascar

Distribution: Madagascar

Host: Lepidolemur ruficaudatus (Primates: Lepilemuridae)

Lemuralges propithecus Bochkov, Klompen, Junge et Williams

Lemuralges propithecus Bochkov, Klompen, Junge & Williams, 2015: 2

Type locality: Madagascar

Distribution: Madagascar

Host: Propithecus diadema (Primates: Indridae)

Genus Coendalges Fain et Méndes, 1979 was transferred from Lobalgidae (Coendalginae) to Psoroptidae (Psoralginae) by Bochkov & OConnor, 2010. The family Lobalgidae has been left with two genera (Echimytricalges and Lobalges)

Genus Psoroptes Gervais, 1844. According to Rinder et al. (2000), “…P. cervinus Ward, 1915, P. cuniculi (Delafond, 1859), P. natalensis Hirst, 1919 and P. ovis (Hering, 1838) are seen as synonyms of P. equi (Hering, 1838)”

Fam. Listrosporalgidae

Didelphialges metachirus Bochkov et Valim

Didelphialges metachirus Bochkov & Valim, 2013: 67

Type locality: Peru

Distribution: Peru

Host: Metachirus nudicaudatus (Didelphimorphia: Didelphidae)

Listropsoralges brevisetosus Bochkov et Valim

Listropsoralges brevisetosus Bochkov & Valim, 2013: 39

Type species: Peru

Distribution: Peru

Host: Marmosa murina (Didelphimorphia: Didelphidae)

Listropsoralges similis Bochkov et Valim

Listropsoralges similis Bochkov & Valim, 2013: 19

Type locality: Panama

Distribution: Panama

Host: Caluromys derbianus (Didelphimorphia: Didelphidae)

Listropsoralges thylamys Bochkov et Valim

Listropsoralges thylamys Bochkov & Valim, 2013: 24

Type locality: Bolivia

Distribution: Bolivia

Host: Thylamys venustus (Didelphimorphia: Didelphidae)

Listropsoralges vossi Bochkov et Valim

Listropsoralges vossi Bochkov & Valim, 2013: 35

Type locality: Brazil

Distribution: Brazil

Host: Monodelphis domestica (Didelphimorphia: Didelphidae)

Listropsoralges caenolestes Bochkov et Valim

Listropsoralges caenolestes Bochkov & Valim, 2013: 26

Type locality: Ecuador

Distribution: Ecuador

Host: Caenolestes fuliginosus (Paucituberculata: Caenolestidae)

The female of Listropsoralges faini Bochkov et Wauthy, 2009 is described in the revision of Bochkov & Valim (2016) for the first time.

Fam. Bochkovocoptidae

Bochkovocoptidae OConnor, 2019: 146

Type genus: Bochkovocoptes OConnor, 2019

Genus Bochkovocoptes OConnor

Bochkovocoptes OConnor, 2019: 146

Type species: Bochkovocoptes otomys OConnor, 2019

Bochkovocoptes otomys OConnor

Bochkovocoptes otomys OConnor, 2019: 146

Type locality:Kigezi Province, Kisoro Dstrict, Mgahinga Gorilla Nat. Park

Distribution: Uganda

Host: Otomys denti kempi (Rodentia: Muridae)

Fam. Pyroglyphidae

Genus Tachornithoglyphus Klimov, Bochkov et OConnor

Tachornithoglyphus Klimov, Bochkov & OConnor, 2015: 97

Type species: Guatemalichus tachornis Cruz, Cuervo et Dusbábek, 1984

Tachornithoglyphus tachornis (Cruz, Cuervo et Dusbábek, 1984)

Guatemalichus tachornis Cruz, Cuervo & Dusbábek, 1984: 1; Cruz 1988: 14; Fain 1988: 22

Tachornithoglyphus tachornis (Cruz, Cuervo et Dusbábek, 1984): Klimov, Bochkov & OConnor, 2015: 97

Type locality: Ciudad de La Habana, Playa El Salado, Bauta

Distribution: Cuba

Host: nest of Tachornis phoenicobia (Apodiformes: Apodidae)

Fam. Dermationidae

Genus Carcinopodacarus Hernandes, Pedroso et Bochkov

Carcinopodacarus Hernandes, Pedroso & Bochkov, 2015: 2

Type species: Carcinopodacarus polymorphus Hernandes, Pedroso et Bochkov, 2015

Carcinopodacarus polymorphus Hernandes, Pedroso et Bochkov

Carcinopodacarus polymorphus Hernandes, Pedroso & Bochkov, 2015: 3

Type locality: Campinas

Distribution: Brazil

Host: Guira guira (Cuculiformes: Cuculidae)

Neodermation actitis Bochkov et Mironov

Neodermation actitis Bochkov & Mironov, 2015: 612; Matyukhin & Yatsuk, 2022

Type locality: the Far East of Russia

Distribution: Russia

Host: Actitis hypoleucos (Charadriiformes: Scolopacidae)

Fam. Epidermoptidae

Metamicrolichus pici (Fain) – new for Egypt (Halawa et al., 2019, sub “Myialges pici”)

Promyalges falconis (Fain) – new for Egypt (Halawa et al., 2019, sub “Myialbes falconis”)

Promyalges italicus Moradi Faradonbeh, Ostovan, OConnor, Gheibi, Joharchi et Macchioni

Promyialges italicus Moradi Faradonbeh, Ostovan, OConnor, Gheibi, Joharchi & Macchioni, 2019: 36

Type locality: Tuscany, Pisa, 43° 42’ N, 10° 24’ E

Distribution: Italy

Host: Pseudolynchia canariensis (Diptera: Hippoboscidae) from Columba livia

Distribution of Sarcoptoidea

(suppl. 2016 - 2021)

EUROPE

Italy (Moradi Faradonbeh et al., 2019) - Promyialges italicus

Russia (Bochkov & Mironov, 2015) – Dermation actitis (the Far East)

AFRICA

Egypt (Nageiba et al., 2001; Halawa et al., 2019) – Metamicrolichus pici, Promyalges falconis

Madagascar (Bochkov, 2018) - Gaudalges averianovi

ASIA

India (Modak et al., 2004; Podder, Biswas & Kumar Saha, 2021) – Pyroglyphidae

NORTH AMERICA

Mexico (Rodríguez-Ortega et al., 2018) - Knemidocoptes mutans

SOUTH and CENTRAL AMERICA

Bolivia (Bochkov & Valim, 2013) - Listropsoralges thylamys

Brazil (Bochkov & OConnor, 2010; Bochkov & Valim, 2013; Hernandes, Pedroso & Bochkov, 2015) – Carcinopodacarus polymorphus, Echimytricalges lonchothrix, E. mesomys, Listropsoralges vossi

Colombia (Bochkov & OConnor, 2010) – Echimytricalges surinamensis

Costa Rica (Bochkov & OConnor, 2010) – Echimytricalges hoplomys

Cuba (Cruz, Cuervo & Dusbabek, 1984) - Tachornithoglyphus tachornis

Ecuador (Bochkov & OConnor, 2010; Bochkov & Valim, 2013) - Echimytricalges hoplomys, E. proechimys, Listropsoralges caenolestes

Panama (Bochkov & Valim, 2013) - Listropsoralges similis

Peru (Bochkov & OConnor, 2010; Bochkov & Valim, 2013) – Echimytricalges guianensis, Listropsoralges brevisetosus

Hosts

AVES

Charadriiformes

Fam. Scolopacidae

Neodermation actitis - Actitis hypoleucos

Cuculiformes

Fam. Cuculidae

Guira guira - Carcinopodacarus polymorphus

Passeriformes

Fam. Passeridae

Passer domesticus niloticus – Promyalges falconis, Metamicrolichus pici

MAMMALIA

Didelphimorphia

Fam. Didelphidae

Caluromys derbianus - Listropsoralges similis

Marmosa murina - Listropsoralges brevisetosus

Metachirus nudicaudatus - Didelphialges metachirus

Monodelphis domestica - Listropsoralges vossi

Thylamys venustus - Listropsoralges thylamys

Erinaceomorpha

Fam. Erinaceidae

Atelerix albiventris - Caparinia ictonyctis

Rodentia

Fam. Echimyidae

Hoplomys gymnurusEchimytricalges hoplomys

Lonchothrix emiliaeEchimytricalges lonchothrix

Mesomys hispidusEchimytricalges mesomys

Proechimys brevicaudaEchimytricalges proechimys

P. oconneli – Echimytricalges surinamensis

P. semispinosus - Echimytricalges hoplomys

P. simonsi – Echimytricalges guianensis

Trinomys albispinus – Echimytricalges brasiliensis

T. iheringi - Echimytricalges brasiliensis

Fam. Muridae

Otomys denti kempi - Bochkovocoptes otomys

Primates

Fam. Indriidae

Propithecus diadema - Lemuralges propithecus

Fam. Lepilemuridae

Lepidolemur ruficaudatus - Gaudalges averianovi

Publications on Sarcoptoidea

(suppl. 2015 - 2021)

  • Aykut M., Ö.K. Erman, S. Doğan. 2016. Variability in population density of house dust mites of Bitlis and Mus, Turkey. - Journal of Medical Entomology, 53(3): 513-518. doi: 10.1093/jme/tjw009
  • Bochkov A.V. 2018. Gaudalges averianovi sp.nov. (Acariformes: Psoroptidae)–a new parasite of Malagasy lemurs.- Systematic and Applied Acarology, 23(4): 593–596. DOI http://doi.org/10.11158/saa.23.4.1
  • Bochkov A. V., P.B. Klimov, Da-Hee, K., M. Skoracki. 2018. Validation of the status of a species with high CO1 and low nuclear genetic divergences: the scab mite Caparinia ictonyctis stat. res. (Acariformes: Psoroptidae) parasitizing the African hedgehog Atelerix albiventris. - Zootaxa, 4544(4): 523–547. DOI: 10.11646/zootaxa.4544.4.4
  • Bochkov A.V., H. Klompen, R.E. Junge, C.V. Wiliams. 2015. Lemuralges propithecus sp. n. (Acariformes: Psoroptidae), an ectoparasite of the diademed sifaka Propithecus diadema (Primates: Indriidae). - Folia Parasitologica, 62: 11. DOI: 10.14411/fp.2015.011
  • Bochkov A.V., S.V. Mironov. 2015. A new species of the genus Neodermation Fain (Acariformes: Dermationidae) and the first record of the precopulatory mate-guarding among dermationids. - Systematic and Applied Acarology, 20: 612–628. DOI: 10.11158/saa.20.6.4.
  • Bochkov A.V., B.M. OConnor. 2010. External morphology and systematics of the family Lobalgidae (Acari: Astigmata) with resolution of the question of family monophyly. - Insect Systematics & Evolution, 41(3): 187 – 229.
  • Bochkov A.V., B.M. OConnor, P. Grootaert. 2013. Revision of the family Listropsoralgidae Fain, 1965 (Acariformes: Sarcoptoidea) - skin parasites of marsupials and rodents. – Zootaxa, 3611: 1 – 69.
  • Bochkov A.V., M.P. Valim. 2016. New species and records of mites of the superfamily Sarcoptoidea (Acariformes: Psoroptidia) from mammals in Brazil. - Acta Parasitologica, 61: 22 – 44.
  • Brooks A.J., R. Wall. 2001. Infection of Psoroptes Mites with the Fungus Metarhizium Anisopliae. - Experimental and Applied Acarology, 25: 869–880.
  • Halawa A.M., S.A.A. Osman, H.A.A. Ahmed. 2019. Diversity and infestation rate of mites associated with some birds in upper and lower Egypt. – Annals of Agric. Sci., Moshtohor, 57(2): 517 – 524.
  • Hernandes F.A., L.G.A. Pedroso, A.V. Bochkov. 2015. Carcinopodacarus polymorphus gen. n. et sp. n. from Guira guira (Cuculiformes: Cuculidae) in Brazil: a first example of male polymorphism in the family Dermationidae (Acariformes: Analgoidea). - Folia Parasitologica, 62: 9. DOI: 10.14411/fp.2015.009.
  • Klimov P.B., A.V. Bochkov, B.M. OConnor. 2015a. Phylogenetic position of the house dust mite subfamily Guatemalichinae (Acariformes: Pyroglyphidae) based on integrated molecular and morphological analyses and different measures of support. - Cladistics, 32(3): 261–275. DOI: 10.1111/cla.12126
  • Klimov P.B., A.V.Bochkov, B.M. OConnor. 2015b. Tachornithoglyphus gen. nov.–a new genus of nidicolous Pyroglyphidae (Acariformes: Astigmata). Zootaxa, 3956: 97–112. DOI: 10.11646/zootaxa.3956.1.5
  • Ludman S. 2016. House dust mite allergy. – Neodoctor. Modak A., G. K. Saha, N. Tandon, S. K. Gupta. 2004. Faunal diversity and habitat preference of house dust mites in West Bengal in relation to nasobronchial allergic disorders. - Rec. zoo!. Surv. India, 102(1-2): 137-146.
  • Matyukhin A.V., A.A. Yatsuk. 2022. The first record of Promyialges italicus Faradonbeh et al. 2019 (Acariformes: Epidermoptidae) in European Russia. – Acarologia, 62(1): 94 – 98.
  • Moradi Faradonbeh M., H. Ostovan, B. M. OConnor, M. Gheibi, O. Joharchi, F. Macchioni. 2019. Promyialges italicus sp. nov. (Astigmata: Epidermoptidae) with redescriptions of mites of the families Epidermoptidae and Cheyletidae (Prostigmata) associated with Pseudolynchia canariensis (Diptera: Hippoboscidae) from Iran and Italy. – Persian Journal of Acarology, 8(1): 27 – 46.
  • Nageiba I,S., S.T. Nagi, H.I. Mohamed, S.O. Eman. 2001. Mites associated with some birds in El-Minia governorate, Upper Egypt. – Egyptian Journal of Biology, 3: 124 – 135.
  • OConnor B.M. 2019. Bochkovocoptidae, an unusual new family of psoroptidian mites (Acari: Astigmata) associated with Otomys denti (Rodentia: Muridae) from Uganda. - Acarina, 27(2): 145 – 150.
  • Podder S., H. Biswas, G. Kumar Saha. 2021. A faunistic survey of house dust mites of Kolkata, West Bengal, India. - Acarological Studies, 3(1): 22 – 31.
  • Reese N.E., W. M. Boyce, I. A Gardner, D. M. Nelson. 1996. Fixation Affects Morphometric Characters of Psoroptes cuniculi Mites (Acari: Psoroptidae). – Journal of Medical Entomology, 33(5): 835-838.
  • Rettenmund C.L., R.J. Ossiboff, D. McAloose, W. Knee, S.E. Wade, J.A. Paré. 2015. Microlichus americanus acariasis in saffron finches (Sicalis flaveola) with dermatitis and feather loss. - Journal of Veterinary Diagnostic Investigation, 27(3): 382–386.
  • Rinder M. et al. 2000. Species of the genus Psoroptes (Acari: Psoroptidae): A taxonomic consideration. - Experimental and Applied Acarology, 24(3): 213-225.
  • Rodríguez-Ortega A., A. Equihua-Martínez, R. Nieto-Aquino, M. P. Chaires-Grijlava, A. Pro-Martínez, L. T. Rodríguez-Ortega. 2018. Presencia de Knemidocoptes mutans (Robin & Lanquetin) (Astigmata: Knemidocoptidae) en gallos minuatura en el estado de Hidalgo, México. - Folia entomologica Mexicana, 4(1): 28-31.
  • Sajid M.S., M. Ahsan, A. Kausar, M. Jawad, M. K. Saleemi. 2017. Sarcoptes scabiei (Acari: Sarcoptidae) infestation in rabbits (Oryctolagus cuniculus): A case study. - Revista Colomb. Entomol., 43(1): 51 - 54.

Update 2020 – 2021 to Vol. VI

Beron P. 2020. Acarorum Catalogus. VI. Rhinonyssidae, Spinturnicidae. Pensoft & Nat. Mus. Natur. Hist. Sofia: 265 pp.

Introduction

As a result of the recent contributions, the number of genera and species (some of them uncertain status) is updated as follows:

Rhinonyssidae – 16 genera, 457 species

Spinturnicidae – 12 genera, 111 species

New contributions to the Catalogue

Fam. Rhinonyssidae

Ptilonyssus dimi Dimov

Ptilonyssus dimi Dimov, 2020b: 6

Type locality: Elez, Lipezk region

Distribution: Russia

Host: Hirundo rustica (Passeriformes: Hirundinidae)

Ptilonyssus harko Dimov

Ptilonyssus harko Dimov, 2020b: 2

Type locality: Viazovka, Orenburg region

Distribution: Russia

Host: Riparia riparia (Passeriformes: Hirundinidae)

Ptilonyssus poriazovi Dimov

Ptilonyssus poriazovi Dimov, 2020b: 9

Type locality: Kohma, Ivanovo region

Distribution: Russia

Host: Emberiza citrinella (Passeriformes: Emberizidae)

Rhinonyssus derojasi Dimov

Rhinonyssus derojasi Dimov, 2020a: 2

Type locality: Revda, Sverdlovsk Region

Distribution: Russia

Host: Anser anser (Anseriformes: Anatidae)

Rhinonyssus karelinae Dimov

Rhinonyssus karelinae Dimov, 2020a: 11

Type locality: Tihvin, Leningrad region

Distribution: Russia

Host: Charadrius dubius (Charadriiformes: Charadriidae)

Rhinonyssus mileni Dimov

Rhinonyssus mileni Dimov, 2020a: 8

Type locality: Dubovka, Volgograd Region

Distribution: Russia

Host: Charadrius dubius (Charadriiformes: Charadriidae)

Rhinonyssus sixli Dimov

Rhinonyssus sixli Dimov, 2020a: 5

Type locality: Mosolovo, Kursk region

Distribution: Russia

Host: Podiceps ruficollis (Podicipediformes: Policipedidae)

Sternostoma alexmironi Dimov

Sternostoma alexmironi Dimov, 2020c: 2

Type locality: Opochka, Pskov region

Distribution: Russia

Host: Oenanthe oenanthe (Passeriformes: Muscicapidae)

Sternostoma antoni Dimov

Sternostoma antoni Dimov, 2020c: 5

Type locality: Liski, Voronej region

Distribution: Russia

Host: Dendrocopos major (Piciformes: Picidae)

Sternostoma chakarovae Dimov

Sternostoma chakarovae Dimov, 2020c: 8

Type locality: Hvalovo, Leningrad region

Distribution: Russia

Host: Oenanthe oenanthe (Passeriformes: Muscicapidae)

Sternostoma delianovae Dimov

Sternostoma delianovae Dimov, 2020c: 11

Type locality: Kirishi, Leningrad region

Distribution: Russia

Host: Turdus pilaris (Passeriformes: Turdidae)

Sternostoma enevi Dimov

Sternostoma enevi Dimov, 2020c: 14

Type locality: Budenovsk, Stavropol region

Distribution: Russia

Host: Turdus pilaris (Passeriformes: Turdidae)

Fam. Spinturnicidae

Ancystropus aethiopicus Hirst, 1923 – Egypt, on Rousettus aegyptiacus in Qalyubiyah Governorate (Hafez et al., 1994)

Meristaspis kenyaensis (Radford, 1947) - Egypt, on Rousettus aegyptiacus in Qalyubiyah Governorate (Hafez et al., 1994)

Periglischrus empheresotrichus Morales-Malacara, Castaño-Meneses et Klompen

Periglischrus empheresotrichus Morales-Malacara, Castaño-Meneses & Klompen in Morales-Malacara, Castaño-Meneses, Klompen & Mancina, 2019: 421

Type locality: Jamaica, St. Ann Parish; ½ mi. S, ½ mi, W. Runaway Bay

Distribution: Cuba, Dominica, Jamaica, Haiti, Guadeloupe

Hosts: Monophyllus redmani, M. plethodon (Chiroptera: Phyllostomatidae)

Periglischrus paracaligus Herrin et Tipton, 1975 - Mexico, on Leptonycteris

yerbabuenae (Zamora-Mejias, Morales - Malacara et al., 2020)

Distribution of Rhinonyssidae and Spinturnicidae

(suppl. 2020 - 2021)

Fam. Rhinonyssidae

EUROPE

Russia (Dimov, 2017, 2020a, b, c, d; Dimov & Zhigulin, 2020; Veiga et al., 2021) – Ptilonyssus dimi, P. harko, P. poriazovi, Rhinonyssus derojasi, Rh. karelinae, Rh. mileni, Rh. sixli, Sternostoma alexmironi, S. antoni, S. chakarovae, S. delianovae, S. enevi

SOUTH AMERICA

Fam. Spinturnicidae

EUROPE

Russia (Orlova, Klimov et al., 2021; Orlova, Vu Dinh Thong et al., 2021)

Crimea (Orlova & Klimov, 2020) – Spinturnix psi

AFRICA

Egypt (Hafez et al., 1994) - Meristaspis kenaensis, Ancystropus aethiopicus

ASIA

Iran (Yousefi et al., 2018) – Spinturnix emarginata

NORTH and CENTRAL AMERICA

Mexico (Zamora-Mejias, Morales – Malacara et al., 2020; Morales-Malacara et al., 2020) – Periglischrus caligus, P. empheresotrichus

SOUTH AMERICA

Paraguay (Morales - Malacara et al., 2020) – Periglischrus herrerai

Hosts

Rhinonyssidae

Aves

Anseriformes

Fam. Anatidae

Anser anser Rhinonyssus derojasi

Charadriiformes

Fam. Charadriidae

Charadrius dubiusRhinonyssus karelinae, Rh. mileni

Passeriformes

Fam. Emberizidae

Emberiza citrinellaPtilonyssus poriazovi

Fam. Hirundinidae

Hirundo rusticaPtilonyssus dimi

Riparia ripariaPtilonyssus harco

Fam. Muscicapidae

Oenanthe oenanthe - Sternostoma alexmironi, S. chakarovae

Fam. Turdidae

Turdus pilarisSternostoma delianovae, S. enevi

Piciformes

Fam. Picidae

Dendrocopos majorSternostoma antoni

Podicipediformes

Fam. Podicipedidae

Podiceps ruficollisRhinonyssus sixli

Chiroptera

Fam. Spinturnicidae

Fam. Pteropodidae

Rousettus aegyptiacus - Meristaspis kenaensis, M. lateralis, Ancystropus aethiopicus, A. zeleborii

Fam. Vespertilionidae

Myotis emarginatusSpinturnix emarginata

Fam. Phyllostomatidae

Periglischrus herrerai Machado-Allison, 1965 is recorded also on Anoura sp., Sturnira ludovici, S. lilium by Morales-Malacara et al. (2020).

Publications on Rhinonyssidae and Spinturnicidae

(suppl. 2020 - 2021)

  • Rhinonyssidae
  • De Rojas M., J. Doña , I. Dimov. 2020. A comprehensive survey of Rhinonyssid mites (Mesostigmata: Rhinonyssidae) in Northwest Russia: New mite-host associations and prevalence data. - Biodiversity Data Journal 8: e49535, 1- 37.
  • Dimov I. 2017. Nasal rhinonyssid mites of birds. Second International Parasitological Symposium, Saint Petersburg, 6 – 8 November 2017, 81 – 83.
  • Dimov I. 2020a. Four new species of nasal mites Rhinonyssus (Mesostigmata: Rhinonyssidae) from Russia. – Arhimed, Journal of Science and Practice, 9: 26 – 41.
  • Dimov I. 2020b. Three new species of nasal mites Ptilonyssus (Mesostigmata: Rhinonyssidae) from Russia. – Arhimed, Journal of Science and Practice, 9: 41 – 52.
  • Dimov I. 2020c. Five new species of nasal mites Sternostoma (Mesostigmata, Rhinonyssidae) from Russia. Arhimed - Journal of Science and Practice, 9: 3-19.
  • Dimov I. 2020d. Taxonomic diversity and morphology of mites of the family Rhinonyssidae of the Northwest of Russia. Saint Petersburg, 214 pp.
  • Dimov I., L.L.C. Zhigulin. 2020. Morphology of nasal mites of the family Rhinonyssidae – scanning electron microscopy. Saint Petersburg, 128.
  • Veiga J., I. Dimov, M. de Rojas. 2021.Endoparasitic Mites (Rhinonyssidae) on Urban Pigeons and Doves: Updating Morphological and Epidemiological Information. - Diversity, 13: 11. https: //doi.org/10.3390/d13010011
  • Spinturnicidae
  • Hafez S. M., N.M. Ebaid, M.E. Tharwat, A.E. Abdel-Megeed. 1994. Bat mites of Egypt (Acari: Spinturnicidae).- Annals of Agricultural Science (Cairo), 39(1): 463-471.
  • Morales-Malacara J.B., Y.M. L. Aldana, E. Reyes-Novelo, C. E. Almazán-Marín, H. A. Ruiz-Piña, A. Cuxim-Koyoc, Á. Aguilar-Setién, H. Colín-Martínez, C. García-Estrada, M. Ojeda. 2018. Redescription of Periglischrus herrerai (Acari: Spinturnicidae) associated to Desmodus rotundus (Chiroptera: Phyllostomidae: Desmodontinae), with a description of adult female heteromorphism and an analysis of its variability throughout the Neotropics. - Journal of Medical Entomology, 55(2): 300–316.
  • Morales-Malacara J.B., G. Castaño-Meneses, H. Klompen, C. A Mancina. 2020. New Species of the Genus Periglischrus (Acari: Spinturnicidae) from Monophyllus Bats (Chiroptera: Phyllostomidae) in the West Indies, Including a Morphometric Analysis of Its Intraspecific Variation. - Journal of Medical Entomology, 57(2): 418–436.
  • Nguyen Thanh Luong, M.V.Orlova, Vu Quang Mu, Dinh Thong. 2021. First record of Eyndhovenia (Mesostigmata: Gamasina: Spinturnicidae) from Vietnam. - Parasitology International, 82.
  • Orlova M.V., P.B. Klimov. 2020. Local host-parasite co-extinction: the bent-wing bat Miniopterus schreibersii (Chiroptera: Miniopteridae) and Spinturnix psi. (Mesostigmata: Spinturnicidae) in Crimea. – Plecotus et al., 23: 80 – 87.
  • Orlova M.V., M.K. Stanyukovich, O.L. Orlov. 2015. Gamasid mites (Mesostigmata: Gamasina) associated with bats (Chiroptera: Vespertilionidae, Rhinolophidae, Molossidae) of boreal Palaearctic zone (Russia and adjacent countries). – Tomsk: Publishing House of Tomsk State University, 150 р.
  • Orlova M.V., P.B. Klimov, O.L. Orlov, S. V. Kruskop, V.S. Lebedev. 2021. New geographic and host records of spinturnicid mites (Mesostigmata: Spinturnicidae) in Asia, with description of the protonymph of Spinturnix tylonycterisi. - Internat. Journal of Acarology, 47(4): 361 – 365.
  • Orlova M.V., Vu Dinh Thong, N.V. Anisimov, D.G. Smirnov, O.L. Orlov. 2021. New findings of spinturnicid mites (Mesostigmata: Gamasina: Spinturnicidae) from Caucasus. – International Parasitology, 85.
  • Yousefi Sh. et al. 2018. Postnatal variation in ectoparasite (Spinturnix emarginata) load in neonates of Geoffroy’s Bat (Myotis emarginatus): how fast do young bats become infested with ectoparasites? - Acta Chiropterologica, 20(1): 187-194.
  • Zamora-Mejías D., J. B. Morales-Malacara, B. Rodríguez-Herrera, M. Ojeda, R. A. Medellín. 2020. Does latitudinal migration represent an advantage in the decrease of ectoparasitic loads in Leptonycteris yerbabuenae (Chiroptera)? - Journal of Mammalogy, 101(4): 979–989, https://doi.org/10.1093/jmammal/gyaa075

Update 2020 – 2021 to Vol. VII

Beron P. 2020. Acarorum Catalogus. VII. Raphignathoidea. Pensoft & Nat. Mus. Natur. Hist. Sofia: 306 pp.

Introduction

As a result of the recent contributions, the number of genera and species in Raphignathoidea (some of them uncertain status) is updated as follows:

Barbutiidae – 1 genus, 7 species

Caligonellidae – 5 genera, 66 species

Camerobiidae – 7 genera, 175 species

Cryptognathidae – 3 genera, 66 species

Dasythyreidae – 2 genera, 3 species

Dytiscacaridae – 1 genus, 3 species

Eupalopsellidae – 7 genera, 50 species

Homocaligidae – 2 genera, 8 species

Mecognathidae – 2 genera, 7 species

Raphignathidae – 3 (?) genera, 83 species

Stigmaeidae – 33 genera, 640 species

Xenocaligonellidae – 2 genera, 5 species

Raphignathoidea total - 68 genera, 1113 species

New contributions to the Catalogue

Fam. Barbutiidae

Barbutia arasbaraniensis Mohammad - Doustaresharaf et Bagheri

Barbutia arasbaraniensis Mohammad - Doustaresharaf & Bagheri, 2021a: 10

Type locality:

Distribution: Iran

Barbutia theroni Khaustov, Vorontsov, Perkovsky et Klimov

Barbutia theroni Khaustov, Vorontsov, Perkovsky & Klimov, 2021: 975

Type locality: Rovno

Distribution: Ukraine

Fam. Caligonellidae

Caligonella quinqueocellata Khaustov

Caligonella quinqueocellata Khaustov, 2021: 911

Type locality: Arabat Spit

Distribution: Crimea

Caligonella astragalusi Pishehvar et Khanjani

Caligonella astragalusi Pishehvar & Khanjani, 2020: 1989

Type locality: Nahavad, Hamedan Province

Distribution: Iran

Molothrognathus sibiricus Khaustov

Molothrognathus sibiricus Khaustov, 2022: 99

Type locality: Altai Republic, Kosh-Agach, 1765 m

Distribution: Russia (Altai Republic)

Molothrognathus tauricus Khaustov

Molothrognathus tauricus Khaustov, 2021: 917

Type locality: Arabat Spit

Distribution: Crimea

Neognathus sibiricus Khaustov

Neognathus sibiricus Khaustov, 2022: 99

Type locality: Tyumen Region, vicinity of lake Kushak

Distribution: Russia (Siberia)

Fam. Camerobiidae

Neophyllobius cibyci Paredes-León, Corona-López, Flores-Palacios et Toledo

Neophyllobius cibyci Paredes-León, Corona-López, Flores-Palacios & Toledo, 2016: 5

Type locality: dry forest 1 km south of San Andres de la Cal, Tepotzlán, Morelos

Distribution: Mexico

Host: inhabiting bromeliads (Tillandsia spp.) and living on two tree species (Quercus obtusata and Sapium macrocarpum)

Neophyllobius denizliensis Akyol

Neophllobius denizliensis Akyol, 2020b: 88

Type locality: Denizli Province

Distribution: Turkey

Neophyllobius electrus Zmudzinski

Neophyllobius electrus Zmudzinski, 2020: 698

Type locality: Eocene Baltic Amber

Distribution: Baltic Amber

Neophyllobius glaesus Zmudzinski

Neophyllobius glaesus Zmudzinski, 2020: 706

Type locality: Eocene Baltic Amber

Distribution: Baltic Amber

Neophyllobius tepoztlanensis Paredes-León, Corona-López, Flores-Palacios et Toledo

Neophyllobius tepoztlanensis Paredes-León, Corona-López, Flores-Palacios & Toledo, 2016: 14

Type locality: dry forest 1 km south of San Andres de la Cal, Tepotzlán, Morelos

Distribution: Mexico

Host: inhabiting bromeliads (Tillandsia spp.) and living on two tree species (Quercus obtusata and Sapium macrocarpum)

Neophyllobius tescalicola Paredes-León, Corona-López, Flores-Palacios et Toledo

Tillandsobius Bolland, 1985 = Tycherobius Bolland, 1986 (after Mirza, Kamran & Alatawi, 2022: 5)

Tillandsobius floridensis Bolland, 1986 = Tycherobius floridensis (Bolland) - Mirza et al., 2022: 5

Neophyllobius texanus McGregor, 1950 = Neophyllobius (Monophyllobius Mirza, 2022:6) texanus McGregor

Monobius Alatawi et Kamran, 2022: 4

Neophyllobius electrus Zmudzinski, 2020 = Monobius electrus (Zmudzinski, 2020) - Mirza et al., 2022: 5

Neophyllobius meyerae Bolland, 1991 = Monobius meyerae (Boland) - Mirza et al., 2022: 5

Neophyllobius tescalicola Paredes-León, Corona-López, Flores-Palacios & Toledo, 2016: 17

Type locality: dry forest 1 km south of San Andres de la Cal, Tepotzlán, Morelos

Distribution: Mexico

Host: in soil and litter under Quercus obtusata

Fam. Cryptognathidae

Cryptognathus karabagiensis Akyol

Cryptognathus karabagiensis Akyol, 2020c: 1644

Type locality: Karabag

Distribution: Turkey

Favognathus rosulatus Doğan et Doğan

Favognathus rosulatus Doğan & Doğan, 2020b: 71

Type locality: Turkey

Distribution: Turkey

Fam. Raphignathidae

Raphignathus kulaensis Akyol

Raphignathus kulaensis Akyol, 2020a: 1096

Type locality: Manisa and Denizli provinces

Distribution: Turkey

Raphignathus rakhshandehi Pishehvar et Khanjani

Raphignathus rakhshandehi Pishehvar & Khanjani, 2021: 185

Type locality: along Gamasiab river, Nahavand, Hamedan Prov.

Distribution: Iran

Raphignathus seraji Pishehvar et Khanjani

Raphignathus seraji Pishehvar & Khanjani, 2021: 186

Type locality: along Gamasiab river, Nahavand, Hamedan Prov.

Distribution: Iran

Fam. Stigmaeidae

Agistemus piquinnus Monjarás-Barrera et Johann

Agistemus piquinnus Monjarás-Barrera & Johann, 2020: 472

Type locality: Ciudad Victoria, Tamaulipas

Distribution: Mexico

Agistemus brasiliensis Matioli, Ueckermann & Oliveira from Mexico (Monjarás-Barrera et al., 2020)

Agistemus vulgaris Soliman et Gomaa from Egypt

Eustigmaeus crassifolius Bizarro et Johann

Eustigmaeus crassifolius Bizarro & Johann, 2020: 826

Type locality: Belém city (1°26’09.2”S, 48°26’28.6”W), Pará state

Distribution: Brazil

Eustigmaeus johnstoni Zhang et Gerson, 1995 – recorded also from Phlebotomus alexandri and from Sergentomyia (Sintonius) clydei and S. sintoni by Majidi, Hajiqanbar & Saboori (2019)

Eustigmaeus segnis (C.L. Koch) - for Spain (Santamaría et al., 2012)

Mediolata similans González-Rodríguez - for Spain (Santamaría et al., 2012)

Prostigmaeus amplius Doğan, Doğan et Türk

Prostigmaeus amplius Doğan, Doğan & Türk, 2020: 1076

Type locality: Turkey

Distribution: Turkey

Stigmaeus akimovi Kharadov

Stigmaeus akimovi Kharadov, 2021a: 43

Type locality: Altai Republic,

Distribution: Russia

Stigmaeus bagherii Rostami et Doustaresharaf

Stigmaeus bagherii Rostami & Doustaresharaf, 2021: 222

Type locality: East Azerbaijan Province, north-western Iran

Distribution: Iran

Stigmaeus exilis Doğan et Doğan

Stigmaeus exilis Doğan & Doğan, 2021: 20

Type locality: Sansa Gorge

Distribution: Turkey

Stigmaeus furcatus Dönel et Doğan, 2011 – new for Iran, recorded on Phlebotomus alexandri by Majidi, Hajiqanbar & Saboori (2019)

Stigmaeus fusus Summers, 1962 - new for Russia (Khaustov, 2021)

Stigmaeus lorestaniensis Ahmad-Hosseini et Jafari

Stigmaeus lorestaniensis Ahmad-Hosseini & Jafari, 2022: 238

Type locality: Shorab Vill., Lorestan Prov.

Distribution: Iran

Stigmaeus pulumurensis Doğan et Doğan

Stigmaeus pulumurensis Doğan & Doğan, 2020a: 42

Type locality: Pülümür Valley

Distribution: Turkey

Stigmaeus sinai Swift, 1987 – recorded also from Phlebotomus alexandri by Majidi, Hajiqanbar & Saboori (2019)

Stigmaeus strobiculatus Kharadov

Stigmaeus strobiculatus Kharadov, 2021a: 248

Type locality: Western Siberia

Distribution: Russia

Stigmaeus youngi (Hirst, 1926) - new for Iran, recorded on Phlebotomus alexandri, Sergentomyia theodori and S. sintoni (Majidi, Hajikanbar & Saboori, 2019)

Distribution of Raphignathoidea (suppl.)

EUROPE

Russia (Khaustov, 2021a,b) – Stigmaeus akimovi (Altai), S. scrobiculatus (Siberia)

Crimea (Khaustov, 2021c) – Caligonella quinqueocellata, Molothrognathus turcicus

Spain (Santamaría et al., 2012) – Eustigmaeus [Ledermulleria] segnis, Mediolata similans

Turkey (Doğan, Doğan & Erman. 2018; Akyol, 2020a, 2020b, 2020c, 2021a, 2021b; Akyol & Koc, 2020; Cobanoglu, Ögreten & Sade, 2021; Doğan, 2020; Doğan & Doğan, 2020a, b, c; Doğan & Doğan, 2021; Doğan, Doğan & Türk, 2018, 2020) – Cryptognathus karabagiensis, Favognathus rosulatus, Molothrognathus shirazicus, Paraneognathus wangae, Prostigmaeus amplius, P. kannjanii, Raphignathus gracilis, R. kulaensis, Stigmaeus exiis, S. pulumurensis, S. shabestariensis

Ukraine (Khaustov et al., 2021) – Barbutia theroni

ASIA

Iran (Majidi, Hajikanbar & Saboori, 2019; Siahkamari et al., 2020; Khosravinedzad, Jafari & Paktinat-Saeij, 2020; Pishehvar & Khanjani, 2021; Rostami, 2021; Gassemi & Hadjizadeh, 2021; Doustaresharaf & Bagheri, 2021a, 2021b, Ahmad-Hosseini & Jafari, 2022) – Barbutia arasbaraniensis, Eustigmaeus johnstoni, Raphignathus rakhshandehi, R. seraji, Stigmaeus bagherii, S. furcatus, S. lorestaniensis, S. sinai, S. youngi

Saudi Arabia (Mirza, Kamran & Alatawi. 2022) - Camerobia southcotti, Neophyllobius combreticola, N. denizliensis, N. lorestanicus, Tycherobius emadi

NORTH AMERICA

Mexico (Monjarás-Barrera et al., 2020) – Agistemus brasiliensis, A. piquinnus

SOUTH AMERICA

Brazil (Bizarro et al., 2020; Toldi et al., 2021 - Eustigmaeus crassifolius

Hosts

Diptera

Fam. Psychodidae

Phlebotomus (Paraphlebotomus) alexandri – Eustigmaeus johnstoni, Stigmaeus furcatus, S. sinai

P. (Phlebotomus) papatasi - Eustigmaeus johnstoni, S. sinai, S. youngi

Sergentomyia (Sintonius) clydei - Eustigmaeus johnstoni

Se. (Se.) sintoni - Eustigmaeus johnstoni, Stigmaeus youngi

Se. (Se.) theodori - Stigmaeus youngi

Publications on Raphignathoidea

(suppl. 2020 - 2021)

  • Abdelgayed S. A., N. M Abd El-Wahed, A. M. Ali, S. Eraky. 2019. Species Composition and Diversity of Mites Inhabiting Pomegranate Orchards at Assiut, Upper Egypt. - Acarines, 13: 29-36.
  • Ahmad – Hosseini M., Sh. Jafari. 2022. Stigmaeus lorestaniensis, a new species of the genus Stigmaeus (Acari: Stigmaeidae) from Southwest of Iran. - Persian Journal of Acarology, 11(2), serial No41: 237 – 243.
  • Akyol M. 2020a. A new species of the genus Raphignathus Dugès (Acari: Raphignathidae) from the Aegean region of Turkey. – Syst. Appl. Acarol., 25(6): 1095 – 1101.
  • Akyol M. 2020b. A new species of the genus Neophyllobius Berlese (Acari: Camerobiidae) from Denizli Province, Turkey. – Acarological Studies, 2(2): 88 –93.
  • Akyol M. 2020c. A new species of the genus Cryptognathus Kramer (Acari: Cryptognathidae) from the Aegaean Region of Turkey. - Syst. Appl. Acarol., 25(9): 1643 – 1648.
  • Akyol M. 2021a. Two New Records of Stigmaeid Mites (Acari: Stigmaeidae) for the Turkish Fauna. – KSU J. Agric. Nat., 24(2): 430 – 434.
  • Akyol M. 2021b. A new record for the mite fauna of Turkey: Molothrognathus shirazicus (Acari, Caligonellidae) and the first description of its protonymph. – Acarol. Stud., 3(1): 43 – 47.
  • Akyol M., K. Koc 2020. Erratum: M. Akyol & K.Koc, A new species and a new record of the genus Tycherobius (Acari: Camerobiidae) for the Turkish Fauna. – Syst. Appl. Acarol., 22(2): 21 – 27 (2017). – Syst. Appl. Acarol., 25(4): 773 – 774.
  • Bizarro G. L., A. C. da Silva Noronha, G.L. Da Silva, N. J. Ferla, L. Johanna. 2020. A new species of Eustigmaeus Berlese (Acari: Stigmaeidae) from Brazil. – Acarologia, 60(4): 825-830.
  • Doğan S. 2019. Raphignathoidea (Acari: Trombidiformes) of Turkey: A review of progress on the systematics, with an updated checklist. - Acarological Studies, 1(2): 129–151.
  • Doğan S. 2020. Correction: Raphignathoidea (Acari: Trombidiformes) of Turkey: A review of progress on the systematics, with an updated checklist. - Acarological Studies, 2(1): 58.
  • Doğan S., S. Doğan. 2018. The stigmaeid mites (Acari: Stigmaeidae) from Yedigoller, Ecence Mountains (Turkey). Abstr. Ecol. Intern. 2018 Symp., Kastamonu, 19-23 June 2018: 771.
  • Doğan Sibel, S. Doğan. 2020a. A new species of the genus Stigmaeus Koch (Acari, Stigmaeidae) from Pülümür Valley (Turkey)[Orig. Turk.]. – Acarological Studies, 2(1): 41 – 45.
  • Doğan S., S. Doğan. 2020b. A new species of the genus Favognathus (Acariformes: Cryptognathidae) from Turkey, with some taxonomic comments on other members of the genus. [Orig. Turk.]. – Acarological Studies, 2(1): 69 – 76.
  • Doğan S., S. Doğan. 2020c. Newly recorded stigmaeid mites (Acariformes: Raphignathoidea: Stigmaeidae) for the fauna of Turkey. – Acarological Studies, 2(2): 94 – 118.
  • Doğan S., S. Doğan. 2021. The first report of malformation in a stigmaeid mite species, Storchia robusta (Berlese) (Acariformes: Stigmaeidae). - International Journal of Acarology, 47(3): 270 – 271.
  • Doğan Salih, Sibel Dikaraoğlu Doğan. 2021 Stigmaeus exilis, a new fusiform species of Stigmaeus Koch (Acariformes: Stigmaeidae) from Sansa, Turkey. - Persian Journal of Acarology, 10(1): 19 – 28.
  • Doğan S., S. Doğan, O. Erman. 2018. Cryptognathid mites (Acari: Cryptognathidae) of Harsit Valley and Örümcek Forests (Turkey). - Abstr. Ecol. Intern. 2018 Symp., Kastamonu, 19-23 June 2018: 671.
  • Doğan S., S. Doğan, O. Erman. 2019a. Mites of Raphignathus Dugès (Acari: Raphignathidae) from Harsit Valley (Turkey), with taxonomic notes on other members of the genus. – Plant Prot. Bull., 59(1): 25 – 36.
  • Doğan S., S. Doğan, O. Erman. 2019b. New record of Eustigmaeus setiferus (Acari, Stigmaeidae) in Turkey. – Eurasia 3rd Intern. Congr. of Appl. Studies 28-30 June 2019 Sivas: 98 – 109.
  • Doğan S., S. Doğan, B. Kabasakal. 2021. Peritremal variations in Caligonella haddadi Bagheri & Maleki (Acariformes: Caligonellidae). – Syst. Appl. Acarology, 26(6): 1021 – 1025.
  • Doğan S., S. Doğan, M. Khanjani. 2021. On some species of the genus Ledermuelleriopsis Willmann (Acariformes: Stigmaeidae) in Turkey. - Syst. Appl. Acarol., 26(2): 455 – 463.
  • Doğan S., S. Doğan, M.B. Türk. 2018. Numerical variations in the body setae of Stigmaeus elongatus Berlese (Acari: Stigmaeidae) - Abstr. Ecol. Intern. 2018 Symp., Kastamonu, 19-23 June 2018: 438.
  • Doğan S., S. Doğan, M.B. Türk. 2020. A new species of the genus Prostigmaeus Kuznetzov (Trombidiformes: Stigmaeidae) from Turkey. – Syst. Appl. Acarol., 25(6): 1075 – 1084.
  • Doustaresharaf M.M., M. Bagheri. 2019. First report nof the Stigmaeus diversus (Acari: Stigmaeidae) from Iran. – Persian J. Acarol., 8(3): 277 – 279.
  • Mohammad-Doustarisharaf M., M. Bagheri. 2021a. Description of Barbutia arasbaraniensis sp. nov. (Acari: Trombidiformes: Barbutiidae) based on the deutonymph and male specimens from Iran. – Persian J. Acarol., 10(1): 9 – 17.
  • Mohammad-Doustarisharaf M., M. Bagheri. 2021b. Raphignathoid mites (Acariformes: Raphignathoidea) in parts of the Azerbaijan provinces of Iran. – Acarol. Stud., 3(2): 56 – 65.
  • Ghasemi A., J. Hajizadeh. 2021. Fauna and identification key for prostigmatid mites (Acari: Prostigmata) associated with greenhouses in Rasht city. – Plant Pest Research. A Quotarly Journal, 11(1): 1 – 14.
  • Gupta S.K., S. Ghoshal, A. Choudhury, B. Mukherjee. 2004. Phytophagous and predatory mite fauna of Sundarbans Biosphere Reserve: II. Some predatory mites occurring on mangrove vegetation and agri-horticultural crops. - Rec. zool. Surv. India, 103(3-4): 33-45.
  • Khaustov A.A. 2021a. A new species and a new record of Stigmaeus (Acari: Prostigmata: Stigmaeidae) from Western Siberia, Russia. - International Journal of Acarology, 47(3): 248-261.
  • Khaustov A.A. 2021b. Contribution to the Stigmaeidae (Acari: Prostigmata) fauna of the Altai Republic, Russia. – Acarina, 29(1): 43 – 66.
  • Khaustov A.A. 2021c. Two new species of Caligonellidae (Acari: Raphignathoidea) from Crimea. – Acarologia, 61(4): 910-927.
  • Khaustov A.A. 2022. Two new species of Caligonellidae (Acari: Raphignathoidea) from Western Siberia, Russia. – Acarologia, 62(1): 99 – 112.
  • Khaustov A.A., V.V. Abramov. 2021. A revision of Postumius (Acari, Prostigmata, Stigmaeidae). – Acarina, 29(1): 115 – 133.
  • Khaustov A.A., D.D. Vorontsov, E.E. Perkovski, P.B. Klimov. 2021. First fossil record of mite family Barbutiidae (Acari: Raphignathoidea) from late Eocene Rovno Amber, with a replacement name Hoplocheylus neosimilis nomen novum (Tarsocheylidae). – Syst. Appl. Acarol., 26(5): 973 – 980.
  • Khosravinedzad M., S. Jafari, S. Paktinat-Saeij. 2020. A part of the Bdelloidea and Raphignathoidea fauna (Trombidiformes: Prostigmata) in Lorestan Province, with Cyta kreiteri recorded for the first time from Iran. – Persian J. Acarol., 9(1): 87 – 92.
  • Majidi M., H. Hajikanbar, A. Saboori. 2019. Parasitic stigmaeid mites (Acari: Stigmaeidae) of phlebotomine sandflies (Diptera: Psychodidae) in Fars Province, southern Iran. - International Journal of Acarology, 45(1-2): 41-47.
  • Mirza J.H., M. Kamran, F. Alatawi. 2022. New Genus and New Subgenera of Camerobiid Mites (Acari: Prostigmata: Camerobiidae) with a Key to World Species of the Genus Neophyllobius. – Insects, 13, 344: 1 – 34. http://doi.org/10.3390/insects 13040344
  • Monjarás-Barrera J.I., G.L. Da Silva, J.C. Chacon-Hernandez, O.S. Da Silva, N.J. Ferla, L. Johann. 2020. A new species and new record of Agistemus Summers (Acari: Stigmaeidae) associated with Capsicum annuum L. var. glabriusculum (Solanaceae) from Northeastern of Mexico. – Acarologia, 60(2): 470 – 480.
  • Oudemans A.C. 1927. Acarologische aanteekeningen LXXXVIII. - Entomologische Berichten, 7(158): 257-263.
  • Paredes-León R., A. M. Corona-López, A. Flores-Palacios, V. H. Toledo. 2016. Camerobiid mites (Acariformes: Raphignathina: Camerobiidae) inhabiting epiphytic bromeliads and soil litter of tropical dry forest with analysis of setal homology in the genus Neophyllobius. - European Journal of Taxonomy, 202: 1-25.
  • Pishehvar Sh., M. Khanjani. 2020. Caligonella astragalusi n.sp. (Acari, Trombidiformes, Caligonellidae) from Western Iran. – Syst. Appl. Acarology, 25(11): 1988 – 1993.
  • Pishehvar Sh., M. Khanjani. 2021. Two new species of the genus Raphignathus (Acari: Raphignathidae) from Hamedan, Iran, with a key to the known species. – Syst. & Applied Acarology, 26(1): 185 – 198.
  • Rostami A., M. Mohammad - Doustaresharaf. 2021. Description of Stigmaeus bagherii n.sp. (Acari: Stigmaeidae), with a key to the known species of Stigmaeus from Iran. – International Journal of Acarology, 47: 222 – 232.
  • Santamaría J.M., M. L. Moraza, D. Elustondo, E. Baquero, R. Jordana, E. Lasheras, R. Bermejo, A. H. Ariño. 2012. Diversity of Acari and Collembola along a pollution gradient in soils of a pre-pyrenean forest ecosystem. - Environmental Engineering and Management Journal, 11(6): 1159 -1169.
  • Siahkamari S., N. Moeininaghadeh, M. Darbemamieh. 2020. Fauna of Prostigmatic mites (Trombidiformes: Prostigmata) associated with Quercus spp. in Bisotun natural preserved area and List of reported species from Oak in Iran (in Farsi). Toldi M., M.E. de Freitas, V.L. da Silva, M. Caumo, J.J. Ferla, C.R. Orlandi, T. Da- Costa, L. Johann, N.J. Ferla. 2021. Communities of predatory mites (Phytoseiidae and Stigmaeidae) in different environments of the Brazilian Pampa. – Acarologia, 61(1): 20 – 30.
  • Zmudzinski M. 2020. New fossil stilt-legged mites of Neophyllobius Berlese, 1886 (Acariformes, Camerobiidae) from Eocene Baltic amber. – J. of Palaeontology, 94(4): 696 – 715.

Update 2020 - 2021 to Vol. VIII

Beron P. 2020. Acarorum Catalogus. VIII. Cheyletoidea, Cloacaroidea. Pensoft & Nat. Mus. Natur. Hist. Sofia: 465 p.

Introduction

As a result of the recent contributions, the number of genera and species in and (some of them uncertain status) is updated as follows:

Cheyletidae – 80 genera, 474 species

Psorergatidae – 3 genera, 77 species

Demodecidae – 8 genera, 122 species

Harpyrhynchidae - 16 genera, 135 species

Syringophilidae - 63 genera, 416 species

Cloacaridae - 6 genera, 15 species

Epimyodicidae – one genus, four species

New contributions to the Catalogue

Fam. Cheyletidae

Bak deleoni Yunker, 1961 – new for India (Modak et al., 2004)

Chelacaropsis moorei Baker, 1949 – India, W. Bengal (A. Gupta et al., 2006), India, Andaman Isl. (Gupta & Ghosh, 1980)

Cheyletus cacahuamilpensis Baker, Cheyletomimus congensis (Cunliffe) for Egypt (Abdelgayed et al., 2020), Cheyletus cacahuamilpensis - for Brazil (Pellegrini & Ferreira, 2013)

Ornithocheyletia lukoschusi Smiley, O. pingus Smiley, O. hallae Smiley, O. gersoni Smiley – new for Egypt (Halawa, Osman & Ahmed, 2019)

Samsinakia trilobites Bochkov et Mironov - new for Iran (Kazemi & Pektinat-Saeij, 2021)

Fam. Demodecidae – Pormann et al., 2021

Fam. Harpyrhynchidae

Harpyrhynchoides [Harpyrhynchus] vercammeni (Lawrence) – new for Egypt (Halawa, Osman & Ahmed, 2019)

Fam. Syringophilidae

Syringophiloidus minor (Berlese) – new for Egypt (Halawa, Osman & Ahmed, 2019)

Bubophilus aegolius Skoracki, Kosicki et Kwiecinski

Bubophilus aegolius Skoracki, Kosicki & Kwiecinski, 2021: 355

Type locality: Poland, Zielenogora, Stary Kisielin

Distribution: Poland, Finland, Switzerland

Host: Aegolius funereus (Strigiformes: Strigidae)

Distribution of Cheyletoidea and Cloacoidea (suppl.)

ASIA

India (Modak et al., 2004; Gupta et al, 2004; A. Gupta et al., 2006; Gupta & Ghosh, 1980) – Chelacaropsis moorei, Bal deleoni

Iran (Kazemi & Paktinat-Saeij, 2021) – Samsinakia trilobitus

AFRICA

Egypt (Abdelgayed et al., 2020; Atwa et al., 2018; Halawa et al., 2019) – Cheyletus cacahuamilpensis, Cheyletomimus congensis, Harpyrhynchoides [Harpyrhynchus] vercammeni, Ornithocheyletia lukoschusi, O. pingus, O. hallae, O. gersoni, Syringophiloidus minor

SOUTH AMERICA

Brazil (Pellegrini & Ferreira, 2013) - Cheyletus cacahuamilpensis

Hosts

Aves

Columbiformes

Fam. Columbidae

Streptopelia senegalensis aegyptiaca - Harpyrhynchoides [Harpyrhynchus] vercammeni, Ornithocheyletia lukoschusi, O. hallae, O. gersoni, O. pinguis

Columba livia domestica - Ornithocheyletia lukoschusi

Passeriformes

Fam. Passeridae

Passer domesticus niloticus - Syringophiloidus minor

Strigiformes

Fam. Stigidae

Aegolius funereusBubophilus aegolius

Publications on Cheyletoidea and Cloacoidea

(suppl. 2020 – 2021 and ommited)

  • Abdelgayed S. A., N. M Abd El-Wahed, A. M. Ali, S. Eraky. 2020. Species Composition and Diversity of Mites Inhabiting Pomegranate Orchards at Assiut, Upper Egypt. - Acarines, 13: 29-36.
  • Atwa W.A., M.E. El-Naggar, A.M. Khalil, M.E. El-Shaer, Z.M.M. Mostafa. 2018. Biological studies on cheyletid predator mite, Cheletogenes ornatus (Canestrini & Fanzago) when fed on the different preys. – Egypt. Acad. J. Biol. Sci., A. Entomology, 11(5): 21 – 29.
  • Gupta A., A. Chalterjee, A. K. Sanyal, S. K. Gupta. 2006. On a collection of soil prostigmatid mites (Acari) from southern parts of West Bengal, India. - Rec. Zool.. Surv. India: 106(4): 61-71.
  • Gupta S.K., S.K. Ghosh. 1980. Some prostigmatid mites (Acarina) from Andaman and Nicobar islands. - Rec. Zool. Surv. India, 77: 189 - 121.
  • Halawa A.M., S. A. A. Osman, H. A. A. Ahmed. 2019. Diversity and infestation rate of mites associated with some birds in upper and lower Egypt. – Annals of Agric. Sci., Moshtohor, 57(2): 517 – 524.
  • Kazemi S., S. Paktinat-Saeij. 2021. First record of Samsinakia trilobites (Prostigmata, Cheyletidae) from Iran. – Persian J. Acarol., 10(1): 107 – 110.
  • Khaustov A.A. Review of free-living predatory cheyletid mites (Acari: Cheyletidae) of Western Siberia, Russia. – Syst. Appl. Acarology, 26(1): 199 – 239.
  • Modak A., G. K. Saha, N. Tandon, S. K. Gupta. 2004. Faunal diversity and habitat preference of house dust mites in West Bengal in relation to nasobronchial allergic disorders. - Rec. zoo!. Surv. India, 102(1-2): 137-146.
  • Pellegrini Th, G., R. L. Ferreira. 2013. Structure and interactions in a cave guano-soil continuum community. - European Journal of Soil Biology, 57: 19-26.
  • Portmann A.N., L. Vieira, F. Majolo, G.L. da Silva. 2021. Demodex folliculorum and Demodex brevis (Acari: Demodecidae) and their association with facial and non- facial pathologies. - International Journal of Acarology, 47(5): 396-403.
  • Skoracki M., J.Z. Kosicki, Z. Kwiecinski. 2021. Distribution of the parasitic mite Bubophilus aegolius sp.n. (Acariformes: Syringophilidae) on the boreal owl Aegolius funereus (L.)(Strigiformes: Strigidae) and the low effectiveness of infestation. – Eur. Zool. J., 88(1): 352 – 362.
  • Zmudzinski M., M. Skoracki, B. Sikora. 2020. An updated checklist of quill mites of the family Syringophilidae (Acariformes: Prostigmata): v2020. Available at: https://sites.google.com/site/syringophilidae/

Beron P. 2021. Acarorum Catalogus. IX. Schizoglyphoidea (Schizoglyphidae) Histiostomatoidea (Histiostomatidae, Guanolichidae) Canestrinioidea (Canestriniidae, Chetochelacaridae, Lophonotacaridae, Heterocoptidae) Hemisarcoptoidea (Chaetodactylidae, Hyadesiidae, Algophagidae, Hemisarcoptidae, Carpoglyphidae, Winterschmidtiidae). Pensoft & Nat. Mus. Natur. Hist. Sofia: 396 p.

The recent article of R. Haitlinger altered drastically the system of fam. Canestriniidae with the description of 9 new genera and the change of the taxonomy of 24 species.

  • Haitlinger R. 2022. A review of host-commensal associations between canestriniid mites (Astigmata: Canestriniidae) and Insects with keys and descriptions of new genera. – The European Zoological Journal, 89(1).

The new genera are: Balearella, (Ibizella Haitlinger et Šundić, 2016, syn.), Cercediphotia, Ciprusenia, Ismaeilienia, Javanellina, Kuramansenia, Midiphotia, Phalerotusellus, Sumatranella.

24 species are transferred to another or new genera: Balearella asilahica, B. balearica, B. samsinaki, Ciprusenia carabicola, C. izabelae, C. kamelskyi, C. sellnicki, C. viviannae, Imaielienia egypti, I.maroccana, Javanellina macgillavryi, Kuramasenia illegalis, K. paavoi, K. pekingensis, K. pictura, K. ramoni, Midiphotia hispanica, M. jureceki, Phaleratusellus alatus, Psedocanestrinia damiani, P. roberti, Sumatranella batocerae, S. elae and S. phoebinae.

As a result of the recent contributions, the number of genera and species in this volume (some of them uncertain status) is updated as follows:

Schizoglyphidae – 1 genus, 1 species

Histiostomatidae – 58 genera, 522 species

Guanolichidae – 3 genera, 4 species

Canestriniidae – 100 genera, 348 species

Chetochelacaridae – 1 genus, 1 species

Lophonotacaridae – 1 genus, 1 species

Heterocoptidae – 12 genera, 28 species

Chaetodactylidae – 5 genera, 110 species

Hyadesiidae – 2 genera, 47 species

Algophagidae – 8 genera, 22 species

Hemisarcoptidae – 8 genera, 24 species

Carpoglyphidae – 4 genera, 10 species

Winterschmidtiidae – 25 genera, 138 species

The series Acarorum Catalogus

Vol. I (2008) – Erythraeidae, Smarididae, Calyptostommatidae – 3 fam., 68 genera, 959 sp.

II (2011) – Listrophoroidea - 5 fam., 100 genera, 925 sp.

III (2014) - Opilioacarida, Holothyrida, Mesostigmata - 13 fam., 85 genera, 422 sp.

IV (2015) – Myobiidae, Pterygosomatidae - 2 fam., 68 genera, 707 sp.

V (2015) – Sarcoptoidea, Analgoidea, 19 fam., 145 genera, 509 sp.

VI (2020) – Rhinonyssidae, Spinturnicidae - 2 fam., 28 genera, 568 sp.

VII (2020) - Raphignathoidea - 12 fam., 68 genera, 1113 sp.

VIII (2020) – Cheyletoidea, Cloacaroidea - 13 fam., 177 genera, 1243 sp.

IX (2021) – Schizoglyphoidea, Histiostomatoidea, Canestrinioidea, Hemisarcoptoidea - 13 fam., 233 genera, 1240 sp.

X (2022) - Labidostommatoidea, Eupodoidea, Tydeoidea, Paratydeoidea, Anystoidea, Caeculoidea, Adamystoidea, Pomerantzioidea – 20 fam., 230 genera, 967 sp. Supplements: 1 fam., 11 genera, 149 sp. Added: 9 new genera, described by Haitlinger (2022).

Total: 102 fam., 1222 genera, 8815 sp.

Acknowledgements

Our gratitude is due to many specialists, sharing their experience, providing the nessessary publications or material. First of all we thank the arachnologists A.Fain, Fan Quing-Hai, A. Bochkov, Z. Feider, A. Khaustov, M. Bertrand, M.Shiba, C.Willmann, H.E.Vistorin, K.Schmölzer, P.Robaux, J.Cooreman, M. Zacharda, M.K.P.Meyer, H.Franz, I.Coineau, R.W.Strandtmann, B.Halliday, E.A.Ueckermann. For the updating of the previous volumes the author is grateful to R. Haitlinger, M. Šundiċ, A. Saboori, Si-Yuan Xu, R. Bassini-Silva, G. Gabryś, A. Bochkov, B. OConnor, M. Skoracki, M.M. Vázquez, H. Klompen, M.S. Araújo, M.V. Orlova, M. Fajfer, M. Akyol, Salih Doğan, S. Doğan, M. Zmudzinski, E.A.Ueckermann, A. Khaustov, Fan Quing-Hai, L.F.d.O. Bernardi, J. Noei, M. Hakimitabar, A. Labrzycka, J. Makol, P.B. Klimov, I. Dimov.

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