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© 2022 Petar Beron.
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Citation:
Beron P (2022) ACARORUM CATALOGUS X. Trombidiformes, Prostigmata, Superfamilia Labidostommatoidea (Labidostommatidae), Superfamilia Eupodoidea, (Eupodidae, Dendrochaetidae, Rhagidiidae, Eriorhynchidae, Pentapalpidae, Penthalodidae, Penthaleidae, Proterorhagiidae, Strandtmanniidae), Superfamilia Tydeoidea, Ereynetidae, Superfamily Paratydeoidea, Paratydeidae, Superfamilia Anystoidea, (Anystidae, Erythracaridae,Teneriffiidae, Pseudocheylidae, Stigmocheylidae), Superfamilia Caeculoidea (Caeculidae), Superfamilia Adamystoidea (Adamystidae), Superfamilia Pomerantzioidea (Pomerantziidae). Advanced Books. https://doi.org/10.3897/ab.e68612
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This is the last volume of the series Acarorum Catalogus. We tried to follow the system generally accepted. After the publication of the first volume of the series (Erytraeoidea, Calyptostomatoidea) in 2008 appeared a comprehensive treatment of mites (Manuel of Acarology III of Krantz & Walter, Eds. 2009), in which many changes in the system of Acari are proposed. Acariformes are already a superorder, Prostigmata – suborder, but within the order Trombidiformes.
Here is treated first the infraorder Labidostommatina with its only superfamily Labidostommatoidea and the only family Labidostommatidae with five genera and 75 species. The other infraorder, treated in the present volume (Eupodoidea) includes the superfamily Eupodoidea with nine families, including five described after 1979. Treated are also the superfamilies Tydeoidea (fam. Ereynetidae), Paratydeoidea (Paratydeidae), Anystoidea (five families), Caeculoidea (Caeculidae), Adamystoidea (Adamystidae), and Pomerantzioidea (Pomerantziidae). In this volume are analysed 230 genera and 967 species of mites.
Labidostommatidae – 75 sp. (+4 fossils)
Eupodidae and Cocceupodidae – 95 sp.
Dendrochaetidae – 1 sp.
Rhagidiidae – 150 sp.
Eriorhynchidae – 5 sp.
Pentapalpidae – 1 sp.
Penthalodidae – 44 sp.
Penthaleidae – 27 sp.
Proterorhagiidae – 1 sp.
Strandtmanniidae – 1 sp.
Ereynetidae – 180 sp.
Paratydeidae – 13 sp.
Anystidae – 102 sp.
Erythracaridae – 68 sp.
Teneriffiidae – 28 sp.
Pseudocheylidae – 23 sp.
Stigmocheylidae – 5 sp.
Caeculidae – 114 sp.
Adamystidae – 28 sp.
Pomerantziidae – 6 sp.
This last volume of the series Acarorum Catalogus ends with an update of the former nine volumes. Added to these volumes are another genera and species.
The recent survey of order Trombidiformes (Zhang, Ed., 2011) defined suborder Prostigmata as containing four infraorders and 40 superfamilies. This volume concerns 20 families of prostigmatid mites, mostly soil and plant inhabitants. Some (Speleognathinae, Riccardoella) are parasites of vertebrates and snails.
Labidostommatoidea Ewing, 1934: 64
Phalanx Stomatostigmata Oudemans, 1906
The superfamily contains only one family (Labidostommatidae) with five genera and 75 species.
Nicoletiellidae G. Canestrini, 1891
Labidostomidae Oudemans, 1904
Labidostommatidae Oudemans, 1906
Strongly hitinized free living mites (600 – 1500 µm long), usually yellow and with one large lense-formed organ behind the side eyes. About 75 species, living in moss, leaf litter and other organic materials. Feed on small insects and mites.
The names of the main genus have varied since their description and are used differently even at present by various authors: Labidostoma, Labidostomma, Nicoletiella (as genus or subgenus of Labidostomma), Nicoletiellidae, Labidostomidae, Labidostommatidae.
The first species has been described very early (Acarus denticulatus Schrank, 1776), but the base of the taxonomy of the family was laid with the descriptions of Labidostomma Kramer, 1879 and Nicoletiella Canestrini, 1882, than several other authors contributes with more new species and discussions: Stoll, Berlese, Lombardini, Coineau, Ewing, Feider, Greenberg, Meyer & Ryke, Oudemans, Grandjean, Robaux, Sellnick, Shiba, Štorkán, Willmann, Womersley, Iavorschi, and others. Some authors should be mentioned specially. Atyeo & Crossley (1961) described six new species from New Zealand. Revision of the family and several new descriptions are due to the Romanian authors Feider & Vasiliu, including generic/subgeneric taxa (Grandjeanellina, Atyeonella, Sellnickiella, Dicastriella, Cornutella, Mahunkiella).
The modern treatment of the family has been started by M. Bertrand since 1980 and so far he published (partly with co-authors) 24 new species and the subgenera Pseudocornutella and Pselistoma.
Fam. Labidostommatidae - 4 gen., 75 recent sp.
Akrostomma Robaux, 1977 - 6 sp.
Eunicolina Berlese, 1911 - 4 sp. (plus one fossil)
Labidostomma Kramer, 1879 - 44 sp
Atyeonella Feider et Vasiliu, 1969 - 8 sp.
Labidostomma Kramer, 1879 – 16 sp.
Nicoletiella Canestrini, 1882 – 17 sp. (plus one fossil)
Cornutella Feider et Vasiliu, 1969 – 2 sp.
Pselistoma Bertrand, 1990 – 1 sp.
Pseudocornutella Sidorchuk et Bertrand, 2013 – 1 sp. (fossil)
Sellnickiella Feider et Vasiliu, 1969 – 20 sp. (plus one fossil)
Dicastriella Feider et Vasiliu, 1970 - 7 sp.
Sellnickiella Feider et Vasiliu, 1969 – 13 sp.
Nicoletiellidae G.Canestrini, 1891: (Type genus: Nicoletiella Canestrini, 1882)
Labidostomidae Oudemans, 1904; Bertrand, 1990a: 31
Labidostommatidae Oudemans, 1906
Sellnickiellinae Feider & Vasiliu, 1969: 206
Nicoletiellinae Can.: Feider & Vasiliu, 1969: 204
Eunicolinae Feider & Vasiliu, 1969: 204
Type genus: Labidostomma Kramer, 1879
Akrostomma Robaux, 1977: 453
Type species: Akrostomma grandjeani Robaux, 1977
Akrostomma coineaui Bertrand, 1983: 160
Type locality: Isla de Ons (Provincia Pontevedra)
Distribution: Spain
Akrostomma coralloides Bertrand & Coineau, 1979: 101
Type locality: Banyuls (valley of Baillaurie, Pouade Road)
Distribution: France
Akrostomma gabriella Crossley & Simmons: 2015: 459
Type locality: California, Los Angeles County, San Gabriel Mts., Glendale Ridge Road, San Dimas Experimental Forest
Distribution: USA (California)
Akrostomma grandjeani Robaux, 1977: 453
Type locality: H. Cowell Redwoods State Park (Sta. Cruz Co.), California
Distribution: USA (California)
Labidostomma plumosum Greenberg, 1952: 201
Labidostoma (Labidostoma) plumosum Greenberg: Bertrand, 1990a: 36
Akrostomma plumosum (Greenberg): Crossley & Simmons, 2015: 459
Type locality: Gale’s Canyon, Winters, California
Distribution: USA (California)
Labidostomma zangherii Lombardini, 1943: 21
Akrostomma zangherii (Lombardini)(?): Bertrand, 1990a: 35
Type locality: Italy
Distribution: Italy
Eunicolina Berlese, 1911: 430
Type species: Eunicolina tuberculata Berlese, 1911. Redescription, Vistorin 1980: 376
Grandjeanellina Feider & Vasiliu, 1969: 206 - Type species: Eunicolina nova Sellnick, 1931; syn. by Bertrand, 1990a: 35
Eunicolina glaesi Bertrand, Sidorchuk & Hoffeins, 2015: 323
Type locality: Baltic amber
Distribution: Baltic amber (Eocene)
Eunicolina nova Sellnick, 1931: 772; Willmann,1941: 51; Beron, 1995: 5
Grandjeanellina nova (Sellnick): Feider & Vasiliu, 1969: 206
Type locality: Korfu (Greece)
Distribution: Bulgaria, Croatia, Greece, Romania
Eunicolina porifera Greenberg, 1932: 201; Feider & Vasiliu, 1969: 206; Bertrand, 1990a: 35
Type locality: Cape Henry, Virginia
Distribution: USA
Eunicolina travei Coineau, 1964: 153; Feider & Vasiliu, 1969: 206; Bertrand, 1990a: 35
Type locality: Saint-Paul d’Eygeaux (Haute-Vienne), France
Distribution: France
Eunicolina tuberculata Berlese, 1911: 430; Feider & Vasiliu, 1969: 206; Vistorin, 1978a: 279; Bertrand, 1990a: 35
Type locality: “in agro Tridentino”,”Tiarno” (Italy)
Distribution: Croatia, Italy
Labidostomma Kramer, 1879: 13
Labidostoma Michael, 1880:
Labidostoma Bertrand, 1990a: 36
Panoplia von Heyden, 1826
Type species: Acarus denticulatus Schrank, 1776
Ceratoacarus Ewing, 1913: 128
Type species: Ceratoacarus pacificus Ewing, 1913
Atyeonella Feider & Vasiliu, 1969: 205 (genus)
Labidostoma (Atyeonella): Bertrand, 1990a: 36 (subgenus)
Type species: Labidostoma fictiluteum Atyeo et Crossley, 1961
Labidostoma fictiluteum Atyeo & Crossley, 1961b: 40
Atyeonella fictiluteum (Atyeo et Crossley): Feider & Vasiliu, 1969: 206
Labidostoma (Atyeonella) fictiluteum Atyeo & Crossley: Bertrand, 1990a: 36
Type locality: Ngongotaha, Rotorua, Auckland
Distribution: New Zealand
Labidostoma glandula Atyeo & Crossley, 1961b: 44
Atyeonella fictiluteum (Atyeo et Crossley): Feider & Vasiliu, 1969: 206
Labidostoma (Atyeonella) glandula Atyeo et Crossley: Bertrand, 1990a: 36
Type locality: Ngongotaha, Rotorua, Auckland
Distribution: New Zealand
Labidostoma malleolus Atyeo & Crossley, 1961b: 37
Atyeonella maleolus (Atyeo et Crossley): Feider & Vasiliu, 1969: 206
Labidostoma (Atyeonella) malleolus Atyeo et Crossley: Bertrand, 1990a: 36
Type locality: Kapiti Island, Wellington
Distribution: New Zealand
Labidostoma multifarium Atyeo & Crossley, 1961b: 33
Atyeonella multifarium (Atyeo et Crossley): Feider & Vasiliu, 1969: 206
Labidostoma (Atyeonella) multifarium (Atyeo et Crossley): Bertrand, 1990a: 36
Type locality: Orepuke, Southland
Distribution: New Zealand
Labidostoma ocellatum Atyeo & Crossley, 1961b: 42
Atyeonella ocelatum (Atyeo et Crossley): Feider & Vasiliu, 1969: 206
Labidostoma (Atyeonella) ocellatum Atyeo & Crossley: Bertrand, 1990a: 36
Type locality: Coromandel Peninsula, Auckland
Distribution: New Zealand
Labidostoma (Atyeonella) peruviana Bertrand, 1990c: 182
Type locality: Sierra Garavito, Prov. de Cusco
Distribution: Peru
Atyeonella simplex Shiba, 1972: 48
Labidostoma (Atyeonella) simplex (Shiba): Bertrand, 1990a: 36
Type locality: Hokkaido, Poroshiri-sanso at the foot of Mt. Poroshiri Hadaka
Distribution: Japan (Hokkaido)
Labidostoma striatum Atyeo & Crossley, 1961b: 46
Atyeonella striatum (Atyeo et Crossley): Feider & Vasiliu, 1969: 206
Labidostoma (Atyeonella) striatum Atyeo et Crossley: Bertrand, 1990a: 36
Type locality: Takahe Valley, Te Anau, Otago
Distribution: New Zealand
Labidostomma Kramer, 1879: 13
Labidostoma Michael, 1888
Type species: Acarus denticulatus Schrank, 1776 (= Labidostomma luteum Kramer, 1879 - Redescription, Lehtinen & Niemi 1995: 217)
Labidostoma (Labidostoma) aethiopica Meyer & Ryke, 1959: 417; Bertrand, 1990a: 36; Bertrand & Theron, 1990: 248
Type locality: South Africa, Bathurst
Distribution: South Africa
Labidostomma amasonica Feider, Vasiliu & Calugar, 1974: 424
Labidostoma (Labidostoma) amasonica Feider, Vasiliu et Calugar: Bertrand, 1990a: 36
Type locality: Brazil [Amazonia]
Distribution: Brazil
Labidostoma (Labidostoma) angolensis Bertrand & Theron, 1990a: 248
Type locality: Luachine, Luangue, Tchitentga
Distribution: Angola
Labidostomma barbae Greenberg, 1952: 198
Labidostoma (Labidostoma) barbae Greenberg: Bertand, 1990a: 36
Type locality: 3 mi southeast of Thomasville, Thomas County, Georgia
Distribution: USA (Georgia)
Labidostoma (Labidostoma) fissurata Bertrand & Theron, 1990a: 248
Type locality: Potchefstroom
Distribution: South Africa
Labidostomma guadalupense Bertrand & Corpuz-Raros, 1997: 363
Type locality: Leyte, Guadalupe
Distribution: Philippines
Labidostoma integrum (Berlese): Berlese, 1911: 429; 1913: 138; Costa, 1961: 259
Labidostoma (Labidostoma) integrum (Berlese): Bertrand, 1990a: 36; Bertrand et al., 2012: 235
Labidostomma integrum : Thor, 1931: 73; Feider & Vasiliu, 1970: 371
Type locality: Umbria (Italy)
Distribution: France (incl. Corsica), Israel, Italy (incl. Sardinia), Kazakhstan, Romania, Spain
Labidostomma integrum : Thor, 1931: 73; Feider & Vasiliu, 1970: 371; Bloszyk et al., 2003: 173.
Labidostomma (L.) caucasicum Reck, 1940: 452; Feider & Vasiliu, 1969: 205 and Bloszyk et al., 2003: 173 consider this species as synonym of L. integrum
Type locality: D.R.Congo
Distribution: D.R.Congo
Labidostoma (Labidostoma) intermedia Bertrand, Bagheri, Akbari, Yazdanian, Irani-Nejad, Mohajer et Saboori, 2012: 237; Jalaeian, Kamali & Bertrand, 2013: 16
Type locality: Golestan province, Gorgan
Distribution: Iran
Labidostoma (Labidostoma) mirax Bertrand & Theron, 1990a: 255
Type locality: Tsitsikama, Cape Province
Distribution: South Africa
Labidostomma motasi Iavorschi, 1992: 47
Type locality: cave Peštera de la Movile, Dobroudja
Distribution: Romania
Labidostomma himalayensis Bhattacharyya, 1966: 46
Type locality: Rohla Valley
Distribution: NW Himalaya (India)
Labidostoma (Labidostoma) neotropica Stoll, 1891: 9; 1993: 27; Bertrand, 1990a: 36
Type locality: Retalhuleu
Distribution: Guatemala
Labidostomma nepalense Feider & Vasiliu, 1968a: 630
Labidostoma (Labidostoma) nepalense Feider & Vasiliu: Bertrand, 1990a: 36
Type locality: Nepal, 27o19’ N, 87o31’E
Distribution: Nepal
Labidostoma (Labidostoma) percilliata Bertrand & Theron, 1990a: 248
Type locality: Luachine, Luangue, Tchitentga
Distribution: Angola
Nicoletiella Canestrini, 1882: 65
Type species: Nicoletia cornuta G. Canstrini et Fanzago, 1877
Mahunkiella asiatica Feider & Vasiliu, 1972: 382
Labidostoma (Nicoletiella) asiatica Feider & Vasiliu: Bertrand, 1990a: 36
Type locality: Prov. South Phenan, Bong ha ri
Distribution: North Korea
Labidostomma caloderum Greenberg, 1952: 197
Labidostoma (Nicoletiella) caloderum Greenberg: Bertrand, 1990a: 36
Type locality: Univ. of Kansas Natural History Reservation, Douglas County, Kansas
Distribution: USA (Kansas)
Mahunkiella coreana Feider & Vasiliu, 1972: 380
Labidostoma (Nicoletiella) coreana (Feider et Vasiliu): Bertrand, 1990a: 36
Type locality: Prov. Kanwon, Kum-gang san
Distribution: N. Korea
Labidostoma (Nicoletiella) corsicum Bertrand, 1981: 36
Type locality: Corsica
Distribution: France (Corsica)
Acarus denticulatus Schrank, 1776: 125; 1781: 520
Labidostoma denticulatum : Oudemans, 1904: 132; 1916: 250; Kontschán, 2008: 361
Labidostoma (Nicoletiella) denticula [sic](Schrank): Bertrand, 1990a: 36
Labidostoma (Nicoletiella) denticulata (Schrank): Vistorin, 1978a: 279; Bertrand et al., 2015: 331
Nicoletiella denticulata (Schrank): Sellnick, 1940: 31; Willmann, 1951: 143; 1955: 217; Franz, 1954: 363; Mihelčič, 1958b: 274
Nicoletiella (Cornutella) denticulata (Schrank): Bloszyk, 1980: 72
Nicoletiella lyra Willmann, 1932: 160; 1934: 50; 1939: 433; 1941: 47; 1955: 217; Dudich, Kolosvary & Szalay, 1940: 53; Bloszyk et al., 2003: 172. Type loc.: Postoina (Slovenia).
Nicoletiella corcyraea Berlese, 1911 (Corfu)
Labidostomma (Nicoletiella) denticulatum (Schrank): Újváry & Kontschán, 2010; Pfliegler & Bertrand, 2011: 178
Type locality: Austria
Distribution: Austria, Germany, Hungary, Island, Italy, Poland, Spain, Slovenia
Nicoletiella fergusoni Robaux, 1977: 443
Labidostoma (Nicoletiella) fergusoni (Robaux): Bertrand, 1990a: 36
Type locality: Mount Madona (Ridge Road)(Sta. Clara Co.), California
Distribution: USA (California)
Labidostomma franzi Bertrand, 1983: 163
Labidostoma (Nicoletiella) franzi Bertrand: Bertrand, 1990a: 36
Type locality: Galicia, La Ramosa
Distribution: Spain
Labidostomma glymma Grandjean, 1942: 119
Labidostoma (Nicoletiella) glymma Bertrand, 1990a: 36
Type locality: France
Distribution: France (cont. and Corsica),
Nicoletiella (Nicoletiella) luteoides Bertrand, 1990c: 181
Type locality: Osorno (40o34’S, 73o09’W)
Distribution: Chile
Labidostomma luteum Kramer, 1879: 13; Thor, 1931: 72; Cooreman, 1943: 2; 1959: 17; Franz, 1954: 363; Bottazzi, 1950; Schweizer & Bader, 1963: 247; Lehtinen & Niemi, 1995: 211
Labidostoma luteum Kramer: Mihelčič, 1958a: 40; 1958b: 273; Hallas, 1978: 31; Schmölzer, 1993: 58
Nicoletiella (Nicoletiella) luteum (Kramer): Vistorin, 1978: 272; Bloszyk et al., 2003: 171
Labidostoma (Nicoletiella) luteum Kramer: Bertrand, 1990a: 36
Labidostomma (Nicoletiella) luteum elongatum Grandjean, 1942: 189 (Périgueux, France). Listed by Bertrand (1990: 37) as species.
Nicoletiella vejdovskyi Štorkán, 1938 – syn. by Lehtinen & Niemi, 1995
Type locality: Germany
Distribution: Austria, Belgium, Denmark, England, Finland, France, Germany, Holland, Ireland, Italy, Sicily, Poland, Spain, Sweden, Switzerland
Labidostoma multisetosa Shiba, 1969: 85
Labidostoma (Nicoletiella) multisetosa Shiba: Bertrand, 1990a: 36
Type locality: Shiga Heights IBP Area
Distribution: Japan
Ceratoacarus pacificus Ewing, 1913: 127
Nicoletiella pacifica (Ewing): Thor, 1931: 76
Labidostoma (Nicoletiella) pacifica (Ewing): Bertrand, 1990a: 36
Type locality: Oregon
Distribution: USA
Labidostoma (Nicoletiella) paleoluteum Dunlop & Bertrand, 2011: 193
Type locality: Baltic camber (loc. idet.), Palaeogene (Eocene)
Distribution: Baltic area
Labidostomma luteum repetitor Grandjean, 1942: 122
Labidostomma repetitor : Lehtinen & Niemi, 1995: 220
Labidostoma (Nicoletiella) repetitor Bertrand, 1990a: 36
Labidostoma Jacquemarti Coineau, 1964: 543 – Syn. by Lehtinen & Niemi, 1995 (Type: Banyuls-sur-Mer, France)
Labidostoma (Nicoletiella) jacquemarti Coineau: Bertrand, 1990a: 36
Type locality: France
Distribution: France
Nicoletiella romanica Feider & Vasiliu, 1968: 31
Labidostoma (Nicoletiella) romanica (Feider & Vasiliu): Bertrand, 1990a: 36
Type locality: the regions of Jassy, Suceava, Ploiesti, Banat, 200 – 800 m
Distribution: Romania
Nicoletiella spelaeophila Willmann, 1940: 209; Feider & Vasiliu, 1969: 205 consider “Nicoletiella speleofila [sic]” and “N. carpaticum” to be synonymes of “N. (C.) cornutum”.
L. spelaeophila (Willmann, 1940) – Cooreman, 1951: 10
Type locality: «Carniolia, Höhle bei Radna«
Distribution: Romania, Slovenia
Labidostomma vialeae Bertrand, 1981: 28
Labidostoma (Nicoletiella) vialeae Bertrand: Bertrand, 1990a: 36
Type locality: Galeria (Corsica)
Distribution: France (Corsica)
Caligonas virescens Targioni Tozzetti, 1886
Labidostomma virescens (Targioni Tozzetti)
Type locality: Italy
Distribution: Italy
Nicoletiella (Cornutella) Feider & Vasiliu, 1969: 205
Type species: Nicoletia cornutum Canestrini et Fanzago, 1877
Mahunkiella Feider & Vasiliu, 1972: 375, synonymized by Bertrand et al., 2012: 242
Nicoletia cornutum Canestrini & Fanzago, 1877: 120
Nicoletiella cornuta (Canestrini et Fanzago): R. Canestrini, 1882: 65; Vistorin, 1978a: 272
Labidostomma cornuta (Canestrini & Fanzago): Cooreman, 1954: 11; 1959: 16
Nicoletiella (Cornutella) cornuta (Canestrini et Fanzago): Feider & Vasiliu, 1970: 371
Labidostomma (Nicoletiella) cornuta (Canestrini et Fanzago): Beron, 1995: 5
Nicoletiella (Cornutella) cornutum (Canestrini et Fanzago): Pfliegler & Bertrand, 2011: 178
Type locality:
Distribution: Algeria, Austria, Bulgaria, Croatia, France, Hungary, Poland, Romania, Slovenia
Nicoletiella (Cornutella) ezoensis Shiba, 1972: 46
Type locality: Hokkaido, Kita Cirque, Mt. Poroshiri, Hidaka
Distribution: Japan (Hokkaido)
Labidostoma multisetosa Shiba, 1969: 86
Mahunkiella multisetosa (Shiba): Feider et Vasiliu, 1972: 377
Labidostomma (Cornutella) multisetosa: Bertrand et al., 2012: 234, 242
Type locality: Shiga Heights (Japan)
Distribution: Japan, North Korea
The species Labidostoma corcyraeum Berlese, 1911 from “Corcyra”(Corfu) and Nicoletiella (Cornutella) absoloni Willmann, 1932 () are listed by Feider & Vasiliu, 1969, but not in the revision of Bertrand, 1990.
Pseudocornutella Sidorchuk & Bertrand, 2013: 31
Type species: Labidostomma (Pseudocornutella) electri Sidorchuk et Bertrand, 2013
Labidostoma (Pseudocornutella) electri Sidorchuk & Bertrand, 2013: 31 (fossil)
Type locality: Baltic Amber
Distribution: Baltic Amber
Pselistoma Bertrand, 1990: 183
Type species: Labidostomma (Pselistoma) legendrei Bertrand, 1990
Labidostomma (Pselistoma) legendrei Bertrand, 1990c: 183
Type locality: Osorno region
Distribution: Chile
Sellnickiella Feider & Vasiliu, 1969: 206
Type species: Labidostoma brasiliense Sellnick, 1922
Dicastriella Feider & Vasiliu, 1970a: 283 (genus)
Sellnickiella (Dicastriella): Bertrand, 1990a: 35 (subgenus)
Type species: Dicastriella covarubiasi Feider et Vasiliu, 1970
Dicastriella caeca Feider, Vasiliu & Calugar, 1974: 420
Sellnickiella (Dicastriella) caeca (Feider, Vasiliu et Calugar): Bertrand, 1990c: 35
Type locality: Misitumi (reg. Tarapaca)
Distribution: Chile
Dicastriella coineaui Feider & Vasiliu, 1970a: 303; Feider, Vasiliu & Calugar, 1974: 422
Sellnickiella (Dicastriella) coineaui (Feider et Vasiliu): Bertrand, 1990c: 35
Type locality: Polpaico (Santiago)
Distribution: Chile
Dicastriella covarubiasi Feider & Vasiliu, 1970a: 297; Feider, Vasiliu & Calugar, 1974: 423
Sellnickiella (Dicastriella) covarubiasi (Feider et Vasiliu): Bertrand, 1990c: 35
Type locality: Limpache (Valparaiso)
Distribution: Chile
Dicastriella fungifera Feider & Vasiliu, 1970a: 306
Sellnickiella (Dicastriella) fungifera (Feider et Vasiliu): Bertrand, 1990c: 35
Type locality: Limache (Valparaiso)
Distribution: Chile
Labidostomma (Dicastriella) fusiformis Bertrand, 1990c: 179
Type locality: Osorno (40o34’S, 73o09’W)
Distribution: Chile
Dicastriella mediohexagonata Feider, Vasiliu & Calugar, 1974: 419
Sellnickiella (Dicastriella) mediohexagonata (Feider, Vasiliu et Calugar): Bertrand, 1990c: 35
Type locality: Farellones (Prov. Santiago)
Distribution: Chile
Sellnickiella Feider & Vasiliu, 1969: 206
Type species: Labidostoma brasiliense Sellnick, 1922
Labidostomma adelaideae Womersley, 1935: 152; Atyeo & Crossley, 1961b: 83
Sellnickiella (Sellnickiella) adelaidae (Womersley): Bertrand, 1990a: 35
Sellnickiella adelaideae (Womersley): Halliday, 1998: 66
Type locality: Adelaida
Distribution: Australia
Labidostoma brasiliense Sellnick, 1922: 45
Sellnickiella brasiliense (Sellnick): Feider & Vasiliu, 1969: 206; 1970: 289
Sellnickiella (Sellnickiella) brasiliensis (Sellnick): Bertrand, 1990a: 35
Type locality: Passa Quatro, Sul de Minas
Distribution: Brazil
Sellnickiella (Sellnickiella) capensis Bertrand & Theron, 1992: 46
Type locality: Hawaquas Montains nr. Paarl, Cape Province
Distribution: South Africa
Labidostoma circinus Atyeo & Crossley, 1961b: 35
Sellnickiella (Sellnickiella) circinus (Atyeo et Crossley): Bertrand, 1990a: 36
Type locality: Ngongotaha, Rotorua, Auckland
Distribution: New Zealand
Sellnickiella decemlabeculata Feider & Vasiliu, 1970: 297; Feider, Vasiliu & Calugar, 1974: 418
Sellnickiella (Sellnickiella) decemlabeculata: Bertrand, 1990c: 35
Type locality: Brazil, Nova Teutonia
Distribution: Brazil, Chile
Labidostoma höegi Thor, 1931a: 57
Labidostomma höegi Thor, 1931b: 73; Meyer & Ryke, 1959: 419
Sellnickiella (Sellnickiella) hoegi (Thor): Bertrand, 1990a: 35
Type locality: Port Elizabeth
Distribution: South Africa
Sellnickiella (Sellnickiella) macroscopa Bertrand, 1990c: 179
Type locality: Cantin (39o22’ S, 72o38’W)
Distribution: Chile
Sellnickiella mediumreticulata Feider & Vasiliu, 1970a: 289; Feider, Vasiliu & Calugar, 1974: 417
Type locality: Nova Teutonia
Distribution: Brazil
Sellnickiella (Sellnickiella) natalensis Bertrand & Theron, 1992: 48
Type locality: St. Lucia (Natal)
Distribution: South Africa
Sellnickiella (Sellnickiella) pulchra Bertrand, 1990c: 178
Type locality: Floresta de Tijuca, reg. of Rio de Janeiro
Distribution: Brazil
Sellnickiella schusteri Feider, Vasiliu & Calugar, 1974: 414
Sellnickiella (Sellnickiella) schusteri Feider, Vasiliu et Calugar: Bertrand, 1990a: 35
Type locality: Serra de Cantareira (Sao Paulo)
Distribution: Brazil
Labidostomma womersleyi Atyeo & Crossley, 1961a: 84
Type locality: Remarkable Creek, Wilmington, South Australia
Distribution: Australia
Sellnickiella balticae Sidorchuk & Bertrand, 2013: 27
Type locality: Baltic Amber
Distribution: Baltic Amber
EUROPE
Austria (Schrank, 1781; Willmann, 1939, 1951; Franz, 1954; Vistorin, 1978; Schmölzer, 1993) –Labidostomma cornuta, L. denticulatum, L. luteum, L. lyra
Bulgaria (Beron, 1995) – Eunicolina nova, Labidostomma (Cornutella) cornutum
Croatia (Willmann, 1940, 1941; Vistorin, 1978а) – Eunicolina tuberculata, Labidostomma (Cornutella) cornutum
Czech Rep. (Willmann, 1939)
Denmark (Luxton, 1966, 1981; Hallas, 1978) – Labidostomma luteum
Finland (Lehtinen & Niemi, 1995) – Labidostomma (L.) luteum
France (contin.) (Cooreman, 1954, 1959; Coineau, 1964a, 1964b; Bertrand, 1980; Bertrand & Coineau, 1979) – Akrostomma coralloides, Labidostomma (Cornutella) cornutum, Labidostomma (Nicoletiella) repetitor (= L. (Nicoletiella) jacquemarti), L. (N.) luteum, Labidostomma (L.) integrum
Corsica (Bertrand, 1981; Bertrand et al., 2012) - Labidostomma (Nicoletiella) corsicum, L. (N.) glymma, L. (N.) vialeae, Labidostomma (L.) integrum
Georgia (Reck, 1940) – Labidostomma caucasicum (= L. integrum)
Germany (C.L.Koch, 1838; Oudemans, 1916; Willmann, 1955; Vistorin, 1978a) – Labidostomma denticulatum, L. luteum, L. lyra
Great Britain (Michael, 1880) – Labidostomma luteum
Corfu (Berlese, 1911; Sellnick, 1931) – “Labidostoma corcyraeum””(= L. denticulatum), Eunicolina nova
Hungary (Willmann, 1937; Dudich, Kolosváry & Szalay, 1940; Újváry & Kontschán, 2010; Pfliegler & Bertrand, 2011) – Labidostomma denticulata, L. lyra, L. (Cornutella) cornutum
Iceland (Sellnick, 1940) – Labidostomma denticulata (actually L. luteum, see Lehtinen & Niemi, 1995)
Ireland (Halbert, 1915) – Nicoletiella cornuta (actually L. luteum, see Lehtinen & Niemi, 1995)
Italy (contin.) (Berlese, 1887; 1911; Lombardini, 1943; Bottazzi, 1956; Vistorin, 1978a) – Akrostomma zangherii (sp. dubia), Eunicolina tuberculata, Labidostomma denticulatum, L. integrum, L. luteum
Sicily (Berlese, 1883; Bottazzi, 1956) – Labidostomma luteum
Sardinia (Bertrand et al., 2012) – Labidostomma repetitor, L. (L.) integrum
Poland (Bloszyk, 1980; Błoszyk & Czarnota, 1998; Bloszyk, Konwerski, Napierala, Goldyn & Gawronska, 2007; Bloszyk et al., 2017; Bloszyk, Adamski & Napierała, 2018) – Labidostomma (Nicoletiella) denticulatum, L. (N.) luteum, L. (Cornutella) cornutum
Romania (Cooreman, 1951; Feider & Vasiliu, 1968, 1970; Iavorschi, 1992; Kontschán, 2008) - Labidostomma motasi, L. (Cornutella) cornutum, L. (L.) integrum (= L. caucasicum), L.(Nicoletiella) denticulata, L. (Nicoletiella) romanica, Eunicolina [Grandjeanellina] nova
Slovakia (Bloszyk et al., 2022) – Labidostomma luteum
Slovenia (Willmann, 1932; Willmann, 1940, 1941; Vistorin, 1978a) – Labidostomma (Nicoletiella) lyra, Labidostomma cornuta, L. spelaeophila
Spain (cont.) (Mihelčič, 1958a,b; Bertrand, 1983 ; Bertrand et al., 2012) – Akrostomma coineaui, Labidostomma (Nicoletiella) luteum, [L. colomerense], L. franzi, L. (L.) integrum
Sweden - Labidostomma (Nicoletiella) luteum
Switzerland (Schweizer & Bader, 1963) – Labidostomma (Nicoletiella) luteum
Turkey (all) (Urhan & Özmen, 2006) – Labidostomma sp.
Baltic Area (Dunlop & Bertrand, 2011; Sidorchuk et Bertrand, 2013) - Labidostomma (Nicoletiella) paleoluteum (fossil), Labidostomma (Pseudocornutella) electri (fossil), Sellnickiella balticae (fossil)
AFRICA
Algeria (Grandjean, 1942) – Labidostomma integrum, L. (Cornutella) cornutum
Angola (Bertrand & Theron, 1990a) – Labidostomma (L.) angolensis, L. (L.) percilliata
DR Congo (Cooreman, 1954) – Labidostomma integrum schoutedeni
South Africa (Thor, 1931a; Meyer & Ryke, 1959; Bertrand & Theron, 1990a, 1992) - Labidostomma (L.) aethiopica, L. (L.) fissurata, L.(L.) mirax, L.(L.) percilliata, Sellnickiella (S.) capensis, S.(S.) hoegi, S.(S.) natalensis
ASIA
India (Bhattacharyya, 1966) – Labidostomma himalayensis
Iran (Bertrand et al., 2012; Jalaeian, Kamali & Bertrand, 2013) – Labidostomma (L.) intermedia
Israel and Palestina - Labidostomma (L.) intermedia
Japan (Shiba, 1969, 1972, 2004) – Labidostomma (Atyonella) simplex, Labidostomma (Cornutella) ezoensis, L. (Cornutella) mutisetosa
Kazakhstan (Bertrand et al., 2012) - Labidostomma (L.) integrum
Korea (N.) (Feider & Vasiliu, 1972) – Labidostomma asiatica, L. (Cornutella) mutisetosa, M. multisetosa
Nepal (Feider & Vasiliu, 1968a) - Labidostomma nepalense
Philippines (Bertrand & Corpus Raros, 1997) - Labidostomma (L.) guadalupense
NORTH AMERICA
USA (Ewing, 1913; Greenberg, 1952; Robaux, 1977; Crossley & Simmons, 2015) - Akrostomma gabriella, A. grandjeani, A. plumosum, Eunicolina porifera, Labidostomma (L.) barbae, Labidostomma (Nicoletiella) caloderum, L. (N.) fergusoni, L. (N.) pacifica,
SOUTH and CENTRAL AMERICA
Brazil (Sellnick, 1922; Feider & Vasiliu, 1970; Feider, Vasiliu & Calugar, 1974; Bertrand, 1990) – Labidostomma amasonica, Sellnickiella (S.) brasiliensis, S.(S.) decemlabeculata, S. (S.) mediumreticulata, S. (S.) schusteri, S. (S.) pulchra
Chile (contin.) (Feider & Vasiliu, 1970; Feider, Vasiliu & Calugar, 1974; Bertrand, 1990c) – Labidostomma (Nicoletiella) luteoides, L. (Pselistoma) legendrei, Sellnickiella (S.) decemlabeculata, Sellnickiella (Dicastriella) caeca, S. (D.) coineaui, S. (D.) covarubiasi, S. (D.) fungifera, S.(D.) fusiformis, S. (D.) mediohexagonata, Sellnickiella (S.) macroscopa,
Guatemala (Stoll, 1891) – Labidostomma (Labidostomma) neotropica
Peru (Bertrand, 1990c) – Labidostomma (Atyeonella) peruviana
OCEANIA
Australia (continent) (Wommersley, 1935; Atyeo & Crossley Jr., 1961a; Bertrand, 1997; Halliday, 1998) – Sellnickiella adelaideae, S. biunguiculata, S. womersleyi
New Zealand (Atyeo & Crossley Jr. 1961b) – Sellnickiella (S.) circinus, Labidostomma Atyonella fictiluteum, L. (A.) glandula, L. (A.) malleolus, L. (A.) multifarium, L. (A.) ocellatum, L. (A.) striatum
Eupodoidea Banks, 1894
According to Zhang, Ed. (2011), the superfamily consists of nine families (four of them described recently), with 85 genera and 309 species. The four “old” families (Eupodidae, Penthaleidae, Penthalodidae and Rhagidiidae) account for 277 of these species.
Eupodides C.L. Koch, 1842: 60
Eupodidae: Murray, 1877: 117
Small or middle-sized (160 – 1000 µm) mites, living in moss, leaf litter or on plants. Predators or plant parasites. Members of Benoinyssus Fain have been found in the nasal mucus of Naja melanoleuca (Serpentes: Elapidae).
Longtime after the descriptions of genus Eupodes Koch, 1835 and the family Eupodidae again by C.L. Koch (1842), the family was left with this genus plus Protereunetes Berlese, 1923, Linopodes C.L. Koch, 1835 and later Cocceupodes Sig Thor, 1934 (the last two being considered after 2010 to belong to a separate family Cocceupodidae Jesionowska, later synonymized with Eupodidae and restaured). Followed the description of five further genera by Fain (1958), A. Baker (1987), Barilo (1991), Fain & Camerik (1994) and Baker & Lindquist (2002). The important revision of Thor & Willmann (1941) was followed by modern contributions by Meyer & Ryke (1960, South Africa), Strandtmann (1971, Alasca), Strandtmann & Goff (1978, Hawaii), Shiba (1978, Japan), Qin Ting-Kui (1998, Australia and New Zealand), Olivier & Theron (1997, 1998, Southern Africa), and others. Several other authors contributed considerably to the study of these mites: Thor in Norway, Strandtmann in Antarctica, Abou-Awad in Egypt, Corpuz-Raros in the Philippines. Most of the modern research is being done in countries outside Europe and the identity of the 26 “Eupodes” species, described by C.L. Koch from Germany and enumerated by Thor & Willmann (1941), remains unclear. Only eight of them have been discussed by modern authors.
Cocceupodidae is a small family of mites, detached from Eupodidae and consisting of three genera and 23 valid species (Jesionowska, 2010). The study of the family starts with Acarus [Linopodes] motatorius Linnaeus, 1758. Linopodes C.L. Koch, 1835 is the oldest genus, includes mites with very long front legs. They live in wet places, rotten wood and under stones, often in caves. Koch (1838) describes from Germany 12 “Linopodes” species, their identity is still unclear. Some Cocceupodes are known from the far North or from Antarctica.
Khaustov (2014) synonymized Cocceupodidae under Eupodidae and Filieupodes under Cocceupodes and described from Crimea a new genus and species Pseudeupodes porosus. Later Khaustov (2015) added another two new genera and species Pseudopenthaleus tauricus and Echinoeupodes echinus, thus the families Eupodidae and Cocceupodidae (combined) seem to include 14 genera and 95 species. Other Polish specialists, based on molecular phylogeny, restaured the validity of the family Cocceupodidae and of the genus Filieupodes Jesionowska. They concluded that “(1) the genera Linopodes, Filieupodes and Cocceupodes are a separate monophyletic lineage of familial rank and the sister group of Eupodidae s.s., and (2) the genus Filieupodes should be considered a separate genus being a sister group of the Linopodes-Cocceupodes clade” (Szudarek-Trepto, Kaźmierski, Dabert et al. 2020).
Fam. Eupodidae C.L. Koch, 1842 - 14 gen., 95 sp.
Aethosolenia Baker et Lindquist, 2002 – 1 sp.
Benoinyssinae Fain, 1958 – 10 sp.
Caleupodes A. Baker, 1987 – 1 sp.
Claveupodes Strandtmann et Prasse, 1977 – 1 sp.
Cocceupodes Sigh Thor, 1934 – 8 sp.
Echinoeupodes Khaustov, 2017 – 2 sp.
Eupodes C.L. Koch, 1835 – 47 sp.
Filieupodes Jesionowska, 2010 – 9 sp.
Linopodes C.L. Koch, 1835 - 10 sp.
Neoprotereunetes Fain et Camerik, 1994 – 1 sp.
Niveupodes Barilo, 1991 – 1 sp.
Pseudeupodes Khaustov, 2014 – 1 sp.
Pseudopenthaleus Khaustov, 2015 – 2 sp.
Xerophiles Jesionowska, 2003 – 1 sp.
Eupodides C.L. Koch, 1842: 60
Eupodini Canestrini, 1886: 693
Eupodidae Murray, 1877: 117
Type genus: Eupodes C.L. Koch, 1835
Benoinyssinae Fain, 195. Type genus: Benoinyssus Fain, 1958: 178.
Type genus: Cocceupodes Sig Thor, 1934 (sensu Jesionowska, 2005)
Aethosolenia Baker & Lindquist, 2002: 2
Type species: Aethosolenia laselvensis Baker et Lindquist, 2002
Aethosolenia laselvensis Baker & Lindquist, 2002: 4
Type locality: Heredia Province, La Selva Biological Station, 50-150 m
Distribution: Costa Rica
Benoinyssus Fain, 1958: 178
Type species: Benoinyssus najae Fain, 1958
Benoinyssus celatus Olivier & Theron, 1997a: 317
Type locality: Cafunfe river
Distribution: Angola
Eupodes ereynetoides Strandtmann & Prasse, 1976: 2; Santamaría et al., 2012: 1164
Benoinyssus ereynetoides (Strandtmann et Prasse): Olivier & Theron, 1997a: 302
Type locality: Etzsdorf/ Saalkreis (East Germany)
Distribution: Germany, Spain
Benoinyssus falcatus Olivier & Theron, 1997a: 311
Type locality: Northern Province, Louis Trichardt
Distribution: South Africa
Eupodes momeni Abou-Awad, 1984: 333
Benoinyssus momeni (Abou-Awad): Olivier & Theron, 1997a: 302
Type locality: Ain Shams Univ., El-qaliubiya Region
Distribution: Egypt
Benoinyssus najae Fain, 1958: 138; 1959: 358
Type locality: Kaninya
Distribution: Rwanda
Host: Naja melanoleuca (Serpentes: Elapidae), nasal mucus
Benoinyssus oconnori Bizarro Wurlitzer & Da Silva, 2020: 3
Type locality: municipality of Novo Mundo, Mato Grosso state
Distribution: Brazil
Host: from soil and litter
Benoinyssus ornatus Olivier & Theron, 1997a: 317
Type locality: Northern Province, Giyani (South Africa)
Distribution: Angola, Swaziland, South Africa
Benoinyssus pusillus Olivier & Theron, 1997a: 314
Type locality: Northern Province, Warmbaths
Distribution: South Africa
Benoinyssus serratus Olivier & Theron, 1997a: 302
Type locality: Northern Province, Turfloop (South Africa)
Distribution: Botswana, Namibia, South Africa, Swaziland
Benoinyssus sp. – Iran (Darbemamieh, Hajiqanbar & Khanjani, 2013: 454)
Opsereynetes tuberculatus Baker, 1945: 18
Benoinyssus tuberculatus (Baker): Hunter, 1964: 181
Ereynetes (Opsereynetes) tuberculatus Baker: Fain, 1964b: 106
Type locality: Desierto de los Leones
Distribution: Mexico
Caleupodes A. Baker, 1987: 103
Type species: Caleupodes reticulatus A. Baker, 1987
Caleupodes reticulatus A. Baker, 1987: 104
Type locality: Suffolk, Hollesley Common
Distribution: England
Claveupodes Strandtmann & Prasse, 1977: 10
Type species: Claveupodes delicatus Strandtmann et Prasse, 1977
Claveupodes delicatus Strandtmann & Prasse, 1977: 11; Martin, 1983: 15
Type locality: Etzdorf/Saalkreis, East Germany
Distribution: Germany, New Zealand
Cocceupodes Sig Thor, 1934: 128 (sensu Jesionowska, 2005)
Type species: Eupodes clavifrons R. Canestrini, 1886
Cocceupodes australis Strandtmann & Tilbrook, 1968: 51; Shiba, 1971: 222; Jesionowska, 2010: 643. Strandtmann & Prasse (1977: 9) synonymize it with C. mollicellus.
Type locality: Deception Island
Distribution: Deception Island (South Shetland Islands), Japan
Cocceupodes breweri Strandtmann, 1971: 87; Jesionowska, 2010: 641
Type locality: Anaktuvik Pass, Alaska
Distribution: USA (Alaska), Greenland
Cocceupodes communis Shiba, 1969c: 68; Jesionowska, 2010: 643
Type locality: Shiga Heights IBP Area, Nagano Pref.
Distribution: Japan
Cocceupodes gracilongus Olivier & Theron, 2003: 186; Jesionowska, 2010: 641
Type locality: S. Afr., Northern Province, Ga-Selati River, near Tzaneen
Distribution: South Africa
Cocceupodes longisolenidiatus Jesionowska, 2007: 352; 2010: 641
Type locality: Poland
Distribution: Poland
Eupodes mollicellus C.L. Koch, 1838: fa. 19, t. 23
Eupodes clavifrons R. Canestrini, 1886: 716 (type loc. Trentino); Haarløv, 1942 : 28
Cocceupodes clavifrons (R. Canestrini): Sig Thor, 1934: 128; Schweizer & Bader, 1963: 218; Strandtmann, 1971: 85
Cocceupodes mollicellus : Oudemans, 1937: 891; Thor & Willmann, 1941: 36; Mihelčič, 1957;
Macfadyen, 1954: 283; Haarløv, 1957: 26; Strandtmann & Prasse, 1977: 8; Strandtmann &
Goff, 1978: 133; Jesionowska, 2010: 641
Cocceupodes curticlava Thor, 1934: 178; Thor & Willmann, 1941: 36; Macfadyen, 1954: 283;
Haarløv, 1957: 27 (syn. by Strandtmann & Prasse, 1977: 8)
Type locality: Germany
Distribution: Austria, Denmark, Germany, Italy, Norway, Hawaii, Greenland, Switzerland, Yan Mayen I.
Cocceupodes planiticus Shiba, 1978: 141; Jesionowska, 2010: 643
Type locality: Japan
Distribution: Japan
Cocceupodes stellatus Strandtmann & Prasse, 1977: 6; Jesionowska, 2010: 641
Type locality: Etzsdorf/ Saalkreis (East Germany)
Distribution: Germany, Greenland (cf. stellatus)
Echinoeupodes Khaustov, 2017: 29
Type species: Echinoeupodes Khaustov, 2017: 29
Echinoeupodes echinus Khaustov, 2017: 31
Type locality: nr. Yalta
Distribution: Crimea
Protereunetes turgidus Shiba, 1978: 145
Echinoeupodes turgidus (Shiba): Khaustov, 2017: 31
Type locality: Gobo, Wakayama
Distribution: Japan
Eupodes C.L. Koch, 1835: fa. 1, t. 18 - 19
Megamerus Dugès, 1834: 50 (non Mac Leay, 1827). Enumerated are: longipes, inflatus, ovalis, celer, roseus, castaneus, fallax
Egypteupodes B.A. Abou-Awad, 1984: 329 (syn. by Olivier & Theron,1997: 58). Type: E. strandtmanni Abou-Awad, 1984
Type species: Eupodes striola C.L. Koch, 1835
Eupodes acanthus Olivier & Theron, 1998: 284
Type locality: Northern Province, Debegeni Falls
Distribution: South Africa
Eupodes alaskanensis Strandtmann, 1971: 79
Type locality: Alaska, Cape Beaufort
Distribution: USA (Alaska)
Eupodes angardi Strandtmann & Sømme, 1977: 138
Type locality: Queen Maud Land, Brattskarvet
Distribution: Antarctica
Eupodes bakeri Abou-Awad, El-Sawaf & Abdel-Khalek, 2006: 46
Type locality: Egypt
Distribution: Egypt
Eupodes berensis Shiba, 1976: 85
Type locality: Malaysia
Distribution: Malaysia
Eupodes berlesei Thor, 1912: 236; Schweizer & Bader, 1963: 217
Type locality: Italy
Distribution: Denmark, Italy, Switzerland
Eupodes brevipes Banks, 1916: 14
Type locality: Laguna Beach, California
Distribution: USA (California)
Eupodes callejae Corpuz- Raros, 2005: 135
Type locality: Baybay, Leyte Is.
Distribution: Philippines (Leyte Is., Luzon Is.)
Eupodes cordiformis Shiba, 1978: 135
Type locality: Kuwabara, Matsuyama, Ehime
Distribution: Japan
Eupodes crozetensis Strandtmann & Davies, 1972: 50; Darbemamieh, Hajiqanbar & Khanjani, 2013: 454
Type locality: French Base Buildings, Possession Island, Crozet Islands
Distribution: Iran, Possession Island, Crozet Islands (sub-antarctic)
Eupodes exiguus Booth, Edwards & Usher, 1985: 392
Type locality: South-Orkney-Islands
Distribution: Antarctica (South-Orkney-Islands)
Eupodes fusifer R. Canestrini, 1886: 714
Type locality: San Remo, Veneto
Distribution: Italy
Eupodes fusiferellus Meyer & Ryke, 1960: 483
Type locality: Bathurst
Distribution: South Africa
Eupodes garciai Corpuz- Raros, 2005: 137
Type locality: Luzon (UPLB campus, College, Laguna)
Distribution: Philippines (Luzon, Leyte Isls)
Type locality: Egypt
Distribution: Egypt
Eupodes gilvus C.L. Koch, 1838: fa. 20, t. 1
Type locality: Regensburg
Distribution: Germany
Megamerus haltica Haller, 1880: 364
Eupodes haltica (Haller): Thor & Willmann, 1941: 25
Type locality: “Moos am Thunersee”
Distribution: Switzerland
Eupodes hamatus Olivier & Theron, 1997: 72
Type locality: Western Cape, Lamberts Bay
Distribution: South Africa
Eupodes hawaiiensis Strandtmann & Goff, 1978: 129; Goff, 1987: 31
Type locality: Hawaii Isl., Kilauea Forest Reserve, 1620 m
Distribution: Hawaii
Eupodes hiemalis C.L. Koch, 1838: fa. 19, t. 4; Thor & Willmann, 1941: 20; Lindroth et al., 1973: 125
Type locality: Regensburg (Germany)
Distribution: Germany, Iceland
Eupodes hjartdaliae Thor, 1934a: 213
Type locality: Hjartdal, Thelemarken
Distribution: Norway
Eupodes iconicus C.L. Koch, 1838: fa.19, t. 7; 1842: fa. 3, p.69; Walckenaer, 1847: 537; Oudemans, 1937: 884; Thor & Willmann, 1941: 12
Type locality: Regensburg (Germany)
Distribution: Germany, Italy
Eupodes indentatus Olivier & Theron, 1997: 69
Type locality: Mt. Inn Hotel, Soutpansberg
Distribution: South Africa
Eupodes infirmus Shiba, 1978: 133
Type locality: Mt. Saragamine, Shigenobu-cho, Ehime
Distribution: Japan
Eupodes leucomelas C.L. Koch, 1838: fa.19, t.14
Type locality: Regensburg
Distribution: Germany
Eupodes littoralis Shiba, 1978: 141
Type locality: Wake, Matsuyama, Ehime
Distribution: Japan
Eupodes longipilus Thor, 1934: 178; Meyer & Ryke, 1960: 487
Type locality: Cape Town
Distribution: South Africa
Eupodes longisetatus Strandtmann, 1964: 148; Shiba, 1969c: 66; 1969a: 149
Type locality: Campbell Island
Distribution: Japan, New Zealand (Campbell Island)
Eupodes lumbarus Shiba, 1976: 88
Type locality: Malaysia
Distribution: Malaysia
Eupodes lutatus Olivier & Theron, 1997: 72
Type locality: Zululand, St Lucia
Distribution: South Africa
Eupodes marinus Banks, 1896: 75; 1907: 596; Thor, 1934: 206; Thor & Willmann, 1941: 27
Type locality: Sea Cliff, New York
Distribution: USA
Eupodes milvinus C.L. Koch, 1838: fa. 19, t.1; 1842: fa. 3, p.70; Walckenaer, 1847: 537; Oudemans, 1937: 890; Thor & Willmann, 1941: 22
Type locality: Oberpfalz
Distribution: Germany, Italy, Holland, Sweden
Eupodes minipilus Olivier & Theron, 1998: 281
Type locality: Northern Province, Giyani (South Africa)
Distribution: Botawana, Cabo Verde Islands, South Africa, Zimbabwe
Protereunetes minutus Strandtmann, 1967: 55
Eupodes minutus (Strandtmann): Strandtmann, 1970: 89; Luxton, 1990: 400
Tydeus antipodus : Womersley (1937)
Eupodes antipodus : Baker (1970)
Type locality: South Georgia, Bay of Isles
Distribution: South Georgia, Macquarie I., New Zealand (South Island)
Eupodes niloticus Abou-Awad & El-Bagouri, 1985: 5; Abdelgayed, 2019: 32
Type locality: Egypt
Distribution: Egypt
Eupodes ocellatus Willmann, 1952: 159
Type locality: Wanderooge I.
Distribution: Germany
Eupodes okinoshimaensis Shiba, 1978: 138
Type locality: Nakamura, Okinoshima, Shimane
Distribution: Japan
Eupodes onoi Shiba, 1976: 92
Type locality: Malaysia
Distribution: Singapore, Malaysia
Eupodes parafusifer Meyer & Ryke, 1960: 483; Olivier & Theron, 1998: 276
Type locality: Mooi River, Potchefstroom (South Africa)
Distribution: Angola, South Africa
Eupodes parvus Booth, Edwards & Usher, 1985: 394
Type locality: South-Orkney-Islands
Distribution: Antarctica (South-Orkney-Islands)
Eupodes parvus grahamensis Booth, Edwards & Usher, 1985: 395
Type locality: Antarctica, Takaki Promontory, Brabant Island
Distribution: Antarctica
Еupodes pasohensis Shiba, 1976: 86
Type locality: Pasoh Forest Reserve, Negeri Sembilan, W. Malaysia
Distribution: Malaysia
Eupodes pseudoclavifrons R. Canestrini, 1886: 717; Thor, 1934: 205; Schmölzer, 1993: 59
Cocceupodes pseudoclavifrons (R. Canestrini): Schweizer & Bader, 1963: 218
Type locality: Italy (Trentino)
Distribution: Austria, Italy, Switzerland
Eupodes pseudolumbarus Shiba, 1976: 90; Corpuz-Raros, 2005: 140
Type locality: Pasoh Forest Reserve, Negeri Sembilan, W. Malaysia
Distribution: W.Malaysia, Philippines (Luzon, Leyte)
Eupodes pusillus Shiba, 1976: 93; Corpuz-Raros, 2005: 143c
Type locality: Pasoh Forest Reserve, Negeri Sembilan, W. Malaysia
Distribution: W.Malaysia, Philippines (Luzon, Leyte)
Eupodes riedli Schuster, 1965
Type locality: Possession I.
Distribution: Possession I., (Crozet Isls, Antarctica)
Eupodes sigmoidensis Strandtmann & Goff, 1978: 123; Goff, 1987: 31; Gupta et al., 2004: 44 Darbemamieh, Hajiqanbar & Khanjani, 2013: 454
Type locality: Hawaii I., east slope of Mauna Loa
Distribution: Hawaii, India, Iran, Ivory Coast
Eupodes signatus C.L. Koch, 1835: fasc. 1, t.19
Type locality: Pleistein
Distribution: Deutschland
Eupodes skiaaki Thor, 1934: 213
Type locality: Linseim, Grjotli, Videsaeter in Skiaak (Gudbrandsdalen)
Distribution: Norway
Eupodes strandtmanni Coineau, 1976: 56
Type locality: Banyuls-sur-Mer
Distribution: France
Eupodes striola C.L. Koch, 1836: fa.1, t.18
Type locality: Germany
Distribution: Germany
Eupodes temperatus Shiba, 1978: 135; Abou-Awad, 1984: 335
Type locality: Yuyama, Matsuyama
Distribution: Egypt, Japan
E. tottanfjella Strandtmann, 1967: 54
Type locality: Queen Maud Land
Distribution: Antarctica
Eupodes trifasciatus C.L. Koch, 1838: fa.19, t.15
Type locality: Regensburg
Distribution: Germany
Eupodes unifasciatus C.L. Koch, 1838: fa.19, t. 11; 1841: 436; 1842: fa.3, p.69; Walckenaer, 1847: 537; Oudemans, 1937: 897; Thor & Willmann, 1941: 15; Turk & Turk, 1952: 497
Type locality: Regensburg (Germany)
Distribution: England, Germany,
Eupodes vallombrosae Thor, 1934: 212; Mihelčič, 1958b: 269; Schweizer & Bader, 1963: 217
Type locality: “Vallombrosa, Firenze”, Italy
Distribution: Italy, Schwitzerland, Spain
Eupodes variegatus C.L. Koch, 1838: fa.19, t.2; 1841: v.3, 435; 1842: fa.3, p. 69; Walckenaer, 1847: 537; Thor, 1912: 236; Halbert, 1915: 110; 1920: 139 (var. halophilus); Hull, 1918: 34; Sellnick, 1940: 32; Haarløv, 1942; Thor & Willmann, 1941: 20; Macfadyen, 1954: 283; Bottazzi, 1950: 37; Mihelčič, 1958a: 104; 1958b: 269; Sellnick, 1958: 38; Meyer & Ryke, 1960: 487; Lipkin & Safriel, 1971: 7 (var. halophila)
Type locality: Regensburg (Germany)
Distribution: Austria, England, Germany, Greenland, Ireland, Island, Israel, Italy, Norway, South Africa, Spain, Sweden, Yan Mayen I.
Eupodes versicolor C.L. Koch, 1838: fa.19, t.13
Type locality: Regensburg
Distribution: Germany
Eupodes viridis Oudemans, 1906: 40
Type locality: Greenland
Distribution: Greenland
Eupodes voxencollinus Thor, 1934: 208; Weis-Fogh, 1948; Evans, 1952: 666; Hallas, 1978: 31; Strandtmann & Goff, 1978: 126; Abou-Awad, 1984: 333; Goff, 1987: 31; Darbemamieh, Hajiqanbar & Khanjani, 2013: 454; Hagstrum D. et al. 2013: 546
Eupodes alaskanensis Strandtmann, 1971: 79; Strandtmann & Prasse, 1977: 4
Eupodes acuminatus Willmann, 1952 – Wangerooge Is.
Type locality: “Voksenkollen bei Oslo” (Norway)
Distribution: Alaska, Canada, China, Denmark, Egypt, England, Germany Hawaii, Iran, Norway
Eupodes winsnesi Strandtmann & Sømme, 1977: 140
Type locality: Queen Maud Land, Brattskarvet, Sverdrupfjella
Distribution: Antarctica
Eupodes wisei Womersley & Strandtmann, 1963: 452
Type locality: Hallett Station
Distribution: Antarctica
Eupodes zaheri Abou-Awad, El-Sawaf & Abdel-Khalek, 2006: 44
Type locality: Egypt
Distribution: Egypt
Filieupodes Jesionowska, 2010: 641
Type species: Filieupodes filiformis Jesionowska, 2010
Cocceupodes aegyptiacus Abou-Awad & El-Bagouri, 1984: 68
Filieupodes aegyptiacus (Abou-Awad et El-Bagouri): Jesionowska, 2010: 642
Type locality: Egypt (on Vitis vinifera)
Distribution: Egypt
Filieupodes filiformis Jesionowska, 2010: 645
Type locality: The Bialowieza Forest. Hajnówka (Podlaskie District)
Distribution: Poland
Filieupodes filistellatus Jesionowska, 2010: 651
Type locality: Ciechocinek near Toruń (Kujawsko-Pomorskie Distr.)
Distribution: Poland
Cocceupodes fusiformis Olivier & Theron, 2003: 165
Filieupodes fusiformis (Olivier et Theron): Jesionowska, 2010: 652
Type locality: S.Afr., Northern Province, near Nylstroom
Distribution: South Africa
Cocceupodes paradoxus Weis-Fogh, 1948: 263; Evans, 1952: 666; Haarløv, 1957: 27
Filieupodes paradoxus (Weis-Fogh): Jesionowska, 2010: 652
Type locality: Denmark
Distribution: England, Denmark, Greenland
Cocceupodes sharkiensis Abou-Awad, El-Sawaf & Abdel-Khalek, 2006: 48
Filieupodes sharkiensis : Jesionowska, 2010: 642
Type locality: Sharki
Distribution: Egypt
Cocceupodes shepardi Strandtmann, 1971: 88
Filieupodes shepardi (Strandtmann): Jesionowska, 2010: 642
Type locality: Cape Beaufort (Alaska)
Distribution: USA (Alaska)
Cocceupodes strandtmanni Abou-Awad & El-Bagouri, 1986: 68
Filieupodes strandtmanni (Abou-Awad et El-Bagouri): Jesionowska, 2010: 642
Type locality: Egypt
Distribution: Egypt
Cocceupodes trisetatus Strandtmann & Prasse, 1977: 7; Strandtmann & Goff, 1978: 130; Goff, 1987: 31
Filieupodes trisetatus (Strandtmann et Prasse): Jesionowska, 2010: 641
Type locality: Etzsdorf/ Saalkreis (East Germany)
Distribution: Germany, Hawaii
Linopodes C.L. Koch, 1836: fa. 1, t. 16
Type species: Linopodes ravus C.L. Koch, 1836
Linopodes antennaepes Banks, 1894: 221; Miller, 1925: 93; Darbemamieh, Hajiqanbar & Khanjani, 2013: 454
Type locality: USA
Distribution: Australia, Canada, Great Britain, Iran, Italy, USA
Linopodes barnufi Abou - Awad Badawi, El-Sawaf & Abdel-Khalek, 2006: 50
Type locality: Egypt
Distribution: Egypt
Linopodes cameronensis Shiba, 1976: 96; Darbemamieh, Hajiqanbar & Khanjani, 2013: 454
Type locality: Malaysia
Distribution: Iran, Malaysia
Linopodes eupodoides R. Canestrini, 1886: 701; Schweizer & Bader, 1963: 219
Type locality: Italy, Trentino
Distribution: Italy, Switzerland
Linopodes pubescens iwatensis Morikawa, 1963: 15
Linopodes iwatensis : Shiba, 1969c: 69
Type locality: Uchimagi, Yamagata-mura, Iwate Pref.
Distribution: Japan
Type locality: Germany
Distribution: Germany
Acarus motatorius Linnaeus, 1758: 616
Gamasus motatorius : Fabricius, 1805: 360
Linopodes motatorius (Linnaeus): Canestrini, 1886: 699; Oudemans, 1897: 112; 1937: 916; Willmann, 1951a: 146; 1954: 234; Bottazzi, 1956: 38; Haarløv, 1957: 30; Cooreman, 1959: 17; Schmölzer, 1962: 283; Schweizer & Bader, 1963: 219 (full synonymy until 1938 in Thor & Willmann, 1941).
Type locality: Europe
Distribution: Austria, Bulgaria, Belgium, Croatia, Czech Rep., Denmark, Germany, Great Britain, Hungary, Iceland, Italy, Norway, Spain, Switzerland
Linopodes motatorius africanus Meyer & Ryke, 1960: 489
Type locality: Bathurst
Distribution: South Africa
Linopodes obsoletus C.L. Koch, 1838: fasc. 18, t.18
Type locality: Regensburg
Distribution: Germany
Linopodes pubescens Morikawa, 1963: 13
Type locality: Shiroyama, Matsuyama City, Ehime Pref.
Distribution: Japan
Linopodes ravus C.L. Koch, 1836: fa. 1, t. 17
Type locality: Regensburg
Distribution: Germany
Neoprotereunetes Fain & Camerik, 1994: 153
Type species: Ereynetes lapidarius Oudemans, 1906
Ereynetes lapidarius Oudemans, 1906: 40
Protereunetes lapidarius : Bujtendijk, 1945: 319; Thor & Willmann, 1941: 58
Neoprotereunetes lapidarius (Oudemans): Fain & Camerik, 1994: 153
Type locality: Arnhem
Distribution: Holland
Niveupodes Barilo, 1991: 140
Type species: Niveupodes niveus Barilo, 1991
Niveupodes niveus Barilo, 1991: 140; Khaustov, 2016: 258
Type locality: Zarafshan Range, 20 km south of Samarkand
Distribution: Uzbekistan
Pseudeupodes Khaustov, 2014: 13
Type species: Pseudeupodes porosus Khaustov, 2014
Pseudeupodes porosus Khaustov, 2014: 13
Type locality: Yalta mountain-forest Nature Reserve
Distribution: Crimea
Pseudopenthaleus Khaustov, 2015: 931
Type species: Pseudopenthaleus tauricus Khaustov, 2015
Penthaleus ciliatus Shiba, 1978: 147
Pseudopenthaleus ciliatus (Shiba): Khaustov, 2015: 927
Type locality: Mt. Sagaramine, Shigenobu-sho, Ehime
Distribution: Japan
Pseudopenthaleus tauricus Khaustov, 2015: 931
Type locality: Crimea
Distribution: Crimea
Xerophiles Jesionowska, 2003: 12
Type species: Xerophiles ereynetoidalis Jesionowska, 2003
Xerophiles ereynetoidalis Jesionowska, 2003: 12
Type locality: Szczecin (Poland)
Distribution: Poland, Slovakia
Among the species described by Müller, 1776, Schrank, 1776, Hermann, 1804, Dugès, 1834, C.L. Koch, 1838, Wankel, 1861, Karsch, 1881, Banks, 1894, 1897, Berlese, 1917, but with unclear identity, are Eupodes cerinus Koch, E. celerrimus Koch, E. variabilis Banks, E. formosulus Koch, E. dugesi Sig Thor (as nom. nov.), E. dilectus Koch, E. cornifer (Müller) (Acarus c.), E. cinctus Koch, E. chloromelas Koch, E. celer (Dugès) (Megamerus c.), E. lineatus Koch, E. lineola Koch, E. fasciola Koch, Е. ochlochlorus Koch, E. saltatorius Schrank (Acarus s.), E. macropus Koch, E. melanurus Koch, E. pallescens Koch, Linopodes ambustus Koch, L. decoloratus Koch, L. decoloreus Koch L. longipes , L. macropus Herm. (Trombidium m.), L. gracilipes Karsch, L. flexuosus Koch, L. flavipes Koch, L. riparius Koch, L. subterraneus Wankel, L. mammuthia Banks (Eupodes m.), L. maculatus Koch, L. melaleucus Koch, L. rubiginosus Koch, L. agitatorius Berlese.
EUROPE
Austria (Irk, 1939; Willmann, 1951; Mihelčič, 1957, 1958; Schmölzer, 1962, 1993) – Cocceupodes mollicellus, Eupodes pseudoclavifrons, E. variegatus, Linopodes motatorius
Belgium (Cooreman, 1943) – Claveupodes sp., Linopodes motatorius
Bulgaria (Beron, 2011) – Linopodes motatorius
Croatia (Matočec, 2002) - Linopodes motatorius
Czech Rep. (Willmann, 1954) – Eupodes sp., Linopodes motatorius, [? L. subterraneus Wankel]
Denmark (Weis-Fogh, 1948; Haarløv, 1957; Hallas, 1978) – Cocceupodes mollicellus, Eupodes berlesei, E. viridis, E. voxencollinus, Filieupodes paradoxus
Greenland (Haarløv, 1942; Makarova, 2015) – Cocceupodes breweri, C. mollicellus (= Eupodes clavifrons R. Canestrini, 1886 = Cocceupodes curticlava Thor, 1934), C. paradoxus, C. cf. stellatus
Estonia (Thor & Willmann, 1941) – Protereunethes striatellus
France (contin.)(Coineau, 1976) – Eupodes strandtmanni
Germany (C.L. Koch, 1835, 1838; Willmann, 1952; Strandtmann & Prasse, 1977) – Benoinyssus ereynetoides, Claveupodes delicatus, Cocceupodes stellatus, Eupodes leucomelas, E. ocellatus, E. striola, E. signatus, E. trifasciatus, E. voxencollinus, Filieupodes trisetatus, Linopodes motatorius, L. ravus, Protereunethes striatellus
Great Britain (Hull, 1918; Evans, 1952; Turk & Turk, 1952; A. Baker, 1987) – Caleupodes reticulatus,Claveupodes sp., Eupodes clavifrons, E. unifasciatus, E. variegatus, E. voxencollinus, Filieupodes paradoxus, Linopodes antennaepes, L. motatorius
Holland (Oudemans, 1906; Thor & Willmann, 1941) - Eupodes milvinus, E. viridis, Protereunethes striatellus
Hungary (Dudich et al., 1940) – Linopodes motatorius
Iceland (Sellnick, 1940) – Eupodes variegatus, Linopodes motatorius
Surtsey (Lindroth, Andersson, Bödvarsson & Richter, 1973) – Cocceupodes clavifrons, Eupodes hiemalis, E. variegatus, Linopodes motatorius
Ireland (Halbert, 1915, 1920) – Eupodes variegatus var. halophilus
Italy (contin.)(Canestrini, 1886; Thor, 1934, Thor & Willmann, 1941; Bottazzi, 1950) - Cocceupodes mollicellus (= clavifrons), Eupodes fusifer, E. iconicus, E. milvinus, E. vallombrosae, E. variegatus, Linopodes antennaepes, L. eupodoides, L. motatorius
Sicily (Berlese, 1923) – Protereunetes agilis
Norway (Thor, 1912, 1934a, 1934c; Thor & Willmann, 1941) – Cocceupodes mollicellus (=curviclava), Eupodes pseudoclavifrons, E. decoloratus, E. hjartdaliae, E. skiaaki, E. variegatus, E. voxencollinus, Linopodes motatorius
Svalbard (Spitzbergen)(Oudemans, 1926; Thor, 1934b, 1934c) – Eupodes clavifrons, Protereunetes boerneri
Poland (Jesionowska, 2005, 2006, 2007a, 2007b, 2008, 2010; Szudarek-Trepto, Kaźmierski, Dabert et al., 2020) – Claveupodes sp., Cocceupodes longisolenidiatus, Filieupodes filiformis, F. filistellatus, Eupodes voxencollinus, Xerophiles ereynetoidalis
Romania (Ivan et al., 2016) – Eupodes sp.
Russia (cont.)
Crimea (Khaustov, 2014, 2015, 2017) – Echinoeupodes echinus, Pseudeupodes porosus, Pseudopenthaleus tauricus
Slovakia (Jesionowska, 2003) - Xerophiles ereynetoidalis
Spain (cont.)(Mihelčič, 1958b; Santamaría et al., 2012) - Cocceupodes sp., Benoinyssus ereynetoides, E. valombrosae, E. variegatus, Linopodes motatorius
Sweden (Thor & Willmann, 1941; Sellnick, 1958) - Eupodes milvinus, E. variegatus
Switzerland (Haller, 1880; Schweizer, 1922; Cooreman, 1954,1959, Schweizer & Bader, 1963) – Eupodes berlesei, E. haltica, E. vallombrosae, Linopodes eupodoides, L. motatorius
AFRICA
Angola (Meyer & Ryke, 1960; Olivier & Theron, 1997a, 1998) – Benoinyssus celatus, B. ornatus, Eupodes parafusifer
Botswana (Olivier & Theron, 1997a; 1998) - Benoinyssus serratus, Eupodes minipulus
Cabo Verde (Olivier & Theron, 1998) - Eupodes minipulus
Egypt (Trägårdh, 1904; Abou-Awad, 1984: 333; Abou-Awad & El-Bagouri, 1984, 1986; Abou-Awad, El-Sawaf & Abdel-Khalek, 2006) – Benoinyssus momeni, Cocceupodes sharkiensis, Filieupodes aegyptiacus, F. strandtmanni, Eupodes bakeri, E. momeni, E. niloticus, E. strandtmanni, E. temperatus, E. zaheri, Linopodes barnufi, L.curtipes
Ivory Coast - Eupodes sigmoidensis
Namibia (Olivier & Theron, 1997a) - Benoinyssus serratus
Rwanda (Fain, 1958, 1959) – Benoinyssus najae
South Africa (Thor, 1934c; Meyer & Ryke, 1960; Olivier & Theron, 1997a, 1997b, 1998, 2003) – Benoinyssus falcatus, B. ornatus, B. pusillus, B. serratus, C. gracilongus, Eupodes acanthus, E. fusiferellus, E. hamatus, E. indentatus, E. lutatus, E. minipilus, E. parafusifer, E. variegatus, Filieupodes fusiformis
Swaziland (Olivier & Theron, 1997a) – Benoinyssus ornatus, B. serratus
Zimbabwe (Olivier & Theron, 1998) - Eupodes minipulus
“East Africa” (Berlese, 1916) – “Linopodes agitatorius”
ASIA
China (Hagstrum et al., 2013) – Eupodes voxencollinus
India (Ghosh & Gupta, 2003; Gupta et al., 2004, Indranil et al., 2011) – Eupodes sigmoidensis
Indonesia (Corpuz-Raros, 2005) – Eupodes pusillus [Java]
Iran (Darbemamieh, Hajiqanbar & Khanjani, 2013) - Benoinyssus sp., Claveupodes sp., Cocceupodes sp., Eupodes crozetensis, E. sigmoidensis, E. voxencollinus, Linopodes antennaepes, L. cameronensis
Israel and Palestina (Lipkin & Safriel, 1971) – Eupodes variegatus var. halophila
Japan (Karsch, 1881; Morikawa, 1963; Shiba, 1969a, 1969c, 1971, 1978) – Cocceupodes communis, Echinoeupodes turgidus, Eupodes cordiformis, E. infirmus, E. longisetatus, E. okinoshimaensis, Linopodes iwatensis, L. pubescens, Protereunetes perforates, P. villosus, Pseudopenthaleus ciliatus
Malaysia (Shiba, 1976) – Eupodes berensis, E. lumbarus, E. onoi, E. pasohensis, E. pusillus, E. pseudolumbarus, Linopodes cameronensis
Philippines (Corpuz-Raros, 2005) – Eupodes callejae, E. garciai, E. onoi, E. pusillus, E. pseudolumbarus
Singapore (Corpuz-Raros, 2005) – Eupodes onoi
United Arab Emirates (Negm, 2014) – Eupodes sp.
Uzbekistan (Barilo, 1991; Khaustov, 2016) – Niveupodes nivalis
Yemen (Ueckermann et al., 2006) – Eupodes sp.
NORTH AMERICA
Canada (Miller, 1925; Behan, 1978; Moseley, 2007; Beaulieu et al., 2019 – 13 sp.) - Eupodes voxencollinus, Linopodes antennaepes
Mexico (Baker, 1945) - Benoinyssus tuberculatus
USA (Banks, 1894; 1896; 1897, 1907; 1916; Miller, 1925; Thor, 1934) – Eupodes brevipes, E. marinus, Linopodes antennaepes
Alaska (Strandtmann, 1971) – Cocceupodes breweri, Eupodes alaskanensis, E. voxencollinus, Filieupodes shepardi, Protereunetes boerneri
Hawaii (Strandtmann & Goff, 1978) - Eupodes hawaiiensis, E. sigmoidensis, E. voxencollinus
SOUTH and CENTRAL AMERICA
Argentina (Bedano, 2004) – 1 sp.
Brazil (Bizarro et al., 2020) – Benoinyssus oconnori
Costa Rica (Baker & Lindquist, 2002) – Aethosolenia laselvensis
OCEANIA
Australia (continent)(Clancy, 1981) - Linopodes sp.
New Zealand (Spain & Luxton, 1971; Martin, 1983; Luxton, 1990) – Claveupodes delicatulus, Linopodes spp., Eupodes minutus,
Campbell Island (Strandtmann, 1964) - Eupodes longisetatus
Macquarie Island (Womersley, 1937; Luxton, 1990) – Eupodes minutus
ANTARCTICA
Antarctic Continent (Strandtmann & Womersley, 1963; Strandtmann, 1967; Strandtmann & Davies, 1972; Gless, 1972; Strandtmann & Sømme, 1977; Marshall & Pugh, 1996; Llano & Gless, 2013) – Eupodes angardi, E. winsnesi, E. wisei
South Georgia (Strandtmann, 1970) – Eupodes minutus
South-Orkney-Islands (Booth, Edwards & Usher, 1985) – Eupodes exiguus, E. parvus
South Shetland Islands (Strandtmann & Tilbrook, 1968) – Cocceupodes australis
SUB-ANTARCTIC
Crozet Islands (Strandtmann & Davies, 1972) – Eupodes crozetensis, E. riedli, Possession I.
Dendrochaetidae Olivier, 2008: 16
The family contains so far only one genus and species (Dendrochaetus acarus Olivier) from South Africa. These mites live in the soil.
Dendrochaetidae Olivier, 2008: 16
Type genus: Dendrochaetus Olivier, 2009 [repl. name for Dendrodus Olivier, 2008, praeocc.]
Dendrodus Olivier, 2008: 19 (praeocc.)
Dendrochaetus Olivier, 2009: 68
Type species: Dendrochaetus acarus Olivier, 2009 [for Dendrodus acarus Olivier, 2008]
Dendrodus acarus Olivier, 2008: 19
Dendrochaetus acarus (Olivier): Olivier, 2009: 68
Type locality: South Africa
Distribution: South Africa
AFRICA
South Africa (Olivier, 2008, 2009) – Dendrochaetus [Dendrodus] acarus
Rhagidiidae Oudemans, 1922: 83
The Rhagidiids are soft bodied, predaceous soil mites. Thorell (1872) described the genus Rhagidia and Cambridge (1876) described from the far away Kerguelen Island another genus – Poecilophysis. Described by Oudemans (1922), the family Rhagidiidae has been studied until 1980 by several researchers (C.L. Koch, Canestrini, Thorell, Berlese, Banks, Hull, Sig Thor, Trägårdh, Willmann, Elliot, Strandtmann, Ehrnstberger, Abou-Awad and El-Bagoury, Womersley, Morikawa, Shiba, Baltac, Halliday, Khaustov, and others). The first review of the family (Thor & Willmann, 1941) contained three genera, 37 certain and 8 uncertain species and five “varieties”. R.W. Strandtmann tried to distinguish some more reliable diagnostic characters, but only the world revision of the family (Zacharda, 1980) achieved the modern system of the family. From the 20 genera present in the monograph Zacharda described 16 as new and from 74 species 35 were described as new.
In the 40 years elapsed since this monograph have been described the new genera Krantzia Zaharda, 1983, Neothoria Abou-Awad et El-Bagoury, 1986 Lindquistula Zacharda, 1986, Kouchibouguacia Zacharda, 1986, Arhagidia Lindquist et Zacharda, 1987, Dongnaia Zacharda, 1994, Janes Jesionowska, 2002, Paracrassocheles Khaustov, 2015 and several new subgenera and species. Now (September 2021) the family Rhagidiidae comprises 28 genera and 150 species of whitish mites, living in the soil and often in caves.
Family Rhagidiidae Oudemans, 1922 - 28 gen., 150 sp.
Arctorhagidia Zacharda, 1980 – 2 sp.
Arhagidia Lindquist et Zacharda, 1987 – 1 sp.
Brevipalpia Zacharda, 1980 - 1 sp.
Coccorhagidia Sig Thor, 1934 – 8 sp.
Crassocheles Zacharda, 1980 - 2 sp.
Elliotta Zacharda, 1980 - 1 sp.
Eskimaia Zacharda, 1980 – 1 sp.
Evadorhagidia Zacharda, 1980 - 4 sp.
Flabellorhagidia Elliott, 1976 - 1 sp.
Foveacheles Zacharda, 1980 - 33 sp.
Foveacheles Zacharda, 1980 - 11 sp.
Hirschmannetta Zacharda, 1980 - 1 sp.
Mediostella Zacharda, 1980 - 5 sp.
Propriorhagidia Zacharda, 1980 - 2 sp.
Proxistella Zacharda, 1980 - 2 sp.
Spelaeocheles Zacharda, 1988 – 1 sp.
Ternirhagidia Zacharda, 1980 - 1 sp.
Trofocheles Zacharda, 1982 - 3 sp.
Usitorhagidia Zacharda, 1980 – 7 sp.
Hammenia Zacharda, 1980 - 1 sp.
Janes Jesionowska, 2002 – 1 sp.
Kouchibouguacia Zacharda, 1986 – 2 sp.
Latoempodia Zacharda, 1980 - 2 sp.
Lindquistula Zacharda, 1986 – 1 sp.
Neothoria Abou-Awad et El-Bagoury, 1986 – 2 sp.
Paracrassocheles Khaustov, 2015 – 1 sp.
Parallelorhagidia Zacharda, 1980 - 3 sp.
Poecilophysis Cambridge, 1876 - 16 sp.
Dentocheles Zacharda, 1980 - 5 sp.
Poecilophysis Cambridge, 1876 - 3 sp.
Procerocheles Zacharda, 1980 - 5 sp.
Soprocheles Zacharda, 1980 - 2 sp.
Wankelia Zacharda, 1980 - 1 sp.
Rhagidia Thorell, 1872 - 28 sp.
Austrorhagidia Zacharda, 1980 – 3 sp.
Deharvengiella Zacharda, 1987 – 3 sp.
Zachardaia Özdikmen, 2008 (= Krantzia Zacharda, 1983) - 1 sp.
Dongnaia Zacharda, 1994 – 1 sp.
Noerneria Canestrini, 1886 – 14 sp.
Rhagidia Thorell, 1872 - 7 sp.
Robustocheles Zacharda, 1980 - 19 sp.
Amoveocheles Zacharda, 1980 - 3 sp.
Lewia Zacharda, 1980 - 5 sp.
Mexia Zacharda, 1997 - 3 sp.
Robustocheles Zacharda, 1980 - 8 sp.
Shibaia Zacharda, 1980 - 6 sp.
Thoria Zacharda, 1980 - 2 sp.
Traegaardhia Zacharda, 1980 - 10 sp.
Troglocheles Zacharda, 1980 - 10 sp.
Tuberostoma Zacharda, 1980 – 3 sp.
Rhagidiidae Oudemans, 1922: 83
Type genus: Rhagidia Thorell, 1872
Arctorhagidia Zacharda, 1980: 719
Type species: Rhagidia mariehammerae Strandtmann, 1971
Rhagidia mariehammerae Strandtmann, 1971: 113
Arctorhagidia mariehammerae (Strandtmann): Zacharda, 1980: 719
Type locality: Alaska, Cape Beaufort
Distribution: USA (Alaska)
Coccorhagidia sateri Strandtmann, 1971: 95
Arctorhagidia sateri (Strandtmann): Zacharda, 1980: 721
Type locality: Alaska, Barrow
Distribution: USA (Alaska)
Arhagidia Lindquist et Zacharda, 1987: 2149
Type species: Arhagidia monothrix Zacharda, 1987
Arhagidia monothrix Lindquist & Zacharda, 1987: 2156
Type locality: Jornada ecological research site of the Chihuahuan Desert in southern New Mexico
Distribution: USA (New Mexico)
Brevipalpia Zacharda, 1980: 712
Type species: Brevipalpia minima Zacharda, 1980
Brevipalpia minima Zacharda, 1980: 712; 2012; Zacharda & Daniel, 1988: 58; Darbemamieh, Hajiqanbar & Khanjani, 2013: 455
Type locality: Bohemia centr., Závist-Knoll near Zbraslav
Distribution: Czech Republic, Iran, Nepal
Coccorhagidia Sig Thor, 1934: 318
Type species: Nörneria clavifrous Canestrini, 1886
Nörneria clavifrons Canestrini, 1886: 710
Rhagidia clavifrons : Berlese, 1921: 90; Bedano, 2004
?Rhagidia mordax: Willmann, 1936: 296
Coccorhagidia clavifrons : Sig Thor, 1934: 318; Dudich et al., 1940: 54; Thor & Willmann, 1941: 128; Willmann, 1951b: 143; Turk & Turk, 1952: 496; Macfadyen, 1954: 284; Haarløv, 1957: 27; Meyer & Ryke, 1960: 493; Shiba, 1969: 81; Strandtmann & Prasse, 1976: 13; Strandtmann & Goff, 1978: 143; Zacharda, 1980: 704; 2013; Abou-Awad, 1985: 258; Goff, 1987: 34; Qin, 1998: 300; Darbemamieh, Hajiqanbar & Khanjani, 2013: 455
Type locality: Doss (Tavon, Trento), Italy
Distribution: Argentina, Austria, Czech Republic, Denmark, Egypt, England, Germany, Hungary, Iran, Italy, Japan, Slovakia; Canada, Hawaii, South Africa, New Zealand, Yan Mayen I.
Coccorhagidia cubana Baltac, 1977a: 197
Type locality: Cueva de México, Province de Camagüey
Distribution: Cuba
Coccorhagidia montana Bannova, Sarmanova & Zhuravleva, 1977: 59
Type locality: Romania
Distribution: Romania
Coccorhagidia divergens Mihelčič, 1957: 103
Type locality: Ötztaler Alpen: Hochfirst, Nivale Polsterpflanzen aus der O-Wand, ca. 3400 m.
Distribution: Austria
Coccorhagidia pittardi Strandtmann, 1971: 98; Zacharda, 1980: 708; 2012: ; Zacharda & Daniel, 1988: 58; Darbemamieh, Hajiqanbar & Khanjani, 2013: 455
Type locality: Alaska, Wainwright
Distribution: Slovakia, Tatra (2200-2367 m); Alaska, Iran, Nepal (3900 m),
Coccorhagidia semiclavifrons Shiba, 1969: 83
Type locality: Shiga Heigths, IBP Area
Distribution: Japan
Rhagidia subterranea Berlese, 1905: 232
Coccorhagidia berlesei Thor & Willmann, 1941: 129 (as nom. nov. pro Scyphius subterraneus nom. nud. Absolon, 1899)
Coccorhagidia subterranea (Berlese): Evans, 1952: 666; Haarløv, 1957: 27
Type locality: Florence (Italy)
Distribution: Denmark, England, Italy
Crassocheles Zacharda, 1980: 550
Type species: Crassoheles muralis Zacharda, 1980
Crassocheles muralis Zacharda, 1980: 551
Type locality: Velký Berdĕz-Hill, Czech Republic
Distribution: Czech Republic
Crassocheles virgo Zacharda, 1980: 554
Type locality: Provodin, Provodingké kameny, Czech Republic
Distribution: Czech Republic
Elliotta Zacharda, 1980: 597
Type species: Flabellorhagidia howarthi Elliott, 1976
Flabellorhagidia howarthi Elliott, 1976: 5-10
Elliotta howarthi : Zacharda, 1980: 597
Type locality: Klickitat Co., Washington, Cheese Cave, 2 km SW Trout Lake, USA
Distribution: USA (Washington, Hawaii, Idaho)
Eskimaia Zacharda, 1980: 723
Type species: Coccorhagidia capitata Strandtmann, 1971
Coccorhagidia capitata Strandtmann, 1971: 98
Eskimaia capitata (Strandtmann): Zacharda, 1980: 723
Type locality: Alaska, Anaktuvik Pass, Brooks Range
Distribution: USA (Alaska)
Evadorhagidia Zacharda, 1980: 725
Type species: Evadorhagidia oblikensis Zacharda, 1980
Evadorhagidia bezdezensis Zacharda, 1980: 734; Ružička, Hajer & Zacharda, 1995: 47
Type locality: Bohemia bor., Malý Bezdĕz-Hill
Distribution: Czech Republic
Evadorhagidia oblikensis corcontica Zacharda, 1993: 57
Evadorhagidia corcontica Zacharda:
Type locality: Krkonose Mts.
Distribution: Czech Republic
Rhagidia janetscheki Willmann, 1953: 218; Strandtmann, 1971: 103
Evadorhagidia janetscheki : Zacharda, 1980: 733
Type locality: Tyrol, the Zillertall Alps, Rossrücken, 3000 m
Distribution: Austria
Evadorhagidia oblikensis Zacharda, 1980: 726
Type locality: Bohemia bor., Oblik-Hill near Louny
Distribution: Czech Republic
Evadorhagidia quinqueseta Zacharda, 1980: 731
Type locality: Canada, N.W. Territory, Atkinson Point
Distribution: Canada (N.W. Territory)
Flabellorhagidia Elliott, 1976: 1
Type species: Flabellorhagidia pecki Elliott, 1976
Flabellorhagidia pecki Elliott, 1976: 2; Zacharda, 1980: 657
Type locality: Butte Co., Idaho, Boy Scout Cave (lava)
Distribution:USA (Idaho)
Foveacheles Zacharda, 1980: 661
Type species: Rhagidia osloensis Sig Thor, 1934
Achaetorhagidia Zacharda, 1994: 36
Type species: Foveacheles (Achaetorhagidia) tropica Zacharda, 1994
Foveacheles (Achaetorhagidia) namcatiensis Zacharda, 1994: 38
Type locality: Nam Cat Tien National Park, South Vietnam
Distribution: Vietnam
Foveacheles (Achaetorhagidia) tropica Zacharda, 1994: 44
Type locality: Nam Cat Tien National Park, South Vietnam
Distribution: Vietnam
Foveacheles Zacharda, 1980: 661
Type species: Rhagidia osloensis Sig Thor, 1934
Foveacheles (Foveacheles) auricularia Zacharda & Elliott, 1985: 468
Type locality: California
Distribution: USA
Foveacheles (Foveacheles) brevichelae Zacharda, 1980: 666; 2012: ; Darbemamieh, Hajiqanbar & Khanjani, 2013: 455; Zacharda & Kučera, 2010: 5
Type locality: Bohemia centr., Karlštejn
Distribution: Austria, Canada, Czech Republic, Finland, Iran, Russia
Foveacheles (Foveacheles) cannadasi Barranco & Amate, 2000: 15
Type locality: Cueva Larga, Sierra de los Filabres, Almeria
Distribution: Spain
Foveacheles (Foveacheles) caucasica Zacharda, 1983b: 305; 1993: 60
Type locality: Kabardino-Balkaria, Poljana Azau, 2 km W of Terskol, N-slope of Ceget Mt., 2600 m
Distribution: Czech Republic, Russia (Caucasus)
Foveacheles (Foveacheles) cegetensis Zacharda, 1983b: 307; 1993: 60; Zacharda & Kučera, 2010: 5; Darbemamieh, Hajiqanbar & Khanjani, 2013: 455
Type locality: Kabardino-Balkaria, Poljana Azau, 2 km W of Terskol, N-slope of Ceget Mt., 2600 – 2800 m
Distribution: Czech Republic, Finland, Iran, Russia (Caucasus)
Foveacheles (Foveacheles) gigantea Zacharda, 2000: 248
Type locality: Ceske Stredohori Mts,
Distribution: Czech Rep.
Foveacheles (Foveacheles) halltalensis Zacharda, 2000: 253
Type locality: Northern Alps,
Distribution: Austria
Foveacheles (Foveacheles) incognita Zacharda, 1980: 670
Type locality: Patria incognita
Distribution: Patria incognita
Foveacheles (Foveacheles) lemoni Abou-Awad, El-Sawaf & Abdel-Khalek, 2009: 147
Type locality: El - Kassasin, El – Sharkiya
Distribution: Egypt
Rhagidia osloensis Sig Thor, 1934: 314; Thor & Willmann, 1941: 120
Foveacheles (Foveacheles) osloensis: Zacharda, 1980: 662; Hippa et al., 1989: 61; Darbemamieh, Hajiqanbar & Khanjani, 2013: 455
Type locality: Norway
Distribution: Austria, Czech Republic, Germany, Iran, Norway, Sweden
Foveacheles (Foveacheles) proxima Zacharda, 2000: 258
Type locality: Northern Alps
Distribution: Austria
Foveacheles (Foveacheles) thaleri Zacharda, 2010: 58
Type locality: Cedar Valley, Troodos
Distribution: Cyprus
Hirschmannetta Zacharda, 1980: 694
Type species: Foveacheles (Hirschmannetta) magna Zacharda, 1980
Foveacheles (Hirschmannetta) magna Zacharda, 1980: 695
Type locality: Harz, Bielshöhle Cave
Distribution: Germany
Mediostella Zacharda, 1980: 672
Type species: Foveacheles (Mediostella) rupestris Zacharda, 1980
Noerneria canestrini Berlese & Trouessart, 1889: 142
Rhagidia gigas var. canestrini: Thor & Willmann, 1941: 112
?Rhagidia fragosa Schuster, 1958: 95
Foveacheles (Mediostella) canestrini: Zacharda, 1980: 679
Type locality: Plage du Croisic (Loire Tufe)
Distribution: France
Foveacheles (Mediostella) pribilovi Zacharda, 1996: 344
Type locality: Alaska, Pribilof Islands, St. Paul’s Island
Distribution: USA (Alaska)
Foveacheles (Mediostella) rupestris Zacharda, 1980: 672
Type locality: the High Tatras, Koprovský štit, 2376 m
Distribution: Slovakia
Foveacheles (Mediostella) vancouverensis Zacharda, 1986: 2542
Type locality: British Columbia, Vancouver Island, SE end of Long Beach
Distribution: Canada
Foveacheles (Mediostella) willmanni Zacharda, 1980: 676
Type locality: Bösensteingipfel
Distribution: Austria
Propriorhagidia Zacharda, 1980: 690
Type species: Foveacheles (Propriorhagidia) mira Zacharda, 1980
Foveacheles (Propriorhagidia) mira Zacharda, 1980: 690
Type locality: Wien
Distribution: Austria
Foveacheles (Propriorhagidia) tuktoyaktukensis Zacharda, 1980: 692; 1985: 78
Type locality: Canada, N.W. Territory, Tuktoyaktut, Beaufort Sea
Distribution: Canada (N.W. Territory), USA
Proxistella Zacharda, 1980: 700
Type species: Scyphius terricola C. L. Koch, 1835
Foveacheles (Proxistella) simulata Zacharda, 2000: 106
Type locality: Berounka River Valley
Distribution: Czech Rep.
Scyphius terricola C. L. Koch, 1835: Fasc. 1, 15
Scyphius terricolus C. L. Koch, 1836: Heft 1, Nr 15
Rhagidia terricola : Willmann, 1936: 294; 1954: 234; Thor, 1912: 235; Halbert, 1915: 110; Vitzthum, 1925: 153; Irk, 1939: 153; Thor & Willmann, 1941: 102; 1951: 146; Macfadyen, 1854: 284; Mihelčič, 1958: 102; Cooreman, 1959: 26; Schmölzer, 1962: 282; 1993: 59; Rack, 1974: 133;
Foveacheles (Proxistella) terricola Zacharda, 1980: 701
Type locality: Germany
Distribution: Austria, Czech Rep., France, Germany, Norway, Switzerland, Yan Mayen I.
Foveacheles (Spelaeocheles) Zacharda, 1988: 75
Type species: Foveacheles (Spelaeocheles) troglodyta Zacharda, 1988
Foveacheles (Spelaeocheles) troglodyta Zacharda, 1988: 76
Type locality: East Slovakia, Stratenska vysočina
Distribution: Slovakia
Ternirhagidia Zacharda, 1980: 696
Type species: Foveacheles (Ternirhagidia) alpina Zacharda, 1980
Foveacheles (Ternirhagidia) alpina Zacharda, 1980: 697
Type locality: the Zillertall Alps, Tetterjoch, 3000 m, Schnoarzensteingipfel, 3320 m
Distribution: Austria
Foveacheles (Ternirhagidia) emendata Zacharda, 1980: 699
Type locality: Patria incognita
Distribution: Patria incognita
Trofocheles Zacharda, 1982: 275
Type species: Foveacheles (Trofocheles) tenorioae Zacharda, 1982
Foveacheles (Trofocheles) goffi Zacharda, 1982: 277
Type locality: Hawaii I., Kau Forest Es., Manu Cave, 2100 m
Distribution: USA (Hawaii)
Foveacheles (Trofocheles) paralleloseta Zacharda, 1985: 77
Type locality: Sam Six Cave, 4 mi. W Wytheville, Wythe Co., Virginia
Distribution: USA (Virginia)
Foveacheles (Trofocheles) tenorioae Zacharda, 1982: 278
Type locality: Hawaii I., Hawaii Volcanoes Nat. Park, Kipuka Puaulu, Cave No 3
Distribution: USA (Hawaii)
Usitorhagidia Zacharda, 1980: 682
Type species: Rhagidia arenaria Willmann, 1952
Rhagidia arenaria Willmann, 1952: 155; Ehrnsberger, 1977: 67
Foveacheles (Usitorhagidia) arenaria Zacharda, 1980: 682
Type locality: Germany, Wangerooge Islands
Distribution: Germany (Wangerooge Islands)
Foveacheles (Usitorhagidia) bentonensis Zacharda, 1983: 1000
Type locality: Oregon
Distribution: USA (Oregon)
Rhagidia clavacrinita Ehrnsberger, 1977a: 95; 1977b: 67
Foveacheles (Usitorhagidia) clavacrinita Zacharda, 1980: 685
Type locality: Kiel, beach forest litter
Distribution: Germany
Foveacheles (Usitorhagidia) harzensis Zacharda, 1980: 688
Type locality: Harz, Einhornhöhle-Cave
Distribution: Germany
Foveacheles (Usitorhagidia) mexicana Zacharda, 1996: 350
Type locality: Mexico, 15 mi SW El Salto, Durango
Distribution: Mexico
Foveacheles (Usitorhagidia) titanica Zacharda & Elliott, 1985: 471
Type locality: California,
Distribution: USA (California)
Foveacheles (Usitorhagidia) unguiculata Zacharda, 1994: 86
Type locality: North Tyrol, the Ötztal Alps, Pitztal, Wildgrat 2974 m
Distribution: Austria
Hammenia Zacharda, 1980: 760
Type species: Hammenia macrostella Zacharda, 1980
Hammenia macrostella Zacharda, 1980: 760
Type locality: Bohemia bor., Velký Bezdĕz-Hill
Distribution: Czech Republic
Janes Jesionowska, 2002: 174
Type species: Janes rafalskii Jesionowska, 2002
Janes rafalskii Jesionowska, 2002: 174
Type locality: Poland
Distribution: Poland
Kouchibouguacia Zacharda, 1986: 2532
Type species: Kouchibouguacia cryptosolenidiata Zacharda, 1986
Kouchibouguacia cryptosolenidiata Zacharda, 1986: 2533
Type locality: New Brunswick, Kouchibouguac Nat. Park, Kollock Creek trail
Distribution: Canada
Kouchibouguacia transcendata Zacharda, 1995: 589
Type locality: Canada, Ontario, St. Lawrence Islands Nat. Park, Grenadier Island
Distribution: Canada (Alberta, Ontario, New Brunswick), USA (New Mexico)
Latoempodia Zacharda, 1980: 585
Type species: Latoempodia macroempodiata Zacharda, 1980
Latoempodia macroempodiata Zacharda, 1980: 585
Type locality: Oblik-Hill near Louny
Distribution: Czech Republic
Latoempodia similis Zacharda, 1980: 589
Type locality: the High Tatra, Mengusovská dolina Valley, 1730 m
Distribution: Slovakia
Lindquistula Zacharda, 1986: 2531
Type species: Lindquistula multisolenidiata Zacharda, 1980
Lindquistula multisolenidiata Zacharda, 1986: 2531
Type locality: Ontario, Pt. Pelee Nat. Park
Distribution: Canada
Neothoria Abou-Awad & El-Bagoury, 1986: 59
Type species: Neothoria niloticus Abou-Awad et El-Bagoury, 1986
Neothoria niloticus Abou-Awad & El-Bagoury, 1986: 59
Type locality: Lower Egypt
Distribution: Egypt
Neothoria zacharda Abu-Awad, 2010: 141
Type locality: Egypt
Distribution: Egypt
Paracrassocheles Khaustov, 2015: 619
Type species: Paracrassocheles clavisetosus Khaustov, 2015
Paracrassocheles clavisetosus Khaustov, 2015: 619
Type locality: Crimea
Distribution: Crimea
Parallelorhagidia Zacharda, 1980: 590
Type species: Coccorhagidia evansi Strandtmann et Prasse, 1976
Coccorhagidia evansi Strandtmann & Prasse, 1976: 12
Parallelorhagidia evansi (Strandtmann et Prasse): Zacharda, 1980: 591
Type locality: Etzsdorf/ Saalkreis (East Germany)
Distribution: Czech Republic, Germany
Parallelorhagidia hawaiiensis Zacharda, 1980: 594; Goff, 1987: 35
Type locality: Hawaii, Big Island
Distribution: USA (Hawaii)
Parallelorhagidia parva Zacharda, 1986: 2538
Type locality: New Brunswick, Kouchibouguac Nat. Park, Kouchibouguac Campground
Distribution: Canada
Poecilophysis Cambridge, 1876: 261
Type species: Poecilophysis kerguelenensis Cambridge, 1876
Dentocheles Zacharda, 1980: 621
Type species: Scyphius pratensis C. L. Koch, 1835
Poecilophysis (Dentocheles) affinis Zacharda, 1983: 996
Type locality: Oregon, Coffin Butte
Distribution: USA (Oregon)
Poecilophysis (Dentocheles) extraneostella Zacharda, 1985: 73
Type locality: Bowling Cave near Ben Hur, Lee Co., Virginia
Distribution: USA (Virginia)
Poecilophysis (Dentocheles) melanoseta Zacharda & Elliot, 1985: 646
Type locality: California
Distribution: USA (California)
Scyphius pratensis C. L. Koch, 1835: fasc. 1, 14
Poecilophysis pratensis : Oudemans, 1905: 202; Goff, 1987: 35; Zacharda & Kučera, 2010: 5
Rhagidia pratensis : Willmann, 1932: 5; 1936: 229; 1951b: 143; 1954: 234; Sellnick, 1940: 31; Haarløv, 1957: 31; Cooreman, 1959: 20
Rhagidia japonica Morikawa, 1963: 15; Shiba, 1969: 76
Rhagidia whartoni Strandtmann, 1971: 107
Poecilophysis (Dentocheles) pratensis Zacharda, 1980: 621; 1983b: 312; 1985: 74
Type locality: Germany
Distribution: Alaska, Austria, Azerbaidzhan, Belgium, Czech Republic, Denmark, Finland, France, Germany, Hawaii, Iceland, Iran, Japan, Russia, Slovakia, Switzerland, USA
Rhagidia reflexa (part.?): Wankel, 1936: 301; 1953: 218
Rhagidia wankeli Zacharda, 1978: 226
Poecilophysis (D.) wankeli Zacharda, 1980: 630
Type locality: Moravia
Distribution: Czech Rep., Slovakia, Austria
Bryobia ? (or Penthaleus ?) weyerensis Packard, 1888: 42
Poecilophysis (D.) weyerensis Zacharda, 1980: 627; 1985: 74; Hippa et al., 1989: 61
Rhyncholophus cavernarum Packard, 1888: 42
Type locality: Augusta County, Virginia, Weyer’s Cave, USA
Distribution: Sweden, USA
Poecilophysis Cambridge, 1876: 261; Zacharda, 1980: 601
Type species: Poecilophysis kerguelenensis Cambridge, 1876
Poecilophysis kerguelenensis Cambridge, 1876: 261
Rhagidia megalochela Trägårdh, 1907: 20
Rhagidia kerguelensis : André, 1947: 78
Rhagidia kerguelensis (part.): Strandtmann & Davies, 1972: 39
Poecilophysis (P.) kerguelenensis: Zacharda, 1980: 602
Type locality: Subantarctic region
Distribution: Possession I., Crozet I., Saint-Paul I., Kerguelen Islands,
Rhagidia macquariensis Womersley & Strandtmann, 1963: 465
Poecilophysis macquariensis Zacharda: Qin, 1998: 300
Poecilophysis (P.) macquariensis Zacharda, 1980: 606
Type locality: Macquarie Island
Distribution: Macquarie Island, Antarctica
Procerocheles Zacharda, 1980: 609
Type species: Rhagidia gelida faeroensis Trägårdh, 1931
Rhagidia gelida faeroensis Trägårdh, 1931: 42
Rhagidia reflexa Willmann, 1936: 300; Thor & Willmann, 1941: 104 (part.?); Cooreman, 1959: 22
Poecilophysis (Procerocheles) faeroensis Zacharda, 1980: 609; Hippa et al., 1989: 61
Type locality: Faroe Isl., Thorshawn (Denmark)
Distribution: Denmark (Faroe Is.), Germany, Czech Republic, Iran, Slovakia, Switzerland
Poecilophysis (Procerocheles) oregonensis Zacharda, 1983: 995
Type locality: Oregon, Washington Co.
Distribution: USA (Oregon)
Poecilophysis (Procerocheles) pseudoreflexa Zacharda, 1980: 614; 1983: 311; Hippa et al., 1989: 61; Zacharda & Kučera, 2010: 5
Type locality: Oblik-Hill near Louny (Czech Rep.)
Distribution: Canada, Czech Republic, Finland, Kirghizstan, Russia, Slovakia, Sweden
Rhagidia recussa Sig Thor, 1909: 14; Leruth, 1939: 412
Poecilophysis (Procerocheles) recussa (Sig Thor): Zacharda, 1993: 52
Type locality: Shara-Ullah Mts., North Siberia
Distribution: Belgium, Czech Republic, Russia
Scyphius spelaeus Wankel, 1861: 258
Rhagidia recussa Sig Thor, 1909: 14 ?
Rhagidia spelaea (Wankel): Willmann, in Thor & Willmann, 1941: 108; Willmann, 1954: 232
Poecilophysis (Procerocheles) spelaea: Zacharda, 1980: 617; Zacharda & Kučera, 2010: 5
Type locality of Scyphius spelaeus: Moravia
Type locality of Rhagidia recussa: Shara-Ullah-Mountains, North Siberia
Distribution: Czech Republic, England, Finland, France, Germany, Belgium, Poland, Romania, Switzerland; Siberia ?
Soprocheles Zacharda, 1980: 634
Type species: Rhagidia saxonica Willmann, 1934
Scyphius elongatus : Valle, 1955: 17, nec Scyphius elongatus C. L. Koch, 1838
Poecilophysis (Soprocheles) arena Zacharda, 1980: 639
Type locality: Moravia Mer., Valtice
Distribution: Czech Republic, Iran, Italy, Hawaii
Rhagidia saxonica Willmann, 1934: 306; 1936: 297
? Coccorhagidia semiclavifrous Shiba, 1969: 83
Rhagidia shibai Strandtmann, 1971: 111
Poecilophysis (Soprocheles) saxonica: Zacharda, 1980: 635; 1983b: 312
Poecilophysis saxonica : Zacharda & Daniel, 1988: 58; Zacharda & Kučera, 2010: 5
Type locality: Leipzig, Anwald b. Dölitz (Germany)
Distribution: Czech Republic, Finland, Germany, Nepal (3900 m), Slovakia, Kirghizstan
Wankelia Zacharda, 1980: 642
Type species: Rhagidia reflexa wolmsdorfensis Willmann, 1936
Rhagidia terricola longipes : Pax, Maschke, 1935: 13
Rhagidia wolmsdorfensis : Willmann, 1941: 53; Willmann, 1954: 232; Baltac, 1973: 167
Rhagidia reflexa wolmsdorfensis Willmann, 1936: 302
Poecilophysis (Wankelia) wolmsdorfensis: Zacharda, 1980: 642; 1985: 74
Type locality: Wolmsdorf, Germany
Distribution: Czech Republic, Germany, Poland, Romania, USA, “Yugoslavia”
Megamerus Dugès, 1834: 50, 53 (nec Megamerus W.S. Mac Leay, 1827)
Scyphius C. L. Koch, 1835: fasc. 1, 14
Rhagidia Thorell, 1872: 700; Zacharda, 1980: 558
Nörneria Canestrini, 1886: 697
Scyphioides Berlese, 1886: 16, nom. nov. for Scyphius C. L. Koch nec Risso
Scyphoides Banks, 1900: 31 (lapsus calami?)
Type species: Rhagidia gelida Thorell, 1872
Austrorhagidia Zacharda, 1980: 560
Type species: Rhagidia mildredae Strandtmann, 1964
Rhagidia mildredae (part.): Strandmann, 1964: 157-160
Rhagidia campbellensis Zacharda: Qin, 1998: 300
Rhagidia (Austrorhagidia) campbellensis Zacharda, 1980: 563
Type locality: Auckland I.
Distribution: Auckland I.
Rhagidia gigas gerlachei Trouessart, 1903: 4-5
Rhagidia gerlachei Trouessart: Womersley & Strandtmann, 1963: 460; Strandtmann, 1967: 65; 1970: 98
Rhagidia (Austrorhagidia) gerlachei Zacharda, 1980: 563
Type locality: Antarctica, Détroit de Gerlache, South Georgia
Distribution: Antarctica (South Georgia)
Rhagidia mildredae Strandtmann, 1964: 157-160; Qin, 1998: 300
Rhagidia (Austrorhagidia) mildredae Zacharda, 1980: 561
Type locality: New Zealand, Beeman Hill, Campbell Island
Distribution: Subantarctic region (Auckland Is., Campbell I.)
Deharvengiella Zacharda, 1987: 2051
Type species: Rhagidia (Deharvengiella) troglomorphica Zacharda, 1987
Rhagidia (Deharvengiella) paraleloseta Zacharda, 2011: 669
Type locality: France
Distribution: France
Rhagidia (Deharvengella) serpentiforma Zacharda, 2011: 672
Type locality:
Distribution:
Rhagidia (Deharvengiella) troglomorphica Zacharda, 1987: 2051
Type locality: the Pyrenees, Haute-Garonne, Couledoux, Grotte de la Buhadére
Distribution: France
Dongnaia Zacharda, 1994: 49
Type species: Dongnaia edaphica Zacharda, 1994
Rhagidia (Dongnaia) edaphica Zacharda, 1994: 51
Type locality: Nam Cat Tien National Park, South Vietnam
Distribution: Vietnam
Nörneria Canestrini, 1886: 708; Zacharda, 1980: 565
Type species: Nörneria gigas Canestrini, 1886
Rhagidia (Noerneria) arizonensis Zacharda, 1995: 1259
Type locality: Arizona, Madera Canyon above Roundup Camp
Distribution: USA
Rhagidia breviseta Zacharda, 1995: 1248; Zacharda & Kučera, 2010: 5
Type locality: Quebec
Distribution: Canada, Finland
Rhagidia (Noerneria) compressa Zacharda, 1986: 2538
Type locality: British Columbia, Domonion Lake
Distribution: Canada
Rhagidia (Noerneria) distisolenidiata Zacharda, 1995: 1258
Type locality: New-Brunswick
Distribution: Canada
Nörneria gigas Canestrini, 1886: 708 -709
Rhagidia gigas : Trägårdh, 1904b: 3; Hull, 1918: 34; Thor & Willmann, 1941: 111-112; Ružička, Hajer & Zacharda, 1995: 47; Zacharda & Kučera, 2010: 5
Rhagidia intermedia Willmann, 1936: 293-294; 1951: 146; Haarløv, 1957: 27; Schmölzer, 1993: 59
Rhagidia danica Ehrnberger, 1977b: 95-100
Rhagidia carpatica Baltac, 1973: 168-171
Rhagidia (Nörneria) gigas Zacharda, 1980: 566
Type locality: Italy, Trentino
Distribution: Austria, Czech Republic, Denmark, Finland, Germany, Italy, Slovakia,
Rhagidia (Noerneria) longiseta Zacharda, 2000: 113; Zacharda & Kučera, 2010: 5
Type locality: Austria
Distribution: Austria, Finland
Rhagidia (Noerneria) parvilobata Zacharda, 1995: 1252; Zacharda & Kučera, 2010: 5
Type locality: Quebec
Distribution: Canada, Finland
Rhagidia (Noerneria) pasadenensis Zacharda, 1986: 2535
Type locality: Newfoundland, Pasadena
Distribution: Canada
Rhagidia (Noerneria) ponderosae Zacharda, 1983: 1000
Type locality: Baker Co., S Fork Forest Camp, 9 mi SW Unity
Distribution: USA (Oregon)
Rhagidia (Noerneria) punkva Zacharda, 1980: 575; 1983b: 314
Type locality: Moravia, Skalni mlýn, Czech Republic
Distribution: Czech Republic, Russia (Siberia)
Rhagidia (Noerneria) quadrisolenidiata Zacharda, 1995: 1262
Type locality: 53 mi north of Oaxaca, km 77, Oaxaca Highway 175, 3200 m
Distribution: Mexico
Rhagidia (Noerneria) rackae Zacharda, 1980: 582; Zacharda & Daniel, 1988: 58
Type locality: Velký-Bezdĕz-Hill, Czech Republic
Distribution: Czech Republic, Nepal (3900 m)
Rhagidia (Noerneria) ruseki Zacharda, 1980: 578; Hippa et al., 1989: 61
Type locality: Oblik-Hill near Louny, Czech Republic
Distribution: Czech Republic, Sweden
Rhagidia (Noerneria) varia Zacharda, 1985: 68
Type locality: Butler Cave, N of Burnsville, Both Co., Virginia
Distribution: USA
Rhagidia Thorell, 1872: 700; Zacharda, 1980: 558
Type species: Rhagidia gelida Thorell, 1872
Rhagidia comelicensis Lombardini, 1962: 191
Type locality: Comelico (Alpi orientali)
Distribution: Italy
Rhagidia crenata Shiba, 1969a: 77; Shiba, 1971a: 158; 1971b: 226
Type locality: Shiga Heights IBP Area
Distribution: Japan
Scyphius diversicolor C. L. Koch, 1838: fasc. 17, 22
Rhagidia diversicolor : Willmann, 1936: 297-298; 1954: 234; Cooreman, 1954: 10; 1959: 17; Zacharda & Kučera, 2010: 5
Rhagidia (Nörneria) diversicolor: Zacharda, 1980: 570
Rhagidia (Noerneria) diversicolor: Zacharda, 1985: 71
Type locality: France, Germany
Distribution: Czech Republic, Finland, Germany, Poland, Switzerland, USA
Rhagidia gelida : Thorell, 1872: 701; Trägårdh, 1904: 44; Thor, 1912: 235; Banks, 1919: 11; Oudemans, 1926: 106; Sellnick, 1940: 31; Thor & Willmann, 1941: 109-111; Macfadyen, 1954: 284; Cooreman, 1959: 27 (var.); Strandtmann, 1971: 106-107; Seniczak et al., 2020: 1
Rhagidia (R.) gelida Zacharda, 1980: 559
Type locality: Norway, Beeren Eiland, Svalbard
Distribution: Canada, Czech Rep., Greenland, Yan Mayen I., Iceland, Norway, Russia, Spitzbergen, Switzerland, USA (Alaska)
Rhagidia orghidani Bannova, Sarmanova & Zhuravleva, 1977: 57
Type locality: Romania
Distribution: Romania
Rhagidia (Rhagidia) qaliubiensis Abou-Awad, El-Sawaf & Abdel-Khalek, 2009: 150
Type locality: El-Kaliubiya region
Distribution: Egypt
Rhagidia buettneri Willmann, 1955: 209; Zacharda, 1980 (sp. inc. sed., “The holotype fragmentary, a tritonymph”)
Type locality: SW Saxony, Brummeisenmoor
Distribution: Germany
Rhagidia meyerae Halliday, 2005: 42
Type locality: South Africa
Distribution: South Africa
Rhagidia uenoi Morikawa, 1963: 18; Shiba, 1971: 224
Type locality: Japan
Distribution: Japan
Robustocheles Zacharda, 1980: 527
Type species: Robustocheles mountana Zacharda, 1980
Amoveocheles Zacharda, 1980: 540
Type species: Robustocheles (Amoveocheles) tricuspidata Zacharda, 1980
Robustocheles (Amoveocheles) californiensis Zacharda, 1986: 2535
Type locality: California, Albany
Distribution: USA (California)
Robustocheles (A.) dentata Zacharda, 1980: 543
Type locality: Prague, Czech Republic
Distribution: Czech Republic
Robustocheles (A.) tricuspidata: Zacharda, 1980: 540
Type locality: Závist-Knoll near Zbraslav, Czech Republic
Distribution: Czech Republic
Lewia Zacharda, 1980: 547
Type species: Rhagidia hilli Strandtmann, 1971
Rhagidia hilli Strandtmann, 1971: 113
Rhagidia grahami Elliott, 1976: 11-14
Robustocheles (Lewia) hilli Zacharda, 1980: 547; 1985: 79
Type locality of Rhagidia hilli: Alaska, Fairbanks
Type locality of Rhagidia grahami: Unita Co., Little Brush Creek Cave, Utah
Distribution: Canada, N-W Territorium, Bathurst I.; USA
Robustocheles (Lewia) lacustris Zacharda, 1997: 1090
Type locality: Cypress Hills Provincial Park (Alberta – Saskatchewan)
Distribution: Canada
Robustocheles (Lewia) obscuritata Zacharda, 1997: 1094; Zacharda & Kučera, 2010: 5
Type locality: Gaspésie Provincial Park (Quebec)
Distribution: Canada, Finland
Robustocheles (Lewia) spinaea Zacharda, 1997: 1094
Type locality: 20 m SE Gallup, New Mexico
Distribution: USA
Mexia Zacharda, 1997: 1084
Type species: Robustocheles (Mexia) longichelae Zacharda, 1996
Robustocheles (Mexia) longichelae Zacharda, 1997: 1085
Type locality: Chiapas, 7 m S El Bosque
Distribution: Mexico
Robustocheles (Mexia) occulta Zacharda & Pugsley, 1988: 646
Type locality: Alberta (Banff Nat. Park, Castleguard Cave); Iowa, Washington
Distribution: Canada, USA
Robustocheles Zacharda, 1980: 528
Type species: Robustocheles montana Zacharda, 1980
Robustocheles (R.) deltacus Abou-Awad, El-Sawat & Abdel-Khalek, 2009: 145
Type locality: Ain Shams Univ., El Qaluibiya
Distribution: Egypt
Robustocheles hamedanensis Zacharda, 2012: 31
Type locality: Hamedan-Shokriyeh
Distribution: Iran
Robustocheles (R.) montana Zacharda, 1980: 529
Type locality: Velká Fatra-Mountains, Černy Kameň, Slovakia
Distribution: Germany, Slovakia
Rhagidia mucronata Willmann, 1936: 297; 1951b: 143; Sellnick, 1940: 31; Evans, 1952: 666; Cooreman, 1954: 10; Haarløv, 1957: 31; Strandtmann & Prasse, 1977: 14
Robustocheles (R.) mucronata (Willmann): Zacharda, 1980: 532; Abou-Awad, 1985: 258; Zacharda & Kučera, 2010: 5; Darbemamieh, Hajiqanbar & Khanjani, 2013: 456
Rhagidia neoshibai Shiba, 1976: 132; syn.: Zacharda, 1984: 870
Robustocheles mucronata (Willmann): Zacharda, 1994: 53
Type locality: Nuremberg, Germany
Distribution: Brazil, Corsica, Czech Republic, Danemark, Egypt, England, Finland, Germany, Iceland, Iran, Poland, Russia (Caucasus), Slovakia, Thailand, USA (Alaska), Vietnam
Robustocheles (R.) pascuaria Zacharda, 1997: 1076
Type locality: Cypress Hills Provincial Park
Distribution: Canada
Rhagidia mordax grandis Willmann, 1935: 28; 1936: 296; Cooreman, 1959: 19; Leruth, 1939: 411
Robustocheles (R.) robusta Zacharda, 1980: 537
Type locality: Czech Republic, Oblik Hill
Distribution: Belgium, Germany, Czech Republic, Switzerland
Robustocheles (R.) similis Zacharda, 1997: 1080
Type locality: Nova Scotia, Cape Breton Highlands
National Park, Paquettes Lake Trail
Distribution: Canada
Shibaia Zacharda, 1980: 737
Type species: Rhagidia longisensilla Shiba, 1969
Rhagidia coreaensis Shiba, 1969: 608
Shibaia coreaensis Zacharda, 1980: 742; 1985: 598
Type locality: Samsheong-gun, Kangweon-do, Yongyeon-gul Cave near Yongyeon-gog, Changseong-eub
Distribution: South Korea
Rhagidia heteropoda Berlese, 1910: 243
Rhagidia longisensilla Shiba, 1969a: 79; Baltac, 1974: 37
Type locality: “in muscis Etruriae”, Toiano, Piza
Distribution: Italy, Japan, Romania
Rhagidia longisensilla Shiba, 1969: 79; 1969b: 608; Strandtmann, 1971: 104; ? Elliott & Strandtmann, 1971: 475; Baltac, 1974: 41; Strandtmann & Goff, 1978: 135; Goff, 1987: 35; Darbemamieh, Hajiqanbar & Khanjani, 2013: 456
Shibaia longisensilla Zacharda, 1980: 738; Zacharda & Daniel, 1988: 58
Shibaia longicornis (Shiba, 1959) - Iran
Type locality: Japan
Distribution: Austria (3133 m), Czech Republic; Germany, Iran, Japan, Alaska, Hawaii, Korea, Nepal (3900 m)
Shibaia shereefi Abou-Awad, 1985: 253
Type locality: Giza region
Distribution: Egypt
Shibaia spiraesolenidiata Zacharda, 1995: 594
Type locality: Nova Scotia, Kejim Kujuk Nat. Park
Distribution: Canada
Shibaia tatrica Zacharda, 1980: 744
Type locality: High-Tatras, Mlynická dolina, 1450 m
Distribution: Slovakia
Thoria Zacharda, 1980: 747
Type species: Thoria brevisensilla Zacharda, 1980
Rhagidia uniseta : Strandtmann, 1971: 105 (not R. uniseta Sig Thor, 1909)
Thoria brevisensilla Zacharda, 1980: 748; Goff, 1987: 35
Type locality: Bohemia bor., Oblik-Hill near Luny
Distribution: Czech Republic, Hawaii
Rhagidia uniseta Sig Thor, 1909: 10; 1912: 236; Sellnick, 1940: 31; Thor & Willmann, 1941: 117; Mihelčič, 1958a: 103; Strandtmann, 1971: 105; Schmölzer, 1993: 59;
?Rhagidia crassipes Schweizer & Bader, 1963: 246
Thoria uniseta Zacharda, 1980: 752; Zacharda & Kučera, 2010: 5
Type locality: Norway
Distribution: Alaska, Austria, Czech Republic, Finland, Iceland, Norway, Switzerland
Traegaardhia Zacharda, 1980: 715
Type species: Rhagidia dalmatina Willmann, 1939
Traegaardhia cavernarum Zacharda, 2010: 26
Type locality: Buso del Marmo Cave (1202 V/BL), Quero (Belluno), Massiccio del Monte Grappa, Veneto, Italy (340 m)
Distribution: Italy
Traegaardhia cavernicola Zacharda, 2010: 13
Type locality: Grotte Vecchia Diga Cave, Barcis (Pordenone), Monte Cavallo
mountain range, the Stubai Alps, Friuli
Distribution: Italy
Rhagidia dalmatina Willmann, 1939: 244; 1951: 146; Mihelčič, 1958a: 103; 1958c: 38; Strandtmann, 1971: 103
? Rhagidia gigas var. longipes Trägårdh, 1912: 608
? Rhagidia longipes: Cooreman, 1959: 18
Rhagidia terricola var. longipes (Trägårdh): Verhoeff, 1912: 155
Traegaardhia dalmatina Zacharda, 1980: 715
Type locality: Croatia, Dalmatia, cave near Krivošije
Distribution: Austria, Croatia, ?Germany, France, Spain
Traegaardhia distosolenidia Zacharda, 2010: 36; Çetin et al., 2021: 232
Type locality: Arcugnano (Vicenza), the Monti Berici Mts, Veneto
Distribution: Italy, Turkey
Traegaardhia gracilis Zacharda, 2010: 31
Type locality: the Boia valley, Valdagno(Vicenza), the Lessinian Mts, Venetia
Distribution: Italy
Foveacheles (Mediostella) holsingeri Zacharda, 1980: 677
Foveacheles cf. holsingeri: Zacharda, 1985: 78
Traegaardhia holsingeri (Zacharda): Zacharda et al., 2010: 45
Type locality: Kentucky, Mammoth Cave
Distribution: USA
Traegaardhia nasuta Zacharda, 2010: 21
Type locality: Buso dei Zambon Cave, San Pietro di Feletto (Treviso), the Hills of
Conegliano, Veneto, northeastern Italy
Distribution: Italy
Traegaardhia similis Zacharda, 2010: 40
Type locality: Toney Bend Mine No. 2, Marion Co., Arkansas
Distribution: Italy
Traegaardhia subterranea Zacharda, 2010: 52
Type locality: Amarcord Cave (6391 V/VI), Valdagno (Vicenza), Lessinian Mts
Distribution: Italy
Traegaardhia vicenzaensis Zacharda, 1994b: 93
Type locality: Veneto, M. Lessini, Torrebelvicino (Vicenza); quota 386 in a cave, Buso del Monte Naro
Distribution: Italy
Troglocheles Zacharda, 1980: 646
Type species: Rhagidia strasseri Willmann, 1932
Troglocheles aggerata Zacharda, 2000: 472
Type locality: Tyrol, Oetztal Alps, Obergurgl
Distribution: Austria
Troglocheles archetypica Zacharda, 2000: 464
Type locality: Tyrol, Oetztal Alps, Obergurgl
Distribution: Austria
Troglocheles christiani Zacharda, Isaia & Piva, 2011: 656
Type locality: Türkenloch Cave near Kleinzell, 721 m
Distribution: Austria
Rhagidia gineti Cooreman, 1959: 23; Strandtmann, 1971: 103; Dobat, 1976
?Rhagidia conciana Lombardini, 1951: 247
Troglocheles gineti : Zacharda, 1980: 655
Type locality: Ain, Grotte de la Buire
Distribution: France, Germany, Switzerland
Troglocheles lanai Zacharda, Isaia & Piva, 2011: 649
Type locality: Vallone dell’Infernotto
Distribution: Italy
Rhagidia odontochela Turk, 1972: 190
Troglocheles odontochela (Turk): Zacharda, 1984: 869
Type locality: England
Distribution: England
Troglocheles quinquesolenidiata Zacharda, Isaia & Piva, 2011: 643
Type locality: Buso Brutto Cave, 918 m
Distribution: Italy
Rhagidia strasseri Willmann, 1932: 3; Thor & Willmann, 1941: 119; Mihelčič, 1958; Cooreman, 1954: 11; 1959: 22
Rhagidia gineti : Rack, 1974: 129, not gineti Cooreman
Troglocheles strasseri Zacharda, 1980: 650; 2000:
Type locality: shaft near Na Lazi, Tarnowaner Wald -Trnovski Gozd-Forest, Slovenia (lectotype)
Distribution: Austria, France, Slovenia, Switzerland
Rhagidia strasseri tirolensis Willmann (unpubl.); Zacharda, 1980: 650
Type locality: Austria, the Kaisergebirge-Mts., Estendorfer Schacht-shaft
Distribution: Austria
Troglocheles vandeli Zacharda, 1987: 2052
Type locality: The Pyrenees, Haute-Garonne, Herran, Trou Mille
Distribution: France
Rhagidia vornatscheri Willmann, 1953: 218
Troglocheles vornatscheri Zacharda, 1980: 653; 2000:
Type locality: Austria, Herdengelhöhle Cave
Distribution: Austria
Tuberostoma Zacharda, 1980: 763
Type species: Coccorhagidia gressitti Womersley et Strandtmann, 1963
Coccorhagidia gressitti Womersley & Strandtmann, 1963: 467; Strandtmann, 1967: 66
Tuberostoma gressitti (Womersley et Strandtmann): Zacharda, 1980: 764
Type locality: Antarctica, W.S.W. from Hallet Station
Distribution: Antarctica
Rhagidia leechi Strandtmann, 1963: 463; Strandtmann, 1970: 100
Tuberostoma leechi (Strandtmann): Zacharda, 1980: 768
Type locality: Livingstone L, South Shetland Is., 62°42‘S, by 60°26/W
Distribution: Antarctica, South Shetland, South Georgia
Coccorhagidia keithi Strandtmann, 1967: 67
Tuberostoma keithi (Strandtmann) Zacharda, 1980: 766
Type locality: Antarctica, Nunatak, 3 mi. NE, Mt. Mirray, Mowson Ge. area
Distribution: Antarctica
Krantzia Zacharda, 1983: 993 (nomen praeoccupatum)
Zachardaia Özdikmen, 2008: 220
Type species: Krantzia quadriseta Zacharda, 1983
Krantzia quadriseta Zacharda, 1983: 993
Type locality: Oregon, Jefferson Co., Camp Sherman
Distribution: USA
In the monograph of Zacharda (1980) are listed 48 species as «Species incertae sedis et inquirenda«. Here is this list (shortly):
1. Scyphius subterraneus Absolon, 1899 - The type unknown.
2. Rhagidia margaretae Baltac, 1974 – Type not seen by Zacharda.
3. Rhagidia strandtmanni Baltac, 1976 - Type not seen by Zacharda.
4. Rhagidia grandjeani Baltac, 1976 - Type not seen by Zacharda.
5. Rhagidia pallida Banks, 1894 – The type unknown. Published also by Miller, 1925.
6. Rhagidia cavicola Banks, 1897 - The type unknown.
7. Nörneria ? maxillata Berlese, 1893 - Type not seen by Zacharda.
8. Rhagidia subterranea Berlese, 1905 - The type unknown.
9. Rhagidia heteropoda Berlese, 1910 - The type unknown.
10. Megamerus fallax Dugès, 1834 - The type unknown.
11. Megamerus roseus Dugès, 1834 - The type unknown. France.
12. Rhagidia trisetata Elliot et Strandtmann, 1971 - Type not seen by Zacharda.
13. Rhagidia muscicola Hull, 1918 - The type unknown. England.
14. Scyphoides maxillatus Karpeles, 1891 - The type unknown.
15. Scyphius pyrrholeucus Koch, 1838 – The type unknown.
16. Scyphius cerinus Koch, 1838 – The type unknown.
17. Scyphius coarctatus Koch, 1838 – The type unknown.
18. Scyphius cylindricus Koch, 1838 – The type unknown.
19. Scyphius diaphanus Koch, 1838 – The type unknown.
20. Scyphius albellus Koch, 1838 – The type unknown.
21. Scyphius oblitteratus Koch, 1838 – The type unknown.
22. Scyphius elongatus Koch, 1838 – The type unknown.
23. Scyphius reflexus Koch, 1838 – The type unknown.
24. Scyphius hamatus Kramer et Neuman, 1883 - The type unknown.
25. Scyphius japonicus Kramer et Neuman, 1883 - The type unknown.
26. Nörneria cylindrica Kramer, 1898 (nec Scyphius cylindricus Koch, 1838) - The type unknown.
27. Gamasus halophilus Laboulbene, 1851 - The type unknown.
28. Rhagidia conciana Lombardini, 1951 - The type unknown.
29. Rhagidia comelicensus Lombardini, 1962 - The type unknown.
30. Rhagidia japonica Morikawa, 1963 - Type not seen by Zacharda.
31. Rhagidia mordax Oudemans, 1906 - Type not seen by Zacharda.
32. Rhagidia fragosa Schuster, 1958 – The holotype unknown.
33. Rhagidia raetica Schweizer et Bader, 1963 - Type not seen by Zacharda.
34. Rhagidia crassipes Schweizer et Bader, 1963 - Type not seen by Zacharda.
35. Coccorhagidia clavifrons Shiba, 1969 - Type not seen by Zacharda.
36. Coccorhagidia semiclavifrons Shiba, 1969 - Type not seen by Zacharda.
37. Rhagidia crenata Shiba, 1969 - Type not seen by Zacharda.
38. Scyphius maniacus Stoll, 1890 - The type unknown.
39. Rhagidia bidens Sig Thor, 1909 - The type unknown.
40. Rhagidia birulai Sig Thor, 1909 - The type unknown. Yan Mayen I. (Macfadyen, 1954)
41. Rhagidia megalochela Trägårdh, 1907 - The type unknown.
42. Rhagidia gigas var. longipes Trägårdh, 1907 - The type unknown.
43. Rhagidia mordax var. grandis Willmann, 1936 - The type unknown.
44. Rhagidia unidentata Willmann, 1939 - The type unknown.
45. Rhagidia dalmatina gigantea Willmann, 1941 - The type unknown.
46. Rhagidia paxi Willmann, 1942 - The holotype fragmentary, probably a nymph.
47. Rhagidia strouhali Willmann, 1953 - The type unknown.
48. Rhagidia pseudocrenata Shiba, 1976 Malaysia Pasoh Forest Reserve
EUROPE
Austria (Irk, 1939; Franz, 1943, 1954; Willmann, 1951a, 1951b, 1953; Janetschek, 1959, 1993; Mihelčič, 1957, 1958; Schmölzer, 1962, 1993; Zacharda, 1980, 1994, 2000d, Zacharda, Isaia & Piva, 2011; Zacharda & Kučera, 2006) – Coccorhagidia clavifrons, C. berlesei, C. divergens (inquir.), Evadorhagidia janetscheki, Foveacheles brevichelae, F. mira, F. osloensis, F. proxima, F. unguiculata, F. willmanni, Foveacheles (Proxistella) terricola, Foveacheles (Ternirhagidia) alpina, Poecilophysis (D.) pratensis, P. (D.) wankeli, Rhagidia gigas (= intermedia), R. longiseta, R. reflexa, Robustocheles mucronata, Shibaia longisensilla, Thoria uniseta, Traegaardhia dalmatina, Troglocheles aggerata, T. archetypica, T. christiani, T. strasseri, T. strasseri tirolensis, T. vornatscheri,
Azerbaidzhan (Zacharda, 1983b) - Poecilophysis (Dentocheles) pratensis
Belgium (Leruth, 1939; Cooreman, 1943) – Foveacheles terricola, Poecilophysis spelaea, Poecilophysis (Dentocheles) pratensis, Robustocheles (R.) robusta,
Bulgaria (Turk, 1970) – Poecilophysis spelaea, P. wolmsdorfensis; Inc. sedis: Rhagidia strouhali
Croatia (Willmann, 1939; Zacharda, 1980) – Traegaardhia dalmatina
Czech Rep. (Wankel, 1861; Willmann, 1954; Zacharda, 1980, 1993, 2000d; Zacharda et al., 2005) – Brevipalpia minima, Coccorhagidia clavifrons, Crassocheles muralis, C. virgo, Evadorhagidia bezdezensis, E. oblikensis, E. corcontica, Foveacheles osloensis, F. brevichelae, Foveacheles (Proxistella) terricola, F. (P.) simulate, Hammenia macrostella, Latoempodia macroempodiata, Poecilophysis arena, P. spelaea, P. pratensis, P. recussa, P. saxonica, P. wolmsdorfensis, P. (D.) wankeli, Rhagidia diversicolor, R. gelida, R. gigas, R. punkva, R. rackae, R. ruseki, [R. reflexa], Robustocheles mucronata, R. robusta, R. dentata, R. tricuspidata, Shibaia longisensilla, Thoria brevisensilla, Th. uniseta, [R. berlesei]
Denmark (Haarløv, 1957) – Coccorhagidia clavifrons, C. subterranea, Poecilophysis (Dentocheles) pratensis, Robustocheles mucronata, Rhagidia intermedia (?)
Greenland (Trägårdh, 1904) – Rhagidia gelida
Faeroe Is. (Trägårdh, 1931) – Poecilophysis faeroensis
Yan Mayen I. (Macfadyen A. 1954) – Coccorhagidia clavifrons, Rhagidia birulai, Rh. gelida, Rh. terricola
Finland (Zacharda & Kučera, 2010) – Rhagidia breviseta, Rh. diversicolor, Rh. gigas, Rh. longiseta, Rh. parvilobata, Poecilophysis pratensis, P. pseudoreflexa, P. spelaea, P. saxonica, Foveacheles brevichela, F. cegetensis, Robustocheles mucronata, R. obscuritata, Thoria uniseta, Evadorhagidia sp.
France (contin.) (Berlese & Trouessart, 1889; Cooreman, 1954, 1959; Zacharda, 1980; 1987, Zacharda, Grafitti & Piva, 2011) – Foveacheles canestrini, Foveacheles (Proxistella) terricola, Poecilophysis (Dentocheles) pratensis, P. spelaea, Rhagidia (Deharvengiella) troglomorphica, Rhagidia (Noerneria) diversicolor, Rh. longipes, Traegaardhia dalmatina, Troglocheles gineti, T. strasseri, T. vandeli, Rh. halophila
Corsica (Cooreman, 1954) - Robustocheles mucronata
Germany (C.L. Koch, 1835; Strandtmann & Prasse, 1977; Ehrnsberger, 1977a, 1977b; Willmann, 1934, 1955; Dobat, 1976; Zacharda, 1980) – Coccorhagidia clavifrons, Foveacheles osloensis, F. magna, F. harzensis, F. clavacrinita, Poecilophysis faeroensis, P. (Dentocheles) pratensis, Rhagidia Buettneri, Rh. gigas (= danica), R. diversicolor, R. clavicrinita, R. longipes, Robustocheles montana, R. mucronata, R. robusta, Shibaia longisensilla, Parallelorhagidia evansi, Troglocheles gineti
Great Britain (Hull, 1918; Evans, 1952; Turk & Turk, 1952; Turk, 1972) – Coccorhagidia clavifrons, C. subterranea, Poecilophysis spelaea, Rhagidia gigas, Robustocheles mucronata, Troglocheles odontochela, “Rhagidia vitzthumi”, “Rhagidia muscicola”
Hungary (Dudich et al., 1940) - Coccorhagidia clavifrons
Iceland (Sellnick, 1940) – Poecilophysis faeroensis = reflexa, P. mucronata, P. pratensis, P. recussa, Rhagidia birulai, Rh. gelida, Rh. terricola, Thoria uniseta,
Surtsey (Lindroth, Andersson, Bödvarsson & Richter, 1973) - Rhagidiidae
Ireland (Halbert, 1915) – Rhagidia halophila, Rh. terricola
Italy (contin.) (Canestrini, 1886; Berlese, 1905, 1910; Bottazzi, 1950; Lombardini, 1962; Zacharda, 2010, 2011a, Zacharda, Grafitti & Piva, 2011; Zacharda, Isaia & Piva, 2011,) – Rhagidia comelicensis, Rh. gigas, Coccorhagidia clavifrons, C. subterranea, Foveacheles terricola, Traegaardhia cavernarum, T. cavernicola, T. distosolenidia, T. gracilis, T. nasuta, T. subterranea, Troglocheles lanai, T. quinquesolenidiata
Sardinia (Zacharda, Grafitti & Piva, 2011) -
Norway (Sig Thor, 1909, 1912, 1934; Hippa & Koponen, 1989; Hippa et al., 1989; Zacharda & Kučera, 2010) – Foveacheles (F.) osloensis, Foveacheles (Proxistella) terricola, Poecilophysis spelaea, Thoria uniseta
Beeren Eiland (Thorell, 1872) – Rhagidia gelida
Spitzbergen (Svalbard)(Thorell, 1971; Oudemans, 1926; Szeniczak et al., 2020) – Rhagidia gelida
Poland (Jesionowska, 2002) – Janes rafalskii, Rhagidia diversicolor, Robustocheles mucronata
Romania (Cooreman, 1951; Baltac, 1973, 1974, 1976, 1977b; Bannova, Sarmanova, Zhuravleva, 1977) – Coccorhagidia montana, Poecilophysis spelaea, P. wolmsdorfensis, Rhagidia longipes, Rh. orghidani, Rh. margaretae (sp.inquir.), Rh. strandtmanni (sp.inquir.), Rh. grandjeani (sp.inquir.), Shibaia heteropoda (sub «Rhagidia longisensilla)
Russia (all, continent.)(Thor, 1909; Mac Lean, Behan & Fjellbeg, 1978; Zacharda, 1983b) – [Rh. birulae Thor] Coccorhagidia clavifrons, C. pittardi, Evadorhagidia bezdezensis, Foveacheles (F.) brevichelae, F. (F.) caucasica, F. (F.) cegetensis, Poecilophysis (Procerocheles) pseudoreflexa, Poecilophysis (Procerocheles) recussa, P. (Sophrocheles) saxonica, P. (Dentocheles) pratensis, Rhagidia (Noerneria) punkva, Thoria brevisensilla, Th. uniseta
Crimea (Khaustov, 2015) – Paracrassocheles clavisetosus
Slovakia (Zacharda, 1980; 1988; Zacharda & Kučera, 2006; Kalúz, 2017) – Coccorhagidia clavifrons, C. pittardi, Foveacheles rupestris, Foveacheles (Spelaeocheles) troglodyta, Latoempodia similis, Poecilophysis pratensis, P. saxonica, Poecilophysis (D.) wankeli, Robustrocheles montana, R. mucronata, Rhagidia gigas, Shibaia tatrica
Slovenia (Willmann, 1932) – Troglocheles strasseri
Spain (cont.)(Mihelčič, 1958c; Barranco & Amate, 2000) - Foveacheles cannadasi, F. terricola, Traegaardhia dalmatina
Sweden (Hippa et al., 1989; Zacharda & Kučera, 2010) – Evadorhagidia sp., Foveacheles osloensis, Poecilophysis faeroensis, P. pseudoreflexa, P. spelaea, P. weyerensis, Rhagidia gigas, Rh. ruseki, Robustocheles robusta
Switzerland (Cooreman, 1954, 1959) – Foveacheles (Proxistella) terricola, Poecilophysis (Procerocheles) faeroensis (= Rh. reflexa), Poecilophysis spelaea, Rhagidia diversicolor, Rh. gelida, Robustocheles (R.) robusta,Troglocheles gineti, T. strasseri
Turkey (Çetin et al., 2021) – Traegaardhia distosolenidia
AFRICA
Egypt (Trägårdh, 1904b; Abou-Awad, 1985; Abou-Awad & El-Bagouri, 1986; Abou-Awad, El-Sawaf & Abdel-Khalek, 2009) - Coccorhagidia clavifrons, Foveacheles (F.) lemoni, Neothoria niloticus, Robustocheles (R.) deltacus, R. (R.) mucronata, Shibaia shereefi, Thoria sp., Neothoria zacharda, Robustocheles (R.) deltacus, Rhagidia (R.) qualiubeinsis
Nigeria – Rhagidiidae gen.sp.
South Africa (Meyer & Ryke, 1960) - Coccorhagidia clavifrons
ASIA
Cyprus (Zacharda, 2010) – Foveacheles thaleri
Iran (Zacharda et al., 2012; Darbemamieh, Hajiqanbar & Khanjani, 2013; Yazanpanah, Kazemi & Zacharda, 2014) – Brevipalpia minima, Coccorhagidia clavifrons; C. pittardi; Foveacheles brevichelae, F. cegetensis, F. osloensis, Poecilophysis arena, P. faeroensis, P. pratensis, Rhagidia cf. breviseta, Robustocheles hamedanensis, R. mucronata
Japan (Ehara, 1960; Morikawa, 1963; Shiba, 1969, 1971a, 1971b) – Poecilophysis pratensis (= Rhagidia japonica Morikawa), [Rhagidia crenata, Rh. uenoi], Shibaia heteropoda (= longisensilla)
Kirghizstan (Zacharda, 1983b) - Poecilophysis (Procerocheles) pseudoreflexa, P. (Dentocheles) pratensis, P. (Sophrocheles) saxonica, Robustocheles mucronata, Thoria brevisensilla
Korea (Shiba, 1969b; Zacharda, 1980) – Shibaia coreanensis,
Malaysia (Shiba, 1976) - Rhagidia whartoni = Poecilophysis pratensis, Robustocheles (R.) mucronata (sub Rhagidia neoshibai), Rhagidia strandtmanni, Rh. pseudocrenata (species inquirendae)
Nepal (Zacharda & Daniel, 1987) – Brevipalpia minima, Coccorhagidia pittardi, Poecilophysis saxonica, Rhagidia rackae, Shibaia longisensilla
Thailand (Zacharda, 1994c) – Robustocheles mucronata
Vietnam (Zacharda, 1994c) – Foveacheles (Achaetorhagidia) namcatiensis, F. (A.) tropica, Rhagidia (Dongnaia) edaphica, Robustocheles mucronata
NORTH AMERICA
Canada (Banks, 1919; Zacharda, 1980; 1985, 1986, 1995, 1997; Zacharda & Pugsley, 1988; Beaulieu et al., 2019) – Coccorhagidia clavifrons, Evadorhagidia quinqueseta, Foveacheles arenaria, F. brevichelae, F. tuktoaktukensis, Foveacheles (Mediostella) vancouverensis, Kouchibouguacia cryptosolenidiata, K. transcendata, Lindquistula multisolenidiata, Parallelorhagidia parva, Poecilophysis pseudoreflexa, Rhagidia gelida, Rhagidia (Noerneria) compressa, Rh. (N.) distisolenidiata, Rh. (N.) parvilobata, Rh. (N.) pasadenensis, Robustocheles (Lewia) lacustris, R. (L.) obscuritata, R. (R.) similis, Robustocheles (Mexia) occulta, R. pasquaria, Shibaia spiraesolenidiata
Mexico (Elliott & Strandtmann, 1971; Zacharda & Elliott, 1986; Zacharda, 1995, 1997) - Rhagidia (N.) quadrisolenidiata, ?Robustocheles infernalis, R. longichelae, ?R. trisetatus
USA (Banks, 1894, 1897, 1904; Call, 1897; Holsinger, 1965; Strandtmann, 1971; Elliott, 1976; Zacharda, 1980; 1983; 1985, 1986, 1995, 2010, 2011; Zacharda & Elliott, 1983; Lindquist & Zacharda, 1987; Zacharda et al., 2010) – Arhagidia monothrix, Coccorhagidia clavifrons, Elliotta howarthi, Foveacheles bentonensis, Foveacheles (Trofocheles) paralleloseta, Foveacheles (Propriorhagidia) tuktoyaktukensis, Zachardaia quadriseta, Poecilophysis (Dentocheles) affinis, P. (D.) extraneostella, P. (D.) pratensis, P. (D.) weyerensis, Poecilophysis (Procerocheles) oregonensis, P. melanoseta, P. saxonica, P. (Wankelia) wolmsdorfensis, Rhagidia cavicola, Rh. diversicolor, Rh. (Noerneria) arizonensis, Rh. (Noerneria) ponderosae, R. varia, Robustocheles (R.) pascuaria, Robustocheles (Amoveocheles) californiensis, Robustocheles (Lewia) hilli, R. mucronata, Traegaardhia holsingeri, T. similis
Alaska (Strandtmann, 1971; Zacharda, 1980, 1996; Zacharda & Elliott, 1985, Zacharda & Pugsley, 1988) – Arctorhagidia mariehammerae, A. sateri, Coccorhagidia pittardi, Eskimaia capitata, Foveacheles pribilovi, Kouchibouguacia transcendata, Robustocheles mucronata, R. hilli, R. occulta, Rhagidia gelida, Thoria uniseta [Rh. whartoni, Rh. shibai]
Hawaii (Strandtmann & Goff, 1978; Zacharda, 1980; 1982; Goff, 1987) – Coccorhagidia clavifrons, Elliotta howarthi, Foveacheles goffi, F. tenorioae, Parallelorhagidia hawaiiensis
SOUTH and CENTRAL AMERICA
Argentina (Bedano, 2004) - Coccorhagidia clavifrons
Brazil – Robustocheles mucronata
Chile (Kramer, 1898) – Rhagidia krameri Sig Thor, 1941= Nörneria cylindrica Kramer, 1898
Cuba (Baltac, 1977a) – Coccorhagidia cubana (sp. inquir.)
Nicaragua (Maes et al., 1989) - Robustocheles (R.) mucronata
OCEANIA
Australia (continent)(Wood, 1970, Qin, 1998) – Rhagidia sp.
New Zealand (Qin, 1998) – Coccorhagidia clavifrons
Macquarie Island (Womersley & Strandtmann, 1963; Luxton, 1990) – Poecilophysis macquariensis
Campbell Island (Strandtmann, 1964) – Rhagidia mildredae
ANTARCTICA
Antarctida (Trouessart, 1903; Zacharda, 1980) – Rhagidia (Austrorhagidia) gerlachei
Possession I. (Strandtmann & Davies, 1972) – Poecilophysis kerguelenensis
South Georgia (Strandtmann, 1970) - Rhagidia (Austrorhagidia) gerlachei, Tuberostoma leechi
Patria incognita (Zacharda, 1980) – Foveacheles emendata
Eriorhynchidae Qin & Halliday, 1997: 152
Small family of five Australian species, described by Womersley (1941) and Qin & Halliday (1997) and recently detached from the Penthalodidae.
Eriorhynchidae Qin & Halliday, 1997: 152
Type genus: Eriorhynchus Qin et Halliday, 1997
Eriorhynchus Qin & Halliday, 1997: 153
Type species: Eriorhynchus womersleyi Qin et Halliday, 1997
Penthalodes australicus Womersley, 1941: 294
Eriorhynchus australicus (Womersley): Qin & Halliday, 1997: 103
Type locality: Victoria, Acheron Way
Distribution: Australia (Victoria)
Eriorhynchus hades Qin & Halliday, 1997: 156
Type locality: Victoria, Rokewood Cemetery
Distribution: Australia (Victoria)
Eriorhynchus ramosus Qin & Halliday, 1997: 157
Type locality: Tasmania, Big Sassy Creek
Distribution: Australia (Tasmania)
Eriorhynchus walteri Qin & Halliday, 1997: 161
Type locality: Miala Nat. Park, Brisbane
Distribution: Australia (Queensland)
Eriorhynchus womersleyi Qin & Halliday, 1997: 161
Type locality: New South Wales, 34.6 km south of Bombala
Distribution: Australia (N.S.Wales)
OCEANIA
Australia (continent)(Womersley, 1941; Qin Ting-Kui & Halliday, 1997) – Eriorhynchus australicus, E. hades, E. waltheri, E. womersleyi
Tasmania (Qin Ting-Kui & Halliday, 1997) – Eriorhynchus ramosus
Pentapalpidae Olivier & Theron, 2000: 386
Small family of one known genus and species from South Africa.
Fam. Pentapalpidae Olivier et Theron, 2000 - 1 gen., 1 sp.
Pentapalpus Olivier et Theron, 2000 - 1 sp.
Pentapalpidae Olivier & Theron, 2000: 386
Type genus: Pentapalpus Olivier et Theron, 2000
Pentapalpus Olivier & Theron , 2000: 386
Type species: Pentapalpus unguempodius Olivier et Theron, 2000
Pentapalpus unguempodius Olivier & Theron, 2000: 386
Type locality: Eastern Cape Province, soil, Summerstrand, Porth Elizabeth, 33.58S, 25.38E
Distribution: South Africa
AFRICA
South Africa (Olivier & Theron, 2000) - Pentapalpus unguempodius
Penthalodidae Sig Thor, 1933: 274
Small or medium-sized (330 – 1500 µm) strongly sclerotisezed mites living in soil or moss. Most (39) species are within the two “old” genera - Penthalodes Murray, 1877 and Stereotydeus Berlese, 1901. Further contributions are due to Baker, Trägårdh, Khaustov, Mihelčič, Strandtmann, Goff, Qin, Thor, Jesionowska, Shiba, Womersley, Olivier, Trouessart, starting with Dugès (1834).
Family Pentalodidae Thor, 1933 - 5 gen., 44 sp.
Callipenthalodes Qin, 1998 – 1 sp.
Hawaiieupodes Strandtmann et Goff, 1978 – 1 sp.
Penthalodes Murray, 1877 - 10 sp.
Protopenthalodes Jesionowska, 1989 – 2 sp.
Stereotydeus Berlese, 1901 - 30 sp.
Penthalodidae Sig Thor, 1933: 272
Type genus: Megamerus Dugès, 1834 (nec McLeay, 1827), renamed Penthalodes Murray, 1877
Callipenthalodes Qin, 1998: 221
Type species: Callipenthalodes pennyae Qin, 1998
Callipenthalodes pennyae Qin, 1998: 224
Type locality: New Zealand
Distribution: New Zealand
Hawaiieupodes Strandtmann & Goff, 1978: 133
Type species: Hawaiieupodes thermophilus Strandtmann et Goff, 1978
Hawaiieupodes thermophilus Strandtmann & Goff, 1978: 133; Jesionovska, 2010: 337
Type locality: Hawaii I., HVNP, Steaming Bluff Trail
Distribution: Hawaii
Megamerus Dugès, 1834: 50 (nec McLeay, 1827)
Penthalodes Murray, 1877: 123
Type species: Megamerus ovalis Dugès, 1834
Penthalodes alaskaensis Jesionowska, 2010: 31
Type locality: Alaska
Distribution: USA (Alaska)
Penthalodes boneti Baker, 1946: 424
Type locality: Desierto de los Leones, Vera Cruz
Distribution: Mexico
Penthalodes carinatus Shiba, 1978: 150
Type locality: Kuroki, Oozu, Ehime
Distribution: Japan
Chromotydaeus columbiae Berlese, 1921: 193
Penthalodes columbiae : Thor & Willmann, 1941: 193; Strandtnann, 1979: 48
Type locality: “Columbia”
Distribution: North America
Penthalodes ovalis sensu Strandtmann & Goff, 1978: 121
Penthalodes hawaiiensis Jesionowska, 2010: 32
Type locality: Hawaii
Distribution: Hawaii
Penthalodes oregonensis Baker, 1946: 424
Type locality: Corvallis
Distribution: USA (Oregon)
Megamerus ovalis Dugès, 1834: 50; 1836: 518; Lucas, 1840: 472; 1850: 472
Trombidium ovale Gervais, 1844: 170
Penthaleus ovatus : C.L. Koch, fa. 18, t.14; Halbert, 1915: 111; Starkoff & Starkoff, 1950: 98
Penthalodes ovalis (Dugès): Oudemans, 1905: 241; 1937: 861; Irk, 1939; Sellnick, 1940: 32; Thor & Willmann, 1941: 60; Willmann, 1951a: 147; 1951b: 142; 1954: 235; Macfadyen, 1954: 283; Bottazzi, 1950: 38; Haarløv, 1957: 30; Mihelčič, 1958a: 104; 1958b: 270; Strandtmann, 1971: 92; Strandtmann & Goff, 1978: 135; Schmölzer, 1962: 283; 1993: 59; Kaluz, 2000: 477
Chromotydaeus ovalis : Berlese, 1903: 252
Chromotydaeus ovatus (Koch): Sig Thor, 1912: 237; Halbert, 1920: 139
Penthaleus arcticus Trägårdh, 1907: 18
Chromotydaeus arcticus (Trägårdh): Summerhayes & Elton, 1928: 249
Type locality: Europe
Distribution: Alaska, Austria, Canada, Czech Rep., Denmark, Hawaii, Iceland, Ireland (ovatus), Italy, Slovakia, Spain, Svalbard, Turkey, Yan Mayen I.
Penthalodes ovasimilis Schweizer, 1951: 62
Type locality: Swiss Alps, many loc. at 2200 – 2850 m
Distribution: Switzerland
Penthalodes polonicus Jesionowska, 2010: 32; Hajizadeh & Noei, 2012; Darbemamieh, Hajiqanbar & Khanjani, 2013: 455
Type locality: Zakopane, The Tatra Mountains Nat. Park (Poland)
Distribution: Iran, Poland
Penthaleus tristiculus Koch & Berendt, 1854: 108
Penthalodes tristiculus (Koch et Berendt): Dunlop et al., 2013
Type locality: amber
Distribution: amber
Penthalodes turneri Baker, 1946: 425
Type locality: Stoddard County, Mo.
Distribution: USA (Missouri, South Carolina, Texas)
Protopenthalodes Jesionowska, 1989: 57
Type species: Protopenthalodes Jesionowska, 1989
Protopenthalodes coniunctus Jesionowska, 1989: 59
Type locality: Poland
Distribution: Poland
Protopenthalodes reticulatus Khaustov, 2015: 139
Type locality: Yalta mountain – forest Nature Reserve
Distribution: Crimea
Stereotydeus Berlese, in Berlese & Leonardi, 1901: 14
Type species: Stereotydeus notophalloides Leonardi, in Berlese & Leonardi 1901
Tectopenthalodes Trägårdh, 1907: 11 (syn. by Womersley, 1941)
Type species: Penthaleus villosus Trouessart, 1902
Stereotydeus areolatus Womersley, 1935: 79; Thor & Willmann, 1941: 70; Strandtmann, 1979: 49; Qin, 1994: 1308; 1998: 299; Fredes, 2015: 33
Type locality: South Australia (Adelaide)
Distribution: Australia, Argentina
Stereotydeus australicus Thor, 1934: 65; Thor & Willmann, 1941: 69; Strandtmann, 1979: 49; Qin, 1994: 1310; 1998: 299
Type locality: Western Australia, Crawley
Distribution: Australia
Penthaleus belli Trouessart, 1902: 225
Chromotydaeus belli : Thor & Willmann, 1941: 66
Stereotydeus belli (Trouessart): Womersley & Strandtmann, 1863: 458; Strandtmann, 1979: 49
Type locality: North Victoria Land
Distribution: Antarctica
Stereotydeus brevipalpus Qin, 1994: 1313; 1998: 299
Type locality: South Australia
Distribution: Australia
Stereotydeus capensis Womersley, 1935: 81
Type locality: Cape Province
Distribution: South Africa
Stereotydeus delicatus Strandtmann, 1967: 63
Type locality: Balleny Is: Sabrina I.
Distribution: Antarctica
Stereotydeus gamasoides Leonardi, 1901: 15
Type locality: Temuco
Distribution: Chile
Stereotydeus ineffabilis Brunetti & Siepel, 2021: 118
Type locality: Inexpressible Island
Distribution: Central Victoria Land, Continental Antarctica
Penthalodes intermedius Trouessart, 1907: 35
Chromotydaeus intermedius (Trt.): Thor & Willmann, 1941: 66
Stereotydeus intermedius (Trouessart): Strandtmann, 1979: 50
Type locality: South Orkney Islands, Laurie Island
Distribution: Antarctica
Stereotydeus lattimorei Baker, 1946: 423
Type locality: Brownsville, Tx, from orchid roots from the Canal Zone
Distribution: Panama
Stereotydeus longipes Strandtmann, 1970: 94
Type locality: South Georgia, Bird I., Wanderer Valley
Distribution: South Georgia (Subantarctic)
Stereotydeus mexicanus Baker, 1946: 422
Type locality: Patzcuaro, Michoacán, Mt. Popocatepetl (at 3000 m)
Distribution: Mexico
Stereotydeus meyeri Strandtmann, 1967: 63
Type locality: Enderby Land: Molodozhnaya
Distribution: Antarctica
Stereotydeus mollis Womersley & Strandtmann, 1963: 453; Wise, 1964: 522; Pittard, Robers & Strandtmann, 1971: 88
Type locality: Observation Hill, Ross I.
Distribution: Antarctica, southern Victoria Land from about 74° to 79° S lat., Balleny Islands
Stereotydeus notophalloides Leonardi, in Berlese & Leonardi, 1901: 15; Thor & Willmann, 1941: 69
Type locality: Temuco
Distribution: Chile
Stereotydeus nudisetatus Strandtmann, 1964: 152; Qin, 1998: 299
Type locality: Campbell Island
Distribution: New Zealand (Campbell Island)
Stereotydeus nunatakensis Brunetti, 2021: 124
Type locality: Starr Nunatak
Distribution: South Victoria Land, Continental Antarctica
Stereotydeus occidentale Womersley, 1935: 80; Strandtmann, 1964: 49; 1979: 49
Stereotydeus occidentalis Womersley: Thor & Willmann, 1941: 71; Qin, 1994: 1315; 1998: 299
Type locality: South Australia
Distribution: Australia
Stereotydeus oregonensis Baker, 1946: 424
Type locality: Oregon, Corvallis
Distribution: USA
Stereotydeus pallotus Olivier, 2006: 55
Type locality: Eastern Cape Province: soil, Hogsback near Alice
Distribution: South Africa
Stereotydeus pseudopulcher Qin, 1994: 1318; 1998: 299
Type locality: Tasmania
Distribution: Australia (Tasmania)
Stereotydeus pulcher Strandtmann, 1964: 154; Qin, 1998: 299
Type locality: Campbell Island
Distribution: New Zealand (Campbell Island)
Stereotydeus punctatus Strandtmann, 1967: 62
Type locality: North Victoria Land: W side of Islands Point
Distribution: Antarctica
Stereotydeus reticulatus Strandtmann, 1970: 90
Type locality: South Georgia, Bird I., Macaroni Coulm, 200-230 m
Distribution: South Georgia
Stereotydeus rupicolamartimus Shiba, 1976: 97
Type locality: Malaysia
Distribution: Malaysia
Stereotydeus shoupi Strandtmann, 1967: 60
Type locality: Queen Maud Mts: Shackleton Glacier area: E side of Massam Glacier
Distribution: Antarctica
Stereotydeus tasmanicus Qin, 1994: 1320; 1998: 299
Type locality: Tasmania
Distribution: Australia (Tasmania)
Stereotydeus undulatus Strandtmann, 1964: 150; Qin, 1998: 299
Type locality: Campbell Island
Distribution: New Zealand (Campbell Island)
Stereotydeus vaginatus Olivier, 2006: 55
Type locality: KwaZulu-Natal: soil, mouth of Tugela River
Distribution: South Africa
Penthaleus villosus Trouessart, 1902: 227
Penthalodes villosus , Trouessart, 1912: 86
Tectopenthalodes villosus (Trouessart): Trägårdh, 1907: 18; Thor & Willmann, 1941: 71
Stereotydeus (Tectopenthalodes) villosus: Berlese, 1917: 8
Stereotydeus villosus : Womersley & Strandtmann, 1963: 457; Strandtmann, 1979: 50
Type locality: South Shetland Islands, South Orkney Islands, Antarctic Peninsula
Distribution: Antarctica
EUROPE
Austria (Irk, 1939; Franz Beier, 1948; Franz, 1954; Willmann, 1951a; Mihelčić, 1958; Schmölzer, 1962, 1993) – Penthalodes ovalis
Bulgaria (Beron, 2011) - Penthalodes ovalis
Czech Rep. (Willmann, 1954) – Penthalodes ovalis
Denmark (Haarløv, 1957) - Penthalodes ovalis
Yan Mayen I. (Macfadyen, 1954) – Penthalodes ovalis
France (contin.) (Dugès, 1834; Kontschan et al., 2018) – Penthalodes ovalis, Inc. sedis: Penthalodes inflatus
Great Britain (Hull, 1918) – Penthalodes ovalis
Iceland (Sellnick, 1940) – Penthalodes ovalis
Ireland (Halbert, 1915) – Penthalodes ovalis
Italy (contin.) (Thor & Willmann, 1941; Bottazzi, 1950) – Penthalodes ovalis
Norway (Mehl, 1979) – Penthalodes ovalis
Svalbard (Spitzbergen)(Thor & Willmann, 1941) – Penthalodes ovalis
Poland (Jesionowska, 1989, 2010) – Penthalodes polonicus, Protopenthalodes coniunctus
Russia (all, continent.)
Crimea (Khaustov, 1915) – Protopenthalodes reticulatus
Slovakia (Kalúz, 2000) – Penthalodes ovalis
Spain (cont.)(Mihelčič, 1958b) - Penthalodes ovalis
Sweden (Strandtnann, 1979) – Penthalodes ovalis
Switzerland (Schweizer, 1951; Schweizer & Bader, 1963) - Penthalodes ovalis
Turkey (all)(Kalúz, 2000) - Penthalodes ovalis, P. ovasimilis
AFRICA
South Africa (Womersley, 1935; Olivier, 2006) – Stereotydeus pallotus, S. vaginatus
ASIA
Iran (Hajizadeh & Noei, 2012; Hajizadeh & Noei, 2012; Darbemamieh, Hajiqanbar & Khanjani, 2013) – Penthalodes polonicus
Japan (Shiba, 1978) – Penthalodes carinatus
Malaysia (Shiba, 1976) - Stereotydeus rupicolamartimus
NORTH AMERICA
Canada (Strandtmann, 1979; Behan, 1978; Beaulieu et al., 2019) – Penthalodes ovalis
Mexico (Baker, 1946) - Penthalodes boneti
USA (Baker, 1946) – Penthalodes oregonensis, P. turneri
Alaska (Strandtmann, 1971; Jesionowska, 2010) – Penthalodes alascaensis, P. ovalis
Hawaii (Strandtmann & Goff, 1978; Goff, 1983, 1987; Jesionowska, 2008, 2010) – Hawaiieupodes termophilus, Penthalodes hawaiiensis, P. ovalis
SOUTH and CENTRAL AMERICA
Argentina (Fredes, 2015) – Stereotydeus areolatus
Chile (contin.) (Leonardi, 1901, 1903) – Stereotydeus notophalloides, S. gamasoides
Panama (Baker, 1946) - Stereotydeus lattimorei
OCEANIA
Australia (continent)(Womersley, 1935; Qin, 1994) – Stereotydeus areolatus, S. australicus, S. brevipalpus, S. occidentalis
Tasmania (Qin, 1994) – Stereotydeus pseudopulcher, S. tasmanicus
New Zealand (Qin, 1998) - Callipenthalodes pennyae
Campbell Island (Strandtmann, 1964) – Stereotydeus nudisetatus, S. pulcher, S. undulatus
ANTARCTICA
Antarctida (Trouessart, 1902; Womersley & Strandtmann, 1963; Strandmann, 1967; Wise, 1964; Pittard, 1971; Pittard, Roberts & Strandtmann, 1971; Brunetti et al., 2021a, b) – Stereotydeus belli, S. delicates, S. meyeri, S. mollis, S. nunatakensis, S. punctatus, S. shoupi, S. villosus
South Georgia (Strandtmann, 1970) – Stereotydeus longipes, S. reticulatus
South Shetland Islands – Stereotydeus villosus
South Orkney Islands (Trouessart, 1912) - Stereotydeus villosus
Inexpressible Island (Brunetti et al., 2021a) – Stereotydeus ineffabilis
Penthaleidae Oudemans, 1931: 252; Sig Thor, 1933: 2
Middle sized (800 – 1000 µm) mites, black with red legs, living on plants and sometimes pests on some agricultural crops.
Family Penthaleidae - 7 gen., 27 sp.
Chromotydaeus Berlese, 1903 - 3 sp.
Halotydeus Berlese, 1891 - 8 sp.
Linopenthaleus Willmann, 1951 - 1 sp.
Linopenthaloides Strandtmann, 1981- 1 sp.
Penthaleus C.L. Koch, 1836 - 12 sp.
Propenthalodes Jesionowska, 1989 – 1 sp.
Turanopenthalodes Barilo, 1988 – 1 sp.
Penthaleidae Oudemans, 1931: 252
Type genus: Penthaleus C.L. Koch, 1836
Chromotydaeus Berlese, 1903: 252
Type species: Penthaleus ovalis (Dugès, 1834)
Penthaleus anauniensis R. Canestrini, 1886: 720
Chromotydaeus anauniensis (R. Canestrini): Thor & Willmann, 1941: 65
Type locality: Trient, Dosso di Tavon
Distribution: Italy
Penthaleus egregius Berlese, 1891(1888 ?): 3; Starkoff & Starkoff, 1950: 98
Chromotydaeus egregius (Berlese): Berlese, 1903: 252
Type locality: Italy
Distribution: Italy
Penthaleus illustris Berlese, 1916: 294; Bernini et al., 1994-96: 34
Chromotydaeus illustris (Berlese): Thor & Willmann, 1941: 67
Type locality: Garden “Boboli”, Florence
Distribution: Italy
Chromotydaeus quartus Qin & Halliday, 1996: 1834
Type locality: Western Australia
Distribution: Australia
Chromotydaeus similis Mihelčič, 1958a: 39
Type locality: Sierra Nevada
Distribution: Spain
NB. Linopodes globosus Canestrini, 1884 was listed under Chromotydeus by Thor & Willmann, 1941: 65, but under line was noted that it could belong also to the [still undescribed] genus Linopenthaleus.
Halotydeus Berlese, 1891: 9
Type species: Notophallus hydrodromus Berlese et Trouessart, 1889
Halotydeus anthropus Qin & Halliday, 1996: 442
Type locality: South Africa
Distribution: South Africa
Halotydeus egregius (Berlese, 1891) [syn. Baker 1996:373]
Halotydeus bakerae Qin & Halliday, 1996: 445
Type locality: South Australia
Distribution: Australia (South Australia)
Halotydeus castellus Qin & Halliday, 1996: 445
Type locality: New-South-Wales
Distribution: Australia (New-South-Wales)
Penthaleus destructor R.W. Jack, 1908; Tucker, 1925: 21
Halotydeus destructor (Tucker): Womersley, 1933: 108; Thor & Willmann, 1941: 91; Macfadyen, 1954: 283; Meyer & Ryke, 1960: 489. Redescription, Qin & Halliday 1996: 447; Robinson & Hoffmann, 2001: 699; Maino, Binns & Umina, 2018: 216
Type locality: South Africa
Distribution: Australia, South Africa, Yan Mayen I. (? introduced)
Notophallus hydrodromus Berlese & Trouessart, 1889: 141
Halotydeus hydrodromus (Berlese & Trouessart): Berlese, 1891: Fasc. 60, No10; Halbert, 1915: 111 (var. albolineatus), 1920: 139; Hull, 1918: 33; Thor & Willmann, 1941: 90
Type locality: France
Distribution: England, France, Ireland, Italy
Halotydeus mollis Luxton, 1990: 668
Type locality: Hong Kong, marine
Distribution: China (Hong Kong)
Halotydeus signiensis Strandtmann & Tilbrook, 1968: 54
Type locality: Robin Peak, Signy Island
Distribution: South Orkney Islands (Signy I.)
Halotydeus spectatus Qin & Halliday, 1996: 449
Type locality: New-South-Wales
Distribution: Australia (New-South-Wales)
Genus Linopenthaleus Willmann, 1951a: 159
Type species: Linopenthaleus irki Willmann, 1951
Linopenthaleus irki Willmann, 1951a: 160; Schweizer & Bader, 1963: 220
Type locality: Grossglockner area (Austria)
Distribution: Austria, Switzerland
Linopenthaloides Strandtmann, 1981: 392
Type species: Linopenthaloides novazealandicus Strandtmann, 1981
Linopenthaloides novazealandicus Strandtmann, 1981: 392
Type locality: N.Z., South I.: W coast, Charleton, ex Metro Cave, Ananui catchement
Distribution: New Zealand
Penthaleus C.L. Koch, 1836: fa.1, t.12 and 13
Type species: Penthaleus haematopus C.L. Koch, 1836 = P. maior (Dugès)
Notophallus Canestrini, 1886: 697
Penthaleus arcticus Trägårdh, 1901: 19
Type locality: Bären Island
Distribution: Norway
Trombidium bipustulatum Hermann, 1804: 40
Penthaleus bipustulatus (Hermann): C.L. Koch, 1838: fa.18, t.7; Macfadyen, 1954: 283
Penthaleus bipustulatum (Hermann): Oudemans, 1937: 852
Type locality: France
Distribution: France, Germany, Yan Mayen I.
Penthaleus crinitus Narayan, 1957: 111
Type locality: Mexico
Distribution: Mexico
Notophallus dorsalis Banks, 1902: 172; Considered synonym of Penthaleus major by Baker (1946: 421).
Penthaleus dorsalis : Bruce, 2021
Type locality: USA
Distribution: USA
Penthaleus erythrocephalus C.L. Koch, 1838: fa.18, t.8; Mihelčič, 1958a: 40; 1958b: 269
Type locality: Germany
Distribution: Germany, Spain
Penthaleus falcatus Qin & Halliday, 1996: 1837; Halliday, 1998: 77; Robinson & Hoffmann, 2001: 699
Type locality: New South Wales
Distribution: Australia
Megamerus fallax Dugès, 1834: 50, 53
Penthaleus fallax (Dugès): Oudemans, 1937: 854
Type locality: France
Distribution: France
Penthaleus guttatus C.L. Koch: fasc. 18, nr. 13; Oudemans, 1937: 855; Thor & Willmann, 1941: 83
Type locality: Regensburg
Distribution: Germany
Notophallus longipilis Canestrini, 1886: 706
Penthaleus longipilis (Canestrini): Shiba, 1969: 71
Type locality: Italy
Distribution: England, Italy, Japan
Tétranyque majeur Dugès, 1834: 53 (nom. nud.)
Rhyncholophus major : Murray, 1877: 126
Penthaleus haematopus C.L. Koch, 1836: fasc. 1, nr.12; Marchal, 1908: 13; Thor, 1912: 237; Hull, 1918: 33
Penthaleus erythropus C.L. Koch, 1838: fasc. 18, nr.6
Penthaleus insulanus Thorell, 1872: 702
Tetranychus borealis L. Koch, 1879: 129
Notophallus insulanus : Trägårdh, 1900: 16
Notophallus haematopus : R. Canestrini, 1886: 704
Notophallus bicolor Froggatt, 1921: 33
Penthaleus bicolor Womersley, 1933: 110
Penthaleus major (Dugès): Meyer & Ryke, 1960: 491; Balevski & Kontev, 1969; Rekk, 1976: 49; Zhang & Hong, 2010: 6
Penthaleus maior (Dugès): Sellnick, 1940: 32; Thor & Willmann, 1941: 80; Schmölzer, 1993: 59; Halliday, 1998: 77; Darbemamieh, Hajiqanbar & Khanjani, 2013: 455
Type locality: Europe
Distribution: Argentina, Australia, Austria, Brazil, Bulgaria, Canada, China, France, Georgia, Germany, Great Britain, Greece, Holland, Hungary (cf. P. major), Iran, Iceland, Italy, Mexico, New Zealand, Norway, Poland, Portugal, Russia, Slovakia, Slovenia, South Africa, Spain, USA
Notophallus minor Canestrini, 1886: 705; Berlese, 1892: fasc. 61, No5
Penthaleus minor (Canestrini): Oudemans, 1905: 241; Thor & Willmann, 1941: 85; Qin & Halliday, 1996; Halliday, 1998: 77; Zhang & Hong, 2010: 6; Darbemamieh, Hajiqanbar & Khanjani, 2013: 455
Type locality: Padua (Italy)
Distribution: Australia, China, Great Britain, Germany, Iran, Italy, Switzerland
Penthaleus saxatilis C.L. Koch, 1836: fa.1, t.13; Thor & Willmann, 1941: 81
Type locality: Regensburg
Distribution: Germany
Penthaleus tectus Halloway, 2005b: 144
Type locality: East Australia
Distribution: Australia
Turanopenthalodes Barilo, 1988: 619
Type species: Turanopenthalodes polytrichus Barilo, 1988
Turanopenthalodes polytrichus Barilo, 1988: 619; Khaustov, 2016: 674
Type locality: Samarkand, settl. Agalyk
Distribution: Uzbekistan
As species with uncertain affinity are listed by Thor & Willmann, 1941 or described before or after this monograph: Penthaleus nemoralis (Müller) (Acarus n.), P. agilis Berlese, 1905, P. hermanni Sig Thor, Notophallus viridis Banks, 1917 (synonymized with Penthaleus major by Baker (1946: 421), Penthaleus guayavicola (Stoll)(Trombidium g.), P. trilineatus (Stoll)(Trombidium g.), P. albicollis (Stoll)(Trombidium a.). Here could be added also some species, described by C.L. Koch (1838): Penthaleus amictus, P. militaris, P. virellus, P. rhodomelas and the described by Ludvig Koch from Novaja Zemlja Penthaleus borealis, P. crassipes.
EUROPA
Austria (Irk, 1942; Willmann, 1951a, 1951b; Mihelčič, 1958c; Schmölzer, 1993) – Linopenthaleus irki, Penthaleus bipustulatus, P. major
Bulgaria (Balevski & Kontev, 1969; Stoeva, 2006; Atanasov, Marinova & Atanasova, 2008; Maneva & Atanasova, 2016) - Penthaleus major
Denmark (Müller O.F. 1776) - ? Penthaleus nemoralis
Yan Mayen I. (Macfadyen, 1954) - Penthaleus bipustulatus
France (contin.)(Hermann, 1804; Dugès, 1834; Qin & Halliday, 1996) – Penthaleus bipustulatus, P. fallax, P. major
Georgia (Rekk, 1976) - Penthaleus major
Germany (C.L. Koch, 1835, 1836, 1838; Qin & Halliday, 1996) – Penthaleus bipustulatus, P. erythrocephalus, P. major, P. saxatilis
Great Britain (Hull, 1918; Wallace & Mahon, 1971) - Halotydeus hydrodromus, Penthaleus longipilis, P. major
Greece (Hadzinicolis, 1986) - Penthaleus major
Holland (Wallace & Mahon, 1971) - Penthaleus major
Hungary (Kontschán et al., 2018) - Penthaleus cf. major
Iceland (Sellnick, 1940; Gudleifsson et al., 2002; Hallas & Gudleifsson, 2005) – Penthaleus major
Ireland (Halbert, 1915, 1920) - Halotydeus hydrodromus, Penthaleus major
Italy (contin.)(Canestrini, 1886, Berlese, 1916; Bernini F., M. Castagnoli, R. Nannelli. 1994-1996. Checklist delle specie de la fauna italiana. Fascicolo 24 Arachnidi Acari. – Bologna, 132 pp. Wallace & Mahon, 1971) – Chromotydaeus anauniensis, Ch. illustris, Penthaleus longipilis, P. major, P. minor
Norway (Thor, 1912; Johansen & Haug, 2002) – Penthaleus major (= haematopus)
Poland (Kontschán et al., 2018) - Penthaleus major
Portugal (cont.)(Wallace & Mahon, 1971; Bernini et al., 1994–96) - Penthaleus major
Russia (all, continent.) (L. Koch, 1879; Stuxberg, 1886; Kornilov & Kositsyna, 1981) – [Penthaleus borealis, P. crassipes], Penthaleus major
Slovenia (Devetak et al., 2019) - Penthaleus major
Spain (cont.)(Mihelčič, 1958a, 1958b; Wallace & Mahon, 1971) – Chromotydaeus similis, Penthaleus major
Switzerland (Schweizer & Bader, 1963) – Linopenthaleus irki, Penthaleus longipilis, P. major, P. minor
AFRICA
South Africa (Jack, 1908; Tucker, 1925; Meyer & Ryke, 1960; Qin & Halliday, 1996; Halliday, 2005) – Halotydeus anthropus, H. destructor, Penthaleus major
ASIA
China (Narayan, 1962; Zhang & Hong, 2010: 6) - Penthaleus major, P. minor
Iran (Darbemamieh, Hajiqanbar & Khanjani, 2013) - Penthaleus major, P. minor
Japan (Shiba, 1969; Kanda & Hirai, 1990) – Penthaleus longipilis, P. major
Uzbekistan (Barilo, 1988; Khaustov, 2016) - Turanopenthalodes polytrichus
NORTH AMERICA
Canada (Behan, 1978; Qin & Halliday, 1996; Beaulieu, 2019) – Penthaleus major,
Mexico (Baker, 1946; Narayan, 1957) – Penthaleus major, Stereotydeus mexicanus
USA (Banks, 1902; Ewing, 1910; Brust, 2018, 2021a, b) – Penthaleus dorsalis, P. major
SOUTH and CENTRAL AMERICA
Argentina (Qin & Halliday, 1996) - Penthaleus major
Brazil (Valle da Silva Pereira et al., 2017) - Penthaleus major
Guatemala (Stoll, 1886 - 1893) - ? Penthaleus trilineatus, ? Penthaleus albicollis
OCEANIA
Australia (continent)(Newman, 1923, 1924, 1925a, 1925b, 1936; Swan, 1934; Womersley, 1941; Wallace, 1970; Wallace & Mahon, 1971; Halliday, 1991, 1998, 2005b; Qin & Halliday, 1995, 1996a, 1996b; Weeks, Fripp & Hoffmann, 1995; Qin, 1998; Weeks & Hoffmann, 1999; James, 1987, 2000; Robinson & Hoffmann, 2001; Umina & Hoffmann, 2003, 2004; Umina, Hoffmann & Weeks, 2004) – Chromotydaeus quartus, Halotydeus bakerae, H. castellus, H. destructor, H. spectatus, Penthaleus major, P. minor, P. tectus
New Zealand (Strandtmann, 1981; Wallace & Mahon, 1971) – Linopenthaloides novazealandicus, Penthaleus major
ANTARCTICA
South Orkney Islands (Strandtmann & Tilbrook, 1968) – Halotydeus signiensis
Proterorhagiidae Lindquist & Palacios-Vargas, 1991: 341
Family containing only one species from Mexico.
Proterorhagiidae Lindquist & Palacios-Vargas, 1991: 341
Type genus: Proterorhagia Lindquist et Palacios-Vargas, 1991
Proterorhagia Lindquist & Palacios-Vargas, 1991: 341
Type species: Proterorhagia oztotloica Lindquist et Palacios-Vargas, 1991
Proterorhagia oztotloica Lindquist & Palacios-Vargas, 1991: 344
Type locality: border area between the states of Colima and Jalisco, Pozo La Escondida near Lagunitas Village, 2200 m
Distribution: Mexico
NORTH AMERICA
Mexico (Lindquist & Palacios-Vargas, 1991) – Proterorhagia oztotloica
Strandtmanniidae Zacharda, 1979: 77
Monotypical family known from the Chech Republic.
Strandtmanniidae Zacharda, 1979: 77
Type genus: Strandtmannia Zacharda, 1979
Pilorhagidia Strandtmann & Goff, 1978: 141
Strandtmannia Zacharda, 1979: 78
Type species: Pilorhagidia hirsuta Strandtmann et Goff, 1978
Strandtmannia celtarum Zacharda, 1979: 78
Pilorhagidia celtarum (Zacharda):
Type locality: Bohemia centr., Zavist-knoll near Zbraslav
Distribution: Czech Rep.
Pilorhagidia hirsuta Strandtmann & Goff, 1978: 141
Type locality: Hawaii I: HVNP, Mauna Loa Trail, 2040 rn
Distribution: Hawaii Islands
EUROPE
Czech Rep. (Zacharda, 1979) - Pilorhagidia celtarum
OCEANIA
Hawaii Islands (Strandtmann & Goff, 1978) – Pilorhagidia hirsuta
Tydeoidea Cunliffe, 1955
The Superfamily contains the families Ereynetidae, Iolinidae, Triophtydeidae and Tydaeidae.
The family Ereynetidae contains three subfamilies: Ereynetinae (free living in the soil), Speleognathinae (parasites in the respiratory passages of molluscs, birds and mammals) and Lawrencarinae (parasites in the respiratory passages of frogs and toads). Until 1936 only the Ereynetinae were known, described by several early acarologists, starting with Schrank (1776) and C.L. Koch (1836). In 1883 A. Berlese described the genus Ereynetes and in 1931 Oudemans created the family Ereynetidae. Some other specialists who contributed to the study of Ereynetinae are Vitzthum (1931), André & Lamy (1930, 1931), Thor (1932a, 1932b, 1936), E.W. Baker (1945), Turk & Phillips (1945), R.A. Baker (1970a, b), Cooreman (1947), Willmann (1936, 1949, 1953), Mihelčic (1958), Hunter (1964), Corpuz-Raros (1975), Shiba (1976), Zabludovskaya (1994) and again Berlese (1910, 1923). Many publications were devoted to the peculiar mites parasitizing the snails and slugs (Riccardoella). The modern arrangement of this interesting group is due to A. Fain and his co-authors J. L. van Goethem, A.M. Camerik and G.M. Barker.
The mighty presence of A. Fain in the study of Ereynetidae began in 1955. Firstly he submitted part of his collection from Rwanda to J. Cooreman, who described the first African member of the family as new genus and species Speleognathopsis galli Cooreman, 1955. This was the end of the pre-Fain era in the study of Ereynetidae. Only from the present day species have been known by this time from the parasitic subfamilies Speleognathinae and Lawrencarinae, and they were wrongly positioned in the system. In 1936 H. Womersley described the peculiar mite Speleognathus australis, for which he created the new family (now subfamily) Speleognathidae. Followed E. Boyd with “Speleognathus” sturni Boyd, 1948, the first among the many nasal parasites of birds and mammals we know now. Womersley (1953) named this mite Boydaia sturni. Meanwhile Lawrence (1952) described the first mite endoparasite of Anura, but named it Riccardoella eweri. Followed several other parasites of frogs and toads, now 12 species in the genera Lawrencarus Fain, 1957, Batracarus Fain, 1961 and Xenopacarus Fain, Baker & Tinsley, 1969. Fain (1962) summarized the data on the mites parasitizing Anura. Kethley (1971) described one new genus (Hydranetes) with two new species of Ereynetid mites, living under elytrae of Hydrophilid beetles in Florida.
The rich collection of endoparasitic mites, brought by Fain from Rwanda and Congo, was the basis for the many descriptions of new taxa in the subfamily Speleognathinae. The mites of this subfamily in the Afrotropical Region have been reviewed by Skoracki et al. (2018).
Very few people interfered in the stream of papers by Fain and co-authors (Aitken, Bafort, Tinsley, Baker, Barker, Cameric, Goff, Herin, Puylaert, Hyland, Klompen, Lukoschus, Mortelmans, Nadchatram, Prasse, van Goethem, Vercammen-Grandjean, Wagner, Flechtmann, Leal). Dubinin (1957) was the author of the new genus Ophthalmognathus. Other specialists describing Speleognathinae were Crossley (1952), Womersley (1936, 1953), Van Eyndhoven (1955), Ford (1959, 1962), Clark (1958, 1960, 1964, 1967a, 1967b), Domrow (1965, 1969, 1991), Pence (1973), Pence & Casto (1976), OConnor (1978).
The family Ereynetidae contains now (end 2021) 180 species of 35 genera and three subfamilies. The progress since the first general paper on the family (Thor, 1933) is obvious: by this time only 15 species of 3 (actually two – Ereynetes and Riccardoella) genera have been recognized in Ereynetidae. Most species are endoparasites.
Fam. Ereynetidae – 35 genera, 180 sp.
Subfam. Ereynetinae – 78 sp.
Tribe Austrereynetini
Austrereynetes Fain et Barker, 2003 – 1 sp.
Tribe Ereynetini
Ereynetes Berlese, 1883 – 55 sp.
Anereynetes Fain, 1964 - 24
Ereynetes Berlese, 1883 - 21
Gymnereynetes Fain, 1964 - 4
Huntereynetes Fain, 1964 - 3
Opsereynetes Thor, 1932 - 3
Hanriccardoella André, Ducarme et Lebrun, 2004 - 1
Hydranetes Kethley, 1971 - 2
Protereunetes Berlese, 1923 - 9
Pseudotydeus Baker et Delfinado, 1974 - 2
Riccardoella Berlese, 1923 - 8
Proriccardoella Fain et van Goethem, 1986 - 6
Riccardoella Berlese, 1923 – 2
Subfam. Speleognathinae - 90 sp.
Tribe Boydaiaini
Boydaia Womersley, 1953 - 41
Coboydaia Fain, 1971 - 5
Coboydaia (Apodiboydaia) Fain, 1985 - 1
Coboydaia (Coboydaia) Fain, 1971 - 4
Tribe Trispeleognathini
Astrida Fain, 1955 - 4
Astrida Fain, 1955 - 2
Cerylonyssus Fain et Aitken, 1970 – 1
Aureliania Fain, 1958 – 1
Meropiboydaia Fain, 1985 - 1
Metaboydaia Fain, 1962 - 1
Neastrida Fain, 1962 - 1
Neoboydaia Fain, 1958 - 3
Ophthalmognathus Dubinin, 1957 - 2
Phoenicopteriella Fain, 1970 - 2
Picinyssus Fain, 1969 - 1
Psittaboydaia Fain, 1985 - 2
Pterniboydaia Fain, 1985 - 1
Trispeleognathus Fain, 1958 -
Tribe Paraspeleognathopsini
Paraspeleognathopsis Fain, 1958 - 4
Tribe Speleochirini
Hipposideroptes Fain, 1970 - 1
Neospeleognathopsis Fain, 1958 – 3
Neospeleognathopsis (Neospeleognathopsis) Fain, 1958 – 1
Neospeleognathopsis (Speleomyotis) - 1
Neospeleognathopsis (Pteropignathus) - 1
Ralliboydaia Fain, 1962 – 3
Speleochir Fain, 1966 - 5
Neospeleochir Fain, 19 - 1
Speleognathopsis Cooreman, 1954 - 6
Speleomys Fain, 1970 – 1
Speleorodens Fain, 1962 - 6
Tribe Speleognathini
Domrownetes OConnor, 1978 - 1
Speleognathus Womersley, 1936 – 4
Subfamily Lawrencarinae Fain, 1957 - 12 sp.
Tribe Lawrencarini Fain
Batracarus Fain, 1961 - 1
Lawrencarus Fain, 1957 - 8
Tribe Xenopacarini Fain, Baker et Tinsley, 1969
Xenopacarus Fain, Baker et Tinsley, 1969 - 3
Ereynetidae Oudemans, 1931: 253
Type genus: Ereynetes Berlese, 1883
Speleognathidae Womersley, 1936 (syn. by Fain, 1957: 250)
Type genus: Speleognathus Fain, 1936: 313
Microereunetidae Bottazzi, 1950: 38; syn. by Fain, 1954a: 110
Type genus: Microereunetes Berlese, 1924: syn. by Fain, 1954a: 110
Ereynetinae Fain, 1957: 250
Type genus: Ereynetes Berlese, 1883
Pseudotydeinae Baker & Delfinado, 1974: 444 – syn. by André & Ducarme, 2003: 373
Type genus: Pseudotydeus Baker et Delfinado, 1974
Austrereynetini Fain & Barker, 2003: 233
Type genus: Austrereynetes Fain et Barker, 2003
Austrereynetes Fain & Barker, 2003: 234
Type species: Austrereynetes maudensis Fain et Barker, 2003
Austrereynetes maudensis Fain & Barker, 2003: 235
Type locality: Maud Island, Marlborough Sounds, New Zealand
Distribution: New Zealand
Host: Pseudaneitea schauinslandi (Gastropoda: Pulmonata: Athoracophoridae)
Type genus: Ereynetes Berlese, 1883
Ereynetes Berlese, 1883: 5.1
Ereunetes Berlese, 1883: fasc.V, 1, tav.22 (non Ereunetes Illiger, 1911: Aves)
Microereunetes Berlese, 1921: 24; 1923: 243
Type species: Tydeus polymitus C.L. Koch, 1836 [Ereynetes galeatus Berlese, 1883], see Fain, 1985b: 257
Neoereunetes Bottazzi, 1950: 44 (n.nov.), type sp. polymytis. Syn. by Fain, 1964a: 111
Ereynetoides Fain et Nadchatram, 1962a: 69: type sp. Ereynetoides malayi Fain et Nadchatram, 1962a: 72. Syn. by Fain, 1964a: 110
Ereynetes Berlese, 1883: 5.1; Fain, 1985b: 257
Type species: Tydeus polymitus C.L. Koch, 1836 [Ereynetes galeatus Berlese, 1883]
Micrereunetes (Protereunetes) agilis Berlese, 1923: 245; Fain, 1964a: 99
Ereynetes (Ereynetes) agilis Berlese: Fain, 1964a: 106; Fain & Camerik, 1994: 150
Protereunetes agilis (Berlese): Thor & Willmann, 1941: 54; Strandtmann, 1971: 82
Type locality: Palermo
Distribution: Sicily
Host: moss
Ereynetoides amplectorus Hunter, 1964: 86; Siahkamari et al., 2020:
Ereynetes (Ereynetes) amplectorus Hunter: Fain, 1964a: 110; Fain & Camerik, 1994: 156
Type locality: American Fork, Utah
Distribution: Mexico, USA (Utah), South Africa, Iran
Host: on celery
Micrereunetes arcuatus Berlese, 1923: 244
Ereynetes arcuatus (Berlese): Thor, 1933: 72
Ereynetes (Ereynetes) arcuatus Berlese: Fain, 1964a: 106
Type locality: Portici
Distribution: Italy
Ereynetes (Ereynetes) boharti Hunter & Cross, 1968: 156; Fain & Camerik, 1994: 151
Type locality: Shoshoni, Wyoming
Distribution: USA (Wyoming)
Host: from artificial nest of bee, Nomia melanderi (Hymenoptera, Halictidae)
Ereynetes (Ereynetes) bolitophilus Fain & Camerik, 1994: 156; Abdelgayed et al., 2019: 32
Type locality: Sandton (South Africa)
Distribution: South Africa, Egypt
Host: cow dung
Ereynetes cavernarum Shiba, 1971: 226
Type locality: Kaminari-ana Pit, Kansu-yama, Fujiwara, Susono-shi, Shizuoka Pref.
Distribution: Japan
Micrereunetes galeatus var. centralis Berlese, 1923: 243
Ereynetes centralis (Berlese): Thor, 1933: 74
Ereynetes (Ereynetes) galeatus centralis (Berlese): Fain, 1964a: 106
Neoereunetes centralis (Berlese): Bottazzi, 1950: 40
Type locality: Italy (“in agro Florentino”)
Distribution: Italy
Host: manure, humus, chestnut wood
Micrereunetes excellens Berlese, 1923: 243
Ereynetes excellens Berlese: Thor, 1933: 73
Ereynetes (Ereynetes) excellens Berlese: Fain, 1964a: 106
Type locality: Japan
Distribution: Japan
Ereynetoides faini Hunter, 1964: 188
Ereynetes (Ereynetes) faini Hunter: Fain, 1985b: 257; Fain & Camerik, 1994: 151
Type locality: Boston
Distribution: USA (on beet root from Denmark and lily bulbs from England), Mexico
Micrereunetes galeatus Berlese, 1923: 243
Acarus limacum , Berlese, 1883 (non Schrank, 1776)
Ereynetes polymitus , R. Canestrini, 1886 (non Koch, 1838)
Ereynetes ministralis , Oudemans, 1928: 382 (non Koch, 1838)
Micrereunetes polymitus , Berlese, 1920 (non Koch, 1838)
Ereynetes galeatus Berlese: Thor, 1933: 74
Ereynetes (Ereynetes) berlesei Oudemans, 1928: 382
Ereynetes (Ereynetes) galeatus (Berlese): Fain, 1964a: 106; Zabludovskaya, 1994: 35
Type locality: Palermo
Distribution: Sicily, Crimea
Host: in soil
Ereynetes (Ereynetes) gandensis Fain & Bafort, 1973: 409; Fain, 1985b: 257; Fain & Camerik, 1994: 151; Zabludovskaya, 1994: 35
Type locality: Gand (Belgium)
Distribution: Belgium, Ukraine
Host: in humus
Ereynetes (Ereynetes) husatoi Shiba, 1976: 139
Type locality: Malaya, Pasoh Forest Reserve
Distribution: Malaysia
Ereynetes (Ereynetes) luriei Fain & Camerik, 1994: 157
Type locality: Sandton
Distribution: South Africa
Host: cow dung
Ereynetoides malayi Fain & Nadchatram, 1962a: 72
Ereynetes (Ereynetes) malayi Fain & Nadchatram: Fain, 1964a: 106; 1985b: 257; Fain & Camerik, 1994: 150
Type locality: Fort Betis, Ulu Kelantan, Malaya
Distribution: Malaysia
Host: nasal cavity of Arachnothera l. longirostra (Passeriformes: Nectariniidae)
Ereynetes (Ereynetes) pegazzanoae Fain, 1964a: 95; Zabludovskaya, 1994: 35
Type locality: Tiarno (Italy)
Distribution: Italy, Ukraine
Host: in dung
Ereynetes (Ereynetes) philippinensis Corpuz-Raros, 1975: 326; Fain & Camerik, 1994: 151
Type locality: College, Laguna
Distribution: Philippines
Host: Scarabaeid beetle (Coleoptera)
Ereynetes (Ereynetes) punctatus Zabludovskaya, 1994: 37
Type locality: Migya Village, Pervomayskiy Rayon, Nikolaevskaya oblast
Distribution: Ukraine
Host: in dung
Micrereunetes galeatus var. septentrionalis Berlese, 1923: 244
Ereynetes septentrionalis (Berlese): Thor, 1933: 73; Çobanoglu et al., 2020:
Ereynetes (Ereynetes) galeatus septentrionalis (Berlese): Fain, 1964a: 106
Ereynetes (Ereynetes) septentrionalis (Berlese): Fain & Camerik, 1994: 150
Type locality: Nord Italy
Distribution: Italy, Turkey
Host: manure
Ereynetes (Ereynetes) tadakii Shiba, 1976: 136
Type locality: Malacca
Distribution: Malaysia
Ereynetes (Ereynetes) tshernogoriensis Zabludovskaya, 1994: 36
Type locality: Karpatskiy Zapovednik [Reserve], Tshernogorskiy Rayon, Transkarpathia; paratypes from Karadagskiy Zapovednik, Crimea
Distribution: Crimea, Ukraine
Ereynetes (Ereynetes) ueckermanni Fain & Camerik, 1994: 154
Type locality: Cluny Farm, Transvaal
Distribution: South Africa
Host: cow and horse dung
Ereynetoides watsoni Fain, 1962a: 925
Ereynetes (Ereynetes) watsoni (Fain): Fain, 1964a: 106; 1985b: 257; Fain & Camerik, 1994: 150
Type locality: Macquarie Id., Antarctica
Distribution: Macquarie I., Antarctica
Host: Stilbocarpa litter
Anereynetes Fain, 1964a: 104; 1985b: 257
Type species: Ereynetes hydrophilus Cooreman, 1947
Ereynetes beauchampi Hunter, 1964: 189
Ereynetes (Anereynetes) beauchampi Hunter: Fain & Camerik, 1994: 151
Type locality: on ginger root, from China at Boston
Distribution: ? China
Host: on ginger roots
Ereynetes bipilosus Willmann, 1953: 482; Franz, 1954: 365
Ereynetes (Anereynetes) bipilosus Willmann: Fain, 1964a: 107; Fain & Camerik, 1994: 153 (with ?)
Type locality: Germany
Distribution: Austria, Germany
Host: soil
Micrereunetes biscutatus Berlese, 1923: 243
Ereynetes biscutatus (Berlese): Thor, 1933: 75; Franz, 1954: 365
Ereynetes (Anereynetes) biscutatus (Berlese): Fain, 1964a: 107; Fain & Camerik, 1994: 152 (with ?)
Type locality: “in Agro Veneto”
Distribution: Austria, Italy
Host: dung
Ereynetes (Anereynetes) botswanensis Fain & Camerik, 1994: 158
Type locality: Mahalapye
Distribution: Botswana
Host: Metacatharsius sp. (Coleoptera: Scarabaeidae)
Ereynetes (Anereynetes) coenobitus Hunter & Poe, 1971: 135; Fain, 1985b: 257; Fain & Camerik, 1994: 151
Type locality: Puerto Rico
Distribution: Puerto Rico
Host: Coenobita clypeatus (Crustacea, Decapoda, Brachiura: Coenobitidae)
Ereynetes (Anereynetes) copridis Fain & Camerik, 1994: 158
Type locality: Mahalapye
Distribution: Botswana
Host: Copris subsidens (Coleoptera: Scarabaeidae)
Micrereunetes corticalis Vitzthum, 1923: 98
Ereynetes corticalis (Vitzthum): Vitzthum, 1930: 425; Mihelčič, 1958: 274
Ereynetes (Anereynetes) copridis (Vitzthum): Fain & Camerik, 1994: 152 (with ?)
Type locality: Waidhofen an der Thaya (Austria)
Distribution: Austria, Spain
Host: “im Mittelgang von Hylesinus und Myelophilus”, under Alisma plantago
Ereynetes cruciatus Berlese, 1910: 243
Micrereunetes cruciatus (Berlese): Fain, 1964a: 96
Ereynetes (Anereynetes) cruciatus (Berlese): Fain, 1964a: 107; Fain & Camerik, 1994: 151
Type locality: Java
Distribution: Indonesia (Java)
Host: Copris molossus (Coleoptera: Scarabaeidae)
Ereynetes davisi Hunter, 1964: 190
Ereynetes (Anereynetes) davisi Hunter: Fain & Camerik, 1994: 151
Type locality: Gadsden, Arizona
Distribution: USA (Arizona)
Host: “Under bark, cottonwood”
Ereynetes hydrophilus Cooreman, 1947: 6
Ereynetes (Anereynetes) hydrophilus Cooreman: Fain, 1964a: 107, 1985b: 257; Zabludovskaya, 1994: 36; Fain & Camerik, 1994: 151
Type locality: Kalmhoute (Belgium)
Distribution: Belgium, Ukraine
Host: in dung; described from the nest of Larus ridubundus
Ereynetes (Gymnereynetes) inermis Fain, 1964a: 96; Fain & Camerik, 1994: 151
Type locality: Florence
Distribution: Italy
Host: humus
Ereynetes insularis Willmann, 1952: 158
Ereynetes (Anereynetes) insularis Willmann: Fain, 1964a: 107; Fain & Camerik, 1994: 153 (with ?)
Type locality: Wangerooge Is.
Distribution: Germany
Host: soil near the coast
Ereynetes (Anereynetes) lyratus Fain & Prasse, 1973b: 233; 1985b: 257; Fain & Camerik, 1994: 151
Type locality: Etzdorf/Saalkreis
Distribution: Germany
Host: soil
Ereynetes (Gymnereynetes) meliponae Fletchtmann, Fain & Leal, 1985c: 95
Ereynetes (Aneynetes) meliponae: Fain & Camerik, 1994: 151
Type locality: Recife, Pernambuco
Distribution: Brazil
Host: Melipona scutellaris (Hymenoptera: Apidae, Meliponinae)
Tydeus ministralis C. L. Koch, 1838: fasc. 20, No 12
Ereynetes ministralis (C. L. Koch): Oudemans, 1928: 382; Thor, 1932: 28; Schweizer & Bader, 1963: 248
Ereynetes (? Anereynetes) ministralis (C. L. Koch): Fain & Camerik, 1994: 152
Type locality: Germany
Distribution: Germany, Switzerland
Ereynetes minutus Willmann, 1949: 127
Ereynetes (Anereynetes) minutus Willmann: Fain, 1964a: 107; Fain & Camerik, 1994: 153 (with ?)
Type locality: Ciechocinek (Poland, not Germany!)
Distribution: Poland
Host: soil
Ereynetes ornatus Mihelčič, 1958: 274
Ereynetes (Anereynetes) ornatus Mihelčič: Fain, 1964a: 107; Fain & Camerik, 1994: 153 (with ?)
Type locality: Spain
Distribution: Spain
Host: under Alisma plantation
Ereynetes (Anereynetes) papuanus Fain & Van Goethem, 1978: 125; 1985b: 257; Fain & Camerik, 1994: 151
Type locality: Laing Island, Morobe Prov.
Distribution: Papua New Guinea
Host: Coenobita rugosa (Crustacea, Decapoda, Brachiura: Coenobitidae)
Ereynetes potator Vitzthum, 1932: 78; Thor, 1933: 75
Ereynetes (Anereynetes) potator Vitzthum: Fain, 1964a: 107; Fain & Camerik, 1994: 152 (with ?)
Type locality: S. Sumatra (Ranau)
Distribution: Indonesia (Sumatra)
Host: in bamboo stumps
Ereynetes proximus Mihelčič, 1958: 275
Ereynetes (Anereynetes) proximus Mihelčič: Fain, 1964a: 107; Fain & Camerik, 1994: 153 (with ?)
Type locality: Spain
Distribution: Spain
Host: under Alisma plantation
Ereynetes sabinensis Baker, 1945: 19
Ereynetes (Anereynetes) sabinensis Baker: Fain, 1964a: 107; Fain & Camerik, 1994: 153 (with ?)
Type locality: Mexico
Distribution: Mexico
Host: bat guano
Ereynetes simplex Willmann, 1936: 196; Turk, 1970: 40
Ereynetes (Anereynetes) simplex Willmann: Fain, 1964a: 107; Fain & Camerik, 1994: 153 (with ?)
Type locality: graphite mine nr. Mährisch-Altstadt (Stare Mesto), Glatzer Schneeberg
Distribution: Bulgaria, Czech Rep. (on the border with Poland)
Host: unknown
Ereynetes sittardiensis Oudemans, 1912: 247; 1914: 120; Vitzthum, 1931: 81; Thor, 1933: 73
Ereynetes (? Anereynetes) sittardiensis Oudemans: Fain, 1964a: 107; Fain & Camerik, 1994: 152; Zabludovskaya, 1994: 36
Type locality: Sittard (Holland)
Distribution: Holland, Ukraine
Host: in the nests of rodents and insectivorans
Ereynetes (Anereynetes) upelbensis Corpuz-Raros, 1975: 329; Fain & Camerik, 1994: 151
Type locality: Mt. Makiling, Laguna
Distribution: Philippines
Gymnereynetes Fain, 1964a: 104; 1985b: 257
Type species: Ereynetes macquariensis Fain, 1962
Ereynetes (Gymnereynetes) aurantipes Fain & Bafort, 1973a: 411; 1985b: 257; Zabludovskaya, 1994: 36; Fain & Camerik, 1994: 152
Type locality: De Pinte nr. Gand (Belgium)
Distribution: Belgium, Ukraine, Crimea
Host: in humus and moss
Micrereunetes (Protereunetes) brevipes Berlese, 1923: 245
Protereunetes brevipes (Berlese): Macfadyen, 1954: 283
Ereynetes (Gymnereynetes) brevipes (Berlese): Fain, 1964a: 107; 1985a: 275; Fain & Camerik, 1994: 152
Type locality: Chianti (in the description: “habitat in Etruria”)
Distribution: Italy, Yan Mayen I.
Host: unknown
Ereynetes (Gymnereynetes) exilis Fain & Prasse, 1973b: 230; 1985b: 257; Zabludovskaya, 1994: 36; Fain & Camerik, 1994: 152
Type locality: Etzdorf/Saalkreis, Germany
Distribution: Germany, Ukraine, Crimea
Host: in soil
Ereynetes macquariensis Fain, 1962a: 921; Strandtmann, 1970: ; Crooker, 1977: 489; Luxton, 1990: 397
Ereynetes (Gymnereynetes) macquariensis: Fain, 1964a: 107; 1976: 319; 1985b: 257; Strandtmann & Tilbrook, 1968: 55; Fain & Camerik, 1994: 152
Type locality: Macquarie I.
Distribution: Macquarie I., Signy I., Kerguelen I., Marion I., Antarctic Peninsula
Huntereynetes Fain, 1964a: 110
Type species: Ereynetoides scutulis Hunter, 1964
Ereynetes (Huntereynetes) propescutulis Hunter et Rosario in Hunter, Rosario & Moser 1989: 16; Penttinen, Viiri & Moser, 2013: 3
Type locality: Georgia (USA)
Distribution: Finland, USA
Host: pine beetles (Coleoptera: Curculionidae, Ips) and their tunnels
Ereynetoides scutulis Hunter, 1964: 181
Ereynetes (Huntereynetes) scutulis (Hunter): Fain, 1964a: 110; 1985b: 258; Fain & Camerik, 1994: 152; Pernek et al., 2008:
Ereynetes scutulis (Hunter): Fernandes et al., 2013: 359
Type locality: Athens, Georgia
Distribution: Croatia, USA (Georgia)
Host: “from Ips tunnel in pine log”; phoretic on Pityokteines spinidens and P. vorontzowi (Col., Scolytidae)
Ereynetes (Huntereynetes) sinescutulis Hunter, Rosario & Moser, 1989: 18; Fain & Camerik, 1994: 152 (with ?)
Type locality: Idaho
Distribution: USA
Host: on pine beetles (Ips pini)
Opsereynetes Thor, 1932: 27(as genus)
Ereynetes (Opsereynetes): Fain, 1964a: 106
Type species: Opsereynetes norvegicus Thor, 1932
Opsereynetes norvegicus Thor, 1932: 27 – 28; 1933: 77; Shiba, 1969b: 87
Ereynetes (Opsereynetes) norvegicus (Thor): Fain, 1964a: 106; 1985b: 258; Fain & Camerik, 1994: 152; Donczyk, 2006: 165
Type locality: South Norway
Distribution: Japan, Norway, Poland
Opsereynetes robustus Baker, 1945: 18
Ereynetes (Opsereynetes) robustus Baker: Fain, 1964a: 106; Fain & Camerik, 1994: 152
Type locality: Desierto de los Leones
Distribution: Mexico
Ereynetes (Opsereynetes) simplexus Baker, 1945: 17
Ereynetes (Opsereynetes) simplexus Baker: Fain, 1964a: 106
Type locality: Desierto de los Leones
Distribution: Mexico
Fain & Cemeric (1994) indicate that: “Several other species have been included in the genus Ereynetes as species of Incertae sedis by Thor (1933): i.e. Acarus padi Schrank, 1803, Acarus junci Hermann, 1804, Acarus mammilaris Canestrini & Fanzago, 1876 and Acarus armatus Canestrini & Fanzago, 1876
Hanriccardoella André, Ducarme & Lebrun, 2004: 81
Type species: Hanriccardoella faini André, Ducarme et Lebrun, 2004
Hanriccardoella faini André, Ducarme & Lebrun, 2004: 83
Type locality: Han Cave (Rochefort)
Distribution: Belgium
Host: free living
Hydranetes Kethley, 1971: 176
Type species: Hydranetes tropisternus Kethley, 1971
Hydranetes citrinensis Kethley, 1971: 181
Type locality: Citra, Marion County, Florida
Distribution: USA (Florida
Host: Tropisternus lateralis nimbatus (Col.: Hydrophilidae)
Hydranetes tropisternus Kethley, 1971: 181
Type locality: Citra, Marion County, Florida
Distribution: USA (Florida
Hosts: Tropisternus lateralis nimbatus, T. quadristriatus, T. striolatus (Col.: Hydrophilidae)
Micrereunetes (Protereunetes) Berlese, 1923: 245 (subgenus)
Protereunetes Berlese: Oudemans, 1931: 252 (genus)
Type species: Micrereunetes (Protereunetes) agilis Berlese, 1923
Protereunetes boerneri Thor, 1934: 129; Macfadyen, 1954: 283; Strandtmann, 1971: 83
Type locality: Svalbard
Distribution: Svalbard, USA (Alaska), Yan Mayen I.
Eupodes clavifrons R. Canestrini, 1886: 716
Protereunetes clavifrons : Strandtmann, 1971: 85
Eupodes mollicellus C.L. Koch, 1838: fasc. 19, nr. 23
Cocceupodes mollicellus : Sig Thor, 1934: 178; Haarløv, 1957: 26
Type locality: Italy
Distribution: Denmark, Germany, Italy, Norway, Swalbard, Sweden
Protereunetes crozeti Strandtmann, 1972: 48
Type locality: Crozet Island
Distribution: Crozet Island
Protereunetes maudae Strandtmann, 1967: 56
Type locality: Victoria Land, Antarctica
Distribution: Antarctica
Protereunetes minutus Strandtmann, 1967: 55
Type locality: Antarctic Peninsula
Distribution: Antarctica
Protereunetus paulinae Llano & Gless, 2013: 299
Type locality: Hallett Station
Distribution: Antarctica
Protereunetes perforatus Shiba, 1978: 147
Type locality: Mt. Sagaramine, Shigenobu-sho, Ehime
Distribution: Japan
Eupodes striatellus C.L. Koch, 1838: fa. 19, t.16
Protereunetes striatellus (C.L. Koch): Oudemans, 1938: 870; Thor & Willmann, 1941: 55
Type locality: Regensburg (Germany)
Distribution: Germany, Estonia, Holland
Protereunetes villosus Shiba, 1978: 145
Type locality: Tomisato, Betshimura, Ehime
Distribution: Japan
Pseudotydeus Baker & Delfinado, 1974: 444
Type species: Pseudotydeus perplexus Baker et Delfinado, 1974
Pseudotydeus lebruni André & Ducarme, 2003: 375
Type locality: Nou Moulin Cave, Wallonia
Distribution: Belgium
Pseudotydeus perplexus Baker & Delfinado, 1974: 447
Type locality: Columbus, Ohio
Distribution: USA (Ohio)
Micrereunetes (Riccardoella) Berlese, 1923: 244
Type species: Acarus limacum Schrank, 1776
Proricardoella Fain & van Goethem, 1986: 129
Type species: Riccardoella oudemansi Thor, 1932
Riccardoella (Proriccardoella) canadensis Fain & Van Goethem, 1986: 134; Zabludovskaya, 1995: 90; André, Ducarme & Lebrun, 2004: 81
Type locality: Morgan Arboretum (Canada)
Distribution: Canada, Ukraine
Host: humus
Riccardoella (Proriccardoella) novaezealandiae Fain & Barker, 2004: 44
Type locality: Mt. Hikurangi, Poverty Bay
Distribution: New Zealand
Host: undescribed genus and species of Athoracophoridae (Gastropoda: Pulmonata)
Riccardoella oudemansi Thor, 1932a: 249; Sellnick, 1940: 33; Willmann, 1941: 58; 1951: 142; Franz, 1954: 365; Varga & Kontschan, 2004: 95
Riccardoella jenynsi Thor, 1933: 66
Riccardoella (Proriccardoella) oudemansi Thor: Fain & Van Goethem, 1986: 129; Fain & Barker, 2004: 48
Type locality: Holland
Distribution: Austria, Bulgaria, Croatia, England, Holland, Hungary, Island, Italy, Mexico, ?Slovakia, South Africa
Hosts: Arion ater, A. lusitanicus, A. rufus (Gastropoda: Arionidae), Deroceras reticulatum, Limax agrestis, L. cinereoniger, L. cinereus, L. flavus, L. maximus, (Gastropoda: Limacidae), Tandonia budapestensis (Gastropoda: Milacidae), Oxychilus draparnaudi, Meledella werneri (Gastropoda: Zonitidae), Cepaea nemoralis (Gastropoda: Helicidae)
Riccardoella (Proriccardoella) reaumuri Fain & Van Goethem, 1986: 134; Fain & Barker, 2004: 48; Waki & Shimano, 2019: 97
Type locality: Jura, nr. Ponarlier
Distribution: France, Japan
Host: Arianta arbustorum (Gastropoda: Helicidae)
Riccardoella (Proriccardoella) triodopsis Fain & Klompen, 1990: 187
Type locality: Bankhead Nat. Forest, Lawrence Co., Alabama
Distribution: USA (Alabama), Japan
Hosts: Xolotrema obstructa [Triodopsis obstricta] (Gastropoda: Polygyridae),
Acusta despecta sieboldtiana (Gastropoda: Bradybaenidae)
Riccardoella zadielensis André, Ducarme & Lebrun, 2004: 85
Type locality: Zádiel Gorge, Zádiel and Silica (Slovakia), Amateur’s Cave (Czech Rep.)
Distribution: Czech Rep., Slovakia
Host: free living
Riccardoella Berlese, 1923: 244 (subgenus); Fain & van Goethem, 1986: 129
Type species: Acarus limacum Schrank, 1776
Acarus limacum Schrank, 1776: 13; 1781: 521
?Acarus helicis Schrank, 1803: 209
Ereynetes limacum (Schrank): Berlese, 1883 (in part); Mégnin, 1889
Micrereunetes limacum : Fain, 1964a: 100
Micrereunetes (Ricardoella) limacum: Berlese, 1923: 244
Riccardoella limacum (Schrank): Thor, 1929: 145; Oudemans, 1929: 254; 1931: 253; Turk & Phillips, 1945: 448 (in part); Schweizer & Bader, 1963: 249; Arutyunyan, 1972: 108; Haralabidis et al., 1983: 144
Riccardoella (R.) limacum (Schrank): Fain & van Goethem, 1986: 130; Fain & Klompen, 1990: 190
Type locality: France
Distribution: Armenia, Belgium, Bulgaria, England, France, Greece, Italy, N. Macedonia (“S. Yugoslavia”), Poland, New Zealand, Serbia, South Africa, Switzerland, USA (recorded also from other countries, but, according to Fain & van Goethem, 1986, these data may concern Riccardoella oudemansi)
Hosts: Arianta arbustorum, Helix aspersa, H. lucorum, H. pomatia (Gastropoda: Helicidae), Cernuella aginnica (Gastropoda: Hygromiidae), Athoracophorus bitentaculatus (Gastropoda: Athoracophoridae) and other snails
Riccardoella tokyoensis Waki, Hiruta & Shimano, 2018: 163
Type locality: Tokyo
Distribution: Japan
Hosts: Tauphaedusa tau (Gastropoda: Clausiliidae)
Species inquirenda:
Hypopus concolor Haldeman, 1851
Ereynetes concolor : Banks, 1915: 22
Riccardoella concolor : Thor, 1933
Distribution: ?North America
Spelaeognathidae Womersley, 1936: 312 (see the comments of Domrow, 1969: 380)
Speleognathinae: Fain, 1957: 251 and auct.
Type genus: Speleognathus Womersley, 1936: 313
Boydaiini Fain, 1985a: 144
Type genus: Boydaia Womersley, 1953
Boydaia Womersley, 1953: 82; Fain, 1962 : 7; 1963: 17
Type species: Speleognathus sturni Boyd, 1948
Boydaia (Boydaia) agelaii Fain & Aitken, 1968: 82; Fain & Lukoschus, 1972a: 90
Boydaia agelaii Fain & Aitken: Bernardon et al., 2015, 2018
Type locality: La Paille Village, Trinidad
Distribution: Brazil, Suriname, Trinidad, USA
Hosts: Agelaius icterocephalus, A. phoeniceus, A. ruficapillus, Chrysomus ruficapillus (Passeriformes: Icteridae)
Boydaia spatulata americana Fain, 1963c: 46
Boydaia americana : Fain & Aitken, 1968: 80; 1970: 330
Boydaia (Boydaia) americana Fain: Fain, 1971a: 108
Type locality: South America, Brazil (Zoo birds)
Distribution: Brazil, South Africa, Trinidad
Hosts: Guiraca coerulea coeruela, Sporophila americana, Cyanocompsa cyanoides rotschildii, Volatinia jacarina, Piranga rubra rubra, Tangara gyrola, Rhamphocelus carbo, Coereba flaveola
Boydaia aratingae Fain, 1963: 42; Fain & Aitken, 1969: 34
Boydaia (Boydaia) aratingae Fain: Fain, 1971a: 111
Type locality: Anvers (coming from Brazil)
Distribution: Brazil
Host: Aratinga jandaya (Psittaciformes: Psittacidae, Psittacinae)
Boydaia bradornis Fain, 1956c: 654; Zabludovskaya, 1998, 2012; Skoracki et al., 2018: 421
Boydaia (Boydaia) bradornis Fain, 1958a: 176
Type locality: Akanyaru (Rwanda)
Distribution: Rwanda, Ukraine
Hosts: Agricola [Bradornis] pallidus griseus, Muscinia muscinia (Passeriformes: Muscicapidae)
Boydaia (Boydaia) buphagi Fain: Fain, 1961: 58; 1963d: 56, 1971a: 110; Skoracki et al., 2018: 422
Type locality: Rwanda
Distribution: Rwanda
Hosts: Buphagus africanus (Passeriformes: Buphagidae), Lamprocolius chloropterus (Passeriformes: Sturnidae)
Boydaia (Boydaia) cecropis Fain, 1969c: 374; Skoracki et al., 2018: 423
Type locality: Akanyaru
Distribution: Rwanda
Host: Cecropis abyssinica unitatis (Passeriformes: Hirundinidae)
Boydaia cinnyris Fain, 1969c: 373, 1971: 98; Skoracki et al., 2018: 423
Type locality: Akanyaru
Distribution: Rwanda
Hosts: Cinnyris cupreus cupreus (= septentrionalis), Chalcomitra senegalensis lamperti (= aequatorialis) (Passeriformes: Nectariniidae)
Boydaia clarki Fain, 1963: 41
Boydaia (Boydaia) clarki Fain: Fain, 1971a: 110
Type locality: Zoo d’Anvers (from the USA)
Distribution: USA
Host: Callipepla squamata (Galliformes: Odontophoridae)
Boydaia colini Clark, 1958: 78; Pence, 1973a: 365
Boydaia (Boydaia) colini Clark: Fain, 1971a: 110
Type locality: Maryland
Distribution: USA (Louisiana, Maryland)
Host: Colinus virginianus (Galliformes: Odontophoridae)
Boydaia cyanerpes Fain, 1963c: 45; Fain & Aitken,1969: 42
Type locality: Recife (dead in Anvers)
Distribution: Brazil
Host: Cyanerpes cyaneus (Passeriformes: Thraupidae)
Boydaia faini Dusbábek & Černý, 1970: 276
Type locality: Havana
Distribution: Cuba
Host: Dendroica cerulea (Passeriformes: Parulidae)
Boydaia falconis Fain, 1956c: 657, 1963c: 60, 1972: 610; Domrow, 1969: 381; Skoracki et al., 2018: 423
Type locality: Akanyaru (Rwanda)
Distribution: Australia, Rwanda, Chad (Tibesti)
Hosts: Falco berigora, F. cuvieri, F. t. tinnunculus (Falconiformes: Falconidae)
Boydaia formicarii Fain & Aitken, 1970: 331
Type locality: Moscueiro Ferry, Benavides
Distribution: Brazil
Hosts: Formicarius analis, F. colma (Passeriformes: Formicariidae)
Boydaia hirundoae Fain, 1956c: 661; 1960: 114; 1963d: 179; Domrow, 1965: 43; 1969: 381; Pence, 1973a: 365; 1973 : 887; 1975: 18; Fain & Aitken, 1968: 81; Fain & Hyland, 1970: 40; Skoracki et al., 2018: 423
Boydaia (Boydaia) hirundoae Fain: Fain, 1958a: 176; 1960 : 114;1963 : 56, 1971: 100; Fain, Herin & Puylaert, 1977: 393
Type locality: Akanyaru (Rwanda)
Distribution: Australia, Belgium, Brazil, Mexico, Rwanda, South Africa, Trinidad & Tobago, USA
Hosts: Hirundo abyssinica, H. neoxena, H. rustica, Petrochelidon [Hylochelidon] ariel, Progne chalybea, P. subis, Stelgidopteryx ruficollis (Passeriformes: Hirundinidae)
Boydaia indica Fain: 1969c: 373; Skoracki et al., 2018: 424
Boydaia (Boydaia) indica Fain: Fain, 1971 : 101
Type locality: Anvers Zoo
Distribution: DR Congo, India, East Africa
Hosts: Acridotheres tristis, Gracupita [Sturnopastor] contra, Sturnus malabaricus, S. pagodarum, Lamprocolius chloropterus, L. chalybaeus, Spreo superbus (Passeriformes: Sturnidae)
Boydaia jordani Van Eyndhoven, 1955: 203
Boydaia (Boydaia) jordani Van Eyndhoven: Fain, 1958a: 176
Boydaia turdi Clark, 1967: 294 – Type: ex Turdus migratorius, Bitterroot Valley, Montana, USA
Type locality: Kampen, Holland
Distribution: Holland, USA
Hosts: Turdus migratorius, T. pilaris (Passeriformes: Turdidae)
Boydaia laticoxa Fain & Aitken, 1969: 34; 1970: 328
Type locality: APEG Forest nr. Belem
Distribution: Brazil
Host: Glyphorynchus spirurus (Passeriformes: Dendrocolaptidae)
Boydaia (Boydaia) laticoxa philydori Fain & Aitken, 1970: 328
Type locality: APEG Forest (Station A), IPEAN, Belém
Distribution: Brazil
Host: Philydor pyrrhodes (Passeriformes: Furnariidae)
Boydaia locustellae Fain, 1970b: 320
Type locality: Hoboken nr. Anvers
Distribution: Belgium
Host: Locustella luscinioides (Passeriformes: Locustellidae)
Boydaia loxiae Fain, 1963d: 180
Type locality: Belgium
Distribution: Belgium
Host: Loxia curvirostra (Passeriformes: Fringillidae)
Boydaia maluri Domrow, 1969: 386
Type locality: Cowley Beach, Kowanyama
Distribution: Australia
Hosts: Malurus amabilis, M. melanocephalus (Passeriformes: Sylviidae)
Boydaia mimi Fain & Hyland, 1970: 40; Pence, 1973: 365
Type locality: Vera Cruz
Distribution: Mexico
Hosts: Mimus gilvus, M. polyglottos (Passeriformes: Mimidae)
Boydaia morenoi Dusbábek & Černý, 1970: 274
Type locality: El Dorado, Prov. of Las Villas
Distribution: Cuba
Host: Mimus polyglottos orpheus (Passeriformes: Mimidae)
Boydaia myzomelae Domrow, 1969: 383
Type locality: Esk
Distribution: Australia
Host: Myzomela sanguinolenta (Passeriformes: Meliphagidae)
Boydaia (Boydaia) nectarinia Fain, 1958a: 180, 1969
Type locality: Akanyaru
Distribution: Rwanda
Hosts: Cyanomitra verticalis viridisplendens, Chalcomitra senegalensis lamperti (= aequatorialis), Cinnyris cupreus cupreus (= septentrionalis) (Passeriformes: Nectariniidae)
Boydaia pheucticola Pence & Casto, 1976: 468; Spicer, 1977b: 498
Type locality: Texas, Jeff Davis Co., Fort Davis
Distribution: Mexico, USA
Hosts: Pheucticus melanocephalus, Spiza americana (Passeriformes: Cardinalidae)
Boydaia podargi Fain & Lukoschus, 1979a: 21; Koch, 1986: 61
Type locality: W. Australia
Distribution: Australia
Host: Podargus strigoides (Caprimulgiformes: Podargidae)
Boydaia psalidoprocnei Fain, 1956c: 660, 1963 : 57, 1963c: 56; Pence, 1973a: 365
Boydaia (Boydaia) psalidoprocnei Fain, 1958a: 176, 1971: 100
Type locality: Akanyaru (Rwanda)
Distribution: Canada, Russia, Rwanda, USA (Louisiana)
Hosts: Iridoprocne bicolor, Psalidoprocne albiceps, Riparia riparia, Tachycineta bicolor (Passeriformes: Hirundinidae)
Boydaia (Boydaia) psittaci Fain & Mortelmans, 1959: 16
Boydaia psittaci : Fain & Aitken, 1969: 42
Type locality: Antwerp Zoo [Peru]
Distribution: Brazil, Mexico, Peru
Hosts: Pionites melanocephala, P. leucogaster, Bolborhynchus lineola, Forpus passerinus, Brotogeris versicolorus, Amoropsittaca aymara (Psittaciformes: Psittacidae)
Boydaia pycnonoti Fain, 1956c: 655; Skoracki et al., 2018: 425
Boydaia (Boydaia) pycnonoti Fain: Fain, 1958a: 176, 1963c: 57; 1971: 98; Fain, Herin & Puylaert, 1977: 393
Type locality: Rwanda
Distribution: India, Kenya, Malaysia, Rwanda, South Africa, Zimbabwe
Hosts: Pycnonotus barbatus tricolor, P. goiavier personatus, P. jocosus, P. leucogenys leucotis, P. leucotis, P. xanthopygus (Passeriformes: Pycnonotidae)
Boydaia quiscali Clark, 1960: 108; Fain & Aitken, 1970: 330; Pence, 1973a: 365
Boydaia (Boydaia) quiscali Clark, 1960: Fain, 1971 : 109
Type locality: Louisiana
Distribution: Brazil, Mexico, USA
Hosts: Quiscalus quiscula (Passeriformes: Icteridae), Saltator maximus, S. coerulescens (Passeriformes: Thraupidae)
Boydaia rosickyi Dusbábek & Černý, 1970: 273
Type locality: Havana
Distribution: Cuba
Host: Passerina cyanea (Passeriformes: Fringillidae)
Boydaia saxicolae Fain, 1969: 370, 1971: 98; Skoracki et al., 2018: 425
Boydaia (Boydaia) saxicolae Fain: Fain, 1971a: 100
Type locality: Astrida (= Bucavu), Rugege Forest (Rwanda)
Distribution: Holland, Rwanda
Hosts: Saxicola torquatus axillaris, Monticola angolensis, Myrmecocichla nigra (Passeriformes: Muscicapidae), Turdus pilaris (Passeriformes: Turdidae)
Boydaia (Boydaia) sinensis Fain & Bafort, 1963: 80
Type locality: China
Distribution: China
Host: Leothrix lutea (Passeriformes: Timaliidae)
Boydaia sturni var. spatulata Fain, 1955b: 695
Boydaia spatulata Fain, 1956a: 47; 1956c: 651; 1956a: 33; 1958a: 10; Domrow 1969: 381; Skoracki et al., 2018: 426
Boydaia (Boydaia) spatulata Fain, 1958a: 176; 1960 : 114, 1963c: 56; 1971: 100
Type locality: Akanyaru, Butare (= Astrida)(Rwanda)
Distribution: Australia, Botswana, Guinea, Rwanda, South Africa, USA
Hosts: Melaniparus leucomelas insignis (Passeriformes: Paridae), Erythropygia [Cercotrichos] hartlaubi, Pogonocichla stellata ruwenzorii, Sheppardia polioptera, Cercomela tractrac (Passeriformes: Muscicapidae), Sialia sialis (Passeriformes: Turdidae), Tiaris canorus, Tangara sp. (Passeriformes: Thraupidae), Cissomela pectoralis, Philemon citreogularis, P. corniculatus (Passeriformes: Melliphagidae), Iduna similis (Passeriformes: Acrocephalidae), Schistolais leucopogon reichenowi (Passeriformes: Cisticolidae), Macronyx croceus (Passeriformes: Motacillidae),
Speleognathus sturni Boyd, 1948: 9; 1951: 79
Boydaia sturni : Womersley, 1953: 82; Cooreman, 1954: 426; Fain, 1956a: 42; 1956c: 651; 1961: 57; 1963c: 56; Domrow, 1969: 381; Pence, 1973a: 365
Boydaia (Boydaia) sturni: Fain, 1958a: 176
Type locality: Ithaca, N.Y.
Distribution: [Australia], Belgium, USA
Hosts: Acridotheres tristis, Ampeliceps coronatus, Sturnus vulgaris, (Passeriformes: Sturnidae), Quiscalus [Cassidix] mexicanus (Passeriformes: Icteridae)
Boydaia (Boydaia) synallaxis Fain, Hyland & Aitken, in Fain, 1969c: 376; Fain & Aitken, 1970: 332
Type locality: Musqueiro Ferry, Para, Belém (Brazil), Vera Cruz Cruz (Mexico)
Distribution: Brazil, Mexico
Hosts: Synallaxis rutilans, S. gujanensis, S. erythrothorax (Passeriformes: Furnariidae), Empidonax sp. (Passeriformes: Tyrannidae)
Boydaia tyrannis Ford, 1959: 380; Fain & Aitken, 1970: 330
Boydaia (Boydaia) tyrannis Ford: Fain, 1971a: 109
Type locality: Hickory Corners, Michigan
Distribution: Brazil, Mexico, Trinidad, USA (Michigan)
Hosts: Tyrannus tyrannus, Elaenia flavogaster, Myiophobus fasciatus, Pyrocephalus rubinus (Passeriformes: Tyrannidae), Manacus manacus, Pipra pipra, P. fasciicauda, P. iris (Passeriformes: Pipridae), Pachyramphus [=Platypsaris] aglaiae (Passeriformes: Tytiridae)
Boydaia willmanni Fritsch, 1962: 237
Type locality: Еrlangen
Distribution: Germany
Host: Gallinago [Capella] gallinago (Charadriiformes: Scolopacidae)
Boydaia zosteropis Fain, 1963c: 43; 1970b: 320; Domrow, 1969: 381; Skoracki et al., 2018: 426
Boydaia (Boydaia) zosteropis Fain: Fain, 1971a: 110
Type locality: Rugege (Rwanda)
Distribution: Australia, Rwanda
Hosts: Zosterops lateralis, Z. senegalensis stuhlmanni (= scotti) (Passeriformes: Zosteropidae)
Boydaia zumpti Fain, 1955b: 695; 1956a: 45; 1956c: 654; Skoracki et al., 2018: 427
Boydaia (Boydaia) zumpti Fain, 1958a: 176
Type locality: Astida (= Butare)
Distribution: Rwanda
Hosts: Euryllus [Andropadus] latirostris latirostris [= eugenius], Bleda syndactyla woosnami, (Passeriformes: Pycnonotidae), Schoenicola brevirostris alexinae (Passeriformes: Locustellidae)
Boydaia (Coboydaia) Fain, 1971: 98 (subgenus)
Coboydaia Fain: Fain, 1985a: 146 (genus)
Type species: Boydaia clavata Fain, 1955
Coboydaia (Apodiboydaia) Fain, 1985a: 146
Type species: Boydaia trochila Fain, 1958
Boydaia trochila Fain, 1958: 11
Boydaia (Coboydaia) trochila Fain: Fain, 1971a: 112
Boydaia (Apodiboydaia) trochila Fain: Fain, 1985a: 146
Type locality: Antwerp Zoo
Distribution: Brazil
Host: Chlorestes notatus (Apodiformes: Trochilidae)
Boydaia (Coboydaia) Fain, 1971a: 98 (subgenus)
Coboydaia Fain: Fain, 1985a: 146 (genus)
Type species: Boydaia clavata Fain, 1955
Boydaia amandavae Fain, 1962b: 363, 1963 : 58, 1970a: 319
Coboydaia amandavae : Fain, 1970b: 319; 1971: 101; Skoracki et al., 2018: 427
Type locality: Antwerp Zoo
Distribution: Central Africa, Indonesia (Java)
Hosts: Amandava subflava, A. amandava, Erythrura prasina, Sporaeginthus subflavus (Passeriformes: Estrildidae)
Boydaia clavata Fain, 1955b: 694; 1956c: 655, 1956a: 44, 1960b: 101, 1963 : 371, 1963c: 59; Skoracki et al., 2018: 428
Boydaia (Apodiboydaia) clavata Fain: Zabludovskaya, 2002: 93
Boydaia (Coboydaia) clavata Fain: Fain, Herin & Puylaert, 1977: 392
Type locality: Rwanda
Distribution: Burundi, Cameroon, Ghana, Guinea, Rwanda, Senegal, South Africa, Zambia, Zimbabwe
Hosts: Euplectes hordeaceus, E. ardens, E. macroura, Othyphantes baglafecht stuhlmanni, Ploceus baglafecht stuhlmanni, P. [Textor] cucullatus, P. [Textor] cucullatus graueri, P. pelzelni, P. velatus, P. vitellinus, P. xanthops (= P. x. camburni), Quelea quelea, Q. quelea centralis, Q. quelea aethiopica, Q. erythrops (Passeriformes: Ploceidae), Saxicola rubetra (Passeriformes: Turdidae) , Vidua chalybeata, V. chalybeata amauropteryx (Passeriformes: Viduidae), Streptopelia vinacea (Columbiformes: Columbidae)
Boydaia nigra Fain, 1955b: 695; 1956a: 44; 1956c: 658; 1959: 33; 1962: 266; 1963a: 168, 1963c: 58, 1963d: 179; Fain & Aitken, 1968: 81; Pence 1973a: 365, 1973c: 888
Boydaia (Coboydaia) nigra Fain, 1955b: 695; 1971: 101; Fain & Goff, 1980: 506; Fain, Herin & Puylaert, 1977: 393
Boydaia (Coboydaia) nigra nigra: Fain & Hyland 1975: 207
Boydaia nigra icteri Fain & Hyland, 1970: 38
Ereynetes crassipes Berlese & Trouessart, 1889: 141; Fain, 1957: 253
Boydaia crassipes (Berlese et Trouessart, 1889): Domrow 1969: 386 (Australia): Riccardoella crassipes: Thor, 1933: 66
Boydaia sturni , sensu Porter & Strandtmann, 1952 not Speleognathus sturni Boyd, 1948; Fain, 1958a: 179
Coboydaia nigra (Fain): Skoracki, 2018: 430
Type locality: Rwanda
Distribution: Belgium, Botswana, Burundi, Cameroon, Guinea, Kazakhstan, Mexico, Russia, Rwanda, South Africa, Trinidad & Tobago, Turkmenistan, Ukraine, USA (Hawaii)
Hosts: Icterus galbula, I. spurius (Passeriformes: Icteridae), Motacilla capensis, M. flava (Passeriformes: Motacillidae), Carduelis carduelis, Carpodacus erythrinus, Euphonia violacea, Fringilla coelebs, F. montifringilla, Haemorhous mexicanus, Serinus sulphuratus shelleyi, S. citrinelloides frontalis, S. mozambicus barbatus, S. [Poliospiza] burtoni tanganyicae, Serinus [Poliospiza] gularis (Passeriformes: Fringillidae), Emberiza aureola, E. citrinella, E. pusilla, E. rustica, E. rutila, E. spodocephala, E. tristrami, Spizella passerina (Passeriformes: Emberizidae), Passer ammodendri, P. domesticus, P. griseus, P. hispaniolensis, P. montanus (Passeriformes: Passeridae), Tanagra violacea (Passeriformes: Thraupidae), Oenanthe isabellina (Passeriformes: Turdidae), Lalage tricolor (Passeriformes: Campophagidae)
Boydaia nigra motacillae Fain, 1969c: 375
Coboydaia nigra motacillae Fain: Zabludovskaya, 2002: 93
Type locality: Nyakibanda, Butare (= Astrida)(Rwanda), Busumbura (Burundi)
Distribution: Belgium, Burundi, Rwanda, Russia, South Africa
Hosts: Anthus trivialis, Motacilla aguimp vidua, M. alba, M. capensis, M. flava (Passeriformes: Motacillidae), Acrocephalus arundinaceus, A. s. scirpaceus (Passeriformes: Acrocephalidae)
Boydaia sturnellae Clark, 1960: 104
Boydaia (Coboydaia) sturnellae: Fain, 1971a: 112
Type locality: Laurel, Maryland
Distribution: USA
Host: Sturnella magna (Passeriformes: Icteridae)
Trispeleognathini Fain, 1985a: 144
Type genus: Trispeleognathus Fain, 1958
Astrida Fain, 1955b: 696
Type species: Astrida caprimulgi Fain, 1955
Astrida Fain, 1955b: 696
Type species: Astrida caprimulgi Fain, 1955
Astrida caprimulgi Fain, 1955b: 696; 1956a: 33; 1956c: 646, 1962a: 9, 1963c: 62; Pence, 1973b: 879, 1973c: 886; Skoracki et al., 2018: 403
Type locality: Muheri nr. Astrida (= Butare)(Rwanda)
Distribution: Malaysia, Rwanda, USA
Hosts: Antrostomus carolinensis, Scotornis fossii welwitschi (Caprimulgiformes: Caprimulgidae), Bubo africanus, Glaucidium perlatum, Otus senegalensis, O. spilocephalus vulpes (Strigiformes: Strigidae)
Astrida onychognathi Fain, 1964c: 37
Type locality: Butare (= Astrida)
Distribution: Rwanda
Host: Onychognathus walleri elgonensis (Passeriformes: Sturnidae)
Cerylonyssus Fain & Aitken, 1970: 326
Type species: Astrida (Cerylonyssus) chlorocerylei Fain et Aitken, 1970
Astrida (Cerylonyssus) chlorocerylei Fain & Aitken, 1970: 327
Type locality: Sapucajuba, IPEAN, Belém
Distribution: Brazil
Host: Chloroceryle aenea (Coraciiformes: Alcedinidae, Cerylinae)
Aureliania Fain, 1958a: 177 (subgenus)
Aureliania Fain, 1970b: 321 (genus)
Type species: Boydaia aureliani Fain, 1955
Boydaia aureliani Fain, 1955b: 694; 1956a: 36
Boydaia (Aureliania) aureliani Fain: Fain, 1958a: 177
Neoboydaia (Aureliania) aureliani (Fain): Fain, 1963 : 61, 1963d: 181, 1963c: 52; Domrow, 1969: 389.
Aureliania aureliani (Fain): Fain 1962a: 9, 1985a: 146; Domrow 1975: 18, 1991: 1289; Skoracki et al., 2018: 404
Type locality: Butare (= Astrida)(Rwanda)
Distribution: Australia, Belgium, Rwanda
Hosts: Tyto alba alba, T. alba affinis (Strigiformes: Tytonidae)
Speleognathus longipilis Rosas Costa, 1955: 79
Boydaia (Aureliania) longipilis (Rosas Costa): Fain, 1958a: 179
Neoboydaia (Aureliania) longipilis (Rosas Costa): Fain, 1963c: 61
Aureliania longipilis (Rosas Costa):
Type locality: Peia. de Buenos-Aires
Distribution: Argentina
Host: Guira guira (Cuculiformes: Cuculidae)
Meropiboydaia Fain, 1985a: 145
Type species: Boydaia merops Fain, 1955
Boydaia merops Fain, 1955b: 694; 1956a: 41, 1956d: 30
Aureliania merops , Fain 1958b: 177
Neoboydaia (Aureliania) merops: Fain, 1963c: 61
Neoboydaia merops (Fain): Domrow, 1966: 210, 1969: 389
Meropiboydaia merops (Fain): Fain, 1985a: 146; Domrow, 1991: 1294
Type locality: Astrida (= Butare), Rwanda
Distribution: Australia, Rwanda
Hosts: Merops apiaster, M. ornatus (Coraciiformes: Meropidae)
Metaboydaia Fain, 1962 : 7; 1985a: 149
Type species: Speleognathus poffei Fain, 1955
Speleognathus poffei Fain, 1955b: 695; 1956a: 23
Speleognathus (Neospeleognathus) poffei: Fain, 1958a: 178
Neoboydaia (Metaboydaia) poffei: Fain 1962a: 7
Trispeleognathus (Neospeleognathus) poffei: Fain, 1963c: 24
Metaboydaia poffei : Fain, 1985a: 149
Type locality: Akanyaru, Astrida (= Butare)
Distribution: Rwanda
Hosts: Scopus umbretta (Pelecaniformes: Scopidae), Coracias caudatus caudatus (Coraciiformes: Coraciidae)
Neastrida Fain, 1962a: 9
Type species: Astrida parrae Fain, 1956
Astrida parrae Fain, 1956e: 113
Astrida (Neastrida) parrae: Fain, 1962a: 9, 1963c: 26
Neastrida parrae : Fain 1985a: 148.
Type locality: Butare (Rwanda)
Distribution: Rwanda, South Africa
Host: Actophilornis africanus (Charadriiformes: Jacanidae)
Boydaia (Neoboydaia) Fain, 1958a: 177 (subgenus)
Neoboydaia Fain, 1961b: 162, 1962a: 7, 1963c: 52
Type species: Boydaia philomachi Fain, 1956
Neoboydaia colymbiformi Clark, 1964: 158; Domrow, 1969: 189; Pence, 1973a: 364, 1975: 18; Fain, 1985a: 148
Type locality: California
Distribution: USA (California, Louisiana)
Hosts: Podiceps [Colymbus] nigricollis, P. ruficollis, Podilymbus podiceps (Podicipediformes: Podicipedidae)
Neoboydia galachrysiae Fain, 1961b: 129; 1985a: 148; Skoracki et al., 2018: 409
Neoboydaia (Neoboydaia) galachrysiae: Fain, 1963c: 19; 1964 : 137
Type locality: Kasongo (Maniema Prov.)
Distribution: Democratic Republic of the Congo
Hosts: Glareola [Galachrysia] cinerea, G. nuchalis (Aves, Glareolidae)
Boydaia philomachi Fain, 1956a: 41; 1956d: 27; 1962e: 300
Boydaia (Neoboydaia) philomachi Fain, 1958a: 177, 1962a: 7, 1964b: 141
Neoboydaia (Neoboydaia) philomachi: Fain, 1962a: 7, 1963c: 18, 1964b: 141; Domrow, 1965: 43
Neoboydaia philomachi eroliae Fain et Hyland, 1970: 42, 1975: 204; Zabludovskaya, 1998: 48
Neoboydaia philomachi : Clark, 1964: 16, fig. 2; Domrow, 1965: 43, 1969: 388; Fain & Aitken 1968: 84; Pence 1973a: 364, 1973c: 884, 885, 891; Fain, 1985a: 148; 1985b: 254
Neoboydaia philomachi thalasseus Fain & Hyland 1975: 204
Type locality: Butare (= Astrida), Rwanda
Distribution: Australia, Russia, Rwanda, USA
Hosts: Actitis macularius, Arenaria interpres, Gallinago gallinago, Limnodromus griseus, L. scolopaceus, Limosa lapponica, Calidris [Philomachus] pugnax, C. melanotos, C. minutilla, Tringa flavipes, T. melanoleuca, T. glareola, T. solitaria (Charadriiformes: Scolopacidae), Sterna hirundo, Thalasseus maximus (Charadriiformes: Laridae, Sterninae)
Ophthalmognathus Dubinin, 1957: 66
Neospeleognathus Fain, 1958a: 178 (subgenus of Speleognathus); 1985a: 147 (syn.)
Type species of Ophthalmognathus: Ophthalmognathus dogieli Dubinin, 1957
Type species of Neospeleognathus: Speleognathus schoutedeni Fain, 1955
Speleognathus schoutedeni Fain, 1955b: 695; 1956b: 18; 1962e: 301
Speleognathus (Neospeleognathus) schoutedeni: Fain, 1958a: 178
Ophthalmognathus dogieli Dubinin, 1957: 68 – Russia, syn. by Domrow, 1969: 392
Neoboydaia (Neospeleognathus) schoutedeni: Fain, 1962 : 7; 1963c: 62, 1963d: 23
Trispeleognathus (Neospeleognathus) schoutedeni: Fain, 1963c: 23
Ophthalmognathus schoutedeni (Fain): Domrow, 1969: 392; 1991: 1295; Fain & Goff, 1980: 506; Fain, 1985a: 147; Halliday, 1998: 50; Skoracki et al., 2018: 418
Type locality: Butare (= Astrida)
Distribution: Australia, Rwanda, Russia
Hosts: Ardea cinerea, Ardeola idae, Nycticorax caledonicus, N. nycticorax (Pelecaniformes: Ardeidae)
Ophthalmognathus tenorioae Fain & Goff, 1980: 506; Goff, 1983: 167; Fain, 1985a: 147
Type locality: Oahu I., Honolulu, Campbell Estate Road
Distribution: Rwanda, USA (Hawai, Oahu I.)
Hosts: Nycticorax nycticorax, N. n. hoactli (Pelecaniformes: Ardeidae)
Phoenicopteriella Fain, 1970b: 323
Type species: Phoenicopteriella mirabilis Fain, 1970
Phoenicopteriella mirabilis Fain, 1970b: 323; 1985a: 152
Type locality: North Africa
Distribution: North Africa
Host: Phoenicopterus ruber (Phoenicopteriformes: Phoenicopteridae)
Speleognathus womersleyi Fain, 1955b: 695; 1956a: 21; Clark, 1958: 84
Trispeleognathus womersleyi : Fain 1958b: 178, 1962a: 9; Fain & Hyland 1975: 205; Kaneko, Matsudaira & Prince Masahito, 1978: 147
Trispeleognathus (Trispeleognathus) womersleyi: Fain, 1963c: 23
Ophthalmognathus womersleyi : Domrow, 1969: 390; Spicer, 1977: 498
Phoenicopteriella womersleyi (Fain): Fain, 1985a: 152, 1985b: 254; Skoracki et al., 2018: 412
Type locality: Butare (= Astrida)(Rwanda)
Distribution: Japan, Rwanda, USA
Hosts: Authya affinis, Anas acuta, A. clypeata, A. discors, A. strepera, Dendrocygna viduata, Sarkidiornis melanotus (Anseriformes: Anatidae)
Picinyssus Fain, 1969c: 369
Type species: Picinyssus buccanodon Fain, 1969
Picinyssus buccanodon Fain, 1969c: 370; 1985a: 151
Type locality: Butahiro and Kamahia (Nord Kivu)
Distribution: DR Congo
Host: Buccanodon duchaillui (Piciformes: Lybiidae)
Psittaboydaia Fain, 1985a: 145
Trichoglossiella Fain, 1985a: 145
Type species of Psittaboydaia: Neoboydaia psittaculae Fain, 1962
Type species of Trichoglossiella: Aureliania trichoglossi Fain, 1970
Psittaboydaia (P.) amazona (Fain): Fain, 1985a: 145
Type locality: Meerzorg
Distribution: Suriname
Host: Amazona amazonica (Psittaciformes: Psittacidae)
Neoboydaia psittaculae Fain, 1962: 7; 1962 : 318; Domrow, 1969: 388
Neoboydaia (Neoboydaia) psittaculae Fain, 1963c: 19
Psittaboydaia (Psittaboydaia) psittaculae (Fain): Fain, 1985a: 147; Skoracki et al., 2018: 415
Type locality: Central Africa
Distribution: Australia, Central Africa
Hosts: Agapornis roseicollis, Trichoglossus moluccanus, Psitteuteles versicolor, Psittacula krammeri (Psittaciformes: Psittaculidae)
Aureliania trichoglossi Fain, 1970b: 321
Psittaboydaia (Trichoglossiella) trichoglossi (Fain): Fain, 1985a: 145
Type locality: Anvers Zoo
Distribution: New Guinea
Host: Trichoglossus haematod forsteni (Psittaciformes: Psittaculidae)
Pterniboydaia Fain, 1985a: 146
Type species: Boydaia pternistis Fain, 1955
Boydaia pternistis Fain, 1955a: 145, 1956c: 646
Aureliania pternistis (Fain): Fain, 1963 : 373
Neoboydaia pternistis (Fain): Fain, 1963c: 21; Kaneko, Kadosaka & Toshioka, 1977: 163
Neoboydaia (Aureliania) pternistis (Fain): Fain, 1963c: 61
Pterniboydaia pternistis (Fain): Fain, 1985a: 146; Skoracki et al., 2018: 416
Type locality: Astrida (= Butare) (Rwanda)
Distribution: Belgium, Japan, Rwanda, Zambia
Hosts: Bambusicola thoracica, Pternistis afer, P. swainsoni, Perdix p. perdix (Galliformes: Phasianidae)
Neoboydaia (Ralliboydaia) Fain, 1962b: 365 (subgenus)
Ralliboydaia Fain: Fain, 1985a: 151 (genus); Fain, 1963: 52 is synonymizing Ralliboydaia with Neoboydaia.
Type species: Neoboydaia (Ralliboydaia) latiralli Fain, 1962
Astrida coccyzae Pence, 1973a: 366
Neoboydaia (Ralliboydaia)
Ralliboydaia coccyzae (Pence): Fain, 1985a: 151
Type locality: Mandeville, La.
Distribution: USA (Louisiana)
Host: Coccyzus americanus (Cuculiformes: Cuculidae)
Neoboydaia (Ralliboydaia) latiralli Fain, 1962b: 365
Neoboydaia (Neoboydaia) latiralli Fain, 1963c: 20
Ralliboydaia latiralli (Fain): Fain, 1985a: 151
Type locality: Brazil
Distribution: Brazil
Host: Latirallus melanophaius oenops (Ralliformes: Rallidae)
Speleognathopsis porphyrionis Domrow, 1965: 44; 1969: 393
Ralliboydaia porphyrionis (Domrow): Fain, 1985a: 151
Type locality: Samford, S. E. Queensland
Distribution: Australia (Queensland)
Host: Porphyrio melanotus (Gruiformes: Rallidae)
Speleognathopsis Cooreman, 1954: 428
Type species: Speleognathopsis galli Cooreman, 1954
Speleognathopsis Cooreman, 1954: 428
Type species: Speleognathopsis galli Cooreman, 1954
Ophthalmognathus accipitris Domrow, 1969: 390; Fain, 1985a: 146
Speleognathopsis accipitris (Domrow): Domrow, 1991: 1301
Type locality: Kowanyama
Distribution: Australia
Host: Accipiter fasciatus (Falconiformes: Accipitridae)
Speleognathopsis benoiti Fain, 1955b: 696; 1956a: 29; 1962 : 9; 1963c: 25; 1985a: 150; Domrow, 1966: 210; 1969: 393; 1991: 1301; McDaniel, 1967: 94
Speleognathopsis wai Fain, Vercammen-Grandjean & Wagner 1966: 115. Type locality: Bloemhof, Transvaal, South Africa (syn. by Domrow, 1969: 393)
Type locality: Butare (= Astrida), Rwanda
Distribution: Australia, Rwanda, USA (Texas)
Hosts: Charadrius melanops, Erythrogonys cinctus, Lobibyx miles, Pluvialis squatarola, Vanellus [Hemiparra] crassirostris, V. senegallus (Charadriiformes: Charadriidae)
Speleognathopsis charadricola Fain, 1964c: 35; 1985a: 150
Type locality: Zoo Anvers
Distribution: Central Africa
Host: Charadrius (Leucopolius) pecuarius (Charadriiformes: Charadriidae)
Speleognathopsis galli Cooreman, 1954: 426; Fain, 1963 : 53; 1963c: 62; 1985a: 149; Domrow, 1969: 392; Talbot, 1969: 582; Faccini, 1981: 135; Panda et al., 1987: 507
Type locality: Astrida (= Butare), Rwanda
Distribution: Brazil, India, Papua New Guinea, Rwanda
Hosts: Gallus gallus domesticus (Galliformes: Phasianidae), Numida meleagris intermedia (Galliformes: Numididae)
Speleomyotis Fain, 1970: 513.
Type species: Speleognathopsis bastini Fain, 1958
Speleognathopsis bastini Fain, 1958b: 342
Speleognathopsis (Speleomyotis) bastini Fain: Fain, 1966: 108; 1970c: 513
Speleognathopsis (Speleomyotis) bastini bastini Fain: Fain & Lukoschus, 1971c: 285
Type locality: Belgium
Distribution: Belgium, Japan, Suriname
Hosts: Eptesicus fuscus, E. melanopterus, Myotis myotis, M. nigricans (Chiroptera: Vespertilionidae)
Speleognathopsis bastini didelphis Fain & Aitken, 1969: 41
Type locality: Jandiai, Caratateua, Bragança (Pará)
Distribution: Brazil
Host: Didelphis marsupialis (Didelphimorphia: Didelphidae)
Speleognathopsis (Speleomyotis) molossus Fain & Lukoschus, 1971c: 285
Type locality: Paramaribo
Distribution: Suriname
Hosts: Cynomops planirostris, Molossus ater, M. molossus (Chiroptera: Molossidae)
Speleognathus (Trispeleognathus) Fain, 1958a: 178
Trispeleognathus Fain: 1962: 7
Type species: Speleognathus striatus Crossley, 1952
Trispeleognathus euricoi Amaral, 1965: 31
Ophthalmognathus euricoi (Amaral): Domrow, 1969: 391
Trispeleognathus (Trispeleognathus) euricoi: Amaral, 1968: 124; Fain & Aitken, 1969: 43
Type locality: Itaporanga (Estado de São Paulo)
Distribution: Brazil
Hosts: Leptotila verreauxi, Nothura m. maculosa (Columbiformes: Columbidae)
Speleognathus striatus Crossley, 1952: 385; Fain, 1955b: 695, 1956b: 18,
Boydaia striatus : Womersley, 1953: 82; Cooreman, 1954: 426; Fain, 1955 : 695; 1962 : 300
Speleognathus (Trispeleognathus) striatus: Fain 1958a: 178
Ophthalmognathus striatus : Domrow 1969: 389; Pence & Casto, 1976: 466
Trispeleognathus (Trispeleognathus) striatus: Fain, 1963c: 22; Amaral 1963: 91; Fain & Aitken, 1969: 43, 1970: 328
Trispeleognathus striatus : Fain 1962a: 9, 1985a: 149; Amaral 1963: 91; Fain & Hyland, 1970: 42; Skoracki et al., 2018: 420
Type locality: Garza Co., Lubbock Co., Texas
Distribution: Brazil, Mexico, Rwanda, Trinidad & Tobago, USA,
Hosts: Columba livia domestica, Columbigallina passerina, Geotrygon montana, Leptotila verreauxi, Streptopelia s. semitorquata (Columbiformes: Columbidae)
Speleochirini Fain, 1985a: 144
Type genus: Speleochir Fain, 1966
Hipposideroptes Fain, 1970c: 514
Type species: Hipposideroptes kimuenzae Fain, 1970
Hipposideroptes kimuenzae Fain, 1970c: 514
Type locality: Kimuenza
Distribution: DR Congo
Host: Hipposideros commersoni gigas (Chiroptera: Hipposideridae)
Hipposideroptes saccopteryx Fain & Lukoschus, 1971c: 289
Type locality: Lelydorp
Distribution: Suriname
Hosts: Saccopteryx bilineata, S. leptura (Chiroptera: Emballonuridae)
Speleochir (Neospeleochir) Fain, 1970c: 514
Neospeleochir Fain:
Type species: Boydaia duboisi Fain, 1955
Boydaia Duboisi Fain, 1955b: 691
Boydaia (Aureliania) duboisi Fain: Fain, 1958a: 176
Speleochir duboisi Fain: Fain, 1966: 108
Speleochir (Neospeleochir) duboisi Fain: Fain, 1970c : 514
Neospeleochir duboisi Fain: Fain, 1970c: 520
Type locality: Astrida (= Butare)
Distribution: Rwanda
Host: Nycteris sp. (Chiroptera: Nycteridae)
Speleognathopsis (Neospeleognathopsis): Fain, 1958a: 177
Neospeleognathopsis Fain: Fain, 1962 : 10
Type species: Speleognathopsis chiropteri Fain, 1955
Neospeleognathopsis Fain, 1958a: 177
Type species: Speleognathopsis chiropteri Fain, 1955
Speleognathopsis chiropteri Fain, 1955b: 692; 1956a: 31
Neospeleognathopsis (N.) chiropteri (Fain): Fain, 1966: 108; 1970c: 513
Type locality: Astrida (Butare)(Rwanda)
Distribution: D.R.Congo, Egypt, Rwanda
Hosts: Eidolon helvum, Epomophorus labiatus minor, Rousettus aegyptiacus, R. aegyptiacus leachi (Chiroptera: Pteropodidae)
Pteropignathus Fain & Lukoschus, 1979: 58
Type species: Neospeleognathopsis (Pteropignathus) pteropus Fain et Lukoschus, 1979
Neospeleognathopsis (Pteropignathus) pteropus Fain & Lukoschus, 1979: 58
Pteropignathus pteropus (Fain et Lukoschus):
Type locality: Brooking Springs
Distribution: Australia
Host: Pteropus scapulatus (Chiroptera: Pteropodidae)
Speleochir Fain, 1966: 103
Type species: Speleochir aitkeni Fain, 1966
Speleochir aitkeni Fain, 1966: 107
Speleochir (S.) aitkeni Fain: Fain, 1970c: 517
Type locality: Aripo Cave, Northern Range
Distribution: Trinidad (W.I.)
Host: Anoura g. geoffroyi (Chiroptera: Phyllostomatidae)
Speleochir (Speleochir) barbulata Fain & Aitken, 1970: 334
Type locality: APEG Forest, IPEAN, Belém
Distribution: Brazil
Host: Chiroptera indet.
Speleochir brasiliensis Fain & Aitken, 1969: 39
Type locality: Mucambo Forest, IPEAN, Brazil
Distribution: Brazil
Hosts: Artibeus jamaicensis, Vampyrodes caraccioloi (Chiroptera: Phyllostomatidae)
Neospeleognathopsis phyllostomi Clark, 1967: 241
Speleochir phyllostomi Clark: Fain & Aitken, 1969: 39
Type locality: Colombia
Distribution: Colombia
Host: Phyllostomus hastatus (Chiroptera: Phyllostomatidae)
Speleochiroides Fain, 1985a: 146
Type species: Speleochir carollia Fain et Liukoschus, 1971
Speleochir carollia Fain & Lukoschus, 1971: 293
Speleochiroides carollia (Fain et Lukoschus): Fain, 1985a: 146
Type locality: Lelydorp
Distribution: Suriname
Host: Carollia perspicillata (Chiroptera: Phylostommatidae)
Paraspeleognathopsini Fain, 1985a: 145
Type genus: Paraspeleognathopsis Fain, 1958
Speleognathopsis (Paraspeleognathopsis) Fain, 1958a: 177
Paraspeleognathopsis Fain: Fain, 1970c: 510 (genus). According to Domrow (2997), Paraspeleognathopsis Fain, 1958 = Speleorodens Fain, 1962 = Speleomys Fain, 1971
Type species: Speleognathopsis bakeri Fain, 1955
Speleognathopsis bakeri Fain, 1955b: 148; 1955a: 695; 1956a: 26; 1956d: 159
Speleognathopsis (Paraspeleognathopsis) bakeri Fain: Fain, 1958a: 177
Paraspeleognathopsis bakeri (Fain): Domrow, 1967: 775; 1991: 1296
Paraspeleognathopsis (P.) bakeri (Fain): Fain & Lukoschus, 1968: 85; Beron, 1973: 189
Type locality: Butare (= Astrida), Rwanda
Distribution: Australia, Belgium, Bulgaria, Holland, Japan, Korea, Rwanda, Spain
Hosts: Apodemus agrarius coreae, A. speciosus, Rattus natalensis (sub Mastomys coucha ugandae), Melomys cervinipes, Sylvaemus flavicollis, S. sylvaticus, S. sylvaticus callipides (Rodentia: Muridae)
Paraspeleognathopsis exaeta Domrow, 1991: 1298
Type locality: S.E.New South Wales, Bondo State Forest
Distribution: Australia
Host: Petaurus breviceps (Diprotodontia: Petauridae)
Speleognathopsis galagoensis Fain, 1956a: 32; 1956: 167
Paraspeleognathopsis (P.) galagoensis Fain, 1963c: 64
Type locality: Akanyaru
Distribution: Rwanda
Host: Galago senegalensis moholi (Primates: Galagonidae)
Paraspeleognathopsis graphiuri Fain, 1970c: 510
Type locality: Butare (= Astrida)
Distribution: Rwanda
Host: Graphiurus (Claviglis) microtis (Rodentia: Gliridae)
Speleomys Fain, 1970c: 513
Type species: Speleognathopsis galliardi Fain, 1955
Speleognathopsis galliardi Fain, 1955b: 149; 1956a: 26; 1956: 163
Speleognathus galliardi Fain, 1956: 163
Speleomys galliardi (Fain): 1970c: 513
Type locality: Kansi and Butare (= Astrida)
Distribution: Rwanda
Hosts: Arvicanthis abyssinicus, Gerbilliscus boehmi (Rodentia: Muridae)
Paraspeleognathopsis (Speleorodens) neotropicalis Fain & Aitken, 1969: 37
Speleomys neotropicalis (Fain et Aitken): Fain, 1970c: 515
Type locality: Jandiai, Caratateua, Bragança (Pará)
Distribution: Brazil
Host: Zygodontomys lasiurus fuscinus (Rodentia: Cricetidae, Sigmodontinae)
Speleorodens Fain, 1962: 10
Paraspeleognathopsis (Speleorodens) Fain, 1963c: 55
Type species: Speleognathopsis strandtmanni Fain, 1956
Paraspeleognathopsis cricetidarum Clark, 1967: 240
Paraspeleognathopsis (Speleorodens) cricetidarum: Fain & Aitken, 1969: 36
Type locality: Panama
Distribution: Brazil, Panama, Trinidad
Hosts: Oryzomys talamancae, O. laticeps velutinus, Zygodontomys cherriei, Heteromys desmarestianus, Neracomys guianae (Rodentia: Cricetidae)
Boydaia derricki Womersley, 1954: 65; Domrow & Smith, 1956: 203
Astrida derricki (Womersley): Fain, 1956: 167
Speleognathopsis derricki (Womersley): Domrow, 1961: 376
Paraspeleognathopsis (Speleorodens) derricki: Fain, 1963: 64
Speleomys derricki (Womersley):
Paraspeleognathopsis derricki (Womersley): Domrow, 1967: 775
Speleorodens derricki (Womersley): Fain, 1971c: 516
Type locality: Mt. Glorious, S.E. Queensland
Distribution: Australia
Hosts: Hydromys chrysogaster, Mus musculus, Rattus assimilis, R. conatus, R. fuscipes (Rodentia: Muridae)
Speleognathopsis michigensis Ford, 1962: 104
Paraspeleognathopsis (Speleorodens) clethrionomys Fain & Lukoschus, 1968: 86 – Holland, ex Myodes [Clethrionomys] glareolus
Speleorodens michigensis (Ford): OConnor, 1978: 124
Type locality: Ingham County, Michigan
Distribution: Holland, USA
Hosts: Microtus agrestis, M. pennsylvanicus, Myodes [Clethrionomys] gapperi, Myodes [C.] glareolus (Rodentia: Arvicolidae)
Paraspeleognathopsis (Speleorodens) neotropicalis Fain & Aitken, 1969: 37
Type locality: Jandiai, Caratateua, Bragança (Pará), Brazil
Distribution: Brazil
Host: Zygodontomys lasiurus fuscinus (Rodentia: Cricetidae)
Speleorodens ruandae Fain, 1962b: 367
Type locality: Butare (= Astrida)
Distribution: Rwanda
Host: «dans une termitière«, accidental
Speleognathopsis strandtmanni Fain, 1955b: 696; 1956c: 645
Speleognathopsis sciuri Clark, 1960: syn. by Fain, 1961a: 160
Speleognathopsis (Neospeleognathopsis) strandtmanni (Fain): Fain, 1961a: 160
Paraspeleognathopsis (Speleorodens) strandtmanni (Fain): Fain, 1963c: 64
Speleorodens strandtmanni (Fain): Fain, 1970c: 518
Type locality: Astrida (= Butare), Rwanda
Distribution: Belgium, DR Congo, Panama, Rwanda, USA
Hosts: Sciurus granatensis, S. vulgaris, S. carolinensis, Funisciurus carruthersi, Tamiscus emini (Rodentia: Sciuridae)
Speleognathini Fain, 1985a: 144
Type genus: Speleognathus Womersley, 1936
Domrownetes OConnor, 1978: 128
Type species: Paraspeleognathopsis exul Domrow, 1975
Paraspeleognathopsis exul Domrow, 1975: 97; 1991: 1298
Domrownetes exul (Domrow): OConnor, 1978: 97
Type locality: Mount Sabine, Otway Ranges, Victoria
Distribution: Australia (Victoria)
Host: Antechinus stuartii (Dasyuromorphia: Dasyuridae)
Speleognathus Womersley, 1936: 313
Type species: Speleognathus australis Womersley, 1936
Speleognathus australis Womersley, 1936: 313; 1954; Domrow, 1961: 374; 1991: 1302; Fain, 1955b: 695; 1956c: 662; 1961; 1962 : 9; 1963c: 54; 1971 : 515; Drummond & Medley, 1964; Southcott, 1986: 201; Halliday, 1998: 51
Speleognathus (Speleognathus) australis Womersley: Fain, 1958a: 178
Speleognathus bovis Fain, 1955; 1956d: 156; 1956b: 18; 1956c: 662 (syn. de S. australis) - Type locality: Astrida (= Butare, Rwanda), ex Artiodactyla: Bovidae
Type locality: on water surface (not “in moss”, see Fain, 1956c: 662, at Glen Osmond, Adelaide, South Australia
Distribution: Australia, USA
Hosts: Bos taurus, Bison bison (Artiodactyla: Bovidae)
Lawrencarinae Fain, 1957: 251
Type genus: Lawrencarus Fain, 1957
Lawrencarini Fain
Type genus: Lawrencarus Fain, 1957
Batracarus Fain, 1961a: 254
Type species: Batracarus hylaranae Fain, 1961
Batracarus hylaranae Fain, 1961a: 254; 1962b: 58
Type locality: Bogor (= Buitenzorg), Java
Distribution: Indonesia (Java)
Host: Hylarana erythraea (Anura: Ranidae)
Lawrencarus Fain, 1957: 250
Type species: Riccardoella eweri Lawrence, 1952
Cernycarus de la Cruz, 1971: 2
Type species: Lawrencarus (Cernycarus) hollandsae de la Cruz, 1971
Lawrencarus (Cernycarus) hollandsae de la Cruz, 1971: 2
Type locality: Culandrillo
Distribution: Cuba
Host: Eleutherodactylus sp. (Anura: Ceratophryidae)
Lawrencarus Fain, 1957: 250
Type species: Riccardoella eweri Lawrence, 1952
Boydaia angelae Womersley, 1953: 83; Fain, 1955b: 694; 1956c: 647
Lawrencarus angelae (Womersley): Fain, 1961a: 246; Domrow, 1961: 379; 1991: 1293; Fain, 1962 :
Type locality: South Australia
Distribution: Australia
Hosts: Limnodynastes tasmaniensis, L. peronii (Anura: Ceratophryidae, Cycloraninae)
Lawrencarus americanus Fain, 1961a: 250
Type locality: Florida
Distribution: USA (Florida)
Host: Hyla septentrionalis (Anura: Hylidae)
Lawrencarus brasiliensis Fain, 1961a: 252; 1962: 52
Type locality: Alto da Serra, nr. Cubatan, State of São Paulo
Distribution: Brazil
Hosts: Cyclorhamphus asper, Eleutherodactylus guntheri (Anura: Ceratophryidae,
Ceratophryinae)
Lawrencarus brasiliensis desantisi Mauri & Bischoff, 1984: 338
Type locality: Averias, Prov. De Santiago del Estero
Distribution: Argentina
Hosts: Bufo arenarum, B. paracnemis (Anura: Bufonidae)
Lawrencarus ceratobatrachi Fain, 1961a: 251; 1962 : 47
Type locality: Buin
Distribution: Bougainville I. (Papua New Guinea)
Host: Ceratobatrachus guntheri (Anura: Ranidae)
Lawrencarus domrowi Fain, 1961a: 253; 1962 : 54; Domrow, 1991: 1293
Type locality: Dinner Ck, Nord Queensland
Distribution: Australia (Queensland)
Host: frog (Anura)
Riccardoella eweri Lawrence, 1952: 747; Lufty, 1960: 183
Boydaia angelae : Fain, 1956: 647
Lawrencarus eweri : Fain, 1957: 250; 1958a: 183; 1961: 253; 1962: 20
Lawrencarus eweri cubanus de la Cruz, 1971: 3
Type locality: Soledad
Distribution: Cuba
Host: Bufo peltacephalus (Anura: Bufonidae)
Lawrencarus eweri dahuricus Zabludovskaya, 1992: 66
Type locality: Chitinskaya oblast, Borzyanskiy Rayon, Borzya River, loc. Arshan
Distribution: Russia
Host: Bufo raddei (Anura: Bufonidae)
Lawrencarus eweri eweri (Lawrence): Fain, 1962 : 21
Type locality: Natal Museum, Pietermaritzburg, South Africa
Distribution: DR Congo, Egypt, Italy, Morocco, Rwanda, South Africa
Hosts: Bufo regularis, B. mauretanicus, B. carens, B. kisoloensis, B. superciliaris, Bufotes v. viridis (Anura: Bufonidae)
Lawrencarus eweri thailandiae Fain, 1961a: 252; 1962: 24
Type locality: Pak Shong
Distribution: Thailand
Host: Bufo melanostictus (Anura: Bufonidae)
Lawrencarus hylae Fain, 1961a: 247; Domrow, 1974: 18; 1991: 1293
Lawrencarus (L.) hylae hylae Fain
Lawrencarus hylae hylae : Fain, 1962: 33
Type locality: Queensland
Distribution: Australia, North America, Brazil
Hosts: Hyla coerulea, H. hayi, H. nasuta (Anura: Hylidae)
Lawrencarus afrixali Fain, 1961a: 251
Lawrencarus hylae afrixali : Fain, 1962: 37
Type locality: Virunga Nat. Park (formerly Albert N.P.)
Distribution: DR Congo
Hosts: Afrixalus fulvovittatus leptosomus, Hyperolius castaneus, Phlyctimanus verrucosus (Anura: Hyperoliidae)
Lawrencarus hylae intermedius Fain, 1962: 39; Mauri & Bischoff, 1984: 338
Type locality: Alto da Serra, Cubatan, Sao Paolo Prov. (Brazil)
Distribution: Argentina, Brazil
Hosts: Hyla c. cinerea, H. hayii (Anura: Hylidae), Pleurodema sp. (Anura: Leptodactylidae)
Lawrencarus hylae brevicipitis Fain, 1962: 41
Type locality: Amari, Usumbara (DR Congo)
Distribution: DR Congo, India
Hosts: Probreviceps macrodactylus, Ramanella triangularis (Anura: Microhylidae, Microhylinae, Brevicipitinae)
Lawrencarus lechriodi Fain, 1962: 31
Type locality: New Guinea
Distribution: New Guinea
Host: Lechriodus platyceps (Anura: Ceratophryidae, Cycloraninae)
Xenopacarini Fain, Baker & Tinsley, 1969: 341
Type genus: Xenopacarus Fain, Baker et Tinsley, 1969
Xenopacarus Fain, Baker & Tinsley, 1969: 341
Type species: Xenopacarus africanus Fain, Baker et Tinsley, 1969.
Xenopacarus africanus Fain, Baker & Tinsley, 1969: 341
Type locality: South Africa (imported into Leeds, England)
Distribution: South Africa
Host: Xenopus laevis (Amphibia: Pipidae, Xenopodinae)
Xenopacarus kenyensis Fain & Tinsley, 1975: 854
Type locality: Kiambu, Nairobi; Marsabit
Distribution: Kenya
Host: Xenopus borealis (Amphibia: Pipidae, Xenopodinae)
Xenopacarus kivuensis Fain & Tinsley, 1993: 513
Type locality: Ebisha, about 8 km from R.St. Irangi, Kivu Prov.
Distribution: D.R. Congo
Host: Xenopus sp. group fraseri (Amphibia: Pipidae, Xenopodinae)
EUROPE
Armenia (Arutyunyan, 1972, 1981) – Boydaia sturni, Riccardoella limacum,
Austria (Willmann, 1951; Franz, 1954; Sixl, 1972) – Ereynetes papilosus, E. biscutatus, Paraspeleognathopsis bakeri, Riccardoella oudemansi
Belgium (Cooreman, 1947; Fain, 1958b, 1962 , 1963c; Fain & Bafort, 1973; Fain & Lukoschus, 1968: 85; Fain & van Goethem, 1986; André & Ducarme, 2003; André et al., 2004) – Ereynetes aurantipes, E. gandensis, E. hydrophilus, Aureliania aureliani, Boydaia loxiae, B. nigra motacillae, B. n. nigra, B. sturni, Hanriccardoella faini, Paraspeleognathopsis bakeri, P. strandtmanni, Pseudotydeus lebruni, Pterniboydaia pternistis, Riccardoella oudemansi, R. limacum
Bulgaria (Turk, 1970; Beron, 1973) – Ereynetes (Anereynetes) simplex, Paraspeleognathopsis bakeri, Riccardoella limacum, R. oudemansi
Croatia (Willmann, 1941; Pernek et al., 2008) – Ereynetes scutulis, Riccardoella oudemansi
Czech Rep. (Willmann, 1936; Zacharda, 1978; André et al., 2004;) – Ereynetes (Anereynetes) simplex, Riccardoella oudemansi, R. zadielensis
Yan Mayen I. (Macfadyen, 1954) – Ereynetes [Protereunetes] brevipes
Finland (Penttinen, Viiri & Moser, 2013) – Ereynetes propescutulis
France (contin.) (Megnin, 1889; Fain & van Goethem, 1986; Morand & Deguzan, 1986) – Riccardoella reaumuri, R. limacum
Germany (Willmann, 1952, 1953; Fritsch, 1962; Schmidt, 1987) – Boydaia willmanni, Ereynetes bipilosus, E. insularis, E. lyratus
Great Britain (Turk & Phillips, 1945; White, 1959; Baker, 1970a, b; Fain & van Goethem, 1986; Graham et al., 1993) – Riccardoella limacum, R. oudemansi,
Greece (Haralabidis et al., 1983) - Riccardoella limacum
Holland (Oudemans, 1912, 1928; Thor, 1932; Van Eyndhoven, 1955; Fain & Lukoschus, 1968: 85; OConnor, 1978; Mienis, 1984; Fain & van Goethem, 1986) – Boydaia saxicolae, Ereynetes sittardiensis, Paraspeleognathopsis bakeri, Ricardoella oudemansi, Speleorodens michigensis (= clethrionomys)
Hungary (András & Jenö, 2004; Varga & Kontschan, 2004) - Riccardoella
oudemansi
Iceland (Sellnick, 1940) – Riccardoella oudemansi
Ireland (Fauna Europaea) – 1 sp.
Italy (contin.)(Berlese, 1923; Bottazzi, 1950; Fain, 1964b, 2004) – Ereynetes arcuatus, E. biscutatus, E. brevipes, E. centralis, E. inermis, E. ministralis, E. pegazzanoae, E. septentrionalis, Protereunetes brevipes, Riccardoella limacum, R. oudemansi
Sicily (Berlese, 1923) – Ereynetes agilis, E. galeatus
N. Macedonia (Haralabidis et al., 1983) – Riccardoella limacum
Norway (Thor, 1932) – Ereynetes norvegicus
Poland (Karbarz-Wiktorowicz, 1973; Dończyk, 2006) – Ereynetes (Opsereynetes) norvegicus, Ereynetes (Gymnereynetes), Riccardoella limacum
Russia (all, continent.)(Dubinin, 1957; Zabludovskaya, 1985, 1986, 1990a, 1990b, 1991, 1992, 1998, 2012) – Boydaia sturni, B. bradornis, B. jordani, B. psalidoprocnei, Coboydaia nigra nigra, C. nigra motacillae, Neoboydaia phylomachi eroliae, Ophthalmognathus schoutedeni (= dogieli), Phoenicopteriella womersleyi, Lawrencarus eweri dahuricus
Crimea (Zabludovskaya, 1994) - Ereynetes (E.) g. galeatus, E. (G.) exilis, E. (Gymnereynetes) aurantipes, E. (E.) tshernogoriensis
Serbia (Stojnić et al., 2016) – Riccardoella limacum, R. oudemansi
Slovakia (André et al., 2004; Shoaib & Cagáň, 2004) – Riccardoella zadielensis
Spain (cont.) (Mihelčič, 1958; Fain & Lukoschus, 1968: 85) – Ereynetes (Anereynetes) corticalis, E. (A.) ornatus, E. (A.) proximus, Paraspeleognathopsis bakeri
Sweden (Moser et al., 1989) – Ereynetes propescutulis
Switzerland (Schweizer, 1922, 1963) – Ereynetes ministralis, Riccardoella limacum
Turkey (all) (Çobanoğlu, Cilbircioğlu & Zabludovskaya, 2020) – Ereynetes septentrionalis
Ukraine (Zabludovskaya, 1994, 1995, 2002, 2005, 2012) – Boydaia psalidoprocnei, Coboydaia nigra, Ereynetes (E.) gandensis, E. (E.) tshernogoriensis, E. (E.) pegazzanoe, E. (E.) punctatus, E.(Anereynetes) hydrophilus, E. (A.) sittardiensis, E. (Gymnereynetes) aurantipes, E. (G.) exilis, Paraspeleognathopsis bakeri, Riccardoella canadensis
AFRICA
Botswana (Fain, 1959, 1960, 1969; Fain & Camerik, 1994; Skoracki et al., 2018) – Boydaia spatulata, Coboydaia nigra, Ereynetes copridis, E. botswanensis
Burundi (Fain, 1963 ; Skoracki et al., 2018) - Coboydaia clavata, C. nigra
Cameroon (Fain, Herin & Puylaert, 1977; Zabludovskaya, 2002) – Boydaia hirundoae, B. pycnonoti, Coboydaia clavata, C. nigra
Chad (Fain, 1963) – Boydaia falconis
Congo DR (Fain, 1961a, 1964 ; 1966: 108; 1970b; Fain & Tinsley, 1993; Skoracki et al., 2018) – Hipposideroptes kimuenzae, Neoboydaia galachrysiae, Neospeleognathopsis chiropteri, Xenopacarus kivuensis
Egypt (Fain, 1963c, 1966: 108; Lufty, 1960; Abdelgayed et al., 2019) – Ereynetes bolitophilus, Lawrencarus eweri eweri, Neospeleognathopsis chiropteri
Ghana (Skoracki et al., 2018) - Coboydaia clavata
Guinea (Zabludovskaya, 2002) – Coboydaia clavata, C. nigra, Pycnonotus barbatus
Kenya (Fain & Tinsley, 1975; Skoracki et al., 2018) – Boydaia pycnonoti, Xenopacarus kenyensis
Rwanda (Cooreman, 1954; Fain, 1955b, 1956a, 1956b, 1956d, 1962, 1963, 1969, 1971, 1972, 1985a, 1985b; Pence & Casto, 1976, Fain & Goff, 1980; Skoracki et al., 2018) – Astrida caprimulgi, Boydaia bradornis, B. buphagi, B. cecropis, B. cinnyris, B. duboisi, B. falconis, B. psalidoprocnei, B. saxicolae, B. spatulata, B. zosteropis, B. zumpti, Coboydaia clavata, C. nigra, Meropiboydaia merops, Neastrida poffei, Neoboydaia philomachi, Ophthalmognathus tenorioae, Phoenicopteriella womersleyi, Pterniboydaia pternistis, Speleorodens strandtmanni
Senegal (Fain, 1963) – Coboydaia clavata
South Africa (Fain, 1956, 1958, 1959, 1963c, 1971; Fain, Baker & Tinsley, 1969; Fain, Herin & Puylaert, 1977; Fain & Camerik, 1994; Ueckermann & Tiedt, 2003; Skoracki et al., 2018) – Boydaia pycnonoti, Ereynetes aplectorus, E. bolitophilus, E. luriei, E. ueckermanni, Lawrencarus eweri eweri, Coboydaia nigra, Neastrida parrae, Neoboydaia psittaculae, Riccardoella limacum, R. oudemansi, Xenopacarus africanus
Zambia (Fain, 1960 , 1963) – Coboydaia clavata, Pterniboydaia pternistis
Zimbabwe (Skoracki et al., 2018) - Boydaia pycnonoti, Coboydaia clavata
ASIA
China (Li, Xuan & Fan, 1992;) – Ereynetes sp.
India (Fain, 1969; Bhat, Kulkarni & Mishra, 1983; Panda, Mishra & Panda, 1987) – Boydaia indica, Speleognathopsis galli
Indonesia (Berlese, 1910; Vitzthum, 1932, Fain, 1961 , 1963) - Batracarus hylaranae, Coboydaia amandavae [Java], Ereynetes cruciatus [Java], E. potator [Sumatra]
Iran (Siahkamari et al., 2020) – Ereynetes [Ereynetoides] amplectorus
Israel and Palestina (Mienis, 1990) – Riccardoella sp.
Japan (Berlese, 1923; Kaneko, 1968; Shiba, 1969а, 1971; Kaneko, Kadosaka & Toshioka, 1977; Kaneko, Matsudaira & Prince Masahito, 1978; Uchikawa & Suzuki, 1984; Kadosaka, Kaneko & Asanuma, 1983, 1987; Waki, Hiruta & Shimano, 2018; Waki et al., 2018; Waki & Shimano, 2019; Waki, Shimano & Asami, 2019; Waki, Shimano & Shiba, 2019; Waki et al., 2020) – Ereynetes cavernarum, E. excellens, E. (Opsereynetes) norvegicus, Speleognathopsis (Speleomyotis) bastini, Paraspeleognathopsis bakeri, Pterniboydaia pternistis, Riccardoella (Proriccardoella) triodopsis, R. tokyoensis, Trispeleognathus womersleyi
Okinawa (Waki et al., 2018) - Riccardoella reaumuri
Kazakhstan (Zabludovskaya, 1998) - Coboydaia nigra
Kyrghyzstan (Zabludovskaya, 2002) - Coboydaia nigra motacillae
Korea (Fain & Lukoschus, 1968; Shiba, 1969b) – Paraspeleognathopsis bakeri
Malaysia (Fain, 1963c, Fain & Nadchatram, 1962; Shiba, 1976; McClure, Ratanaworadhan et al., 1973) – Astrida caprimulgi, Boydaia pycnonoti, B. spatulata, Ereynetes husatoi, E. malayi, E. tadakii
New Guinea (Fain, 1962) - Lawrencarus lechriodi
Philipppines (Corpuz-Raros, 1975) – Ereynetes philippinensis, Ereynetes (Anereynetes) upelbensis
Turkmenistan (Zabludovskaya, 1998) – Coboydaia nigra
Thailand (Fain, 1961a) – Lawrencarus eweri thailandiae
NORTH AMERICA
Canada (Fain & van Goethem, 1986; Knee, Proctor & Galloway, 2008) – Boydaia aratingae, B. faini, B. psalidoprocnei, B. sturni, Neoboydaia colymbiformi, Riccardoella canadensis,
Mexico (Baker, 1945, Clark, 1960; Fain, 1963c; Fain & Hyland, 1970; Polaco & Mendl, 1988; Vásquez-Rojas & Estrada-Venegas, 2010) – Boydaia mimi, Ereynetes (Opsereynetes) robustus, E. (O.) simplexus, E. (Opsereynetes) tuberculatus, E. (Anereynetes) sabinensis, E. (Ereynetes) amplectorus, E. (E.) faini, Riccardoella oudemansi, Trispeleognathus striatus
USA (Boyd, 1948, 1951; Clark, 1958, 1960, 1964, 1967a, 1967b; Hunter, 1964; McDaniel, 1967; Hunter & Cross, 1968; Kethley, 1971; Pence, 1973a, 1973b, 1973c; Baker & Delfinado,1974; Pence & Casto, 1976, Spicer, 1977; Fain & Klompen, 1990; Hilario-Pérez & Danel, 2016; Hilario-Pérez & Dowling, 2018) – Astrida caprimulgi, A. coccyzae, Boydaia agelaii, B. colini, B. hirundoae, B. mimi, B. pheucticola, B. psalidoprocnei, B. quiscali, B. spatulata, B. sturnellae, B. sturni, B. turdi, Coboydaia nigra, Ereynetes amplectorus, E. boharti, E. davisi, E. scutulis; Hydranetes citrinensis, H. tropisternus; Neoboydaia philomachi, Ophthalmognathus womersleyi, Pseudotydeus perplexus, Riccardoella (Proriccardoella) triodopsis, Ricardoella (R.) limacum, Trispeleognathus striatus, Speleognathopsis benoiti
Hawai Is. (Fain & Goff, 1980) – Coboydaia nigra, Ophthalmognathus tenorioae,
SOUTH and CENTRAL AMERICA
Argentina (Rosas Costa, 1955; Mauri & Bischoff, 1984) – Aureliania longipilis,
Lawrencarus brasiliensis desantisi, L. hylae intermedius
Brazil (Amaral, 1963, 1965, 1967; Fain, 1962, 1963 ; Fain & Aitken, 1969, 1970; Faccini et al., 1981; Flechtmann & Baggio, 1985; Mascarenhas & Brum, 2006; Bernardon et al., 2015; Bernardon, Mascarenhas et al., 2018; Mendoza – Roldan, 2019) – Boydaia agelaii, B. americana, B. aratingae, B. cyanerpes, B. laticoxa, B. psittaci, B. quiscali, B. synallaxis, B. tyrannis, Coboydaia trochila, Lawrencarus brasiliensis, Neoboydaia lateralli, Speleochir barbulata, S. brasiliensis, Speleomys neotropicalis, Neospeleognathopsis (Speleomyotis) bastini didelphis, Riccardoella limacum, Speleognathopsis galli, Trispeleognathus euricoi, T. striatus
Cuba (Dusbábek & Černy, 1970; de la Cruz, 1971) – Boydaia faini, B. morenoi, B. rosickyi, Lawrencarus (L.) eweri cubanus, L. (Cernycarus) hollandsae
Panama (Clark, 1967a) - Speleorodens strandtmanni
Puerto Rico (Hunter & Poe, 1971) – Ereynetes coenobitus
Suriname (Fain & Lukoschus, 1971, 1972) – Boydaia agelaii, Hipposideroptes saccopteryx, Neospeleognathopsis (Speleomyotis) bastini bastini, N. (S.) molossus, Speleochiroides carollia, Psittaboydaia amazona
Trinidad and Tobago (Fain & Aitken, 1968) - Boydaia agelaii, B. americana, B. hirundoae, B. tyrannis, Trispeleognathus striatus
OCEANIA
Australia (continent) (Rainbow, 1906; Domrow, 1960, 1962, 1965, 1966, 1967, 1969, 1975, 1991; OConnor, 1978; Fain & Lukoschus, 1968, 1979; Womersley, 1936, 1953, 1954) – Aureliania aureliani, Boydaia spatulata, Domrownetes exul, Meropiboydaia merops, Neoboydaia philomachi, N. psittaculae, Paraspeleognathopsis bakeri, P. exaeta, Speleorodens derricki, Pteropignathus pteropi, Speleognathopsis benoiti, Riccardoella limacum (probably R. oudemansi, see Fain, 2004)
New Zealand (Barker & Ramsay, 1978; Fain & Barker, 2003, 2004) – Austreynetes maudensis, Riccardoella (Riccardoella) limacum, Riccardoella (Proriccardoella) novaezealandiae
Macquarie I. (Fain, 1962d; Luxton, 1990) – Ereynetes (Gymnereynetes) macquariensis
Papua New Guinea (Domrow, 1969; Talbot, 1969; Fain & Van Goethem, 1978) – Ereynetes (Anereynetes) papuanus, Lawrencarus ceratobatrachi, Speleognathopsis galli
ANTARCTICA
Kerguelen I. (Fain, 1976) - Ereynetes macquariensis
Marion I. (Fain, 1976) - Ereynetes macquariensis
Signy I. (Strandmann & Tilbrook, 1968) - Ereynetes macquariensis
Antarctic Peninsula (Strandtmann, 1967; Crooker, 1977; Llano & Gless, 2013) - Ereynetes macquariensis, Protereunetes minutus, P. paulinae
Mollusca
Gastropoda
Pulmonata
Fam. Arionidae
Arion ater - Riccardoella oudemansi
A. lusitanicus - Riccardoella oudemansi
A. rufus - Riccardoella oudemansi
Fam. Athoracophoridae
Pseudaneitea schauinslandi - Austreynetes maudensis
Athoracophorus bitentaculatus – Riccardoella limacum
Athoracophoridae gen. sp. – Riccadoella (Proriccardoella) novaezealandiae
Fam. Clausiliidae
Tauphaedusa tau – Riccardoella tokyoensis
Fam. Bradybaenidae
Acusta despecta sieboldtiana – Riccardoella (Proriccardoella) triodopsis
Fam. Helicidae
Cepaea nemoralis - Riccardoella oudemansi
Helix lucorum - Riccadoella limacum
H. pomatia - Riccadoella limacum
H. aspersa - Riccadoella limacum
Fam. Limacidae
Deroceras reticulatum - Riccardoella oudemansi
Limax agrestis - Riccardoella oudemansi
L. cinereoniger - Riccardoella oudemansi
L. cinereus - Riccardoella oudemansi
L. flavus - Riccardoella oudemansi
L. maximus - Riccardoella oudemansi
L. cinereoniger - Riccardoella oudemansi
Monochroma brunneum – Riccardoella limacum
Fam. Milacidae
Tandonia budapestensis - Riccardoella oudemansi
Fam. Polygiridae
Xolotrema obstructa [Triodopsis obstrictus] – Riccardoella (Proriccardoella) triodopsis
Fam. Zonitidae
Meledella werneri – Riccardoella oudemansi
Oxychilus draparnaudi - Riccardoella oudemansi
Phylum Arthropoda
Subphylum Crustacea
Order Decapoda
Infraorder Brachiura
Fam. Coenobitidae
Coenobita clypeatus – Ereynetus coenobitus
Insecta
Order Coleoptera
Fam. Hydrophilidae
Tropisternus lateralis nimbatus - Hydranetes citrinensis
Tropisternus lateralis nimbatus - Hydranetes tropisternus
T. striolatus - Hydranetes tropisternus
T. quadristriatus – Hydranetes tropisternus
Fam. Scarabaeidae
Copris subsidens - Ereynetes (Anereynetes) copridis
C. molossus – Ereynetes cruciatus
Metacatharsius sp. - Ereynetes (Anereynetes) botswanensis
Scarabaeidae indet. – Ereynetes philippinensis
Order Hymenoptera
Fam. Apidae
Melipona scutellaris - Ereynetes (Gymnereynetes) meliponae
Subphylum Vertebrata
Class Amphibia
Order Anura
Fam. Bufonidae
Bufo arenarum - Lawrencarus brasiliensis desantisi
B. melanostictus - Lawrencarus eweri thailandiae
B. regularis - Lawrencarus eweri eweri
B. mauretanicus - Lawrencarus eweri eweri
B. carens - Lawrencarus eweri eweri
B. kisoloensis - Lawrencarus eweri eweri
B. paracnemis – Lawrencarus brasiliensis desantisi
B. peltacephalus - Lawrencarus eweri cubanus
B. raddei - Lawrencarus eweri dahuricus
B. superciliaris - Lawrencarus eweri eweri
Bufotes v. viridis - Lawrencarus eweri eweri
Fam. Ceratophryidae
Cyclorhamphus asper - Lawrencarus brasiliensis
Eleutherodactylus guntheri - Lawrencarus brasiliensis
Eleutherodactylus sp. – Lawrencarus (Cernycarus) hollandsae
Lechriodus platyceps - Lawrencarus lechriodi
Limnodynastes tasmaniensis - Lawrencarus angelae
L. peronii - Lawrencarus angelae
Fam. Hyperoliidae
Afrixalus fulvovittatus leptosomus - Lawrencarus hylae afrixali
Hyperolius castaneus - Lawrencarus hylae afrixali
Phlyctimanus verrucosus - Lawrencarus hylae afrixali
Fam. Hylidae
Hyla cinerea cinerea - Lawrencarus hylae intermedius
H. coerulea - Lawrencarus hylae hylae
H. hayii - Lawrencarus hylae intermedius
H. nasuta - Lawrencarus hylae hylae
H. septentrionalis – Lawrencarus americanus
Fam. Leptodactylidae
Pleurodema sp. - Lawrencarus hylae intermedius
Fam. Microhylidae
Probreviceps macrodactylus – Lawrencarus hylae brevicipitis
Ramanella triangularis - Lawrencarus hylae brevicipitis
Fam. Pipidae
Xenopus borealis – Xenopacarus kenyensis
X. laevis – Xenopacarus africanus
Xenopus sp. group fraseri – Xenopacarus kivuensis
Fam. Ranidae
Hylarana erythraea - Batracarus hylaranae
Ceratobatrachus guntheri - Lawrencarus ceratobatrachi
Class Aves
Order Anseriformes
Fam. Anatidae
Aythia affinis - Phoenicopteriella womersleyi
Anas acuta – Phoenicopteriella womersleyi
A. clypeata - Phoenicopteriella womersleyi
A. discors – Phoenicopteriella womersleyi
A. strepera – Phoenicopteriella womersleyi
Dendrocygna viduata – Phoenicopteriella womersleyi
Sarkidiornis melanotos – Phoenicopteriella womersleyi
Order Caprimulgiformes
Fam. Caprimulgidae
Antrostomus carolinensis - Astrida caprimulgi
Scotornis fossii welwitschi - Astrida caprimulgi
Fam. Podargidae
Podargus strigoides – Boydaia podargi
Order Charadriiformes
Fam. Charadriidae
Afribyx senegallus - Speleognathopsis benoiti
Charadrius melanops - Speleognathopsis benoiti
Ch. pecuarius – Speleognathopsis charadricola
Erythrogonys cinctus - Speleognathopsis benoiti
Hemiparra crassirostris – Speleognathopsis benoiti
Lobibyx miles - Speleognathopsis benoiti
Philomachus pugnax – Neoboydaia philomachi
Pluvialis squatarola - Speleognathopsis benoiti
Tringa glareola - Neoboydaia philomachi
Fam. Glareolidae
Glareola [Galachrysis] cinerea – Neoboydaia galachrysiae
G. nuchalis - Neoboydaia galachrysiae
Rhinoptilus africanus – Speleognathopsis benoiti (= wai)
Fam. Jacanidae
Actophilornis africanus – Neastrida parrae
Fam. Scolopacidae
Actitis macularius - Neoboydaia philomachi
Arenaria interpres - Neoboydaia philomachi
Calidris melanotos - Neoboydaia philomachi
C. minutilla - Neoboydaia philomachi
Gallinago gallinago - Neoboydaia philomachi, Boydaia willmanni
Limnodromus griseus - Neoboydaia philomachi
L. scolopaceus - Neoboydaia philomachi
Limosa lapponica – Neoboydaia philomachi
Neoboydaia philomachi - Neoboydaia philomachi
Philomachus pugnax - Neoboydaia philomachi
Tringa flavipes - Neoboydaia philomachi
T. melanoleuca - Neoboydaia philomachi
T. glareola - Neoboydaia philomachi
T. solitaria - Neoboydaia philomachi
Fam. Laridae
Sterna hirundo - Neoboydaia philomachi
Thalasseus maximus - Neoboydaia philomachi
Order Columbiformes
Fam. Columbidae
Columba livia domestica – Trispeleognathus striatus
Columbigallina passerina – Trispeleognathus striatus
Geotrygon montana – Trispeleognathus striatus
Leptotila verreauxi - Trispeleognathus striatus
Streptopelia s. semitorquata – Trispeleognathus striatus
S. vinacea - Coboydaia clavata
Order Coraciiformes
Fam. Coraciidae
Coracias caudatus caudatus – Metaboydaia poffei
Fam. Meropidae
Merops apiaster – Meropiboydaia merops
M. ornatus - Meropiboydaia merops
Order Cuculiformes
Fam. Cuculidae
Guira guira - Carcinopodacarus polymorphus
Order Falconiformes
Fam. Accipitridae
Accipiter fasciatus – Speleognathopsis accipitris
Fam. Falconidae
Falco berigora - Boydaia falconis
F. cuvieri – Boydaia falconis
F. t. tinnunculus – Boydaia falconis
Order Galliformes
Fam. Odontophoridae
Callipepla squamata - Boydaia clarki
Colinus virginianus – Boydaia colini
Fam. Phasianidae
Bambusicola thoracica – Pterniboydaia pternistis
Gallus gallus domesticus – Speleognathopsis galli
Pternistis afer – Pterniboydaia pternistis
P. swainsoni – Pterniboydaia pternistis
Perdix p. perdix – Pterniboydaia pternistis
Fam. Numididae
Numida meleagris intermedia - Speleognathopsis galli
Order Gruiformes
Fam. Rallidae
Laterallus melanophaius oenops - Neoboydaia lateralli
Order Passeriformes
Fam. Acrocephalidae
Acrocephalus a. arundinaceus - Coboydaia nigra
A. s. scirpaceus - Coboydaia nigra
Iduna similis - Boydaia spatulata
Fam. Buphagidae
Buphagus africanus - Boydaia buphagi
Fam. Campophagidae
Lalage tricolor – Coboydaya nigra (sub Boydaia crassipes)
Fam. Cardinalidae
Spiza americana – Boydaia pheucticola
Fam. Cisticolidae
Schistolais leucopogon reichenowi - Boydaia spatulata
Fam. Dendrocolaptidae
Glyphorynchus spirurus - Boydaia laticoxa
Fam. Emberizidae
Emberiza aureola – Coboydaia nigra
E. citrinella – Coboydaia nigra
E. pusilla – Coboydaia nigra
E. rustica – Coboydaia nigra
E. rutila – Coboydaia nigra
E. spodocephala – Coboydaia nigra
E. tristrami – Coboydaia nigra
Spizella passerina – Coboydaia nigra
Fam. Estrildidae
Amandava amandava - Coboydaia amandavae
A. subflava - Coboydaia amandavae
Erythrura prasina - Coboydaia amandavae
Sporaeginthus subflavus - Coboydaia amandavae
Fam. Formicariidae
Formicarius analis - Boydaia formicarii
F. colma - Boydaia formicarii
Fam. Fringillidae
Carduelis c. carduelis - Coboydaia nigra
Carpodacus erythrinus – Coboydaia nigra
Euphonia violacea - Coboydaia nigra
Fringilla coelebs – Coboydaia nigra
F. montifringilla - Coboydaia nigra
Haemorhous mexicanus - Coboydaia nigra
Loxia curvirostra – Boydaia loxiae
Passerina cyanea – Boydaia rosickyi
Serinus burtoni tanganjicae - Coboydaia nigra
S. frontalis - Coboydaia nigra
S. [Poliospiza] gularis – Coboydaia nigra
S. mozambicus - Coboydaia nigra
S. sulphuratus shelleyi - Coboydaia nigra
Fam. Furnariidae
Philydor pyrrhodes - Boydaia laticoxa philydori
Synallaxis rutilans - Boydaia synallaxis
S. gujanensis - Boydaia synallaxis
S. erythrothorax - Boydaia synallaxis
Fam. Hirundinidae
Cecropis abyssinica unitatis – Boydaia cecropis
Hirundo abyssinica – Boydaia hirundoae
H. neoxena - Boydaia hirundoae
H. rustica - Boydaia hirundoae
Hylochelidon ariel - Boydaia hirundoae
Progne chalybea - Boydaia hirundoae
P. subis - Boydaia hirundoae
Psalidoprocne albiceps - Boydaia psalidoprocnei
Riparia riparia - Boydaia psalidoprocnei
Stelgidopteryx ruficollis - Boydaia hirundoae
Tachycineta bicolor - Boydaia psalidoprocnei
Fam. Icteridae
Agelaius icterocephalus – Boydaia agelaii
A. phoeniceus - Boydaia agelaii
A. ruficapillus - Boydaia agelaii
Chrysomus ruficapillus - Boydaia agelaii
Icterus galbula - Coboydaia nigra
I. spurius – Coboydaia nigra icteri
Quiscalus quiscula - Boydaia buphagi, B. quiscali
Q. mexicanus – Boydaia sturni
Sturnella magna – Boydaia buphagi, Coboydaia sturnellae
Fam. Laniidae
Corvinella corvina - Coboydaia nigra
Fam. Locustellidae
Locustella luscinioides - Boydaia locustellae
Schoenicola brevirostris – Boydaia zumpti
Fam. Meliphagidae
Cissomela pectoralis - Boydaia spatulata
Myzomela sanguinolenta – Boydaia myzomelae
Philemon citreogularis – Boydaia spatulata
P. corniculatus - Boydaia spatulata
Fam. Mimidae
Mimus gilvus – Boydaia mimi
M. polyglottos - Boydaia mimi
M. polyglottos orpheus – Boydaia morenoi
Fam. Motacillidae
Anthus spinoletta - Coboydaia nigra
A. trivialis – Coboydaia nigra motacillae
Macronyx croceus - Boydaia spatulata
Motacilla aguimp vidua - Coboydaia nigra motacillae
M. alba - Coboydaia nigra motacillae
M. capensis - Coboydaia nigra motacillae
M. flava - Coboydaia nigra motacillae
Fam. Muscicapidae
Agricola pallidus griseus - Boydaia bradornis
Cercomela tractrac - Boydaia spatulata
Erythropygia [Cercotrichos] hartlaubi – Boydaia spatulata
Luscinia luscinia - Boydaia bradornis
Muscicapa rufigastra - Boydaia spatulata
Pogonocichla stellate ruwenzorii - Boydaia spatulata
Sheppardia polioptera - Boydaia spatulata
Fam. Nectariniidae
Arachnothera l. longirostris - Ereynetes malayi
Chalcomitra senegalensis lamperti (=aequatorialis) - Boydaia cinnyris, B. nectarinia
Cinnyris cupreus cupreus (= septentrionalis) - Boydaia cinnyris, B. nectarinia
Cyanomitra verticalis viridisplendens - Boydaia nectarinia
Fam. Paridae
Melaniparus leucomelas insignis [as Parus niger insignis] – Boydaia spatulata
Fam. Parulidae
Dendroica cerulea - Boydaia faini
Fam. Passeridae
Passer ammodendri - Coboydaia nigra
P. domesticus – Coboydaia nigra
P. griseus - Coboydaia nigra nigra
P. hispaniolensis - Coboydaia nigra
P. montanus - Coboydaia nigra
Fam. Pipridae
Manacus manacus - Boydaia tyrannis
Pipra fasciicauda – Boydaia tyrannis
P. iris - Boydaia tyrannis
P. pipra - Boydaia tyrannis
Fam. Ploceidae
Euplectes hordeaceus – Coboydaia clavata
E. ardens - Coboydaia clavata
E. macroura - Coboydaia clavata
Hypochera amauropteryx - Coboydaia clavata
H. chalybeata - Coboydaia clavata
Othyphantes baglafecht stuhlmanni - Coboydaia clavata
Ploceus baglafecht stuhlmanni - Coboydaia clavata
P. [Textor] cucullatus - Coboydaia clavata
P. [Textor] cucullatus graueri - Coboydaia clavata
P. pelzelni - Coboydaia clavata
P. velatus - Coboydaia clavata
P. vitellinus - Coboydaia clavata
P. xanthops (= P. x. camburni) - Coboydaia clavata
Quelea quelea - Coboydaia clavata
Q. quelea centralis - Coboydaia clavata
Q. quelea aethiopica - Coboydaia clavata
Q. erythrops - Coboydaia clavata
Fam. Pycnonotidae
Bleda syndactyla woosnami – Boydaia zumpti
Euryllas latirostris latirostris - Boydaia zumpti
Pycnonotus barbatus tricolor - Boydaia pycnonoti
P. erythrophthalmus - Boydaia pycnonoti
P. jocosus - Boydaia pycnonoti
P. goiavier personatus - Boydaia pycnonoti
P. leucogenys leucotis - Boydaia pycnonoti
P. leucotis - Boydaia pycnonoti
P. simplex - Boydaia pycnonoti
P. xanthopygus - Boydaia pycnonoti
Fam. Sturnidae
Ampeliceps coronatus – Boydaia sturni
Acridotheres tristis – Boydaia indica, B. sturni
Lamprocolius chloropterus – Boydaia buphagi, B. indica
L. chalybaeus - Boydaia indica
Onychognathus walleri elgonensis - Astrida onychognathi
Spreo superbus - Boydaia indica
Sturnopastor contra - Boydaia indica
Sturnus malabaricus – Boydaia indica
S. pagodarum - Boydaia indica
S. vulgaris - Boydaia indica
Fam. Sylviidae
Malurus amabilis – Boydaia maluri
M. melanocephalus - Boydaia maluri
Fam. Thraupidae
Cyanerpes cyanea – Boydaia cyanerpes
Tangara sp. - Boydaia spatulata
Tiaris canorus - Boydaia spatulata
Fam. Turdidae
Oenanthe isabellina - Coboydaia nigra
Saxicola rubetra - Coboydaia clavata
Sialia sialis – Boydaia spatulata
Turdus migratorius – Boydaia jordani
T. pilaris - Boydaia jordani (= turdi), B. saxicolae
Fam. Tyrannidae
Empidonax sp. – Boydaia synallaxis
Pyrocephalus rubinua - Boydaia tyrannis
Tyrannus tyrannus – Boydaia tyrannis
Fam. Tytiridae
Pachyramphus [=Platypsaris] aglaiae - Boydaia tyrannis
Fam. Viduidae
Vidua chalybeata - Coboydaia clavata
V. chalybeata amauropteryx - Coboydaia clavata
Fam. Zosteropidae
Zosterops lateralis – Boydaia zosteropis
Z. senegalensis stuhlmanni (= scotti) - Boydaia zosteropis
Order Pelecaniformes
Fam. Ardeidae
Ardea cinerea - Ophthalmognathus schoutedeni (= O. dogieli)
Ardeola idae – Ophthalmognathus schoutedeni
Nycticorax caledonicus - Ophthalmognathus schoutedeni
N. nycticorax - Ophthalmognathus schoutedeni, O. tenorioae
N. n. hoactli - Ophthalmognathus tenorioae
Fam. Scopidae
Scopus umbretta – Metaboydaia poffei
Order Piciformes
Fam. Lybiidae
Buccanodon duchaillui - Picinyssus buccanodon
Order Podicipidiformes
Fam. Podicipedidae
Podiceps nigricollis – Neoboydaia colymbiformi
P. ruficollis - Neoboydaia colymbiformi
Podilymbus podiceps - Neoboydaia colymbiformi
Order Psittaciformes
Fam. Psittacidae
Amazona amazonica - Psittaboydaia amazona
Aratinga jandaya – Boydaia aratingae
Fam. Psittaculidae
Agapornis roseicollis - Psittaboydaia psittaculae
Psittacula krameri – Psittaboydaia psittaculae
Agapornis roseicollis - Psittaboydaia psittaculae
Trichoglossus haematod forsteni – Psittaboydaia trichoglossi
Trichoglossus moluccanus - Psittaboydaia psittaculae
Order Ralliformes
Fam. Rallidae
Latirallus melanophaius oenops – Ralliboydaia latiralli
Porphyrio melanotus – Ralliboydaia porphirionis
Order Strigiformes
Fam. Strigidae
Bubo africanus – Astrida caprimulgi
Glaucidium perlatum – Astrida caprimulgi
Otus senegalensis – Astrida caprimulgi
O. spilocephalus vulpes - Astrida caprimulgi
Fam. Tytonidae
Tyto alba affinis – Aureliana aureliani
T. alba alba - Aureliana aureliani
Class Mammalia
Order Dasiuromorphia
Fam. Dasyuridae
Antechinus stuartii - Domrownetes exul
Order Diprotodontia
Fam. Petauridae
Petaurus breviceps - Paraspeleognathopsis exaeta
Order Didelphimorphia
Fam. Didelphidae
Didelphis marsupialis – Speleomyotis bastini
Order Chiroptera
Fam. Pteropodidae
Eidolon helvum – Neospeleognathopsis chiropteri
Epomophorus labiatus minor – Neospeleognathopsis chiropteri
Pteropus scapulatus – Pteropignathus pteropus
Rousettus aegyptiacus - Neospeleognathopsis chiropteri
R. aegyptiacus leachi - Neospeleognathopsis chiropteri
Fam. Hipposideridae
Hipposideros commersoni gigas - Hipposideroptes kimuenzae
Fam. Nycteridae
Nycteris sp. – Neospeleochir duboisi
Fam. Phyllostomatidae
Anoura g. geoffroy – Speleochir aitkeni
Artibeus jamaicensis – Speleochir brasiliensis
Carollia perspicillata – Speleochiroides carollia
Phyllostomus hastatus – Speleochir phyllostomi
Vampyrodes carracioloi - Speleochir brasiliensis
Fam. Rhinolophidae
Rhinolophus cornutus – Neospeleognathopsis (Speleomyotis) bastini
Fam. Vespertilionidae
Eptesicus fuscus - Neospeleognathopsis (Speleomyotis) bastini
E. melanopterus - Neospeleognathopsis (Speleomyotis) bastini
Myotis myotis – Neospeleognathopsis (Speleomyotis) bastini
M. nigricans - Neospeleognathopsis (Speleomyotis) bastini
Fam. Molossidae
Cynomops planirostris – Neospeleognathopsis (Speleomyotis) molossi
Molossus ater - Neospeleognathopsis (Speleomyotis) molossi
M. molossus - Neospeleognathopsis (Speleomyotis) molossi
Fam. Emballonuridae
Saccopteryx bilineata – Hipposideroptes saccopteryx
S. leptura - Hipposideroptes saccopteryx
Order Rodentia
Fam. Arvicolidae
Microtus agrestis – Speleorodens michigensis (= clethrionomys)
M. pennsilvanicus - Speleorodens michigensis
Myodes [Clethrionomys] gapperi - Speleorodens michigensis
M. glareolus - Speleorodens michigensis (= clethrionomys)
Fam. Cricetidae
Neacomys guianae - Speleorodens cricetidarum
Zygodontomys lasiurus fuscinus - Speleorodens neotropicalis
Fam. Gliridae
Claviglis murinus microtis – Paraspeleognathopsis graphiuri
Fam. Muridae
Murinae
Apodemus agrarius coreae - Paraspeleognathopsis bakeri
A. flavicollis - Paraspeleognathopsis bakeri
A. speciosus – Paraspeleognathopsis bakeri
Arvicanthis abysinicus – Speleomys galliardi
Dasymys incomtus medius – Speleomys galliardi
Hydromys chrysogaster – Speleorodens derricki
Mastomys coucha ugandae - Paraspeleognathopsis bakeri
Melomys cervinipes - Paraspeleognathopsis bakeri
Mus musculus - Speleorodens derricki
Pelomys fallax concolor – Speleomys galliardi
Rattus assimilis - Speleorodens derricki
R. conatus – Speleorodens derricki
R. rattus – Speleorodens derricki
Sylvaemus sylvaticus – Paraspeleognathopsis bakeri
S. sylvaticus callipides – Paraspeleognathopsis bakeri
Gerbillinae
Gerbilliscus boehmi – Speleomys galliardi
Fam. Sciuridae
Funisciurus carruthersi – Speleorodens strandtmanni
Sciurus carolinensis – Speleorodens strandtmanni (= P. sciuri)
S. granatensis - Speleorodens strandtmanni
S. vulgaris - Speleorodens strandtmanni
Tamiscus emini – Speleorodens strandtmanni
Order Primates
Fam. Galagonidae
Galago senegalensis moholi - Paraspeleognathopsis galagoensis
Order Artiodactyla
Fam. Bovidae
Bos taurus – Speleognathus australis (= bovis)
Bison bison - Speleognathus australis
Paratydeoidea Baker, 1949
The superfamily contains only the family Paratydeidae.
Paratydeidae Baker, 1949: 119
Small family of 13 known species of six genera, described by Berlese (1910), Baker (1949, 1950), Theron, Meyer & Ryke (1969), Kuznetzov (1973), Soliman (1974), Kandeel (1992), Kandeel & Hoda (1992), Flechtmann (1992), Seeman & Walter (2000).
Paratydeidae Baker, 1949: 119
Type genus: Paratydeus Baker, 1949
Hexatydeus Soliman, 1974: 198
Type species: Hexatydeus aegyptiacus Soliman, 1974
Hexatydeus aegyptiacus Soliman, 1974: 198
Type locality: Dakahlia, El Manzala
Distribution: Egypt
Hexatydeus amabilis Kandeel, 1992: 6
Type locality: Egypt
Distribution: Egypt
Paratydeus Baker, 1949: 119
Type species: Paratydeus alexanderi Baker, 1949
Paratydeus alexanderi Baker, 1949: 120
Type locality: intercepted at Brownsville, Texas on twig with bananas from Mexico
Distribution: Mexico
Sacotydeus Theron, Meyer & Ryke, 1969: 698
Type species: Sacotydeus lootsi Theron, Meyer et Ryke, 1969
Sacotydeus lootsi Theron, Meyer & Ryke, 1969: 699
Type locality: Potchefstroom, Transvaal
Distribution: South Africa
Scolotydaeus Berlese, 1910: 214
Type species: Scolotydaeus bacillus Berlese, 1910
Neotydeus Baker, 1950: 289. Syn. by Delfinado & Baker, 1974: 207
Type species: Neotydeus ardisanneae Baker, 1950
Neotydeus ardisannae Baker, 1950: 291
Scolotydeus ardisannae (Baker): Syn. by Delfinado & Baker, 1974: 207
Type locality: Sanburn, Johnson Co., Illinois
Distribution: USA
Scolotydaeus bacillus Berlese, 1910: 214; Thor, 1932: 90; 1933: 44; Theron, Meyer & Ryke, 1969: 708
Type locality: Palermo (“in agri Panormitani”)
Distribution: Norway, Italy (incl. Sicily)
Scolotydaeus corticicola Flechtmann, 1992: 299
Type locality: Brazil
Distribution: Brazil
Scolotydaeus simplex Delfinado & Baker, 1974: 207
Type locality: New York, Long Island, Adirondack Oark
Distribution: U.S.A. (New York)
Tanytydeus Theron, Meyer & Ryke, 1969: 703
Type species: Tanytydeus cristatus Theron, Meyer et Ryke
Tanytydeus cristatus Theron, Meyer & Ryke, 1969: 703
Type locality: Potchefstroom, Transvaal
Distribution: South Africa, Crimea
Tanytydeus kakadu Seeman & Walter, 2000: 399
Type locality: NT, Kakadu Nat. Park, Baroalba Gorge, Gubara Walk
Distribution: Australia (Northern Territory)
Tanytydeus lamington , Seeman & Walter, 2000: 395
Type locality: Lamington Nat. Park, Mt Bithongabel
Distribution: Australia (Queensland)
Tanytydeus neocristatus Kandeel & Hoda, 1992: 311
Type locality: Egypt
Distribution: Egypt
Walytydeus Kuznetzov, 1973: 12
Type species: Walytydeus tauricus Kuznetzov, 1973
Walytydeus tauricus Kuznetzov, 1973: 13
Type locality: Crimea
Distribution: Crimea
EUROPE
Crimea (Kuznetzov, 1973) – Tanytydeus cristatus, Walytydeus tauricus
Italy (contin.)(Berlese, 1882)
Sicily (Berlese, 1910) - Scolotydaeus bacillus
Norway (Thor, 1932) - Scolotydaeus bacillus
AFRICA
Egypt (Kandeel, 1992; Kandeel & Hoda, 1992; Soliman, 1974) – Hexatydeus
aegypticus, H. amabilis, Tanytydeus neocristatus
South Africa (Theron, Meyer & Ryke, 1969) – Sacotydeus lootsi, Tanytydeus
cristatus
NORTH AMERICA
Mexico (Baker, 1949) - Paratydeus alexanderi
USA (Baker, 1950; Delfinado & Baker, 1974) - Scolotydaeus (= Neotydeus) ardisannae, S. simplex
SOUTH and CENTRAL AMERICA
Brazil (Flechtmann, 1992) - Scolotydaeus corticicola
OCEANIA
Australia (continent.)(Seeman & Walter, 2000) – Tanytydeus lamington, T. kakadu
The cohort includes (Walter et al, in Krantz & Walter, 2009) the superfamilies Anystoidea, Caeculoidea, Adamystoidea, Paratydeoidea, and Pomerantzioidea, all treated in the present volume.
Anystoidea Cunliffe, 1955: 213; Southcott, 1957: 173
Phalanx Hamatostomata Wainstein, 1965: 77
In this superfamily are included the families Anystidae, Erythracaridae, Teneriffiidae, Pseudocheylidae, and Stigmocheylidae. They are predators.
The Anystids (s.lat.) are fast moving predators, moderately large (500 – 1500 µm), preying upon small mites and insects. Most often they are found on the soil surface or on plants. As early as in 1927 it was shown by Hirst that they can reach the highest mountains (Tibet, Mount Everest, 4950 m).
Acarus baccarum Linnaeus, 1758 and A. salicinus (now Anystis) were the first members of Anystidae to be described. Since the description of the genus Anystis von Heyden, 1826 on the basis of Trombidium cornigerum Hermann, 1804, many people contributed with new descriptions of Anystids: Haller (1882), Berlese (1882, 1886, 1903, 1905, 1923, described the genera Tarsotomus, Erythracarus), Thor (1912) described Tarsolarkus, Banks (1916), Schweizer (1922), Hirst (1927, 1931 - described Anandia), Trägårdh (1905, 1931). Among the early students of Anystidae are also worth mentioning Dugès (1834), C.L. Koch (1836, 1847), Stoll (1886, 1887).
It was Oudemans, who revised the family in 1936, describing nine new genera (Autenriethia, Barellea, Walzia, Scharfenbergia, Snartia, Tencateia, Chaussieria, Bechsteinia, Siblyia) and new species and dividing the family into two subfamilies Anystinae and Erythracarinae, the only recognized ever since.
After 1936 the most important contributions were the partial revision of Meyer & Ueckermann (1987) in South Africa (with the descriptions of the new genera Namandia and Paranandia and of 7 new species) and the revision of Erythracarinae of Otto (1999b), of Erythracarus of Otto (1999a), of Chaussieria (Otto, 1999c) and of Tarsotomus and Paratarsotomus (Otto, 1999d), in Australia. Other acarologists who contributed to the system or the biology of Anystidae after 1936 were Evans & Browning, Barilo, Naudo, Willmann (described the genus Absoloniana), Reda, Kuznetsov (described the genus Paratarsotomus), Womersley and others.
Finally. Pepato & Klimov (2015) upgraded the subfamily Erythracarinae on family level.
Fam. Anystidae Oudemans, 1902 - 16 gen., 102 sp. and Fam. Erythracaridae Oudemans – 8 gen., 68 sp.
Subfam. Anystinae Oudemans, 1936 - 8 gen., 34 sp.
Anystis von Heyden, 1826 - 21 sp.
(= Actineda C.L. Koch, 1836)
Autenriethia Oudemans, 1936 - 1 sp.
Barellea Oudemans, 1936 - 1 sp.
Mesoanystis Zacharda, 1985 - 1 sp. (fossil)
Scharfenbergia Oudemans, 1936 - 2 sp.
Snartia Oudemans, 1936 - 1 sp.
Tencateia Oudemans, 1936 - 3 sp.
Walzia Oudemans, 1936 - 5 sp.
Fam. Erythracaridae Oudemans: Pepato & Klimov, 2015 - 8 gen., 68 sp.
Chaussieria Oudemans, 1937 - 10 sp.
Erythracarus Berlese, 1903 - 19 sp.
Erythrocheylus Berlese, 1903 - 2 sp.
Namadia Meyer & Ueckermann, 1987 - 1 sp.
Paratarsotomus Kuznetsov, 1983 – 5 sp.
Siblyia Oudemans, 1936 - 1 sp.
Tarsolarcus Thor, 1912 - 3 sp.
Tarsotomus Berlese, 1882 - 27 sp.
(= Absoloniana Willmann, 1940)
(= Paranandia Meyer еt Ueckermann, 1987)
(= Anandia Hirst, 1927)
(= Parabsolonia Barilo, 1984)
(= Bechsteinia Oudemans, 1936)
(= Chabrieria Oudemans, 1936)
Anystinae Oudemans, 1902:
Anystidae Thor, 1903; Oudemans, 1909; 1936: 364
Erythracaridae Miller, 1925: 97
Type genus: Anystis von Heyden, 1826
Anystinae: Oudemans, 1936: 383
Type genus: Anystis von Heyden, 1826
Anystis von Heyden, 1826: 609
Type species: Trombidium cornigerum Hermann, 1804
Actineda C.L. Koch, 1836: 6
Type species: Actineda hilaris C.L. Koch, 1836
Actineda agilis Banks, 1894: 211; 1895: 432; 1903: 275
Anystis agilis (Banks): Banks, 1907: 598; Ewing, 1909: 432; Miller, 1925: 98; Cuthbertson & Murchie, 2007: 276
Type locality: USA
Distribution: Bermuda Is., Great Britain, Italy, USA
Anystis andrei Oudemans, 1936: 408
Type locality: Beausoleil, France, 350 m
Distribution: France
Acarus baccarum Linnaeus, 1758: 106
Actineda baccarum : Stoll, 1886
Anystis baccarum (L.): Trägårdh, 1905: 63; 1931: 624; Thor, 1912 : 388; Hull, 1918: 29; Vitzthum, 1926: 121; André, 1927: 168; Oudemans, 1906: 85; 1936: 387; Irk, 1939: 154; Willmann, 1939: 13; 1951b: 144; 1954: 236; Irk,1939: 154; Duduch et al., 1940: 54; Evans, 1953: 280; Franz, 1954: 367; Mihelčič, 1958a: 40; 1958b: 278; Shiba, 1969: 101; 2006: 87; Hardman et al., 1885; Smith-Meyer & Ueckermann, 1987: 2 (redescription), Gupta, 2002: 12. Detailed synonymy in Vitzthum, 1926
Anystes baccarum (L.): Irk, 1939; Schweizer, 1951: 66
Actineda coccinea Targioni-Tozzetti, 1878: 258.
Type locality: Sweden
Distribution: Algeria, Australia, Austria, Canada, Chile (Juan Fernandes Is.), China, Egypt, Faroes Is., Germany, Great Britain, Hungary, Iceland, India, Iran, Israel, Italy, Japan, Korea, Madeira, Malaysia, Mexico, New Guinea, Norway, St-Helena, South Africa, Spain, Sweden, Switzerland, Tanzania, Tunisia, Turkey, U.S.A.
Anystis berlesei : Oudemans, 1936: 395
Type locality: Italy
Distribution: Italy, St. Helena
Anystis borrusica : Oudemans, 1936: 405
Type locality: Königsberg, now Kaliningrad, Russia
Distribution: Russia (Kaliningrad District)
Anystis flaveola Stoll, 1887 (nec flaveola C.L. Koch, 1836)
Anystis citreola : Oudemans, 1936: 410
Type locality: Guatemala
Distribution: Guatemala
Trombidium cornigerum Hermann, 1804: 47
Actineda cornigera (Hermann): C.L. Koch, 1836: Heft 17. Considered by Vitzthum (1926) who listed some names of C.L. Koch as synonyms of A. baccarum: Actineda flaveola, A. pini, A. triangularis, A. robuscula, A. rubis, and others. In his revision of Anystidae Oudemans (1936) indicates that “Kochschen “Arten” wahrscheinlich Nymphen sind”(p.365, 368). as synonym of Anystis baccarum.
Erythraeus cornigerus: Dugès, 1834: 16, 44.
Anystis cornigerum (Hermann): von Heyden, 1826. Type of genus Anystis von Heyden, 1826.
Type locality: France
Distribution: France, Great Britain, Italy
Trombidium cornutum Hermann, 1804: 47
Anystis sp.: Thor & Willmann, 1947: 513
Anystis cornutum (Hermann): Otto, 1999a: 895 (with “?”)
Type locality: France
Distribution: France
Trombidium cursorium Gervais, 1844: 187
Anystis cursorium (Gervais):
Type locality: France
Distribution: France
Anystis germanica : Oudemans, 1936: 403
Type locality: Königsberg, now Kaliningrad, Russia
Distribution: Russia (Kaliningrad District)
Anystis indica Gupta, 1992: 144; 2002:
Type locality: India
Distribution: India
Actineda jabanica Berlese, 1905: 156
Anystis jabanica (Berlese): Holm & Wallace, 1989: 77
Type locality: Buitenzorg (Java)
Distribution: Australia, Indonesia (Java)
Anystis lahorense Khan, 1967: 400
Type locality: Lahor
Distribution: Pakistan
Anystis langei Barilo, 1984c: 452
Type locality: Zeravshan Range, 20 km south of Samarkand
Distribution: Uzbekistan
Anystis nagalandensis Gupta, 1991: 235; 1992:
Type locality: Botsa, Nagaland
Distribution: India
Actineda pallescens C.L. Koch, 1838: 6
Type locality: Germany
Distribution: Germany, Holland, Italy, Sainte-Helene
Acarus salicinus : Linnaeus, 1758: 618
Anystis salicinus (L.): Oudemans, 1936: 393; Smith – Meyer & Ueckermann, 1987: 8; Otto, 1992: 33; Shiba, 2006: 87; Zhang & Hong, 2010: 8
Anystis rosae Oudemans, 1936: 397- Type locality: Arnhem, Holland
Anystis kochi Oudemans, 1936: 399 - Type locality: Prov. Gelderland, Holland
Type locality: (?) Sweden
Distribution: China, France, Germany, Great Britain, Holland, Japan, Sweden; Sainte-Helene, South Africa (introduced)
Anystis sellnicki : Oudemans, 1936: 401
Type locality: Königsberg, now Kaliningrad, Russia
Distribution: Russia (Kaliningrad District)
Actineda venustula Koch & Berendt, 1854: Plate XIII, fig. 106
Anystis venustula (Koch et Berendt): Oudemans, 1936: 411
Type locality: Baltic Amber (Paleogene)
Distribution: Baltic Amber (Paleogene)
Acarus vitis Schrank, 1781: 519
Actineda vitis (Schrank): Halbert, 1915: 114
Anystis vitis (Schrank): Oudemans, 1936: 464; Bottazzi, 1950: 29; Starkoff & Starkoff, 1950: 95
Type locality: Austria
Distribution: Austria, Ireland, Italy
Anystis voigtsi : Oudemans, 1936: 405
Type locality: Bremen, Germany
Distribution: Germany
Anystis salicinus : Meyer & Ueckermann, 1987: 8, in part; Holm & Wallace, 1989: 79 (nec Acarus salicinus Linnaeus)
Anystis wallacei Otto, 1992: 25; Halliday, 2005: 7; Barbar, 2018
Type locality: France
Distribution: China, France, Portugal, Spain, Morocco, Syria; South Africa, Australia (introduced)
Autenriethia Oudemans, 1936: 425
Type species: Actineda velox Berlese, 1905
Actineda velox : Berlese, 1905: 156
Autenriethia velox (Berlese): Oudemans, 1936: 427
Type locality: Johore, Malaysia
Distribution: Malaysia
Barellea Oudemans, 1936: 427
Type species: Anystis sinensis Berlese, 1923
Anystis sinensis : Berlese, 1923: 237
Barellea sinensis (Berlese): Oudemans, 1936: 427; Zhang & Hong, 2010: 8
Type locality: Peking
Distribution: China
Mesoanystis Zacharda, 1985: 150
Type species: Mesoanystis taymirensis Zacharda, 1985
Mesoanystis taymirensis Zacharda, 1985: 150
Type locality: Taymir Peninsula, Jantardach (Upper Cretaceous amber)
Distribution: Russia (Taymir)
Scharfenbergia Oudemans, 1936: 412
Type species: Actineda hilaris C.L.Koch, 1836
Actineda hilaris : C.L. Koch, 1836: 6
Scharfenberhia hilaris (C.L.Koch): Oudemans, 1936: 412
Type locality: Esslarn, Germany
Distribution: Germany, France
Scharfenbergia gauthieri Oudemans, 1936: 414
Type locality: Reghaia, Algeria
Distribution: Algeria
Snartia Oudemans, 1936: 417
Type species: Snartia nepenthus Oudemans, 1936
Snartia nepenthus Oudemans, 1936: 417
Type locality: Haranggaol, Lake Toba, Sumatra
Distribution: Sumatra (Indonesia)
Tencateia Oudemans, 1936: 422
Type species: Tencateia besselingi Oudemans, 1936
Tencateia besselingi : Oudemans, 1936: 424; Willmann, 1954: 236; Meyer & Ueckermann, 1987: 14
Type locality: Roosendaal, Noord-Brabant
Distribution: Czech Rep., Holland
Tencateia kanthiensis Gupta, 1992: 146
Type locality: W. Bengal, Distr. Medinpur, Kanthi
Distribution: India
Tencateia toxopei : Oudemans, 1936: 425; Willmann, 1951a: 147; Shiba, 2006: 87
Type locality: Oisterwijk, Noord-Brabant (Holland)
Distribution: Austria, Holland, Japan
Tencateia villosa : Meyer & Ueckermann, 1987: 10
Type locality: South Africa (10 km from Postmasburg, Cape Prov., and many other places)
Distribution: South Africa
Walzia Oudemans, 1936: 419
Type species: Actineda antiguensis Stoll, 1886
Actineda antiguensis : Stoll, 1886: 7
Walzia antiguensis (Stoll): Oudemans, 1936: 419
Type locality: Antigua, Guatemala
Distribution: Guatemala, Venezuela
Walzia australica : Womersley, 1942: 18; Holm & Wallace, 1989; Su et al., 2006
Type locality: South Australia, Queensland
Distribution: Australia (South Australia, Queensland), China
Walzia darjeelingensis Gupta, 1992: 148; 2002:
Type locality: West Bengal, Darjeeling Botanical Garden
Distribution: India
Walzia hammeni : Meyer & Ueckermann, 1987: 16
Type locality: South Africa, Blyde River Canyon, Transvaal, and many other places
Distribution: South Africa
Walzia indiana : Meyer & Ueckermann, 1987: 14; Gupta, 2002:
Type locality: Midnapur, West Bengal, India
Distribution: India
Subfamilia Erythracarinae Oudemans
Erythracarinae Oudemans, 1936: 427
Erythracaridae: Pepato & Klimov, 2015
Type genus: Erythracarus Oudemans, 1936
Schellenbergia Oudemans, 1936: 433 (praeocc.)
Chaussieria Oudemans, 1937: 662 (replacement name); Meyer & Ueckermann, 1987: 18; Otto, 1999b: 254
Chausseria (misspelling): Willmann, 1951b: 147; Hutterer & Rack, 1977: 421
Type species: Erythraeus domesticus C.L. Koch, 1847
Chaussieria aprica Otto, 1999b: 258
Type locality: 10 miles south of Larrimah, Northern Territory
Distribution: Australia
Erythraeus parietinus (Hermann): Berlese, 1882: (not Hermann, 1804)
Tarsotomus berlesei Oudemans, 1906: 87
Schellenbergia berlesei (Oudemans): Oudemans, 1936: 434
Chausseria berlesei (Oudemans): Willmann, 1951a: 147
Chaussieria berlesei (Oudemans): Meyer & Ueckermann, 1987: 18
Type locality: Italy
Distribution: Austria, Italy
Chaussieria capensis Meyer & Ryke, 1960: 185; Ueckermann & Smith Meyer, 1988: 41 (synonymized it with Ch. venustissima, but Otto, 1999b: 264 restored it as full species); López-Campos & Vázquez-Rojas, 2010: 157
Type locality: South Africa, Bathurst
Distribution: South Africa
Chaussieria dissimilis : Naudo, 1977: 298; Otto, 1999b: 265
Type locality: St. Helena, Prosperous Bay Plain
Distribution: St. Helena
Erythraeus domesticus : C.L.Koch, 1847: 257
Tarsotomus domesticus : Oudemans, 1921: 358
Schellenbergia domesticus (C.L.Koch): Oudemans, 1936: 433
Schellenbergia berlesei (Oudemans): Oudemans, 1936: 434
Chaussieria domestica (C.L.Koch): Meyer & Ueckermann, 1987: 22; Otto, 1999b: 266; Shiba, 2006: 87
Erythraeus venustissimus : Berlese, 1882: fasc. 2(3); 1888: 3; Canestrini, 1885: 1652; 1886: 164; Schweizer, 1922: 81
Erythracarus venustissimus (Berlese): Type: Sicily. Vitzthum, 1928: 47; Schweizer, 1951: 66; Mihelčič, 1958a: 40; Schweizer & Bader, 1963: 266
Schellenbergia venustissima (Berlese): Oudemans, 1936: 436
Chaussieria venustissima (Berlese): Naudo, 1977: 300; Meyer & Ueckermann, 1987: 20 (in part, European material only); Ueckermann & Smith Meyer, 1988: 41; Otto & Olomski, 1994: 287
Chausseria venustissima (Berlese)(misspelling): Hutterer & Rack, 1976: 434
Tarsotomus berlesei Oudemans, 1905: 87 (syn. by Otto, 1999b: 266)
Chaussieria berlesei (Oudemans): Meyer & Ueckermann, 1987: 18
Chaussieria maritima : Evans & Browning, 1953: 419. Type: Babbacombe, South Devon, England. Meyer & Ueckermann, 1987: 22. Syn. by Otto, 1999b: 266
Type locality: Germany
Distribution: Austria, Germany, Great Britain, Greece (Crete), Holland, Iran, Italy (incl. Sicily), Japan, Russia, Spain, Switzerland, Uzbekistan
Chaussieria pilifera Otto, 1999b: 270
Type locality: South Australia, 31 km NW of Renmark
Distribution: Australia (South Australia, Victoria, Western Australia)
Chaussieria pulleni Otto, 1999b: 274
Type locality: 79 km NNW of Renmark, South Australia
Distribution: Australia
Chaussieria pumila Otto, 1999b: 275
Type locality: NSW, 43 km E of Balranald
Distribution: Australia (NSW, Northern Territory, South Australia, Victoria, Western Australia)
Chaussieria sanctaehelenae : Naudo, 1977: 292; Meyer & Ueckermann, 1987: 22; Otto, 1999b: 282
Chaussieria brevis Naudo, 1977: 298; Meyer & Ueckermann, 1987: 22. Syn. by Otto, 1999b: 283
Chaussieria benoiti Naudo, 1977: 297; Meyer & Ueckermann, 1987: 22.
Type locality: Prosperous Bay Plain
Distribution: St. Helena
Tarsotomus warregense : Hirst, 1931: 562; Oudemans, 1936: 442
Schellenbergia warregense (Hirst): Oudemans, 1936: 442; Womersley, 1942: 20
Chaussieria warregense (Hirst): Naudo, 1977: 300; Meyer & Ueckermann, 1987: 18; Holm & Wallace, 1989; Otto, 1999b: 286
Type locality: Barrigun, near Warrego River, New South Wales
Distribution: Australia (New South Wales, ACT, Northern Territory, South Australia, Western Australia, Tasmania, Queensland)
Erythracarus Berlese, 1903: 26
Type species: Trombidium ruricola (not parietinum Hermann, 1804, fide Otto, 1999a)
Palaeoerythracarus Zacharda, 1985: 150 – syn. by Otto, 2000: 457
Type species: Palaeoerythracarus sachalinensis Zacharda, 1985
Bechsteinia Oudemans, 1936: 436 – syn. by Otto, 1999a: 828
Chabrieria Oudemans, 1936: 446 – syn. by Otto, 1999a: 828
Type species of Bechsteinia Oudemans: Bechsteinia schneideri Oudemans, 1936
Type species of Chabrieria Oudemans: Tarsotomus terminalis Banks, 1916
Erythracarus amnicolus Otto, 1999a: 860
Type locality: 67 km S of Menindee, N.S.W.
Distribution: Australia (N.S.W., Western Australia)
Erythracarus avius Otto, 1999a: 864
Type locality: Tasmania, Peters Link Road
Distribution: Australia (Tasmania)
Erythracarus barbarus Otto, 1999a: 865
Type locality: McNary (Arizona)
Distribution: USA (Arizona)
Erythracarus ciliatus Otto, 1999a: 868
Type locality: 5 km W of Cowangie, Victoria
Distribution: Australia (Vtoria, Western Australia)
Erythracarus decoris Otto, 1999a: 831
Type locality: Bethungra, N.S.W.
Distribution: Australia (N.S.W., A.C.T., Victoria, Northern Territory, Western Australia)
Erythracarus elegans Otto, 1999a: 835
Type locality: 13 km NW of Goulbourn, N.S.W., road to Crookwell
Distribution: Australia (N.S.W., A.C.T., Victoria)
Erythraeus epigeus C.L. Koch, 1837: 16.24; 1842: 53
Erythracarus ? epigeus (C.L. Koch): Oudemans, 1936: 431
Erythracarus epigeus (C.L. Koch): Otto, 1999a: 876
Type locality: along the river Danube near Regensburg
Distribution: Germany
Bechsteinia festina : Meyer & Ueckermann, 1987: 28
Erythracarus festinus (Meyer et Ueckermann): Otto, 1999a: 840
Type locality: Transvaal, near Golela, Pongola River
Distribution: South Africa
Erythracarus flavipes Otto, 1999a: 878
Type locality: A.C.T., Canberra, Mount Ainslie
Distribution: Australia
Bechsteinia grahami : Meyer & Ryke, 1960: 184; Meyer & Ueckermann, 1987: 28
Erythracarus grahami (Meyer et Ryke): Otto, 1999a: 883
Type locality: Bathurst, near Albany, Cape Province, South Africa
Distribution: Angola, South Africa
Erythraeus hirsutus Menge, 1854: 107
Erythracarus ? hirsutus (Menge): Oudemans, 1936: 432; Otto, 1999a: 884
Type locality: amber
Distribution: amber
Erythraeus lagopus Menge, 1854: 107
Erythracarus ? lagopus (Menge): Oudemans, 1936: 432; Otto, 1999a: 885
Type locality: amber
Distribution: amber
Erythracarus katherinae Otto, 1999a: 841
Type locality: Burrewarra Point, N.S.W.
Distribution: Australia, incl. Tasmania
Bechsteinia longisetae (Reda), 1991: 728
Erythracarus longisetae (Reda): Otto, 1999a:
Type locality: Egypt
Distribution: Egypt
Bechsteinia festina : Meyer & Ueckermann, 1987: 28 (in part, misidentification)
Erythracarus nasutus Otto, 1999a: 844; Lopez-Campos & Vazquez-Rojas, 2010: 157; de Oliveira Bernardi et al., 2010: 119
Type locality: Transvaal, Loskop Dam
Distribution: Brazil, Mexico, South Africa
Bechsteinia nobarii Reda, 1991: 729
Type locality: Egypt
Distribution: Egypt
Trombidium parietinum : Hermann, 1804: 37
Erythraeus parietinus Latr.: Haller, 1882: 314
Erythraeus parietinus (Hermann): C.L. Koch, 1837: 16.23; 1842: 53 + figure 25
Tarsotomus parietinus (Herm.): Oudemans, 1906: 85
Erythracarus parietinus (Herm.): Thor, 1912 : 388; Schweizer, 1951: 66; Franz, 1954: 367; Bottazzi, 1950: 29; Mihelčič, 1958a: 40; 1958b: 278
Erythraeus parietinus Herm.: Hull, 1918: 29
Erythracarus parietinum (Hermann): Oudemans, 1936: 429
Erythraeus comes Berlese, 1886: 6; syn. by Oudemans, 1906: 87 (as Tarsotomus comes)
Tarsotomus comes (Berlese): André, 1927: 169
Erythracarus comes (Berlese): Berlese, 1903: 26; 1920: 8; Starkoff & Starkoff, 1950: 96
Anystis parietinum : Oudemans, 1936:
Bechsteinia solarii Ueckermann, 1987: 56. Type: Hamburg, Germany, Syn. by Otto, 1999a: 886
Erythraeus spinatus Banks, 1894: 210. Type: New York State, Long Island, Sea Cliff Village. Oregon. Syn. by Otto, 1999a: 886
Type locality of E. parietinus: France
Distribution: Algeria, Austria, France, Germany, Great Britain, Holland, Italy, Norway, Spain, Switzerland, Ukraine, USA
Erythraeus proavus Menge, 1854: 107
Type locality: amber
Distribution: amber
Trombidium pyrrholeucum Hermann, 1804: 58
Erythraeus ruricola Dugès, 1834: 40; C.L. Koch, 1837: 1.5, 1842: 53; Canestrini, 1885: 1650; Berlese, 1887: No2,3. Synonymy by Oudemans, 1936: 432
Erythracarus ruricola (Dugès): Berlese, 1903: 26; Oudemans, 1936: 432; Schweizer, 1951: 66; Mihelčič, 1958a: 40; Schweizer & Bader, 1963: 266 (in part, specimen from the locality ‘Scanf’ only, fide Otto, 1999a: 846)
Erythracarus pyrrholeucus : Oudemans, 1936: 432; Otto, 1999a: 850 (redescription); Barbar, 2018
Tarsotomus terminalis : Banks, 1916: 13. Type: Claremont, California. Syn. by Otto, 1999a: 847
Chabrieria terminalis (Banks): Oudemans, 1936: 446
Bechsteinia schneideri : Type: San Remo, Italy; Oudemans, 1936: 436; Cooreman, 1946: 4000 (Italy, Cyprus); Ueckermann, 1987: 57. Syn. by Otto, 1999a: 847
Bechsteinia californica McGregor, 1956: 23; Ueckermann, 1987: 56. Type: Whittier, California. Syn. by Otto, 1999a: 847
Type locality of Erythracarus pyrrholeucus: France
Distribution: Algeria, Australia, Cyprus, Iran, Italy, France, Spain, Ukraine, Crimea, U.S.A. (California)
Erythraeus raripilus Menge. 1854: 107
Erythracus [sic] raripilus (Menge): Oudemans, 1936: 433
Erythracarus raripilus (Menge): Otto, 1999a: 894
Type locality: amber
Distribution: amber
Palaeoerythracarus sachalinensis Zacharda, in Zacharda & Krivoluckiy, 1985: 150
Erythracarus sachalinensis (Zacharda)
Type locality: South Sachalin I., Starodubskoe (Paleogene amber)
Distribution: Russia (Sachalin)
Erythraeus tricolor Lucas,1846: 311
Erythracarus tricolor (Lucas): Oudemans, 1936: 433; Otto, 1999b: 895 (with “?”)
Type locality: Algeria
Distribution: Algeria
Erythrocheylus Berlese, 1903: 27
Type species: Pseudocheylus erythraeoides Leonardi, 1901
Pseudocheylus erythraeoides Leonardi in Berlese & Leonardi, 1901: 110
Erythrocheylus erythraeoides (Leonardi, 1901): Berlese, 1903:
Type locality: Funchal
Distribution: Chile
Erythrocheylus littoralis Berlese, 1903: 27
Type locality: Italy
Distribution: Italy
Lacteoscythis Pogrebnyak, 1995: 11
Type species: Lacteoscythis arenaria Pogrebnyak, 1995
Lacteoscythis arenaria Pogrebnyak, 1995: 11
Type locality: Black Sea Natural Reserve (south part of Ukraine)
Distribution: Ukraine
Namadia Meyer & Ueckermann, 1987: 32
Type species: Namadia floreata Meyer et Ueckermann, 1987
Namadia floreata Meyer & Ueckermann, 1987: 32
Type locality: Gamoep
Distribution: South Africa
Paratarsotomus Kuznetsov, 1983: 91
Type species: Paratarsotomus scutellatus Kuznetsov, 1983
Tarsotomus behningii Thor, 1912a: 471 (the species is named after A. Behning); Oudemans, 1936: 441
Paratarsotomus behningii (Thor): Otto, 1999c: 797
Type locality: “in einem Nebenteich des Salzsees Baskuntschak”, Russia
Distribution: Russia
Tarsotomus callunae Oudemans, 1936: 438; Meyer & Ueckermann, 1987: 26
Paratarsotomus callunae (Oudemans): Otto, 1999c: 797
Type locality: Bergen-op-Zoom, Noord - Brabant
Distribution: Holland
Tarsotomus macropalpis : Banks, 1916: 13
Paratarsotomus macropalpis (Banks): Otto, 1999c: 798; Abdelgayed et al., 2019: 31
Type locality: Claremont, California
Distribution: California (USA), Egypt
Erythraeus sabulosus Berlese, 1885: No 7; 1886: fa. 29; Schweizer, 1922: 81
Tarsotomus sabulosus (Berlese): Oudemans, 1936: 411
Erythracarus sabulosus (Berl.): Berlese, 1903: 26; Mihelčic, 1958b: 278
Erythracarus sabulosus var. oelandicus Trägårdh, 1915: 126. Oeland Insel. Syn. by Otto, 1999c: 800
Tarsotomus rackae : Meyer & Ueckermann, 1987: 26 -Type locality: Oberweiden, Austria. Syn. by Otto, 1999c: 800
Tarsotomus oelandicus (Trägårdh): Oudemans, 1936: 442. Syn. by Otto, 1999c: 800
Paratarsotomus sabulosus (Berlese): Otto, 1999c: 800; Ueckermann et al., 2006: 255
Type locality: Italy
Distribution: Austria, Germany, Italy, Spain, Sweden, Ukraine, Yemen
Paratarsotomus scutellatus Kuznetsov, 1983: 91; Otto, 1999c: 802; Lopez-Campos & Vazquez-Rojas, 2010: 157
Type locality: Uzbekistan
Distribution: Mexico, Uzbekistan
Pedidromus Otto, 2000: 458
Type species: Pedidromus peliculus Otto, 2000
Pedidromus agitatus Otto, 2000: 459
Type locality: N.S. Wales, 80 km East of Hay
Distribution: Australia (NSW, Northern Territory, Western Australia)
Pedidromus curiosus Otto, 2000: 460
Type locality: 12 km SW of Calperum, South Australia
Distribution: Australia
Pedidromus durongensis Otto, 2000: 461
Type locality: 8 km NW of Durong, Queensland
Distribution: Australia
Pedidromus peliculus Otto, 2000: 462
Type locality: N.S. Wales, 43 km East of Balranald
Distribution: Australia (NSW, South Australia)
Pedidromus pilotrix Otto, 2000: 462
Type locality: 10 km SSW of Warren, NSW
Distribution: Australia
Pedidromus velox Otto, 2000: 464
Type locality: Taplan, South Australia
Distribution: Australia (South Australia, Western Australia, Queensland, Victoria)
Siblyia Oudemans, 1936: 445
Type species: Erythraeus ignipes Dugès, 1834
Erythraeus ignipes : Dugès, 1834: 43
Siblyia ignipes (Dugès): Oudemans, 1936: 445
Type locality: ? Montpellier, France
Distribution: France
Tarsolarkus Thor, 1912a: 466
Tarsolarcus Thor, 1912a: 465
Neotarsolarcus Kuznetsov, 1983: 87. Syn. by Oto, 2000: 470. Type: Neotarsolarcus clypeatus Kuznetsov, 1983, Tadjikistan and Russia.
Type species: Tarsolarkus articulosus Thor, 1912
Tarsolarkus articulosus : Thor, 1912a: 466; Irk, 1939: 154; Sellnick, 1949: 128; Willmann, 1951b: 147; Hull, 1918: 30; Irk, 1939: 154; Oudemans, 1936: 444; Sellnick, 1949: 128; Franz, 1954: 367; Schmölzer, 1956: 548; Mehl, 1979: 36; Otto, 2000: 470
Tarsolarcus articulosus : Willmann, 1951b: 147
Erythracarus cluozzai Schweizer & Bader, 1963: 265. Type: Switzerland, Val Cluozza, 2100 m; syn. by Otto, 1999a: 896
Tarsolarkus oudemansi Barilo, 1984a: 1006 from Uzbekistan. Syn. by Otto, 2000: 470
Type locality: Norway (many places)
Distribution: Austria, France, Great Britain, Norway, Sweden, Switzerland, Uzbekistan
Neotarsolarcus clypeatus Kuznetsov, 1983: 88
Tarsolarkus clypeatus (Kuznetsov): Otto, 2000: 470
Type locality: Russia
Distribution: Russia, Tajikistan
Tarsolarkus longisetus Barilo, 1984a: 1004; Otto, 2000: 472
Type locality: 8 km East of Samarkand
Distribution: Uzbekistan
Tarsotomus Berlese, 1882: 2
Type species: Erythraeus hercules Berlese, 1882
Anandia Hirst, 1927: 445 – syn. by Otto, 1999c: 751
Type species: Anandia alticola Hirst, 1927
Absoloniana Willmann, 1940: 215 – syn. by Otto, 1999c: 751
Type species: Absoloniana diversipes Willmann, 1940
Paranandia Meyer & Ueckermann, 1987: 34 – syn. by Otto, 1999c: 751
Type species: Anandia bathurstensis Meyer et Ryke, 1960
Parabsolonia Barilo, 1984: 1736 – syn. by Otto, 1999c: 751
Type species: Parabsolonia ignicola Barilo, 1984
Tarsotomus abruptus Otto, 1999c: 756
Type locality: 10 km SSW of Warren, NSW
Distribution: Australia (NSW, ACT, Victoria)
Tarsotomus aiolos Otto, 1999c: 758
Type locality: 64 km SE of Cooya Pooya Homestead, Western-Australia
Distribution: Australia (WA)
Tarsotomus aleantis Otto, 1999c: 759
Type locality: Millstream Chichester National Park; Western-Australia
Distribution: Australia (WA)
Anandia alticola : Hirst, 1927: 445; Oudemans, 1936: 445; Meyer & Ryke, 1960: 180; Meyer & Ueckermann, 1987: 22
Tarsotomus alticolus (Hirst): Otto, 1999c: 785
Type locality: “Upper Rombok, Mount Everest, 4950 m”, Tibet
Distribution: Tibet (China)
Tarsotomus ambitus Otto, 1999c: 762
Type locality: Millstream Chichester National Park; Western-Australia
Distribution: Australia (WA)
Absoloniana amentata : Barilo, 1984a: 1009; Meyer & Ueckermann, 1987: 24
Tarsotomus amentatus (Barilo): Otto, 1999c: 786; Ueckermann et al., 2006: 255
Type locality: Dzhambayski Region, Samarkandskaya oblast (Uzbekistan)
Distribution: Uzbekistan, Yemen
Tarsotomus anubis Otto, 1999c: 763
Type locality: 8 km W of Underbool, road to Murrayville; Victoria
Distribution: Australia (Victoria, Northern Territory, South Australia)
Tarsotomus argillus Otto, 1999c: 767
Type locality: 3 km east of Swan Hill; Victoria
Distribution: Australia (Victoria, NSW, South Australia)
Anandia bathurstensis : Meyer & Ryke, 1960: 182
Paranandia bathurstensis (Meyer et Ryke): Meyer & Ueckermann, 1987: 35
Tarsotomus bathurstensis (Meyer et Ryke): Otto, 1999c: 751
Type locality: Albany District, Bathurst, Cape Province
Distribution: South Africa
Absoloniana bestiola Barilo, 1984b: 1734; Meyer & Ueckerman, 1987: 24
Tarsotomus bestiolus (Barilo): Otto, 1999c: 786
Type locality: Uzbekistan
Distribution: Uzbekistan
Tarsotomus colossus Otto, 1999c: 769
Type locality: Mount Newman, Balfour Downs; Western-Australia
Distribution: Australia (WA, Western Australia)
Tarsotomus comosus Otto, 1999c: 771
Type locality: 10 km SSW of Warren, NSW
Distribution: Australia (NSW)
Tarsotomus desertorus Otto, 1999c: 772
Type locality: 32 km south of Murrayville, road to Nhill; Victoria
Distribution: Australia (Victoria, NSW, Northern Territory, South Australia)
Tarsotomus dianellus Otto, 1999c: 775
Type locality: south of Walpeup, road to Patchewollock; Victoria
Distribution: Australia (Victoria, NSW, South Australia)
Tarsotomus dioxis Otto, 1999c: 777
Type locality: 80 km east of Hay (NSW)
Distribution: Australia (NSW)
Tarsotomus erraticus : Banks, 1912: 441; Oudemans, 1936: 441; Otto, 1999c: 787
Type locality: Springer, New Mexico
Distribution: USA (New Mexico)
Tarsotomus heliophilus Otto, 1999c: 778
Type locality: 14 km north of Balranald, road to Ivanhoe; New-South-Wales
Distribution: Australia (NSW, Victoria, South Australia, Western Australia)
Erythraeus hercules : Berlese, 1882: fa. 2; 1888: 3
Tarsotomus hercules (Berlese): Trägårdh, 1905: 64; Oudemans, 1936: 441; Starkoff & Starkoff, 1950: 96; Cooreman, 1955: 123; Bottazzi, 1950: 30; Mihelčič, 1958a: 40; Ueckermann et al., 2006: 255
Absoloniana diversipes : Willmann, 1940: 215; 1941: 56; Barilo, 1984a: 1007; Meyer & Ueckermann, 1987: 24 -Type locality: Cave Čančarica, Bjelashnica planina, South Bosna. Syn. by Otto, 1999c: 787
Type locality of Erythraeus hercules: Sicily, Acireale
Distribution: Bosnia, Egypt, France, Iran, Italy, Salamin Is. (Greece), Sicily, Spain, Sudan, Switzerland, Ukraine, Uzbekistan, Yemen
Parabsolonia ignicola Barilo, 1984: 1738
Tarsotomus ignicolus (Barilo): Otto, 1999c: 789
Type locality: Uzbekistan
Distribution: Ukraine, Uzbekistan
Erythraeus hercules var. nivalis Schweizer, 1922: 80
Tarsotomus nivalis : Oudemans, 1936: 441; Bottazzi, 1951: 222
Type locality: Alps (2760 - 3000 m)
Distribution: Italy, Switzerland (Cima del Pezzacul, 2300 m)
Absoloniana pentagonea : Meyer & Ueckermann, 1987: 24
Tarsotomus pentagoneus Otto, 1999c: 789
Type locality: Malelane and Pongola, Transvaal, South Africa
Distribution: South Africa
Tarsotomus primitivus Otto, 1999c: 790
Type locality: 32 km south of Murrayville, road to Nhill; Victoria
Distribution: Australia (Victoria)
Tarsotomus pusillus Otto, 1999c: 779
Type locality: 64 km SE of Cooya Pooya Homestead, Western-Australia
Distribution: Australia (WA)
Tarsotomus pygmaeus Otto, 1999c: 781
Type locality: Kurunjini (Hamersley Range) National Park; Western Australia
Distribution: Australia (WA)
Tarsotomus solatus Otto, 1999c: 781
Type locality: Uluru National Park, Mount Olga, Northern Territory
Distribution: Australia (NT)
Diversipes undulatus Mahunka & Mahunka-Papp, 1993: 326
Tarsotomus undulatus (Mahunka et Mahunka-Papp):
Type locality: Madagascar
Distribution: Madagascar
Tarsotomus velopes Otto, 1999d: 783
Type locality: north of Roebourne; Western-Australia
Distribution: Australia (WA)
EUROPE
Austria (Irk, 1939; Willmann, 1951a, 1951b; Hutterer & Rack, 1976; Meyer & Ueckermann, 1987; Otto, 1999b) – Anystis baccarum, Chaussieria berlesei, Ch. domestica (= venustissima), Tarsolarkus articulosus, Paratarsotomus sabulosus (= Tarsotomus rackae), Tencateia toxopei
Belgium (Cooreman, 1943) – Anystis baccarum
Bosna and Herzegovina (Willmann, 1940, 1941) – Tarsotomus hercules (= Absoloniana diversipes)
Bulgaria (Beron, in prep.)
Czech Rep. (Willmann, 1954) – Anystis baccarum, Tencateia besselingi
Denmark (Hallas, 1978) -
Feroes Is. (Trägårdh, 1931) – Anystis baccarum
France (contin.)(Hermann, 1804; Dugès, 1834; Gervais, 1844; Oudemans, 1936; Schmölzer, 1956; Otto, 1992, 1999b) – Anystis andrei, A. cornigerum, A. cornutum, A. cursorium, A. salicinus, A. wallacei, Erythracarus parietinus, Scharfenbergia hilaris, Siblyia ignipes, Tarsolarkus articulosus, Tarsotomus hercules
Germany (C.L. Koch, 1836, 1842, 1847; Haller, 1882; Oudemans, 1936; Ueckermann, 1987; Otto, 1999b) - Anystis baccarum, A. salicinus, A. voigtsi, Chaussieria domestica (= venustissima), Erythracarus epigeus, E. parietinus (= Bechsteinia solarii), Paratarsotomus sabulosus, Scharfenbergia hilaris
Great Britain (Thor, 1912a; Hull, 1918; Evans & Browning, 1953; Otto, 1999b; Cuthbertson, Bell & Murchie, 2003; Cuthbertson & Murchie, 2004a, b, 2007, 2010) – Anystis baccarum, A. salicinus, A. cornigerum, A. cursorium, Chaussieria domestica (= maritima), Erythracarus parietinus, Tarsolarkus articulosus
Greece (Sellnick, 1931) - Anystis baccarum
Crete (Hutterer, 1978) - Chaussieria domestica (= venustissima)
Salamine (Cooreman, 1955) – Tarsotomus hercules
Holland (Oudemans, 1906, 1936) – Anystis salicinus, Chaussieria domestica, Erythracarus parietinus, Tencateia besselingi, Paratarsotomus callunae
Hungary (Dudich et al., 1940; Ripka et al., 2002) – Anystis baccarum
Iceland (Sellnick, 1940) – Anystis baccarum
Ireland (Halbert, 1915) – Anystis vitis
Italy (contin.) (Berlese, 1886; Oudemans, 1936; Starkoff & Starkoff, 1950; Bottazzi, 1950, 1951; Castagnoli & Nannelli, 1995; Otto, 1999b) – Anystis agilis, A. baccarum, A. conigerum, A.. pallescens, A. vitis, Erythracarus parietinus [= comes], E. pyrrholeucus (= E. [Bechsteinia] schneideri), Chaussieria domestica, Tarsotomus hercules, T. nivalis
Sicily (Berlese, 1882; Bottazzi, 1950; Otto, 1999b) – Chaussieria domestica, Erythracarus pyrrholeucus,Tarsotomus hercules
Latvia (Kuznetsov & Petrov, 1984; Petrova, Salmane & Čudare, 2004) - Anystis baccarum
Norway (Thor, 1912a, 1912b, 1912c) – Anystis baccarum, Erythracarus parietinus (= comes), Tarsolarkus articulosus
Portugal (cont.) (Halliday, 2005) – Anystis wallacei
Madeira (Willmann, 1939) – Anystis baccarum
Romania (Călugăr & Ivan, 2016) - Anystidae
Russia (all, continent.)(Thor, 1912; Oudemans, 1936; Lange, Drozdovskij & Bushkovskaya. 1974a, b; Kuznetsov, 1983; Meyer & Ueckermann, 1987) – Anystis borrusica, A. germanica, A. sellnicki (from Kaliningrad District), Chaussieria domestica (= venustissima), Paratarsotomus behningii, Tarsolarkus clypeatus
Crimea (Otto, 1999a) - Erythracarus pyrrholeucus
Sachalin (Zacharda, 1985; Otto, 2000) – Erythracarus [Palaeoerythracarus] sachalinensis
Serbia (Mladenović et al., 2013) – Anystis sp.
Spain (cont.) (Mihelčič, 1958a, 1958b; Otto, 1992) – Anystis baccarum, A. wallacei, Chaussieria domestica (= venustissima), Erythracarus parietinus, E. pyrrholeucus (= ruricola), Tarsotomus hercules
Sweden (Linnaeus, 1758; Trägårdh, 1915; Sellnick, 1949) – Anystis baccarum, A. salicinus, Paratarsotomus sabulosus [Öland, as “var. oelandicus Träg.], Tarsolarkus articulosus
Switzerland (Schweizer, 1922; Schweizer & Bader, 1963) – Anystis baccarum, Chaussieria domestica (= venustissima), Erythracarus parietinus, E. pyrrholeucus (= ruricola), Tarsolarkus articulosus (= Erythracarus cluozzai Schweizer et Bader), Tarsotomus hercules (= T. nivalis)
Turkey (Ülgentürk, Szentkirályi et al. 2013) – Anystis baccarum
Ukraine (Pogrebnyak, 1995; Otto, 1999a, 1999b) – Erythracarus parietinus, E. pyrrholeucus, Lacteoscythis arenaria, Paratarsotomus sabulosus, Tarsotomus hercules, T. ignicolus
AFRICA
Algeria (Lucas, 1946; Berlese, 1886; André, 1927; Oudemans, 1936) – Anystis baccarum, Scharfenbergia gauthieri, Erythracarus parietinus [Tarsotomus comes], E. tricolor
Angola – Erythracarus [Bechsteinia] grahami
Egypt (Trägårdh, 1905; Reda, 1991; Reda & Gomma, 1991; Abdelgayed, Abd El-Wahed, Ali & Eraky, 2019) – Anystis baccarum, Paratarsotomus macropalpis, Erythracarus nobarii, Tarsotomus hercules, Erythracarus nobarii, Neoaegyptocus chelicifurcata
Malawi (Meyer & Ueckermann, 1987) – Chaussieria domestica (= venustissima)
Morocco (Otto, 1992) – Anystis wallacei
Reunion (Quilici, Kreiter, Ueckermann & Vincenot, 1997) – 2 sp.
Sainte-Helene (Naudo, 1977; Meyer & Ueckermann, 1987; Otto, 1999b: 182) – Anystis baccarum, A. berlesei, A. kochi, Chaussieria dissimilis, Ch. sanctaehelenae (= brevis, = benoiti)
South Africa (Oudemans, 1936; Meyer, 1973; Meyer & Ueckermann, 1987; Ueckermann, 1987; Ueckermann & Meyer Smith, 1988; Otto, 1999a, 1999b; Halliday, 2005) – Anystis baccarum, A. wallacei (introduced), Erythracarus [Bechsteinia] grahami, Erythracarus festinus (= Bechsteinia festina), Chaussieria domestica (= venustissima), Ch. capensis, Namadia floreata, Tarsotomus bathurstensis, Tarsotomus [Absoloniana] pentagonea, Tencateia villosa, Walzia hammeri
Sudan (Trägårdh, 1905) – Tarsotomus hercules
Tanzania (Evans, 1953) – Anystis baccarum
Tunisia (Kreiter et al., 2002) - Anystis baccarum
ASIA
China (Berlese, 1923; Hirst, 1927; Su et al., 2006; Zeng et al., 2007; Zhang & Hong, 2010) – Anystis baccarum, A. salicinus, A. wallacei,Tarsotomus [Anandia] alticola (Tibet), Barellea sinensis, Walzia australica
Cyprus (Cooreman, 1946) – Erythracarus [Bechsteinia] pyrrholeucus (= schneideri)
India (Meyer & Ueckermann, 1987; Gupta, 1991, 1992, 2002) – Anystis baccarum, A. indica, A. nagalandensis, Tencateia kanthiensis, Walzia derjeelingensis, W. indiana
Indonesia (Berlese, 1905; Oudemans, 1936; Holm & Wallace, 1989;) – Anystis [Actineda] jabanica (Jawa), Snartia nepenthus (Sumatra)
Iran (Otto, 1999a, 1999b) – Erythracarus pyrrholeucus, Chaussieria domestica, Tarsotomus hercules
Israel and Palestina – Anystis baccarum
Japan (Shiba, 1969, 1976, 2006) - Anystis baccarum, A. salicinus, Tencateia toxopei, Chaussieria domestica, Erythracarus [Bechsteinia] sp.
Okinawa (Kishida, 1959) – Erythracarus simku (nomen nudum)
Korea (Lee Kim & Kim, 1995) - Anystis baccarum
Malaysia (Berlese, 1905; Vitzthum, 1926) – Anystis baccarum, Autenriethia velox
Syria (Barbar, 2018) – Anystis wallacei, Erythracarus pyrrholeucus
Tajikistan (Kuznetsov, 1983) - Tarsolarkus clypeatus
Uzbekistan (Barilo, 1984a, 1984b; Otto, 1999b) – Paratarsotomus scutellatus, Chaussieria domestica, Tarsotomus [Absoloniana] amentata, T. [A.] bestiola, T. hercules (= Absoloniana diversipes), T. [Parabsolonia] ignicola, Tarsolarkus longisetus
Yemen (Ueckermann et al., 2006) – Anystis baccarum, Erythracarus sp. near nasutus, E. sp. near parietinus, Paratarsotomus sabulosum Tarsolarkus sp., Tarsotomus amentatus, T. hercules
NORTH AMERICA
Canada (Hardman et al., 1885) - Anystis baccarum
Mexico (Vitzthum, 1933; López- Campos & Vázquez-Rojas, 2010) – Anystis baccarum, Chaussieria capensis, Erythracarus nasutus, Paratarsotomus sculellatus, Tarsotomus sp., new genera 1 and 2
USA (Banks, 1894, 1895, 1907, 1912, 1916; Ewing, 1909; Miller, 1925; McGregor, 1956; Sorensen, Kinn, Doutt & Cate, 1976; Otto, 1999a) – Anystis agilis, A. baccarum, Erythracarus barbarus, E. parietinus, Erythracarus pyrrholeucus [= Tarsotomus [Bechsteinia] terminalis, = Bechsteinia californica], Tarsotomus erraticus, Paratarsotomus macropalpis
SOUTH and CENTRAL AMERICA
Bermuda Isls (Banks, 1903) – Anystis agilis
Brazil (de Oliveira Bernardi et al., 2010) – Erythracarus nasutus
Chile (contin.)(Leonardi, in Berlese & Leonardi, 1901) - Anystidae
Juan Fernandez Is. (Trägårdh, 1931) – Anystis baccarum
Costa Rica (Banks, 1916) - Anystidae
Guatemala (Stoll, 1886, 1887) – Anystis citreola, Walzia antiguensis
Venezuela (Oudemans, 1936) – Walzia antiguensis
OCEANIA
Australia (continent)(Hirst, 1931; Womersley, 1933, 1942; Holm & Wallace, 1989; Otto & Halliday, 1991; Otto, 1992, 1999b,c; Halliday, 1998) – Anystis baccarum, A. jabanica, A. wallacei [intr.], Chaussieria pilicola, Ch. puleni, Ch. pumila, Ch. warregense, Erythracarus amnicolus, E. ciliatus, E. decoris, E. elegans, E. flavipes, E. katherinae, E. pyrrholeucus, Pedidromus agitatus, P. curiosus, P. durongensis, P. peliculus, P. pilotrix, P. velox, Tarsotomus anubis, T. argillus, T. colossus, T. comosus, T. desertorus, T. dianellus, T. dioxis, T. heliophilus, T. primitivus, T. pusillus, T. solatus, Walzia australica
Tasmania (Otto, 1999a,b) – Erythracarus avium, Chaussieria warregense
Papua New Guinea (Canestrini, 1898) – “Actineda vitis”
Teneriffiidae Thor, 1911: 179
Recorded first by Thor (1911) from Canary Is. and Paraguay, this small family contains now 26 species of predaceous free-living mites, living from the dunes of Namibia and Somalia and the intertidal zone of Japan to the high altitude in Alps (3207 m) and Himalaya (5050 m). Their generic taxonomy remains subject of discussions, especially after the revision of McDaniel, Morihara & Lewis (1976), in which only the two “initial” genera were recognized for the family Teneriffiidae: Teneriffia Thor, 1911 and Parateneriffia Thor, 1911. All genera, established between 1911 and 1976 by various authors: Neoteneriffiola Hirst, 1924, Heteroteneriffia Hirst, 1925, Austroteneriffia Womersley, 1935, Mesoteneriffia Irk, 1939, as well as the name Teneriffiola Strand, 1911 and the family name Teneriffiolidae, based on it, are considered by the McDaniel et al. (1976) as synonyms. Mesoteneriffiolla Schmölzer, 1956 and “Neoteneriffiola” tadjikistanica Wainstein, 1969 were not considered. Other acarologists added to the taxonomy (and sometimes to the confusion) of Teneriffiidae: Vitzthum (1942), Irk (1939a, b), Radford (1950), Schmölzer (1952, 1956), Tibbetts (1958), Besch (1962), Eller & Strandtmann (1963), Ehara (1965), Wainstein (1969), Shiba & Furukawa (1975), and others.
New genera and species described after 1976 by Schmölzer (2002 – Himalteneriffia from Ladakh), Luxton (1993), Yin, Bei & Li (1995 - Sinoteneriffia from China), Luo et al. (1996 – Sinoteneriffia from China), Luo, Yin et al. (2007, Neoteneriffiola from China), Bernardi et al. (2012, Brazil), Ueckermann & Khanjani (2002), Khanjani, Asali Fayaz & Ueckermann (2011), Khanjani, Yazdanpanah & Asali Fayaz (2013), Khanjani, Hoseini & Asali Fayaz (2014), Ueckermann & Durucan (2020, Teneriffia from Turkey), Zmudzinski, Skoracki & Friedrich (2021, Teneriffia from Ethiopia).
Judson (1994, 1995) not only described new and redescribed the older species of Teneriffiidae, but questioned the lumping of several genera by McDaniel et al. (1976) into two original units. Based on very detailed morphological studies, he reviewed and removed from synonymy the genera Neoteneriffiola and Austroteneriffia and described new species from Namibia and Australia.
In our collection in Sofia we have many specimens of Teneriffiidae from Greece, Afghanistan, Chinese Karakorum, Somalia, Nigeria, Tanzania, Zambia, Peru, and Mexico, which represent new species and are being described.
Fam. Teneriffiidae – 28 sp.
Austroteneriffia Womersley, 1935 - 10
Heteroteneriffia Hirst, 1925 - 2
Mesoteneriffia Irk, 1939 - 1
Mesoteneriffiola Schmölzer, 1956 - 1
Himalteneriffia Schmölzer, 2002 - 1
Neoteneriffiola Hirst, 1924 - 5
Parateneriffia Thor, 1911 - 1
Sinoteneriffia Yin, Bei et Li, 1995 – 2
Teneriffia Thor, 1911 - 5
Teneriffiidae Thor, 1911: 179
Teneriffiolidae Hirst, 1924: 1078
Teneriffiinae Womersley, 1935: 334
Type genus: Teneriffia Thor, 1911
Austroteneriffia Womersley, 1935: 334
Type species: Austroteneriffia hirsti Womersley, 1935
Austroteneriffia hirsti : Womersley, 1935: 335; Vitzthum, 1942: 808; Radford, 1950: 76; Judson, 1995: 830
Parateneriffia hirsti : McDaniel, Morihara & Lewis, 1976: 532
Type locality: Menindie (S. Australia)
Distribution: Australia
Neoteneriffiola hojoensis : Shiba & Furukawa, 1975: 118
Austroteneriffia hojoensis : Judson, 1995: 833
Type locality: Kashima, Hojo, Ehime Pref., near the sea shore, Japan
Distribution: Japan
Neoteneriffiola japonica : Ehara, 1965: 226; Shiba & Furukawa, 1975: 111
Austroteneriffia japonica (Ehara): Judson, 1995: 833
Type locality: Sapporo, Japan
Distribution: Japan
Austroteneriffia kamalii Ueckermann & Khanjani, 2002: 168
Type locality: Famenin, Hamadan prov., litter of Medicago sativa
Distribution: Iran
Austroteneriffia khorramabadiensis Khanjani, Hoseini & Asali Fayaz, 2014: 70
Type locality: Kaka Reza region, Khorramabad vic., Lorestan Province
Distribution: Iran
Austroteneriffia leei : Judson, 1995: 835
Type locality: Coomandook, S. Australia
Distribution: South Australia (Australia)
Austroteneriffia littorina : Shiba & Furukawa, 1975: 123; Judson, 1995: 837
Type locality: Wake, Matsuyama, Ehime Pref., Japan
Distribution: Japan
Austroteneriffia shiraziensis Khanjani, Yazdanpanah & Asali Fayaz, 2013: 36
Type locality: Koohmare Sorkhi region, Fars province
Distribution: Iran
Neoteneriffiola tadjikistanica Wainstein, 1969: 1250; 1978: 202
Austroteneriffia tadjikistanica (Wainstein): Judson, 1995: 838; Ueckermann, van Harten & Smith Meyer, 2006: 256
Type locality: Tadjikistan
Distribution: Tadjikistan, Yemen (incl. Socotra Island)
Austroteneriffia zamaniani Khanjani, Asali Fayaz & Ueckermann, 2011: 551
Type locality: Sorkh Abad Village, Hamedan vic., Hamedan Prov.
Distribution: Iran
Austroteneriffia Andréi: Besch, 1962: 236
“Austroteneriffia” andrei Besch: Judson, 1995: 827 (another genus!)
Type locality: “Arroyo Lopez, Gletscherbach bei Bariloche”, Prov. Rio Negro, Distribution: Argentina
Heteroteneriffia Hirst, 1925: 1278; synonymized by McDaniel et al., 1976: 529, but restaured by Ueckermann & Durucan, 2020
Type species: Heteroteneriffia marina Hirst, 1925
Heteroteneriffia marina : Hirst, 1925: 1278
Heteroteneriffiolla marina : Shiba, 1976: 134
Teneriffia marina (Hirst): McDaniel et al., 1976: 529
Type locality: Teluk Kemang, near Port Dickson, Negri Sembilan, Malaysia
Distribution: Japan, Malaysia
Heteroteneriffia mortoni Luxton, 1993: 104
Teneriffia mortoni (Luxton): Judson, 1994: 115
Type locality: Hong Kong marine littoral
Distribution: China (Hong Kong)
Himalteneriffia Schmölzer, 2002: 133
Type species: Himalteneriffia riccabonai Schmölzer, 2002
Himalteneriffia riccabonai Schmölzer, 2002: 133
Type locality: “Basislager am Markha – Fluss ca. 50 km nördlich von Leh, ca. 77o öL., 34o nBr., nur wenige km vom Kongmaru-La entfernt; ca. 5050 m Höhe”
Distribution: India (Ladakh)
Mesoteneriffia Irk, 1939: 220
Type species: Mesoteneriffia steinbocki Irk, 1939
Mesoteneriffia steinbocki : Irk, 1939: 222; Strandtmann, 1965: 261
Mesoteneriffia steinboecki : Schmölzer, 1952: 22; 2002: 124; Schweizer & Bader, 1963: 263
Type locality: “Ötztaler und Stubaier Alpen”, 2719 - 3050 m, Austria
Distribution: Austria, Switzerland
Mesoteneriffiola Schmölzer, 1956: 549
Type species: Mesoteneriffiola alpina Schmölzer, 1956
Mesoteneriffiola alpina Schmölzer, 1956: 549; 2002: 124
Type locality: “Unterhalb d. Roche d’Alvau, 2900 m”, Daufiné
Distribution: France
Neoteneriffiola Hirst, 1924: 1078
Parateneriffia Thor: McDaniel et al., 1976: 532 (in part)
Type species: Neoteneriffiola luxoriensis Hirst, 1924
Neoteneriffiola coineaui : Judson, 1994: 117
Type locality: Hamilton Range, Namibia
Distribution: Namibia
Neoteneriffiola luxoriensis : Hirst, 1924: 1078; Judson, 1994: 115
Parateneriffia luxoriensis :
Type locality: Assuan and Luxor, Egypt
Distribution: Egypt
Neoteneriffiola uta : Tibbetts, 1958: 44; López-Campos & Vázquez-Rojas, 2010: 157
Parateneriffia uta (Tibbetts): McDaniel et al., 1976: 532
Type locality: Utah
Distribution: Mexico, U.S.A. (Utah)
Neoteneriffiola xerophila De Oliveira Bernardi, Pellegrini & Ferreira, 2012: 411
Type locality: Brazil
Distribution: Brazil
Neoteneriffiola yunnanensis Luo Youzhen, Yin Suigong, Chen Bin & Zhang Yidong, 1997: 87
Type locality: Yunnan
Distribution: China (Yunnan)
Distribution: South Africa (Meyer et al., 1973)
Parateneriffia Thor, 1911: 176
Type species: Parateneriffia bipectinata Thor, 1911
Parateneriffia bipectinata Thor, 1911: 177
Type locality: Paraguay
Distribution: Paraguay
Sinoteneriffia Yin, Bei & Li, 1995: 433
Type species: Sinoteneriffia nuda Yin, Bei et Li, 1995
Sinoteneriffia kunmingensis Luo, Yin, Zhang & Bin, 1997: 243
Type locality: Kunming
Distribution: China
Sinoteneriffia nuda Yin, Bei & Li, 1995: 434 (Engl. 437)
Type locality: Liaoning Prov., from Sabina chinensis
Distribution: China
Teneriffia Thor, 1911: 172
Teneriffiola Strand, 1911: 287
Type species: T. quadripapillata Thor, 1911
Teneriffia aethiopica Zmudzinski, Skoracki & Friedrich, 2021: 318
Type locality:
Distribution: Ethiopia
Teneriffia mexicana : McDaniel et al., 1976: 529
Type locality: Sonora, Mexico
Distribution: Mexico
Teneriffia quadripapillata Thor, 1911: 173; Schmölzer, 2002: 131
Teneriffiola quadripapillata (Thor): Hirst, 1924: 1080
Type locality: Orotava, Teneriffa
Distribution: Teneriffa (Canary Is., Spain)
Teneriffia sebahatae Ueckermann & Durucan, 2020: 1141
Type locality: sea shore nr. Antalya (Kundu)
Distribution: Turkey
Teneriffia hajiqanbari Paktinat-Saeij & Kazemi, 2022: 263
Type locality: Queshm Island, southern Hormozgan Province
Distribution: Iran
EUROPE
Austria (Irk, 1939; Schmölzer, 1952) - Mesoteneriffia steinbocki
France (Schmölzer, 1956) - Mesoteneriffiola alpina
Greece (Beron, in prep.) - Teneriffiidae
Switzerland (Schweizer & Bader, 1963) - Mesoteneriffia steinbocki
Turkey (Ueckermann & Durucan, 2020) – Teneriffia sebahatae
AFRICA
Canary Is.(Thor, 1911) - Teneriffia quadripapillata
Egypt (Hirst, 1924) - Neoteneriffiola luxoriensis
Ethiopia (Zmudzinsk, Skoracki & Friedrich, 2021) – Teneriffia aethiopica
Namibia (Judson, 1994) - Neoteneriffiola coineaui
Nigeria (Beron, in prep.) - Teneriffiidae
Somalia (Beron, in prep.) - Teneriffiidae
South Africa (Meyer et al., 1973) - Neoteneriffiola sp.
Tanzania (Beron, in prep.) - Teneriffiidae
Zambia (Beron, in prep.) - Teneriffiidae
ASIA
Afghanistan (Beron, in prep.) - Teneriffiidae
China (Yin Suigong, Bei Naxir, Li Xuejin, 1995; Luo Youzhen, Yin Suigong, Zhang Yidong, Chen Bin, 1996; Luo Youzhen, Yin Suigong, Chen Bin, Zhang Yidong, 1997; Beron, in prep.) – Neoteneriffiola yunnanica, Sinoteneriffia kunmingensis, S. nuda
India (Schmölzer, 2002; Paktinat-Saeij & Kazemi, 2022 Zmudzinski et al., 2021) – Himalteneriffia riccabonai
Iran (Ueckermann & Khanjani, 2002; Khanjani, Asali Fayaz & Ueckermann, 2011; Khanjani, Yazdanpanah & Asali Fayaz, 2013; Khanjani, Hoseini & Asali Fayaz, 2014) – Austroteneriffia kamalii, A. khorramabadiensis, A. shiraziensis, A. zamaniensis
Japan (Ehara, 1965; Shiba, 1976; Shiba & Furukawa, 1975; Judson, 1995) – Austroteneriffia hojoensis, A. japonica, A. littorina, Teneriffia marina
Malaysia (Hirst, 1925) - Teneriffia marina
Tadjikistan (Wainstein, 1969) - Austroteneriffia tadjikistanica
Yemen (incl. Socotra)(Ueckermann, van Harten & Smith Meyer, 2006) – Austroteneriffia tadjikistanica, Parateneriffia sp.
NORTH and CENTRAL AMERICA
Mexico (McDaniel, Morihara et Lewis, 1976; López-Campos & Vázquez-Rojas, 2010; Beron, in prep.) - Neoteneriffiola uta, Teneriffia mexicana
USA (Tibbetts, 1958) - Neoteneriffiola uta
SOUTH AMERICA
Argentina (Besch, 1962) - “Austroteneriffia” andrei
Brazil (Bernardi et al., 2012) – Neoteneriffiola
Paraguay (Thor, 1911) - Parateneriffia bipectinata
Peru (Beron, in prep.) - Teneriffiidae
OCEANIA
Australia (Womersley, 1935; Judson, 1995) - Austroteneriffia hirsti, A. leei
Pseudocheylidae Oudemans, 1909
Small free - living mites, found under bark, in moss, soil or in bird’s nest. Three genera with 24 known species. Widely distributed, but rarely found.
Pseudocheylidae Oudemans, 1909:
Type genus: Pseudocheylus Berlese, 1888
Anoplocheylus Berlese, 1910: 210 (subgenus)
Anoplocheylus (Berlese): Baker & Atyeo, 1964: 262 (genus)
Type species: Pseudocheylus (Anoplocheylus) europaeus Berlese, 1910
Rhagina Womersley, 1935: 336 (Syn. by Baker & Wharton, 1952)
Type species: Rhagina protea Womersley, 1935
Anoplocheylus aegypticus Baker & Atyeo, 1964: 267; Gerson, 1967: 367; van Dis & Ueckermann, 1991: 113
Type locality: Ismaelia - Port Said Road, 50 kilometers
Distributon: Egypt
Anoplocheylus bonabjadidiensis Navaei-Bonab et al., 2011: 421; Navaei-Bonab & Kazazi, 2013: 28
Type locality: Bonabjadid, Marand, East Azerbaijan Province, Iran
Distributon: Iran
Anoplocheylus brasiliensis Khaustov & Tolstikov, 2015: 60
Type locality: Rio de Janeiro, Leme, Atlantic rain forest
Distribution: Brazil
Anoplocheylus brevisetosus Ueckermann & Khanjani, 2004: 59
Type locality: Rustenburg, North West Province
Distributon: South Africa
Anoplocheylus clavatus Baker & Atyeo, 1964: 265; van Dis & Ueckermann, 1991: 106
Type locality: Gnangara (16 miles north of Perth), Western Australia
Distributon: Australia
Anoplocheylus corticicola Fungarworn & Butcher, 2015: 2
Type locality: Bangkok, Pathumwan Dist., Chulalongkorn Univ. Campus
Distribution: Thailand
Pseudocheylus (Anoplocheylus) europaeus Berlese, 1910: 210
Anoplocheylus europaeus (Berlese): Baker & Atyeo, 1964: 263; van Dis & Ueckermann, 1991: 106; Livshitz & Mitrofanov, 1973: 771; Ueckermann & Khanjani, 2004: 55
Type locality: Palermo
Distributon: Italy (Sicily)
Anoplocheylus hapsiscutus Flechtmann, 1971: 49
Type locality: Piracicaba, São Paulo
Distributon: Brazil
Anoplocheylus kazemii Bagheri, in Bagheri et al., 2013: 292
Type locality: Iran
Distributon: Iran
Anoplocheylus malayeriensis Ueckermann & Khanjani, 2004: 55; Navaei-Bonab & Kazazi, 2013: 28
Type locality: Mey, 23 km from Malãyer to Arãk
Distributon: Iran
Anoplocheylus marivaniensis Khanjani, Hoseini & Amini, 2014: 186
Type locality: Merivan, Kurdistan Province
Distributon: Iran
Anoplocheylus paraclavatus van Dis & Ueckermann, 1991: 106; Ueckermann & Khanjani, 2004: 55
Type locality: Edenville, Pongola
Distributon: South Africa
Rhagina protea Womersley, 1935: 336
Anoplocheylus protea (Womersley): Baker & Atyeo, 1964: 264; van Dis & Ueckermann, 1991: 111; Ueckermann & Khanjani, 2004: 57
Type locality: Glen Osmond, South Australia
Distributon: Australia
Anoplocheylus qorvehiensis Khanjani, Hoseini & Amini, 2014: 189
Type locality: Qorveh, Kurdistan Province
Distributon: Iran
Anoplocheylus sinai Bagheri, in Bagheri et al., 2013: 290; Navaei-Bonab & Kazazi, 2013: 28
Type locality: Iran
Distributon: Iran
Anoplocheylus tauricus Livshitz & Mitrofanov, 1973: 770; van Dis & Ueckermann, 1991: 113; Ueckermann & Khanjani, 2004: 65; Bayram & Cobanoğlu, 2006:
Type locality: cape Kazantip, Crimea
Distributon: Turkey, Crimea, Iran
Anoplocheylus tellustrus van Dis & Ueckermann, 1991: 109
Anoplocheylus reticulatus van Dis & Ueckermann, 1991: 112. Type: Pongola, Edenville, South Africa. Syn. by Ueckermann & Khanjani, 2004: 61
Type locality: Pongola, Edenville
Distributon: South Africa
Anoplocheylus transiens Delfinado & Baker, 1974: 209; van Dis & Ueckermann, 1991: 113; Ueckermann & Khanjani, 2004: 61
Type locality: Heckscher Park, Long Islang, New York
Distributon: USA
Neocheylus Trägårdh, 1906: 870
Type species: Neocheylus natalensis Trägårdh, 1906
Neocheylus collis Baker & Atyeo, 1964: 270
Type locality: Mount Toolbrunup (Stirling Ranges, 300 miles south from Perth), W. Australia
Distributon: Australia
Neocheylus natalensis Trägårdh, 1906: 870
Type locality: Natal
Distributon: South Africa
Neocheylus nidicolus Lawrence, 1954: 74
Type locality: Karamoja (Uganda)
Distributon: Uganda
Pseudocheylus Berlese, 1888: 189
Type species: Pseudocheylus biscalatus Berlese, 1888
Cheyletiella americana Ewing, 1909: 429
Pseudocheylus americanus (Ewing): Baker & Wharton, 1952: 226; Skvarla et al., 2013: 401
Type locality: Urbana, Illinois
Distributon: USA (Illinois)
Pseudocheylus biscalatus Berlese, 1888: 189; Flechtmann, 1971: 33
Type locality: Rio-Apa (Paraguay), Mato Grosso (Brazil)
Distribution: Brazil, Paraguay
EUROPE
Italy (contin.)
Sicily (Berlese, 1910) – Anoplocheylus europaeus
Russia
Crimea (Livshitz & Mitrofanov, 1973) – Anoplocheylus tauricus
Turkey (all) (Bayram & Cobanoğlu, 2006) - Anoplocheylus tauricus
AFRICA
Egypt (Baker & Atyeo, 1964) – Anoplocheylus aegyptiacus
South Africa (Trägårdh, 1906; van Dis & Ueckermann, 1991; Ueckermann & Khanjani, 2004) – Anoplocheylus brevisetosus, A. paraclavatus, A. tellustrus (= reticulatus), Neocheylus natalensis
Uganda (Lawrence, 1954) – Neocheylus nidicolus
ASIA
Iran (Ueckermann & Khanjani, 2004; Navaei-Bonab et al., 2011; Bagheri et al., 2013; Navaei-Bonab & Kazazi, 2013: 28; Khanjani, Hoseini & Amini, 2014) – Anoplocheylus bonabjadidiensis, A. kazemii, A. malayeriensis, A. marivaniensis, A. qorvehiensis, A. sinai, A. tauricus
Israel (Gerson, 1967) - Anoplocheylus aegyptiacus
Thailand (Fuangarworn & Butcher, 2015) – Anoplocheylus corticicola
NORTH AMERICA
Canada (Smith & Lindquist,1979) – one species
USA (Ewing, 1909; Delfinado & Baker, 1974; Skvarla et al., 2013) – Anoplocheylus transiens, Pseudocheylus americanus
SOUTH and CENTRAL AMERICA
Brazil (Berlese, 1888; Flechtmann, 1971; Khaustov & Tolstikov, 2015) – Anoplocheylus brasiliensis, A. hapsiscutus, Pseudocheylus biscalatus
Paraguay (Berlese, 1888) - Pseudocheylus biscalatus
OCEANIA
Australia (continent) (Womersley, 1935; Baker & Atyeo, 1964) – Anoplocheylus clavatus, A. protea, Neocheylus collis
Stigmocheylidae Berlese, 1910: 209
Small family (five known species) of soil mites.
Stigmocheylidae Berlese, 1910b: 209; Kethley, 1990: 698
Type genus: Stigmocheylus Berlese, 1910
Stigmocheylus Berlese, 1910a: 209
Type species: Stigmocheylus brevisetus Berlese, 1910
Stigmocheylus americanus Bochkov, 2008: 315
Type locality: Las Cruces, Jornada, New Mexico
Distribution: USA (New Mexico)
Stigmocheylus brevisetus Berlese, 1910a: 209
Type locality: Pisa
Distribution: Italy
Stigmocheylus oberon Bochkov, 2008: 314
Type locality: New Mexico
Distribution: USA (New Mexico)
Stigmocheylus persicus Damavandian & Paktinat-Saeij, 2018: 2160
Type locality: Noor, Mazandaran province, northern Iran
Distribution: Iran
Hemitarsocheylus pilosus Soliman & Zaher, 1975: 105
Stigmocheylus pilosus (Soliman et Zaher): Bochkov, 2008: 314
Type locality: Giza
Distribution: Egypt
EUROPE
Italy (Berlese, 1910) - Stigmocheylus brevisetus
AFRICA
Egypt (Soliman & Zaher, 1975) – Stigmocheylus pilosus
ASIA
Iran (Naghibinejad, Ahadiyat, & Ostovan, 2009; Damavandian & Paktinat-Saeij, 2018) – Stigmocheylus persicus
NORTH AMERICA
USA (Bochkov, 2008) – Stigmocheylus americanus, S. oberon
Superfamily Caeculoidea Ewing, 1934
Phalanx Caeculata Wainstein, 1965: 77
The superfamily contains one family – Caeculidae.
Family Caeculidae Berlese, 1883
Dufour (1832) started the study of these peculiar mites, called after the mythical son of Vulcan. They prefer the open rocky habitats and are known already from all continents. The second genus in the family (Procaeculus) was described by Jacot (1936), more than a century after the first. The low intensity of the studies of Caeculidae of many countries is shown by the multitude of species, found in the USA by several authors, mostly by Mulaik (1945), and (after 1952) in Spain, Morocco, N. Macedonia and Chad by H. Franz, who revised the family (Franz, 1952), describing the new genera Allocaeculus and Microcaeculus, and compiling a list of all known Caeculidae. Very profound research on the family was enterprised by Y. Coineau since 1963. He described several new species, mainly from Southern Africa, and the new genera Calocaeculus, Andocaeculus, and Neocaeculus. Three new species have been described from Australia (Taylor, Gunawardene & Kinnear, 2013, Taylor, 2014) and one from Thailand (Fuangarworn & Butcher, 2015) and Canada (Bernard et al., 2020). One new Allocaeculus was described from Slovakia (Mangová et al., 2014), another one from Turkey (Per et al., 2017). New checklist of the family was published by Taylor et al. (2013).
Fossil species have been described from amber from Poland, Myanmar, Mexico, and Dominican Republic (Coineau & Magowski, 1994, Coineau & Poinar, 2000, Rivas et al., 2016, Porta et al., 2019).
Now the family is considered to contain at least 114 sp. in seven genera.
Fam. Caeculidae – 114 sp.
Allocaeculus Franz, 1952 – 36 sp.
Andocaeculus Coineau, 1974 - 5
Caeculus Dufour, 1832 - 21
Calocaeculus Coineau, 1974 - 1
Microcaeculus Franz, 1952 - 25
Neocaeculus Coineau, 1967 – 12
Procaeculus Jacot, 1936 - 13
Incertae sedis - 1
Hoplopini Canestrini & Fanzago, 1877: 481
Caeculidae Berlese, 1883
Type genus: Caeculus Dufour, 1832
Allocaeculus Franz, 1952: 108
Type species: Allocaeculus relictus Franz, 1952
Allocaeculus andalusiacus Franz, 1952: 113
Type locality: Sevilla, Cinca de Pino, Andalusia
Distribution: Spain
Allocaeculus catalanus Franz, 1952: 2; 1954: 210; 1955: 12
Type locality: Gerona (Spain), Eastern Pyrenees (France)
Distribution: France, Kazachstan, Russia, Spain
Allocaeculus circinatus Coineau, 1974b: 278
Type locality: Ghaub Farm in Ovidi highlands, 25 km S Tsumeb
Distribution: Namibia
Caeculus dubius Kulczynski, 1901: 367
Allocaeculus dubius (Kulczynski): Franz, 1952: 122
Type locality: Algeria
Distribution: Algeria
Allocaeculus echinatus Franz, 1952: 118
Type locality: Sevilla, Andalusia
Distribution: Spain
Allocaeculus erinaceus Coineau, 1974b: 277
Type locality: Ingwavuma
Distribution: South Africa
Allocaeculus grandjeani Franz, 1957: 10
Type locality: Khenifra
Distribution: Morocco
Allocaeculus hirsutus Coineau, 1974b: 277
Type locality: Glencoe
Distribution: South Africa
Caeculus hoggarensis André, 1936: 111
Allocaeculus hoggarensis (André): Franz, 1952: 122; Piffl, 1959: 54
Type locality: Hoggar, Mt Ilaman
Distribution: Algeria
Allocaeculus hystriciformis Franz, 1952: 120
Type locality: Mte Arastepa in Sierra Ronda
Distribution: Spain
Caeculus hystrix Lawrence, 1939: 542
Allocaeculus hystrix (Lawrence): Franz, 1952: 122
Type locality: Zululand, Hluhluve Game Reserve
Distribution: South Africa
Allocaeculus indicus Piffl, 1959: 52
Type locality: Saidu Sharif, Swat Area, NW Pakistan
Distribution: Pakistan
Allocaeculus kalahariensis Coineau, 1974b: 277
Type locality: Kalahari Gemsbok National Park
Distribution: South Africa
Allocaeculus kenyae Franz, 1964: 82
Type locality: 8 km N Nairobi
Distribution: Kenya
Allocaeculus kocheri Franz, 1964: 84
Type locality: Oukaimeden, 2300 m
Distribution: Morocco
Allocaeculus logonensis Franz, 1957: 6
Type locality: Logon, Deressia nr. Lai
Distribution: Chad
Allocaeculus meseticola Franz, 1952: 114; 1955: 14
Type locality: Alcala de Henares
Distribution: Spain
Caeculus mosambicensis André, 1936: 115
Allocaeculus mosambicensis (André): Franz, 1952: 122; 1957: 12; Piffl, 1959: 54
Type locality: Vila Pery
Distribution: Mozambique
Allocaeculus multispinosus Franz, 1955: 11; 1960: 86
Type locality: Puerto de Perales, Sierra de Gata
Distribution: Spain, Turkey
Allocaeculus nigeriensis Coineau, 1974b: 278
Type locality: Ibadan
Distribution: Nigeria
Allocaeculus ouadaiensis Franz, 1957: 7
Type locality: Mont Kelinguen nr. Abecher
Distribution: Chad
Caeculus pilosus Lawrence, 1938: 211
Allocaeculus pilosus (Lawrence): Franz, 1952: 122; 1957: 12; Piffl, 1959: 54
Type locality: N. Damaraland
Distribution: Namibia
Allocaeculus relictus Franz, 1952: 111; 1954b: 369; 1955: 15; Winkler, 1953: 55
Type locality: “Kalvarienberg b. Gumpoldskirchen”, Austria
Distribution: Austria, Czech Rep.
Allocaeculus sandbergensis Mangová, Krumpál & Ľuptáčik, 2014: 214
Type locality: National nature reserve Devínska Kobyla near Sandberg
Distribution: Slovakia
Allocaeculus sarhroi Franz, 1964: 86
Type locality: Tizi-n-Tazazerte, Jbel Sarhro, 2100 m
Distribution: Morocco
Allocaeculus schusteri Franz, 1960: 83
Type locality: Ohrid
Distribution: N. Macedonia
Caeculus sclerodermatus André, 1936: 106
Allocaeculus sclerodermatus (André): Franz, 1952: 122; 1957: 12; Piffl, 1959: 54
Type locality: South Tunisia, Nefta
Distribution: Tunisia
Hoplopus sculptus Karpelles, 1893: 39
Caeculus sculptus Karpelles, 1893: 39
Allocaeculus sculptus (Karpelles): Franz, 1952: 122; Piffl, 1959: 54
Type locality: Hungary
Distribution: Hungary
Caeculus senegalensis André, 1938: 264
Allocaeculus senegalensis (André): Taylor et al., 2013: 448
Type locality: Dakar
Distribution: Senegal
Allocaeculus sirwani Paktinat-Saeij, Ghobari & Samani, 2020: 834
Type locality: Marivan city, Kurdistan Province
Distribution: Iran
Caeculus spatulifer Michael, 1890: 432; Vitzthum, 1933: 87
Allocaeculus spatulifer (Michael): Franz, 1952: 122
Type locality: Algeria
Distribution: Algeria, Croatia, Spain
Allocaeculus spinosissimus Franz, 1952: 116
Allocaeculus spinosissimus rondae Franz, 1952: 118. Type locality: Mte Arastepa in Sierra Ronda
Type locality: Sierra Guadarrama above Escorial
Distribution: Spain
Caeculus sudanensis Trägårdh, 1904: 70
Allocaeculus sudanensis : Franz, 1952: 122
Type locality: “Gebelein am weissen Nil”
Distribution: Sudan
Allocaeculus tenerifae Franz, 1965: 67
Type locality: Las Madrigueras (S. Tenerife)
Distribution: Spain (Canary Is.)
Allocaeculus tschadensis Franz, 1957: 8; 1964: 86
Type locality: Mont Kelinguen nr. Abecher (Chad)
Distribution: Chad, Morocco, Senegal
Allocaeculus tschadensis rifensis Franz, 1960: 84
Type locality: Zoco el Arba, road Tetuan - Xauen
Distribution: Morocco
Allocaeculus turcicus Per, Doğan, Zeytun & Ayyıldız, 2017: 371
Type locality: Karamlıkdere Valley, Yozgat, 844 m
Distribution: Turkey
Andocaeculus Coineau, 1974b: 279
Type species: Microcaeculus brundini Franz, 1962
Andocaeculus beatrizrosso Porta, Pizarro-Araya & Ramírez, 2021: 42
Type locality: Argentina
Distribution: Argentina
Microcaeculus Brundini Franz, 1962: 99
Andocaeculus brundini (Franz): Coineau, 1974b: 279
Type locality: Arroyo Lopez, W from Bariloche (Prov. Rio Negro)
Distribution: Argentina
Andocaeculus burmeisteri Porta, Pizarro-Araya & Ramírez, 2021: 28
Type locality: Argentina
Distribution: Argentina
Andocaeculus caioi Ott & Ott, 2014: 356
Type locality: Fazenda Rincão dos Moraes, Área de Proteção Ambiental do Rio Ibirapuitã, Sant’Ana do Livramento, Rio Grande do Sul
Distribution: Brazil
Microcaeculus weinrauchi Franz, 1964: 89
Andocaeculus weinrauchi :
Type locality: Puna “El Infernillo”, 3000 m
Distribution: Argentina
Caeculus Dufour, 1832: 289
Hoplopus Canestrini & Fanzago, 1877: 480 (replacement name)
Type species: Caeculus echinipes Dufour, 1832
Pseudocaeculus americanus (Banks): Taylor et al., 2013: 448.
Caeculus americanus Banks, 1899: 221; McDaniel & Boe, 1990: 722
Type locality: Palm Springs, California
Distribution: USA (California)
Caeculus archeri Mulaik, 1945: 13; Franz, 1952: 121; McDaniel & Boe, 1990: 723
Type locality: Monte Sano, Madison Co., Alabama
Distribution: USA (Alabama, Tennessee)
Caeculus calechius Mulaik, 1945: 5; Franz, 1952: 121; McDaniel & Boe, 1990: 723
Type locality: Edinbu.rg, Hidalgo Co., Texas
Distribution: USA (Montana, Texas, Utah)
Caeculus cassiopeiae Bernard & Lumley, in: Bernard, Lumley, Buck & Cobb. 2020: 2
Type locality: Alberta, Medicine Hat, near Gas City Campground; 50°2.23’N, 110°43.56’W
Distribution: Canada
Caeculus clavatus Banks, 1906: 135; Franz, 1952: 121; McDaniel & Boe, 1990: 722
Type locality: USA (California)
Distribution: USA (California)
Caeculus cremnicolus Enns, 1958: 107; Skvarla, Fisher & Dowling, 2013: 401; Bernard et al., 2020:
Type locality: Baskett Wildlife Research and Education Area, Devil’s Backbone; Easley; Wilton, Missouri
Distribution: Canada, USA (Arkansas, Missouri)
Caeculus crosleyi Coineau, 1974b: 276
Type locality: DeKalb & Rockdale Counties
Distribution: USA (Georgia)
Caeculus dorotheae Mulaik, 1945: 9; Franz, 1952: 121; McDaniel & Boe, 1990: 722 (dortheae)
Type locality: Lockes, Nye Co., Nevada
Distribution: USA (Arizona, Nevada)
Caeculus echinipes Dufour, 1832: 289; André, 1927: 168; 1932: 485; Irk, 1939: 154; Willmann, 1951: 148; Franz, 1952: 98; 1954a: 212; 1954b: 369; Kratochvil, 1962: 174; Schmölzer, 1962: 286; Coineau & Haupt, 1976
Hoplopus echinipes : Canestrini & Fanzago, 1877: 477
Type locality: Moxente (Valencia)
Distribution: Algeria, Austria, Bulgaria, France, Germany, Greece, Italy, Spain, Switzerland
Caeculus echinipes crosbyi Jacot, 1936: 18; Franz, 1952: 121
Caeculus echinipes : Franz, 1955: 14
Type locality: Pic du Midi de Bigore, Pyrenees
Distribution: France
Caeculus gertschi Mulaik, 1945: 8; Franz, 1952: 121
Type locality: Rio Grande City, Starr Co., Texas
Distribution: USA (Texas)
Caeculus hardyi Mulaik & Allred, 1954: 36; McDaniel & Boe, 1990: 723
Type locality: Laredo, Harlingen (New Mexico, Texas), San Pedro (Mexico)
Distribution: Mexico, USA
Caeculus hypopachus Mulaik, 1945: 10; Franz, 1952: 122; McDaniel & Boe, 1990: 723
Type locality: Edinburg, Hidalgo Co., Texas, in stomach of toad (Hypopachus cuneus)
Distribution: USA (Texas)
Caeculus janetae Higgins & Mulaik, 1957b: 267
Type locality: SW USA
Distribution: USA
Caeculus kerrulius Mulaik, 1945: 8; Franz, 1952: 122; McDaniel & Boe, 1990: 722
Type locality: Kerrville, KerrCo., Texas
Distribution: USA (Texas)
Caeculus krantzi Coineau, 1974b: 276
Type locality: Manzanita, 32 mi E of Sweet Home, Oregon
Distribution: USA (Oregon)
Caeculus lewisi McDaniel & Boe, 1990: 716
Type locality: South Dakota
Distribution: USA (South Dakota)
Caeculus mariae Higgins & Mulaik, 1957a: 27
Type locality: Oregon
Distribution: USA (Oregon)
Caeculus pettiti Nevin, 1943: 389; 1945: 195; Franz, 1952: 122; McDaniel & Boe, 1990: 723
Type locality: Virginia
Distribution: USA
Caeculus tipus Mulaik, 1945: 6; Mulaik & Allred, 1954: 38; Franz, 1952: 122; McDaniel & Boe, 1990: 723
Type locality: Edinburg, Hidalgo Co., Texas
Distribution: USA (Utah, Texas)
Caeculus valverdius Mulaik, 1945: 6; Franz, 1952: 122; McDaniel & Boe, 1990: 72
Type locality: Edinburg, Hidalgo Co., Texas
Distribution: USA (Arizona, Texas)
Caeculus veracruzensis Porta & Vázquez Rojas, 2020: 473
Type locality: La Mancha, Veracruz
Distribution: Mexico
Calocaeculus Coineau, 1974b: 279
Type species: Calocaeculus lawrencei Coineau, 1974
Calocaeculus lawrencei Coineau, 1974b: 279
Type locality: Pearston in Somerset East
Distribution: South Africa
Microcaeculus Franz, 1952: 101
Type species: Microcaeculus austriacus Franz, 1952
Type species: Sclerocaeculus namibensis (Piffl, 1965)
Caeculus Andrei Lawrence, 1939: 540
Microcaeculus andrei (Lawrence): Franz, 1952: 122; Halliday, 2005: 28
Type locality: South Africa
Distribution: South Africa
Microcaeculus austriacus Franz, 1952: 103; 1954b: 369; Pieper, 1970: 106; Thaler et al., 1993: 311
Type locality: “Kalvarienberg b. Gumpoldskirchen a. O-Hang des Anningers südl. v. Wien” (Austria)
Distribution: Austria, Czech Rep., Germany
Microcaeculus canariensis Franz, 1965: 68
Type locality: Ortes Los Cristianos (Tenerife)
Distribution: Spain (Canary Is.)
Microcaeculus castrii Franz, 1964: 87
Type locality: Cordillera de la Costa, Prov. De Coquimbo, Alcones (and several other localities)
Distribution: Chile
Caeculus deserticolus Lawrence, 1939: 537
Microcaeculus deserticolus (Lawrence): Franz, 1952: 122
Sclerocaeculus deserticolus (Lawrence) – Coineau, 1974: 59
Type locality: Kakamas, Namaqualand
Distribution: South Africa
Microcaeculus franzi Coineau, 1968: 143
Type locality: Ghisonaccia, Corsica
Distribution: Corsica
Microcaeculus graecus Franz, 1964: 91
Type locality: Arachova, 2000 m
Distribution: Greece
Microcaeculus hispanicus Franz, 1952: 105; Coineau, 1966: 23
Type locality: Alcalá de Henares (Spain)
Distribution: France, Spain
Caeculus insolitus André, 1935: 113
Microcaeculus insolitus (André): Franz, 1952: 122
Type locality: Grande Salvage
Distribution: Salvages Islands (Portugal)
Microcaeculus insularis Franz, 1964: 90
Type locality: The Big Island against Essaouira (Mogador)
Distribution: Morocco
Caeculus laoshanensis Jacot, 1936: 20
Microcaeculus laoshanensis (Jacot): Franz, 1952: 122
Type locality: Laoshan
Distribution: China
Caeculus liguricus Vitzthum, 1933: 87; Willmann, 1954: 237
Microcaeculus liguricus (Vitzthum): Franz, 1952: 122
Type locality: Coast of Liguria
Distribution: Czech Rep., Italy, Morocco
Caeculus monticolus Lawrence, 1939: 539
Microcaeculus monticolus (Lawrence): Franz, 1952: 122; Halliday, 2005: 28
Type locality: South Africa
Distribution: South Africa
Caeculus muscorum Lucas, 1849: 307
Microcaeculus muscorum (Lucas): Franz, 1952: 122
Type locality: Algeria
Distribution: Algeria
Microcaeculus namibensis Piffl, 1965: 1
Sclerocaeculus namibensis (Pifl): Taylor et al., 2013: 449
Type locality: Namibia
Distribution: Namibia
Microcaeculus nicoleae Coineau, 1974b: 276
Type locality: Bloeddrif
Distribution: South Africa
Microcaeculus nudus Franz, 1964: 88
Type locality: Serro El Roble, Cordillera de la Costa, Provincias de Valparaiso y .Santiago, Gipfel del Cerro
Distribution: Chile
Microcaeculus pica Otto, 2009: 6
Type locality: New South Wales, Guerilla Bay
Distribution: Australia (NSW)
Caeculus pisanus André, 1936: 123; Franz, 1952: 122 (Inc. sed.)
Microcaeculus pisanus (André): Taylor et al., 2013: 449
Type locality: Italy
Distribution: Italy
Microcaeculus sabulicola Franz, 1952: 107; Coineau, 1969b: 207
Type locality: “Andalusien, am Sandstrand westl. Algeciras”(Spain)
Distribution: France, Morocco, Portugal, Spain
Caeculus sinensis Jacot, 1936: 22
Microcaeculus sinensis (Jacot): Franz, 1952: 122
Type locality: China
Distribution: China
Caeculus sinensis taishanicus Jacot, 1936: 22
Microcaeculus sinensis taishanicus (Jacot): Franz, 1952: 122
Type locality: Taishan
Distribution: China
Microcaeculus hispanicus steineri Franz, 1955: 9
Microcaeculus steineri Franz:
Type locality: Sierra Nevada, 2700 m
Distribution: Spain
Microcaeculus steineri delamarei Coineau, 1974b: 276
Type locality: Font Estramer, 12 km N from Perpignan
Distribution: France
Microcaeculus stephani Coineau & Haupt, 1978: 622
Type locality: Formentera, Playa Mitjora
Distribution: Spain (Formentera, Pityusas)
Caeculus tibbettsi Higgins & Mulaik, 1958: 17
Microcaeculus tibbettsi (Higgins et Mulaik): Franz, 1962: 100
Type locality: Seoul
Distribution: Korea
Caeculus uchidai Asanuma, 1940: 271
Microcaeculus uchidai (Asanuma): Taylor et al., 2013: 449
Type locality: Japan
Distribution: Japan
Neocaeculus Coineau, 1967: 58
Type species: Neocaeculus luxtoni Coineau, 1967
Neocaeculus bornemisszai Coineau & Enns, 1969: 690
Type locality: Kimberley, 100 km S Wyndham
Distribution: Australia
Caeculus bruchi Berlese, 1916: 289; Franz, 1952: 122 (Inc. sed., Microcaeculus ?)
Neocaeculus bruchi : Coineau, 1974: Taylor et al., 2013: 449
Type locality: “La Plata”
Distribution: “La Plata”(Argentina)
Neocaeculus imperfectus Taylor, Gunawardene & Kinnear, 2013: 440
Type locality: Barrow Island, Western Australia
Distribution: Australia
Neocaeculus johnstoni Coineau, 1974: 68
Type locality: Lord Howe Island
Distribution: Lord Howe Island
Neocaeculus kinnearae Taylor, 2014: 348
Type locality: Barrow Island, Western Australia
Distribution: Australia
Neocaeculus knoepffleri Coineau & Enns, 1969: 679
Type locality: Kimberley, 100 km S Wyndham
Distribution: Australia
Neocaeculus lamorali Coineau, 1974b: 278
Type locality: De Waal farm, 10 km N of Leonardville
Distribution: Namibia
Neocaeculus luxtoni Coineau, 1967a: 58
Type locality: Alexandra (Otago)
Distribution: New Zealand
Neocaeculus nudonates Taylor, 2014: 343
Type locality: Barrow Island, Western Australia
Distribution: Australia
Neocaeculus orientalis Fuangarworn & Butcher, 2015: 251
Type locality: Thailand
Distribution: Thailand
Neocaeculus setecidades Ott & Ott, 2018: 2
Type locality: Parque Nacional Sete Cidades (04°06’44.2”S - 41°41’47.7”W), sample site PNC7C 0323, Piracuruca, Piauí
Distribution: Brazil
Neocaeculus womersleyi Coineau, 1974: 68
Type locality: Risdon
Distribution: Tasmania
Procaeculus Jacot, 1936: 25
Type species: Procaeculus bryani Jacot, 1936
Procaeculus aitkeni Coineau, 1967b: 719
Type locality: Trinidad
Distribution: Trinidad
Caeculus brevis Mulaik, 1945: 11; Franz, 1952: 121; McDaniel & Boe, 1990: 723
Procaeculus brevis (Mulaik): Taylor et al., 2013: 449.
Type locality: Houston, Harris Co., Texas
Distribution: USA (Texas)
Procaeculus bryani Jacot, 1936: 25; Franz, 1952: 121; Coineau, 1967b: 713
Type locality: Peale Island, Wake Island, Western end of the Hawaiian Island group
Distribution: Hawaiian Islands
Procaeculus coineaui Porta et al., 2016: 38
Type locality: Myanmar
Distribution: Myanmar (Burma)
Procaeculus dominicensis Coineau & Poinar, 2000: 143
Type locality: Mine in Northern Cordillera, El Mamey Formation (Upper Eocene)
Distribution: Dominican Republic
Procaeculus eridanosae Coineau & Magowski, 1994: 243
Type locality: Baltic amber (Eocene)
Distribution: Poland
Procaeculus magnus Coineau, 1974b: 279
Type locality: Mazatlan, Sin.
Distribution: Mexico
Caeculus mexicanus Mulaik & Allred, 1954: 34; McDaniel & Boe, 1990: 723
Procaeculus mexicanus (Mulaik et Allred): Coineau, 1967b: 718
Type locality: San Luis Potosi (also intercepted in Texas)
Distribution: Mexico
Caeculus orchidicolis Mulaik & Allred, 1954: 30
Procaeculus orchidicolis (Mulaik et Allred): Coineau, 1967b: 718
Type locality: Chilpancingo (also intercepted in Texas)
Distribution: Mexico
Caeculus oregonus Mulaik & Allred, 1954: 28; McDaniel & Boe, 1990: 723
Procaeculus oregonus (Mulaik et Allred): Coineau, 1967b: 718
Type locality: Oak Ridge, Oregon
Distribution: USA (Oregon)
Caeculus potosi Mulaik & Allred, 1954: 32
Procaeculus potosi (Mulaik et Allred): Coineau, 1967: 718
Type locality: Tamazunchale (also intercepted in Texas)
Distribution: Mexico
Caeculus puertoricus Mulaik, 1945: 12; Franz, 1952: 122
Procaeculus puertoricus (Mulaik): Taylor et al., 2013: 450
Type locality: Como Springs
Distribution: Puerto Rico
Caeculus willmannii Vitzthum, 1933: 86
Procaeculus willmanni (Vitzthum): Franz, 1952: 121
Procaeculus aitkeni Coineau, 1967a: 719; syn. by Taylor et al., 2013: 450
Type of P. aitkeni: Bush-Bush Forest, Trinidad
Type locality: Guatemala
Distribution: Guatemala
Caeculus hirtipes Berlese, 1910: 369; Franz, 1952: 122 (Inc. sed.)
Type locality: “Mechieria”
Distribution: Algeria
Caeculus hirtipes var. graecus Berlese, 1910: 369; Franz, 1952: 122 (Inc. sed.)
Type locality: Naxos
Distribution: Greece
EUROPE
Austria (Irk, 1939; Willmann, 1951; Franz, 1952, 1955; Thaler, 1979; Thaler, Knoflach & Meyer, 1993) – Allocaeculus relictus, Caeculus echinipes, Microcaeculus austriacus
Bulgaria (Kratochvil, 1962) – Caeculus echinipes
Croatia (Vitzthum, 1933) – Allocaeculus spatulifer
Czech Rep. (Franz, 1952; Winkler, 1953; Willmann, 1954) – Allocaeculus relictus, Microcaeculus austriacus, M. liguricus
France (Moniez, 1893; André, 1932; Jacot, 1936; Franz, 1954; Coineau, 1966, 1969b, 1974) – Allocaeculus catalanus, Caeculus echinipes, C. echinipes crosbyi, Microcaeculus hispanicus, M. sabulicola, M. steineri delamarei
Corsica (Coineau, 1968) - Microcaeculus franzi
Germany (Pieper, 1970; Coineau & Haupt, 1976) – Caeculus echinipes, Microcaeculus austriacus
Greece (Berlese, 1910; Sellnick, 1931; Franz, 1964) – Caeculus echinipes, Microcaeculus graecus; inc. sed.: Caeculus hirtipes var. graecus
Italy (Vitzthum, 1933; Franz, 1954) – Allocaeculus sculptus, Caeculus echinipes, Microcaeculus liguricus, M. pisanus
Sicily (Berlese, 1883) – Caeculus echinipes
Hungary (Karpelles, 1893) - Caeculus
N. Macedonia (Franz, 1960) – Allocaeculus schusteri
Poland (Coineau & Magowski, 1994) - Procaeculus eridanosae (fossil)
Portugal (Coineau, 1969b) – Microcaeculus sabulicola
Salvages Islands (André, 1935) – Microcaeculus insolitus
Slovakia (Mangová, Krumpál & Ľuptáčik, 2014) – Allocaeculus sandbergensis
Spain (Dufour, 1832; Vitzthum, 1933; Franz, 1952, 1954, 1955, 1960; Mihelčič, 1958) – Allocaeculus andalusiacus, A. catalanus, A. echinatus, A. hystriciformis, A. meseticola, A. multispinosus, A. spatulifer, A. spinosissimus (incl. A. spinosissimus rondae), Caeculus echinipes, Microcaeculus hispanicus, M. sabulicola, M. steineri
Canary Is. (Franz, 1965) – Allocaeculus tenerifae, Microcaeculus canariensis
Balear and Pityusas Islands (Coineau & Haupt, 1977) – Microcaeculus stephani [Formentera]
Switzerland (Schweizer, 1951; Schweizer & Bader, 1963; Pieper, 1970) – Caeculus echinipes
Turkey (Karaca et al., 2012; Per, Doğan, Zeytun & Ayyıldız, 2017; Per, Doğan & Ayyıldız, 2019) – Allocaeculus multispinosus, A. turcicus
AFRICA
Algeria (Lucas, 1849; Michael, 1890; Kulczynski, 1901; André, 1927; 1935) – Allocaeculus dubius, A. hoggarensis, A. muscorum, A. spatulifer, Caeculus echinipes
Chad (Franz, 1957, 1964) – Allocaeculus logonensis, A. ouadaiensis, A. tschadensis
Kenya (Franz, 1964) - Allocaeculus kenyae
Morocco (Coineau, 1969b; Franz, 1960, 1964) – Allocaeculus grandjeani, A. kocheri, A. sarhroi, A. tschadensis rifensis, Microcaeculus insularis, M. liguricus, M. sabulicola
Mozambique (André, 1936) - Allocaeculus mosambicensis
Namibia (Lawrence, 1938; Piffl, 1965; Coineau, 1974b) – Allocaeculus circinatus, A. pilosus, Microcaeculus namibensis, Neocaeculus lamorali
Nigeria (Coineau, 1974b) – Allocaeculus nigeriensis
Senegal (André, 1938; Franz, 1964) – Allocaeculus sp. cf. andalusiacus, A. tschadensis, A. senegalensis
Somalia (Berlese, 1916) – “Caeculus pr. spathuliger Michael”
South Africa (Lawrence, 1939; Coineau, 1974b; Halliday, 2005) – Allocaeculus erinaceus, A. hirsutus, A. hystrix, A. kalahariensis, Calocaeculus lawrencei, Microcaeculus andrei, M. deserticolus, M. monticolus, M. nicoleae
Sudan (Trägårdh, 1904) - Allocaeculus sudanensis
Tunisia (André, 1936) – Allocaeculus sclerodermatus
ASIA
China (Jacot, 1936) - Microcaeculus laoshanensis, M. sinensis
Iran (Paktinat-Saeij, Ghobari & Samani, 2020) – Allocaeculus sirwani
Japan (Asanuma, 1940) – Microcaeculus uchidai
Kazachstan – Allocaeculus catalanus
Korea (Higgins & Mulaik, 1958) – Caeculus tibbettsi
Myanmar (Burma)(Porta et al., 2019) – Procaeculus coineaui
Pakistan (Piffl, 1959) – Allocaeculus indicus
Thailand (Fuangarworn & Butcher, 2015) – Neocaeculus orientalis
NORTH AMERICA
Canada (Bernard, Lumley, Buck & Cobb, 2020) - Caeculus cremnicolus, C. cassiopeiae
Mexico (Mulaik & Allred, 1954; Coineau, 1974b; Rivas et al., 2016; Porta & Vázquez Rojas, 2020) – Caeculus veracruzanus, Procaeculus magnus, P. mexicanus, P. orchidicolis, P. potosi
United States (Banks, 1899, 1906; Nevin, 1943, 1945; Mulaik, 1945; Mulaik & Allred, 1954; Higgins & Mulaik, 1957a, 1957b; Enns, 1958; Coineau, 1974b; Hagan, 1985; McDaniel & Boe, 1990; Skvarla, Fisher & Dowling, 2013) – Caeculus americanus, C. archeri, C. calechius, C. clavatus, C. cremnicolus, C. dorotheae, C. janetae, C. gertschi, C. krantzi, C. lewisi, C. mariae, Procaeculus brevis, P. oregonus
Hawaii (Jacot, 1936) – Procaeculus bryani
CENTRAL and SOUTH AMERICA
Argentina (Berlese, 1916; Franz, 1962, 1964; Porta, Pizarro-Araya & Ramírez, 2021) – Andocaeculus beatrizrosso, A. brundini, A. burmeisteri, A. weinrauchi, Neocaeculus bruchi
Brazil (Ott & Ott, 2014, 2018) – Andocaeculus caioi, Neocaeculus setecidades
Chile (Franz, 1964) – Microcaeculus castrii, M. nudus
Dominican Republic (Coineau & Poinar, 2000) – Procaeculus dominicensis (fossil)
Guatemala (Vitzthum, 1933) – Procaeculus willmanni
Puerto Rico (Mulaik, 1945) – Procaeculus puertoricus
Trinidad (Coineau, 1967b) – Procaeculus aitkeni
OCEANIA
Australia (Coineau & Enns, 1970; Otto, 1993; Taylor, Gunawardene & Kinnear, 2013; Taylor, 2014) – Microcaeculus pica, Neocaeculus bornemisszai, N. imperfectus, N. kinnearae, N. knoepffleri, N. nudonates
Tasmania (Coineau, 1974) - Neocaeculus womersleyi
New Zealand (Coineau, 1967a) – Neocaeculus luxtoni
Lord Howe Island (Coineau, 1974) - Neocaeculus johnstoni
Saxidromoidea Coineau, 1974a: 1059
Adamystoidea Cunliffe, 1957
The mite Superfamily Adamystoidea contains only one family – Adamystidae.
Rare or little studied free-living mites, first annouced by Cunliffe (1957) and Coineau (1974a, 1974b, 1976). They are known so far only from 14 countries, but are certainly much more widespread. Ueckermann (1989) revised the family and described six new species, five of them sympatric in the area of Edenville, Orange Free State, South Africa. Another two new genera have been described fom South Africa by Coineau, Theron & Fernandez (2006). Rafalski (1982) described one very high living species in Hindu Kush (Adamystis coineaui, 5100 m). Other researchers described species from Middle Asia (Uzbekistan, Tajikistan, Barilo, 1986), USA (McDaniel & Bolen, 1983), Ukraine (Pogrebnyak, 2007), Cuba (Palacios-Vargas & Prieto, 1995), Mexico (López-Campos, 1996), Iran (Beyzavi, Ueckermann & Ostovan 2012; Fuangarworn & Beyzavi, 2012; Khanjani, Alvandy, Asalifayaz & Ueckermann, 2012), and Thailand (Fuangarworn & Lekprayoon, 2010). So far only 28 species are known in this family, from the genera Adamystis Cunliffe, 1957 (21 sp.), Nannodromus (1 sp.), Saxidromus Coineau, 1974 (4 sp.), Bovidromus Coineau, Theron et Fernandez, 2006 (1 sp.) and Rhinodromus Coineau, Theron et Fernandez, 2006 (1 sp.)
Adamystinae Cunliffe, 1957: 172
Saxidromidae Coineau, 1974a: 1059
Adamystidae: Coineau, 1974b: 431
Type genus: Adamystis Cunliffe, 1957
Adamystinae Cunliffe, 1957: 172
Type genus: Adamystis donnae Cunliffe, 1957
Adamystis Cunliffe, 1957: 172
Type species: Adamystis donnae Cunliffe, 1957
Adamystis alvandicus Khanjani, Alvandy, Asali Fayaz & Ueckermann, 2010: 32
Type locality: Spiny Hawthorn trees, Crataegus aronia (L.) (Rosaceae), highland of Alvand Mountain, Hamedan, Hamedan province (34°39.8’27”N, 48°23’0.35”E).
Distribution: Iran
Adamystis beckyanneae McDaniel & Bolen, 1983: 177; Ueckermann, 1989: 236
Type locality: Olton Town, Lamb County, Texas
Distribution: USA (Texas)
Adamystis buchelis Ueckermann, 1989: 229
Type locality: Edenville, Orange Free State
Distribution: South Africa
Adamystis coineaui Rafalski, 1982: 135; Ueckermann, 1989: 236
Type locality: Valley of Mandaras, Valley of Darum, High Hindu Kush, 5100 m
Distribution: Pakistan
Adamystis donnae Cunliffe, 1957: 172; Ueckermann, 1989: 236
Type locality: Tahoe City, California
Distribution: USA (California)
Adamystis doumengei Coineau, 1974: 1061; Ueckermann, 1989: 239
Type locality: Ermelo Road, Transvaal
Distribution: South Africa
Adamystis ekaterinae Ghasemi-Moghadam & Saboori, 2019: 14
Type locality: Chitgar Forest Park
Distribution: Iran
Adamystis fonsi Coineau, 1974: 1061; Ueckermann, 1989: 239; Pogrebnyak, 2007: 13
Type locality: Banyuls-sur-mer, Eastern Pyrenees
Distribution: France, Ukraine
Adamystis iranoturanianensis Beyzavi, Ueckermann & Ostovan, 2012: 28
Type locality: from soil and humus under oak trees (Quercus sp., Fagaceae), Kamfiruze, Fars province (30°7.15’55”N, 52°15’56.17”E).
Distribution: Iran
Adamystis monomaculata Ueckermann, 1989: 232
Type locality: Edenville, Orange Free State; Potchefstroom, Transvaal
Distribution: South Africa
Adamystis nudis Ueckermann, 1989: 231
Type locality: Edenville, Orange Free State
Distribution: South Africa
Adamystis oculeusis Ueckermann, 1989: 226
Type locality: Edenville, Orange Free State
Distribution: South Africa
Adamystis persiangulfensis Paktinat-Saeij & Kazemi, 2021: 260
Type locality: Qeshm Island
Distribution: Iran
Adamystis saboorii Paktinat-Saeij & Ahaniazad, 2019: 133
Type locality: Yarshahr village, Maragheh city, East Azerbaijan Province
Distribution: Iran
Adamystis sarae Hunter & Crossley, 1968: 181; Ueckermann, 1989: 229
Type locality: Savannah River Plant Reservation near Aiken, South Carolina
Distribution: USA
Adamystis sartorum Barilo, 1986: 1414
Type locality: Samarkand, Buhara (Uzbekistan)
Distribution: Tadjikistan, Uzbekistan
Adamystis thailandensis Fuangarworn & Lekprayoon, 2010: 61
Type locality: Pang-Nga Province, Thaimuang District, Hard-Thaimuang Khao-Champee National Park, 8°29’38”N, 98°13’39”E
Distribution: Thailand
Adamystis theroni Fuangarworn & Beyzavi, 2012: 449
Type locality: Zagros mountain, south of Kamfiruz region, Fars province (30°8’N, 52°17’E).
Distribution: Iran
Adamystis trimaculata Ueckermann, 1989: 239
Type locality: Edenville, Orange Free State
Distribution: South Africa
Adamystis ueckermanni Fuangarworn & Beyzavi, 2012: 452
Type locality: Zagros mountain, south of Kamfiruz region, Fars province (30°8’N, 52°17’E).
Distribution: Iran
Adamystis yvesi Ueckermann, 1989: 236
Type locality: Pongola, Transvaal
Distribution: South Africa
Saxidrominae Coineau, 1974a: 1059
Saxidromidae: Coineau, 1974a: 1059; 1974b: 431
Type genus: Saxidromus Coineau, 1974
Bovidromus Coineau, Theron & Fernandez, 2006: 67
Type species: Bovidromus roussouwi Coineau, Theron et Fernandez, 2006
Bovidromus roussouwi Coineau, Theron et Fernandez, 2006: 67
Type locality: South Africa
Distribution: South Africa
Nannodromus Fernandez, Coineau, Theron & Tiedt, 2014: 19
Type secies: Nannodromus reveilleti Fernandez, Coineau, Theron et Tiedt, 2014
Nannodromus reveilleti Fernandez, Coineau, Theron & Tiedt, 2014: 19
Type locality: Kruger National Park, N’Wanetsi: S24°27’30.56”, E31°58’35.30”
Distribution: South Africa
Rhinodromus Coineau, Theron & Fernandez, 2006: 73
Type species: Rhinodromus lootsi Coineau, Theron et Fernandez, 2006
Rhinodromus lootsi Coineau, Theron & Fernandez, 2006: 73
Type locality: South Africa
Distribution: South Africa
Saxidromus Coineau, 1974: 1060
Type species: Saxidromus delamarei Coineau, 1974
Saxidromus annahoffmannae Lopes-Campos, 1996: 215
Type locality: Veracruz
Distribution: Mexico
Saxidromus caribeus Palacios-Vargas & Prieto, 1995: 213
Type locality: Santiago de las Vegas, Habana region
Distribution: Cuba
Saxidromus delamarei Coineau, 1974: 1060; Coineau & Naudo, 1986: 303; Ueckermann, 1989: 227
Type locality: Banyuls-sur-Mer, Eastern Pyrenees
Distribution: France
Saxidromus knoepffleri Coineau, 1974: 1061; Ueckermann, 1989: 227
Type locality: Ermelo Road, Melspruit, Transvaal
Distribution: South Africa
EUROPE
France (Coinneau, 1974) – Adamystis fonsi, Saxidromus delamarei
Ukraine (Pogrebnyak, 2007) - Adamystis fonsi
AFRICA
South Africa (Coineau, 1974; Ueckermann, 1989; Coineau, Theron & Fernandez, 2006; Fernandez, Coineau, Theron & Tiedt, 2014) – Adamystis buchelis, A. doumengei, A. monomaculata, A. nudis, A. oculensis, A. trimaculata, A. yvesi, Bovidromus roussouwi, Nannodromus reveilleti, Rhinodromus lootsi, Saxidromus knoepffleri
ASIA
Iran (Beyzavi, Ueckermann & Ostovan 2012; Fuangarworn, Beyzavi & Ostovan, 2012; Khanjani, Alvandy, Asalifayaz & Ueckermann, 2012; Paktinat-Saeij et al., 2019; Paktinat-Saeij & Kazemi, 2021) – Adamystis alvandicus, A. ekaterinae, A. iranoturanianensis, A. persiangulfensis, A. saboorii, A. theroni, A. ueckermanni
Japan (Shiba, 1999) – Adamystidae gen. sp.
Pakistan (Rafalski, 1982) – Adamystis coineaui
Tadjikistan (Barilo, 1986) – Adamystis sartorum
Thailand (Fuangarworn & Lekprayoon, 2010) - Adamystis thailandensis
Uzbekistan (Barilo, 1986) - Adamystis sartorum
NORTH AMERICA
Mexico (López-Campos, 1996) – Saxidromus annahoffmannae
United States (Cunliffe, 1957; Hunter & Crossley, 1968; McDaniel & Bolen, 1983) – Adamystis beckyanneae, A. donnae, A. sarae
Hawaii (Swift, 2000) – Adamystidae gen. sp.
CENTRAL and SOUTH AMERICA
Cuba (Palacios-Vargas & Prieto, 1995) – Saxidromus caribeus
French Antilles (Saint-Martin)(Coineau, 1979) – Adamystidae gen. sp.
OCEANIA
Australia (Walter & O’Dowd, 1995) – Adamystidae gen. sp.
The subfamily contains only one family – Pomerantziidae.
Small family (six known species), established by Baker (1949) for one species, found In Georgia (USA). Price (1971, 1974, 1975) added new species from the USA.
Quing-Hai Fan & Yan Chen (2005) revised the family and separated two of the species of Price in a new genus Apomerantzia. One new species (Pomerantzia subterranea) was found in deep humus in Fuzhou, China, another one (Pomerantzia philippina Bochkov et Walter, 2007) – in the Philippines.
Pomerantziidae Baker, 1949: 269
Type genus: Pomerantzia Baker, 1949
Apomerantzia Quing-Hai Fan & Yan Chen, 2005: 8
Type species: Pomerantzia prolata Price, 1971
Pomerantzia kethleyi Price, 1975: 487; Kethley, 1989: 101
Apomerantzia kethleyi (Price): Quing-Hai Fan & Yan Chen, 2005: 7
Type locality: Indiana, Lake Co., St. John
Distribution: USA (Indiana, Illinois, Minnesota)
Pomerantzia prolata Price, 1971: 395
Apomerantzia prolata (Price): Quing-Hai Fan & Yan Chen, 2005: 5
Type locality: California, Nevada Co.
Distribution: USA (California)
Pomerantzia Baker, 1949: 269
Type species: Pomerantzia charlesi Baker, 1949
Pomerantzia benhami Price, 1974: 425
Type locality: California, Fresno Co.
Distribution: USA (California)
Pomerantzia charlesi Baker, 1949: 269; Price, 1974: 420
Type locality: Upson County, Georgia
Distribution: USA (Georgia)
Pomerantzia philippina Bochkov & Walter, 2007: 159
Type locality: Romblon Prov., Sibuyan Isl, Magdiwang, beach near of Puto
Point, 12°26’N, 122°41’E, soil
Distribution: the Philippines
Pomerantzia subterranea Quing-Hai Fan & Yan Chen, 2005: 8
Type locality: Fuzhian
Distribution: China
ASIA
China (Quing-Hai Fan & Yan Chen, 2005) – Pomerantzia subterranea
Philippines (Bochkov & Walter, 2007) – Pomerantzia philippina
NORTH AMERICA
United States (Baker, 1949; Price, 1971, 1974, 1975; Price & Benham, 1976) – Apomerantzia kethleyi, A. prolata, Pomerantzia benhami, P. charlesi
In the period of 14 years (2008 - 2021) of publication of my 9 volumes of the series Acarorum Catalogus, the fast development of the Acarology brought to light many new taxa of Acari and many other mites have been re-arranged or synonymized. This task requires a supplement to the published volumes (one, concerning Vol.I, has been already published). These supplements will be sertainly followed by many other changes, but it would be proper to fix by 2021 the situation within the 97 families of mites considered in the series.
First update of Volume I:
Beron P. 2017. Acarorum Catalogus I Acariformes: Calyptostomatoidea (Calyptostomatidae), Erythraeoidea (Smarididae, Erythraeidae). Pensoft & Nat. Mus. Natur. Hist. Sofia: supplement 1. - Historia naturalis bulgarica, 24: 131 – 154.
Beron P. 2008. Acarorum Catalogus I Acariformes: Calyptostomatoidea (Calyptostomatidae), Erythraeoidea (Smarididae, Erythraeidae). Pensoft & Nat. Mus. Natur. Hist. Sofia: 271 pp.
Many specialists from Poland, Montenegro, Iran, Turkey, China, Brazil, and other countries added new information to the knowledge on these groups. As a result of the recent contributions, the number of genera and species in Calyptostomatoidea and Erythraeoidea (some of them of uncertain status) is updated as follows:
Calyptostomatoidea
Calyptostomatidae – one genus, ten species
Erythraeoidea
Smarididae – 10 genera, 59 species
Erythraeidae – 57 genera, ca. 890 species
Fam. Calyptostomatidae
Calyptostoma seemani Masoumi et Saboori
Calyptostoma seemani Masoumi & Saboori, 2020: 1823
Type locality: Kandovan region, Alborz province
Distribution: Iran
Unlike most terrestrial parasitengone mites (chiggers, erythraeids), known mostly from parasitic and easy-to-collect larvae, smaridid systematics are mainly based on the post larval instar (Dos Santos, Welbourn, Klimov & Pepato, 2021).
Clavismaris maquine Ott & Ott, 2018: 2
Type locality: Atlantic forest area at FEPAGRO Maquiné, Rio Grande do Sul
Distribution: Brazil
Sphaerotarsus sadafae Amin, Khanjani & Nadri, 2020: 1965
Type locality:
Distribution: Iran
Southcottiana Dos Santos, Welbourn, Klimov & Pepato, 2021: 104
Type species: Sphaerotarsus monticolus Southcott, 1997
Sphaerotarsus monticolus Southcott, 1997: 6
Southcottiana monticola (Southcott, 1997): Dos Santos, Welbourn, Klimov & Pepato, 2021: 104
Type locality: Mt. Lofty, S. Australia
Disribution: Australia (S. Australia)
Trichosmaris calcarensis Dos Santos, Welbourn, Klimov & Pepato, 2021: 108
Type locality: cave, 5 km E Pain Municipality, Minas Gerais
Distribution: Brazil
Trichosmaris paulensis Dos Santos, Welbourn, Klimov & Pepato, 2021: 112
Type locality: Americana Municipality, São Paulo State
Distrinution: Brazil
Collemboerythraeus izadpanahi Noei, Sabbori, Sedghi & Hakimitabar, 2017: 2
Type locality: Simkan region, Jahrom city, Fars province
Distribution: Iran
Host: Collembola: Sminthuridae
Brachycaudus sp., Hyperomyzus lactucae, Macrosiphoniella sanborni (Homoptera: Aphidae) by Oner et al. (2021).
Erythraeus (E.) hubeiensis Xu, Yi, Guo & Jin, 2019: 55
Type locality: Qizimeishan National Natural Reserve, Hubei Province
Distribution: China
Host: Typhlocybini (Cicadellidae: Typhlocybinae)
Erythraeus (Parerythraeus) thomasi Roland & Gabryś, 2020: 326
Type locality: Biogeographic province: Varsinais-Suomi, loc.:670:25 V, Kaarina Kuusiston Kartano, Kuiva keto
Distribution: Finland
Note: the subgenus Parerythraeus is reinstalled as subgenus, out of synonymy with Erythraeus, by Roland & Gabryś (2020).
E. (Zaracarus) hainanensis Xu, Yi, Guo & Jin, 2019: 64
Type locality: Bawangling National Natural Reserve, Hainan Province
Distribution: China (Hainan)
Host: planthoppers (Hemiptera: Delphacidae)
Lasioerythraeus jessicae Dos Santos Costa 2019: 184
Type locality: 19°05’46.0”S, 43°21’25.0”W, in an iron ore terrain cave, Conceição do Mato Dentro municipality, Minas Gerais State
Distribution: Brazil
Leptus (L.) andongensis Hakimitabar, Joharchi & Jung, 2020: 1
Type locality: Andong
Distribution: S. Korea
Host: Meimuna sp. (Insecta: Homoptera: Cicadidae)
Leptus (Leptus) brasilicus Haitlinger, Šundiċ & Pompermaier. 2017: 874
Type locality: 30 km northeast of Brasilia
Distribution: Brazil
Leptus (L.) cameroonicus Haitlinger, Šundiċ & Nkwala, 2020: 608
Type locality: forest reserve of the town Mbalmayo (3º28’34.218” N; 11º29’37.416E)
Distribution: Cameroon
Host: Eyprepocinemis plorans ibandana (Orthoptera: Acrididae)
Leptus candangus Šundiċ, Haitlinger & Pompermaier, 2017: 90
Type locality: Serrado-Pianaltinade (pitfall traps)
Distribution: Brazil
Leptus (Leptus) chingombensis Bernard, Felska & Makol, 2019: 794
Type locality: Mikwa stream, Michinga Escarpment, 1160 m
Distribution: Zambia
Hosts: Heliaeschna fuliginosa (Odonata: Aeschnidae), Orthetrum julia (Libellulidae), Paragomphus cognatus (Gomphidae), Pseudagrion spernatum (Coenagrionidae)
Leptus (Leptus) darvishi Saboori, Hakimitabar & Khademi, 2018: 95
Type locality: Komaijan and Mahallat cities, Markazi Province
Distribution: Iran
Host: undetermined grasshoppers (Orthoptera: Acrididae) and undetermined beetles (Coleoptera: Buprestidae and Tenebrionidae)
Leptus (L.) flechtmanni Bassini-Silva, Jacinavicius & Barros-Battesti, 2020: 2
Type locality: Serra dos Carajás,Parauapebas, Pará, Brazil, (6° 6’29’’S, 50° 18’16’’W)
Distribution: Brazil
Host: Unidentified opilion (Opiliones: Cosmetidae)
Leptus (Leptus) grancanaricus Haitlinger & Šundiċ, 2020: 836
Type locality: Gran Canaria, Tejada: Mirador de Timagada, altitude 1,258 m
Distribution: Canary Islands
Host: Herpisticus guanarteme (Coleoptera: Curculionidae)
Leptus (L.) guarani Haitlinger, Šundiċ, Ázara & Bernardi, 2020: 1923
Type locality: State Parana, Morretes
Distribution: Brazil
Hosts: Acanthogonyleptes editus (Opiliones: Gonyleptidae)
Leptus (L.) haitlingeri Jacinavicius, Bassini-Silva & Barros-Battesti, 2018: 975; Bassini-Silva et al., 2020: 8
Type locality: Serra da Cantareira, Săo Paulo municipality, State of Săo Paulo
Distribution: Brazil
Hosts: Fidena (Fidena) sp. (Diptera: Tabanidae); Panstrongylus megistus (Hemiptera: Reduviidae), Dysdercus sp. (Hemiptera: Pyrrhocoridae), Obiricodesmus rosascens (Diplopoda, Polydesmida: Chelodesmidae)
Leptus (Leptus) machadoi Haitlinger & Šundiċ, 2020: 839
Type locality: Fuerteventura, Jandia, Pico de la Zarza, 807 m
Distribution: Canary Islands
Host: Laparocerus maxorata (Coleoptera: Curculionidae)
Leptus planaltensis Haitlinger, Šundiċ & Pompermaier, 2017: 879
Type locality: Brazil
Distribution: Brazil
Leptus pouryayevalii Hakimitabar, Saboori & Fadaei, 2021: 138
Type locality: Koushk Bala Village, Alborz Prov., Chalous road, Karaj, 1824 m
Distribution: Iran
Host: shaking the leaves of weeds
Leptus (L.) sidorchukae dos Santos Costa, 2019: 159
Type locality: 19°34’36.0”S, 44°01’13.0”W, in an unnamed limestone cave, Pedro Leopoldo municipality, Minas Gerais State
Distribution: Brazil
Leptus sp. on Chrysops brunneus and Hybomitra lasiophthalma (Diptera: Tabanidae, Canada)(Mullen et al., 1989)
Abrolophus bochkovi Hakimitabar, Saboori & Fadaei, 2020: 108
Type locality: Qazvin province
Distribution: Iran
Abrolophus diaoluoensis Xu & Jin, 2021: 24
Type locality: Hainan, Diaoluoshan National Natural Reserve
Distribution: China (Hainan)
Abrolophus hajikanbar Noei, 2022: 226
Type locality: Chahardeh and Chahkand villages, Birjand, South Khorasan province
Distribution: Iran
Abrolophus karamani Haitlinger & Šundiċ, 2019: 108; the species is considered as a junior synonym of A. aitapensis (Southcott 1948)(Hakimitabar, Saboori & Fadaei, 2020b).
Type locality: Port Louis
Distribution: Guadeloupe
Abrolophus marianopolicus Haitlinger & Šundiċ, 2018b: 354.
Type locality: 5 km north of Marianopoli (prov. Caltainisetta)
Distribution: Sicily
Abrolophus poljankensis Haitlinger & Šundiċ, 2021: 39
Type locality: Nižna Poljanka n. Bardejov (Slovakia)
Distribution: Poland, Slovakia
Abrolophus zelandicus Luxton, 1989: 416; 1990: 398
Type locality: Auckland, Nelson (on a sea shore)
Distribution: New Zealand
Charletonia elbasani Šundić, Haitlinger & Milosević, 2017: 564
Type locality: Elbasan
Distribution: Albania
Host: on grasses
Caeculisoma allopenlineata Xu & Jin, 2020: 286
Type locality: Diaoluoshan National Natural Reserve, Hainan Province
Distribution: China
Hosts: Hemiptera: Cicadellidae, Delphacidae
Caeculisoma penlineatus Xu & Jin, 2019c: 511; Xu et al., 2020: 295
Type locality: Chongqing Municipality
Distribution: China (Chongqing Municipality, Guizhou Prov., Hainan Prov.)
Hosts: Mileewa margheritae (Hemiptera: Cicadellidae), Gen. sp. (Hem.: Delphacidae)
Caeculisoma semispinus Xu & Jin, 2019d: 561
Type locality: Qizimeishan National Reserve, Hubei Province
Distribution: China
Host: Shaddai sp. (Hemiptera: Cicadellidae)
Dambullaeus adonis Makol & Moniuszko, 2014: 5
Type locality: Western Madagascar
Distribution: Madagascar
Host: Latois nigrolineata (Hemiptera: Fulgoromorpha: Flatidae)
Dambullaeus hainanensis Xu & Jin, 2019: 456
Type locality: Hainan
Distribution: China
Dambullaeus jianfengensis Xu & Jin, 2020: 297
Type locality: Jianfengling National Natural Reserve, Hainan Province
Distribution: China (Hainan)
Iguatonia seemani Xu & Jin, 2020: 306
Type locality: Jianfengling National Natural Reserve, Hainan Province
Distribution: China (Hainan)
Iguatonia xinfengi Xu & Jin, 2020: 315
Type locality: Jianfengling National Reserve, Hainan Province
Distribution: China (Hainan)
Balaustium akramii Noei, 2017: 2218; Oner et al., 2021: 2
Type locality: Sari city, Mazandaran province, Iran
Distribution: Iran, Turkey
Host: Uroleucon sanchi (Homoptera: Aphidae)
Balaustium izmirensis Noei & Ersin, 2019: 31
Type locality: Bornova, Izmir
Distribution: Turkey
Balaustium ryszardi Šundiċ & Noei, 2021: 2609
Type locality: Peloponese
Distribution: Greece
EUROPE
Albania (Šundiċ, Haitlinger & Milošević, 2017; Haitlinger & Šundiċ, 2018a) - Abrolophus quisquiliarus, Charletonia elbasani, Erythraeus (Zaracarus) lancifer, E. (E.) moeritzensis, E. (E.) southcotti, Marantelophus iranicus, Charletonia kalithensis
Finland (Roland & Gabryś, 2020) – Erythraeus (Parerythraeus) thomasi
Greece (Haitlinger & Šundiċ, 2018a; Šundiċ & Noei, 2021) – Balaustium ryszardi, Abrolophus anzelmi, A. petanovicae, A. wratislaviensis, Erythraeus (E.) regalis
Hungary (Haitlinger & Šundiċ, 2018) - Charletonia cardinalis, Moldoustium baltiensis
Italy (Haitlinger & Šundiċ, 2018a) - Abrolophus wratislaviensis, A. anzelmi, A. dagmarae
Sicily (Haitlinger & Šundiċ, 2018b) - Abrolophus marianopolicus, A. wratislaviensis
Montenegro (Haitlinger & Šundiċ, 2018a) - Erythraeus (E.) cinereus, E. (E.) regalis, Charletonia kalithensis
Poland (Haitlinger & Šundiċ, 2021) – Abrolophus poljankus
Slovakia (Haitlinger & Šundiċ, 2021) - Abrolophus poljankus, Leptus molochinus
Turkey (Gencsoylu, 2007; Noei et al., 2017; Noei, Ersin & Çakmak, 2019; Oner et al., 2021) – Balaustium akrami, B. izmirensis, E.(E.) phalangoides, E.(E.) uhadi, E. (Zaracarus) coleopterus, E.(Z.) iranicus, E. (Z.) kurdistaniensis. Charletonia krendowskyi
AFRICA
Canary Islands (Haitlinger, 2019, Haitlinger & Šundiċ, 2020) - Leptus (Leptus) grancanaricus, L.(L.) machadoi, Abrolophus quisquiliarus
Cameroon (Haitlinger, Šundiċ & Nkwala, 2020) – Leptus cameroonicus
Madagascar (Makol et al., 2014) – Dambullaeus adonis
Zambia (Bernard, Felska & Makol, 2019) – Leptus (L.) chingombensis
ASIA
China (Xu, Yi, Guo & Jin, 2019a, b, c, d, 2020, 2021) – Abrolophus diaoluoensis, Caeculisoma allopenlineatus, C. penlineatus, C. semispinus, Dambullaeus jianfengensis, Erythraeus (E.) hubeiensis
Hainan (Xu, Yi, Guo & Jin, 2020) - Caeculissoma penlineatus, Erythraeus (Z.) hainanensis, Dambullaeus hainanensis, Iguatonia seemani, I. xinfengi
India (Gupta et al., 2004; Gupta A. et al., 2006) - Erythraeidae
Iran (Noei et al., 2017; Noei, Maroufpoor, Faizi & Ostovan, 2017; Noei & Asadollahzadeh et al., 2017; Saboori et al., 2018; Hakimitabar, Saboori & Saaveh, 2018; Noei, Saboori, Sedghi & Hakimitabar, 2018; Noei & Rabieh, 2019; Noei & Honarmand. 2020; Hakimitabar, Saboori & Fadaei, 2020, 2021; Masoumi, Saboori & Seiedy, 2020; Amin, Khanjani & Nadri, 2020; Saboori & Shirvani, 2021, Noei, 2022) – Calyptostoma seemani; Smarididae: Sphaerotarsus sadafae; Erythraeidae: Abrolophus bochkovi, Balaustium akrami, Collemboerythraeus izadpanahi, Leptus darwishi, L. pouryayevalii
South Korea (Hakimitabar, Johardii & Jong, 2020) – Leptus andongensis
NORTH AMERICA
Canada (Mullen et al. 1989) – Leptus sp.
Mexico (Montiel-Parra & Pérez-Ortiz, 2007; Chaires-Grijalva et al., 2021) – Leptus
indianensis, L. lomani, L. onnae, L. ruginus
SOUTH and CENTRAL AMERICA
Argentina (Cédola, Nolasco & Minardi, 2018) – Balaustium sp.
Bolivia (Torrico-Bazoberry et al., 2020) – Leptus hringuri
Brazil (Pinto et al., 2014; Šundić et al., 2017; Ott & Ott, 2018; Jacinavicius, 2019; Haitlinger, Šundiċ & Pompermaier, 2017; Šundiċ, Haitlinger & Pompermaier, 2017; Guimarraes et al., 2019; Bassini-Silva et al., 2019, 2020; Haitlinger et al., 2020; Dos Santos et al., 2021) – Smarididae: Clavismaris maquine, Trichosmaris calcarensis, T. paulensis; Erythraeidae: Leptus brasilicus, L. candangus, L. guarani, L. flechtmanni, L. haitlingeri, L. planaltensis, L. simonettae,
Costa Rica (Dos Santos et al., 2021) – Trichosmaris longirostris
OCEANIA
Australia (Dos Santos et al., 2021) - Southcottiana monticola
New Zealand (Luxton, 1989, 1990; Xu & Zhang, 2020) – Abrolophus zelandicus, Balaustium medicagoense
Arachnida
Opiliones
Fam. Cosmetidae
Acromares vittatum - Leptus ca. indianensis
Cynorta sp. - Leptus ruginus
Holovonones compressus - Leptus ruginus
Heterovonones incrassatus – Leptus indianensis
Vonones pelaezi - Leptus ruginus
Unidentified opilion – Leptus flechtmanni
Fam. Gonyleptidae
Acanthogonyleptes editus – Leptus (Leptus) guarani
Caldassius nigripes – Leptus sp.
Gonyleptes gonyleptoides - Leptus sp.
Moreiranula mamillata - Leptus sp.
Fam. Stygnopsidae
Paramitraceras granulatus - Leptus ruginus
Fam. Sclerosomatidae
Metopilio sp. - Leptus ruginus
Leiobunum nycticorpum - Leptus ruginus
Paranelima sp. - Leptus ruginus
Prionostemma sp. - Leptus ruginus, L. nr. indianensis
Krusa sp. - Leptus ruginus
Scorpiones
Fam. Buthidae
Centruroides limpidus limpidus – Leptus ruginus
Fam. Caraboctonidae
Hadrurus gerstchi - Leptus ruginus
Fam. Vejovidae
Vejovis mexicanus – Leptus sp.
Vejovis spinigerus – Leptus sp.
Vejovis variegatus – Leptus ruginus
Amblypygi
Fam. Phrynidae
Phrynus garridoi – Leptus onnae
Paraphrynus pococki – Leptus onnae
Araneae
Fam. Filistatidae
Kukulcania sp. - Leptus ca. indianensis
Ixodida
Fam. Argasidae
Ornithodoros sp. - Leptus (Leptus) simonettae
Myriapoda
Diplopoda - Polydesmida
Fam. Chelodesmidae
Obiricodesmus rosascens - Leptus haitlingerii
Insecta s.l.
Collembola
Fam. Sminthuridae - Collemboerythraeus izadpanahi
Odonata
Fam. Aeschnidae
Heliaeschna fuliginosa - Leptus (Leptus) chingombensis
Fam. Gomphidae
Paragomphus cognatus - Leptus (Leptus) chingombensis
Fam. Libellulidae
Orthetrum julia - Leptus (Leptus) chingombensis
Fam. Coenagrionidae
Pseudagrion spernatum - Leptus (Leptus) chingombensis
Orthoptera
Fam. Acrididae
Eyprepocinemis plorans ibandana – Charletonia braunsi, C. cameroonensis, C. justinae, Leptus cameroonicus
Zonocerus variegatus – Charletonia cameroonensis
Hemiptera Heteroptera
Fam. Reduviidae
Panstrongylus megistus – Leptus haitlingerii
Fam. Pyrrhocoridae
Dysdercus sp. - Leptus haitlingerii
Hemiptera
Fam. Cicadellidae
Mileewa margheritae – Caeculisoma penlineatus
Shaddai sp. – Caeculisoma semispinus
Fam. Delphacidae
Gen. sp. - Caeculisoma penlineatus
Fam. Cicadidae
Meimuna sp. - Leptus (L.) andongensis
Fam. Membracidae
Alchisme grossa - Leptus hringuri
Fam. Flatidae
Latois nigrolineata - Dambullaeus adonis
Homoptera
Fam. Aphididae
Uroleucon (U.) sonchi - Balaustium akrami, Erythraeus ankaraicus, E. uhadi, E. (Z.) coleopterus, E.(Z.) iranicus, E.(Z.) kurdistaniensis
Uroleucon (U.) aeneum - Erythraeus ankaraicus
Uroleucon sp. – Curteria duzgunesae
Aphis craccivora - Erythraeus ankaraicus
A. nerii - Erythraeus ankaraicus, E.(Z.) iranicus
Aulacorthum solani - Erythraeus ankaraicus
Brachycaudus tragopogonis – Erythraeus (Z.) kurdistaniensis
Brachycaudus sp. - Erythraeus ankaraicus
Hyperomyzus lactucae - Erythraeus ankaraicus, Erythraeus (Z.) kurdistaniensis
Hyalopterus pruni - Erythraeus ankaraicus
Macrosiphoniella sanborni – Erythraeus ankaraicus
Myzus sp. – Erythraeus phalangoides
Sitobion sp. - Erythraeus phalangoides
Coleoptera
Fam. Curculionidae
Herpisticus guanarteme – Leptus grancanaricus
Laparocerus maxorata - Leptus machadoi
Fam. Elateridae
Monocrepidius flavangulus - Leptus sp.
Glyphonyx flohri – Leptus sp.
Diptera
Fam. Tabanidae
Fidena (Fidena) sp. - Leptus haitlingeri
Chrysops brunneus – Leptus sp.
Hybomitra lasiophthalma – Leptus sp.
Fam. Drosophilidae
Drosophila subobscura – Leptus mariae
(suppl. 2017 – 2021 and ommited)
Beron P. 2011. Acarorum Catalogus. II. Acariformes Acaridida Listrophoroidea (Listrophoridae, Dromiciocoptidae, Myocoptidae, Chirodiscidae, Atopomelidae). Pensoft & Nat. Mus. Natur. Hist. Sofia: 263 pp.
As a result of the recent contributions (mostly by A. Bochkov and co-authors), the number of genera and species in Listrophoroidea (some of them uncertain status) is updated as follows:
Listrophoridae – 24 genera, 177 species
Dromiciocoptidae – 1 genus, 3 species
Myocoptidae – 5 genera, 86 species
Chirodiscidae - 26 genera, 241 species
Atopomelidae - 44 genera, 413 species
Total 100 genera, 925 sp.
Afrolistrophorus (Afrolistrophorus) laonastes Bochkov, Abramov & Hugot, 2011: 11
Type locality: Kammouane Province, 18 km north of Thakhek, near Ban Doy village, 17º33’26’’N, 104º49’28’’E, 140 m a. s. l.
Distribution: Laos
Host: Laonastes aenigmamus (Rodentia: Diatomyidae)
Afrolistrophorus (Afrolistrophorus) hapalomys Bochkov & Abramov, 2014: 139
Type locality: Binh Phuoc Province, Bu Gia Map National Park, 13
km NE Bu Gia Map Village, 540 m a.l.s., 12°11’37”N, 107°12’21”E
Distribution: Vietnam
Host: Hapalomys delacouri Thomas (Rodentia, Muridae)
Afrolistrophorus (Afrolistrophorus) saxatilomys Bochkov, Abramov & Hugot, 2011: 14
Type locality: Kammouane Province
Distribution: Laos
Host: Saxatilomys paulinae (Rodentia: Muridae)
Amlistrophorus geoxus Sikora & Bochkov, 2012: 390
Type locality: Los Lagos, Lago de Todoslos Santos, 41°06’S 72°12’W
Distribution: Chile
Host: Geoxus valdivianus (Rodentia: Cricetidae, Sigmodontinae)
Dubininetta abramovi Bochkov, 2017: 440
Type locality: 5 km east of Na Hang, Tuyen Quang Province, Vietnam; 22°20’59”N, 105°25’36”E
Distribution: Vietnam
Host: Euroscaptor subanura (Soricomorpha: Talpidae)
Dubininetta euroscaptor Bochkov, 2017: 438
Type locality: 14 km north of Kon Plong Town, Kon Plong District, Kon Tum Province, Vietnam; 14°43’20”N, 108°18’58”E
Distribution: Vietnam
Host: Euroscaptor parvidens (Soricomorpha: Talpidae)
Lynxacarus (Lynxacarus) serrafreirei Bochkov & Valim, 2016: 30
Type locality: Rio de Janeiro
Distribution: Brazil
Host: Galictis cuja (Carnivora: Mustelidae)
Prolistrophorus (Aprolistrophorus) parabidentatus Bochkov, Lareschi & Barreto, 2014: 590
Type locality: Argentina
Distribution: Argentina
Host: Akodon azarae (Rodentia: Cricetidae, Sigmodontinae)
Prolistrophorus (Aprolistrophorus) tylomys Bochkov, Lareschi & Barreto, 2014: 592
Type locality: Guatemala
Distribution: Guatemala
Host: Tylomys nudicaudus (Rodentia: Cricetidae)
Prolistrophorus hylaeamys Bochkov & Valim, 2016: 27
Type locality: Minas Gerais
Distribution: Brazil
Host: Hylaeamys laticeps (Rodentia: Cricetidae, Sigmodontinae)
Pteromychirus menetes Bochkov, Abramov & Hugot, 2011: 17
Type locality: Kammouane Province, 18 km north of Thakhek, near Ban Doy village, 17º33’26’’N, 104º49’28’’E, 140 m a. s. l.
Distribution: Laos
Host: Menetes berdmorei (Rodentia: Sciuridae)
Apocalypsis Bochkov, 2010: 148 (praeoccupied name, not Butler, 1876, Lepidoptera).
Apocalyptoides Bochkov, 2016b: (replacement name)
Type species Criniscansor [not Gliricoptes !] allactaga Fain et Lukoschus, 1979
Criniscansor [not Gliricoptes] allactaga Fain et Lukoschus, 1979: 236
Apocalypsis allactaga (Fain et Lukoschus): Bochkov, 2010: 148
Apocalyptoides allactaga (Fain et Lukoschus): Bochkov, 2016b:
Type locality: Char-nur
Distribution: Mongolia
Host: Allactaga sibirica (Rodentia: Dipodidae)
Myocoptes (M.) lophuromys Bochkov, OConnor & Skoracki, 2016: 588
Type locality: Uganda
Distribution: Uganda
Host: Lophuromys voosnami (Rodentia: Muridae)
Myocoptes (M.) meriones Bochkov, 2016b: 477
Type locality: Kirghizia
Distribution: Kirghizia, Tajikistan
Hosts: Meriones tamariscinus, M. libycus (Rodentia: Muridae)
Myocoptes (Comyoptes) nesokia Bochkov & OConnor, 2017: 47
Type locality: Pakistan
Distribution: Pakistan
Host: Nesokia indica (Rodentia: Muridae)
Trichoecius dubininae Bochkov, 2016b: 462
Type locality: Chelyabinsk Province, Russia
Distribution: Kazakhstan, Kirghizia, Russia
Host: Lasiopodomys gregalis (Rodentia: Arvicolidae)
Trichoecius faini Bochkov, OConnor & Skoracki, 2016: 595
Type locality: Tanzania
Distribution: Tanzania
Host: Hylomyscus arcimontanus (Rodentia: Muridae)
Trichoecius hylomyscus Bochkov, OConnor & Skoracki, 2016: 593
Type locality: Tanzania
Distribution: Gabon, Tanzania
Hosts: Hylomyscus anselli, H. stella (Rodentia: Muridae)
Trichoecius lemmus Bochkov, 2016b: 466
Type locality: Taymyr peninsula
Distribution: Russia
Host: Lemmus sibiricus (Rodetia: Arvicolidae)
Trichoecius meriones Bochkov, 2016b: 468
Type locality: Turkmenia
Distribution: Turkmenia
Host: Meriones meridianus (Rodentia: Muridae)
Schizocarpus azasicus Bochkov & Saveljev, 2012: 12
Type locality: Tuva, Todzhinskii District
Distribution: Russia (Tuva)
Host: Castor fiber tuvinicus (Rodentia: Castoridae)
Schizocarpus faini Bochkov, Labrzycka, Skoracki & Saveljev, 2012: 46
Type locality: Suwalki
Distribution: Poland
Host: Castor fiber belorussicus (Rodentia: Castoridae)
Schizocarpus gozdziewskii Bochkov, Labrzycka, Skoracki & Saveljev, 2012: 51; Bochkov & Saveljev, 2012: 16
Type locality: Suwalki
Distribution: Poland, Russia (Tuva)
Hosts: Castor fiber belorussicus, C.f. orientoeuropaeus, C.f. tuvinicus (Rodentia: Castoridae)
Schizocarpus heatherae Bochkov, Labrzycka, Skoracki & Saveljev, 2012: 49
Type locality: Suwalki
Distribution: Poland
Host: Castor fiber belorussicus (Rodentia: Castoridae)
Schizocarpus heideckei Bochkov & Saveljev, 2012: 9
Type locality: Tuva, Todzhinskii District
Distribution: Russia (Tuva)
Host: Castor fiber tuvinicus (Rodentia: Castoridae)
Schizocarpus klompeni Bochkov, Labrzycka, Skoracki & Saveljev, 2012: 50
Type locality: Suwalki
Distribution: Poland
Host: Castor fiber belorussicus (Rodentia: Castoridae)
Schizocarpus lavrovi Bochkov & Saveljev, 2012: 7
Type locality: Tuva, Todzhinskii District
Distribution: Russia (Tuva)
Host: Castor fiber tuvinicus (Rodentia: Castoridae)
Schizocarpus parahumilis Bochkov, Labrzycka, Skoracki & Saveljev, 2012: 53
Type locality: Suwalki
Distribution: Poland
Host: Castor fiber belorussicus (Rodentia: Castoridae)
Schizocarpus pseudonumerosus Bochkov, Labrzycka, Skoracki & Saveljev, 2012: 47
Type locality: Suwalki
Distribution: Poland
Host: Castor fiber belorussicus (Rodentia: Castoridae)
Schizocarpus saveljevi Bochkov, 2012: 168
Type locality: Leningrad Province
Distribution: Russia
Host: Castor fiber (Rodentia: Castoridae)
Schizocarpus testiculatus Bochkov, Labrzycka, Skoracki & Saveljev, 2012: 44
Type locality: Suwalki
Distribution:. Poland
Host: Castor fiber belorussicus (Rodentia: Castoridae)
Schizocarpus tuvinicus Bochkov & Saveljev, 2012: 10
Type locality: Tuva, Todzhinskii District
Distribution: Russia
Host: Castor fiber tuvinicus (Rodentia: Castoridae)
Schizocarpus unzhakovi Bochkov et Saveljev, 2012: 8
Type locality: Tuva, Todzhinskii District
Distribution: Russia (Tuva)
Host: Castor fiber tuvinicus (Rodentia: Castoridae)
Schizocarpus zurowskii Bochkov, Labrzycka, Skoracki & Saveljev, 2012: 52; Bochkov & Saveljev, 2012: 16
Type locality: Suwalki
Distribution: Poland, Russia
Host: Castor fiber belorussicus (Rodentia: Castoridae)
Schizocoptes daberti Bochkov, 2016: 005
Type locality: South Kivu, Plantation Lemera
Distribution: DR Congo
Host: Chrysochloris stuhlmanni (Afrosoricida: Chrysochloridae)
Soricilichus sylvisorex Bochkov, Mbaltini & Verheyen, 2016: 249
Type locality: DR Congo
Distribution:. DR Congo
Hosts: Sylvisorex granti, S. lunaris, S. vulcanorum (Soricomorpha: Soricidae)
Atopomelus megalotis Bochkov, Abramov & Hugot, 2011: 6
Type locality: Kammouane Province, 18 km north of Thakhek, near Ban Doy village, 17º33’26’’N, 104º49’28’’E, 140 m a. s. l.
Distribution: Laos
Host: Hylomys megalotis (Erinaceomorpha: Erinaceidae)
Didelphoecius micoureus Bochkov & Valim, 2016: 32
Type locality: Minas Gerais
Distribution: Brazil
Host: Micoureus paraguayanus (Didelphimorphia: Didelphidae)
Listrophoroides oconnori Bochkov, Abramov & Hugot, 2011: 3
Type locality: Kammouane Province, 18 km north of Thakhek, near Ban Doy village, 17º33’26’’N, 104º49’28’’E, 140 m a. s. l.
Distribution: Laos
Host: Saxatilomys paulinae (Rodentia: Muridae)
EUROPE
Poland (Bochkov, Labrzycka, Skoracki & Saveljev, 2012) – Schizocarpus brachyurus, Sch. faini, Sch. fedjushini, Sch. gozdziewskii, Sch. heatherae, Sch. insignis, Sch. klompeni, Sch. parahumilis, Sch. pseudonumerosus, Sch. radiatus, Sch. testiculatus, Sch. zurowskii
Russia (entire)(Bochkov & Saveljev, 2012; Bochkov, 2012, 2016b) – Schizocarpus brachyurus (= Sch. intercalatus, = Sch. parabrachyurus), Sch. gozdziewskii, Sch. saveljevi, Schi. azasicus, Sch. daberti, Sch. heidekei, Sch. insignis, Sch. lavrovi, Sch. modestus, Sch. pusillus, Sch. radiatus, Sch. subornatus, Sch. tuvinicus, Sch. unzhakovi, Trichoecius dubininae, T. lemmus
AFRICA
Burundi (Bochkov, OConnor & Skoracki, 2016) - Myocoptes grammomys
DR Congo (Bochkov, Mbaltini & Verheyen, 2016; Bochkov, 2016) - Soricilichus sylvisorex, Schizocoptes daberti
Gabon (Bochkov, OConnor & Skoracki, 2016) - Trichoecius hylomyscus
South Africa (Bochkov, OConnor & Skoracki, 2016) – Trichoecius angolensis
Tanzania (Bochkov, OConnor & Skoracki, 2016) – Criniscansor congolensis, Gliricoptes vulcanorum, Myocoptes spinulatus, Trichoecius faini, T. hylomyscus, T. lootensis
Uganda (Bochkov, OConnor & Skoracki, 2016) - Lophuromys voosnami, Myocoptes grammomys, Trichoecius otomys
ASIA
Kazakhstan (Bochkov, 2016) - Trichoecius dubininae
Kirghizia (Bochkov, 2016) - Trichoecius dubininae
Laos (Bochkov, Abramov & Hugot, 2011) – Afrolistrophorus laonastes, A. saxatilomys, Pteromychirus menetes, Atopomelus megalotis, Listrophoroides oconnori
Vietnam (Bochkov, 2017, 2018) – Dubininetta abramovi, D. euroscaptor, Lutrilichus javanicus
SOUTH and CENTRAL AMERICA
Argentina (Sikora & Bochkov, 2012) - Prolistrophorus (Aprolistrophorus) parabidentatus, Prolistrophorus akodon, P. amazonicus, P. argentines, P. (P.) frontalis, P. (Beprolistrophorus) hirstianus
Bolivia (Sikora & Bochkov, 2012) - Prolistrophorus amazonicus
Brazil (Sikora & Bochkov, 2012, Bochkov & Valim, 2016; Gorza et al., 2018) – Didelphoecius micoureus, Leporacarus gibbus, L. sylvilagi, Lynxacarus serrafreirei, Parakosa tadarida, Prolistrophorus bidentatus, P. ctenomys, P. hylaeamys, Sclerolistrophorus oxymycteris
Chile (Sikora & Bochkov, 2012) - Amlistrophorus geoxus
Colombia (Bochkov, Lareschi & Barreto, 2014) - Prolistrophorus (Prolistrophorus) grassii, P. (P.) argentinus
Far East of Russia (Bochkov, 2017) - Dubininetta taiwanensis
Grenada (Springer et al., 2018) – Afrolistrophorus sp., Atopomelidae
Guatemala (Bochkov, Lareschi & Barreto, 2014) - Prolistrophorus (Aprolistrophorus) tylomys
Panama (Sikora & Bochkov, 2012) - Prolistrophorus nectomys
Peru (Sikora & Bochkov, 2012) - Prolistrophorus nectomys
Suriname (Sikora & Bochkov, 2012) - Prolistrophorus musculinus
Didelphimorphia
Fam. Didelphidae
Metachirus nudicaudatus – Didelphoecius validus
Micoureus paraguayanus - Didelphoecius micoureus
Erinaceomorpha
Fam. Erinaceidae
Hylomys megalotis - Atopomelus megalotis
Afrosoricida
Fam. Chrysochloridae
Chrysochloris stuhlmanni – Schizocoptes daberti
Soricomorpha
Fam. Soricidae
Sylvisorex granti - Soricilichus sylvisorex
S. lunaris - Soricilichus sylvisorex
S. vulcanorum - Soricilichus sylvisorex
Fam. Talpidae
Euroscaptor parvidens – Dubininetta euroscaptor
E. subanura – Dubininetta abramovi
Mogera robusta - Dubininetta taiwanensis
Chiroptera
Fam. Molossidae
Molossus molossus – Parakosa tadarida
Rodentia
Fam. Arvicolidae
Lasiopodomys gregalis – Trichoecius dubininae
Lemmus sibiricus - Trichoecius lemmus
Fam. Cricetidae
Akodon affinis - Prolistrophorus (Prolistrophorus) argentinus
A. azarae - Prolistrophorus (Aprolistrophorus) parabidentatus
A. cursor – Prolistrophorus bidentatus
A. montensis – Prolistrophorus akodon
Calomys callosus - Prolistrophorus amazonicus
C. laucha - Prolistrophorus amazonicus
C. musculinus - Prolistrophorus amazonicus
Geoxus valdivianus – Amlistrophorus geoxus
Hylaeamys laticeps - Prolistrophorus hylaeamys
Melanomys caliginosus - Prolistrophorus argentinus, P. nectomys
Nectomys palmipes – Prolistrophorus nectomys
Oligoryzomys sp. – Prolistrophorus (P.) frontalis
Oryzomys laticeps - Sclerolistrophorus oxymycteris
Scapteromys aquaticus - Prolistrophorus argentines, P. (Beprolistrophorus) hirstianus
Tylomys nudicaudus - Prolistrophorus (Aprolistrophorus) tylomys
Zygodontomys brevicauda - Prolistrophorus (Prolistrophorus) grassii
Fam. Ctenomyidae
Ctenomys torquatus – Prolistrophorus ctenomys
Fam. Muridae
Dendromus mysticalis – Myocoptes spinulatus
Grammomys dryas – Myocoptes grammomys
G. macmillani – Criniscansor congolensis
Hapalomys delacouri – Afrolistrophorus hapalomys
Hylomyscus anselli – Trichoecius hylomyscus
H. arcimontanus – Trichoecius faini
H. stella - Trichoecius hylomyscus
Meriones tamariscinus – Myocoptes meriones
M. libycus - Myocoptes meriones
M. meridianus – Trichoecius meriones
Otomys typus – Trichoecius otomys
Saxatilomys paulinae - Afrolistrophorus saxatilomys, Listrophoroides oconnori
Lagomorpha
Fam. Leporidae
Sylvilagus brasiliensis – Leporacarus sylvilagi
Fam. Castoridae
Castor fiber – Schizocarpus saveljevi
Castor fiber belorussicus – Schizocarpus brachyurus, Sch. heatherae, Sch. faini,
Sch. fedjushini, Sch. gozdziewskii, Sch. insignis, Sch. klompeni, Sch. parahumilis, Sch. pseudonumerosus, Sch. radiatus, Sch. testiculatus, Sch. zurowskii
Castor fiber tuvinicus - Schizocarpus azasicus, Sch. daberti, Sch. gozdziewskii, Sch. grandis, Sch. heidekei, Sch. insignis, Sch. lavrovi, Sch. modestus, Sch. pusillus, Sch. radiatus, Sch. subornatus, Sch. tuvinicus, Sch. unzhakovi, Sch. zurowskii
Carnivora
Fam. Mustelidae
Melogale moschata - Lutrilichus javanicus
Galictis cuja - Lynxacarus (Lynxacarus) serrafreirei
(suppl. 2011 - 2021)
Beron P. 2014. Acarorum Catalogus. III. Opilioacarida, Holothyrida, Mesostigmata (Dermanyssoidea). Pensoft & Nat. Mus. Natur. Hist. Sofia: 286 pp.
As a result of the recent contributions, the number of genera and species in Beron, 2014 (some of them of uncertain status) is updated as follows:
Opilioacarida – 13 genera, 55 recent and 3 fossil species
Holothyrida – 15 genera, 30 species
Macronyssidae – 34 genera, 236 species
Dermanyssidae – 3 genera, 39 species
Halarachnidae – 6 genera, 40 species
Raillietiidae – 1 genus, 7 species
Entonyssidae - 8 genera, 24 species
Manitherionyssidae – 1 genus, 1 species
Hystrichonyssidae - 1 genus, 1 species
Dasyponyssidae - 2 genera, 2 species
Spelaeorhynchidae - 1 genus, 7 species
Amazonacarus Vázquez, de Araújo & Feres, 2014: 152
Type species: Amazonacarus setosus Vázquez, de Araújo et Feres, 2014
Amazonacarus paraensis Vázquez, de Araújo & Feres, 2014: 162
Type locality: Para
Distribution: Brazil
Amazonacarus setosus Vázquez, de Araújo & Feres, 2014: 155
Type locality: Pará, Alter do Chão
Distribution: Brazil
Brasilacarus Vázquez, de Araújo & Feres, 2015: 377
Type species: Brasilacarus cocaris Vázquez, de Araújo et Feres, 2015
Brasilacarus cocaris Vázquez, de Araújo & Feres, 2015: 375
Type locality: Manaus, near Rio Negro and Taruma-Mirim stream
Distribution: Brazil
Neocarus belizensis Vázquez, May, Alamilla & Klompen, 2018: 132
Type locality: Maya Mountains
Distribution: Belize
Neocarus caipora Bernardi, Klompen & Ferreira, 2014: 47
Type locality: Canãa dos Carajas, Gruta Cav11-S11 (06°24’38.3”S 50°19’38.7”W)
Distribution: Brazil
Neocarus chactemalensis Vázquez & Klompen, 2015: 537
Type locality: Quintana Roo, Chetumal, campus Quintana Roo State University, 8 m
Distribution: Mexico
Neocarus comalensis Vázquez & Klompen, 2015: 538
Type locality: Guerrero, El Comal, elevation 1749 m
Distribution: Mexico
Neocarus coronatus : Araújo, Bichuette, Bauchan, Ochoa & Feres, 2018: 303
Type locality: São Desidério county, Bahia state
Distribution: Brazil
Neocarus entrerios Vázquez, Bernardi & Klompen, 2020: 512
Type locality: Misiones, P.N. Iguazu, Area de la Garganta del Diablo, 185 m
Distribution: Argentina
Type locality: Misiones, Iguazu, Puerto Libertad, 173 m, 25.9189°S 54.5818°W
Distribution: Argentina
Neocarus simmonsi Bernardi, Zampaulo & Oliveira, 2020: 14
Type locality: Minas Gerais State, São Gonçalo do Rio Abaixo city, Simmons’ cave, 19°53’55”S, 43°28’22”W
Distribution: Brazil
Neocarus spelaion Vázquez, Bernardi & Klompen, 2020: 3
Type locality: Minas Gerais State, Matozinhos, Cave 013, 19°31’57.21”S, 44°5’30.30”W
Distribution: Brazil
Opilioacarus baeticus Moraza Zorrilla, Prieto & Balanzateguic, 2021: 130
Type locality: Isleta del Moro: Barranco del Paraíso (Níjar, Almería)
Distribution: Spain
Opilioacarus brignolii Araújo & Di Palma, 2018: 138
Type locality: Apulia, Bari, Monopoli
Distribution: Italy, incl. Sardinia
Opilioacarus groehni Dunlop & Bernardi, 2014: 760
Type locality: from the Cretaceous Burmese amber
Distribution: Burma
Opilioacarus thaleri Vázquez, Ávila & Jus, 2021: 552
Type locality: Creta, Georgioupolis
Distribution: Greece (Creta)
Diplothyrus lehtineni Vázquez, Araújo & Feres, 2016: 271
Type locality: Pará, São Geraldo do Araguaia, Serra das Andorinhas
Distribution: Brazil
Parichoronyssus alexanderfaini Morales-Malacara & Guerrero, 2020: 405
Type locality: Loc. near Bartica, Brit. Guiana, Forest Reserve 24 mi. along Potaro Rd.
Distribution: Guiana
Host: Rhinophylla pumilio (Chiroptera: Phyllostomatidae)
Parichoronyssus gettingeri Morales-Malacara & Guerrero, 2020: 410
Type locality:
Distribution:
Hosts: Rhynchonycteris naso (Chiroptera: Embalonuridae), Noctilio leporinus (Chiroptera: Noctilionidae)
Steatonyssus olesovi Orlova, Stanyukovich & Orlov, 2015: 97
Type locality: Russia
Distribution: Russia
Hosts: Myotis dasycneme (probably the main host), M. blythii, M. daubentoni, M. brandtii, M. mystacinus, and Ognev’s long-eared bat.
Opilioacarida
EUROPE
Italy (Araújo, Di Palma & Feres, 2018) – Opilioacarus brignolii
Sardinia (Araújo, Di Palma & Feres, 2018) - Opilioacarus brignolii
Greece
Crete (Vázquez et al., 2021) – Opilioacarus thaleri
Spain (Moraza Zorrilla, Prieto & Balanzateguic, 2021) – Opilioacarus baeticus
NORTH AMERICA
Mexico (Vázquez & Klompen, 2015) - Neocarus chactemalensis, N. comalensis,
SOUTH and CENTRAL AMERICA
Argentina (Vázquez, Bernardi & Klompen, 2020) – Neocarus entrerios, N. misiones
Belize (Vázquez, May, Alamilla & Klompen, 2018) – Neocarus belizensis
Brazil (Araújo, Bichuette, Bauchan, Ochoa & Feres, 2018; Bernardi & Borges-Filho, 2018; Bernardi, Klompen & Ferreira, 2014; Vázquez, de Araújo & Feres, 2014, 2015; Bernardi, Zampaulo & Oliveira, 2020) – Amazonacarus paraensis, A. setosus, Brasilacarus cocaris, Neocarus caipora, N. coronatus, N. simmonsi, N. spelaion
Holothyrida
SOUTH and CENTRAL AMERICA
Brazil (Bernardi, Azara & Ferreira, 2011; Vázquez, Araújo & Feres, 2016) – Diplothyrus lehtineni, D. schubarti According to Bernardi et al., 2011 (fide Lehtinen), Holothyrida are known also from the Philippines, Costa Rica and Panama.
Some Mesostigmata
EUROPE
Italy (Cafiero et al., 2015) - Ornithonyssus bacoti
Russia (Orlova, Smirnov et al., 2017; Orlova, Krilov et al., 2021) – Cryptonyssus flexus, Macronyssus diversipilis, M. leislerianus, Steatonyssus noctulus
Turkey (Ciloglu et al., 2020) – Dermanyssus gallinae
AFRICA
Egypt (Negm et al., 2018; Halawa et al., 2019) – Ornithonyssus sylviarum, Steatonyssus longipes, Dermanyssus
Namibia (Orlova, Laverty et al., 2020) - Steatonyssus (S.) afer
NORTH AMERICA
Mexico (Morales-Malacara & Guerrero, 2020) - Parichoronyssus gettingeri
SOUTH and CENTRAL AMERICA
Argentina (Lareschia et al., 2017) - Ornithonyssus bursa
Guiana (Morales-Malacara & Guerero, 2020) - Parichoronyssus alexanderfaini
Peru (Morales-Malacara & Guerrero, 2020; Angoma et al., 2021) – acari on bats
Venezuela (Morales-Malacara & Guerrero, 2020) – Parichoronyssus gettingeri
Aves
Columbiformes
Fam. Columbidae
Columba livia domestica - Ornithonyssus bursa, O. sylviarum, Steatonyssus longipes
Streptopelia senegalensis aegyptiaca - Ornithonyssus bursa, Steatonyssus longipes
Passeriformes
Fam. Ploceidae
Ploceus philippinus – Pellonysus flabellifer
Fam. Passeridae
Passer domesticus niloticus - Ornithonyssus bursa, Steatonyssus longipes
Fam. Pycnonotidae
Pycnonotus barbatus – Ornithonyssus bacoti
Fam. Sturnidae
Sturnus vulgaris - Ornithonyssus bursa
Mammalia
Chiroptera
Fam. Cistugidae
Cistugo seabrae - Steatonyssus (S.) afer
Fam. Vespertilionidae
Myotis tschuliensis – Macronyssus diversipilis
Nyctalus leisleri – Macronyssus leislerianus
N. noctula – Steatonyssus noctulus
Pipistrellus nathusii – Cryptonyssus flexus
(suppl. 2014 - 2021)
Opilioacarida
Holothyrida
Some Mesostigmata
Beron P. 2015. Acarorum Catalogus. IV. Myobioidea, Pterygosomatoidea. Pensoft & Nat. Mus. Natur. Hist. Sofia: 251 pp.
As a result of the recent contributions, the number of genera and species in Myobiidae and Pterygosomatoidea (some of them with uncertain status) is updated as follows:
Myobiidae – 55 genera, 511 species
Pterygosomatidae – 13 genera, 196 species
Crocidurobia faini Bochkov, Mbalitini & Verheyen, 2016: 496
Type locality: DR Congo
Distribution: DR Congo
Hosts: Crocidura denti, Crocidura cf. niobe, Crocidura cf. littoralis (Soricomorpha: Soricidae)
Crocidurobia lukoschusi Bochkov, 2017: 212
Type locality: Vietnam
Distribution: Vietnam
Host: Crocidura tanakae (Soricomorpha: Soricidae)
Radfordia (Hesperomyobia) akodon Bochkov, 2017b
Type locality: Bolivia
Distribution: Bolivia
Host: Akodon varius (Rodentia: Cricetidae, Sigmodontinae)
Radfordia (Radfordia) mirabilis Bochkov & Abramov, 2014: 142
Type locality: Binh Phuoc Province, Bu Gia Map National Park, 13
km NE Bu Gia Map Village, 540 m a.l.s., 12°11’37”N, 107°12’21”E
Distribution: Vietnam
Host: Hapalomys delacouri (Rodentia: Muridae)
†Protohylomysobia erinaceophilus Sidorchuk et Bochkov, 2018 (2019)
Callopistiella Silva-de la Fuente, Casanueva & Moreno Salas, 2015: 66
Type species: Callopistiella atacamensis Silva-de la Fuente, Casanueva et Moreno Salas, 2015
Callopistiella atacamensis Silva-de la Fuente, Casanueva & Moreno Salas, 2015: 67
Type locality: Caldera: Copiapó Province, Atacama Region, 26°56’S, 70°44’W
Distribution: Chile
Host: Callopistes maculatus (Sauria: Teiidae)
Geckobia africana Fajfer, 2018: 2
Type locality: Irimga Region, Mufindi District, Uzungwa Scarp Forest Reserve
Distribution: Tanzania
Host: Cnemaspis africana (Sauria: Gekkonidae)
Geckobia andina Machado et al., 2019: 335
Type locality: Cerro Verde, Andean highlands
Distribution: Peru
Host: Phyllodactylus gerrhopygus (Sauria: Gekkonidae)
Geckobia circumdata Machado et al., 2019: 342
Type locality: Arequipa, Cerro Verde, 2345 m
Distribution: Peru
Host: Phyllodactylus gerrhopygus (Sauria: Gekkonidae)
Geckobia milii Fajfer, 2019: 6
Type locality: Far North Queensland, Cape York Peninsula
Distribution: Australia (Queensland)
Host: Underwoodisaurus milii (Sauria: Carphodactylidae)
Neopterygozoma Fajfer, 2019: 422
Type species: Pterygosoma chilensis Fajfer et González-Acuña, 2013
Neopterygosoma schroederi Fajfer, 2020: 544
Type locality: Maule region, Curicó Prov., Curicó
Distribution: Chile
The species Pterygosoma chilensis, P. cyanopteri, P. formosus, P. levissima, P. ligare, P. ovata, described by Fajfer & Gonzalez – Acuña in 2013, as well as Pterygosoma patagonica Ditmar de la Cruz, Morando et Avila, 2004, have been transferred by Fajfer (2020a) in the genus Neopterygosoma.
Pterygosoma agamae Peters, 1849 – published again from Tanzania and Mozambique (Fajfer, 2020b: 99)
Pterygosoma balochistani Fajfer, 2016a: 348; 2020b: 26
Type locality: Balochistan, Darzi Chan
Distribution: Pakistan
Host: Landakia nupta nupta (Sauria: Agamidae)
Pterygosoma blanfordi Fajfer, 2016a: 344
Type locality: South India, Kerala State, Palakkad district, Parambiculam Tiger Reserve
Distribution: India
Host: Psammophilus blanfordianus (Sauria: Agamidae)
Pterygosoma gladiator Bertrand, Finkelman et Paperna, 2000 is synonymised with P. neumanni (Berlese, 1910) by Fajfer, 2020b: 111
Pterygosoma livingstonei Bertrand et Modry, 2004 is synonymised with P. circularis Jack, 1962 by Fajfer, 2020b
Pterygosoma pallidum Fajfer, Melnikov & Dabert, 2016: 792; Fajfer, 2020b: 11
Type locality: Aquaba Province, 11 km north-east of Ar- Risha
Distribution: Jordan
Host: Trapelus pallidum (Sauria: Agamidae)
Pterygosoma parasiniatum Fajfer, Melnikov & Dabert, 2016: 801
Type locality: NE of Ar-Risha
Distribution: Jordan
Host: Pseudotrapelus cf. sinaiticus (Sauria: Agamidae)
Pterygosoma rhipidostichata Bertrand, Finkelman and Paperna, 2000 is synonymised with P. mutabilis Jack, 1961 by Fajfer, 2020b
Pterygosoma theobaldi Fajfer, Melnikov & Dabert, 2016: 806
Type locality: North India
Distribution: India
Host: Phrynocephalus teobaldi (Sauria: Agamidae)
Pimeliaphylus hemidactyli Fajfer & Karanth, 2021: 442
Type locality: Karnataka State, Bangalore, IISc campus
Distribution: India
Hosts: Hemidactylus murrayi, H. leschenaultia, H. parvimaculatus (Squamata: Gekkonidae)
New hosts and distribution records are given for Pterygosoma neumanni (Berlese, 1910), P. bibronii Jack, 1962, P. longipalpae Lawrence, 1936, P. gracilipalpis and P. sinaita Jack, 1961.
(suppl. 2015 - 2021)
Myobioidea
ASIA
Vietnam (Bochkov, 2017) – Crocidurobia lukoschusi
NORTH AMERICA
Mexico (Herrera-Mares et al., 2017; Herrera-Mares, Guzmán-Cornejo & Morales-Malacara, 2021) - Acanthophthirius lasiurus, Ioanella mimon, Eudusbabekia jimenezi, E. mimon, E. saguei
SOUTH and CENTRAL AMERICA
Bolivia (Bochkov, 2017) - Radfordia (Hesperomyobia) akodon, R. (H.) vandenberghi
Pterygosomatoidea
AFRICA
Tanzania (Fajfer, 2018) – Geckobia africana
South Africa (OConnor, 2020) - Zonurobia circularis limpopoensis
ASIA
India (Fajfer, 2016a) - Pterygosoma blanfordi, P. theobaldi
Jordan (Fajfer, 2016b) - Pterygosoma pallidum, P. parasiniatum
Pakistan (Fajfer, 2016a) - Pterygosoma balochistani
SOUTH and CENTRAL AMERICA
Argentina (Ditmar de la Cruz, Morando & Avila, 2004, Fajfer, 2020) – Neopterygosoma patagonica
Chile (Fajfer, 2019, 2020) – Neopterygosoma spp.(formerly in Pterygosoma), Neopterygosoma schroederi, Callopistiella atacamensis
Cuba (Borroto Paez et al., 2020) – Geckobia hemidactyli
Panama (Domínguez et al., 2020) - Geckobiella stamii
Peru (Machado et al., 2019) – Geckobia andina, G. circumdata
OCEANIA
Australia (Fajfer, 2018) – Geckobia milii
Myobioidea
Soricomorpha
Fam. Soricidae
Crocidura denti - Crocidurobia faini
Crocidura cf. niobe - Crocidurobia faini
Crocidura cf. littoralis - Crocidurobia faini
Crocidura tanakae - Crocidurobia lukoschusi
Chiroptera
Fam. Phyllostomatidae
Mimon cozumelae - Eudusbabekia mimon, Ioanella mimon
Rodentia
Fam. Cricetidae
Akodon varius - Radfordia (Hesperomyobia) akodon
Oligoryzomys flavescens - Radfordia (Hesperomyobia) vandenberghi
Fam. Muridae
Hapalomys delacouri - Radfordia (Radfordia) mirabilis
Pterygosomatoidea
Reptilia
Sauria
Fam. Agamidae
Landakia nupta nupta - Pterygosoma balochistani
Phrynocephalus teobaldi - Pterygosoma theobaldi
Psammophilus blanfordianus - Pterygosoma theobaldi
Pseudotrapelus cf. sinaiticus - Pterygosoma parasiniatum
Trapelus pallidum - Pterygosoma pallidum
Fam. Carphodactylidae
Underwoodisaurus milii – Geckobia milii
Fam. Gekkonidae
Cnemaspis africana – Geckobia africana
Hemidactylus mabouia – Geckobia hemidactyli
H. murrayi – Pimeliaphilus hemidactyli
H. parvimaculatus - Pimeliaphilus hemidactyli
H. leschenaultia - Pimeliaphilus hemidactyli
Phyllodactylus gerrhopygus – Geckobia andina, G. circumdata
Fam. Teiidae
Callopistes maculatus - Callopistiella atacamensis
(suppl. 2015 - 2021)
Myobioidea
Pterygosomatoidea
Beron P. 2015. Acarorum Catalogus. V. Sarcoptoidea. Pensoft & Nat. Mus. Natur. Hist. Sofia: 380 pp.
As a result of the recent contributions, the number of genera and species in Sarcoptoidea (some of them uncertain status) is updated as follows:
Lemurnyssidae – 2 genera, 4 species
Lobalgidae – 2 genera, 9 species
Psoroptidae – 31 genera, 58 species
Listropsoralgidae - 4 genera, 19 species
Bochkovocoptidae – 1 genus, 1 species
Rhyncoptidae - 5 genera, 13 species
Sarcoptidae - 17 genera, 117 species
Pneumocoptidae - 1 genus, 4 species
Gastronyssidae - 9 genera, 42 species
Chirorhynchobiidae - 1 genus, 3 species
Pyroglyphidae - 22 genera, 51 species
Dermationidae - 12 genera, 54 species
Epidermoptidae - 16 genera, 51 species
Gaudoglyphidae – 1 genus, 1 species
Ptyssalgidae – 1 genus, 1 species
Turbinoptidae - 9 genera, 36 species
Cytoditidae - 2 genera, 14 species
Laminosioptidae – 6 genera, 27 species
Guanolichidae - 3 genera, 4 species
Echimytricalges lonchothrix Bochkov & OConnor, 2010: 218
Type locality: Para, Rio Tapajoz, Boim, 03°05’S, 55°18’W
Distribution: Brazil
Host: Lonchothrix emiliae (Rodentia: Echimyidae)
Echimytricalges proechimys Bochkov & OConnor, 2010: 214
Type locality: Oriente, Rio Jatun Yacu
Distribution: Ecuador
Host: Proechimys brevicauda (Rodentia: Echimyidae)
Echimytricalges whitakeri Fain & Ritzi, 2001 – validated by Bochkov & OConnor, 2010: 221 from nomen nudum
Caparinia ictonyctis Lawrence, 1955: 54
Type locality: South Africa
Distribution: South Africa
Host: Atelerix albiventris (Erinaceomorpha: Erinaceidae)
Gaudalges averianovi Bochkov, 2018: 593
Type locality: Madagascar
Distribution: Madagascar
Host: Lepidolemur ruficaudatus (Primates: Lepilemuridae)
Lemuralges propithecus Bochkov, Klompen, Junge & Williams, 2015: 2
Type locality: Madagascar
Distribution: Madagascar
Host: Propithecus diadema (Primates: Indridae)
Didelphialges metachirus Bochkov & Valim, 2013: 67
Type locality: Peru
Distribution: Peru
Host: Metachirus nudicaudatus (Didelphimorphia: Didelphidae)
Listropsoralges brevisetosus Bochkov & Valim, 2013: 39
Type species: Peru
Distribution: Peru
Host: Marmosa murina (Didelphimorphia: Didelphidae)
Listropsoralges similis Bochkov & Valim, 2013: 19
Type locality: Panama
Distribution: Panama
Host: Caluromys derbianus (Didelphimorphia: Didelphidae)
Listropsoralges thylamys Bochkov & Valim, 2013: 24
Type locality: Bolivia
Distribution: Bolivia
Host: Thylamys venustus (Didelphimorphia: Didelphidae)
Listropsoralges vossi Bochkov & Valim, 2013: 35
Type locality: Brazil
Distribution: Brazil
Host: Monodelphis domestica (Didelphimorphia: Didelphidae)
Listropsoralges caenolestes Bochkov & Valim, 2013: 26
Type locality: Ecuador
Distribution: Ecuador
Host: Caenolestes fuliginosus (Paucituberculata: Caenolestidae)
The female of Listropsoralges faini Bochkov et Wauthy, 2009 is described in the revision of Bochkov & Valim (2016) for the first time.
Bochkovocoptidae OConnor, 2019: 146
Type genus: Bochkovocoptes OConnor, 2019
Bochkovocoptes OConnor, 2019: 146
Type species: Bochkovocoptes otomys OConnor, 2019
Bochkovocoptes otomys OConnor, 2019: 146
Type locality:Kigezi Province, Kisoro Dstrict, Mgahinga Gorilla Nat. Park
Distribution: Uganda
Host: Otomys denti kempi (Rodentia: Muridae)
Tachornithoglyphus Klimov, Bochkov & OConnor, 2015: 97
Type species: Guatemalichus tachornis Cruz, Cuervo et Dusbábek, 1984
Guatemalichus tachornis Cruz, Cuervo & Dusbábek, 1984: 1; Cruz 1988: 14; Fain 1988: 22
Tachornithoglyphus tachornis (Cruz, Cuervo et Dusbábek, 1984): Klimov, Bochkov & OConnor, 2015: 97
Type locality: Ciudad de La Habana, Playa El Salado, Bauta
Distribution: Cuba
Host: nest of Tachornis phoenicobia (Apodiformes: Apodidae)
Carcinopodacarus Hernandes, Pedroso & Bochkov, 2015: 2
Type species: Carcinopodacarus polymorphus Hernandes, Pedroso et Bochkov, 2015
Carcinopodacarus polymorphus Hernandes, Pedroso & Bochkov, 2015: 3
Type locality: Campinas
Distribution: Brazil
Host: Guira guira (Cuculiformes: Cuculidae)
Neodermation actitis Bochkov & Mironov, 2015: 612; Matyukhin & Yatsuk, 2022
Type locality: the Far East of Russia
Distribution: Russia
Host: Actitis hypoleucos (Charadriiformes: Scolopacidae)
Promyialges italicus Moradi Faradonbeh, Ostovan, OConnor, Gheibi, Joharchi & Macchioni, 2019: 36
Type locality: Tuscany, Pisa, 43° 42’ N, 10° 24’ E
Distribution: Italy
Host: Pseudolynchia canariensis (Diptera: Hippoboscidae) from Columba livia
(suppl. 2016 - 2021)
EUROPE
Italy (Moradi Faradonbeh et al., 2019) - Promyialges italicus
Russia (Bochkov & Mironov, 2015) – Dermation actitis (the Far East)
AFRICA
Egypt (Nageiba et al., 2001; Halawa et al., 2019) – Metamicrolichus pici, Promyalges falconis
Madagascar (Bochkov, 2018) - Gaudalges averianovi
ASIA
India (Modak et al., 2004; Podder, Biswas & Kumar Saha, 2021) – Pyroglyphidae
NORTH AMERICA
Mexico (Rodríguez-Ortega et al., 2018) - Knemidocoptes mutans
SOUTH and CENTRAL AMERICA
Bolivia (Bochkov & Valim, 2013) - Listropsoralges thylamys
Brazil (Bochkov & OConnor, 2010; Bochkov & Valim, 2013; Hernandes, Pedroso & Bochkov, 2015) – Carcinopodacarus polymorphus, Echimytricalges lonchothrix, E. mesomys, Listropsoralges vossi
Colombia (Bochkov & OConnor, 2010) – Echimytricalges surinamensis
Costa Rica (Bochkov & OConnor, 2010) – Echimytricalges hoplomys
Cuba (Cruz, Cuervo & Dusbabek, 1984) - Tachornithoglyphus tachornis
Ecuador (Bochkov & OConnor, 2010; Bochkov & Valim, 2013) - Echimytricalges hoplomys, E. proechimys, Listropsoralges caenolestes
Panama (Bochkov & Valim, 2013) - Listropsoralges similis
Peru (Bochkov & OConnor, 2010; Bochkov & Valim, 2013) – Echimytricalges guianensis, Listropsoralges brevisetosus
AVES
Charadriiformes
Fam. Scolopacidae
Neodermation actitis - Actitis hypoleucos
Cuculiformes
Fam. Cuculidae
Guira guira - Carcinopodacarus polymorphus
Passeriformes
Fam. Passeridae
Passer domesticus niloticus – Promyalges falconis, Metamicrolichus pici
MAMMALIA
Didelphimorphia
Fam. Didelphidae
Caluromys derbianus - Listropsoralges similis
Marmosa murina - Listropsoralges brevisetosus
Metachirus nudicaudatus - Didelphialges metachirus
Monodelphis domestica - Listropsoralges vossi
Thylamys venustus - Listropsoralges thylamys
Erinaceomorpha
Fam. Erinaceidae
Atelerix albiventris - Caparinia ictonyctis
Rodentia
Fam. Echimyidae
Hoplomys gymnurus – Echimytricalges hoplomys
Lonchothrix emiliae – Echimytricalges lonchothrix
Mesomys hispidus – Echimytricalges mesomys
Proechimys brevicauda – Echimytricalges proechimys
P. oconneli – Echimytricalges surinamensis
P. semispinosus - Echimytricalges hoplomys
P. simonsi – Echimytricalges guianensis
Trinomys albispinus – Echimytricalges brasiliensis
T. iheringi - Echimytricalges brasiliensis
Fam. Muridae
Otomys denti kempi - Bochkovocoptes otomys
Primates
Fam. Indriidae
Propithecus diadema - Lemuralges propithecus
Fam. Lepilemuridae
Lepidolemur ruficaudatus - Gaudalges averianovi
(suppl. 2015 - 2021)
Beron P. 2020. Acarorum Catalogus. VI. Rhinonyssidae, Spinturnicidae. Pensoft & Nat. Mus. Natur. Hist. Sofia: 265 pp.
As a result of the recent contributions, the number of genera and species (some of them uncertain status) is updated as follows:
Rhinonyssidae – 16 genera, 457 species
Spinturnicidae – 12 genera, 111 species
Ptilonyssus dimi Dimov, 2020b: 6
Type locality: Elez, Lipezk region
Distribution: Russia
Host: Hirundo rustica (Passeriformes: Hirundinidae)
Ptilonyssus harko Dimov, 2020b: 2
Type locality: Viazovka, Orenburg region
Distribution: Russia
Host: Riparia riparia (Passeriformes: Hirundinidae)
Ptilonyssus poriazovi Dimov, 2020b: 9
Type locality: Kohma, Ivanovo region
Distribution: Russia
Host: Emberiza citrinella (Passeriformes: Emberizidae)
Rhinonyssus derojasi Dimov, 2020a: 2
Type locality: Revda, Sverdlovsk Region
Distribution: Russia
Host: Anser anser (Anseriformes: Anatidae)
Rhinonyssus karelinae Dimov, 2020a: 11
Type locality: Tihvin, Leningrad region
Distribution: Russia
Host: Charadrius dubius (Charadriiformes: Charadriidae)
Rhinonyssus mileni Dimov, 2020a: 8
Type locality: Dubovka, Volgograd Region
Distribution: Russia
Host: Charadrius dubius (Charadriiformes: Charadriidae)
Rhinonyssus sixli Dimov, 2020a: 5
Type locality: Mosolovo, Kursk region
Distribution: Russia
Host: Podiceps ruficollis (Podicipediformes: Policipedidae)
Sternostoma alexmironi Dimov, 2020c: 2
Type locality: Opochka, Pskov region
Distribution: Russia
Host: Oenanthe oenanthe (Passeriformes: Muscicapidae)
Sternostoma antoni Dimov, 2020c: 5
Type locality: Liski, Voronej region
Distribution: Russia
Host: Dendrocopos major (Piciformes: Picidae)
Sternostoma chakarovae Dimov, 2020c: 8
Type locality: Hvalovo, Leningrad region
Distribution: Russia
Host: Oenanthe oenanthe (Passeriformes: Muscicapidae)
Sternostoma delianovae Dimov, 2020c: 11
Type locality: Kirishi, Leningrad region
Distribution: Russia
Host: Turdus pilaris (Passeriformes: Turdidae)
Sternostoma enevi Dimov, 2020c: 14
Type locality: Budenovsk, Stavropol region
Distribution: Russia
Host: Turdus pilaris (Passeriformes: Turdidae)
Periglischrus empheresotrichus Morales-Malacara, Castaño-Meneses & Klompen in Morales-Malacara, Castaño-Meneses, Klompen & Mancina, 2019: 421
Type locality: Jamaica, St. Ann Parish; ½ mi. S, ½ mi, W. Runaway Bay
Distribution: Cuba, Dominica, Jamaica, Haiti, Guadeloupe
Hosts: Monophyllus redmani, M. plethodon (Chiroptera: Phyllostomatidae)
yerbabuenae (Zamora-Mejias, Morales - Malacara et al., 2020)
(suppl. 2020 - 2021)
Fam. Rhinonyssidae
EUROPE
Russia (Dimov, 2017, 2020a, b, c, d; Dimov & Zhigulin, 2020; Veiga et al., 2021) – Ptilonyssus dimi, P. harko, P. poriazovi, Rhinonyssus derojasi, Rh. karelinae, Rh. mileni, Rh. sixli, Sternostoma alexmironi, S. antoni, S. chakarovae, S. delianovae, S. enevi
SOUTH AMERICA
Fam. Spinturnicidae
EUROPE
Russia (Orlova, Klimov et al., 2021; Orlova, Vu Dinh Thong et al., 2021)
Crimea (Orlova & Klimov, 2020) – Spinturnix psi
AFRICA
Egypt (Hafez et al., 1994) - Meristaspis kenaensis, Ancystropus aethiopicus
ASIA
Iran (Yousefi et al., 2018) – Spinturnix emarginata
NORTH and CENTRAL AMERICA
Mexico (Zamora-Mejias, Morales – Malacara et al., 2020; Morales-Malacara et al., 2020) – Periglischrus caligus, P. empheresotrichus
SOUTH AMERICA
Paraguay (Morales - Malacara et al., 2020) – Periglischrus herrerai
Rhinonyssidae
Aves
Anseriformes
Fam. Anatidae
Anser anser – Rhinonyssus derojasi
Charadriiformes
Fam. Charadriidae
Charadrius dubius – Rhinonyssus karelinae, Rh. mileni
Passeriformes
Fam. Emberizidae
Emberiza citrinella – Ptilonyssus poriazovi
Fam. Hirundinidae
Hirundo rustica – Ptilonyssus dimi
Riparia riparia – Ptilonyssus harco
Fam. Muscicapidae
Oenanthe oenanthe - Sternostoma alexmironi, S. chakarovae
Fam. Turdidae
Turdus pilaris – Sternostoma delianovae, S. enevi
Piciformes
Fam. Picidae
Dendrocopos major – Sternostoma antoni
Podicipediformes
Fam. Podicipedidae
Podiceps ruficollis – Rhinonyssus sixli
Chiroptera
Fam. Spinturnicidae
Fam. Pteropodidae
Rousettus aegyptiacus - Meristaspis kenaensis, M. lateralis, Ancystropus aethiopicus, A. zeleborii
Fam. Vespertilionidae
Myotis emarginatus – Spinturnix emarginata
Fam. Phyllostomatidae
Periglischrus herrerai Machado-Allison, 1965 is recorded also on Anoura sp., Sturnira ludovici, S. lilium by Morales-Malacara et al. (2020).
(suppl. 2020 - 2021)
Beron P. 2020. Acarorum Catalogus. VII. Raphignathoidea. Pensoft & Nat. Mus. Natur. Hist. Sofia: 306 pp.
As a result of the recent contributions, the number of genera and species in Raphignathoidea (some of them uncertain status) is updated as follows:
Barbutiidae – 1 genus, 7 species
Caligonellidae – 5 genera, 66 species
Camerobiidae – 7 genera, 175 species
Cryptognathidae – 3 genera, 66 species
Dasythyreidae – 2 genera, 3 species
Dytiscacaridae – 1 genus, 3 species
Eupalopsellidae – 7 genera, 50 species
Homocaligidae – 2 genera, 8 species
Mecognathidae – 2 genera, 7 species
Raphignathidae – 3 (?) genera, 83 species
Stigmaeidae – 33 genera, 640 species
Xenocaligonellidae – 2 genera, 5 species
Raphignathoidea total - 68 genera, 1113 species
Barbutia arasbaraniensis Mohammad - Doustaresharaf & Bagheri, 2021a: 10
Type locality:
Distribution: Iran
Barbutia theroni Khaustov, Vorontsov, Perkovsky & Klimov, 2021: 975
Type locality: Rovno
Distribution: Ukraine
Caligonella quinqueocellata Khaustov, 2021: 911
Type locality: Arabat Spit
Distribution: Crimea
Caligonella astragalusi Pishehvar & Khanjani, 2020: 1989
Type locality: Nahavad, Hamedan Province
Distribution: Iran
Molothrognathus sibiricus Khaustov, 2022: 99
Type locality: Altai Republic, Kosh-Agach, 1765 m
Distribution: Russia (Altai Republic)
Molothrognathus tauricus Khaustov, 2021: 917
Type locality: Arabat Spit
Distribution: Crimea
Neognathus sibiricus Khaustov, 2022: 99
Type locality: Tyumen Region, vicinity of lake Kushak
Distribution: Russia (Siberia)
Neophyllobius cibyci Paredes-León, Corona-López, Flores-Palacios & Toledo, 2016: 5
Type locality: dry forest 1 km south of San Andres de la Cal, Tepotzlán, Morelos
Distribution: Mexico
Host: inhabiting bromeliads (Tillandsia spp.) and living on two tree species (Quercus obtusata and Sapium macrocarpum)
Neophllobius denizliensis Akyol, 2020b: 88
Type locality: Denizli Province
Distribution: Turkey
Neophyllobius electrus Zmudzinski, 2020: 698
Type locality: Eocene Baltic Amber
Distribution: Baltic Amber
Neophyllobius glaesus Zmudzinski, 2020: 706
Type locality: Eocene Baltic Amber
Distribution: Baltic Amber
Neophyllobius tepoztlanensis Paredes-León, Corona-López, Flores-Palacios & Toledo, 2016: 14
Type locality: dry forest 1 km south of San Andres de la Cal, Tepotzlán, Morelos
Distribution: Mexico
Host: inhabiting bromeliads (Tillandsia spp.) and living on two tree species (Quercus obtusata and Sapium macrocarpum)
Tillandsobius Bolland, 1985 = Tycherobius Bolland, 1986 (after Mirza, Kamran & Alatawi, 2022: 5)
Tillandsobius floridensis Bolland, 1986 = Tycherobius floridensis (Bolland) - Mirza et al., 2022: 5
Neophyllobius texanus McGregor, 1950 = Neophyllobius (Monophyllobius Mirza, 2022:6) texanus McGregor
Neophyllobius electrus Zmudzinski, 2020 = Monobius electrus (Zmudzinski, 2020) - Mirza et al., 2022: 5
Neophyllobius meyerae Bolland, 1991 = Monobius meyerae (Boland) - Mirza et al., 2022: 5
Neophyllobius tescalicola Paredes-León, Corona-López, Flores-Palacios & Toledo, 2016: 17
Type locality: dry forest 1 km south of San Andres de la Cal, Tepotzlán, Morelos
Distribution: Mexico
Host: in soil and litter under Quercus obtusata
Cryptognathus karabagiensis Akyol, 2020c: 1644
Type locality: Karabag
Distribution: Turkey
Favognathus rosulatus Doğan & Doğan, 2020b: 71
Type locality: Turkey
Distribution: Turkey
Raphignathus kulaensis Akyol, 2020a: 1096
Type locality: Manisa and Denizli provinces
Distribution: Turkey
Raphignathus rakhshandehi Pishehvar & Khanjani, 2021: 185
Type locality: along Gamasiab river, Nahavand, Hamedan Prov.
Distribution: Iran
Raphignathus seraji Pishehvar & Khanjani, 2021: 186
Type locality: along Gamasiab river, Nahavand, Hamedan Prov.
Distribution: Iran
Agistemus piquinnus Monjarás-Barrera & Johann, 2020: 472
Type locality: Ciudad Victoria, Tamaulipas
Distribution: Mexico
Eustigmaeus crassifolius Bizarro & Johann, 2020: 826
Type locality: Belém city (1°26’09.2”S, 48°26’28.6”W), Pará state
Distribution: Brazil
Prostigmaeus amplius Doğan, Doğan & Türk, 2020: 1076
Type locality: Turkey
Distribution: Turkey
Stigmaeus akimovi Kharadov, 2021a: 43
Type locality: Altai Republic,
Distribution: Russia
Stigmaeus bagherii Rostami & Doustaresharaf, 2021: 222
Type locality: East Azerbaijan Province, north-western Iran
Distribution: Iran
Stigmaeus exilis Doğan & Doğan, 2021: 20
Type locality: Sansa Gorge
Distribution: Turkey
Stigmaeus lorestaniensis Ahmad-Hosseini & Jafari, 2022: 238
Type locality: Shorab Vill., Lorestan Prov.
Distribution: Iran
Stigmaeus pulumurensis Doğan & Doğan, 2020a: 42
Type locality: Pülümür Valley
Distribution: Turkey
Stigmaeus strobiculatus Kharadov, 2021a: 248
Type locality: Western Siberia
Distribution: Russia
EUROPE
Russia (Khaustov, 2021a,b) – Stigmaeus akimovi (Altai), S. scrobiculatus (Siberia)
Crimea (Khaustov, 2021c) – Caligonella quinqueocellata, Molothrognathus turcicus
Spain (Santamaría et al., 2012) – Eustigmaeus [Ledermulleria] segnis, Mediolata similans
Turkey (Doğan, Doğan & Erman. 2018; Akyol, 2020a, 2020b, 2020c, 2021a, 2021b; Akyol & Koc, 2020; Cobanoglu, Ögreten & Sade, 2021; Doğan, 2020; Doğan & Doğan, 2020a, b, c; Doğan & Doğan, 2021; Doğan, Doğan & Türk, 2018, 2020) – Cryptognathus karabagiensis, Favognathus rosulatus, Molothrognathus shirazicus, Paraneognathus wangae, Prostigmaeus amplius, P. kannjanii, Raphignathus gracilis, R. kulaensis, Stigmaeus exiis, S. pulumurensis, S. shabestariensis
Ukraine (Khaustov et al., 2021) – Barbutia theroni
ASIA
Iran (Majidi, Hajikanbar & Saboori, 2019; Siahkamari et al., 2020; Khosravinedzad, Jafari & Paktinat-Saeij, 2020; Pishehvar & Khanjani, 2021; Rostami, 2021; Gassemi & Hadjizadeh, 2021; Doustaresharaf & Bagheri, 2021a, 2021b, Ahmad-Hosseini & Jafari, 2022) – Barbutia arasbaraniensis, Eustigmaeus johnstoni, Raphignathus rakhshandehi, R. seraji, Stigmaeus bagherii, S. furcatus, S. lorestaniensis, S. sinai, S. youngi
Saudi Arabia (Mirza, Kamran & Alatawi. 2022) - Camerobia southcotti, Neophyllobius combreticola, N. denizliensis, N. lorestanicus, Tycherobius emadi
NORTH AMERICA
Mexico (Monjarás-Barrera et al., 2020) – Agistemus brasiliensis, A. piquinnus
SOUTH AMERICA
Brazil (Bizarro et al., 2020; Toldi et al., 2021 - Eustigmaeus crassifolius
Diptera
Fam. Psychodidae
Phlebotomus (Paraphlebotomus) alexandri – Eustigmaeus johnstoni, Stigmaeus furcatus, S. sinai
P. (Phlebotomus) papatasi - Eustigmaeus johnstoni, S. sinai, S. youngi
Sergentomyia (Sintonius) clydei - Eustigmaeus johnstoni
Se. (Se.) sintoni - Eustigmaeus johnstoni, Stigmaeus youngi
Se. (Se.) theodori - Stigmaeus youngi
(suppl. 2020 - 2021)
Beron P. 2020. Acarorum Catalogus. VIII. Cheyletoidea, Cloacaroidea. Pensoft & Nat. Mus. Natur. Hist. Sofia: 465 p.
As a result of the recent contributions, the number of genera and species in and (some of them uncertain status) is updated as follows:
Cheyletidae – 80 genera, 474 species
Psorergatidae – 3 genera, 77 species
Demodecidae – 8 genera, 122 species
Harpyrhynchidae - 16 genera, 135 species
Syringophilidae - 63 genera, 416 species
Cloacaridae - 6 genera, 15 species
Epimyodicidae – one genus, four species
Fam. Cheyletidae
Bak deleoni Yunker, 1961 – new for India (Modak et al., 2004)
Chelacaropsis moorei Baker, 1949 – India, W. Bengal (A. Gupta et al., 2006), India, Andaman Isl. (Gupta & Ghosh, 1980)
Cheyletus cacahuamilpensis Baker, Cheyletomimus congensis (Cunliffe) for Egypt (Abdelgayed et al., 2020), Cheyletus cacahuamilpensis - for Brazil (Pellegrini & Ferreira, 2013)
Ornithocheyletia lukoschusi Smiley, O. pingus Smiley, O. hallae Smiley, O. gersoni Smiley – new for Egypt (Halawa, Osman & Ahmed, 2019)
Samsinakia trilobites Bochkov et Mironov - new for Iran (Kazemi & Pektinat-Saeij, 2021)
Fam. Demodecidae – Pormann et al., 2021
Fam. Harpyrhynchidae
Harpyrhynchoides [Harpyrhynchus] vercammeni (Lawrence) – new for Egypt (Halawa, Osman & Ahmed, 2019)
Fam. Syringophilidae
Syringophiloidus minor (Berlese) – new for Egypt (Halawa, Osman & Ahmed, 2019)
Bubophilus aegolius Skoracki, Kosicki et Kwiecinski
Bubophilus aegolius Skoracki, Kosicki & Kwiecinski, 2021: 355
Type locality: Poland, Zielenogora, Stary Kisielin
Distribution: Poland, Finland, Switzerland
Host: Aegolius funereus (Strigiformes: Strigidae)
ASIA
India (Modak et al., 2004; Gupta et al, 2004; A. Gupta et al., 2006; Gupta & Ghosh, 1980) – Chelacaropsis moorei, Bal deleoni
Iran (Kazemi & Paktinat-Saeij, 2021) – Samsinakia trilobitus
AFRICA
Egypt (Abdelgayed et al., 2020; Atwa et al., 2018; Halawa et al., 2019) – Cheyletus cacahuamilpensis, Cheyletomimus congensis, Harpyrhynchoides [Harpyrhynchus] vercammeni, Ornithocheyletia lukoschusi, O. pingus, O. hallae, O. gersoni, Syringophiloidus minor
SOUTH AMERICA
Brazil (Pellegrini & Ferreira, 2013) - Cheyletus cacahuamilpensis
Aves
Columbiformes
Fam. Columbidae
Streptopelia senegalensis aegyptiaca - Harpyrhynchoides [Harpyrhynchus] vercammeni, Ornithocheyletia lukoschusi, O. hallae, O. gersoni, O. pinguis
Columba livia domestica - Ornithocheyletia lukoschusi
Passeriformes
Fam. Passeridae
Passer domesticus niloticus - Syringophiloidus minor
Strigiformes
Fam. Stigidae
Aegolius funereus – Bubophilus aegolius
(suppl. 2020 – 2021 and ommited)
Beron P. 2021. Acarorum Catalogus. IX. Schizoglyphoidea (Schizoglyphidae) Histiostomatoidea (Histiostomatidae, Guanolichidae) Canestrinioidea (Canestriniidae, Chetochelacaridae, Lophonotacaridae, Heterocoptidae) Hemisarcoptoidea (Chaetodactylidae, Hyadesiidae, Algophagidae, Hemisarcoptidae, Carpoglyphidae, Winterschmidtiidae). Pensoft & Nat. Mus. Natur. Hist. Sofia: 396 p.
The recent article of R. Haitlinger altered drastically the system of fam. Canestriniidae with the description of 9 new genera and the change of the taxonomy of 24 species.
The new genera are: Balearella, (Ibizella Haitlinger et Šundić, 2016, syn.), Cercediphotia, Ciprusenia, Ismaeilienia, Javanellina, Kuramansenia, Midiphotia, Phalerotusellus, Sumatranella.
24 species are transferred to another or new genera: Balearella asilahica, B. balearica, B. samsinaki, Ciprusenia carabicola, C. izabelae, C. kamelskyi, C. sellnicki, C. viviannae, Imaielienia egypti, I.maroccana, Javanellina macgillavryi, Kuramasenia illegalis, K. paavoi, K. pekingensis, K. pictura, K. ramoni, Midiphotia hispanica, M. jureceki, Phaleratusellus alatus, Psedocanestrinia damiani, P. roberti, Sumatranella batocerae, S. elae and S. phoebinae.
As a result of the recent contributions, the number of genera and species in this volume (some of them uncertain status) is updated as follows:
Schizoglyphidae – 1 genus, 1 species
Histiostomatidae – 58 genera, 522 species
Guanolichidae – 3 genera, 4 species
Canestriniidae – 100 genera, 348 species
Chetochelacaridae – 1 genus, 1 species
Lophonotacaridae – 1 genus, 1 species
Heterocoptidae – 12 genera, 28 species
Chaetodactylidae – 5 genera, 110 species
Hyadesiidae – 2 genera, 47 species
Algophagidae – 8 genera, 22 species
Hemisarcoptidae – 8 genera, 24 species
Carpoglyphidae – 4 genera, 10 species
Winterschmidtiidae – 25 genera, 138 species
Vol. I (2008) – Erythraeidae, Smarididae, Calyptostommatidae – 3 fam., 68 genera, 959 sp.
II (2011) – Listrophoroidea - 5 fam., 100 genera, 925 sp.
III (2014) - Opilioacarida, Holothyrida, Mesostigmata - 13 fam., 85 genera, 422 sp.
IV (2015) – Myobiidae, Pterygosomatidae - 2 fam., 68 genera, 707 sp.
V (2015) – Sarcoptoidea, Analgoidea, 19 fam., 145 genera, 509 sp.
VI (2020) – Rhinonyssidae, Spinturnicidae - 2 fam., 28 genera, 568 sp.
VII (2020) - Raphignathoidea - 12 fam., 68 genera, 1113 sp.
VIII (2020) – Cheyletoidea, Cloacaroidea - 13 fam., 177 genera, 1243 sp.
IX (2021) – Schizoglyphoidea, Histiostomatoidea, Canestrinioidea, Hemisarcoptoidea - 13 fam., 233 genera, 1240 sp.
X (2022) - Labidostommatoidea, Eupodoidea, Tydeoidea, Paratydeoidea, Anystoidea, Caeculoidea, Adamystoidea, Pomerantzioidea – 20 fam., 230 genera, 967 sp. Supplements: 1 fam., 11 genera, 149 sp. Added: 9 new genera, described by Haitlinger (2022).
Total: 102 fam., 1222 genera, 8815 sp.
Our gratitude is due to many specialists, sharing their experience, providing the nessessary publications or material. First of all we thank the arachnologists A.Fain, Fan Quing-Hai, A. Bochkov, Z. Feider, A. Khaustov, M. Bertrand, M.Shiba, C.Willmann, H.E.Vistorin, K.Schmölzer, P.Robaux, J.Cooreman, M. Zacharda, M.K.P.Meyer, H.Franz, I.Coineau, R.W.Strandtmann, B.Halliday, E.A.Ueckermann. For the updating of the previous volumes the author is grateful to R. Haitlinger, M. Šundiċ, A. Saboori, Si-Yuan Xu, R. Bassini-Silva, G. Gabryś, A. Bochkov, B. OConnor, M. Skoracki, M.M. Vázquez, H. Klompen, M.S. Araújo, M.V. Orlova, M. Fajfer, M. Akyol, Salih Doğan, S. Doğan, M. Zmudzinski, E.A.Ueckermann, A. Khaustov, Fan Quing-Hai, L.F.d.O. Bernardi, J. Noei, M. Hakimitabar, A. Labrzycka, J. Makol, P.B. Klimov, I. Dimov.