ACARORUM CATALOGUS VI. Order Mesostigmata. Gamasina: Dermanyssoidea (Rhinonyssidae, Spinturnicidae)

Petar Beron

doi:10.3897/ab.e54206

Petar Beron^‡

‡ Unaffiliated, Sofia, Bulgaria

Corresponding author: Petar Beron ( beron@mail.bg )

This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.

Citation: Beron P (2020) ACARORUM CATALOGUS VI. Order Mesostigmata. Gamasina: Dermanyssoidea (Rhinonyssidae, Spinturnicidae). Advanced Books. https://doi.org/10.3897/ab.e54206

Superorder Acariformes

This volume contains data on two families, 16 genera and 445 species of the families Rhinonyssidae (parasites in the respiratory system of birds) and 12 genera and 110 species of Spinturnicidae (ectoparasites of bats).

Order Mesostigmata G. Canestrini, 1891

According to Lindquist, Krantz & Walter (2009), Order Mesostigmata is composed of three suborders: Sejida, Trigynaspida and Monogynaspida with 26 superfamilies and about 12000 described species.

Suborder Monogynaspida Camin et Gorirossi, 1955

According to Beaulieu et al. (2011), the suborder consists of the infraorders Uropodina Kramer, 1881 and Gamasina Kramer, 1881, as well as of 15 superfamilies and 78 families. In Lindquist, Krantz & Walter (2009) the superfamilies are 18.

Infraorder (or Cohort ?) Gamasina Kramer, 1881

Cohort Gamasina of Lindquist, Krantz & Walter (2009) includes 10 superfamilies. Here belong most of the Mesostigmata, both predators and parasites.

Within Gamasina are delimitated 11 superfamilies and 42 families. Superfamily Dermanyssoidea remains within the Hyporder Dermanyssiae Evans et Till, 1979, with 17 families.

Beaulieu et al. 1911. Superorder Parasitiformes Reuter, 1909. In: Zhang Z.-Q. (Ed.) Animal biodiversity. An outline of higher-level classification and survey of taxonomic richness. – Zootaxa, 3148: 123–128.

Lindquist, Krantz & Walter 2009. Mesostigmata. In Krantz & Walter (Eds)., Manuel of Acarology, III Edition.

Superfamily Dermanyssoidea Kolenati

Dermanyssoidea Kolenati, 1859

After Krantz & Walter (2009, Eds), 14 families of parasitic mites form this superfamily, after the latest list of Parasitiformes families (Beaulieu et al., 1911) – 17. In this volume are considered the families Rhinonyssidae and Spinturnicidae.

The Mite Family Rhinonyssidae

Almost all Rhinonyssidae live in the nare passages of birds. One (Sternostoma tracheacolum Lawrence) has been found in the tracheae and lungs of birds and some Ptilonyssus have been recorded as living on the bird’s body or even in the nests.

N. Bregetova reviewed in 1951 the most numerous and common mites living in the respiratory system of birds – the Rhinonyssidae. By this time she was able to enumerate only 26 species in 10 genera, six of which are still valid: Rhinonyssus Trouessart, Sternostoma Berlese et Trouessart, Ptilonyssus Berl. et Tr., Rhinoecius Cooreman, Larinyssus Strandtmann and Rallinyssus Strandtmann. Four other generic names have been synonymized since (Sternostomum, Neonyssus, Ptilonyssoides and Rhinonyssoides). This scarce information was contributed mainly by Giebel (who described the first rhinonyssid in 1871), Trouessart, Berlese, Trägårdh, Hirst, Vitzthum, Castro, Cooreman, Strandtmann and Bregetova. Trouessart (1895) considered these mites as subfamily Rhinonyssinae and Vitzthum (1935) raised them in a family. However, some authors (Domrow) have longtime maintained the broader concept of Dermanyssidae and considered Rhinonyssinae as one of it’s subfamilies. Bregetova has splited these mites into two families – Rhinonyssidae and Ptilonyssidae, which was not accepted aside from few authors (Dusbábek & Černý, 1970).

Speaking of the “prehistory” of the studies on Rhinonyssidae we must notice the work of Brazilian specialists Castro & Pereira (1947), Castro (1948) and Pereira & Castro (1949) describing the subfamily Ptilonyssinae and several new species.

As for many other families, the “prehistory” of the research ends with the beginning of the long series of publications of Alex Fain (from 1956 to 1974). Based on his large collection, obtained in Africa, and on many mites from zoo birds, Fain (together with Aitken, Bafort, Bastin, Carpenter, Holland, Hyland, Johnson, Lukoschus, Nadchatram, Sixl, Moritsch, Vercammen-Grandjean) contributed greatly to the modern system of Rhinonyssidae, describing many new genera, species and subdividing the family into subfamilies. In the same time active research was carried out in Australia by R. Domrow (from 1964 to 1992). Domrow synonymised many older genera (Mesonyssus Fain, Mesonyssoides Fain et Nadchatram, Psitanyssus Fain, Falconyssus Fain). Some other genera, described by Castro (1948), were made synonyms by Zumpt & Till (1955) in their important contribution to the Rhinonyssidae of Southern Africa (Flavionyssus, Rochanyssus, Paraneonyssus, Travanyssus, Vitznyssus). Also important contributions to the study of Rhinonyssidae were made by Butenko (many publications on Russian and other mites since 1961, including a book in 1984, partly with Stanjukovich and Lavrovskaya), Shumilo & Lunkashu (incl. a book in 1976), Strandtmann and co-authors (Brooks, Clifford, Furman, Wharton), Knee (2008, together with Proctor and Galloway), Pence (1972–1979, together with Casto, Young and Canaris), Bregetova (from 1950 to 1983), Hyland (1961–1970, partly with Clark, Ford and Galloway), Kadosaka, Kaneko & Asanuma (1987), Feider & Mironescu (1972–1982), Dusbábek (since 1969), Amaral (1962–1978), Spicer (since 1977), George (1961), Furman (1957), Wilson (from 1958 to 1980), Sakakibara (1967, 1968), Crossley (1950, 1952), Grétillat (1969, 1970), Sixl (1969–1971), Sartbaev (1979) and others. Many new studies have been made in Spain by Guevara, Úbeda, Rodriguez, Rojas and other acarologists.

The Studies on Rhinonyssidae

Berlese & Trouessart (1889) describe the important genera Ptilonyssus and Sternostoma, containing more than half of all Rhinonyssidae

Trouessart (1894) – becomes the author of another important genus – Rhinonyssus

Trouessart (1895) describes the family Rhinonyssidae

Between 1895 and the Word War II very few authors study species (no new genera) of Rhinonyssidae: Trägårdh (1904), Vitzthum (1935), Hirst (1921–1923)

The post-war period is very rich in research on endoparasitic acari, incl. Rhinonyssidae. The following authors should be noticed (some of them still working on these mites):

J. Cooreman (1946) – describes the new genus Rhinoecius – start of research on the Rhinonyssidae on owls.

In Brazil M. Castro and C. Pereira try to restructure the family Rhinonyssidae, describing the subfamily Ptilonyssinae and several new species.

In the same time in the United States several researchers start working on Rhinonyssidae. Most prominent among them is R.W. Strandtmann (papers on Rhinonyssidae from 1948 to 1962, partly with C. Clifford, D. Furman, W. Wharton and others). The publications of Strandtmann contributed greatly to the knowledge on these mites. He described many new species and the genera Larinyssus and Rallinyssus.

D.A. Crossley – publications in 1950 and 1952 with descriptions of new mites from pigeons.

The great Belgian Acarologist A. Fain (1912–2009) started publishing on nasal mites in 1956, based on his large collection from Rwanda and Congo. He reorganysed the system of Rhinonyssidae, described (partly together with Aitken, Bafort, Bastin, Carpentier, Herin, Pulyert, Holland, Hyland, Johnston, Lukoschus, Mortelmans, Nadchatram, Sixl, and Vercammen-Grandjean) the new subfamily Rhinoeciinae, five valid new genera, 174 valid new species and several subspecies from many countries.

N. Wilson published papers on Rhinonyssidae from Antarctic and Subantarctic birds, birds of the Philippines, New Guinea, Taiwan, Thailand, Alaska and other Pacific areas (1958–1980). Also from this area many species have been described by I. Sakakibara and by E.W. Baker and M. Delfinado. The nasal mites of Japan were studied by T. Kadosaka and other researches.

Russian school in the study of Rhinonyssidae started with the thesis of Belopolskaya (1951) and is based mainly on the many papers of N. Bregetova and O. Butenko (with K.I. Lavrovskaya, M. Stanyukovich and other co-authors). Butenko worked in the reserve of Oka and described many new taxa of Rhinonyssidae. N. Bregetova considered these mites as being two families – Rhinonyssidae and Ptilonyssidae, what was not accepted but by few authors (Dusbábek & Černý, 1970). She published from 1950 to 1983 10 papers containing data on Rhinonyssidae, including a monograph on Gamasida and attempts to clarify the origin of these mites. S.K. Sartbaev published several papers on Rhinonyssidae of Central Asia from 1961 to 1983. On the Rhinonyssidae of Asia published several papers also T.T. Isakova, D.S. Ivanov and others. On Modavian mites worked R.P. Shumilo and M.I. Lunkashu.

In South Africa F. Zumpt and W.M. Till published in 1955 and 1961 important contribution on Rhinonyssidae, with descriptions of nine new species and a list of African Rhinonyssidae

D.P. Furman – in 1957 published comprehensive revision of genus Sternostoma

S. Gretillat – in 1961 described two new species and in 1959, together with A. Capron and E.R. Brygoo, described new nasal mites from Madagascar. The taxa, created by them (Agapornyssinae subfam. nov., Agapornyssus gen. nov. and Agapornyssus faini sp.n.) were synonymes.

H. G. Ford – in 1961 described Ptilonyssus constrictus

Z. Feider and I. Mironescu described from 1962 to 1982 several new species from Romania and Papua New Guinea.

K.E. Hyland – in series of papers (1959–1970)(patly with H.G. Ford, T.D. Clark and A. Moorhouse) described many new taxa from Mexico and USA

F. Dusbábek and V. Černý – studies of Cuban Rhinonyssidae (1969, 1970)

The series of papers (1972–1979) of D.B. Pence contain important data on the nasal mites from Louisiana, but also of other states.

P. Beron (1975) published the only paper on Bulgarian Rhinonyssidae. W. Sixl published papers on Rhinonyssidae from Austria.

V. Amaral took over the Brazilian school of research on nasal mites and from 1962 to 1974 published many new species and comprehensive list of Rhinonyssidae. This studies have been followed by C.S. Mascarenhas, D.S. Silva and other Brasilian researchers.

In Australia R. Domrow wrote from 1964 to 1992 many papers containing data on Rhinonyssidae of Australia and New Zealand. He reformed the system of these mites, put in synonymy many species and several genera (Sternostomoides, Rhinosterna, Sternoecius, Mesonyssus, Mesonyssoides, Psittanyssus, Falconyssus).

G.S. Spicer published between 1977 and 1987 several papers on the Rhinonyssidae of Guatemala and the USA.

W. Knee, H. Proctor, T. Galloway are publishing actively on the nasal mites of Canada.

I. Dimov published descriptions of several new species from Russia and a monograph (2018) concerning the Rhinonyssidae of birds in the NW of Russia.

The Composition of Rhinonyssidae

Fam. Rhinonyssidae Trouessart, 1894 – 16 gen., 445 sp.

Subfam. Rhinoeciinae Fain, 1957 – 3 gen., 16 sp.

Rhinoecius Cooreman, 1946 – 12 sp.

Ruandanyssus Fain, 1957 – 3 sp.

Zumptnyssus Fain, 1959 – 1 sp.

Subfam. Rhinonyssinae Trouessart, 1895 – 4 gen., 51 sp.

Charadrinyssus Butenko, 1984 – 1 sp.

Larinyssus Strandtmann, 1948 – 5 sp.

Rallinyssus Strandtmann, 1948 – 12 sp.

Rhinonyssus Trouessart, 1894 – 33 sp.

Subfam. Ptilonyssinae (Castro, 1948) Fain, 1957 – 9 gen., 378 sp.

Locustellonyssus Bregetova, 1965 – 2 sp.

Passeronyssus Fain, 1960 – 5 sp.

Pipronyssus Fain et Aitken, 1967 – 1 sp.

Ptilonyssus Berlese et Trouessart, 1889 – 215 sp.

Sternostoma Berlese et Trouessart, 1889 (incl. Rhinosterna Fain, 1964; Sternoecius Fain et Aitken, 1967; Sternostomoides Bregetova, 1965) – 75 sp.

Tinaminyssus Strandtmann et Wharton, 1958 – 68 sp.

Trochilonyssus Fain et Aitken, 1967 – 1 sp.

Tyranninyssus Brooks et Strandtmann, 1960 – 4 sp.

Vitznyssus Castro, 1948 (Astridiella Fain, 1957) – 9 sp.

Catalogue of Rhinonyssidae

Family Rhinonyssidae Trouessart

Rhinonyssinae Trouessart, 1895

Rhinonyssidae Trouessart: Vitzthum, 1935

Type genus: Rhinonyssus Trouessart, 1894

Ptilonyssidae Castro, 1948

Neonyssidae Bregetova, 1964

Subfamily Rhinoeciinae Fain

Rhinoeciinae Fain, 1957a: 156

Type genus: Rhinoecius Cooreman, 1946

Genus Rhinoecius Cooreman

Rhinoecius Cooreman, 1946: 1

Type species: Rhinoecius oti Cooreman, 1946

Rhinoecius aegolii Butenko

Rhinoecius aegolii Butenko, 1971c: 204; 1984: 131; Philips, 1993; 2000: 225; Knee, Proctor & Galloway, 2008: 370; Knee & Galloway, 2017: 96

Type locality: Okskiy Zapovednik, Russia

Distribution: Canada, Russia

Hosts: Aegolius acadicus, A. funereus, Surnia ulula (Strigiformes: Strigidae)

Rhinoecius africanus (Zumpt et Patterson)

Rhinonyssus africanus Zumpt & Patterson, 1951: 90

Rhinoecius africanus (Zumpt et Patterson): Zumpt & Patterson, 1955: 82; Fain, 1957b: 130; 1958b: 294

Type locality: Pietermaritzburg, Natal

Distribution: South Africa

Host: Asio capensis (Strigiformes: Strigidae)

Rhinoecius alifanovi Butenko

Rhinoecius alifanovi Butenko, 1976: 306, 1984: 127; Knee, Proctor & Galloway, 2008: 370

Type locality: Omskaya Obl., Krutinskiy Rayon; Ryazanskaya Obl.

Distribution: Canada, Russia

Host: Asio flammeus (Strigiformes: Strigidae)

Rhinoecius bisetosus Strandtmann

Rhinoecius bisetosus Strandtmann, 1952: 212; 1960: 132; Fain, 1957b: 130; Amaral, 1962b; Úbeda, Rodríguez & Guevara, 2000

Type locality: Lubbock Co., Texas

Distribution: Spain, Thailand, USA (Texas)

Hosts: Athene noctua, Glaucidium cuculoides, G. c. brugeli, Speotyto cunicularia, S. cunicularia grallaria (Strigiformes: Strigidae)

Rhinoecius brikinboricus Butenko

Rhinoecius brikinboricus Butenko, 1976: 305; 1984: 124; Knee, Proctor & Galloway, 2008: 370; Knee & Galloway, 2017: 96

Type locality: Ryazanskaya Obl., Brikin Bor

Distribution: Canada, Russia

Host: Asio otus (Strigiformes: Strigidae)

Rhinoecius cavannus Wilson

Rhinoecius cavannus Wilson, 1968c: 143; McClure, Ratanaworadhan et al., 1973: 37; Butenko, 1984: 130

Type locality: Chiengmai Province, Doi Inthanon

Distribution: Thailand

Host: Otus bakkamoena (Strigiformes: Strigidae)

Rhinoecius cooremani Strandtmann

Rhinoecius cooremani Strandtmann, 1952: 208; Fain, 1957b: 130; Strandtmann & Wharton, 1958: 165; Domrow, 1969b: 333, 423; Pence, 1973a: 359; Knee, Proctor & Galloway, 2008: 370; Knee & Galloway, 2017: 96

Type locality: Luling, Texas

Distribution: Australia, Canada, USA (La., Texas)

Hosts: Ninox connivens, N. novaeseelandiae, Strix nebulosa, S. varia (Strigiformes: Strigidae)

Rhinoecius grandis Strandtmann

Rhinoecius grandis Strandtmann, 1952: 205; Fain, 1957b: 130; Knee, Proctor & Galloway, 2008: 370

Type locality: Lubbock Co., Texas

Distribution: Brazil, Canada, USA (Texas)

Hosts: Bubo virginianus, Pulsatrix melanonota koeniswaldiana (Strigiformes: Strigidae)

Rhinoecius nycteae Butenko

Rhinoecius nycteae Butenko, 1976: 307; 1984: 133; Knee, Proctor & Galloway, 2008: 370

Type locality: Omskaya Obl., Cherlakskiy rayon

Distribution: Canada, Russia

Host: Bubo scandiacus [= Nyctea scandiaca] (Strigiformes: Strigidae)

Rhinoecius oti Cooreman

Rhinoecius oti Cooreman, 1946: 1; Pereira & Castro, 1949: 233; Zumpt & Patterson, 1951: 77; Zumpt & Till, 1955: 82; Fain, 1957b: 130; Butenko, 1984: 124

Type locality: Schooten (Belgium)

Distribution: Belgium

Host: Asio otus otus (Strigiformes: Strigidae)

Rhinoecius subbisetosus Bregetova

Rhinoecius subbisetosus Bregetova, 1965: 214; Butenko, 1984: 130

Type locality: Badhiz, Akar-Cheshme

Distribution: Turkmenistan

Host: Athene noctua (Strigiformes: Strigidae)

Rhinoecius tytonis Fain

Rhinoecius tytonis Fain, 1956b: 394; Fain, 1957b: 131, 138; 1959a: 33; Butenko, 1984: 132

Type locality: Akanyaru (Rwanda)

Distribution: Rwanda, South Africa

Hosts: Tyto alba, T. alba affinis (Strigiformes: Tytonidae)

Genus Ruandanyssus Fain

Ruandanyssus Fain, 1957a: 148

Type species: Ruandanyssus terpsiphonei Fain, 1957

Ruandanyssus artami Domrow

Ruandanyssus sp., Domrow, 1967a: 217

Ruandanyssus artami Domrow, 1969b: 330

Type locality: Kowanyama

Distribution: Australia

Hosts: Artamus cinereus, A. minor (Passeriformes: Artamidae)

Ruandanyssus strandmanni Sakakibara

Ruandanyssus strandmanni Sakakibara, 1968a: 13

Type locality: Enarotali

Distribution: New Guinea

Host: ?Neositta sp. (Passeriformes: Neosittidae)

Ruandanyssus terpsiphonei Fain

Ruandanyssus terpsiphonei Fain, 1957a: 148; 1957b: 131; Strandtmann & Wharton, 1958: 165; Domrow, 1965b: 432; 1967a: 217; 1969b: 329; 1979d: 439; Kadosaka, Kaneko & Asanuma, 1987: 40

Ruandanyssus terpsiphonei echongi Sakakibara, 1968: 15 – type: Enarotali, New Guinea, from Pachycephala (syn. by Domrow, 1969)

Type locality: Astrida (= Butare), Akanyaru (Rwanda)

Distribution: Australia, Japan, New Guinea, Rwanda

Hosts: Artamus personatus (Passeriformes: Artamidae), Coracina novaehollandiae, C. robusta, Pericrocotus roseus divaricatus (Passeriformes: Campephagidae), Lalage tricolor (Passeriformes: Corvidae), Pachycephala griseiceps, P. pectoralis, P. rufiventris (Passeriformes: Pachycephalidae), Sericornis frontalis, S. keri (Passeriformes: Acanthizidae), Struthidea cinerea (Passeriformes: Corcoracidae), Monarcha trivirgata, Myiagra rubecula, Terpsiphone viridis kivuensis (Passeriformes: Monarchidae)

Genus Zumptnyssus Fain

Zumptnyssus Fain, 1959c: 112

Type species: Ruandanyssus buboensis Fain, 1958

Zumptnyssus buboensis (Fain)

Ruandanyssus buboensis Fain, 1958b: 292

Zumptnyssus buboensis (Fain): Fain 1959c: 112; 1960: 113

Type locality: Lephepe (Botswana)

Distribution: Botswana, Mozambique

Host: Bubo africanus (Strigiformes: Strigidae)

Subfamily Rhinonyssinae Trouessart

Rhinonyssinae Trouessart, 1895 – Fain, 1957a: 154

Type genus: Rhinonyssus Trouessart, 1894

Genus Charadrinyssus Butenko

Charadrinyssus Butenko, 1984: 74

Type species: Larinyssus charadrius Butenko, 1984

Charadrinyssus charadrius (Butenko)

Larinyssus charadrius Butenko, 1975b: 951

Charadrinyssus charadrius (Butenko): Butenko, 1984: 74

Type locality: Kurgaldjinskiy Rayon

Distribution: Kazakhstan

Host: Charadrius morinellus (Charadriiformes: Charadriidae)

Genus Larinyssus Strandtmann

Larinyssus Strandtmann, 1948: 128

Type species: Larinyssus orbicularis Strandtmann, 1948

Larinyssus benoiti Fain

Larinyssus benoiti Fain, 1961: 128; 1964: 134; Domrow, 1966: 196; 1969: 322; 1979d: 492; 1988: 864; Sartbaev, 1979: 56; Butenko, 1984: 72

Type locality: Kasongo, DR Congo

Distribution: Australia, DR Congo, Kirghizstan, Russia

Hosts: Galachrysia cinerea, Glareola nordmanni, G. pratincola, Stiltia isabella (Charadriiformes: Glareolidae)

Larinyssus iohanssenae Dimov

Larinyssus iohanssenae Dimov, 2013: 123; 2018: 167

Type locality: Russia

Distribution: Russia

Host: Sterna hirundo (Charadriiformes: Sternidae)

Larinyssus orbicularis Strandtmann

Larinyssus orbicularis Strandtmann, 1948: 507; Pereira & Castro, 1949: 232; Bregetova, 1951: 117; Zumpt & Paterson, 1951: 78; Zumpt & Till, 1955: 68; Fain, 1956b: 157; 1957b: 60; 1958: 9; 1960c: 95; Černy & Dusbábek, 1970: 487; Pence, 1972a: 160; 1975: 21; Domrow, 1967a: 216; 1969: 322; 1973: 84; 1988: 865; Butenko, 1984: 68; Dimov, 2016; 2018: 168

Larinyssus petiti Gretillat, 1961 – syn. by Domrow, 1969: 322

Type locality of L. orbicularis: Galveston, Texas

Distribution: Australia, Brazil, DR Congo, Cuba, Russia, Rwanda, South Africa, USA, Zambia

Hosts: Anous minutus, Chlidonias leucoptera, Ch. nigra, Ch. orbicularis, Gelochelidon nilotica, Sterna fuscata, S. hirundinacea, S. hirundo, S. maxima, Thalasseus acuflavidus, Th. bergii (Charadriiformes: Sternidae), Larus argentatus, L. atricilla, L. delawarensis, L. dominicanus, L. minutus, L. novaehollandiae (Charadriiformes: Laridae), Rynchops nigra (Charadriiformes: Rynchopidae)

Larinyssus sterna Fain et Holland

Larinyssus sterna Fain & Holland, 1972: 30

Type locality: Aves Is., Venezuela

Distribution: Venezuela

Host: Onychoprion [Sterna] fuscatus fuscatus (Charadriiformes: Sternidae)

Larinyssus substerna Butenko

Larinyssus substerna Butenko, 1975: 180; 1984: 71

Type locality: Tomskaya Obl., Kolominskiy Rayon (Russia)

Distribution: Kazakhstan, Kirghizstan, Russia

Host: Sterna hirundo (Charadriiformes: Sternidae)

Larinyssus sp

Distribution: Brazil

Host: Larus dominicanus (Charadriiformes: Laridae)

Genus Rallinyssus Strandtmann

Rallinyssus Strandtmann, 1948: 512

Type species: Rallinyssus caudistigmus Strandtmann, 1948

Rallinyssoides Fain, 1960a: 295 – syn. by Wilson, 1965: 624

Type species: Rallinyssus congolensis Fain, 1956

Rallinyssus amaurornis Wilson

Rallinyssus amaurornis Wilson, 1965: 624; 1966; McClure, Ratanaworadhan et al., 1973: 37

Type locality: Nan t’ou Hsien, Puli (Taiwan)

Distribution: Philippines, Taiwan

Hosts: Amaurornis phoenicurus chinensis, Poliolimnas cinereus collingwoodi (Gruiformes: Rallidae)

Rallinyssus caudistigmus Strandtmann

Rallinyssus caudistigmus Strandtmann, 1948: 512; Bregetova, 1951: 118; 1956: 198; Zumpt & Till, 1955: 68; Strandtmann & Wharton, 1958: 160; Fain, 1957: 58; 1959a: 33; Domrow, 1966b: 196; 1969b: 324; Feider & Mironescu, 1973: 159; Fain, Sixl & Moritsch, 1974: 7; Pence, 1972; 1975: 42; Guevara-Benítez & Úbeda-Ontíveros, 1975a: 492; Butenko, 1976; 1984: 78; Sartbaev, 1977; 1979: 56; Kadosaka, Kaneko & Asanuma, 1987: 39; Knee, Proctor & Galloway, 2008: 368; Knee & Galloway, 2017: 93; Kun & Vega, 2016: 97; Silva et al., 2018; Dimov, 2018: 184

Type locality: Texas

Rallinyssus caspicus Butenko, 1976: 72; 1984: 81; Sartbaev, 1979: 56 – type: Kizil Agachskiy Zapovednik (Azerbaidjan), from Gallinula chloropus. Syn. by Domrow, 1979: 513

Distribution: Argentina, Australia, Azerbaijan, Austria, Canada, Japan, Kazakhstan, Kirghizstan, Romania, Russia, South Africa, Spain, Turkmenistan, Ukraine, USA (Texas)

Hosts: Fulica americana, F. armillata, F. atra, F. cristata, Gallinula chloropus, G. chloropus indica, G. tenebrosa, Rallus elegans (Gruiformes: Rallidae)

Rallinyssus congolensis Fain

Rallinyssus congolensis Fain, 1956b: 396; 1957b: 58; Strandtmann & Wharton, 1958: 160; Domrow, 1973: 84

Rallinyssoides congolensis (Fain): Fain, 1960a: 624

Rallinyssus porzana congolensis Fain: McClure, Ratanaworadhan et al., 1973: 37

Type locality: Akanyaru

Distribution: Australia (incl. Tasmania), DR Congo, Philippines

Host: Porzana tabuensis (Gruiformes: Rallidae)

Rallinyssus cychramus Wilson

Rallinyssus cychramus Wilson, 1965: 627; Butenko, 1984: 88

Type locality: PNG, Western Highlands Distr., Tambul, 2300 m

Distribution: Australia, DR Congo, Papua New Guinea

Hosts: Porzana parva, P. pusilla, P. tabuensis (Gruiformes: Rallidae)

Rallinyssus gallinulae Fain

Rallinyssus gallinulae Fain, 1960a: 295; 1962: 265; Wilson, 1965: 634; 1968: 221; Domrow, 1965b: 450; 1969b: 323; 1972: 99; 1973: 84; Butenko, 1976: 74; 1984: 83

Rallinyssus rallus Wilson, 1965: 631 – syn. by Domrow, 1969: 323

Type locality: Anvers Zoo

Distribution: Australia (incl. Tasmania), Azerbaijan, Belgium, New Zealand

Hosts: Gallinula chloropus, Hypotaenidia philippensis, Porphyrio porphyrio (= melanotus), Rallina tricolor, Rallus pectoralis (Gruiformes: Rallidae)

Rallinyssus limnocoracis Fain

Rallinyssus limnocoracis Fain, 1956b: 397; McClure, Ratanaworadhan et al., 1973: 37

Rallinyssoides limnocoracis (Fain): Fain & Nadchatram, 1962: 271

Type locality: Akanyaru (Rwanda)

Distribution: Malaysia (Selangor), Rwanda

Hosts: Limnocorax flavirostris, Porzana f. fusca (Gruiformes: Rallidae)

Rallinyssus porzanae Wilson

Rallinyssus porzanae Wilson, 1967a: 215

Type locality: Nueva Viskaya, Luzon

Distribution: Philippine Is.

Host: Zapornia [Porzana] tabuensis filipina (Gruiformes: Rallidae)

Rallinyssus rallinae Wilson

Rallinyssus rallinae Wilson, 1966a: 220; McClure, Ratanaworadhan et al., 1973: 37

Type locality: Nueva Viskaya, Luzon

Distribution: Philippine Is.

Host: Rallina eurizonoides eurizonoides (Gruiformes: Rallidae)

Rallinyssus sorae Pence et Young

Rallinyssus sorae Pence & Young, 1979: 791; Knee & Galloway, 2017: 93

Type locality: Maryland, Ohio

Distribution: Canada, USA (Maryland, Ohio)

Host: Porzana carolina (Gruiformes: Rallidae)

Rallinyssus strandtmanni Gretillat

Rallinyssus strandtmanni Gretillat, 1961: 151; Wilson, 1965: 637;1966: 225; Butenko, 1984: 89

Rallinyssoides strandtmanni (Gretillat): Amaral, 1962: 189

Type locality: France

Distribution: France

Host: Gallinula chloropus (Gruiformes: Rallidae)

Rallinyssus trappi (Amaral)

Rallinyssoides trappi Amaral, 1962: 185; 1968: 123

Rallinyssus trappi (Amaral): Wilson, 1965: 637

Type locality: Itaporanga, Estado de Sâo Paulo

Distribution: Brazil

Host: Ortygonax nigricans (Gruiformes: Rallidae)

Rallinyssus verheyeni Fain

Rallinyssus verheyeni Fain, 1963a: 176; Wilson, 1966: 225; Pence, 1972a: 158; 1973: 884; 1975: 21; Sartbaev, 1979: 56; Butenko, 1984: 85

Type locality: Horion – Hozémont, Prov. Liège, Belgium

Distribution: Australia, Belgium, Kirghizstan, Russia, USA (La.)

Hosts: Podiceps furicollis, P. ruficollis (Podicipiformes: Podicipedidae), Rallus aquaticus, R. elegans, R. limicola (Gruiformes: Rallidae)

Genus Rhinonyssus Trouessart

Rhinonyssus Trouessart, 1894: 723

Type species: Rhinonyssus coniventris Trouessart, 1894

Sommatericola Trägårdh, 1904b: 28

Type species: Sommatericola levinseni Trägårdh, 1905

Rhinonyssus afribyx Fain

Rhinonyssus afribyx Fain, 1956a: 150; 1957b: 48

Type locality: Astrida [Butare], Rwanda

Distribution: DR Congo, Rwanda

Hosts: Vanellus [Afribyx] senegallus, Vanellus [Stephanibyx] lugubris (Charadriiformes: Charadriidae)

Rhinonyssus alberti Strandtmann

Rhinonyssus alberti Strandtmann, 1956: 140; Fain, 1957b: 48; Feider & Mironescu, 1973: 415; Butenko, 1965: 131; 1984: 47

Type locality: Longbeach, California

Distribution: Australia, Azerbajdjan, Kazakhstan, Romania, Russia, USA

Hosts: Podiceps auritus, P. nigricolis (= Colymbus caspicus), P. alberti, P. cristatus, P. griseigena, P. ruficollis (Podicipediformes: Podicipedidae)

Rhinonyssus apus Fain

Rhinonyssus apus Fain, 1957a: 151; 1957b: 48

Type locality: Akanyaru (Rwanda)

Distribution: Rwanda

Host: Apus caffer streubeli (Apodiformes: Apodidae)

Rhinonyssus belenopteri Fain

Rhinonyssus belenopteri Fain, 1964: 29

Type locality: Anvers Zoo (Chile)

Distribution: Chile

Host: Belenopterus chilensis (Charadriiformes: Charadriidae)

Rhinonyssus bregetovae Butenko

Rhinonyssus bregetovae Butenko, 1974: 464; 1984: 39; Dimov, 2016; 2018: 137

Type locality: Ryazanskaya Obl., Tomskaya Obl. (Russia), Tselinogradskaya Obl. (Kazakhstan)

Distribution: Kazakhstan, Russia

Host: Charadrius dubius (Charadriiformes: Charadriidae)

Rhinonyssus caledonicus Hirst

Rhinonyssus caledonicus Hirst, 1921: 357; Strandtmann, 1956: 135; 1959; Fain, 1957b: 45; 1960b: 312; Pence, 1975: 20; Butenko, 1984: 33; Dimov, 2018: 142

Sternostomum caledonicum: Vitzthum, 1935: 569; Bregetova, 1951: 116; 1956: 198; Turk, 1953: 13

Sternostoma caledonicus: Zumpt & Till, 1955: 90

Type locality: Glass Pt.

Distribution: Germany, Russia (incl. Far East, Komandor and other islands), Shetland Is., USA

Hosts: Aethia pusilla, Cepphus columbae, Ciceronia pusilla, Cerorhinca monocerata, Uria aalge, U. grylle, U. lomyia (Charadriiformes: Alcidae)

Rhinonyssus chettusiae Butenko

Rhinonyssus chettusiae Butenko, 1973: 1874; 1984: 42

Type locality: Kamýsh (Kazakhstan)

Distribution: Kazakhstan

Host: Chettusia gregaria (Charadriiformes: Charadriidae)

Rhinonyssus clangulae Butenko et Stanyukovich

Rhinonyssus clangulae Butenko & Stanyukovich, 2001: 522

Type locality: Nizhne-Kolymskiy Rayon, Yakutia

Distribution: Russia

Host: Clangula hyemalis (Anseriformes: Anatidae)

Rhinonyssus colymbicola Fain et Bafort

Rhinonyssus colymbicola Fain & Bafort, 1963: 471; Butenko, 1984: 50; Knee & Galloway, 2017: 95

Type locality: Damme (Belgium)

Distribution: Belgium, Canada, Russia

Hosts: Podiceps auritus, P. griseigena, P. nigricollis (Podicipediformes: Podicipedidae)

Rhinonyssus coniventris Trouessart

Rhinonyssus coniventris Trouessart, 1894: 724; Hirst, 1921: 361; Vitzthum, 1935: 572; Castro, 1948: 256; Strandtmann, 1951: 130; 1956: 130; Zumpt & Patterson, 1951: 78; Turk, 1953: 13; Zumpt & Till, 1955: 84; Fain, 1957b: 43; 1960: 312; 1963: 88; Strandtmann & Wharton, 1958: 157; Fain, 1963: 88; Domrow, 1965b: 450; 1967a: 216; 1969b: 327, 423; Černý & Dusbábek, 1970: 484

Type locality: Europe

Distribution: Australia, Belgium, Canada, Cuba, England, Shetland Islands, South Africa, USA (incl. Alaska and Hawaii Is.)

Hosts: Arquatella ptilocnemis, Arenaria interpres, Calidris canutus, C. maritima, Catophrophorus semipalmatus, Charadrius alexandrinus, Ch. hiaticula, Ch. marginatus, Crocethia alba, Rhyacophilus glareola, Totanus flavipes, Erolia ruficollis (Charadriiformes: Charadriidae), Tringa ochropus (Charadriiformes: Scolopacidae)

Rhinonyssus dobromiri Dimov et Spicer

Rhinonyssus dobromiri Dimov & Spicer, 2013: 292; Dimov, 2018: 133

Type locality: Leningrad Province

Distribution: Russia

Host: Vanellus vanellus (Charadriiformes: Charadriidae)

Rhinonyssus echinipes Hirst

Rhinonyssus echinipes Hirst, 1921: 359 – type: Shetland Islands, ex Charadrius hiaticula; Turk, 1953: 13; Vitzthum, 1935: 572; Castro, 1948: 256; Strandtmann, 1951: 132; Butenko, 1984: 28; de Rojas et al., 2007

Rhinonyssus coniventris echinipes: Fain, 1963: 93; Fain, 1963: 179; Černy & Dusbábek, 1970: 484; Amaral & Reboucas, 1974: 55

Type locality: Shetland Islands

Distribution: Belgium, Shetland Islands, Spain, Turkmenistan

Hosts: Charadrius alexandrinus, Ch. hiaticula, Ch. melodus, Ch. wilsonia, Pluvialis squatarola (Charadriiformes: Charadriidae)

Rhinonyssus himantopus Strandtmann

Rhinonyssus himantopus Strandtmann, 1951: 136; 1956: 132; 1959: 134; Strandtmann & Wharton, 1958: 158; Fain, 1956a: 149; 1957b: 44; 1958c: 9; 1960c: 92; Domrow, 1966b: 195; 1969b: 326; 1979d: 493; Černy & Dusbábek, 1970: 484; Pence, 1972a: 157; Butenko, 1984: 46; Kadosaka, Kaneko & Asanuma, 1987

Type locality: Kleberg Co., Texas

Distribution: Australia, D.R. Congo, Cuba, France, Japan, USA, South Africa, Zambia

Hosts: Charadrius melanops, Ch. tricollaris, Erythrogonys cinctus; Lobibyx miles, L. novaehollandiae, Vanellus [sub syn. Hemiparra] crassirostris, V. spinosus (Charadriiformes: Charadriidae), Himantopus himantopus, H. leucocephalus, H. mexicanus, Recurvirostra americana (Charadriiformes: Recurvirostridae)

Rhinonyssus kadrae Dimov

Rhinonyssus kadrae Dimov, 2013а: 118; 2018: 134

Type locality: Leningrad Province

Distribution: Russia

Host: Anas platyrhynchos (Anseriformes: Anatidae)

Rhinonyssus levinseni (Trägårdh)

Sommatericola levinseni Trägårdh, 1904b: 28

Sternostomum levinseni: Vitzthum, 1935: 569; Bregetova, 1951: 114; 1956: 198; Domrow, 1969: 326

Sternostomum levinsi: erratum

Rhinonyssus levinseni (Trägårdh): Hirst, 1921a: 357; van Eyndhoven, 1964: 300; Butenko, 1984: 65; Dimov, 2016 ; 2018: 140

Type locality: Greenland

Distribution: Australia, Greenland, Holand, Russia

Hosts: Anas gibberifrons, A. platyrhynchos, A. querquedula, Anser anser, Polysticta stelleri, Somateria fischeri, S. molissima, S. spectabilis (Anseriformes: Anatidae)

Rhinonyssus marilae Butenko et Stanyukovich

Rhinonyssus marilae Butenko & Stanyukovich, 2001: 524

Type locality: Primorskiy Kray (Far East)

Distribution: Russia

Host: Aythya marila (Anseriformes: Anatidae)

Rhinonyssus minutus (Bregetova)

Sternostomum minutus Bregetova, 1950: 1007; 1951: 116; Fain, 1956: 150

Sternostoma minutus: Furman, 1957: 483

Rhinonyssus minutus: Fain, 1957b: 48;1961a: 514; Strandtmann & Wharton, 1958: 159; Domrow, 1965b: 450; 1967a: 216; 1969b: 326; Butenko, 1984: 36; Dimov, 2018: 141

Type locality: Russia

Distribution: Australia, Azerbaydzhan, Kirghizstan, Russia, Turkmenistan

Hosts: Charadrius alexandrinus, Ch. dubius, Ch. hiaticula (Charadriiformes: Charadriidae), Otis tarda (Otidiformes: Otitidae)

Rhinonyssus minutus leucopolius Fain

Rhinonyssus minutus leucopolius Fain, 1972: 363

Type locality: Anvers Zoo

Distribution: ?

Host: Charadrius [Leucopolius] pecuarius (Charadriiformes: Charadriidae)

Rhinonyssus neglectus Hirst

Rhinonyssus neglectus Hirst, 1921: 359; Vitzthum, 1935: 572; Castro, 1948: 257; Bregetova, 1951: 117; 1953: 332; Strandtmann, 1951: 132; Fain, 1957; Fain, 1960: 312; Domrow, 1969; Pence, 1975; Butenko, 1984: 23; Knee & Proctor, 2010; Dimov, 2016; 2018: 138

Rhinonyssus coniventris neglectus (Hirst): Fain, 1963: 91; Turk, 1953: 13

Type locality: Shetland Islands

Distribution: Australia, Azerbajdjan, Canada, Germany, Shetland Islands, Kazakhstan, Russia (incl. Maritime Distr.), Rwanda

Hosts: Arenaria interpres, Calidris alpina, C. cornutus, C. maritima, Charadrius alexandrinus, Tringa glareola, T. ochropus (Charadriiformes: Charadriidae)

Rhinonyssus pluvialis Fain et Johnston

Rhinonyssus pluvialis Fain & Johnston, 1966: 27

Type locality: 5 km S Wooster, Ohio

Distribution: USA (Ohio)

Host: Pluvialis dominica (Charadriiformes: Charadriidae)

Rhinonyssus podicipedus Feider et Mironescu

Rhinonyssus podicipedus Feider & Mironescu, 1972: 317; Butenko, 1984: 51

Rhinonyssus tschomgae Butenko, 1973: 244 – type: Azerbajdzhan, Kyzyl Agach (syn. by Butenko, 1984: 51)

Type locality: Sfintu Georghe (Tulcea)

Distribution: Azerbajdzhan, Romania, Russia, Uzbekistan

Host: Podiceps cristatus (Podicipiformes: Podicipedidae)

Rhinonyssus podilymbi Pence

Rhinonyssus podilymbi Pence, 1972a: 154

Type locality: Louisiana, Cameron Parish

Distribution: USA (La)

Host: Podilymbus podilymbi (Podicipediformes: Podicipedidae)

Rhinonyssus poliocephali Fain

Rhinonyssus poliocephali Fain, 1956a: 149; 1957b: 45; Strandtmann & Wharton, 1958: 159; Domrow, 1965b: 450; 1967a: 216; 1969b: 325

Rhinonyssus polycephali: Butenko, 1984: 53

Type locality: Astrida (= Butare), Rwanda

Distribution: Australia, Rwanda

Hosts: Podiceps [Poliocephalus] ruficollis capensis, P. ruficollis (Podicipiformes: Podicipedidae)

Rhinonyssus polystictae Butenko

Rhinonyssus polystictae Butenko, 1984: 67; Dimov, 2016; 2018: 139

Type locality: Yakutia, Kitisno

Distribution: Russia (Yakutia)

Hosts: Polysticta stelleri, Somateria mollissima (Anseriformes: Anatidae)

Rhinonyssus rhinolethrum (Trouessart)

Sternostomum rhinolethrum Trouessart, 1895: 393; Berlese, 1912: 71; Hirst, 1922: 89; Vitzthum, 1935; 1941: 656; Bregetova, 1951: 114

Rhinonyssus rhinolethrum: de Castro, 1948; Strandtmann, 1951: 132; 1956: 137; Strandtmann & Wharton, 1958: 159; Fain, 1958c: 9; 1960: 108; 1962b: 267; Mitchell & Rhodes, 1960: 107; Mitchell, 1963a: 506, 1963b; Domrow, 1969b: 324; Wilson, 1970: 74; Pence, 1972: 156; Butenko, 1984: 57; Mascarenhas et al., 2009: 295

Sternostoma rhinolethrum: Zumpt & Till, 1955: 90

Rhinonyssus rhinolethrus: Domrow, 1966: 196; 1967a: 216

Rhinonyssus anatidae Butenko, 1971: 490 – Russia (syn. by Domrow, 1979: 513)

Rhinonyssus rhinolethrum anatidae: Butenko, 1971: 490; 1984: 61

Rhinonyssus subrhinolethrum Butenko, 1971: 491; 1984: 62; Sartbaev, 1977; 1979: 56 – type: Spasskiy Rayon, Ryazanskaya Obl., Russia; Dimov, 2016; 2018: 135 (syn. by Domrow, 1979: 513)

Rhinonyssus nyrocae Butenko, 1971a: 492; 1984: 64; Dimov, 2016; 2018: 135 – type: Danube Delta, from Aythia spp. Syn. by Domrow, 1979: 513

Type locality: Europe

Distribution: Antarctica (South Georgia, etc.), Argentina, Australia, Azerbaijan, Belgium, Botswana, Brazil, Canada, D.R. Congo, Cuba, France, Greenland, Holland, Kazakhstan, Kirghizstan, Papua New Guinea, Russia, Ukraine, USA, Uzbekistan

Hosts: Aix sponsa, Alopochen aegyptiacus, Anas acuta, A. [Mareca] americana, A. carolinensis, A. clypeata, A. crecca, A. discors, A. erythrorhyncha, A. falcata, A. georgicus, A. gibberifrons, A. penelope, A. platyrhynchos, A. querquedula, A. strepera, A. superciliosa, Anser albifrons, A. anser, A. rossii, Aythia affinis, A. baeri, A. ferina, A. fuligula, A. nyroca, Branta canadensis, Callonetta leucophrys, Chenonetta jubata, Cygnus columbianus, C. melanocoryphus, Dendrocygna arcuata, D. bicolor, D. viduata, Metopiana peposaca, Netta rufina, Spatula clypeata, Tadorna tadorna (Anseriformes: Anatidae); Tringa nebularia (Charadriiformes: Scolopacidae)

Rhinonyssus shcherbininae Butenko

Rhinonyssus shcherbininae Butenko, 1984: 25

Type locality: Turkmenistan

Distribution: Kazakhstan, Kirghizstan, Turkmenistan

Host: Charadrius alexandrinus (Charadriiformes: Charadriidae)

Rhinonyssus sphenisci Fain et Mortelmans

Rhinonyssus sphenisci Fain & Mortelmans, 1959: 13; Gastal et al., 2018: 487

Type locality: Peru

Distribution: Brazil, Peru

Hosts: Spheniscus humboldti, S. magellanicus (Sphenisciformes: Spheniscidae)

Rhinonyssus sphenisci schelli Fain et Hyland

Rhinonyssus sphenisci schelli Fain & Hyland, 1963: 4

Rhinonyssus schelli Fain & Hyland: Wilson, 1970: 74; 1971: 17

Type locality: Base Roi Baudouin

Distribution: Antarctica

Hosts: Pygoscelis adeliae, P. papua (Sphenisciformes: Spheniscidae)

Rhinonyssus spinactitis Dusbábek

Rhinonyssus spinactitis Dusbábek, 1969: 213; Černy & Dusbábek, 1970: 485

Type locality: Canal de Vijil, La Gloria, Prov. de Camagüey

Distribution: Cuba

Host: Actitis macularia (Charadriiformes: Scolopacidae)

Rhinonyssus strandtmanni Fain et Johnson

Rhinonyssus strandtmanni Fain & Johnson, 1966: 25

Type locality: 5 km S of Wooster, Ohio

Distribution: Rwanda, USA (Ohio)

Hosts: Charadrius tricollaris, Ch. vociferus (Charadriiformes: Charadriidae)

Rhinonyssus tringae Fain

Rhinonyssus tringae Fain, 1963b: 96; Černy & Dusbábek, 1970: 485; Amaral & Reboucas, 1974: 55; Butenko, 1984: 31; de Rojas, Mora et al., 2002: 676; de Rojas et al., 2007: 1041

Type locality: Astrida (= Butare), Rwanda

Distribution: Cuba, Russia, Rwanda, Spain

Hosts: Tringa glareola, T. melanoleuca, T. ochropus, T. totanus (Charadriiformes: Scolopacidae)

Rhinonyssus vanellochettusiae Butenko

Rhinonyssus vanellochettusiae Butenko, 1984: 43

Type locality: Imam-Baba

Distribution: Turkmenistan

Host: Vanellochettusia leucura (Charadriiformes: Charadriidae)

Rhinonyssus vanellus Fain

Rhinonyssus afribyx vanellus Fain, 1972: 362

Type locality: Brasschaat nr Anvers, Belgium

Rhinonyssus vanellus Butenko, 1973: 1872; Sartbaev, 1979: 56

Type locality: Novosibirskaya Obl., Karasukskiy Rayon

Distribution: Belgium, Kazakhstan, Kirghizstan, Russia

Host: Vanellus vanellus (Charadriiformes: Charadriidae)

Rhinonyssus waterstoni Hirst

Rhinonyssus waterstoni Hirst, 1921: 359; Castro, 1948: 256; Strandtmann, 1956: 134; Fain, 1957: 44; Butenko, 1969: 208, 1984: 34; Dimov, 2018: 143

Sternostomum waterstoni: Vitzthum, 1935: 569; Bregetova, 1951: 116; 1956: 198; Turk, 1953: 13

Type locality: Shetland Is.

Distribution: Germany, Russia, Shetland Is., USA

Hosts: Alca torda, Uria aalge, U. lomyia (Charadriiformes: Alcidae)

Subfam. Ptilonyssinae (Castro) Fain

Ptilonyssinae (Castro, 1948) Fain, 1957: 155

Ptilonyssidae: Castro, 1948

Type genus: Ptilonyssus Berlese et Trouessart, 1889

Genus Passeronyssus Fain

Passeronyssus Fain, 1960: 110

Type species: Ptilonyssus viduae Fain, 1956

Passeronyssus kittacinclae Fain

Passeronyssus kittacinclae Fain, 1964: 300

Type locality: Zoo Anvers

Distribution: South Asia

Host: Kittacincla malabarica (Passeriformes: Muscicapidae)

Passeronyssus megaluri Sakakibara

Passeronyssus megaluri Sakakibara, 1968: 305

Type locality: West Sepik Distr., Telefomin

Distribution: New Guinea

Host: ? Megalurus timoriensis (Passeriformes: Megaluridae)

Passeronyssus urieli do Amaral

Passeronyssus urieli do Amaral, 1973: 68

Type locality: Cipó, Est de São Paulo

Distribution: Brazil

Host: Myiarchus ferox australis (Passeriformes: Tyrannidae)

Passeronyssus viduae (Fain)

Ptilonyssus viduae Fain, 1956a: 147

Passeronyssus viduae (Fain): Fain, 1960: 110

Type locality: Astrida (= Butare)

Distribution: Rwanda

Host: Vidua macroura (Passeriformes: Viduidae)

Passeronyssus zeferinoi Amaral

Passeronyssus zeferinoi Amaral, 1973: 73

Type locality: Hanhaem, Est. de São Paulo

Distribution:Brazil

Host: Cacicus haemorrhow (Passeriformes: Icteridae)

Genus Locustellonyssus Bregetova

Locustellonyssus Bregetova, 1965: 1093

Type species: Locustellonyssus amurensis Bregetova, 1965

Locustellonyssus amurensis Bregetova

Locustellonyssus amurensis Bregetova, 1965: 1093

Type locality: Amuro-Zeyskoe Plato (Amurskaya obl.)

Distribution: Russia

Host: Locustella fasciolata (Passeriformes: Megaluridae)

Locustellonyssus sibiricus Butenko et Stanyukovich

Locustellonyssus sibiricus Butenko & Stanyukovich, 2001: 526

Type locality: Manatka, Tomskaya Oblast

Distribution: Russia

Host: Locustella certhiola (Passeriformes: Megaluridae)

Genus Pipronyssus Fain et Aitken

Pipronyssus Fain & Aitken, 1967: 36

Type species: Pipronyssus manaci Fain et Aitken, 1967

Pipronyssus manaci Fain et Aitken

Pipronyssus manaci Fain & Aitken, 1967: 36; 1970: 349

Type locality: Elgin Trace, Vega de Oropouche

Distribution: Brazil, Trinidad

Hosts: Pipra erythrocephala, P. iris, P. pipra (Passeriformes: Pipridae)

Genus Ptilonyssus Berlese et Trouessart

Ptilonyssus Berlese & Trouessart, 1889: 128

Type species: Ptilonyssus echinatus Berlese et Trouessart, 1889

Neonyssus Hirst, 1922: 771

Type species: Neonyssus intermedius Hirst, 1921

Rhinonyssoides Hirst, 1922: 770 – syn. by Zumpt & Till, 1955: 69

Type species: Rhinonyssoides trouessarti Hirst, 1922

Neonyssoides Hirst, 1923: 976

Type species: Rhinonyssus (Neonyssoides) nucifragae Hirst, 1923

Flavionyssus Castro, 1948: 266 – syn. by Zumpt & Till, 1955: 69

Type species: Ptilonyssus (Flavionyssus) rabelloi Castro, 1948

Rochanyssus Castro, 1948: 272 – syn. by Zumpt & Till, 1955: 69

Type species: Ptilonyssus (Rochanyssus) werneri Castro, 1948

Paraneonyssus Castro, 1948: 274 – syn. by Zumpt & Till, 1955: 69

Type species: Neonyssus (Paraneonyssus) enriettii Castro, 1948

Travanyssus Castro, 1948: 276 – syn. by Zumpt & Till, 1955: 69

Type species: Neonyssus (Travanyssus) paranensis Castro, 1948

Rhinacarus Castro, 1948: 257

Type species: Rhinonyssus (Rhinacarus) angrensis Castro, 1948

[Vitznyssus Castro, 1948: 277 – syn. by Zumpt & Till, 1955: 69, but restaured by Butenko, 1977]

Type species: Dermanyssus nitzschi Giebel, 1871

Cas Baker et Wharton, 1952: 81

Type species: Rhinonyssus (Rhinacarus) angrensis Castro, 1948

Hapalognatha Butenko, 1959: 17; 1960: 1490

Type species: Hapalognatha prima Butenko, 1959 (= Ptilonyssus motacillae Fain, 1956)

Periglischrodes Baker & Delfinado, 1964: 589 – Syn. by Bregetova, 1967: 135

Type species: Periglischrodes gressitti Baker et Delfinado, 1964 – transferred to Rhinonyssidae by Prasad (1969a)

Ptilonyssoides Vitzthum, 1935

Type species: Ptilonyssoides triscutatus Vitzthum, 1935

Spizonyssus Bregetova, 1967

Type species: Neonyssus (Spizonyssus) pygmaeus Bregetova, 1967

Ptilonyssus acanthopneustes Stanyukovich et Butenko

Ptilonyssus acanthopneustes Stanyukovich & Butenko, 2003: 37

Type locality: Tyumenskaya Oblast

Distribution: Russia

Host: Phylloscopus borealis (Passeriformes: Phylloscopidae)

Ptilonyssus acrocephali Fain

Ptilonyssus acrocephali Fain, 1964a: 55; Domrow, 1969b: 358; 1988: 868; Pence, 1972; 1975: 23; Fain, Sixl & Moritsch, 1974: 3; Spicer, 1987: 261; Knee, Proctor & Galloway, 2008: 369; Knee & Galloway, 2017: 95

Type locality: Deurne (Anvers, Belgium)

Distribution: Australia, Austria, Belgium, Canada, USA

Hosts: Acrocephalus arundinaceus, A. australis, A. schoenobaenus, A. s. scirpaceus (Passeriformes: Acrocephalidae), Regulus calendula (Passeriformes: Regulidae)

Ptilonyssus afroturdi Fain

Ptilonyssus afroturdi Fain, 1962c: 130

Type locality: Akanyaru

Distribution: Rwanda

Host: Turdus olivaceus graueri (Passeriformes: Turdidae)

Ptilonyssus agelaii Fain

Ptilonyssus agelaii Fain & Aitken, 1967: 7

Type locality: Biche-Ortoire Lagoon, Biche

Distribution: Trinidad

Host: Agelaius icterocephalus (Passeriformes: Icteridae)

Ptilonyssus ailuroedi Domrow

Ptilonyssus ailuroedi Domrow, 1964a: 31; 1964b: 348; 1964d: 619; 1988: 868

Type locality: Mt. Glorious, S.E. Queensland

Distribution: Australia

Host: Ailuroedus crassirostris (Passeriformes: Ptilonorhynchidae)

Ptilonyssus alaudae (Butenko et Stanyukovich)

Neonyssus (Otocorinyssus) alaudae Butenko & Stanyukovich, 2001: 520

Astridiella alaudae (Butenko):

Type locality: Turkmenistan

Distribution: Turkmenistan

Host: Alauda arvensis (Passeriformes: Alaudidae)

Ptilonyssus amarali Fain

Ptilonyssus amarali Fain, 1964b: 7; Fain & Aitken, 1967: 6

Type locality: Anvers Zoo

Distribution: Brazil, Trinidad

Hosts: Cyanerpes caeruleus, C. cyanea (Passeriformes: Thraupidae)

Ptilonyssus amazonicus Fain

Ptilonyssus amazonicus Fain, 1972: 358

Type locality: Redondo Lake, nr. Manaos

Distribution: Brazil

Host: Aves

Ptilonyssus ammomani Stanyukovich et Butenko

Ptilonyssus ammomani Stanyukovich & Butenko, 2003: 32

Type locality: Karagan, Ashhabad Region

Distribution: Turkmenistan

Host: Ammomanes deserti (Passeriformes: Alaudidae)

Ptilonyssus andropadi Fain

Ptilonyssus andropadi Fain, 1956a: 141; Strandtmann, 1960: 144

Type locality: Rugege Forest (Rwanda)

Distribution: Malaysia (Sabah), Rwanda

Hosts: Andropadus latirostris eugenius, Hypsipetes flavala connectens, Phyllastrephus fischeri sucosus (Passeriformes: Pycnonotidae)

Ptilonyssus angrensis (Castro)

Rhinonyssus (Rhinacarus) angrensis Castro, 1948: 257

Cas angrensis: Strandtmann & Wharton, 1958: 169

Sternostoma angrensis (Castro): Zumpt & Till, 1955: 91; Fain, 1957: 84; Amaral, 1968: 122; Sartbaev, 1983: 84

Ptilonyssus angrensis: Domrow, 1969b: 348; 1988: 868; Pence, 1973: 887; Knee, Proctor & Galloway, 2008: 369

Type locality: Fazenda Japuiba, Angra dos Reis, Brazil

Distribution: Australia, Brazil, Canada, Kirghizstan, USA

Hosts: Hirundo aethiopica, H. neoxena, Progne chalybea domestica, P. subis, Ptyonoprogne rupestris, Riparia riparia (Passeriformes: Hirundinidae)

Ptilonyssus angustirostris Fain et Aitken

Ptilonyssus angustirostris Fain & Aitken, 1968: 12; 1970: 353

Type locality: APEG Forest

Distribution: Brazil

Host: Glyphorynchus spirurus (Passeriformes: Furnariidae, Dendrocolapinae)

Ptilonyssus anthi Stanyukovich et Butenko

Ptilonyssus anthi Stanyukovich & Butenko, 2003: 43

Type locality: Ryazanskaya Oblast

Distribution: Russia

Host: Anthus trivialis (Passeriformes: Motacillidae)

Ptilonyssus astridae Fain

Ptilonyssus astridae Fain, 1956a: 141; 1957b: 98; Fain & Lukoschus, 1979: 10; Domrow, 1988: 868; 1992: 1605; Fain, Herin & Puylaert, 1977: 386

Paraneonyssus astridae Strandtmann & Wharton, 1958: 176

Type locality: Astrida (= Butare), Rwanda

Distribution: Australia, Cameroon, Rwanda

Hosts: Donacola castaneothorax (Passeriformes: Ploceidae), Euschistospiza dybowskii, Lagonosticta senegala ruberrima, L. rubricata congica, Poephila personata (Passeriformes: Estrildidae), Vidua macroura (Passeriformes: Viduidae)

Ptilonyssus aureliani Fain

Ptilonyssus aureliani Fain, 1956a: 146; 1957b: 109; Fain, Herin & Puylaert, 1977: 386

Type locality: Astrida (= Butare, Rwanda)

Distribution: Cameroon, Rwanda

Hosts: Camaroptera brevicaudata, Prinia subflava graueri (Passeriformes: Cisticolidae)

Ptilonyssus balimoensis Sakakibara

Ptilonyssus balimoensis Sakakibara, 1968a: 17; Domrow, 1969: 368; 1988: 869

Type locality: Western District, Balimo (Papua New Guinea)

Distribution: Australia, Papua New Guinea

Hosts: Meliphaga macleayana, Xanthotis flaviventer (Passeriformes: Meliphagidae)

Ptilonyssus belemensis Fain et Aitken

Ptilonyssus belemensis Fain & Aitken, 1968: 4

Type locality: Mucambo Forest

Distribution: Brazil

Host: unknown (Passeriformes: ? Furnariidae, Dendrocolaptinae)

Ptilonyssus bombycillae Fain

Ptilonyssus bombycillae Fain, 1972: 357; Rodríguez, Úbeda & Guevara, 1993: 13; Knee, Proctor & Galloway, 2008: 369

Ptilonyssus bombycillae Pence, 1973c: 360 – type from Bombycilla cedrorum, New Orleans, La. (USA); Spicer, 1978: 891

Type locality: Waarschoot

Distribution: Belgium, Canada, Spain, USA

Hosts: Bombycilla cedrorum, B. garrulus (Passeriformes: Bombycillidae)

Ptilonyssus bradypteri (Fain)

Passeronyssus bradypteri Fain, 1962a: 143

Ptilonyssus bradypteri: Domrow, 1969b: 358; 1988: 869

Type locality: Akanyaru (Rwanda)

Distribution: Australia, Rwanda

Hosts: Bradypterus baboecala centralis (Passeriformes: Megaluridae), Cinclorhamphus mathewsi (Passeriformes: Sylviidae)

Ptilonyssus buloloensis Sakakibara

Ptilonyssus buloloensis Sakakibara, 1968c: 426; Domrow, 1988: 869

Type locality: Bulolo River, Papua New Guinea

Distribution: Australia, Papua New Guinea

Host: Manucodia (Phonygammus) keraudrenii (Passeriformes: Paradisaeidae)

Ptilonyssus cacici Fain

Ptilonyssus cacici Fain, 1964b: 9

Type locality: Anvers Zoo

Distribution: South America

Host: Cacicus sp. (Passeriformes: Icteridae)

Ptilonyssus calamocichlae Fain

Ptilonyssus calamocichlae Fain, 1956a: 142

Type locality: Akanyaru

Distribution: Rwanda

Host: Acrocephalus [Calamocichla] rufescens foxi (Passeriformes: Acrocephalidae)

Ptilonyssus calamocichlae hippolaisi Fain

Ptilonyssus calamocichlae hippolaisi Fain, 1963c: 78

Type locality: Akanyaru

Distribution: Rwanda

Host: Hippolais icterina (Passeriformes: Acrocephalidae)

Ptilonyssus calandrellae (Bregetova)

Neonyssus (Otocorinyssus) calandrellae Bregetova, 1967: 129

Astridiella calandrellae (Bregetova): Úbeda et al., 1986: 167

Type locality: at Zeravshan River, 15 km upstream Penjikent (Tajikistan)

Distribution: Georgia, Spain, Tadjikistan

Host: Calandrella cinerea (Passeriformes: Alaudidae)

Ptilonyssus callinectoides (Brooks et Strandtmann)

Tyranninyssus callinectoides Brooks & Strandtmann, 1960: 420; Hyland & Moorhouse, 1970: 53

Ptilonyssus callinectoides (Brooks et Strandtmann): Pence, 1972c: 790; Knee, Proctor & Galloway, 2008: 369

Type locality: Briscoe County, Texas

Distribution: Canada, Mexico, USA (La., Texas)

Hosts: Myiarchus cinerascens, M. crinitus, M. tyrannulus (Passeriformes: Tyrannidae)

Ptilonyssus calvaria Knee

Ptilonyssus calvaria Knee, 2008: 350; Knee & Galloway, 2017: 93

Type locality: Winnipeg (Manitoba)

Distribution: Canada (Manitoba, Alberta)

Hosts: Passerculus sandwicensis, Spizella passerina (Passeriformes: Passerellidae)

Ptilonyssus camptostoma Hyland et Moorhouse

Ptilonyssus camptostoma Hyland & Moorhouse, 1970: 44

Type locality: Tlacotalpan (Veracruz)

Distribution: Mexico

Host: Camptostoma imberbe (Passeriformes: Tyrannidae)

Ptilonyssus capensis Zumpt et Till

Ptilonyssus capensis Zumpt & Till, 1955: 72; Fain, 1956a: 138; 1957b: 89; Butenko, 1965: 132; Úbeda-Ontíveros & Guevara-Benítez, 1978

Neonyssus capensis (Zumpt et Till): Shumilo & Lunkashu, 1970: 19; Sartbaev, 1983: 84

Paraneonyssus capensis: Strandtmann & Wharton, 1958: 176

Type locality: Cape Province

Distribution: Moldavia, Kirghizstan, Russia, Rwanda, South Africa, Spain

Hosts: Anthus campestris, A. pratensis, A. spinoletta, A. trivialis, Macronyx capensis, M. croceus, Motacilla alba (Passeriformes: Motacillidae)

Ptilonyssus capitatus (Strandtmann)

Paraneonyssus capitatus Strandtmann, 1956: 133; Strandtmann & Wharton, 1958: 177

Astridiella capitatus (Str.): Fain, 1959: 21; Butenko, 1965: 132

Ptilonyssus capitatus Domrow, 1969b: 339; 1988: 869

Type locality: Tooele, Utah

Distribution: Australia, Russia, Rwanda, South Africa, USA (Utah)

Hosts: Alauda arvensis, Mirafra javanica, Otocoris alpestris (Passeriformes: Alaudidae), Teloporus dohertyi, T. zeylonus (Passeriformes: Malaconotidae)

Ptilonyssus carduelis Fain

Ptilonyssus carduelis Fain, 1962b: 253; Wilson & Haas, 1980: 545; Úbeda-Ontíveros, Guevara-Benítez & Morillas-Marquez, 1985: 272; Knee, Proctor & Galloway, 2008: 368

Neonyssus (Fringilonyssus) carduelis (Fain): Bregetova, 1967: 131; Shumilo & Lunkashu, 1970: 20; Beron, 1975: 187; Sartbaev, 1983: 84

Type locality: Anvers (Belgium)

Distribution: Belgium, Bulgaria, Canada, Kirghizstan, Moldavia, Russia, Spain, Tajikistan, USA (Alaska); Australia (introduced)

Hosts: Acanthis flavirostris, Carduelis [Acanthis] c. cannabina, Carpodacus erythrinus, C. flammea, C. rhodochlamys, C. rubicilla, Leucosticte brandti, Loxia curvirostra, L. leucoptera, Pinicola enucleator (Passeriformes: Fringillidae)

Ptilonyssus cerchneis Fain

Ptilonyssus cerchneis Fain, 1957b: 133; Strandtmann, 1962: 100; Bregetova, 1964: 5; Domrow, 1965b: 457; 1969b: 374; 1988: 869; Butenko, 1984: 156; Knee, Proctor & Galloway, 2008: 368; Knee & Galloway, 2017: 95

Rhinonyssoides cerchneis, Strandtmann & Wharton, 1958: 172

Type locality: Rugege (Rwanda)

Distribution: Australia, Canada, Russia, Rwanda, USA

Hosts: Falco berigora, F. cenchroides, F. naumanni, F. sparverius, F. tinnunculus, F. [Cerchneis] tinnunculus rufescens (Falconiformes: Falconidae), Turdus migratorius (Passeriformes: Turdidae)

Ptilonyssus cercomacrae Fain et Aitken

Ptilonyssus cercomacrae Fain & Aitken, 1970: 390

Type locality: APEG (Varsea), IPEAN, Belém

Distribution: Brazil

Host: Cercomacra tyrannina (Passeriformes: Formicariidae)

Ptilonyssus certhiae Fain et Bafort

Ptilonyssus certhiae Fain & Bafort, 1963b: 483; Fain, Sixl & Moritsch, 1974: 3

Type locality: St Laureins (Belgium)

Distribution: Austria, Belgium

Host: Certhia b. brachydactyla (Passeriformes: Certhiidae)

Ptilonyssus certhiaxicola Fain

Ptilonyssus certhiaxicola Fain, 1964: 31; Fain & Aitken, 1967: 6; 1970: 356

Type locality: Anvers Zoo

Distribution: Brazil, Trinidad

Hosts: Certhiaxis cinnamomea, Synallaxis rutilans (Passeriformes: Furnariidae)

Ptilonyssus certhilaudae Fain

Ptilonyssus certhilaudae Fain, 1972: 360

Type locality: Bloemhof, Transvaal

Distribution: South Africa

Host: Certhilauda albofasciata (Passeriformes: Alaudidae)

Ptilonyssus chalybaedomesticae Amaral

Ptilonyssus chalybaedomesticae Amaral, 1967: 169; 1968: 121; Černý & Dusbábek, 1970: 482

Type locality: Angra dos Reis (Rio de Janeiro)

Distribution: Brazil

Host: Progne chalybea domestica (Passeriformes: Hirundinidae)

Ptilonyssus chiroxiphiae Fain et Aitken

Ptilonyssus chiroxiphiae Fain & Aitken, 1970: 361

Type locality: APEG (Station A), IPEAN, Belém

Distribution: Brazil

Host: Chiroxiphia pareola (Passeriformes: Pipridae)

Ptilonyssus chloris Fain

Ptilonyssus carduelis chloris Fain, 1962b: 257

Neonyssus chloris (Fain): Shumilo & Lunkashu, 1970: 20

Ptilonyssus (Frigilonyssus) carduelis chloris Fain: Bregetova, 1967: 130

Ptilonyssus chloris Fain: Rodríguez, Úbeda, Guevara & Ariza, 1989: 152

Type locality: Anvers (Belgium)

Distribution: Belgium, Moldavia, Russia, Spain

Host: Chloris chloris (Passeriformes: Fringillidae)

Ptilonyssus chlorocichlae Fain

Ptilonyssus chlorocichlae Fain, 1956a: 142

Type locality: Akanyaru

Distribution: Rwanda

Host: Chlorcichla flavicollis pallidigula (Passeriformes: Pycnonotidae)

Ptilonyssus chloropsicola Fain

Ptilonyssus chloropsicola Fain, 1966: 90

Type locality: Anvers (Zoo)

Distribution: Southeast Asia

Host: Chloropsis cochinchinensis (Passeriformes: Chloropseidae)

Ptilonyssus cinnyricincli Fain

Ptilonyssus cinnyricincli Fain, 1962c: 132

Type locality: Buzi

Distribution: Mozambique

Host: Cinnyricinclus leucogaster (Passeriformes: Sturnidae)

Ptilonyssus cinnyris Zumpt et Till

Ptilonyssus cinnyris Zumpt & Till, 1955: 78; Fain, 1956a: 146; 1957b: 86; Nadchatram, McClure & Chong, 1964: 112; Domrow, 1969: 374; 1988: 869

Rhinonyssoides cinnyris Strandtmann & Wharton, 1958: 172

Type locality: Cape Prov. (South Africa)

Distribution: Australia, Malaysia, Rwanda, South Africa

Hosts: Arachnothera longirostris, Chalcomitra senegalensis aequatorialis, Cinnyris afra, C. regius kivuensis, Cyrtostomus frenatus, Nectarinia kilimensis (Passeriformes: Nectariniidae)

Ptilonyssus cisticolarum Fain

Ptilonyssus cisticolarum Fain, 1959a: 26; 1960: 109

Type locality: Rustenburg, Transvaal

Distribution: Botswana, Rwanda, South Africa

Hosts: Apalis flavida caniceps, Cisticola robusta nuchalis, C. lais (Passeriformes: Cisticolidae)

Ptilonyssus coccothraustis Fain et Bafort

Ptilonyssus coccothraustis Fain & Bafort, 1963b: 477; Sixl, 1969: 218; Fain, Sixl & Moritsch, 1974: 3

Neonyssus coccothraustis: Bregetova, 1967: 130; Shumilo & Lunkashu, 1970: 20

Type locality: Anvers (Belgium)

Distribution: Austria, Belgium, Canada, Georgia, Kyrghizstan, Moldavia, Ukraine

Hosts: Cocothraustes coccothraustes, C. vespertinus (Passeriformes: Fringillidae)

Ptilonyssus colluricinclae Domrow

Ptilonyssus colluricinclae Domrow, 1964d: 596; 1969b: 339; 1979d: 494; 1988: 870

Type locality: Brisbane (Qu.)

Distribution: Australia

Hosts: Colluricincla phaea (= harmonica), C. megarhyncha, Pachycephala griseiceps, P. pectoralis (Passeriformes: Pachycephalidae)

Ptilonyssus colopteryx Fain et Aitken

Ptilonyssus colopteryx Fain & Aitken, 1971: 32

Type locality: APEG Forest, Station A (IPEAN)

Distribution: Brazil

Host: Colopteryx galeatus (Passeriformes: Tyrannidae)

Ptilonyssus condylocoxa Fain et Lukoschus

Ptilonyssus condylocoxa Fain & Lukoschus, 1979: 11; Koch, 1986: 60; Domrow, 1988: 870

Type locality: Beverley Springs, Western Australia

Distribution: Australia

Host: Ramsayornis fasciatus (Passeriformes: Meliphagidae)

Ptilonyssus conopophagae Fain et Aitken

Ptilonyssus conopophagae Fain & Aitken, 1970: 378

Type locality: Mosqueiro Ferry, Benavides, Belém

Distribution: Brazil

Host: Conopophaga roberti (Passeriformes: Conopophagidae)

Ptilonyssus conopophilae Fain et Lukoschus

Ptilonyssus conopophilae Fain & Lukoschus, 1979: 13; Koch, 1986: 60; 1988: 870

Type locality: Napier Downs, Western Australia

Distribution: Australia

Hosts: Conopophila rufogularis, Grantiella picta (Passeriformes: Meliphagidae), ? Taeniopygia [Poephila guttata] castanotis (Passeriformes: Estrildidae)

Ptilonyssus corcoracis Domrow

Ptilonyssus corcoracis Domrow, 1969: 345; 1988: 870

Type locality: Condamine, Charleville

Distribution: Australia

Host: Corcorax melanorhamphus (Passeriformes: Corcoracidae)

Ptilonyssus corvi Pence

Ptilonyssus corvi Pence, 1972c: 791

Type locality: Laplace, Louisiana

Distribution: USA (La.)

Host: Corvus brachyrhynchos (Passeriformes: Corvidae)

Ptilonyssus corythopicola Fain et Aitken

Ptilonyssus corythopicola Fain & Aitken, 1970: 372

Type locality: Mosqueiro Ferry, Benavides, Belém

Distribution: Brazil

Host: Corythopis torquata (Passeriformes: Conopophagidae)

Ptilonyssus cractici Domrow

Ptilonyssus cractici Domrow, 1964d: 598, 616; 1966b: 196; 1969b: 349; 1972: 100; 1979: 497; Fain & Lukoschus, 1979: 10

Type locality: Brisbane (Qu.), Condamine, Logan Village (Australia)

Distribution: Australia, New Zealand

Hosts: Cracticus mentalis, C. nigrogularis, C. quoyi, C. torquatus (Passeriformes: Cracticidae), Dacelo gigas (Coraciiformes: Alcedinidae), Gymnorhina hypoleuca, G. tibicen, Lalage tricolor (Passeriformes: Compephagidae), Strepera graculina (Passeriformes: Artamidae)

Ptilonyssus cyanosylviae Butenko et Lavrovskaya

Ptilonyssus cyanosylviae Butenko & Lavrovskaya, 1980: 530

Type locality: Novosibirskaya Oblast

Distribution: Russia

Host: Cyanosylvia svecica (Passeriformes: Muscicapidae)

Ptilonyssus daptrii Fain

Ptilonyssus daptrii Fain, 1966: 88

Type locality: Rio Preto da Eva

Distribution: Brazil

Host: Daptrius ater (Falconiformes: Falconidae)

Ptilonyssus degtiarevae Dimov et Mironov

Ptilonyssus degtiarevae Dimov & Mironov, 2012: 169

Type locality: Gavrilovo Village, Leningradskaya Obl.

Distribution: Russia

Host: Passer domesticus (Passeriformes: Passeridae)

Ptilonyssus dendrocittae (Sakakibara et Strandtmann)

Paraneonyssus dendrocitti Sakakibara & Strandtmann, 1968: 43

Ptilonyssus dendrocittae (Sakakibara & Strandtmann)

Type locality: Nan-T‘ou Hsien: Pu-Li

Distribution: Taiwan

Host: Dendrocitta [Crypsirina] occipitalis (Passeriformes: Corvidae)

Ptilonyssus dendrocolapticola Fain et Aitken

Ptilonyssus dendrocolapticola Fain & Aitken, 1968: 8

Type locality: APEG Forest

Distribution: Brazil

Host: Dendrocolaptes certhia (Passeriformes: Furnariidae, Dendrocolaptinae)

Ptilonyssus desfontainei Zumpt et Till

Ptilonyssus desfontainei Zumpt & Till, 1955: 71; Fain, 1957b: 84

Type locality: Transvaal

Distribution: South Africa

Host: Thamnolaea cinnamomeiventris (Passeriformes: Turdidae)

Ptilonyssus diadori Butenko et Lavrovskaya

Ptilonyssus diadori Butenko & Lavrovskaya, 1980: 530

Type locality: Russia

Distribution: Russia

Host: Aves

Ptilonyssus dicaei Domrow

Ptilonyssus dicaei Domrow, 1966a: 201; 1967a: 217; 1969b: 366; 1988: 870

Type locality: Mt. Jukes, near Mackay, N.Q.

Distribution: Australia

Host: Dicaeum hirundinaceum (Passeriformes: Dicaeidae)

Ptilonyssus dicruri Fain

Ptilonyssus dicruri Fain, 1956a: 137; Domrow, 1969b: 348; 1988: 871; Domrow & Nadchatram, 1978: 87; Feider & Mironescu, 1980: 61

Ptilonyssoides dicruri: Strandtmann & Wharton, 1958: 170; Strandtmann, 1960: 140

Passeronyssus dicruri (Fain): Fain, 1960: 110; Domrow, 1965b: 451

Type locality: Astrida (= Butare), Rwanda

Distribution: Australia, Botswana, Malaysia, Papua New Guinea, Rwanda, Thailand

Hosts: Chibia bracteata, Dicrurus adsimilis, D. hottentottus, D. leucophaeus stigmatops, D. macrocercus cathoecus, D. remifer (Passeriformes: Dicruridae)

Ptilonyssus dioptrornis Fain

Ptilonyssus dioptrornis Fain, 1956a: 137; 1959a: 26; Domrow, 1969b: 347; 1988: 871; Domrow & Nadchatram, 1978: 86; Kadosaka, Kaneko & Asanuma, 1983: 221

Ptilonyssoides dioptrornis Strandtmann & Wharton, 1958: 170

Passeronyssus dioptrornis (Fain): Fain, 1960: 110

Type locality: Rugege Forest (Rwanda)

Distribution: Australia, Japan, Malaysia, Rwanda, South Africa

Hosts: Cossypha [Bessonornis] archeri, C. dichroa, Dioptrornis fischeri toruensis (Passeriformes: Turdidae), Copsychus malabaricus, C. saularis musicus, Eopsaltria capito, Muscicapa c. cyanomelana, M. rufigastra (Passeriformes: Muscicapidae)

Ptilonyssus dolicaspis Feider et Mironescu

Ptilonyssus dolicaspis Feider & Mironescu, 1980: 59

Type locality: West Squak

Distribution: Papua New Guinea

Host: Megalurus timonensis (Passeriformes: Megaluridae)

Ptilonyssus domrowi Feider et Mironescu

Ptilonyssus domrowi Feider & Mironescu, 1980: 57

Type locality: Tambul

Distribution: Papua New Guinea

Host: Paramythia montium (Passeriformes: Paramythiidae)

Ptilonyssus donatoi Pereira et Castro

Ptilonyssus (Rhinonyssoides) donatoi Pereira & Castro, 1949: 224

Ptilonyssus donatoi: Zumpt & Till, 1955: 88; Fain, 1957b: 84; Amaral, 1968: 121

Type locality: Estado de Sao Paulo

Distribution: Brazil

Host: Coragyps atratus foetens (Cathartiformes: Catarthidae)

Ptilonyssus dryoscopi Zumpt et Till

Ptilonyssus dryoscopi Zumpt & Till, 1955: 73; Fain, 1956a: 139; 1957b: 84

Paraneonyssus dryoscopi (Zumpt et Till): Strandtmann & Wharton, 1958: 177

Type locality: Cape Prov. (South Africa)

Distribution: Rwanda, South Africa

Hosts: Dryoscopus cubla, Tchagra senegala (Passeriformes: Malaconotidae)

Ptilonyssus echinatus Berlese et Trouessart

Ptilonyssus echinatus Berlese & Trouessart, 1889: 9; Vitzthum, 1935: 578; Pereira & Castro, 1949: 219; Zumpt & Till, 1955: 74; Fain, 1956: 136; 1956: 114; 1957b: 137; 1957c: 218; 1960: 108; 1962b: 262; Strandtmann & Wharton, 1958: 179; Strandtmann, 1960: 146; George, 1961: 114; Domrow, 1964d: 607; 1969b: 349, 423; 1988: 871; Fain & Lukoschus, 1972: 90; Fain, Sixl & Moritsch, 1974: 1; Beron, 1975: 186; Scherbinina & Butenko, 1981: 70; Spicer, 1987: 261; Knee, Proctor & Galloway, 2008: 369; Huhta, 2016: 148 (S. incert.).

Ptilonyssoides echinatus (Berlese et Trouessart): Shumilo & Lunkashu, 1970: 29

Hapalognatha platytrichia Butenko, 1959: 17; 1960: 1494. – Type: Tatarstan (Russia), from Hirundo rustica (syn. by Fain, 1962b: 262)

Type locality: Germany

Distribution: Australia, Austria, Bulgaria, Cameroon, Canada, Cuba, ? Finland, Germany, Moldavia, Russia, South Africa, Spain, Suriname, Thailand, Trinidad, Turkmenistan, Ukraine, USA

Hosts: Atticora melanoleuca, Delichon urbica, Hirundo neoxena, H. nigrita, H. pyrrhonota, H. rustica, H. rustica erythrogaster, H. rustica gutturalis, Hylochelidon ariel, H. nigricans, Riparia riparia, Petrochelidon pyrrhonota, P. spilodera, Stelgidopteryx ruficollis, Tachycineta bicolor (Passeriformes: Hirundinidae)

Ptilonyssus elaeniae Fain et Aitken

Ptilonyssus elaeniae Fain & Aitken, 1967: 11

Type locality: La Horquette Trace, Tumpuna Road

Distribution: Trinidad

Host: Elaenia flavogaster (Passeriformes: Tyrannidae)

Ptilonyssus elbeli (Strandtmann)

Cas elbeli Strandtmann, 1960: 137

Ptilonyssus elbeli (Strandtmann): Domrow, 1968; Feider & Mironescu, 1978: 266; Sartbaev, 1981: 130 [Cas elbelli]

Type locality: Thailand

Distribution: Azerbaidjan, Belgium, Kirghizstan, Romania, Thailand

Hosts: Acridotheres tristis, Sturnus vulgaris (Passeriformes: Sturnidae)

Ptilonyssus elongatus Fain

Ptilonyssus elongatus Fain, 1964e:123

Type locality: Astrida (= Butare)

Distribution: Rwanda

Hosts: Apalis flavida caniceps, Prinia leucopogon reichenowi (Passeriformes: Cisticolidae)

Ptilonyssus emberizae Fain

Ptilonyssus emberizae Fain, 1956a: 140; 1957b: 95; 1964a; Sixl, 1969: 218; Fain, Sixl & Moritsch, 1974: 2; Domrow, 1972: 99; 1988: 871; Kadosaka, Kaneko & Asanuma, 1983: 221; Guevara Benítez, Úbeda Ontíveros & Morillas Marquez, 1986a: 75

Paraneonyssus emberizae Strandtmann & Wharton, 1958: 177

Fain & Lukoschus, 1979: 10

Neonyssus emberizae (Fain): Shumilo & Lunkashu, 1970: 21; Scherbinina & Butenko, 1981: 70

Type locality: Astrida (= Butare), Rwanda

Distribution: Australia, Austria, Belgium, Moldavia, New Zealand, Spain, Russia, Rwanda, Turkmenistan, Ukraine

Hosts: Emberiza bruniceps, E. cia, E. citrinella, E. flaviventris, E. schoeniclus, E. sulphurata (Passeriformes: Emberizidae), Lonchura pectoralis, Steganopleura bichenovii (Passeriformes: Estrildidae), Fringilla coelebs (Passeriformes: Fringillidae), Plectrophenax nivalis (Passeriformes: Calcariidae), Sturnus vulgaris (Passeriformes: Sturnidae)

Ptilonyssus enicuri Fain et Nadchatram

Passeronyssus enicuri Fain & Nadchatram, 1962: 276

Ptilonyssus enicuri Fain et Nadchatram: Kadosaka, Kaneko & Asanuma, 1983: 222

Type locality: Malaysia

Distribution: Japan, Malaysia

Hosts: Enicurus schistaceus, Erithacus a. akahige (Passeriformes: Turdidae)

Ptilonyssus enrietti Castro

Neonyssus (Paraneonyssus) enrietti Castro, 1948: 274; Pereira & Castro, 1949: 227

Ptilonyssus enrietti (Castro): Zumpt & Till, 1955: 89; Fain, 1957b: 89

Type locality: Brazil

Distribution: Brazil

Host: Phylloscartes ventralis ventralis (Passeriformes: Tyrannidae)

Ptilonyssus eremophilae Butenko et Lavrovskaya

Ptilonyssus eremophilae Butenko & Lavrovskaya, 1980: 526; Sartbaev, 1983: 84

Type locality: Ryazanskaya Obl. (Russia)

Distribution: Kirghizstan, Russia

Host: Eremophila alpestris (Passeriformes: Alaudidae)

Ptilonyssus estrildicola Fain

Ptilonyssus estrildicola Fain, 1959a: 22; Domrow, 1965: 453; Sakakibara, 1968b: 309

Type locality: Astrida (= Butare), Rwanda

Distribution: Australia, Botswana, Papua New Guinea, Rwanda, South Africa

Hosts: Amadina erythrocephala, Batis capensis, Granatina granatina, Lagonosticta rhodopareia, L. rubricata, Pytilia melba, Taeniopygia castanotis, Uraeginthus angolensis, U. bengalus ugandae (Passeriformes: Estrildidae)

Ptilonyssus euroturdi Fain et Hyland

Ptilonyssus euroturdi Fain & Hyland, 1963a: 381; Fain & Bafort, 1963b: 485; Fain, 1966: 7; Feider & Mironescu, 1968; Shumilo & Lunkashu, 1970: 29; Pence, 1972: 1162; Fain, Sixl & Moritsch, 1974: 3; Sartbaev, 1983: 85 ; Spicer, 1984: 800; Kadosaka, Kaneko & Asanuma, 1987: 36; Rodríguez, Úbeda & Guevara, 1990: 301; De Rojas et al., 2001: 961; Knee, Proctor & Galloway, 2008: 369; Dimov, 2016; 2018: 88

Ptilonyssus euroturdi mimicola Fain & Hyland, 1963a: 384 – Type: Kalamazoo Co., Michigan, from Dumetella carolinensis (syn. by Spicer, 1984: 800)

Type locality: Lauwe nr. Bruges, Belgium

Distribution: Austria, Belgium, Canada, Guatemala, Japan, Kirghizstan, Moldavia, New Zealand, Romania, Russia, Spain, Trinidad, Ukraine, USA

Hosts: Catharus dryas, Turdus e. ericetorum, T. fumigatus, T. iliacus, T. merula, T. migratorius, T. naumanni euromus, T. nudigenus, T. philomelus, T. pilaris, T. ruficollis, T. v. viscivorus (Passeriformes: Turdidae), Dumetella carolinensis (Passeriformes: Mimidae), Sturnus vulgaris (Passeriformes: Sturnidae)

Ptilonyssus faini (Strandtmann)

Ptilonyssoides faini Strandtmann, 1960: 142

Ptilonyssus faini: Domrow, 1968: 348

Type locality: Trus Madi Mt, Pampang, Ulu Kaingaran, North Borneo

Distribution: Malaysia (Borneo)

Host: Dicrurus hottentotus borneensis (Passeriformes: Dicruridae)

Ptilonyssus fluvicolae (Fain et Aitken)

Neotyranninyssus fluvicolae Fain et Aitken, 1967: 29; Hyland & Moorehouse, 1970: 53

Ptilonyssus fluvicolae (Fain et Aitken)

Type locality: Nariva Road, 3 mile post (Trinidad)

Distribution: Mexico, Trinidad

Hosts: Fluvicola pica, Pyrocephalus rubinus (Passeriformes: Tyrannidae)

Ptilonyssus formicarii Fain et Aitken

Ptilonyssus formicarii Fain & Aitken, 1970: 364

Type locality: Mosqueiro Ferry, Benavides, Belém

Distribution: Brazil

Host: Formicarius analis (Passeriformes: Formicariidae)

Ptilonyssus formicarii colmae Fain et Aitken

Ptilonyssus formicarii colmae Fain & Aitken, 1970: 368

Type locality: Mosqueiro Ferry, Benavides, Belém

Distribution: Brazil

Host: Formicarius colmae (Passeriformes: Formicariidae)

Ptilonyssus fringillae Fain et Sixl

Ptilonyssus fringillae Fain & Sixl, 1971: 90; Fain, Sixl & Moritsch, 1974: 4; Úbeda-Ontíveros & Guevara-Benítez, 1978

Type locality: Brunnsee, Styria; paratypes from Belgium

Distribution: Austria, Belgium, Spain

Hosts: Anthus pratensis (Passeriformes: Motacillidae); Regulus regulus (Passeriformes: Regulidae)

Ptilonyssus fringillicola Fain

Ptilonyssus fringillicola Fain, 1959a: 28; 1962b: 262; Butenko, 1965: 132; Shumilo & Lunkashu, 1970: 30; Fain, Sixl & Moritsch, 1974: 2; Fain, Herin & Pulyaert, 1977: 387

Ptilonyssus pyrrhulae Feider, 1962: 43 – Type: Romania, from Pyrrhula pyrrhula (syn. by Fain, Sixl & Moritsch, 1974: 2)

Type locality: Rugege Forest (Rwanda)

Distribution: Austria, Belgium, Cameroon, Kirghizstan, Romania, Russia, Rwanda, South Africa, Ukraine

Hosts: Carduelis c. carduelis, Pyrrhula pyrrhula, P. pyrrhula coccinea, Serinus atrogularis, S. mozambicus, S. [Poliospiza] striolata kivuensis (Passeriformes: Fringillidae), Emberiza flaviventris (Passeriformes: Emberizidae)

Ptilonyssus gerygonae Domrow

Ptilonyssus sp. Domrow, 1967a: 217

Ptilonyssus gerygonae Domrow, 1969b: 363; 1979d: 500; 1988: 871

Type locality: Queensland

Distribution: Australia

Hosts: Acanthiza reguloides; Gerygone palpebrosa (Passeriformes: Acanthizidae)

Ptilonyssus gilcolladoi Úbeda, Rodríguez, Guevara et Rojas

Ptilonyssus gilcolladoi Úbeda, Rodríguez, Guevara & Rojas, 1989: 157

Type locality: Prov. de Granada

Distribution: Spain

Host: Cettia cetti (Passeriformes: Cettiidae)

Ptilonyssus gliciphilae Domrow

Ptilonyssus gliciphilae Domrow, 1966a: 204; 1967a: 217; 1969b: 370; 1979d: 504; 1988: 872

Type locality: Chelona nr. Sarina, Esk

Distribution: Australia

Hosts: Gliciphila indistincta, Myzomela pectoralis (Passeriformes: Meliphagidae)

Ptilonyssus grallinae Domrow

Ptilonyssus grallinae Domrow, 1964d: 611, 618; 1969b: 366; 1979d: 500; 1988: 872; Fain & Lukoschus, 1979: 10

Type locality: Brisbane, Condamine (Qu.)

Distribution: Australia

Host: Grallina cyanoleuca (Passeriformes: Monarchidae)

Ptilonyssus gressitti (Baker et Delfinado)

Periglischrodes gressitti Baker et Delfinado, 1964: 589

Ptilonyssus gressitti (Baker et Delfinado): Bregetova, 1967: 135

Type locality: Bulolo

Distribution: Papua New Guinea

Host: «Chiroptera indet. » (according to Bregetova, 1967, it is a chance inclusion in a collection of spinturnicid mites, the real host should be a bird)

Ptilonyssus hirsti (de Castro et Pereira)

Neonyssus hirsti de Castro & Pereira, 1947: 129

Neonyssus (Paraneonyssus) hirsti: Černy & Dusbábek, 1970: 483

Ptilonyssus (Paraneonyssus) hirsti: Pereira & Castro, 1949: 227; Porter & Strandtmann, 1952: 393

Paraneonyssus hirsti Strandtmann & Wharton, 1958: 177; Feider, 1962: 58;1964: 268

Ptilonyssus hirsti: Zumpt & Till, 1955: 70; Fain, 1956a: 140; 1957b: 86; 1963: 170; Domrow, 1964d: 608, 1969b: 341; 1988: 872; Sixl, 1969: 218; Amaral, 1968: 119; Butenko, 1971; Fain, Sixl & Moritsch, 1974: 1; Pence, 1975: 24; Fain, Herin & Pulyaert, 1977: 387; Úbeda-Ontíveros & Guevara-Benítez, 1978; Shterbinina & Butenko, 1981; Knee, Proctor & Galloway, 2008: 370; Moodi et al., 2014: 3; Dimov, 2016; 2018: 85

Type locality: Portugal and North America

Distribution: Australia, Austria, Brazil, Cameroon, Canada, Cuba, Hawaii Is., Iran, New Guinea, Portugal, Romania, Rwanda, South Africa, Spain, Thailand, Turkmenistan, USA

Hosts: Parus major (Passeriformes: Paridae), Passer domesticus, P. griseus, P. indicus, P. montanus, P. ugandae griseus, Petronia dentata (Passeriformes: Passeridae), Fringilla coelebs (Passeriformes: Fringillidae)

Ptilonyssus hiyodori Kadosaka, Kaneko et Asanuma

Ptilonyssus hiyodori Kadosaka, Kaneko & Asanuma, 1987: 36

Type locality: Japan

Distribution: Japan

Host: Hypsipetes amaurotis amaurotis (Passeriformes: Pycnonotidae)

Ptilonyssus hoffmannae Luz-Zamudio

Ptilonyssus hoffmannae Luz-Zamudio, 1984: 113

Type locality: Mexico

Distribution: Mexico

Host: Carpodacus mexicanus centralis (Passeriformes: Fringillidae)

Ptilonyssus hoseini (Fain et Aitken)

Passeronyssus hoseini Fain & Aitken, 1967: 21

Passeronyssus havanensis (Fain et Aitken): Pence, 1972c: 793; Černy & Dusbábek, 1970: 486

Ptilonyssus hoseini (Fain et Aitken): Pence, 1972c: 793; 1973b: 876; Spicer, 1977b: 498; 1987: 261

Type locality: Saint Isidore-Plum Road, Biche (Trinidad)

Distribution: Cuba, Trinidad, USA

Hosts: Hylophylax poecilonota, Phlegopsis nigromaculata (Passeriformes: Thamnophilidae), Cyclarchis gujanensis, Vireo bellii, V. flavifrons, V. griseus, V. solitarius (Passeriformes: Vireonidae)

Ptilonyssus hylophylax Fain et Aitken

Ptilonyssus hylophylax Fain & Aitken, 1970: 383

Type locality: Pirelli Plantation, Marituba; Mosqueiro Ferry, Benevides

Distribution: Brazil

Hosts: Hylophylax poecilonota, Phlegopsis nigromaculata (Passeriformes: Formicariidae)

Ptilonyssus hypocnemoides Fain et Aitken

Ptilonyssus hypocnemoides Fain & Aitken, 1970: 389

Type locality: APEG Forest (Station A), Mucambo Forest, IPEAN, Belém

Distribution: Brazil

Host: Hypocnemoides maculicauda (Passeriformes: Formicariidae)

Ptilonyssus icteridius Strandtmann et Furman

Ptilonyssus icteridius Strandtmann & Furman, 1956: 167; Fain, 1957b: 96; Fain & Aitken, 1967: 5; Pence, 1972c: 794; 1975: 24; Cooper, Madden & Crites, 1975: 262; Spicer, 1977b: 498; 1978: 891, 1987: 261; Spicer, 1978: 891; Knee, Proctor & Galloway, 2008: 368; Moodi et al., 2014: 3; Knee & Galloway, 2017: 93

Paraneonyssus icteridius (Str. et Furman): Zemskaya & Ilienko, 1958: 476

Ptilonyssus (Paraneonyssus) icteridius (Str. et Furman): Wallace & Olsen, 1966

Neonyssus (Paraneonyssus) icteridius (Str. et Furman): Černý & Dusbábek, 1970: 483

Type locality: USA

Distribution: Canada, Iran, Russia, Trinidad, USA

Hosts: Agelaius phoeniceus, Euphagus carolinus, E. cyanocephalus, Icterus galbula, I. nigrogularis, Molothrus ater californicus, Quiscalus lugubris, Qu. quiscula, Sturnella magna, S. neglecta, Xanthocephalus xanthocephalus (Passeriformes: Icteridae), Passer domesticus, P. montanus (Passeriformes: Passeridae), Porzana carolina (Gruiformes: Rallidae), Piranga ludoviciana, Spiza americana (Passeriformes: Cardinalidae), Myiarchus crinitus (Passeriformes: Tyrannidae), Carpodacus mexicanus (Passeriformes: Fringillidae)

Ptilonyssus indicatoris Fain

Ptilonyssus indicatoris Fain, 1957b: 134

Type locality: Akanyaru

Distribution: Rwanda

Host: Indicator indicator (Piciformes: Indicatoridae)

Ptilonyssus inflatipalpus Fain et Aitken

Ptilonyssus inflatipalpus Fain & Aitken, 1970: 396

Type locality: Mosqueiro Ferry, Benevides

Distribution: Brazil

Host: Myrmornis torquata (Passeriformes: Formicariidae)

Ptilonyssus inornatus Fain et Aitken

Ptilonyssus inornatus Fain & Aitken, 1968: 14; 1970: 352

Type locality: APEG Forest

Distribution: Brazil

Hosts: Myrmotherula axillaris, M. longipennis, M. menetriesi (Passeriformes: Formicariidae)

Ptilonyssus insularis cubanus Černý

Ptilonyssus insularis cubanus Černý, 1969: 231

Ptilonyssus insularis Černý & Dusbábek, 1970: 482

Type locality: Aguacate, Mantingua, Baracoa, prov. of Oriente

Distribution: Cuba

Host: Dives atroviolaceus (Passeriformes: Icteridae)

Ptilonyssus insularis insularis Černý

Ptilonyssus insularis insularis Černý, 1969: 230

Ptilonyssus insularis Černý & Dusbábek, 1970: 482

Type locality: Cayo Piedra, Isla de Pinos

Distribution: Cuba (Pinos)

Host: Quiscalus niger caribaeus (Passeriformes: Icteridae)

Ptilonyssus intermedius (Hirst)

Neonyssus intermedius Hirst, 1922: 771; Vitzthum, 1935: 578; Castro, 1948: 268; Pereira & Castro, 1949: 228; Zumpt & Patterson, 1951: 77; Zumpt & Till, 1955: 62; Fain, 1957b: 52

Ptilonyssus intermedius (Hirst): Fain & Hyland, 1962: 341

Type locality: Madagascar

Distribution: Madagascar, Russia

Host: Aves («on a bird from Madagascar«)

Ptilonyssus isakovae Butenko et Lavrovskaya

Ptilonyssus isakovae Butenko & Lavrovskaya, 1983: 113

Type locality: Kirghizstan (Ottuk Vill., Issykkul Lake)

Distribution: Kirghizstan

Host: Phylloscopus inornatus (Passeriformes: Phylloscopidae)

Ptilonyssus japuibensis Castro

Ptilonyssus japuibensis Castro, 1948: 264; Pereira & Castro, 1949: 219; Zumpt & Till, 1955: 89; Fain, 1957b: 103; George, 1961: 127; Fain & Aitken, 1967: 3; Amaral, 1968: 120; Hyland & Moorhouse, 1970: 44; Fain & Aitken, 1970: 353; Pence, 1973b: 877; Knee, Proctor & Galloway, 2008: 368; Knee & Galloway, 2017: 93

Type locality: Fazenda Japuiba, Angra dos Reis (Estado do Rio de Janeiro)

Distribution: Brazil, Canada, Mexico, Trinidad, USA

Hosts: Rhamphocelus carbo, Rh. bresilius dorsalis, Sporophila americana, Tachyphonus rufus, T. surinamus, Volatinia jacarina (Passeriformes: Thraupidae), Pheucticus ludovicianus (Passeriformes: Cardinalidae), Junco hiemalis, Pipilo erythrophthalmus, Spizella palida, Zonotrichia albicollis (Passeriformes: Passerellidae), Molothrus ater, Quiscalus quiscala (Passeriformes: Icteridae), Empidonax alnorum (Passeriformes: Tyrannidae)

Ptilonyssus japuibensis cyanocompsae Fain et Aitken

Ptilonyssus japuibensis cyanocompsae Fain & Aitken, 1970: 354

Type locality: Mosqueiro Ferry, Benavides, Belém

Distribution: Brazil

Host: Cyanocompsa cyanoides (Passeriformes: Thraupidae)

Ptilonyssus langei Butenko

Ptilonyssus langei Butenko, 1972: 1074

Type locality: Azerbaidjan, Kazakhstan, Russia

Distribution: Azerbaidjan, Kazakhstan, Russia

Host: Sturnus vulgaris (Passeriformes: Sturnidae)

Ptilonyssus lanii Zumpt et Till

Ptilonyssus lanii Zumpt & Till, 1955: 71; Fain, 1956: 143; 1957c: 217; George, 1961: 119; Shumilo & Lunkashu, 1970: 30; Domrow & Nadchatram, 1978: 87; Sartbaev, 1981: 131

Ptilonyssus angustatus Feider & Mironescu, 1969: 21 – Type from Romania, on Lanius collurio.

Type locality: Transvaal

Distribution: Kirghizstan, Malaysia, Moldavia, Romania, Russia, South Africa, Ukraine, USA

Hosts: Lanius collaris, L. collurio, L. cristatus, L. ludovicianus, L. minor (Passeriformes: Laniidae); Sturnus vulgaris (Passeriformes: Sturnidae), Coracias garrulus (Coraciiformes: Coraciidae)

Ptilonyssus lobatus Strandtmann

Ptilonyssus lobatus Strandtmann, 1960: 147; Fain & Nadchatram, 1962: 279

Type locality: Khao Yoi, Phet Buri (Thailand)

Distribution: Malaysia, Thailand

Hosts: Sturnia sturnina, Sturnus javanicus grandis, S. tristis tristis (Passeriformes: Sturnidae)

Ptilonyssus lovottiae Dimov et Mironov

Ptilonyssus lovottiae Dimov & Mironov, 2012: 171

Type locality: Konzj, Leningrad Province

Distribution: Russia

Host: Passer montanus (Passeriformes: Passeridae)

Ptilonyssus ludovicianus Černý

Ptilonyssus ludovicianus Černý, 1969: 228; Černý & Dusbábek, 1970: 482; Knee & Galloway, 2017: 93

Type locality: Soledad, prov. of Las Villas, Botanical Garden

Distribution: Canada, Cuba

Hosts: Pheucticus ludovicianus, Passerina cyanea (Passeriformes: Cardinalidae)

Ptilonyssus lusciniae (Fain)

Passeronyssus lusciniae Fain, 1962: 140; 1963a: 174; Úbeda, Rodríguez & Guevara, 2000

Ptilonyssus lusciniae (Fain): Shumilo & Lunkashu, 1970: 31

Type locality: Belgium

Distribution: Belgium, Malaysia, Moldavia, Rwanda, Spain

Hosts: Luscinia c. cyanae, L. luscinia, Myrmecocychla nigra, Phoenicurus phoenicurus (Passeriformes: Muscicapidae)

Ptilonyssus lymozemae Domrow

Ptilonyssus lymozemae Domrow, 1965b: 453; 1969b: 371; 1979d: 504; 1988: 872

Type locality: Samford, Logan Village

Distribution: Australia

Host: Myzomela sanguinolenta (Passeriformes: Meliphagidae)

Ptilonyssus macclurei Fain

Ptilonyssus macclurei Fain, 1963c: 72; Domrow, 1964d: 609, 617; 1967: 217; 1969b: 366; 1988: 872

Type locality: Mitchell River Mission, Queensland (???)

Distribution: Australia, Malaysia ?

Host: Rhipidura leucophrys (Passeriformes: Muscicapidae)

Ptilonyssus madagascariensis Grétillat, Capron et Brygoo

Ptilonyssus madagascariensis Grétillat, Capron & Brygoo, 1959: 381

Type locality: Tananarive

Distribution: Madagascar

Host: Hypsipetes [Ixocincla] madagascariensis (Passeriformes: Pycnonotidae)

Ptilonyssus malacoptilae (Fain et Aitken)(new comb.)

Passeronyssus malacoptilae Fain & Aitken, 1971: 40

Type locality: Ananindeua

Distribution: Brazil

Host: Malacoptila rufa (Piciformes: Bucconidae)

Ptilonyssus malaysiae Fain

Ptilonyssus malaysiae Fain, 1964c: 65

Type locality: Bukit Lagong, Selangor

Distribution: Malaysia

Host: Irena puella malayensis (Passeriformes: Irenidae)

Ptilonyssus maluri Domrow

Ptilonyssus maluri Domrow, 1965b: 451; 1969b: 339; 1979d: 493; 1988: 872

Type locality: Esk

Distribution: Australia

Host: Malurus melanocephalus (Passeriformes: Maluridae)

Ptilonyssus mariacastroae Fain

Ptilonyssus mariacastroae Fain, 1961b: 48; Fain & Aitkin, 1967: 6; Spicer, 1977b: 498

Type locality: Anvers Zoo (Brazil)

Distribution: Brazil, Mexico, Trinidad

Hosts: Amazilia chionopectus, A. yucatanensis, Chrysolampis mosquitus, Saucerottia tobaci (Apodiformes: Trochilidae)

Ptilonyssus melanocoryphae (Bregetova)

Neonyssus (Otocorinyssus) melanocoryphae Bregetova, 1967: 127

Astridiella melanocoryphae (Bregetova): Úbeda & Morillas Márquez, 1979: 128

Type locality: at Zeravshan River, 15 km upstream Penjikent

Distribution: Tadjikistan

Host: Melanocorypha bimaculata (Passeriformes: Alaudidae)

Ptilonyssus meliphagae Domrow

Ptilonyssus meliphagae Domrow, 1964d: 606; 1967a: 217; 1969b: 372; 1979: 508; 1988: 873

Type locality: Mt. Cotton, Queensland

Distribution: Australia

Hosts: Meliphaga chrysops; Melithreptus albogularis (Passeriformes: Meliphagidae)

Ptilonyssus melissae Spicer

Ptilonyssus melissae Spicer, 1977a: 597; 1978: 891; 1987: 261

Type locality: Diamond-A Ranch, Hood Co., Fort Worth, Tarrant Co., Texas

Distribution: USA (Texas, Montana, Alasca)

Hosts: Carpodacus mexicanus, C. purpureus, Leucosticte tephrocotis, Pinicola enucleator (Passeriformes: Fringillidae)

Ptilonyssus melittophagi Fain

Ptilonyssus melittophagi Fain, 1956a: 136; 1959a: 26

Type locality: Astrida (= Butare), Rwanda

Distribution: Rwanda, South Africa

Hosts: Melittophagus lafresnayi oreobates, M. pusillus (Passeriformes: Meropidae)

Ptilonyssus microecae Domrow

Ptilonyssus microecae Domrow, 1966a: 196; 1967: 217; 69b: 356; 1988: 873

Type locality: Esk

Distribution: Australia

Hosts: Microeca fascinans; M. flavigaster (Passeriformes: Muscicapidae)

Ptilonyssus mimi George

Ptilonyssus mimi George, 1961: 120

Type locality: Kingsville, Texas

Distribution: USA (Texas)

Host: Mimus polyglottos (Passeriformes: Mimidae)

Ptilonyssus mironovi Dimov

Ptilonyssus mironovi Dimov, 2012: 27

Type locality: Lomonosov, Leningrad Province

Distribution: Russia

Host: Parus caeruleus (Passeriformes: Paridae)

Ptilonyssus missimensis Sakakibara

Ptilonyssus missimensis Sakakibara, 1968c: 429

Type locality: Mt. Missim, Morobe District

Distribution: Papua New Guinea

Host: “Honey eater” (Passeriformes: Meliphagidae)

Ptilonyssus monarchae Domrow

Ptilonyssus monarchae Domrow, 1969b: 360; 1988: 873

Type locality: Innisfail

Distribution: Australia

Host: Monarcha trivirgata (Passeriformes: Monarchidae)

Ptilonyssus montifringillae Butenko et Lavrovskaya

Ptilonyssus montifringillae Butenko & Lavrovskaya, 1983: 118

Type locality: Jaz, Issykkul Region

Distribution: Kirghizstan

Host: Montifringilla nivalis (Passeriformes: Passeridae)

Ptilonyssus morofskyi Hyland

Ptilonyssus morofskyi Hyland, 1963: 146; Fain, 1964a: 61; Pence, 1972; 1975: 25; Spicer, 1977a: 600; 1978: 892; Guevara & Úbeda, 1980: 333; Knee, Proctor & Galloway, 2008: 368; Knee & Galloway, 2017: 93

Neonyssus (Spizonyssus) morofskyi (Hyland): Bregetova, 1967: 131; Černý & Dusbábek, 1970: 482

Neonyssus morofskyi (Hyland): Shumilo & Lunkashu, 1970: 22

Type locality: Kalamazoo Co., Michigan

Distribution: Belgium, Canada, Cuba, Moldavia, Spain, USA

Hosts: Carduelis [Spinus] tristis, C. [Acanthis] cannabina, C. flammea, Loxia curvirostra, Serinus serinus (Passeriformes: Fringillidae), Calcarius ornatus, Plectrophenax nivalis (Passeriformes: Calcariidae), Aimophila ruficeps, Ammodramus lecontei, Ammospiza maritima, Calamospiza melanocorys, Junco caniceps, J. hyemalis, Melospiza georgiana, M. lincolni, M. melodia, Passerella iliaca, Passerculus sandwichensis, Pipilo erythrophthalmus, Pooecetes gramineus, Spizella pusilla, Zonotrichia albicollis, Z. leucophrys, Z. querula (Passeriformes: Passerellidae), Dendroica coronata, D. pinus, Mniotilta varia, Geothlypis trichas, Seiurus motacilla, Wilsonia pusilla (Passeriformes: Parulidae)

Ptilonyssus motacillae Fain

Ptilonyssus motacillae Fain, 1956a: 143; 1957b: 137; 1959a: 24; 1960: 109; 1962a: 128; Strandtmann & Wharton, 1958: 181; Domrow, 1964d: 600; 1965b: 460; 1967: 217; 1969b: 352; 1979: 499; 1988: 873; Shumilo & Lunkashu, 1970: 31; Fain, Sixl & Moritsch, 1974: 2; Pence, 1975: 25; Sartbaev, 1983: 84; Rodriguez, Úbeda & Guevara, 1993: 13 ; Butenko, Stanyukovich & Dimov, 2010: 15; Dimov, 2018: 87

Hapalognatha prima Butenko, 1959: 17; 1960: 1490

Ptilonyssus motacilla: Butenko, 1965

Ptilonyssus estrildicola taeniopygiae Fain, 1963b: 74; Fain, Herin & Puylaert, 1977: 386

Ptilonyssus estrildicola Fain, 1959: 22; Domrow, 1965b: 453

Ptilonyssus fringillicola Fain, 1959: 28

Ptilonyssus cinnyricincti Fain, 1962: 132

Type locality: Astrida (= Butare), Rwanda

Distribution: Australia, Austria, Botswana, Kirghizstan, Moldavia, Russia, Rwanda, South Africa, Spain, Ukraine, USA

Hosts: Anthus australis, A. spinoletta, A. trivialis, Ficedula albicollis, Motacilla aguimp vidua, M. alba, M alba baicalensis, M. capensis wellsi, M. cinerea, M. citreola, M. flava, M. tschutschensis macronyx (Passeriformes: Motacillidae), Lonchura punctulata, L. [Eodice] malabarica, Taeniopygia castanotis (Passeriformes: Estrildidae), Pachycephala pectoralis, P. rufiventris (Passeriformes: Pachycephalidae), Petroica rosea (Passeriformes: Petroicidae), Cercomela familiaris modesta, Erithacus rubecula, Erythropygia coryphaeus, Luscinia svecica, Muscicapa striata, Myrmecocichla formicivora, Oenanthe isabellina, O. oenanthe, Phoenicurus erythronotus, Ph. motacillae (Passeriformes: Muscicapidae), Cyanosylvia svecica, Saxicola rubetra, S. torquata axillaris (Passeriformes: Turdidae), Mirafra africanoides (Passeriformes: Alaudidae), Acridotheres tristis (Passeriformes: Sturnidae), Panurus biarmicus (Passeriformes: Panuridae), Leucosticte nemoricola, Serinus pusillus (Passeriformes: Fringillidae), Scotocerca inquieta (Passeriformes: Cisticolidae), Perdix daurica (Galliformes: Phasianidae)

Ptilonyssus motacillae phoenicuri Fain

Ptilonyssus motacillae phoenicuri Fain, 1966: 90; Shumilo & Lunkashu, 1970: 32

Type locality: Strée (Hainaut), Belgium

Distribution: Belgium, Ukraine

Hosts: Phoenicurus ochruros, Ph. o. gibraltariensis (Passeriformes: Muscicapidae)

Ptilonyssus muscicapae Bregetova

Ptilonyssus muscicapae Bregetova, 1970: 59; Shumilo & Lunkashu, 1970: 32

Type locality: Ribachiy, Kaliningradskaya Obl. (Russia)

Distribution: Moldavia, Russia, Turkmenistan

Hosts: Muscicapa latirostris, M. sibirica, M. striata (Passeriformes: Muscicapidae)

Ptilonyssus muscicapoides Butenko et Lavrovskaya

Ptilonyssus muscicapoides Butenko & Lavrovskaya, 1980b: 351

Type locality: Russia

Distribution: Russia

Host: Ficedula hypoleuca (Passeriformes: Muscicapidae)

Ptilonyssus myiobii Fain et Aitken

Ptilonyssus myiobii Fain & Aitken, 1970: 356

Type locality: Mosqueiro Ferry, Benavides, Belém

Distribution: Brazil

Host: Myiobius barbatus (Passeriformes: Tyrannidae)

Ptilonyssus myrmotherulae Fain et Aitken

Ptilonyssus myrmotherulae Fain & Aitken, 1968: 16; 1970: 356

Type locality: APEG Forest, Belém

Distribution: Brazil

Host: Myrmotherula hauxwelli (Passeriformes: Formicariidae)

Ptilonyssus myzanthae Domrow

Ptilonyssus myzanthae Domrow, 1964d: 603, 617; 1969b: 367; 1979: 500; 1988: 873

Type locality: Condamine, Qu.

Distribution: Australia

Hosts: Anthochaera chrysoptera, Myzantha flavigula, M. melanocephala (Passeriformes: Meliphagidae)

Ptilonyssus myzomelae Domrow

Ptilonyssus myzomelae Domrow, 1965b: 455; 1969b: 370; 1979d: 502; 1988: 874

Type locality: Samford

Distribution: Australia

Hosts: Myzomela obscura, M. sanguinolenta (Passeriformes: Meliphagidae)

Ptilonyssus nadchatrami Fain

Ptilonyssus nadchatrami Fain, 1964c: 63

Type locality: Bukit Lagong, Selangor

Distribution: Malaysia

Hosts: Pycnonotus brunneus, P. e. erythrophthalmus (Passeriformes: Pycnonotidae)

Ptilonyssus neochmiae Domrow

Ptilonyssus neochmiae Domrow, 1969b: 341, 423; 1988: 874; 1992: 1605

Type locality: Innisfail

Distribution: Australia

Hosts: Neochmia albiventer, N. phaeton (Passeriformes: Ploceidae), Erythrura gouldiae, Poephila acuticauda (Passeriformes: Estrildidae)

Ptilonyssus nicatoris Fain

Ptilonyssus nicatoris Fain, 1958b: 294

Type locality: Kivu

Distribution: DR Congo

Host: Nicator chloris (Passeriformes: Pycnonotidae)

Ptilonyssus nitzschi (Giebel)(nomen dubium, see Fain, 1957d)

Dermanyssus nitzschi Giebel, 1871: 32

Neonyssus (Vitznyssus) nitzschi: Castro, 1948: 277

Ptilonyssus nitzschi: Zumpt & Till, 1955: 69; Spicer, 1978: 892

Type locality: ? Europe

Distribution: Australia, [Europe], USA

Hosts: Caprimulgus carolinensis, C. macrurus, Phalaenoptilus nuttallii (Caprimulgiformes: Caprimulgidae)

Ptilonyssus nivalis Knee

Ptilonyssus nivalis Knee, 2008: 359

Type locality: Oak Hammock Marsh, Manitoba

Distribution: Canada

Host: Plectrophenax nivalis (Passeriformes: Fringillidae)

Ptilonyssus novaeguineae (Hirst)

Rhinonyssus novae-guineae Hirst, 1921: 769

Passeronyssus novae-guineae (Hirst): Fain, 1960b: 314

Ptilonyssus novae-guineae (Hirst): Fain & Hyland, 1962: 346

Ptilonyssus novaeguineae (Hirst): Sakakibara, 1968; Domrow, 1969: 374; Feider & Mironescu, 1980: 61

Type locality: Papua New Guinea

Distribution: Papua New Guinea

Hosts: Paradisaea reggiana, Ptiloris magnificus, P. paradiseus, P. victoriae (Passeriformes: Paradisaeidae)

Ptilonyssus nucifragae (Hirst)

Rhinonyssus (Neonyssoides) nucifragae Hirst, 1923: 975; Bregetova, 1965: 133

Neonyssus nucifragae: Bregetova, 1953: 332; Fain, 1957: 52; Isakova, 1965: 89

Ptilonyssus nucifragae (Hirst): Fain, 1960b: 311

Neonyssoides nucifragae (Hirst): Bregetova, 1967: 133; Shumilo & Lunkashu, 1970:

Type locality: Heilsberg

Distribution: [East Prussia – formerly in Germany], Kirghizstan, Latvia, Russia, Ukraine

Hosts: Garrulus glandarius, Nucifraga caryocatactes, Perisoreus infaustus (Passeriformes: Corvidae)

Ptilonyssus nudus Berlese et Trouessart

Ptilonyssus nudus Berlese & Trouessart, 1889: 126; Berlese, 1892: 124; Oudemans, 1905: 237; Hirst, 1916: 73; Vitzthum, 1935: 578; Nordberg, 1936: 49; Castro & Pereira, 1947: 127; Bregetova, 1951: 118; Porter & Strandtmann, 1952: 393; Turk, 1953: 13; Zumpt & Till, 1955: 79; Fain, 1956a: 144; 1957b: 100; 1963: 171; George, 1961: 116; Butenko, 1965; Isakova, 1965: 82; 1968; Domrow, 1969: 372; Shumilo & Lunkashu, 1970: 33; Pence, 1975: 109; Fain, Herin & Pulyaert, 1977: 387; Knee, Proctor & Galloway, 2008: 369; Huhta, 2016: 148 (Sp. incert.), Dimov, 2018: 93

Type locality: Europe (France, Italy)

Distribution: [Australia], Cameroon, Canada, England, ?Finland, France, Germany, Holland, Italy, Moldavia, Portugal, Russia, Rwanda, USA

Hosts: Passer domesticus, P. griseus, P. montanus (Passeriformes: Passeridae); also there are published findings on Hypochera funerea nigerrima, Vidua macroura (Passeriformes: Viduidae)(Fain, 1956) and on Emberiza cirlus (Pass.: Emberizidae) (Berlese & Trouessart, 1889), and Fringilla coelebs (from Vitzthum, 1935), but Fain (1957b) thinks, that this information is doubtful. Also findings on Parus atricapilus, P. major (Passeriformes: Paridae), Sturnus vulgaris (Passeriformes: Sturnidae), Hirundo rustica, Riparia riparia (Passeriformes: Hirundinidae), Carpodacus roseus (Passeriformes: Fringillidae)

Ptilonyssus ohioensis Fain et Johnston

Ptilonyssus ohioensis Fain & Johnston, 1966: 29

Type locality: Ohio

Distribution: USA (Ohio)

Host: Cathartes aura (Cathartiformes: Cathartidae)

Ptilonyssus olaoi Pereira et Castro

Ptilonyssus (Ptilonyssus) olaoi Pereira & Castro, 1949: 219

Ptilonyssus olaoi Pereira et Castro: Zumpt & Till, 1955: 88; Fain, 1957: 104; Strandtmann, 1960: 146; Amaral, 1968: 121

Type locality: Boracéa (Estado de Sao Paulo)

Distribution: Brazil

Host: Ampelion cucullatus (Passeriformes: Cotingidae)

Ptilonyssus orientalis (Ewing)

Sommatericola orientalis Ewing, 1933: 13

Ptilonyssus orientalis (Ewing): Standtmann, 1956: 136

Type locality: Bangkok

Distribution: Thailand

Host: Lanius nigriceps longicaudatus (Passeriformes: Laniidae)

Ptilonyssus orioli Fain

Ptilonyssus oriolus Fain, 1956a: 144; 1962b: 269; Butenko, 1969; Shumilo & Lunkashu, 1970: 33

Type locality: Akanyaru (Rwanda)

Distribution: Belgium, Moldavia, Rwanda, Ukraine

Hosts: Oriolus larvatus angolensis, O. o. oriolus (Passeriformes: Oriolidae)

Ptilonyssus orthonychus Domrow

Ptilonyssus orthonychus Domrow, 1969b: 354; 1988: 874

Type locality: Wilson’s Peak, Queensland

Distribution: Australia

Host: Orthonyx temminckii (Passeriformes: Orthonychidae)

Ptilonyssus paradisaeus (Sakakibara)

Ptilonyssoides paradisaeus Sakakibara, 1968: 298

Ptilonyssus paradisaeus (Sakakibara)

Type locality: Morobe District, Wau, Nakata DRidge, 1515 m

Distribution: Papua New Guinea

Host: Paradisornis rudolphi (Passeriformes: Paradisaeidae)

Ptilonyssus paranensis (Castro)

Neonyssus (Travanyssus) paranensis Castro, 1948: 276; Pereira & Castro, 1949; Zumpt & Till, 1955: 90

Ptilonyssus (Travanyssus) paranensis (Castro): Pereira & Castro, 1949: 227

Ptilonyssus paranensis: Zumpt & Till, 1955: 90; Fain, 1957b: 100; Amaral, 1968: 120

Type locality: Estado de Parana

Distribution: Brazil

Host: Elaenia flavogaster (Passeriformes: Tyrannidae)

Ptilonyssus pari Fain et Hyland

Ptilonyssus pari Fain & Hyland, 1963a: 375; Fain & Bafort, 1963b: 485; Butenko, 1965: 131; Sixl, 1969: 218; Sixl & Reich, 1971: 401; Butenko & Stolbov, 1971; Pence, 1972c: 793; 1975: 90; Fain, Sixl & Moritsch, 1974: 2; Spicer, 1978: 892; Kadosaka et al., 1983; Rodriguez, Úbeda, Guevara & Ariza, 1989: 152; Dimov, 2016; 2018: 81

Neonyssus pari (Fain et Hyland): Shumilo & Lunkashu, 1970: 23

Type locality: Shiawansee, Michigan (USA), Fosse, etc. (Belgium)

Distribution: Austria, Belgium, Japan, Moldavia, Russia, Spain, Ukraine, USA (La., Nevada)

Hosts: Parus ater, P. atricapillus, P. atricapillus subrhenanus, P. caeruleus, P. carolinensis, P. cristatus mitratus, P. gambeli, P. insularis, P. m. major, P. montanus (Passeriformes: Paridae), Sitta europaea (Sittidae)

Ptilonyssus pastoris Butenko

Ptilonyssus pastoris Butenko, 1972: 1075

Type locality: Mihaylovka Vill., Dzhambulskiy Rayon, Kazakhstan

Distribution: Kazakhstan

Host: Pastor roseus (Passeriformes: Sturnidae)

Ptilonyssus pericrocoti (Sakakibara)

Paraneonyssus pericrocoti Sakakibara, 1967: 597

Ptilonyssus pericrocoti (Sakakibara)

Type locality: Liukuei

Distribution: Taiwan

Host: Pericrocotus roseus divaricatus (Passeriformes: Campephagidae)

Ptilonyssus perisorei George

Ptilonyssus perisorei George, 1961: 108; Spicer, 1978: 892; Wilson & Haas, 1980: 545

Type locality: Santa Fe, New Mexico

Distribution: Canada, USA (Alaska, New Mexico, Utah)

Hosts: Aphelocoma coerulescens, Cyanocitta stelleri, Perisoreus canadensis (Passeriformes: Corvidae)

Ptilonyssus phainopeplae George

Ptilonyssus phainopeplae George, 1961: 108; Pence, 1975: 25

Type locality: Val Verde Co., Texas

Distribution: USA (Texas)

Host: Phainopepla nitens (Passeriformes: Ptilogonatidae)

Ptilonyssus philemoni Domrow

Ptilonyssus philemoni Domrow, 1964d: 600, 617; 1965b: 460; 1967: 218; 1969b: 367; 1979: 502; 1988: 874

Ptilonyssus pentagonicus Fain & Lukoschus, 1979: 18; 1986: 60. Type loc.: Mount Hart (Western Australia), ex Manorina flavigula (syn. by Domrow, 1987)

Type locality: Logan Village, Condamine, Qu.

Distribution: Australia

Hosts: Entomyzon cyanotis; Manorina flavigula, Meliphaga gracilis, M. notata, Myzantha flavigula, Philemon citreogularis, P. corniculatis, Plectorhyncha lanceolata (Passeriformes: Meliphagidae)

Ptilonyssus phyllastrephi Zumpt et Till

Ptilonyssus phyllastrephi Zumpt & Till, 1955: 71; Fain, 1957b: 104

Type locality: Cape Prov.

Distribution: South Africa

Host: Phyllastrephus terrestris (Passeriformes: Pycnonotidae)

Ptilonyssus phylloscopi Fain

Ptilonyssus phylloscopi Fain, 1962b: 257; Isakova, 1965; Guevara-Benítez & Úbeda Ontíveros, 1980a: 231

Type locality: Anvers (Belgium)

Distribution: Belgium, Kirghizstan, Russia, Spain

Hosts: Phylloscopus collybita, Ph. griseolus, Ph. inornatus, Ph. t. trochilus (Passeriformes: Phylloscopidae)

Ptilonyssus pinicola Knee

Ptilonyssus pinicola Knee, 2008: 359

Type locality: Coyote Lake Nature Sanctuary, Alberta

Distribution: Canada

Host: Pinicola enucleator (Passeriformes: Fringillidae)

Ptilonyssus pipromorphae Fain et Aitken

Ptilonyssus pipromorphae Fain & Aitken, 1967: 13; 1968: 3; 1970: 355

Type locality: Morne Poui Road, Aripo Valley

Distribution: Brazil, Trinidad

Hosts: Pipromorpha macconnelli, P. oleaginea (Passeriformes: Tyrannidae)

Ptilonyssus pirangae (Černý)

Neonyssus (Paraneonyssus) pirangae Černý, 1969: 232; Černý & Dusbábek, 1970: 484

Ptilonyssus pirangae: Pence, 1972c: 793; 1973: 887; Knee, Proctor & Galloway, 2008: 368; Moodi, 2014: 3

Type locality: Havana, Botanical Garden

Distribution: Canada, Cuba, Iran

Hosts: Piranga ludoviciana, P. rubra rubra (Passeriformes: Thraupidae), Parus bicolor (Passeriformes: Paridae)

Ptilonyssus pittae Domrow

Ptilonyssus pittae Domrow, 1964a: 26; 1964d: 618; 1969d: 349; 1979: 495; 1988: 874

Type locality: Upper Brookfield, S.E. Queensland

Distribution: Australia

Host: Pitta versicolor (Passeriformes: Pittidae)

Ptilonyssus platypsaris Fain et Aitken

Ptilonyssus platypsaris Fain & Aitken, 1971: 34

Type locality: APEG Forest, Station A (IPEAN)

Distribution: Brazil

Host: Platypsaris minor (Passeriformes: Cotingidae)

Ptilonyssus platyrhinchus Fain et Aitken

Ptilonyssus platyrhinchus Fain & Aitken, 1970: 374

Type locality: Mosqueiro Ferry, Benavides, Belém

Distribution: Brazil

Host: Platyrhinchus saturatus (Passeriformes: Tyrannidae)

Ptilonyssus plesiotypicus Knee

Ptilonyssus plesiotypicus Knee, 2008: 367

Type locality: Alberta

Distribution: Canada

Host: Carpodacus purpureus (Passeriformes: Fringillidae)

Ptilonyssus ploceanus Fain

Ptilonyssus ploceanus Fain, 1956a: 139; 1957b: 89; Fain, Herin & Puylaert, 1977: 388; Lindholm, Venter & Ueckermann, 1998: 148

Type locality: Akanyaru (Rwanda)

Distribution: Cameroon, Rwanda, South Africa

Hosts: Anaplectes rubriceps leuconotus, Euplectes ardeus, E. hordeaceus, E. macrourus, E. orix, E. o. nigrifrons, Hyphanturgus ocularius, Ploceus cucullatus, P. vitellinus, Quelea erythrops, Textor atrogularis (Passeriformes: Ploceidae), Petronia dentata (Passeriformes: Passeridae), Serinus mozambicus (Passeriformes: Fringillidae); Chrysococcyx caprius (Cuculiformes: Cuculidae)

Ptilonyssus prunellae Fain et Bafort

Ptilonyssus prunellae Fain & Bafort, 1963b: 480; 1966: 7; Beron, 1975: 186; Guevara-Benítez & Úbeda Ontíveros, 1978a: 259; Sartbaev, 1983: 84

Type locality: Anvers, etc. (Belgium)

Distribution: Belgium, Bulgaria, Kirghizstan, Spain

Hosts: Prunella fulvescens, P. m. modularis (Passeriformes: Prunellidae)

Ptilonyssus psalidoprocnei Fain

Ptilonyssus psalidoprocnei Fain, 1956a: 146

Type locality: Astrida (= Butare)

Distribution: Rwanda

Host: Psalidoprocne albiceps (Passeriformes: Hirundinidae)

Ptilonyssus psaltriparus Spicer

Ptilonyssus psaltriparus Spicer, 1978: 892

Type locality: San Pablo Reservoir, Contra Costa Co., California

Distribution: USA (California)

Host: Psaltriparus minimus (Passeriformes: Paridae)

Ptilonyssus pseudothymanzae Fain et Lukoschus

Ptilonyssus pseudothymanzae Fain & Lukoschus, 1979: 15; Koch, 1986: 60; Domrow, 1988: 874

Type locality: Western Australia (The Kimberley)

Distribution: Australia (WA)

Hosts: Meliphaga flavescens, Melithreptus albogularis (Passeriformes: Meliphagidae)

Ptilonyssus psophodae Domrow

Ptilonyssus psophodae Domrow, 1964a: 29; 1964d: 618; 1969b: 366; 1988: 874

Type locality: Upper Brookfield, S.E. Queensland

Distribution: Australia

Host: Psophodes olivaceus (Passeriformes: Cinclosomatidae)

Ptilonyssus ptyonoprognes Butenko et Lavrovskaya

Ptilonyssus ptyonoprognes Butenko & Lavrovskaya, 1983: 116

Type locality: Kirghizstan

Distribution: Kirghizstan

Host: Ptyonoprogne rupestris (Passeriformes: Hirundinidae)

Ptilonyssus pycnonoti Fain

Ptilonyssus pycnonoti Fain, 1956a: 146; 1957b: 109; 1959a: 24; 1960: 109

Type locality: Astrida (= Butare)

Distribution: Rwanda, South Africa

Hosts: Pycnonotus barbarus tricolor, P. nigricans (Passeriformes: Pycnonotidae)

Ptilonyssus pycnonoti malayi Fain

Ptilonyssus pycnonoti malayi Fain, 1963: 78

Type locality: Bukit Mandol, Selangor

Distribution: Malaysia

Host: Pycnonotus plumosus (Passeriformes: Pycnonotidae)

Ptilonyssus pygiptilae Fain et Aiken

Ptilonyssus pygiptilae Fain & Aiken, 1971: 37

Type locality: Ananindeua

Distribution: Brazil

Host: Pygiptila stellaris (Passeriformes: Thamnophilidae)

Ptilonyssus pygmaeus (Bregetova)

Neonyssus (Spizonyssus) pygmaeus Bregetova, 1967: 131

Neonyssus pygmaeus Bregetova: Shumilo & Lunkashu, 1970: 22

Ptilonyssus pygmaeus (Bregetova): Domrow, 1969: 341

Type locality: Kaliningradskaya Onl. (Russia), 10 km S Penjikent

Distribution: Moldavia, Russia, Tadjikistan

Hosts: Carduelis carduelis, C. carduelis subcaniceps, Spinus spinus (Passeriformes: Fringillidae)

Ptilonyssus pyriglenae Fain et Aitken

Ptilonyssus pyriglenae Fain & Aitken, 1968: 18; 1970: 355

Type locality: Mucambo Forest

Distribution: Brazil

Host: Pyriglena leucoptera (Passeriformes: Formicariidae)

Ptilonyssus pyrrhulinus Stanyukovich et Butenko

Ptilonyssus pyrrhulinus Stanyukovich & Butenko, 2003: 38

Type locality: Ryazanskaya Oblast

Distribution: Russia

Host: Pyrrhula pyrrhula (Passeriformes: Fringillidae)

Ptilonyssus rabelloi (Castro)

Ptilonyssus (Flavionyssus) rabelloi Castro, 1948: 266; Pereira & Castro, 1949: 226

Ptilonyssus rabelloi (Castro): Zumpt & Till, 1955: 88; Fain, 1957: 109; Amaral, 1968: 120

Type locality: Juquiá (Estado de São Paulo)

Distribution: Brazil

Host: Sittasomus griseicapillus sylviellus (Passeriformes: Furnariidae, Dendrocolaptinae)

Ptilonyssus radovskyi Feider et Mironescu

Ptilonyssus radovskyi Feider & Mironescu, 1980: 55

Type locality: Wan Creek, Tambula

Distribution: Papua New Guinea

Hosts: Peneothello cyanus, Tregellasia leucops (Passeriformes: Petroicidae), Amblyornis macgregori (Passeriformes: Ptilonorhynchidae)

Ptilonyssus reguli Fain et Sixl

Ptilonyssus reguli Fain & Sixl, 1969: 264; Fain, Sixl & Moritsch, 1974: 4; Rodriguez, Úbeda & Guevara, 1990: 301

Type locality: Autal Graz (Austria)

Distribution: Austria, Spain

Hosts: Regulus ignicapillus, R. regulus (Passeriformes: Regulidae)

Ptilonyssus richmondenae George

Ptilonyssus richmondenae George, 1961: 122; Pence, 1975: 25; Hilario-Perez & Dowling, 2018: 296

Type locality: Liberty Hill, Texas

Distribution: USA (Texas)

Hosts: Richmondena cardinalis (Passeriformes: Fringillidae), Molothrus ater (Passeriformes: Icteridae)

Ptilonyssus rhipidurae Domrow

Ptilonyssus rhipidurae Domrow, 1966a: 199; 1969b: 366; 1988: 875

Type locality: Esk

Distribution: Australia

Host: Rhipidura fuliginosa (Passeriformes: Muscicapidae)

Ptilonyssus ripariae Stanyukovich et Butenko

Ptilonyssus ripariae Stanyukovich & Butenko, 2003: 35

Type locality: Pologrudino Village, Tyumenskaya Oblast

Distribution: Russia

Host: Riparia riparia (Passeriformes: Hirundinidae)

Ptilonyssus ruandae Fain

Ptilonyssus ruandae Fain, 1956b: 395; 1957b: 109; 1960: 110; Strandtmann & Wharton, 1958: 182; Domrow, 1964d: 608; 1969b: 366; 1988: 875; Kadosaka, Kaneko & Asanuma, 1987: 33

Ptilonyssus ruandae alcippei Fain & Nadchatram, 1962: 279; Nadchatram, McClure

& Chong, 1964: 104 – Type: Malaysia, Mont Brinchang (1700 m), Pahang, from Alcippe nipalensis peracensis; McClure, Ratanaworadhan et al., 1973: 35

Type locality of Ptilonyssus ruandae Fain: Akanyaru (Rwanda)

Distribution: Australia, Botswana, Japan, Malaysia, Rwanda

Hosts: Parisoma sucaeruleum (Passeriformes: Sylviidae), Zosterops lateralis, Z. palpebrosa japonica (Passeriformes: Zosteropidae), Alcippe nipalensis peracensis (Passeriformes: Timaliidae)

Ptilonyssus ruandae sylviae Fain

Ptilonyssus ruandae sylviae Fain, 1963a: 172; Fain, Sixl & Moritsch, 1974: 2

Type locality: Clermont (Namur), Strée (Hainaut)

Distribution: Austria, Belgium

Hosts: Sylvia a. atricapilla, S. c. communis (Passeriformes: Sylviidae)

Ptilonyssus sairae Castro

Ptilonyssus (Ptilonyssus) sairae Castro, 1948: 260; Pereira & Castro, 1949: 219 Ptilonyssus sairae: Zumpt & Till, 1955: 89; Fain, 1956a: 144; 1957b: 102; George, 1961: 125; Fain & Aitken, 1967: 4; 1968: 3; Amaral, 1968: 120; Spicer, 1977a: 598; 1977b: 498; 1978: 892; 1984: 799; 1987: 261; Kadosaka, Kaneko & Asanuma, 1987: 35; Morelli & Spicer, 2007: 183; Mascarenhas, Coimbra et al., 2011: 507; Knee & Galloway, 2017: 94–95; Dimov, 2016; 2018: 89

Ptilonyssus constrictus Ford, 1961: 5 – type from Dendroica petechia, Seiurus noveboracensis (Passeriformes: Parulidae), Trinidad, USA (syn. by Pence, 1973b: 877)

Ptilonyssus constrictus longisetosus Černy & Dusbábek, 1970: 481 – type from Seiurus motacilla (Passeriformes: Parulidae), Havana, Cuba

Ptilonyssus garridoi Cruz, 1971: 1

Type locality of Ptilonyssus (Ptilonyssus) sairae Castro: Brazil

Distribution: Brazil, Canada, Cuba, Guatemala, Japan, Mexico, Russia, Trinidad, USA (incl. Alasca)

Hosts: Emberiza schoeniclus (Passeriformes: Emberizidae), Aimophila ruficeps, Ammodramus savannarum, Arremon taciturnus, Chondestes grammacus, Junco caniceps, J. hyemalis, Melospiza melodia, Paroaria coronata, Pooecetes gramineus, Spizella passerina, S. pallida, S. pusilla, Zonotrichia querula (Passeriformes: Passerellidae), Chlorospingus ophthalmicus, Tangara gyrola, T. seledon, Tiaris fuliginosa (Passeriformes: Thraupidae), Dendroica auduboni, D. coronata, D. erithacorides, D. petechia, D. pinus, Geothlypis philadelphia, Muniotitla varia, Oreothlypis peregrina, O. ruficapilla, Parula americana, Seiurus noveboracensis, Setophaga pensylvanica, S. fusca, Vermivora chrysoptera, V. celata, V. ruficapilla, Wilsonia pusilla (Passeriformes: Parulidae), Passerina ciris, Guiraca caerulea (Passeriformes: Cardinalidae), Parus carolinensis, P. major (Passeriformes: Paridae), Carduelis pinus, Carduelis [Spinus] tristis, Carduelis chlorus, Chloris sinica minor (Passeriformes: Fringillidae), Polioptila albiloris (Passeriformes: Polioptilidae), Euphagus cynocephalus, Molothrus ater (Passeriformes: Icteridae)

P. S. Pence & Casto (1976) consider Ptilonyssus japuibensis Castro, P. japuibensis cyanocompsae Fain et Aitken, P. agelaii Fain, P. teretistris Černy, P. ludovicianus Černy and P. garridoi Cruz, but this was not followed by the Canadian and some other authors.

Ptilonyssus salpinctis George

Ptilonyssus salpinctis George, 1961: 116

Type locality: Brisko Co., Texas

Distribution: USA (Texas)

Host: Salpinctes obsoletus (Passeriformes: Troglodytidae)

Ptilonyssus saltator Fain et Aiken

Ptilonyssus saltator Fain & Aiken, 1971: 37

Type locality: Ananindeua

Distribution: Brazil

Host: Saltator maximus (Passeriformes: Fringillidae)

Ptilonyssus schoenobaeni Fain, Sixl et Moritsch

Ptilonyssus schoenobaeni Fain, Sixl & Moritsch, 1974: 4

Type locality: Illmitz, Burgenland

Distribution: Austria

Host: Acrocephalus schoenobaenus (Passeriformes: Acrocephalidae)

Ptilonyssus schumili Butenko et Lavrovskaya

Ptilonyssus schumili Butenko & Lavrovskaya, 1980: 522; Scherbinina & Butenko, 1981: 70; Dimov, 2016; 2018: 86

Type locality: Ryazanskaya Obl., Okskiy Zap.

Distribution: Russia, Kazahstan, Kirghizstan, Turkmenistan

Hosts: Alauda arvensis, Galerida cristata (Passeriformes: Alaudidae)

Ptilonyssus sclateriae Fain et Aitken

Ptilonyssus schoenobaeni Fain & Aitken, 1970: 401

Type locality: Mucambo Forest, IPEAN, Belém

Distribution: Brazil

Host: Sclateria naevia (Passeriformes: Thamnophilidae)

Ptilonyssus sclerurus Fain et Aitken

Ptilonyssus sclerurus Fain & Aitken, 1970: 395

Type locality: Pirelli Plantation, Marituba

Distribution: Brazil

Host: Sclerurus rufigularis (Passeriformes: Furnariidae)

Ptilonyssus serini Fain

Ptilonyssus serini Fain, 1956a: 139; 1957b: 90

Type locality: Akanyaru, Rwanda

Distribution: Rwanda

Hosts: Serinus canicollis sassii, S. citrinelloides frontalis, S. mozambicus barbatus (Passeriformes: Fringillidae)

Ptilonyssus setosae Domrow

Ptilonyssus setosae Domrow, 1969b: 362; 1988: 875

Type locality: Ella Bay, Queensland

Distribution: Australia

Hosts: Rhipidura rufifrons, R. setosa (Passeriformes: Muscicapidae)

Ptilonyssus sialiae George

Ptilonyssus sialiae George, 1961: 122

Type locality: Lubbock Co., Texas

Distribution: USA (Texas)

Host: Sialia currucoides (Passeriformes: Turdidae)

Ptilonyssus sittae Fain

Ptilonyssus sittae Fain, 1965: 158; 1966: 117; Domrow, 1969b: 348; 1988: 875; Butenko, 1969; Sixl, 1969: 218; Shumilo & Lunkashu, 1970: 33; Sixl & Reich, 1971: 401; Fain, Sixl & Moritsch, 1974: 3

Ptilonyssus strandtmannianus Feider & Mironescu, 1969: 644 (syn. by Domrow, 1972: 575)

Ptilonyssus bregetovae Feider & Mironescu, 1969: 656 (syn. by Domrow, 1972: 575)

Ptilonyssus pelmaspis Feider & Mironescu, 1969: 656 (syn. by Domrow, 1972: 575)

Ptilonyssus maxvachoni Feider & Mironescu, 1970: 17 (syn. by Domrow, 1972: 575)

Type locality: Kappellen, nr. Antwerp (Belgium)

Distribution: Australia, Austria, Belgium, Moldavia, Romania, Ukraine

Hosts: Climacteris picumnus (Passeriformes: Climacteridae), Sitta europaea (Passeriformes: Sittidae), Parus major (Passeriformes: Paridae)

Ptilonyssus souzai Pereira et Castro

Ptilonyssus (Rhinonyssoides) souzai Pereira & Castro, 1949: 84; Zumpt & Till, 1955: 88

Ptilonyssus souzai: Fain, 1957b: 84; Fain & Johnston, 1966: 29

Type locality: Brazil

Distribution: Brazil

Hosts: Milvago chimachima chimachima (Falconiformes: Falconidae, Polyborinae), Rupornis magnirostris magniplumis (Accipitriformes: Accipitridae)

Ptilonyssus sphecotheris Domrow

Ptilonyssus sphecotheris Domrow, 1964d: 613; 1967: 218; 1969b: 372; 1988: 875

Type locality: Brisbane, Qu.

Distribution: Australia

Hosts: Chlamydera nuchalis (Passeriformes: Ptilonorhynchidae), Sphecotheres flaviventris, S. vieillotii (Passeriformes: Oriolidae)

Ptilonyssus spini Stanyukovich et Butenko

Ptilonyssus spini Stanyukovich & Butenko, 2003: 34; Dimov, 2018: 92

Type locality: Ribachiy, Kaliningradskaya Oblast

Distribution: Russia (Kaliningrad)

Host: Spinus spinus (Passeriformes: Fringillidae)

Ptilonyssus spinosus (Brooks et Strandtmann)

Tyranninyssus spinosus Brooks & Strandtmann, 1960: 421; Hyland, 1961: 279; Černý & Dusbábek, 1970: 484

Ptilonyssus spinosus (Brooks et Strandtmann): Mendes et al., 2014: 480

Type locality: Lubbock County, Texas

Distribution: Brazil, Cuba (incl. Pinos), Puerto Rico, USA (Michigan, Texas)

Hosts: Machetornis rixosa, Muscivora forficata, Pitangus sulphuratus, Tyrannus c. caudifasciatus, T. c. flavescens, T. dominicensis, T. melancholicus, T. tyranni, T. tyrannus, T. verticalis (Passeriformes: Tyrannidae)

Ptilonyssus steganurae Fain

Ptilonyssus steganurae Fain, 1967: 150

Type locality: Zoo Anvers

Distribution: Central Africa

Host: Vidua [Steganura] paradisea (Passeriformes: Viduidae)

Ptilonyssus sternostomicus Fain et Aitken

Ptilonyssus sternostomicus Fain & Aitken, 1968: 20; 1970: 353

Type locality: APEG Forest

Distribution: Brazil

Host: Schiffornis turdinus (Passeriformes: Pipridae)

Ptilonyssus stomioperae Domrow

Ptilonyssus stomioperae Domrow, 1966a: 206; 1967a: 217; 1969b: 317; 1979d: 504; 1988: 876

Type locality: Magnificent Creek, Mitchell River

Distribution: Australia

Hosts: Meliphaga flava, Stomiopera unicolor (Passeriformes: Meliphagidae)

Ptilonyssus strandtmanni Fain

Ptilonyssus strandtmanni Fain, 1956a: 148; 1957b: 86; Fain, Herin & Pulyaert, 1977: 387; Guevara-Benítez & Úbeda Ontíveros, 1980a: 231; Sartbaev, 1983: 84; Butenko, 1984: 157

Type locality: Astrida (= Butare), Rwanda

Distribution: Cameroon, Kirghizstan, Russia, Rwanda, Spain

Hosts: Apus affinis, A. apus, A. caffer streubeli (Apodiformes: Apodidae)

Ptilonyssus stresemanni Vitzthum

Ptilonyssus stresemanni Vitzthum, 1935: 579; Castro, 1948: 260; Pereira & Castro, 1949: 219; Zumpt & Till, 1955: 89; Fain, 1957b: 101; 1961a: 511

Type locality: Brazil

Distribution: Brazil

Host: Procnias albus (Passeriformes: Cotingidae)

Ptilonyssus struthideae Domrow

Ptilonyssus struthideae Domrow, 1969d: 343; 1988: 876

Type locality: Condamine, Charleville, Qu.

Distribution: Australia

Host: Struthidea cinerea (Passeriformes: Grallinidae)

Ptilonyssus sturnopastoris Fain

Ptilonyssus sturnopastoris Fain, 1963d: 1; Domrow, 1969b: 372; 1988: 876

Type locality: N. Queensland (described from Anvers, Belgium)

Distribution: Australia

Hosts: Aplonis metallica, Sturnopastor capensis (Passeriformes: Sturnidae)

Ptilonyssus sylviicola Stanyukovich et Butenko

Ptilonyssus sylviicola Stanyukovich & Butenko, 2003: 40

Type locality: Tatarstan

Distribution: Russia (Tatarstan)

Host: Sylvia communis (Passeriformes: Fringillidae)

Ptilonyssus tachycinetae George

Ptilonyssus tachycinetae George, 1961: 112; Butenko, 1965: 132

Type locality: Florida City, Florida

Distribution: Russia, USA (Florida)

Hosts: Riparia riparia, Tachycineta bicolor (Passeriformes: Hirundinidae)

Ptilonyssus terenotricci Fain et Aitken

Ptilonyssus terenotricci Fain & Aitken, 1970: 357

Type locality: Mosqueiro Ferry, Benavides, Belém

Distribution: Brazil

Host: Terenotriccus erythrurus (Passeriformes: Tyrannidae)

Ptilonyssus teretistris Černý

Ptilonyssus teretistris Černý, 1969: 227; Černý & Dusbábek, 1970: 482

Type locality: El Dorado, prov. of Las Villas

Distribution: Cuba

Host: Teretistris fornsi (Passeriformes: Parulidae)

Ptilonyssus terpsiphonei Fain

Ptilonyssus terpsiphonei Fain, 1956a: 145; 1957b: 137; 1959a: 22; Strandtmann & Wharton, 1958: 182; Domrow, 1965b: 453; 1969b: 356; 1988: 876; Sakakibara, 1968b: 309

Type locality: Astrida (= Butare), Rwanda

Distribution: Australia, Papua New Guinea, Rwanda, South Africa

Hosts: Carterornis leucotis, Myagra rubecula, Piesorhynchus alecto, Seisura inquieta (Passeriformes: Muscicapidae), Arses kaupi, Monarcha melanopsis, M. trivirgata, Terpsiphone viridis, T. viridis kivuensis (Passeriformes: Monarchidae)

Ptilonyssus thamnomanes Fain et Aitken

Ptilonyssus thamnomanes Fain & Aitken, 1970: 374

Type locality: Mosqueiro Ferry, Benavides, Belém

Distribution: Brazil

Host: Thamnomanes caesius (Passeriformes: Formicariidae)

Ptilonyssus thamnophili Fain et Aitken

Ptilonyssus thamnophili Fain & Aitken, 1967: 9; 1970: 356

Type locality: Elgin Trace, Vega de Oropouche (Trinidad)

Distribution: Brazil, Trinidad

Hosts: Thamnophilus aethiops, Th. doliatus (Passeriformes: Thamnophilidae), Platyrinchus saturatus (Passeriformes: Tyrannidae)

Ptilonyssus thryothori Pence

Ptilonyssus thryothori Pence, 1972d: 1165; 1973b: 876

Type locality: Laplace, La.

Distribution: USA (Louisiana)

Host: Thryothorus ludovicianus (Passeriformes: Troglodytidae)

Ptilonyssus thymanzae Domrow

Ptilonyssus thymanzae Domrow, 1964d: 604, 617; 1966a: 204, 1969b: 317; 1979: 505; 1988: 876

Ptilonyssus sp. Domrow, 1965b: 548

Type locality: Condamine, Qu.

Distribution: Australia

Hosts: Acanthagenys rufogularis, Anthochaera chrysoptera, Meliphaga chrysops, M. fusca, M. lewinii, M. melanops, M. notata, M. penicillata, Myzantha flavigula, M. melanocephala (Passeriformes: Meliphagidae)

Ptilonyssus tillae Fain

Ptilonyssus tillae Fain, 1959a: 25

Type locality: Sabi Sand Wildtuin, near Newington, Transvaal

Distribution: Rwanda, South Africa

Hosts: Lamprocolius chalybeus, Lamprotornis australis, L. mevesii (Passeriformes: Sturnidae)

Ptilonyssus toxostomae Pence

Ptilonyssus toxostomae Pence, 1972d: 1163; 1973b: 876; Spicer, 1987: 261

Type locality: Louisiana, Covington

Distribution: USA (Louisiana)

Host: Toxostoma rufum (Passeriformes: Mimidae)

Ptilonyssus traubi Strandtmann

Ptilonyssus traubi Strandtmann, 1960: 144

Type locality: Kinabalu Mt., Tenompak (Sabah)

Distribution: Malaysia (Sabah)

Host: Aves indet.

Ptilonyssus travassosfilhoi (Castro)

Neonyssus (Paraneonyssus) travassosfilhoi Castro, 1948: 275

Ptilonyssus (Paraneonyssus) travassosfilhoi (Castro): Pereira & Castro, 1949: 227

Ptilonyssus travassosfilhoi (Castro): Zumpt & Till, 1955: 88 ; Fain, 1957b: 88; Fain & Aitken, 1970: 352

Type locality: Juquiá (Estado de São Paulo)

Distribution: Brazil

Hosts: Philydor lichtensteini (Passeriformes: Furnariidae, Dendrocolaptinae), Automolus infuscatus (Passeriformes: Furnariidae)

Ptilonyssus triscutatus (Vitzthum)

Ptilonyssoides triscutatus Vitzthum, 1935: 581; Strandtmann & Wharton, 1958: 170; Shumilo & Lunkashu, 1970: 28; Butenko, 1984: 161

Neonyssus (Ptilonyssoides) triscutatus (Vitzthum): Castro, 1948: 276

Neonyssus triscutatus (Vitzthum): Zumpt & Till, 1955: 68

Ptilonyssus triscutatus Fain, 1956b: 136; 1957b: 121; Nadchatram, McClure & Chong, 1964: 104; Domrow, 1966b: 196; 1967: 217; 1969b: 347; 1979: 494; 1988: 877; Fain, Herin & Puylaert, 1977: 386; Scherbinina & Butenko, 1981: 70

Type locality: Europe

Distribution: Australia, Cameroon, Kazakhstan, Malaysia, Moldavia, Russia, Rwanda, Turkmenistan, Ukraine

Hosts: Melittophagus bullocki, Merops apiaster, M. nubicus, M. ornatus, M. persicus, M. superciliosus, M. viridis (Coraciiformes: Meropidae)

Ptilonyssus troglodytis Fain

Ptilonyssus troglodytis Fain, 1964a: 60; Knee, Proctor & Galloway, 2008: 370

Type locality: Bois de Oisquercq (Brabant)

Distribution: Belgium, Canada

Host: Troglodytes troglodytes (Passeriformes: Troglodytidae)

Ptilonyssus trouessarti (Hirst)

Rhinonyssoides trouessarti Hirst, 1921: 770; Strandtmann & Wharton, 1958: 172; syn. with Ptilonyssus by Zumpt & Till, 1955: 69

Ptilonyssus trouessarti: Zumpt & Till, 1955: 69; Fain & Hyland, 1962: 346; Domrow, 1964d: 608; 1969b: 373; 1979: 510; 1988: 877; Sakakibara, 1968b: 302

Type locality: Australia

Distribution: Australia, Papua New Guinea

Hosts: Oriolus flavocinctus, O. sagittatus; type from «Sphecotheres maxillaris» (Passeriformes: Oriolidae)

Ptilonyssus trouessarti pseudotrouessarti Sakakibara

Ptilonyssus trouessarti pseudotrouessarti Sakakibara, 1968b: 305

Type locality: Bulolo River

Distribution: Papua New Guinea

Host: “Leatherneck” (Tropidorhynchus sordidus – Meliphagidae)

Ptilonyssus tyrannus (Brooks et Strandtmann)

Tyranninyssus tyrannus Brooks & Strandtmann, 1960: 419; Hyland & Moorhouse, 1970: 48

Tyranninyssus tyrannus caribaeus Černý, 1969: 233. Type: Cuba, Santo Tomas, Cienaga de Zapata, prov. Las Villas, from Contopus caribaeus caribaeus

Tyranninyssus tyrannisoides Brooks & Strandtmann, 1960: 420. Type: Lamb County, Texas, from Empidonax minimus, Sayornis sayus (Passeriformes: Tyrannidae). Syn. by Hyland & Moorhouse, 1970: 48

Ptilonyssus tyrannus Pence, 1972: 790; Knee, Proctor & Galloway, 2008: 370; Knee & Galloway, 2017: 95

Type locality: Lamb County, Texas

Distribution: Guatemala, Mexico, Canada, USA (Texas)

Hosts: Contopus sordlidulus, C. caribaeus caribaeus, Empidonax alnorum, E. flavescens, E. flaviventris, E. minimus, E. traillii, Mitrephanes phaeocercus, Nuttallornis borealis, Sayornis phoebe, S. sayus, Tyrannus melancholicus (Passeriformes: Tyrannidae)

Ptilonyssus viduicola Fain

Ptilonyssus viduicola Fain, 1962c: 135; Fain, Herin & Puylaert, 1977: 388

Type locality: Akanyaru (Rwanda)

Distribution: Cameroon, Rwanda

Host: Vidua macroura (Passeriformes: Viduidae)

Ptilonyssus vireonis (Dusbábek)

Passeronyssus vireonis Dusbábek, 1969: 220; Černy & Dusbábek, 1970: 486

Ptilonyssus vireonis (Dusbábek): Pence, 1972c: 793; Knee, Proctor & Galloway, 2008: 370

Type locality: Havana

Distribution: Canada, Cuba, USA (La.)

Hosts: Vireo o. olivaceus, V. solitarius (Passeriformes: Vireonidae)

Ptilonyssus vossi Spicer

Ptilonyssus vossi Spicer, 1977a: 595; 1987: 261

Type locality: Fort Worth, Tarrant Co., Texas

Distribution: USA (Texas)

Host: Troglodytes aedon (Passeriformes: Troglodytidae)

Ptilonyssus walterberghi Fain

Ptilonyssus walterberghi Fain, 1958: 3

Type locality: Anvers Zoo

Distribution: Japan

Host: Suthora webbiana (Passeriformes: Paridae)

Ptilonyssus werneri (Castro)

Neonyssus (Rochanyssus) werneri Castro, 1948: 272; Pereira & Castro, 1949: 226

Ptilonyssus (Rochanyssus) werneri (Castro): Pereira & Castro, 1949: 226

Ptilonyssus werneri (Castro): Zumpt & Till, 1955: 90; Fain, 1957b: 86; Fain & Aitken, 1967: 3; 1970: 355; Amaral, 1968: 120

Type locality: Botucatû (Estado de São Paulo)

Distribution: Brazil, Trinidad

Hosts: Elaenia flavogaster, Е. obscura sordida (Passeriformes: Tyrannidae), Gnorimopsar chopi chopi, Sturnella [Leistis militares] superciliaris (Passeriformes: Icteridae)

Ptilonyssus wilsoni Sakakibara

Ptilonyssus wilsoni Sakakibara, 1967: 599; Tenorio, 1976: 19; Domrow & Nadchatram, 1978: 87

Type locality: Enarotali, NW New Guinea

Distribution: Malaysia, Papua New Guinea

Hosts: Apus affinis, Collocalia esculenta (Apodiformes: Apodidae)

Ptilonyssus xenops Fain et Aitken

Ptilonyssus xenops Fain & Aitken, 1968: 8; 1971: 30

Type locality: APEG Forest

Distribution: Brazil

Host: Xenops minutus (Passeriformes: Furnariidae)

Ptilonyssus xyphorhynchus Fain et Aitken

Ptilonyssus xyphorhynchus Fain & Aitken, 1970: 369

Type locality: Mosqueiro Ferry, Benavides, Belém

Distribution: Brazil

Host: Xyphorhynchus spixii (Passeriformes: Dendrocolapidae)

Ptilonyssus zeferinoi Amaral

Ptilonyssus zeferinoi Amaral, 1968: 107

Type locality: Estado de São Paulo

Distribution: Brazil

Hosts: Gnorimopsar chopi chopi, Sturnella [Leistis militares] superciliaris (Passeriformes: Icteridae)

Ptilonyssus zumpti Fain

Ptilonyssus zumpti Fain, 1956b: 138; 1957b: 91

Type locality: Akanyaru (Rwanda)

Distribution: Rwanda

Host: Amblyospiza albifrons montana (Passeriformes: Ploceidae)

Genus Sternostoma Berlese et Trouessart

Sternostoma Berlese & Trouessart, 1889: 126

Type species: Sternostoma cryptorhynchum Berlese et Trouessart, 1889

Sternostomum Vitzthum, 1935: 569

Type species: Sternostomum technaui Vitzthum, 1935

Agapornyssus Gretillat et al., 1959: 376

Type species: Agapornyssus faini Grétillat, Capron & Brygoo, 1959

Sternostomoides Bregetova, 1965: 709 – syn. by Domrow, 1969: 374

Type species: Sternostomum technaui Vitzthum, 1935

Rhinosterna Fain, 1964: 125 – syn. by Domrow, 1969: 374

Type species: Rhinosterna aymarae Fain, 1964

Sternoecius Fain et Aitken, 1967: 24 – syn. by Domrow, 1969: 374

Type species: Sternoecius piprae Fain et Aitken, 1967

Sternostoma alexfaini Feider et Mironescu

Sternostoma alexfaini Feider & Mironescu, 1969: 32

Type locality: Sinaia

Distribution: Romania

Host: Lanius excubitor (Passeriformes: Laniidae)

Sternostoma artami Feider et Mironescu

Sternostoma artami Feider & Mironescu, 1982: 87

Type locality: Papua New Guinea

Distribution: Papua New Guinea

Host: Artamus leucorhynchus (Passeriformes: Artamidae)

Sternostoma augei Amaral

Sternostoma augei Amaral, 1962: 69; Pence & Casto, 1975b: 362

Type locality: Itaporanga (Estado do São Paulo)

Distribution: Brazil, USA (Texas)

Host: Speotyto cunicularia grallaria (Strigiformes: Strigidae)

Sternostoma aymarae (Fain)

Rhinosterna aymarae Fain, 1964e: 126

Sternostoma aymarae (Fain): Domrow, 1969: 374

Type locality: Argentina

Distribution: Argentina

Host: Amoropsittaca aymara (Psittaciformes: Psittacidae)

Sternostoma batis Fain

Sternostoma laniorum var. batis Fain, 1957b: 77

Sternostoma batis Fain: Butenko, 1965; Fain & Aitken, 1968: 22; 1970: 349; Sartbaev, 1983: 84; Úbeda, Rodríguez & Guevara, 2000

Type locality: Akanyaru (Rwanda)

Distribution: Brazil, Kirghizstan, Russia, Rwanda, Spain

Hosts: Alauda arvensis, Eremophila alpestris (Passeriformes: Alaudidae), Batis molitor puella, Oenanthe oenanthe (Passeriformes: Muscicapidae), Xiphorhynchos spixii (Passeriformes: Furnariidae, Dendrocolaptinae), Pyriglena leucoptera (Passeriformes: Furnariidae), Loxia curvirostra (Passeriformes: Fringillidae)

Sternostoma boydae Strandtmann (nom. emend.)

Sternostoma boydi Strandtmann, 1951: 138 (named after Elizabeth Boyd); Zumpt & Till, 1955: 86; Fain, 1956a: 151; 1957b: 66; 1960c: 95; 1963a: 176; Fain & Aitken, 1967: 16; Fain & Holland, 1972: 32; Pence, 1972: 781; Feider & Mironescu, 1972: 326; Spicer, 1978: 894; Sartbaev, 1983: 84; Butenko, 1984: 84; Knee, 2008: 9

Type locality: Galveston, Texas, USA

Distribution: Australia, Belgium, Canada, Kazakhstan, Kirghizstan, Russia (incl. Far East), Trinidad, USA, Venezuela, Zambia

Hosts: Actitis macularia, Arenaria interpres, Calidris alba, Crocethia alba, Ereunetes pusillus, Limosa lapponica, Numenius arquatus, Phalaropus lobatus, Philomachus pugnax, Tringa [Totanus] erythropus, T. nebularia, T. ochropus, T. totanus, Rhyacophilus glareola (Charadriiformes: Scolopacidae), Chlidonias leucoptera, Onychophon [Sterna] fuscatus fuscatus, Sterna hirundo (Charadriiformes: Sternidae), Larus atricilla, L. californicus, L. delawarensis, L. glaucescens, L. occidentalis (Charadriiformes: Laridae)

Sternostoma bruxellarum Fain

Sternostoma bruxellarum Fain, 1961b: 53; Butenko, 1965; Sixl, 1969; Shumilo & Lunkashu, 1970: 24; Beron, 1975: 7

Type locality: Liège Prov. and Brussels

Distribution: Austria, Belgium, Bulgaria, Moldavia, Russia, Ukraine

Host: Sturnus vulgaris (Passeriformes: Sturnidae)

Sternostoma callithrix Fain et Aitken

Sternostoma callithrix Fain & Aitken, 1970: 343

Type locality: Ananindeua, Para

Distribution: Brazil

Host: Empidonomus varius (Passeriformes: Tyrannidae)

Sternostoma christinae Guevara – Benítez, Lopez-Roman et Úbeda – Ontíveros

Sternostoma christinae Guevara – Benítez, Lopez-Roman & Úbeda – Ontíveros, 1974: 161

Type locality: Granada

Distribution: Spain

Host: Phylloscopus collybita (Passeriformes: Phylloscopidae)

Sternostoma cisticolae Fain

Sternostoma cisticolae Fain, 1957c: 218; 1959a: 32

Type locality: Debeete

Distribution: Botswana, South Africa

Hosts: Apalis flavida, Cisticola chiniana (Passeriformes: Cisticolidae)

Sternostoma clementei Amaral

Sternostoma clementei Amaral, 1968: 114

Type locality: Boracéa, Estado de Sao Paulo

Distribution: Brazil

Host: Knipolegus cyanirostris (Passeriformes: Tyrannidae)

Sternostoma colii Fain

Sternostoma colii Fain, 1956b: 392; 1957b: 79

Type locality: Astrida (= Butare)

Distribution: Rwanda

Host: Colius striatus kiwuensis (Coliiformes: Coliidae)

Sternostoma constricta Feider et Mironescu

Sternostoma constricta Feider & Mironescu, 1982: 87

Type locality: Papua New Guinea

Distribution: Papua New Guinea

Host: Paradisaea reggiana (Passeriformes: Paradisaeidae)

Sternostoma cooremani Fain

Sternostoma cooremani Fain, 1956a: 154; 1957b: 72; Strandtmann & Wharton, 1958: 166; Domrow, 1965b: 448; 1966b: 192; 1969b: 376; Shumilo & Lunkashu, 1970: 24; Fain, Herin & Puylaert, 1977: 392; Sartbaev, 1981: 130; Butenko, 1984: 151

Type locality: Astrida (= Butare), Rwanda

Distribution: Australia, Cameroon, Kirghizstan, Malaysia, Moldavia, Russia, Rwanda

Hosts: Dacelo gigas, Melittophagus lafresnayi oreobates, Merops apiaster, M. nubicoides, M. orientalis, M. ornatus (Coraciiformes: Meropidae)

Sternostoma cooremani halcyoni Fain et Nadchatram

Sternostoma cooremani halcyoni Fain & Nadchatram, 1962: 280

Sternostoma cooremani: McClure, Ratanaworadhan et al., 1973: 37

Type locality: Rantau Panjang, Selangor

Distribution: Malaysia

Host: Halcyon chloris humii (Coraciiformes: Halcyonidae)

Sternostoma cordiscutata Feider et Mironescu

Sternostoma cordiscutata Feider & Mironescu, 1982: 87

Type locality: Papua New Guinea

Distribution: Papua New Guinea

Host: Chalcites meyeri (Cuculiformes: Cuculidae)

Sternostoma crotophagae Pence et Casto

Sternostoma crotophagae Pence & Casto, 1975: 364

Type locality: La Salle Co., Millett, Texas

Distribution: USA (Texas)

Host: Crotophaga sulcirostris (Cuculiformes: Cuculidae)

Sternostoma cryptorhynchum Berlese et Trouessart

Sternostoma cryptorhynchum Berlese & Trouessart, 1889: 128; Vitzthum, 1935: 571; Castro, 1948: 257; Zumpt & Till, 1955: 86; Fain, 1956a: 152; 1957b: 70; Domrow, 1969: 379; Fain, Hehn & Puylaert, 1977: 392; Knee, Proctor & Galloway, 2008: 369

Type locality: Europe

Distribution: France, Cameroon, Canada

Hosts: Clyptospiza monteiri (Passeriformes: Estrildidae), Passer domesticus (Passeriformes: Passeridae), Pinicola enucleator (Passeriformes: Fringillidae)

Sternostoma cuculorum Fain

Sternostoma cuculorum Fain, 1956a: 155; 1957b: 74; 1957c: 218; Strandtmann & Wharton, 1958: 166; Domrow, 1965b: 449; 1969b: 376; 1979: 510; Shumilo & Lunkashu, 1970: 25; McClure, Ratanaworadhan et al., 1973: 37; Butenko, 1984: 155

Sternostoma cuculorum var. urolestis Fain, 1959a: 32 – South Africa, ex Urolestes melanoleucus (Laniidae)

Sternostoma laniorum Fain: Strandtmann & Wharton, 1958: 167; Domrow, 1965b: 449; 1966b: 193; Shumilo & Lunkashu, 1970: 26

Sternostoma zosteropus Domrow, 1966a: 194

Type locality: Gakoma [Rwanda]

Distribution: Australia, Botswana, Canada, Malaysia, Moldavia, Russia, Rwanda, Ukraine

Hosts: Arses kaupi, Cacomantis pyrrhophanus, C. variolosus, Colluricincla megarhyncha (Passeriformes: Artamidae), Eopsaltria capito, Eudynamys orientalis, Monarcha trivirgata, Myiagra rubecula, Oreoica gutturalis (Passeriformes: Falcunculidae), Rhipidura leucophrys (Passeriformes: Rhipiduridae), Zosterops lateralis (Passeriformes: Zosteropidae), Chrysococcyx caprius, Clamator levaillanti, Cuculus canorus, C. solitarius, Phaenicophaeus chlorophaeus (Cuculiformes: Cuculidae); Urolestes melanoleucus (Passeriformes: Laniidae), Yuhina zantholeuca (Passeriformes: Timaliidae), Sturnus sturninus (Passeriformes: Sturnidae), Catharus ustulatus (Passeriformes: Turdidae)

Sternostoma cuculorum korolevae Bregetova et Slavoshevskaya

Sternostoma cuculorum korolevae Bregetova & Slavoshevskaya, 1983: 234

Type locality: Altai, Troitskiy Rayon, Malaya rechka Vill.

Distribution: Russia

Host: Cuculus sp. (Cuculiformes: Cuculidae)

Sternostoma darlingi Spicer

Sternostoma darlingi Spicer, 1984: 795

Type locality: Guatemala

Distribution: Guatemala

Host: Mitrephanes phaeocercus (Passeriformes: Tyrannidae)

Sternostoma dumetellae Pence

Sternostoma dumetellae Pence, 1972b: 785; Spicer, 1987: 261

Type locality: Laplace, La.

Distribution: USA

Host: Dumetella carolinensis (Passeriformes: Mimidae)

Sternostoma dureni Fain

Sternostoma dureni Fain, 1956a: 153; 1957b: 68; Strandtmann & Wharton, 1958: 166; Domrow, 1969b: 376; Dimov, 2016; 2018: 62

Type locality: Astrida (= Butare), Rwanda

Distribution: Australia, Russia, Rwanda

Hosts: Oreocincla lunulata, Turdoides jardinei emini, T. melanops sharpei, Turdus merula, T. olivaceus graueri (Passeriformes: Turdidae)

Sternostoma epistomata Feider et Mironescu

Sternostoma epistomata Feider & Mironescu, 1973: 333; Butenko, 1984: 139

Sternostoma chlidoniadis Butenko, 1974: 1089 – type: Novosibirskaya obl., Zdvinskiy rayon, Malie Chany Lake, from Chlidonias nigra (syn. by Butenko, 1984)

Type locality: Tulcea (Romania)

Distribution: Romania, Russia

Hosts: Chlidonias hybrida, Ch. nigra (Charadriiformes: Sternidae)

Sternostoma eurocephali Fain

Sternostoma eurocephali Fain, 1960: 111

Type locality: Gravelotte (Transvaal)

Distribution: South Africa

Host: Eurocephalus anguitimens (Passeriformes: Laniidae)

Sternostoma ficedulae Fain et Sixl

Sternostoma eurocephali Fain & Sixl, 1971: 89; Fain, Sixl & Moritsch, 1974: 6

Type locality: Brunnsee (Styria)

Distribution: Austria

Host: Ficedula albicollis (Passeriformes: Motacillidae)

Sternostoma francolini Fain

Sternostoma francolini Fain, 1960c: 95; 1961a: 518; Sartbaev, 1979; 1983: 83; Butenko, 1984: 153

Type locality: Lochinvar, nr. Monze (Zambia)

Distribution: Belgium, Kazakhstan, Kyrghizstan, Russia, Zambia

Hosts: Alectoris chukar, Francolinus coqui, Perdix perdix perdix (Galliformes: Phasianidae)

Sternostoma fulicae Fain et Bafort

Sternostoma fulicae Fain & Bafort, 1963: 474; Domrow, 1969: 380; Feider & Mironescu, 1973: 159; Butenko, 1976: 77, 1984: 149; Sartbaev, 1981: 130; de Rojas et al., 2001: 961; de Rojas, Mora et al., 2002: 676

Type locality: Belgium

Distribution: Australia, Azerbaidjan, Belgium, Kirghizstan, Romania, Russia, Spain

Hosts: Fulica atra, Gallinula chloropus (Gruiformes: Rallidae)

Sternostoma furmani Strandtmann

Sternostoma furmani Strandtmann, 1960: 132

Type locality: Trus Madi Mt., Pampang, Uli Kaingaran, North Borneo

Distribution: Malaysia (Borneo)

Host: Arborophila brunneopectus erythrophys (Galliformes: Phasianidae)

Sternostoma galowayi Knee

Sternostoma galowayi Knee, 2018

Type locality: Winnipeg, Manitoba

Distribution: Canada

Host: Eremophila alpestris (Passeriformes: Alaudidae)

Sternostoma giganteum Fain

Sternostoma giganteum Fain, 1962c: 148

Type locality: Kisenyi

Distribution: Rwanda

Host: Batis molitor puella (Passeriformes: Muscicapidae)

Sternostoma gliciphilae Domrow

Sternostoma gliciphilae Domrow, 1966a: 192; 1969b: 377; Fain & Lukoschus, 1979: 19

Type locality: Esk

Distribution: Australia

Hosts: Gliciphila indistincta, Ramsayornis fasciatus (Passeriformes: Meliphagidae)

Sternostoma graculi (Butenko)(new comb.)

Sternostomoides graculi Butenko, 1999: 47

Type locality: Kyrghizstan

Distribution: Kyrghizstan

Host: Pyrrhocorax [Graculus] graculus (Passeriformes: Corvidae)

Sternostoma guevarai Guevara-Benítez, Úbeda-Ontíveros et Cutillas-Barrios

Sternostoma guevarai Guevara-Benítez, Úbeda-Ontíveros & Cutillas-Barrios, 1979: 29

Type locality: Granada

Distribution: Spain

Host: Alectoris rufa (Galliformes: Phasianidae)

Sternostoma hedonophilum Fain et Aitken

Sternostoma hedonophilum Fain & Aitken, 1970: 348

Type locality: Mosqueiro Ferry, Benavides, Belém

Distribution: Brazil

Host: Platyrhinchus saturatus (Passeriformes: Tyrannidae)

Sternostoma hirundinis Fain

Sternostoma hirundinis Fain, 1956b: 154; 1957b: 72; 1959a: 31; Shumilo & Lunkashu, 1970: 25; Černy & Dusbábek, 1970: 485; Pence, 1973a: 359; Spicer, 1977b: 498; 1987: 261

Type locality: Astrida (= Butare), Rwanda

Distribution: Cuba, Mexico, Moldavia, Rwanda, South Africa, Ukraine, USA (La.)

Hosts: Hirundo rustica, H. r. erythrogaster, H. smithi, Progne subis, Psalidoprocne albiceps, Riparia riparia, Tachycineta albilina (Passeriformes: Hirundinidae), Bombycilla cedrorum (Passeriformes: Bombycillidae), Cisticola aberrans (Passeriformes: Cisticolidae)

Sternostoma hirundo Fain, Herin et Puylaert

Sternostoma hirundo Fain, Herin & Puylaert, 1977: 389

Type locality: Chirundu (South Africa); paratypes from Cameroon and Rwanda

Distribution: Cameroon, Rwanda, South Africa

Hosts: Hirundo aethiopica, H. rustica, H. smithi (Passeriformes: Hirundinidae)

Sternostoma hutsoni Furman

Sternostoma hutsoni Furman, 1957: 477; Fain & Aitken, 1967: 17; 1971: 28; Spicer, 1984: 799

Type locality: Buttowillow, Kern County, California

Distribution: Brazil, Guatemala, Trinidad, USA (California)

Hosts: Catharus dryas, Hylocichla u. ustulata, Turdus albicollis, T. nudigenis, T. rufiventris (Passeriformes: Turdidae)

Sternostoma hylandi Fain et Johnston

Sternostoma hylandi Fain & Johnston, 1966: 32; Shumilo & Lunkashu, 1970: 27; Fain, Sixl & Moritsch, 1974: 6; Butenko, 1984: 148; Knee, Proctor & Galloway, 2008: 370

Type locality: 5 km S of Wooster, Ohio

Distribution: Austria, Canada, Russia, USA (Ohio)

Hosts: Dendrocopos leucotos, D. major, D. [Picoides] pubescens (Piciformes: Picidae)

Sternostoma inflatum Fain

Sternostoma inflatum Fain, 1963c: 83

Type locality: Akanyaru

Distribution: Rwanda

Host: Turdoides jardinei emini (Passeriformes: Timaliidae)

Sternostoma isabelae Ubuda – Ontíveros et Guevara – Benítez

Sternostoma isabelae Ubuda -Ontíveros & Guevara – Benítez, 1980: 366

Type locality: Moraleda de Zafayona (Granada)

Distribution: Spain

Host: Calandrella cinerea (Passeriformes: Alaudidae)

Sternostoma kelloggi Hyland et Clark

Sternostoma kelloggi Hyland & Clark, 1959: 223; Černy & Dusbábek, 1970: 485; Pence, 1972a: 781; 1975: 22

Type locality: Kalamazoo County, Michigan

Distribution: Cuba, USA (Louisiana, Michigan)

Hosts: Dumetella carolinensis, Mimus polyglottos orpheus, Toxostoma rufum (Passeriformes: Mimidae)

Sternostoma kodrensis Shumilo et Lunkashu

Sternostoma kodrensis Shumilo & Lunkashu, 1970: 25

Type locality: Shepot and Tarashani Villages

Distribution: Ukraine

Host: Troglodytes troglodytes (Passeriformes: Troglodytidae)

Sternostoma lagonostictae Fain

Sternostoma lagonostictae Fain, 1956b: 155; 1957b: 72

Type locality: Akanyaru

Distribution: Rwanda

Host: Lagonosticta rubricata congica (Passeriformes: Estrildidae)

Sternostoma longisetosae Hyland

Sternostoma longisetosae Hyland, 1961: 280; Hyland & Moorhouse, 1970: 56; Knee, Proctor & Galloway, 2008: 369

Sternostoma callithrix Fain & Aitken, 1970: 348 – Type from Empidonomus varius, Ananindeua, Pará, Brazil (syn. by Pence & Casto, 1975b)

Type locality: Michigan (USA)

Distribution: Brazil, Canada, Mexico, USA

Hosts: Empidonax alnorum, E. traillii, E. vireonis, Muscivora tyrannus, Pyrocephalus rubinus, Pitangus sulphuratus, Tyrannus tyrannus (Passeriformes: Tyrannidae)

Sternostoma loxiae Fain

Sternostoma loxiae Fain, 1965: 159; 1966: 120; Knee & Proctor, 2010

Type locality: Belgium

Distribution: Belgium, Canada, England

Hosts: Loxia curvirostra (Passeriformes: Fringillidae), Sialia currucoides (Passeriformes: Turdidae)

Sternostoma marchae Dimov

Sternostoma marchae Dimov, 2012: 225; 2018: 59

Type locality: Saint Petersburg, Leningradskaya Obl.

Distribution: Russia

Host: cage Serinus canaria (Passeriformes: Fringillidae)

Sternostoma minutus (Bregetova)

Sternostomum minutus Bregetova, 1950: 1007

Type locality: Vostochniy Murman, Zapovednik “Sem ostrovov”

Distribution: Russia

Host: Charadrius hiaticula (Charadriiformes: Charadriidae)

Sternostoma mortelmansi Fain et Bastin

Sternostoma mortelmansi Fain & Bastin, 1959: 116

Type locality: Anvers Zoo

Distribution: Brazil

Host: Apodiformes – Trochilidae gen. sp.

Sternostoma nectarinia Fain

Sternostoma nectarinia Fain, 1956a: 152; 1957b: 68; Strandtmann, 1960: 135; Fain, Herin & Puylaert, 1977: 392

Type locality: Astrida (= Butare), Rwanda

Distribution: Cameroon, DR Congo, Rwanda, Thailand

Hosts: Chalcomitra senegalensis aequatorialis, Cinnyris cupreus septentrionalis, Dicaeum concolor olivaceum, Nectarinia pulchella, N. purpureiventris (Passeriformes: Nectariniidae)

Sternostoma neosittae Domrow

Sternostoma sp. Domrow, 1967a: 217

Sternostoma neosittae Domrow, 1969b: 377

Type locality: Kowanyama (Queensland)

Distribution: Australia

Host: Neositta striata (Passeriformes: Neosittidae)

Sternostoma numerovi (Butenko)(new comb.)

Sternostomoides numerovi Butenko, 1999: 49

Type locality: Guinea

Distribution: Guinea

Host: Eremopteryx leucotis (Passeriformes: Alaudidae)

Sternostoma opistaspis Feider et Mironescu

Sternostoma opistaspis Feider & Mironescu, 1982: 87

Type locality: Papua New Guinea

Distribution: Papua New Guinea

Host: Cracticus cassicus (Passeriformes: Cracticidae)

Sternostoma paddae Fain

Sternostoma paddae Fain, 1958a: 8; Fain & Bafort, 1963a: 3; Domrow, 1969: 379

Type locality: Antverp Zoo

Distribution: South Asia

Host: Padda oryzivora (Passeriformes: Estrildidae)

Sternostoma pastor Fain

Sternostoma pastor Fain, 1967: 152; Feider & Mironescu, 1978: 265

Type locality: Antverp Zoo

Distribution: Belgium, Romania

Host: Sturnus roseus (Passeriformes: Sturnidae)

Sternostoma pencei Spicer

Sternostoma pencei Spicer, 1984: 797

Type locality: Baja Verapaz, 2 km E, 2.4 km S Purulhá

Distribution: Guatemala, USA (Missouri)

Host: Empidonax flavescens (Passeriformes: Tyrannidae)

Sternostoma piprae (Fain et Aitken)

Sternoecius piprae Fain & Aitken, 1967: 24; 1968: 3

Sternostoma piprae (Fain et Aitken): Domrow, 1969: 374

Type locality: Corneillac Estate, Vega de Oropouche (Trinidad)

Distribution: Brazil, Trinidad

Hosts: Manacus manacus, Pipra erythrocephala (Passeriformes: Pipridae)

Sternostoma pirangae Pence

Sternostoma pirangae Pence, 1973b: 874; Spicer, 1984: 799; Mascarenhas, Coimbra et al., 2011: 507

Sternostoma tangarae Fain & Aitken (in part), Pence, 1972: 785

Type locality: Mandeville. La.

Distribution: Brazil, Guatemala, USA (Louisiana, Texas)

Hosts: Piranga rubra (Passeriformes: Cardinalidae), Chlorospingus ophthalmicus (Passeriformes: Thraupidae), Paroaria coronata (Passeriformes: Emberizidae)

Sternostoma porteri Hyland

Sternostoma porteri Hyland, 1963: 149; Fain & Johnston, 1966: 31; Pence, 1972: 786; 1973: 886; Pence & Casto, 1975b: 362; Butenko, 1984: 145; Knee, Proctor & Galloway, 2008: 370

Type locality: Kalamazoo Co., Michigan

Distribution: Azerbajdjan, Canada, Ukraine, USA (Michigan, Ohio, Texas)

Hosts: Centurus aurifrons, Colaptes auratus, Picoides pubescens, P. villosus, Picus viridis, Sphyrapicus varius (Piciformes: Picidae)

Sternostoma quiscali Fain et Aitken

Sternostoma quiscali Fain & Aitken, 1967: 18; Černy & Dusbábek, 1970: 485

Type locality: El Socorro Estate Road, La Paille Village, San Juan

Distribution: Cuba, Trinidad

Hosts: Quiscalus lugubris, Qu. niger caribaeus (Passeriformes: Icteridae)

Sternostoma sayornis Pence et Casto

Sternostoma sayornis Pence & Casto, 1975b: 366

Type locality: Texas, Presidio Co., Candelaria

Distribution: USA (Texas)

Host: Sayornis nigricans (Passeriformes: Tyrannidae)

Sternostoma schiffornis Fain et Aitken

Sternostoma schiffornis Fain & Aitken, 1970: 343

Type locality: Mosqueiro Ferry, Benavides, Belém

Distribution: Brazil

Host: Schiffornis turdinus (Passeriformes: Pipridae)

Sternostoma setifer Knee

Sternostoma setifer Knee, 2008: 367

Type locality: Delta Marsh, Manitoba

Distribution: Canada

Host: Empidonax minimus (Passeriformes: Tyrannidae)

Sternostoma sialiphilus Hyland et Ford

Sternostoma sialiphilus Hyland & Ford, 1960: 101; Knee, Proctor & Galloway, 2008: 369

Type locality: Kalamazoo County, Michigan

Distribution: Canada, USA (Michigan)

Hosts: Sialia sialis (Passeriformes: Turdidae), Riparia riparia (Passeriformes: Hirundinidae)

Sternostoma sinense Fain et Bafort

Sternostoma sinense Fain & Bafort, 1963c: 7; Fain, Herin & Puylaert, 1977: 392

Type locality: Anvers Zoo

Distribution: Cameroon; type comes from Asian bird (Leiothrix lutea)

Hosts: Phoeniculus purpureus (Coraciiformes: Phoeniculidae), Quelea erythropus, Q. quelea (Passeriformes: Ploceidae), Leiothrix lutea (Passeriformes: Timaliidae)

Sternostoma sternahirundo Butenko

Sternostoma sternahirundo Butenko, 1974: 1090; 1984: 142; De Rojas et al., 961

Type locality: Novosibirskaya Obl., Zdvinskiy rayon, Malie Chany Lake (Russia)

Distribution: Russia, Spain

Hosts: Sterna hirundo (Charadriiformes: Sternidae), Larus argentatus (Charadriiformes: Laridae)

Sternostoma straeleni Fain

Sternostoma straeleni Fain, 1958c: 9

Type locality: Vitshumbi (Virunga Nat. Park)

Distribution: D.R. Congo

Host: Cinnyricinclus leucogaster (Passeriformes: Sturnidae)

Sternostoma strandtmanni Furman

Sternostoma strandtmanni Furman, 1957: 476; Fain & Aitken, 1967: 18; 1971: 28; de Rojas, Mora et al., 2002: 676

Type locality: California

Distribution: Brazil, Spain, Trinidad, USA

Hosts: Agelaius icterocephalus, A. tricolor, Icterus nigrogularis, Chrysomus ruficapillus, Leistes militaris, Molothrus sp., Quiscalus lugubris (Passeriformes: Icteridae); Tringa totanus (Charadriiformas: Scolopacidae)

Sternostoma strigitis Butenko

Sternostoma strigitis Butenko, 1976: 303; 1984: 143

Type locality: Ryazanskaya Obl.

Distribution: Russia

Host: Asio otus (Strigiformes: Strigidae)

Sternostoma sturnicola Fain

Sternostoma sturnicola Fain, 1956a: 151; 1957b: 65; 1959a: 31; 1961b: 52; Strandtmann & Wharton, 1958: 168; Fain, Herin & Puylaert, 1977: 392

Type locality: Astrida (= Butare), Rwanda

Distribution: Cameroon, Botswana, Rwanda

Hosts: Buphagus africanus (Passeriformes: Buphagidae), Lamprocolius chalybaeus, L. purpureus, Lamprotornis purpuropterus (Passeriformes: Sturnidae)

Sternostoma tangarae Fain et Aitken

Sternostoma tangarae Fain & Aitken, 1967: 20; Pence, 1972b: 785

Type locality: La Fortune Estate, Vega de Oropouche

Distribution: Trinidad, USA

Hosts: Piranga rubra (Passeriformes: Cardinalidae), Tangara mexicana (Passeriformes: Thraupidae)

Sternostoma technaui (Vitzthum)

Sternostomum technaui Vitzthum, 1935: 569

Sternostoma technaui: Zumpt & Till, 1955: 91; Fain, 1957b: 62; Furman, 1957: 482; Strandtmann & Wharton, 1958: 168; Domrow, 1969b: 379; Pence, 1972b: 786; Sartbaev, 1983: 84; Kadosaka, Kaneko & Asanuma, 1987: 38; Knee, Proctor & Galloway, 2008: 370

Sternostomoides technaui (Vitzthum): Bregetova, 1965: 709

Rhinonyssus technaui: Castro, 1948: 257

Sternostoma turdi Zumpt & Till, 1955: 85; Fain, 1956: 151; 1957b: 64; 1959a: 31; 1962: 264; 1963: 176; Furman, 1957: 481; Fain & Aitken, 1967: 17; Fain, Sixl & Moritsch, 1974: 5; Úbeda-Ontíveros & Guevara-Benítez, 1979: 291; de Rojas et al., 2001: 961; de Rojas, Mora et al., 2002: 676. – Type: South Africa. Syn. by Domrow, 1969: 380

Sternostomoides turdi (Zumpt et Till): Bregetova, 1965: 712; Shumilo & Lunkashu, 1970: 23

Sternostoma spatulatum Furman, 1957: 480 – type: Buttowillow, Kern County, California, from Hylocichla u. ustulata (Turdidae) ; Knee & Galloway, 2017: 95.

Syn. by Domrow, 1969: 380

Sternostomoides spatulatum (Furman): Bregetova, 1965: 712

Sternostomoides spatulatus (Furman): Černy & Dusbábek, 1970: 486

Sternostoma borceanum Feider & Mironescu, 1968: 105. Type: Romania, from Turdus ericetorum, T. pilaris, T. torcuatus (syn. by Fain, Sixl & Moritsch, 1974: 6); Feider & Mironescu, 1978: 267, from Sturnus vulgaris, Sf. Ghorghe, Tulcea

Sternostomoides orlandoi Dusbábek, 1969: 225; Černy & Dusbábek, 1970: 486 –

Type locality: Santa Fé, Prov. of Havana, Cuba, from Mimocichla plumbea rubripes, M. plumbea schistacea (Passeriformes: Turdidae). Syn. by Kadosaka, Kaneko & Asanuma, 1987: 38

Type locality of Sternostoma technaui: Europe

Distribution: Australia, Austria, Belgium, Brazil, Canada, Germany, Japan, Kirghizstan, Moldavia, Romania, Russia, Rwanda, South Africa, Spain, Tadjikistan, Trinidad, Ukraine, USA

Hosts: Cinclus cinclus aquaticus, C. pallasi (Passeriformes: Cinclidae), Hylocichla u. ustulata, Monticola saxatilis, Oreocincla dauma, O. lunulata, Turdus abyssinicus baraka, T. ericetorum, T. fumigatus, T. Iliacus, T. libonyanus, T. merula, T. migratorius, T. olivaceus, T. olivaceus graueri, T. philomelos, T. pilaris, T. ruficollis, T. viscivorus (Passeriformes: Turdidae), Sturnus vulgaris (Passeriformes: Sturnidae)

Sternostoma thienponti Fain

Sternostoma thienponti Fain, 1956a: 152; 1957b: 68; 1960: 111; Strandtmann & Wharton, 1958: 168; Domrow, 1965b: 449; 1966b: 194; 1969b: 375

Sternostoma thieponti [sic] Fain: Strandtmann, 1960: 135

Type locality: Astrida (= Butare), Rwanda

Distribution: Australia, Botswana, Rwanda, Thailand

Hosts: Dicrurus adsimilis, D. macrocercus cathoecus (Passeriformes: Dicruridae), Cracticus nigrogularis, C. quoyi (Passeriformes: Cracticidae), Gymnorhina tibicen (Passeriformes: Artamidae)

Sternostoma tracheacolum Lawrence

Sternostoma tracheacolum Lawrence, 1948: 366; Zumpt & Till, 1955: 86; Fain, 1956a: 152; 1957b: 65; Strandtmann & Wharton, 1958: 168; Strandtmann, 1960: 137; Fain & Hyland, 1962: 404; Amaral, 1962: 72; Murray, 1966: 262; Domrow, 1966a: 194; 1969b: 377, 423; Mathey, 1967: 777; Shumilo & Lunkashu, 1970: 26; Hyland & Moorhouse, 1970: 56; Fain & Aitken, 1971: 28; Pence, 1972b: 785; McClure, Ratanaworadhan et al., 1973: 37; Fain, Sixl & Moritsch, 1974: 5; Blasco Sabio & Portus Vinyeta, 1974: 37; Guevara-Benítez & Úbeda-Ontíveros, 1974: 176; Knee & Proctor, 2010; Knee & Galloway, 2017: 94

Sternostoma meddai Lombardini, 1953: 187 – Italy (syn. by Fain & Hyland, 1962)

Sternostoma castroae Fain, 1956b: 393 – Rwanda, ex Macronyx croceus (syn. by Fain, 1957)

Agapornyssus faini Grétillat, Capron & Brygoo, 1959: 375 – Madagascar, ex Agapornis cana (syn. by Fain & Hyland, 1962)

Type locality: Pietermaritzburg (South Africa)

Distribution: Australia, Austria, Belgium, Brazil, Canada, DR Congo, France, Great Britain, Italy (incl. Sardinia), Kirghizstan, Madagascar, Mexico, Moldavia, Russia, South Africa, Spain, Thailand, Trinidad, Uruguay, USA

Hosts: Coereba flaveola (Passeriformes: Coerebidae), Chloebia [Poephila] gouldiae, (Passeriformes: Estrildidae), Arachnothera longirostris, Nectarinia jugularis flammaxillaris (Nectariniidae), Sylvia melanocephala (Sylviidae), Chloris chloris, Coccothraustes coccothraustes, Serinus canaria (Fringillidae), Hirundo rustica, Iridoprocne bicolor, Riparia riparia (Hirundinidae), Dendrocincla merula (Furnariidae, Dendrocolaptinae), Cinnyricinclus leucogaster verreauxi (Sturnidae), Myiarchus crinitus, Rhynchocyclus olivaceus, Tyrannus melancholicus (Tyrannidae), Seiurus noveboracensis (Parulidae), Parus major (Paridae), Molothrus ater, Sturnella magna (Icteridae), Thraupidae, Melopsittacus undulatus (Psittaciformes: Psittacidae), Macronyx croceus (Passeriformes: Motacillidae), Melospiza melodia, Pipilo fuscus, Spizella pusilla, Sturnella magna (Passeriformes: Emberizidae), Cardinalis [Pyrrhuloxia] sinuatus, Passerina cyanea (Passeriformes: Cardinalidae), Acrocephalus arundinaceus (Passeriformes: Acrocephalidae)

Sternostoma tyrannus Brooks et Strandtmann

Sternostoma tyrannus Brooks & Strandtmann, 1960: 418; Pence, 1975: 23

Type locality: Lamb County, Texas

Distribution: USA (Texas)

Host: Nuttallornis borealis (Passeriformes: Tyrannidae)

Sternostoma ubedai Úbeda – Ontíveros et Guevara – Benítez

Sternostoma ubedai Úbeda – Ontíveros & Guevara – Benítez, 1981: 176

Type locality: Prov. de Granada

Distribution: Spain

Host: Sylvia melanocephala (Passeriformes: Sylviidae)

Sternostoma ziegleri Feider et Mironescu

Sternostoma ziegleri Feider & Mironescu, 1982: 87

Type locality: Papua New Guinea

Distribution: Papua New Guinea

Host: Cacomantis variolosus (Cuculiformes: Cuculidae)

Sternostoma zini Dimov et Knee

Sternostoma zini Dimov & Knee, 2012: 139

Type locality: Viritsa, Leningrad Province

Distribution: Russia

Host: Cuculus canorus (Cuculiformes: Cuculidae)

Genus Tinaminyssus Strandtmann et Wharton

Tinaminyssus Strandtmann & Wharton, 1958: 161

Type species: Neonyssus (Ptilonyssoides) trappi Peraira et de Castro, 1949: 229

Mesonyssus Fain, 1960: 313 (syn. by Domrow, 1969: 305)

Type species: Neonyssus treronis Fain, 1956: 394

Mesonyssoides Fain & Nadchatram, 1962: 271 (syn. by Domrow, 1969: 305)

Type species: Mesonyssoides malayi Fain & Nadchatram, 1962: 272

Mesonyssoides Strandtmann & Clifford, 1962: 723 (syn. with Mesonyssus by Wilson, 1964b: 357; with Tinaminyssus by Domrow, 1969: 305)

Type species: Mesonyssoides ixoreus Strandtmann & Clifford, 1962: 723

Psittonyssus Fain, 1963: 69 (syn. by Domrow, 1969: 305)

Type species: Psittonyssus baforti Fain, 1963: 70

Falconyssus Fain, 1966: 85 (syn. by Domrow, 1969: 305)

Type species: Falconyssus elani Fain, 1966: 86

Tinaminyssus alexfaini (Amaral)(new comb.)

Mesonyssus alexfaini Amaral, 1967: 135

Type locality: Itaporanga, Est. de São Paulo

Distribution: Brazil

Host: Leptotila verreauxi (Columbiformes: Columbidae)

Tinaminyssus alisteri (Wilson)(new comb.)

Mesonyssus alisteri Wilson, 1964b: 370

Type locality: Morobe District, Wau, Bulolo River

Distribution: Papua New Guinea

Host: «king parrot« (Psittaciformes)

Tinaminyssus angustus (Wilson)

Mesonyssus angustus Wilson, 1966: 607

Tinaminyssus angustus (Wilson): Domrow

Type locality: Morobe District: Wau: Bulolo

Distribution: Papua New Guinea

Tinaminyssus aprosmicti (Domrow)

Mesonyssoides aprosmicti Domrow, 1964b: 26; Fain & Lukoschus, 1979: 20

Mesonyssoides platycerci Domrow, 1964b: 27 (syn. by Domrow, 1969: 306)

Mesonyssus aposmicti (Domrow): Domrow, 1967a: 217; Wilson, 1968: 395

Tinaminyssus aprosmicti Domrow, 1969b: 306; 1979d: 488; 1987: 883

Type locality: Condamine, Kowanyama

Distribution: Australia

Hosts: Aprosmictus erythropterus, A. scapularis, Barnardius barnardi, Platycercus adscitus, Psephotus haematonotus, P. varius (Psittaciformes: Psittaculidae)

Tinaminyssus baforti (Fain)

Psittonyssus baforti Fain, 1963: 70

Tinaminyssus baforti (Fain): Domrow

Type locality: Avicentra (Anvers)

Distribution: East Africa

Host: Psittacula baforti (Psittaciformes: Psittaculidae)

Tinaminyssus belopolskii (Bregetova)

Neonyssus belopolskii Bregetova, 1950: 1005; 1951: 119; 1953: 332; Fain, 1957b: 53; Strandtmann & Wharton, 1958: 162; Mitchell, 1961: 105

Neonyssus ardeae Zumpt & Till, 1955: 63 – type: Benoni, Transvaal, from Ardea melanocephala; Fain, 1956a: 133; 1957b: 137; 1959a: 33 (syn. by Strandtmann, 1956: 137)

Neonyssus belepolskii: Strandtmann, 1956: 137

Mesonyssus belopolskii Fain, 1961a: 515; 1962d: 306; Domrow, 1965b: 433; 1966b: 191; 1967a: 216; Wilson, 1968a: 213; Černy & Dusbábek, 1970: 486; Butenko, 1984: 91

Mesonyssus belopolskii nycticoracis Fain, 1961a: 516; Butenko, 1984: 93. – Type: Astrida (= Butare) Rwanda, from Nycticorax nycticorax (syn. by Domrow, 1969)

Neonyssus ixobrychi Fain, 1956a: 134; Butenko, 1965: 131. – Type: Astrida (= Butare), Rwanda, from Ixobrychus minutus (syn. by Domrow, 1969: 311, but restaured by Butenko, 1984)

Neonyssus marcandrei Gretillat, Capron & Brygoo, 1959: 379 – Type: Lac Itasy, Madagascar, from Melanophyx ardesiaea (Ardeidae). Syn. by Fain, 1961a: 516.

Mesonyssus minuti Feider & Mironescu, 1974: 764 – type: Iasi, Ciric (syn. by Domrow, 1979: 513)

Tinaminyssus belopolskii Domrow, 1969b: 311; 1987: 883; Pence, 1972a: 162; Bernardon, Müller & Mascarenhas, 2013: 674.

Type locality: Russia (Sudzuhinskiy Goszapovednik)

Distribution: Australia, Botswana, Brazil, Cuba, Madagascar, Philippines (Mindanao), Russia, Rwanda, South Africa, Spain, Canary Isl., USA

Hosts: Ardea cinerea, A. melanocephala, A. purpurea manillensis, Bubulcus ibis, Butorides striata, B. virescens, Egretta garzetta, E. thula thula, Florida caerulea, Hydranassa tricolor, Ixobrychus minutus, Leucophoyx thula, Melanophyx ardesiaea, Notophoyx novaehollandiae, N. picata, Nycticorax nycticorax, Syrigma sibilatrix (Pelicaniformes: Ardeidae)

Tinaminyssus bubulci (Zumpt et Till)

Neonyssus bubulci Zumpt & Till, 1955: 66; Fain, 1956a: 134; 1957: 54; 1959: 33; 1960: 108; Černy & Dusbábek, 1970: 486 (Cuba). – Type: Johannesburg, South Africa, from Bubulcus ibis Syn. by Domrow (1969), but revalidated by Pence (1972)

Mesonyssus bubulci (Zumpt et Till): Černy & Dusbábek, 1970: 486 (Cuba); Butenko, 1984: 99

Tinaminyssus bubulci (Zumpt et Till): Pence, 1972: 160; 1973: 883; Rodríguez-Braza, Guevara-Benítez & Úbeda-Ontíveros, 1990: 73; de Rojas et al., 2002

Type locality: Johannesburg (South Africa)

Distribution: Cuba, South Africa

Hosts: Bubulcus ibis, Egretta intermedia brachyrhyncha (Pelecaniformes: Ardeidae)

Tinaminyssus buteonis (Fain)(new comb.)

Neonyssus buteonis Fain, 1956a: 135; 1957b: 51; 1959a: 32

Mesonyssus buteonis Fain, 1962d: 306

Falconyssus buteonis (Fain): Fain, 1966: 87

Type locality: Astrida (= Butare), Rwanda

Distribution: Rwanda, South Africa

Hosts: Buteo rufofuscus augur, Lophaetus occipitalis (Accipitriformes: Accipitridae); Coracias caudata (Coraciiformes: Coraciidae)

Tinaminyssus camargoi (Amaral) (new comb.)

Mesonyssus camargoi Amaral, 1962: 145

Type locality: Pirapora, Est. de São Paulo

Distribution: Brazil

Host: Penelope sp. (Galliformes: Cracidae)

Tinaminyssus carapachibeyus Dusbábek

Tinaminyssus carapachibeyus Dusbábek, 1969: 216; Černy & Dusbábek, 1970: 487

Type locality: Carapachibey, Isla de Pinos

Distribution: Cuba (Isla de Pinos)

Host: Geotrygon chrysia (Columbiformes: Columbidae)

Tinaminyssus castroae (Amaral)(new comb.)

Neonyssus castroae Amaral, 1963: 9

Type locality: Itaporanga, Est. de São Paulo

Distribution: Brazil

Hosts: Oreopeleia montana and Leptotila verreauxi (Columbiformes: Columbidae)

Tinaminyssus chiarellii Amaral et Baquer

Tinaminyssus chiarellii Amaral & Baquer, 1963: 65; Hyland & Moorhouse, 1970: 54

Type locality: Zoo in São Paulo

Distribution: Brazil, Mexico

Hosts: Cyanocorax chrysops chrysops (Passeriformes: Corvidae), Pitangus sulfuratus (Passeriformes: Tyrannidae)

Tinaminyssus columbae (Crossley)

Neonyssus (Neonyssus) columbae Crossley, 1950: 309

Neonyssus columbae Crossley: Zumpt & Till, 1955: 67; Fain, 1956a: 134; Strandtmann & Wharton, 1958: 162

Mesonyssus columbae (Crossley): Fain, 1962: 306; Foulk & Matthysse, 1965: 127; Domrow, 1965b: 440; Wilson, 1964: 368, 1966b: 1211; Černy, 1970: 244; Fain, Sixl & Moritsch, 1974: 7; Butenko, 1969: 208, 1984: 114; Dimov, 2016; 2018: 177

Tinaminyssus columbae (Crossley): Domrow, 1969b: 313; 1987: 884; Pence, 1975: 21; Kadosaka, Kaneko & Asanuma, 1987; Úbeda et al., 2003: 68; Knee, Proctor & Galloway, 2008: 368

Type locality: Alabama

Distribution: Australia, Austria, Belgium, Canada, Czech Rep., Japan, Russia, Rwanda, Spain, Ukraine, USA (Alabama, Georgia, Kentucky)

Hosts: Columba guinea guinea, C. livia, C. livia domestica, Ocyphaps lophotes, Streptopelia o. orientalis, S. semitorquata, Zenaida macroura (Columbiformes: Columbidae), Milvus migrans, M. tenebrosus (Accipitriformes: Accipitridae)

Tinaminyssus crypturelli Fain

Tinaminyssus crypturelli Fain, 1967: 153

Type locality: Anvers Zoo

Distribution: South America

Host: Crypturellus tataupa (Tinamiformes: Tinamidae)

Tinaminyssus cunhai (Amaral)(new comb.)

Mesonyssus cunhai Amaral, 1963: 103

Type locality: Itaporanga, Est. de São Paulo

Distribution: Brazil

Host: Leptotila verreauxi (Columbiformes: Columbidae)

Tinaminyssus daceloae (Domrow)

Mesonyssus daceloae Domrow, 1965b: 445

Falconyssus daceloae (Domrow): Fain, 1966: 87

Tinaminyssus daceloae Domrow, 1969b: 322; 1979d: 492; 1983: 884

Type locality: Samford, Esk

Distribution: Australia

Hosts: Dacelo gigas, D. leachii (Coraciiformes: Halcyonidae)

Tinaminyssus egrettae (Butenko)(new comb.)

Mesonyssus egrettae Butenko, 1984: 96

Type locality: Krasnodarskiy Kray, Primorsko-Ahtarskiy Rayon

Distribution: Russia

Host: Egretta garzetta (Pelicaniformes: Ardeidae)

Tinaminyssus elani (Fain)(new comb.)

Falconyssus elani Fain, 1966c: 86

Type locality: Luluabourg (now Kananga)

Distribution: DR Congo

Host: Elanus coeruleus (Accipitriformes: Accipitridae)

Tinaminyssus epileus (Wilson)

Mesonyssus epileus Wilson, 1964b: 358; 1965: 638; 1968a: 213; Tenorio, 1976

Tinaminyssus epileus (Wilson): Domrow, 1969: 312

Type locality: Finschhafen

Distribution: Papua New Guinea

Host: Aviceda subcristata (Accipitriformes: Accipitridae)

Tinaminyssus geopeliae (Fain)

Mesonyssus geopeliae Fain, 1964b: 33; Domrow, 1965b: 440; 1966b: 192; Wilson, 1966b: 1211; Fain & Lukoschus, 1979: 20

Tinaminyssus geopeliae Domrow, 1969b: 315, 1979d: 490; 1987: 884; McClure, Ratanaworadhan et al., 1973; 35

Type locality: Anvers Zoo (Malaysia)

Distribution: Australia, Hawaii, Malaysia, Papua New Guinea, Philipppines

Hosts: Geopelia cuneata, G. humeralis, G. placida, G. striata striata (Columbiformes: Columbidae)

Tinaminyssus geotrygoni (Dusbábek)

Mesonyssus geotrygoni Dusbábek, 1969: 218; Černy & Dusbábek, 1970: 487

Tinaminyssus geotrygoni: Pence, 1975: 21

Type locality: Carapachibey, Isla de Pinos

Distribution: Cuba (Pinos)

Host: Geotrygon chrysia (Columbiformes: Columbidae)

Tinaminyssus gerschi (Feider et Mironescu) (new comb.)

Mesonyssus gerschi Feider & Mironescu, 1974: 752

Type locality: Delta of Danube

Distribution: Romania

Host: Ardea purpurea (Pelecaniformes: Ardeidae)

Tinaminyssus gourae (Wilson)(new comb.)

Mesonyssus gourae Wilson, 1964b: 361

Type locality: NW New Guinea, Boemi River

Distribution: Papua New Guinea

Hosts: Ducula pinon, Goura cristata (Columbiformes: Columbidae)

Tinaminyssus halcyonus (Domrow)

Mesonyssus halcyonus Domrow, 1965b: 443; Wilson, 1966a: 609; 1968a: 213

Falconyssus halcyonus: Fain, 1966: 86

Tinaminyssus halcyonus Domrow, 1969b: 321, 1972: 99; 1979: 492; 1987: 884; McClure, Ratanaworadhan et al., 1973: 35

Type locality: Logan Village, Brisbane

Distribution: Australia, New Zealand, Papua New Guinea, Philippines

Hosts: Halcyon chloris collaris, H. macleayii, H. pyrrhopygius, H. sanctus (Coraciiformes: Halcyonidae)

Tinaminyssus hirtus (Wilson)

Mesonyssus hirtus Wilson, 1966b: 1210; 1968a: 215

Tinaminyssus hirtus Domrow, 1969b: 313; 1987: 884; McClure, Ratanaworadhan et al., 1973: 35

Type locality: Siaton, Kandugay, Negros Oriental, Philippines

Distribution: Australia, Papua New Guinea, Philippines

Hosts: Chalcophaps chrysochlora, Ch. indica indica, Ch. stephani (Columbiformes: Columbidae)

Tinaminyssus ixobrychi (Fain)

Neonyssus ixobrychi Fain, 1956a: 154; 1957b: 54; Butenko, 1965: 131

Mesonychus ixobrychi (Fain): Feider & Mironescu, 1974: 751; Butenko, 1984: 94

Tinaminyssus ixobrychi (Fain): Domrow, 1969: 311

Mesonyssus minuti Feder & Mironescu, 1974: 764 (syn. by Butenko, 1984: 95)

Type locality: Astrida [Butare], Rwanda

Distribution: Moldavia, Kirghizstan, Rwanda, Russia, South Africa

Host: Ixobrychus minutus (Pelicaniformes: Ardeidae)

Tinaminyssus ixoreus (Strandtmann et Clifford)

Mesonyssoides ixoreus Strandtmann & Clifford, 1962: 723; Isakova, 1965

Tinaminyssus ixoreus (Strandtmann et Clifford): Pence, 1975: 21; Spicer, 1984: 800

Type locality: Lost Horse Canyon, Ravalli County, Montana

Distribution: Canada, Guatemala, Russia, USA (Montana)

Hosts: Catharus dryas, Ixoreus naevius (Passeriformes: Turdidae)

Tinaminyssus juxtamelloi Pence et Canaris

Tinaminyssus juxtamelloi Pence & Canaris, 1976: 116

Type locality: Mimbres, Luna Co., New Mexico

Distribution: USA (New Mexico)

Host: Columba fasciata (Columbiformes: Columbidae)

Tinaminyssus kakatuae (Domrow)

Mesonyssoides kakatuae Domrow, 1964b: 25

Mesonyssus kakatuae Domrow, 1966b: 191; 1967a: 217; Wilson, 1966: 759

Tinaminyssus kakatuae Domrow, 1969b: 306; 1987: 885

Type locality: Kowanyama, Condamine

Distribution: Australia

Hosts: Calyptorhynchus banksi, Cacatua roseicapilla, C. sulphurea occidentalis, Nymphicus [Leptolophus] hollandicus (Psittaciformes: Cacatuidae)

Tinaminyssus macropygiae (Wilson)

Mesonyssus macropygiae Wilson, 1966a: 607; 1968b: 215

Tinaminyssus macropygiae Domrow, 1969b: 319;1987: 885

Type locality: NW New Guinea, Oransbari

Distribution: Australia, Papua New Guinea, Philippines

Hosts: Macropygia amboinensis, M. phasianella (Columbiformes: Columbidae)

Tinaminyssus malayi (Fain et Nadchatram)

Mesonyssoides malayi Fain & Nadchatram, 1962: 272

Tinaminyssus malayi (Fain et Nadchatram): McClure, Ratanaworadhan et al., 1973: 35

Type locality: Mont Brinchang (1700 m), Pahang

Distribution: Malaysia

Host: Cyanops (= Megalaima) franklini minor (Piciformes: Capitonidae)

Tinaminyssus marciae (Amaral)(new comb.)

Mesonyssus marciae Amaral, 1966: 153

Type locality: Boracéa, São Paulo

Distribution: Brazil

Host: Leuchocloris albicolis (Apodiformes: Trochilidae)

Tinaminyssus megaloprepiae Domrow

Tinaminyssus megaloprepiae Domrow, 1969b: 313; 1987: 885

Type locality: Maalan

Distribution: Australia

Host: Megaloprepia magnifica (Columbiformes: Columbidae)

Tinaminyssus melloi (de Castro)

Neonyssus (Neonyssus) melloi de Castro, 1948: 270

Mesonyssus melloi (de Castro): Fain, 1962d: 306; Domrow, 1965b: 434; 1966b: 192; Amaral, 1968: 122; Butenko, 1969; Shumilo & Lunkashu, 1970: 16; Fain, Sixl & Moritsch, 1974: 7; Sartbaev, 1982: 131; Rodriguez-Braza, 1987: 175C; Dimov, 2018: 174

Neonyssus hirsutus Feider, 1962: 58. – Type: Romania, from Columba livia (syn. by Domrow, 1969: 312)

Neonyssus melloi: Zumpt & Till, 1955: 68; Fain, 1957b: 137; 1959a: 33; Strandtmann & Wharton, 1958: 163

Tinaminyssus melloi: Domrow, 1969b: 312 ; 1972: 99; 1987: 885; Zamudio, 1988: 205; Kadosaka, Kaneko & Asanuma, 1987; Úbeda et al., 2003: 66; Knee, Proctor & Galloway, 2008: 368

Type locality: Brazil

Distribution: Australia, Austria, Canary Is., Canada, Czech Rep., Brazil, Hungary, Japan, Kirghizstan, Moldavia, New Zealand, Romania, Russia, South Africa, Spain, Ukraine, USA

Hosts: Columba livia domestica, C. norfolciensis, C. oenas, C. palumbus, Leucosarcia melanoleuca, Stigmatopelis senegalensis, Streptopelia capicola, S. chinensis, S. o. orientalis, Zenaida macroura (Columbiformes: Columbidae), Passer domesticus (Passeriformes: Passeridae)

Tinaminyssus metropeliae (Fain)(new comb.)

Mesonyssus metropeliae Fain, 1972: 361

Type locality: Zoo Anvers

Distribution: Zoo Anvers

Host: Metropelia caeciliae (Columbiformes: Columbidae)

Tinaminyssus milvi (Fain)

Mesonyssus milvi Fain, 1962: 149

Tinaminyssus milvi (Fain): Domrow, 1969: 305

Type locality:

Distribution:

Host: Milvus (Accipitriformes: Accipitridae)

Tinaminyssus minisetosum (Butenko)

Mesonyssus minisetosum Butenko, 1984: 115

Tinaminyssus minisetosum (Butenko): Úbeda et al., 2003: 70

Type locality: Kirghizstan, Alayskaya Dolina (Valley)

Distribution: Kirghizstan, Spain

Host: Columba palumbus (Columbiformes: Columbidae)

Tinaminyssus myristicivorae Domrow

Mesonyssus sp. “A” Domrow, 1967a: 215

Tinaminyssus myristicivorae Domrow, 1969b: 316, 1979d: 490; 1987: 885

Type locality: Kowanyama

Distribution: Australia

Host: Myristicivora spilorrhoa (Columbiformes: Columbidae)

Tinaminyssus navajasi (Pereira et Castro)

Neonyssus (Ptilonyssoides) navajasi Pereira & Castro, 1949: 231

Neonyssus navajasi Pereira et Castro: Zumpt & Till, 1955: 87; Fain, 1957b: 57

Mesonyssus navajasi: Fain, 1962d: 306; Amaral, 1968: 122

Tinaminyssus navajasi: Strandtmann & Wharton, 1958: 161

Type locality: Estados de São Paulo and Mato Grosso

Distribution: Brazil

Host: Nothura maculosa (Tinamiformes: Tinamidae)

Tinaminyssus neoixobrychi Pence

Tinaminyssus neoixobrychi Pence, 1972a: 165; 1975: 22

Type locality: Louisiana, Norco

Distribution: USA

Host: Ixobrychus exilis (Pelicaniformes: Ardeidae)

Tinaminyssus ocyphabus (Domrow)

Mesonyssus ocyphabus Domrow, 1965b: 437

Tinaminyssus ocyphabus Domrow, 1969b: 312; 1987: 885

Type locality: Condamine, Samford

Distribution: Australia

Hosts: Geophaps scripta, Ocyphaps lophotes (Columbiformes: Columbidae).

Tinaminyssus oenae (Fain)(new comb.)

Mesonyssus oenae Fain, 1963c: 61

Type locality: Anvers Zoo

Distribution: South Africa

Host: Oena c. capensis (Columbiformes: Columbidae)

Tinaminyssus peleiae (Wilson)(new comb.)

Mesonyssus pileiae Wilson, 1966a: 609

Type locality: Leyte I., Santa Cruz, Mahaplag

Distribution: Philippines

Host: Phapitreron brevirostris (Columbiformes: Columbidae)

Tinaminyssus phabus (Domrow)

Mesonyssus phabus Domrow, 1965b: 435

Tinaminyssus phabus Domrow, 1969b: 317

Type locality: Esk

Distribution: Australia

Host: Phaps chalcoptera (Columbiformes: Columbidae)

Tinaminyssus phalliger (Fain)(new comb.)

Psittanyssus phalliger Fain, 1965: 160

Type locality: Zoo Anvers

Distribution: Indonesia

Host: Psittacula alexandri (Psittaciformes: Psittaculidae)

Tinaminyssus phassae (Wilson)(new comb.)

Mesonyssus phassae Wilson, 1966: 602

Type locality: Kaohsiung Hsien, Liukuei

Distribution: Taiwan

Host: Sphenurus sieboldii sororius (Columbiformes: Columbidae)

Tinaminyssus prionituri (Wilson)(new comb.)

Mesonyssus prionituri Wilson, 1966: 759; 1968: 397

Type locality: Philippines (Leyte, Palawan)

Distribution: Philippines (Leyte, Palawan)

Hosts: Prioniturus [flavicans] montanus, P. [discurus] platenae, P. discurus whiteheadi (Psittaciformes: Psittaculidae)

Tinaminyssus pteroclesi (Fain)(new comb.)

Mesonyssus pteroclesi Fain, 1963c: 63

Type locality: Lochinvar

Distribution: South Africa

Host: Pterocles gutturalis (Pteroclidiformes: Pteroclididae)

Tinaminyssus ptilinopi (Wilson)

Mesonyssus ptilinopi Wilson, 1964b: 366; 1965: 638; 1966: 1212; 1968: 221; Tenorio, 1976

Tinaminyssus ptilinopi: Domrow, 1969b: 315; McClure, Ratanaworadhan et al., 1973: 36

Type locality: SE New Guinea (Papua), Cape Kileton

Distribution: Australia, Malaysia, Papua New Guinea, Philippines, Taiwan

Hosts: Ptilinopus coronulatus, P. ornatus, P. perlatus, P. superbus (Columbiformes: Columbidae)

Tinaminyssus sartbaevi (Butenko)

Mesonyssus sartbaevi Butenko, 1984: 111

Tinaminyssus sartbaevi (Butenko): Úbeda et al., 2003: 68

Type locality: Kirghizstan, Alayskaya Dolina (Valley)

Distribution: Kirghizstan, Spain

Host: Columba palumbus (Columbiformes: Columbidae)

Tinaminyssus saucerottiae (Fain et Aitken)

Trochilonyssus saucerottiae Fain & Aitken, 1967: 32

Tinaminyssus saucerottiae (Fain et Aitken): Fain & Aitken, 1970: 410

Type locality: La Horquette Trace, Tumpuna Road, Trinidad

Distribution: Trinidad

Hosts: Campylopterus largipennis, Saucerottia tobaci (Apodiformes: Trochilidae)

Tinaminyssus schoutedeni (Fain)

Neonyssus schoutedeni Fain, 1956a: 134; 1957b: 49

Mesonyssus schoutedeni Fain, 1960c: 92

Falconyssus schoutedeni (Fain): Fain, 1966: 86

Tinaminyssus schoutedeni: Domrow & Nadchatram, 1978: 85

Type locality: Astrida (= Butare), Rwanda

Distribution: Malaysia, Rwanda, Zambia

Hosts: Halcyon leucocephala, H. smyrnensis (Coraciiformes: Halcyonidae)

Tinaminyssus senotrusovae (Butenko)(new comb.)

Mesonyssus senotrusovae Butenko, 1984: 103

Type locality: Tyumenskaya Obl., Yalutorovskiy Rayon (Russia)

Distribution: Kazakhstan, Russia

Host: Streptopelia turtur (Columbiformes: Columbidae)

Tinaminyssus serraoi (de Castro)(new comb.)

Neonyssus (Neonyssus) serraoi de Castro, 1948: 270; Zumpt & Till, 1955: 87; Fain, 1957b: 52

Mesonyssus serraoi (de Castro): Fain, 1962d: 306; Amaral, 1968: 122

Type locality: Brazil

Distribution: Brazil

Host: Rhynchotus rufescens (Tinamiformes: Tinamidae)

Tinaminyssus squamosus (Vitzthum)

Rhinonyssus squamosus Vitzthum, 1935: 567; Pereira & de Castro, 1949: 222; Zumpt & Till, 1955: 88

Neonyssus squamosus (Vitzthum): Fain, 1957b: 48

Rhinonyssoides squamosus: Strandtmann & Wharton, 1958: 173

Mesonyssus squamosus: Fain, 1961: 45; 1962: 305

Falconyssus squamosus: Fain & Aitken, 1969: 407

Tinaminyssus squamosus: Domrow

Type locality: West Indies

Distribution: West Indies, Brazil

Hosts: Florisuga melivora, Sericotes holosericeus (Apodiformes: Trochilidae)

Tinaminyssus streptopeliae (Fain)(new comb.)

Mesonyssus melloi streptopeliae Fain, 1962d: 310; Shumilo & Lunkashu, 1970: 17

Mesonyssus streptopeliae: Fain, Herin & Puylaert, 1977: 392; Sartbaev, 1983: 86

Type locality: Musha (near Astrida, Rwanda)

Tinaminyssus melloi streptopeliae: Rodriguez-Braza, Guevara-Benítez & Úbeda-Ontíveros, 1990: 73

Distribution: Cameroon, Kirghizstan, Moldavia, Rwanda, Spain, Ukraine

Hosts: Columba livia, Streptopelia orientalis, S. s. semitorquata, S. turtur (Columbiformes: Columbidae)

Tinaminyssus streptopelioides (Butenko)

Mesonyssus streptopelioides Butenko, 1984: 107

Tinaminyssus streptopelioides (Butenko): de Rojas et al., 2002:

Type locality: Azerbaydzhan (Kizil-Agach Reserve)

Distribution: Azerbaydzhan, Kazakhstan, Russia, Turkmenistan

Host: Streptopelia turtur (Columbiformes: Columbidae)

Tinaminyssus tanysipterae (Wilson)

Mesonyssus tanysipterae Wilson, 1966a: 605

Tinaminyssus tanysipterae Domrow, 1979: 490; 1987: 886

Type locality: NW New Guinea, Oransbary

Distribution: Australia, Papua New Guinea

Hosts: Tanysiptera danae, T. galatea, T. sylvia (Coraciiformes: Halcyonidae)

Tinaminyssus tetae (Wilson)(new comb.)

Mesonyssus tetae Wilson, 1964b: 605; Tenorio, 1976

Type locality: NW New Guinea, Enarotali, 1900 m

Distribution: Papua New Guinea

Host: green fruit dove (Ptilinopus purpuratus)(Columbiformes: Columbidae)

Tinaminyssus tinamicola (Fain)(new comb.)

Mesonyssus tinamicola Fain, 1963c: 66

Type locality: Anvers Zoo

Distribution: Argentina

Host: Eudromia elegans (Tinamiformes: Tinamidae, Nothurinae)

Tinaminyssus trappi (Pereira et Castro)

Neonyssus (Ptilonyssoides) trappi Pereira & Castro, 1949: 229

Mesonyssus trappi: Fain, 1962d: 306; Amaral, 1968: 122

Tinaminyssus trappi: Strandtmann & Wharton, 1958: 161

Type locality: Boracéa (Estado de São Paulo)

Distribution: Brazil

Host: Tinamus solitarius (Tinamiformes: Tinamidae)

Tinaminyssus treronis (Fain)(new comb.)

Neonyssus treronis Fain, 1956b: 394; 1957b: 56

Mesonyssus treronis Fain, 1962d: 314

Type locality: Akanyaru

Distribution: Rwanda

Host: Treron c. calva (Columbiformes: Columbidae)

Tinaminyssus triangulus (Strandtmann)

Neonyssus triangulus Strandtmann, 1961: 323

Mesonyssus triangulus (Strandtmann): Fain, 1962: 307

Tinaminyssus triangulus (Strandtmann): Pence, 1975: 22

Type locality: Edinburg, Texas

Distribution: USA (Texas)

Host: Zenaida asiatica (Columbiformes: Columbidae)

Tinaminyssus trichoglossi (Domrow)

Mesonyssoides trichoglossi Domrow, 1964: 29

Mesonyssus trichoglossi Wilson, 1964b: 378; 1966b: 191; 1967a: 217

Mesonyssus neopsittaci Wilson, 1964: 372; 1966: 759; 1968: 397 – Type: Enarotali, 1900 m, Bulldog Road, 2440 m, from many species of parrots (syn. by Domrow, 1969: 311)

Mesonyssus domicellae Wilson, 1964b: 375; 1968b: 397 – Type: May River, Papua New Guinea, from Domicella lory (syn. by Domrow, 1969: 311)

Mesonyssus chalcopsyttae Wilson, 1968b: 395 – Type: Oriomo River, Western District, Papua New Guinea, from Chalcopsytta scintillata (syn. by Domrow, 1969: 311)

Tinaminyssus trichoglossi Domrow, 1969b: 311; 1979d: 488; 1987: 886

Type locality: Logan & Tamborine Villages, Queensland

Distribution: Australia, Papua New Guinea

Hosts: Chalcopsitta scintillata, Charmosina papou, Glossopsitta pusilla, Micropsitta bruijnii, Neopsittacus musschenbroekii, N. pullicauda, Pseudeos fuscata, Psittacella modesta, Trichoglossus chlorolepidotus, T. haematodus, T. moluccanus (Psittaciformes: Psittaculidae)

Tinaminyssus turturi (Fain)(new comb.)

Mesonyssus turturi Fain, 1962d: 314

Type locality: Astrida (= Butare)

Distribution: Rwanda

Host: Turtur afer (Columbiformes: Columbidae)

Tinaminyssus welchi Domrow

Mesonyssus sp. “B” Domrow, 1967a: 215

Tinaminyssus welchi Domrow, 1969b: 319; 1987: 886

Type locality: Kowanyama

Distribution: Australia

Host: Ducula [Myristicivora] spilorrhoa (Columbiformes: Columbidae, Treroninae)

Tinaminyssus zenaidurae (Crossley)

Neonyssus zenaidurae Crossley, 1952: 386; Zumpt & Till, 1955: 87; Fain, 1956b: 394; 1957b: 55

Mesonyssus zenaidurae (Crossley): Fain, 1962: 314; Wilson, 1964: 369; Fain & Aitken, 1967: 39

Tinaminyssus zenaidurae: Pence, 1973a: 359; 1975: 22; Powders & Coffey, 1983: 107; Knee, Proctor & Galloway, 2008: 368

Type locality: Dickens Co., Texas

Distribution: Canada, Trinidad, USA (Louisiana, Oklahoma, Texas, Alabama, California, Mississippi)

Hosts: Columbigallina passerina, C. talpacoti, Zenaida [Zenaidura] macroura (Columbiformes: Columbidae)

Tinaminyssus zumpti (Fain)

Mesonyssus zumpti Fain, 1960: 92

Falconyssus zumpti (Fain): Fain, 1966: 86

Type locality: Rooiberg, Transvaal

Distribution: South Africa

Host: Halcyon chelicuti (Coraciiformes: Halcyonidae)

Genus Trochilonyssus Fain et Aitken

Trochilonyssus Fain & Aitken, 1967: 32

Type species: Trochilonyssus trinitatis Fain et Aitken, 1967

Trochilonyssus trinitatis Fain et Aitken

Trochilonyssus trinitatis Fain et Aitken, 1967: 32

Type locality: Esperanza Estate, Vega de Oropouche

Distribution: Trinidad

Host: Glaucis hirsuta (Apodiformes: Trochilidae)

Genus Tyranninyssus Brooks et Strandtmann

Tyranninyssus Brooks & Strandtmann, 1960: 418

Type species: Tyranninyssus tyrannus Brooks et Strandtmann, 1960

Tyranninyssus attilae Fain et Aitken

Tyranninyssus attilae Fain & Aitken, 1970: 405; 1971: 43

Type locality: Mosqueiro Ferry, Benavides, Belém

Distribution: Brazil

Hosts: Attila spadiceus (Passeriformes: Cotingidae); Myiozetetes cayenensis (Passerifor