ACARORUM CATALOGUS VI. Order Mesostigmata. Gamasina: Dermanyssoidea (Rhinonyssidae, Spinturnicidae)

Petar Beron

doi:10.3897/ab.e54206

Petar Beron^‡

‡ Unaffiliated, Sofia, Bulgaria

Corresponding author: Petar Beron ( beron@mail.bg )

This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.

Citation: Beron P (2020) ACARORUM CATALOGUS VI. Order Mesostigmata. Gamasina: Dermanyssoidea (Rhinonyssidae, Spinturnicidae). Advanced Books. https://doi.org/10.3897/ab.e54206

Superorder Acariformes

This volume contains data on two families, 16 genera and 445 species of the families Rhinonyssidae (parasites in the respiratory system of birds) and 12 genera and 110 species of Spinturnicidae (ectoparasites of bats).

Order Mesostigmata G. Canestrini, 1891

According to Lindquist, Krantz & Walter (2009), Order Mesostigmata is composed of three suborders: Sejida, Trigynaspida and Monogynaspida with 26 superfamilies and about 12000 described species.

Suborder Monogynaspida Camin et Gorirossi, 1955

According to Beaulieu et al. (2011), the suborder consists of the infraorders Uropodina Kramer, 1881 and Gamasina Kramer, 1881, as well as of 15 superfamilies and 78 families. In Lindquist, Krantz & Walter (2009) the superfamilies are 18.

Infraorder (or Cohort ?) Gamasina Kramer, 1881

Cohort Gamasina of Lindquist, Krantz & Walter (2009) includes 10 superfamilies. Here belong most of the Mesostigmata, both predators and parasites.

Within Gamasina are delimitated 11 superfamilies and 42 families. Superfamily Dermanyssoidea remains within the Hyporder Dermanyssiae Evans et Till, 1979, with 17 families.

Beaulieu et al. 1911. Superorder Parasitiformes Reuter, 1909. In: Zhang Z.-Q. (Ed.) Animal biodiversity. An outline of higher-level classification and survey of taxonomic richness. – Zootaxa, 3148: 123–128.

Lindquist, Krantz & Walter 2009. Mesostigmata. In Krantz & Walter (Eds)., Manuel of Acarology, III Edition.

Superfamily Dermanyssoidea Kolenati

Dermanyssoidea Kolenati, 1859

After Krantz & Walter (2009, Eds), 14 families of parasitic mites form this superfamily, after the latest list of Parasitiformes families (Beaulieu et al., 1911) – 17. In this volume are considered the families Rhinonyssidae and Spinturnicidae.

The Mite Family Rhinonyssidae

Almost all Rhinonyssidae live in the nare passages of birds. One (Sternostoma tracheacolum Lawrence) has been found in the tracheae and lungs of birds and some Ptilonyssus have been recorded as living on the bird’s body or even in the nests.

N. Bregetova reviewed in 1951 the most numerous and common mites living in the respiratory system of birds – the Rhinonyssidae. By this time she was able to enumerate only 26 species in 10 genera, six of which are still valid: Rhinonyssus Trouessart, Sternostoma Berlese et Trouessart, Ptilonyssus Berl. et Tr., Rhinoecius Cooreman, Larinyssus Strandtmann and Rallinyssus Strandtmann. Four other generic names have been synonymized since (Sternostomum, Neonyssus, Ptilonyssoides and Rhinonyssoides). This scarce information was contributed mainly by Giebel (who described the first rhinonyssid in 1871), Trouessart, Berlese, Trägårdh, Hirst, Vitzthum, Castro, Cooreman, Strandtmann and Bregetova. Trouessart (1895) considered these mites as subfamily Rhinonyssinae and Vitzthum (1935) raised them in a family. However, some authors (Domrow) have longtime maintained the broader concept of Dermanyssidae and considered Rhinonyssinae as one of it’s subfamilies. Bregetova has splited these mites into two families – Rhinonyssidae and Ptilonyssidae, which was not accepted aside from few authors (Dusbábek & Černý, 1970).

Speaking of the “prehistory” of the studies on Rhinonyssidae we must notice the work of Brazilian specialists Castro & Pereira (1947), Castro (1948) and Pereira & Castro (1949) describing the subfamily Ptilonyssinae and several new species.

As for many other families, the “prehistory” of the research ends with the beginning of the long series of publications of Alex Fain (from 1956 to 1974). Based on his large collection, obtained in Africa, and on many mites from zoo birds, Fain (together with Aitken, Bafort, Bastin, Carpenter, Holland, Hyland, Johnson, Lukoschus, Nadchatram, Sixl, Moritsch, Vercammen-Grandjean) contributed greatly to the modern system of Rhinonyssidae, describing many new genera, species and subdividing the family into subfamilies. In the same time active research was carried out in Australia by R. Domrow (from 1964 to 1992). Domrow synonymised many older genera (Mesonyssus Fain, Mesonyssoides Fain et Nadchatram, Psitanyssus Fain, Falconyssus Fain). Some other genera, described by Castro (1948), were made synonyms by Zumpt & Till (1955) in their important contribution to the Rhinonyssidae of Southern Africa (Flavionyssus, Rochanyssus, Paraneonyssus, Travanyssus, Vitznyssus). Also important contributions to the study of Rhinonyssidae were made by Butenko (many publications on Russian and other mites since 1961, including a book in 1984, partly with Stanjukovich and Lavrovskaya), Shumilo & Lunkashu (incl. a book in 1976), Strandtmann and co-authors (Brooks, Clifford, Furman, Wharton), Knee (2008, together with Proctor and Galloway), Pence (1972–1979, together with Casto, Young and Canaris), Bregetova (from 1950 to 1983), Hyland (1961–1970, partly with Clark, Ford and Galloway), Kadosaka, Kaneko & Asanuma (1987), Feider & Mironescu (1972–1982), Dusbábek (since 1969), Amaral (1962–1978), Spicer (since 1977), George (1961), Furman (1957), Wilson (from 1958 to 1980), Sakakibara (1967, 1968), Crossley (1950, 1952), Grétillat (1969, 1970), Sixl (1969–1971), Sartbaev (1979) and others. Many new studies have been made in Spain by Guevara, Úbeda, Rodriguez, Rojas and other acarologists.

The Studies on Rhinonyssidae

Berlese & Trouessart (1889) describe the important genera Ptilonyssus and Sternostoma, containing more than half of all Rhinonyssidae

Trouessart (1894) – becomes the author of another important genus – Rhinonyssus

Trouessart (1895) describes the family Rhinonyssidae

Between 1895 and the Word War II very few authors study species (no new genera) of Rhinonyssidae: Trägårdh (1904), Vitzthum (1935), Hirst (1921–1923)

The post-war period is very rich in research on endoparasitic acari, incl. Rhinonyssidae. The following authors should be noticed (some of them still working on these mites):

J. Cooreman (1946) – describes the new genus Rhinoecius – start of research on the Rhinonyssidae on owls.

In Brazil M. Castro and C. Pereira try to restructure the family Rhinonyssidae, describing the subfamily Ptilonyssinae and several new species.

In the same time in the United States several researchers start working on Rhinonyssidae. Most prominent among them is R.W. Strandtmann (papers on Rhinonyssidae from 1948 to 1962, partly with C. Clifford, D. Furman, W. Wharton and others). The publications of Strandtmann contributed greatly to the knowledge on these mites. He described many new species and the genera Larinyssus and Rallinyssus.

D.A. Crossley – publications in 1950 and 1952 with descriptions of new mites from pigeons.

The great Belgian Acarologist A. Fain (1912–2009) started publishing on nasal mites in 1956, based on his large collection from Rwanda and Congo. He reorganysed the system of Rhinonyssidae, described (partly together with Aitken, Bafort, Bastin, Carpentier, Herin, Pulyert, Holland, Hyland, Johnston, Lukoschus, Mortelmans, Nadchatram, Sixl, and Vercammen-Grandjean) the new subfamily Rhinoeciinae, five valid new genera, 174 valid new species and several subspecies from many countries.

N. Wilson published papers on Rhinonyssidae from Antarctic and Subantarctic birds, birds of the Philippines, New Guinea, Taiwan, Thailand, Alaska and other Pacific areas (1958–1980). Also from this area many species have been described by I. Sakakibara and by E.W. Baker and M. Delfinado. The nasal mites of Japan were studied by T. Kadosaka and other researches.

Russian school in the study of Rhinonyssidae started with the thesis of Belopolskaya (1951) and is based mainly on the many papers of N. Bregetova and O. Butenko (with K.I. Lavrovskaya, M. Stanyukovich and other co-authors). Butenko worked in the reserve of Oka and described many new taxa of Rhinonyssidae. N. Bregetova considered these mites as being two families – Rhinonyssidae and Ptilonyssidae, what was not accepted but by few authors (Dusbábek & Černý, 1970). She published from 1950 to 1983 10 papers containing data on Rhinonyssidae, including a monograph on Gamasida and attempts to clarify the origin of these mites. S.K. Sartbaev published several papers on Rhinonyssidae of Central Asia from 1961 to 1983. On the Rhinonyssidae of Asia published several papers also T.T. Isakova, D.S. Ivanov and others. On Modavian mites worked R.P. Shumilo and M.I. Lunkashu.

In South Africa F. Zumpt and W.M. Till published in 1955 and 1961 important contribution on Rhinonyssidae, with descriptions of nine new species and a list of African Rhinonyssidae

D.P. Furman – in 1957 published comprehensive revision of genus Sternostoma

S. Gretillat – in 1961 described two new species and in 1959, together with A. Capron and E.R. Brygoo, described new nasal mites from Madagascar. The taxa, created by them (Agapornyssinae subfam. nov., Agapornyssus gen. nov. and Agapornyssus faini sp.n.) were synonymes.

H. G. Ford – in 1961 described Ptilonyssus constrictus

Z. Feider and I. Mironescu described from 1962 to 1982 several new species from Romania and Papua New Guinea.

K.E. Hyland – in series of papers (1959–1970)(patly with H.G. Ford, T.D. Clark and A. Moorhouse) described many new taxa from Mexico and USA

F. Dusbábek and V. Černý – studies of Cuban Rhinonyssidae (1969, 1970)

The series of papers (1972–1979) of D.B. Pence contain important data on the nasal mites from Louisiana, but also of other states.

P. Beron (1975) published the only paper on Bulgarian Rhinonyssidae. W. Sixl published papers on Rhinonyssidae from Austria.

V. Amaral took over the Brazilian school of research on nasal mites and from 1962 to 1974 published many new species and comprehensive list of Rhinonyssidae. This studies have been followed by C.S. Mascarenhas, D.S. Silva and other Brasilian researchers.

In Australia R. Domrow wrote from 1964 to 1992 many papers containing data on Rhinonyssidae of Australia and New Zealand. He reformed the system of these mites, put in synonymy many species and several genera (Sternostomoides, Rhinosterna, Sternoecius, Mesonyssus, Mesonyssoides, Psittanyssus, Falconyssus).

G.S. Spicer published between 1977 and 1987 several papers on the Rhinonyssidae of Guatemala and the USA.

W. Knee, H. Proctor, T. Galloway are publishing actively on the nasal mites of Canada.

I. Dimov published descriptions of several new species from Russia and a monograph (2018) concerning the Rhinonyssidae of birds in the NW of Russia.

The Composition of Rhinonyssidae

Fam. Rhinonyssidae Trouessart, 1894 – 16 gen., 445 sp.

Subfam. Rhinoeciinae Fain, 1957 – 3 gen., 16 sp.

Rhinoecius Cooreman, 1946 – 12 sp.

Ruandanyssus Fain, 1957 – 3 sp.

Zumptnyssus Fain, 1959 – 1 sp.

Subfam. Rhinonyssinae Trouessart, 1895 – 4 gen., 51 sp.

Charadrinyssus Butenko, 1984 – 1 sp.

Larinyssus Strandtmann, 1948 – 5 sp.

Rallinyssus Strandtmann, 1948 – 12 sp.

Rhinonyssus Trouessart, 1894 – 33 sp.

Subfam. Ptilonyssinae (Castro, 1948) Fain, 1957 – 9 gen., 378 sp.

Locustellonyssus Bregetova, 1965 – 2 sp.

Passeronyssus Fain, 1960 – 5 sp.

Pipronyssus Fain et Aitken, 1967 – 1 sp.

Ptilonyssus Berlese et Trouessart, 1889 – 215 sp.

Sternostoma Berlese et Trouessart, 1889 (incl. Rhinosterna Fain, 1964; Sternoecius Fain et Aitken, 1967; Sternostomoides Bregetova, 1965) – 75 sp.

Tinaminyssus Strandtmann et Wharton, 1958 – 68 sp.

Trochilonyssus Fain et Aitken, 1967 – 1 sp.

Tyranninyssus Brooks et Strandtmann, 1960 – 4 sp.

Vitznyssus Castro, 1948 (Astridiella Fain, 1957) – 9 sp.

Catalogue of Rhinonyssidae

Family Rhinonyssidae Trouessart

Rhinonyssinae Trouessart, 1895

Rhinonyssidae Trouessart: Vitzthum, 1935

Type genus: Rhinonyssus Trouessart, 1894

Ptilonyssidae Castro, 1948

Neonyssidae Bregetova, 1964

Subfamily Rhinoeciinae Fain

Rhinoeciinae Fain, 1957a: 156

Type genus: Rhinoecius Cooreman, 1946

Genus Rhinoecius Cooreman

Rhinoecius Cooreman, 1946: 1

Type species: Rhinoecius oti Cooreman, 1946

Rhinoecius aegolii Butenko

Rhinoecius aegolii Butenko, 1971c: 204; 1984: 131; Philips, 1993; 2000: 225; Knee, Proctor & Galloway, 2008: 370; Knee & Galloway, 2017: 96

Type locality: Okskiy Zapovednik, Russia

Distribution: Canada, Russia

Hosts: Aegolius acadicus, A. funereus, Surnia ulula (Strigiformes: Strigidae)

Rhinoecius africanus (Zumpt et Patterson)

Rhinonyssus africanus Zumpt & Patterson, 1951: 90

Rhinoecius africanus (Zumpt et Patterson): Zumpt & Patterson, 1955: 82; Fain, 1957b: 130; 1958b: 294

Type locality: Pietermaritzburg, Natal

Distribution: South Africa

Host: Asio capensis (Strigiformes: Strigidae)

Rhinoecius alifanovi Butenko

Rhinoecius alifanovi Butenko, 1976: 306, 1984: 127; Knee, Proctor & Galloway, 2008: 370

Type locality: Omskaya Obl., Krutinskiy Rayon; Ryazanskaya Obl.

Distribution: Canada, Russia

Host: Asio flammeus (Strigiformes: Strigidae)

Rhinoecius bisetosus Strandtmann

Rhinoecius bisetosus Strandtmann, 1952: 212; 1960: 132; Fain, 1957b: 130; Amaral, 1962b; Úbeda, Rodríguez & Guevara, 2000

Type locality: Lubbock Co., Texas

Distribution: Spain, Thailand, USA (Texas)

Hosts: Athene noctua, Glaucidium cuculoides, G. c. brugeli, Speotyto cunicularia, S. cunicularia grallaria (Strigiformes: Strigidae)

Rhinoecius brikinboricus Butenko

Rhinoecius brikinboricus Butenko, 1976: 305; 1984: 124; Knee, Proctor & Galloway, 2008: 370; Knee & Galloway, 2017: 96

Type locality: Ryazanskaya Obl., Brikin Bor

Distribution: Canada, Russia

Host: Asio otus (Strigiformes: Strigidae)

Rhinoecius cavannus Wilson

Rhinoecius cavannus Wilson, 1968c: 143; McClure, Ratanaworadhan et al., 1973: 37; Butenko, 1984: 130

Type locality: Chiengmai Province, Doi Inthanon

Distribution: Thailand

Host: Otus bakkamoena (Strigiformes: Strigidae)

Rhinoecius cooremani Strandtmann

Rhinoecius cooremani Strandtmann, 1952: 208; Fain, 1957b: 130; Strandtmann & Wharton, 1958: 165; Domrow, 1969b: 333, 423; Pence, 1973a: 359; Knee, Proctor & Galloway, 2008: 370; Knee & Galloway, 2017: 96

Type locality: Luling, Texas

Distribution: Australia, Canada, USA (La., Texas)

Hosts: Ninox connivens, N. novaeseelandiae, Strix nebulosa, S. varia (Strigiformes: Strigidae)

Rhinoecius grandis Strandtmann

Rhinoecius grandis Strandtmann, 1952: 205; Fain, 1957b: 130; Knee, Proctor & Galloway, 2008: 370

Type locality: Lubbock Co., Texas

Distribution: Brazil, Canada, USA (Texas)

Hosts: Bubo virginianus, Pulsatrix melanonota koeniswaldiana (Strigiformes: Strigidae)

Rhinoecius nycteae Butenko

Rhinoecius nycteae Butenko, 1976: 307; 1984: 133; Knee, Proctor & Galloway, 2008: 370

Type locality: Omskaya Obl., Cherlakskiy rayon

Distribution: Canada, Russia

Host: Bubo scandiacus [= Nyctea scandiaca] (Strigiformes: Strigidae)

Rhinoecius oti Cooreman

Rhinoecius oti Cooreman, 1946: 1; Pereira & Castro, 1949: 233; Zumpt & Patterson, 1951: 77; Zumpt & Till, 1955: 82; Fain, 1957b: 130; Butenko, 1984: 124

Type locality: Schooten (Belgium)

Distribution: Belgium

Host: Asio otus otus (Strigiformes: Strigidae)

Rhinoecius subbisetosus Bregetova

Rhinoecius subbisetosus Bregetova, 1965: 214; Butenko, 1984: 130

Type locality: Badhiz, Akar-Cheshme

Distribution: Turkmenistan

Host: Athene noctua (Strigiformes: Strigidae)

Rhinoecius tytonis Fain

Rhinoecius tytonis Fain, 1956b: 394; Fain, 1957b: 131, 138; 1959a: 33; Butenko, 1984: 132

Type locality: Akanyaru (Rwanda)

Distribution: Rwanda, South Africa

Hosts: Tyto alba, T. alba affinis (Strigiformes: Tytonidae)

Genus Ruandanyssus Fain

Ruandanyssus Fain, 1957a: 148

Type species: Ruandanyssus terpsiphonei Fain, 1957

Ruandanyssus artami Domrow

Ruandanyssus sp., Domrow, 1967a: 217

Ruandanyssus artami Domrow, 1969b: 330

Type locality: Kowanyama

Distribution: Australia

Hosts: Artamus cinereus, A. minor (Passeriformes: Artamidae)

Ruandanyssus strandmanni Sakakibara

Ruandanyssus strandmanni Sakakibara, 1968a: 13

Type locality: Enarotali

Distribution: New Guinea

Host: ?Neositta sp. (Passeriformes: Neosittidae)

Ruandanyssus terpsiphonei Fain

Ruandanyssus terpsiphonei Fain, 1957a: 148; 1957b: 131; Strandtmann & Wharton, 1958: 165; Domrow, 1965b: 432; 1967a: 217; 1969b: 329; 1979d: 439; Kadosaka, Kaneko & Asanuma, 1987: 40

Ruandanyssus terpsiphonei echongi Sakakibara, 1968: 15 – type: Enarotali, New Guinea, from Pachycephala (syn. by Domrow, 1969)

Type locality: Astrida (= Butare), Akanyaru (Rwanda)

Distribution: Australia, Japan, New Guinea, Rwanda

Hosts: Artamus personatus (Passeriformes: Artamidae), Coracina novaehollandiae, C. robusta, Pericrocotus roseus divaricatus (Passeriformes: Campephagidae), Lalage tricolor (Passeriformes: Corvidae), Pachycephala griseiceps, P. pectoralis, P. rufiventris (Passeriformes: Pachycephalidae), Sericornis frontalis, S. keri (Passeriformes: Acanthizidae), Struthidea cinerea (Passeriformes: Corcoracidae), Monarcha trivirgata, Myiagra rubecula, Terpsiphone viridis kivuensis (Passeriformes: Monarchidae)

Genus Zumptnyssus Fain

Zumptnyssus Fain, 1959c: 112

Type species: Ruandanyssus buboensis Fain, 1958

Zumptnyssus buboensis (Fain)

Ruandanyssus buboensis Fain, 1958b: 292

Zumptnyssus buboensis (Fain): Fain 1959c: 112; 1960: 113

Type locality: Lephepe (Botswana)

Distribution: Botswana, Mozambique

Host: Bubo africanus (Strigiformes: Strigidae)

Subfamily Rhinonyssinae Trouessart

Rhinonyssinae Trouessart, 1895 – Fain, 1957a: 154

Type genus: Rhinonyssus Trouessart, 1894

Genus Charadrinyssus Butenko

Charadrinyssus Butenko, 1984: 74

Type species: Larinyssus charadrius Butenko, 1984

Charadrinyssus charadrius (Butenko)

Larinyssus charadrius Butenko, 1975b: 951

Charadrinyssus charadrius (Butenko): Butenko, 1984: 74

Type locality: Kurgaldjinskiy Rayon

Distribution: Kazakhstan

Host: Charadrius morinellus (Charadriiformes: Charadriidae)

Genus Larinyssus Strandtmann

Larinyssus Strandtmann, 1948: 128

Type species: Larinyssus orbicularis Strandtmann, 1948

Larinyssus benoiti Fain

Larinyssus benoiti Fain, 1961: 128; 1964: 134; Domrow, 1966: 196; 1969: 322; 1979d: 492; 1988: 864; Sartbaev, 1979: 56; Butenko, 1984: 72

Type locality: Kasongo, DR Congo

Distribution: Australia, DR Congo, Kirghizstan, Russia

Hosts: Galachrysia cinerea, Glareola nordmanni, G. pratincola, Stiltia isabella (Charadriiformes: Glareolidae)

Larinyssus iohanssenae Dimov

Larinyssus iohanssenae Dimov, 2013: 123; 2018: 167

Type locality: Russia

Distribution: Russia

Host: Sterna hirundo (Charadriiformes: Sternidae)

Larinyssus orbicularis Strandtmann

Larinyssus orbicularis Strandtmann, 1948: 507; Pereira & Castro, 1949: 232; Bregetova, 1951: 117; Zumpt & Paterson, 1951: 78; Zumpt & Till, 1955: 68; Fain, 1956b: 157; 1957b: 60; 1958: 9; 1960c: 95; Černy & Dusbábek, 1970: 487; Pence, 1972a: 160; 1975: 21; Domrow, 1967a: 216; 1969: 322; 1973: 84; 1988: 865; Butenko, 1984: 68; Dimov, 2016; 2018: 168

Larinyssus petiti Gretillat, 1961 – syn. by Domrow, 1969: 322

Type locality of L. orbicularis: Galveston, Texas

Distribution: Australia, Brazil, DR Congo, Cuba, Russia, Rwanda, South Africa, USA, Zambia

Hosts: Anous minutus, Chlidonias leucoptera, Ch. nigra, Ch. orbicularis, Gelochelidon nilotica, Sterna fuscata, S. hirundinacea, S. hirundo, S. maxima, Thalasseus acuflavidus, Th. bergii (Charadriiformes: Sternidae), Larus argentatus, L. atricilla, L. delawarensis, L. dominicanus, L. minutus, L. novaehollandiae (Charadriiformes: Laridae), Rynchops nigra (Charadriiformes: Rynchopidae)

Larinyssus sterna Fain et Holland

Larinyssus sterna Fain & Holland, 1972: 30

Type locality: Aves Is., Venezuela

Distribution: Venezuela

Host: Onychoprion [Sterna] fuscatus fuscatus (Charadriiformes: Sternidae)

Larinyssus substerna Butenko

Larinyssus substerna Butenko, 1975: 180; 1984: 71

Type locality: Tomskaya Obl., Kolominskiy Rayon (Russia)

Distribution: Kazakhstan, Kirghizstan, Russia

Host: Sterna hirundo (Charadriiformes: Sternidae)

Larinyssus sp

Distribution: Brazil

Host: Larus dominicanus (Charadriiformes: Laridae)

Genus Rallinyssus Strandtmann

Rallinyssus Strandtmann, 1948: 512

Type species: Rallinyssus caudistigmus Strandtmann, 1948

Rallinyssoides Fain, 1960a: 295 – syn. by Wilson, 1965: 624

Type species: Rallinyssus congolensis Fain, 1956

Rallinyssus amaurornis Wilson

Rallinyssus amaurornis Wilson, 1965: 624; 1966; McClure, Ratanaworadhan et al., 1973: 37

Type locality: Nan t’ou Hsien, Puli (Taiwan)

Distribution: Philippines, Taiwan

Hosts: Amaurornis phoenicurus chinensis, Poliolimnas cinereus collingwoodi (Gruiformes: Rallidae)

Rallinyssus caudistigmus Strandtmann

Rallinyssus caudistigmus Strandtmann, 1948: 512; Bregetova, 1951: 118; 1956: 198; Zumpt & Till, 1955: 68; Strandtmann & Wharton, 1958: 160; Fain, 1957: 58; 1959a: 33; Domrow, 1966b: 196; 1969b: 324; Feider & Mironescu, 1973: 159; Fain, Sixl & Moritsch, 1974: 7; Pence, 1972; 1975: 42; Guevara-Benítez & Úbeda-Ontíveros, 1975a: 492; Butenko, 1976; 1984: 78; Sartbaev, 1977; 1979: 56; Kadosaka, Kaneko & Asanuma, 1987: 39; Knee, Proctor & Galloway, 2008: 368; Knee & Galloway, 2017: 93; Kun & Vega, 2016: 97; Silva et al., 2018; Dimov, 2018: 184

Type locality: Texas

Rallinyssus caspicus Butenko, 1976: 72; 1984: 81; Sartbaev, 1979: 56 – type: Kizil Agachskiy Zapovednik (Azerbaidjan), from Gallinula chloropus. Syn. by Domrow, 1979: 513

Distribution: Argentina, Australia, Azerbaijan, Austria, Canada, Japan, Kazakhstan, Kirghizstan, Romania, Russia, South Africa, Spain, Turkmenistan, Ukraine, USA (Texas)

Hosts: Fulica americana, F. armillata, F. atra, F. cristata, Gallinula chloropus, G. chloropus indica, G. tenebrosa, Rallus elegans (Gruiformes: Rallidae)

Rallinyssus congolensis Fain

Rallinyssus congolensis Fain, 1956b: 396; 1957b: 58; Strandtmann & Wharton, 1958: 160; Domrow, 1973: 84

Rallinyssoides congolensis (Fain): Fain, 1960a: 624

Rallinyssus porzana congolensis Fain: McClure, Ratanaworadhan et al., 1973: 37

Type locality: Akanyaru

Distribution: Australia (incl. Tasmania), DR Congo, Philippines

Host: Porzana tabuensis (Gruiformes: Rallidae)

Rallinyssus cychramus Wilson

Rallinyssus cychramus Wilson, 1965: 627; Butenko, 1984: 88

Type locality: PNG, Western Highlands Distr., Tambul, 2300 m

Distribution: Australia, DR Congo, Papua New Guinea

Hosts: Porzana parva, P. pusilla, P. tabuensis (Gruiformes: Rallidae)

Rallinyssus gallinulae Fain

Rallinyssus gallinulae Fain, 1960a: 295; 1962: 265; Wilson, 1965: 634; 1968: 221; Domrow, 1965b: 450; 1969b: 323; 1972: 99; 1973: 84; Butenko, 1976: 74; 1984: 83

Rallinyssus rallus Wilson, 1965: 631 – syn. by Domrow, 1969: 323

Type locality: Anvers Zoo

Distribution: Australia (incl. Tasmania), Azerbaijan, Belgium, New Zealand

Hosts: Gallinula chloropus, Hypotaenidia philippensis, Porphyrio porphyrio (= melanotus), Rallina tricolor, Rallus pectoralis (Gruiformes: Rallidae)

Rallinyssus limnocoracis Fain

Rallinyssus limnocoracis Fain, 1956b: 397; McClure, Ratanaworadhan et al., 1973: 37

Rallinyssoides limnocoracis (Fain): Fain & Nadchatram, 1962: 271

Type locality: Akanyaru (Rwanda)

Distribution: Malaysia (Selangor), Rwanda

Hosts: Limnocorax flavirostris, Porzana f. fusca (Gruiformes: Rallidae)

Rallinyssus porzanae Wilson

Rallinyssus porzanae Wilson, 1967a: 215

Type locality: Nueva Viskaya, Luzon

Distribution: Philippine Is.

Host: Zapornia [Porzana] tabuensis filipina (Gruiformes: Rallidae)

Rallinyssus rallinae Wilson

Rallinyssus rallinae Wilson, 1966a: 220; McClure, Ratanaworadhan et al., 1973: 37

Type locality: Nueva Viskaya, Luzon

Distribution: Philippine Is.

Host: Rallina eurizonoides eurizonoides (Gruiformes: Rallidae)

Rallinyssus sorae Pence et Young

Rallinyssus sorae Pence & Young, 1979: 791; Knee & Galloway, 2017: 93

Type locality: Maryland, Ohio

Distribution: Canada, USA (Maryland, Ohio)

Host: Porzana carolina (Gruiformes: Rallidae)

Rallinyssus strandtmanni Gretillat

Rallinyssus strandtmanni Gretillat, 1961: 151; Wilson, 1965: 637;1966: 225; Butenko, 1984: 89

Rallinyssoides strandtmanni (Gretillat): Amaral, 1962: 189

Type locality: France

Distribution: France

Host: Gallinula chloropus (Gruiformes: Rallidae)

Rallinyssus trappi (Amaral)

Rallinyssoides trappi Amaral, 1962: 185; 1968: 123

Rallinyssus trappi (Amaral): Wilson, 1965: 637

Type locality: Itaporanga, Estado de Sâo Paulo

Distribution: Brazil

Host: Ortygonax nigricans (Gruiformes: Rallidae)

Rallinyssus verheyeni Fain

Rallinyssus verheyeni Fain, 1963a: 176; Wilson, 1966: 225; Pence, 1972a: 158; 1973: 884; 1975: 21; Sartbaev, 1979: 56; Butenko, 1984: 85

Type locality: Horion – Hozémont, Prov. Liège, Belgium

Distribution: Australia, Belgium, Kirghizstan, Russia, USA (La.)

Hosts: Podiceps furicollis, P. ruficollis (Podicipiformes: Podicipedidae), Rallus aquaticus, R. elegans, R. limicola (Gruiformes: Rallidae)

Genus Rhinonyssus Trouessart

Rhinonyssus Trouessart, 1894: 723

Type species: Rhinonyssus coniventris Trouessart, 1894

Sommatericola Trägårdh, 1904b: 28

Type species: Sommatericola levinseni Trägårdh, 1905

Rhinonyssus afribyx Fain

Rhinonyssus afribyx Fain, 1956a: 150; 1957b: 48

Type locality: Astrida [Butare], Rwanda

Distribution: DR Congo, Rwanda

Hosts: Vanellus [Afribyx] senegallus, Vanellus [Stephanibyx] lugubris (Charadriiformes: Charadriidae)

Rhinonyssus alberti Strandtmann

Rhinonyssus alberti Strandtmann, 1956: 140; Fain, 1957b: 48; Feider & Mironescu, 1973: 415; Butenko, 1965: 131; 1984: 47

Type locality: Longbeach, California

Distribution: Australia, Azerbajdjan, Kazakhstan, Romania, Russia, USA

Hosts: Podiceps auritus, P. nigricolis (= Colymbus caspicus), P. alberti, P. cristatus, P. griseigena, P. ruficollis (Podicipediformes: Podicipedidae)

Rhinonyssus apus Fain

Rhinonyssus apus Fain, 1957a: 151; 1957b: 48

Type locality: Akanyaru (Rwanda)

Distribution: Rwanda

Host: Apus caffer streubeli (Apodiformes: Apodidae)

Rhinonyssus belenopteri Fain

Rhinonyssus belenopteri Fain, 1964: 29

Type locality: Anvers Zoo (Chile)

Distribution: Chile

Host: Belenopterus chilensis (Charadriiformes: Charadriidae)

Rhinonyssus bregetovae Butenko

Rhinonyssus bregetovae Butenko, 1974: 464; 1984: 39; Dimov, 2016; 2018: 137

Type locality: Ryazanskaya Obl., Tomskaya Obl. (Russia), Tselinogradskaya Obl. (Kazakhstan)

Distribution: Kazakhstan, Russia

Host: Charadrius dubius (Charadriiformes: Charadriidae)

Rhinonyssus caledonicus Hirst

Rhinonyssus caledonicus Hirst, 1921: 357; Strandtmann, 1956: 135; 1959; Fain, 1957b: 45; 1960b: 312; Pence, 1975: 20; Butenko, 1984: 33; Dimov, 2018: 142

Sternostomum caledonicum: Vitzthum, 1935: 569; Bregetova, 1951: 116; 1956: 198; Turk, 1953: 13

Sternostoma caledonicus: Zumpt & Till, 1955: 90

Type locality: Glass Pt.

Distribution: Germany, Russia (incl. Far East, Komandor and other islands), Shetland Is., USA

Hosts: Aethia pusilla, Cepphus columbae, Ciceronia pusilla, Cerorhinca monocerata, Uria aalge, U. grylle, U. lomyia (Charadriiformes: Alcidae)

Rhinonyssus chettusiae Butenko

Rhinonyssus chettusiae Butenko, 1973: 1874; 1984: 42

Type locality: Kamýsh (Kazakhstan)

Distribution: Kazakhstan

Host: Chettusia gregaria (Charadriiformes: Charadriidae)

Rhinonyssus clangulae Butenko et Stanyukovich

Rhinonyssus clangulae Butenko & Stanyukovich, 2001: 522

Type locality: Nizhne-Kolymskiy Rayon, Yakutia

Distribution: Russia

Host: Clangula hyemalis (Anseriformes: Anatidae)

Rhinonyssus colymbicola Fain et Bafort

Rhinonyssus colymbicola Fain & Bafort, 1963: 471; Butenko, 1984: 50; Knee & Galloway, 2017: 95

Type locality: Damme (Belgium)

Distribution: Belgium, Canada, Russia

Hosts: Podiceps auritus, P. griseigena, P. nigricollis (Podicipediformes: Podicipedidae)

Rhinonyssus coniventris Trouessart

Rhinonyssus coniventris Trouessart, 1894: 724; Hirst, 1921: 361; Vitzthum, 1935: 572; Castro, 1948: 256; Strandtmann, 1951: 130; 1956: 130; Zumpt & Patterson, 1951: 78; Turk, 1953: 13; Zumpt & Till, 1955: 84; Fain, 1957b: 43; 1960: 312; 1963: 88; Strandtmann & Wharton, 1958: 157; Fain, 1963: 88; Domrow, 1965b: 450; 1967a: 216; 1969b: 327, 423; Černý & Dusbábek, 1970: 484

Type locality: Europe

Distribution: Australia, Belgium, Canada, Cuba, England, Shetland Islands, South Africa, USA (incl. Alaska and Hawaii Is.)

Hosts: Arquatella ptilocnemis, Arenaria interpres, Calidris canutus, C. maritima, Catophrophorus semipalmatus, Charadrius alexandrinus, Ch. hiaticula, Ch. marginatus, Crocethia alba, Rhyacophilus glareola, Totanus flavipes, Erolia ruficollis (Charadriiformes: Charadriidae), Tringa ochropus (Charadriiformes: Scolopacidae)

Rhinonyssus dobromiri Dimov et Spicer

Rhinonyssus dobromiri Dimov & Spicer, 2013: 292; Dimov, 2018: 133

Type locality: Leningrad Province

Distribution: Russia

Host: Vanellus vanellus (Charadriiformes: Charadriidae)

Rhinonyssus echinipes Hirst

Rhinonyssus echinipes Hirst, 1921: 359 – type: Shetland Islands, ex Charadrius hiaticula; Turk, 1953: 13; Vitzthum, 1935: 572; Castro, 1948: 256; Strandtmann, 1951: 132; Butenko, 1984: 28; de Rojas et al., 2007

Rhinonyssus coniventris echinipes: Fain, 1963: 93; Fain, 1963: 179; Černy & Dusbábek, 1970: 484; Amaral & Reboucas, 1974: 55

Type locality: Shetland Islands

Distribution: Belgium, Shetland Islands, Spain, Turkmenistan

Hosts: Charadrius alexandrinus, Ch. hiaticula, Ch. melodus, Ch. wilsonia, Pluvialis squatarola (Charadriiformes: Charadriidae)

Rhinonyssus himantopus Strandtmann

Rhinonyssus himantopus Strandtmann, 1951: 136; 1956: 132; 1959: 134; Strandtmann & Wharton, 1958: 158; Fain, 1956a: 149; 1957b: 44; 1958c: 9; 1960c: 92; Domrow, 1966b: 195; 1969b: 326; 1979d: 493; Černy & Dusbábek, 1970: 484; Pence, 1972a: 157; Butenko, 1984: 46; Kadosaka, Kaneko & Asanuma, 1987

Type locality: Kleberg Co., Texas

Distribution: Australia, D.R. Congo, Cuba, France, Japan, USA, South Africa, Zambia

Hosts: Charadrius melanops, Ch. tricollaris, Erythrogonys cinctus; Lobibyx miles, L. novaehollandiae, Vanellus [sub syn. Hemiparra] crassirostris, V. spinosus (Charadriiformes: Charadriidae), Himantopus himantopus, H. leucocephalus, H. mexicanus, Recurvirostra americana (Charadriiformes: Recurvirostridae)

Rhinonyssus kadrae Dimov

Rhinonyssus kadrae Dimov, 2013а: 118; 2018: 134

Type locality: Leningrad Province

Distribution: Russia

Host: Anas platyrhynchos (Anseriformes: Anatidae)

Rhinonyssus levinseni (Trägårdh)

Sommatericola levinseni Trägårdh, 1904b: 28

Sternostomum levinseni: Vitzthum, 1935: 569; Bregetova, 1951: 114; 1956: 198; Domrow, 1969: 326

Sternostomum levinsi: erratum

Rhinonyssus levinseni (Trägårdh): Hirst, 1921a: 357; van Eyndhoven, 1964: 300; Butenko, 1984: 65; Dimov, 2016 ; 2018: 140

Type locality: Greenland

Distribution: Australia, Greenland, Holand, Russia

Hosts: Anas gibberifrons, A. platyrhynchos, A. querquedula, Anser anser, Polysticta stelleri, Somateria fischeri, S. molissima, S. spectabilis (Anseriformes: Anatidae)

Rhinonyssus marilae Butenko et Stanyukovich

Rhinonyssus marilae Butenko & Stanyukovich, 2001: 524

Type locality: Primorskiy Kray (Far East)

Distribution: Russia

Host: Aythya marila (Anseriformes: Anatidae)

Rhinonyssus minutus (Bregetova)

Sternostomum minutus Bregetova, 1950: 1007; 1951: 116; Fain, 1956: 150

Sternostoma minutus: Furman, 1957: 483

Rhinonyssus minutus: Fain, 1957b: 48;1961a: 514; Strandtmann & Wharton, 1958: 159; Domrow, 1965b: 450; 1967a: 216; 1969b: 326; Butenko, 1984: 36; Dimov, 2018: 141

Type locality: Russia

Distribution: Australia, Azerbaydzhan, Kirghizstan, Russia, Turkmenistan

Hosts: Charadrius alexandrinus, Ch. dubius, Ch. hiaticula (Charadriiformes: Charadriidae), Otis tarda (Otidiformes: Otitidae)

Rhinonyssus minutus leucopolius Fain

Rhinonyssus minutus leucopolius Fain, 1972: 363

Type locality: Anvers Zoo

Distribution: ?

Host: Charadrius [Leucopolius] pecuarius (Charadriiformes: Charadriidae)

Rhinonyssus neglectus Hirst

Rhinonyssus neglectus Hirst, 1921: 359; Vitzthum, 1935: 572; Castro, 1948: 257; Bregetova, 1951: 117; 1953: 332; Strandtmann, 1951: 132; Fain, 1957; Fain, 1960: 312; Domrow, 1969; Pence, 1975; Butenko, 1984: 23; Knee & Proctor, 2010; Dimov, 2016; 2018: 138

Rhinonyssus coniventris neglectus (Hirst): Fain, 1963: 91; Turk, 1953: 13

Type locality: Shetland Islands

Distribution: Australia, Azerbajdjan, Canada, Germany, Shetland Islands, Kazakhstan, Russia (incl. Maritime Distr.), Rwanda

Hosts: Arenaria interpres, Calidris alpina, C. cornutus, C. maritima, Charadrius alexandrinus, Tringa glareola, T. ochropus (Charadriiformes: Charadriidae)

Rhinonyssus pluvialis Fain et Johnston

Rhinonyssus pluvialis Fain & Johnston, 1966: 27

Type locality: 5 km S Wooster, Ohio

Distribution: USA (Ohio)

Host: Pluvialis dominica (Charadriiformes: Charadriidae)

Rhinonyssus podicipedus Feider et Mironescu

Rhinonyssus podicipedus Feider & Mironescu, 1972: 317; Butenko, 1984: 51

Rhinonyssus tschomgae Butenko, 1973: 244 – type: Azerbajdzhan, Kyzyl Agach (syn. by Butenko, 1984: 51)

Type locality: Sfintu Georghe (Tulcea)

Distribution: Azerbajdzhan, Romania, Russia, Uzbekistan

Host: Podiceps cristatus (Podicipiformes: Podicipedidae)

Rhinonyssus podilymbi Pence

Rhinonyssus podilymbi Pence, 1972a: 154

Type locality: Louisiana, Cameron Parish

Distribution: USA (La)

Host: Podilymbus podilymbi (Podicipediformes: Podicipedidae)

Rhinonyssus poliocephali Fain

Rhinonyssus poliocephali Fain, 1956a: 149; 1957b: 45; Strandtmann & Wharton, 1958: 159; Domrow, 1965b: 450; 1967a: 216; 1969b: 325

Rhinonyssus polycephali: Butenko, 1984: 53

Type locality: Astrida (= Butare), Rwanda

Distribution: Australia, Rwanda

Hosts: Podiceps [Poliocephalus] ruficollis capensis, P. ruficollis (Podicipiformes: Podicipedidae)

Rhinonyssus polystictae Butenko

Rhinonyssus polystictae Butenko, 1984: 67; Dimov, 2016; 2018: 139

Type locality: Yakutia, Kitisno

Distribution: Russia (Yakutia)

Hosts: Polysticta stelleri, Somateria mollissima (Anseriformes: Anatidae)

Rhinonyssus rhinolethrum (Trouessart)

Sternostomum rhinolethrum Trouessart, 1895: 393; Berlese, 1912: 71; Hirst, 1922: 89; Vitzthum, 1935; 1941: 656; Bregetova, 1951: 114

Rhinonyssus rhinolethrum: de Castro, 1948; Strandtmann, 1951: 132; 1956: 137; Strandtmann & Wharton, 1958: 159; Fain, 1958c: 9; 1960: 108; 1962b: 267; Mitchell & Rhodes, 1960: 107; Mitchell, 1963a: 506, 1963b; Domrow, 1969b: 324; Wilson, 1970: 74; Pence, 1972: 156; Butenko, 1984: 57; Mascarenhas et al., 2009: 295

Sternostoma rhinolethrum: Zumpt & Till, 1955: 90

Rhinonyssus rhinolethrus: Domrow, 1966: 196; 1967a: 216

Rhinonyssus anatidae Butenko, 1971: 490 – Russia (syn. by Domrow, 1979: 513)

Rhinonyssus rhinolethrum anatidae: Butenko, 1971: 490; 1984: 61

Rhinonyssus subrhinolethrum Butenko, 1971: 491; 1984: 62; Sartbaev, 1977; 1979: 56 – type: Spasskiy Rayon, Ryazanskaya Obl., Russia; Dimov, 2016; 2018: 135 (syn. by Domrow, 1979: 513)

Rhinonyssus nyrocae Butenko, 1971a: 492; 1984: 64; Dimov, 2016; 2018: 135 – type: Danube Delta, from Aythia spp. Syn. by Domrow, 1979: 513

Type locality: Europe

Distribution: Antarctica (South Georgia, etc.), Argentina, Australia, Azerbaijan, Belgium, Botswana, Brazil, Canada, D.R. Congo, Cuba, France, Greenland, Holland, Kazakhstan, Kirghizstan, Papua New Guinea, Russia, Ukraine, USA, Uzbekistan

Hosts: Aix sponsa, Alopochen aegyptiacus, Anas acuta, A. [Mareca] americana, A. carolinensis, A. clypeata, A. crecca, A. discors, A. erythrorhyncha, A. falcata, A. georgicus, A. gibberifrons, A. penelope, A. platyrhynchos, A. querquedula, A. strepera, A. superciliosa, Anser albifrons, A. anser, A. rossii, Aythia affinis, A. baeri, A. ferina, A. fuligula, A. nyroca, Branta canadensis, Callonetta leucophrys, Chenonetta jubata, Cygnus columbianus, C. melanocoryphus, Dendrocygna arcuata, D. bicolor, D. viduata, Metopiana peposaca, Netta rufina, Spatula clypeata, Tadorna tadorna (Anseriformes: Anatidae); Tringa nebularia (Charadriiformes: Scolopacidae)

Rhinonyssus shcherbininae Butenko

Rhinonyssus shcherbininae Butenko, 1984: 25

Type locality: Turkmenistan

Distribution: Kazakhstan, Kirghizstan, Turkmenistan

Host: Charadrius alexandrinus (Charadriiformes: Charadriidae)

Rhinonyssus sphenisci Fain et Mortelmans

Rhinonyssus sphenisci Fain & Mortelmans, 1959: 13; Gastal et al., 2018: 487

Type locality: Peru

Distribution: Brazil, Peru

Hosts: Spheniscus humboldti, S. magellanicus (Sphenisciformes: Spheniscidae)

Rhinonyssus sphenisci schelli Fain et Hyland

Rhinonyssus sphenisci schelli Fain & Hyland, 1963: 4

Rhinonyssus schelli Fain & Hyland: Wilson, 1970: 74; 1971: 17

Type locality: Base Roi Baudouin

Distribution: Antarctica

Hosts: Pygoscelis adeliae, P. papua (Sphenisciformes: Spheniscidae)

Rhinonyssus spinactitis Dusbábek

Rhinonyssus spinactitis Dusbábek, 1969: 213; Černy & Dusbábek, 1970: 485

Type locality: Canal de Vijil, La Gloria, Prov. de Camagüey

Distribution: Cuba

Host: Actitis macularia (Charadriiformes: Scolopacidae)

Rhinonyssus strandtmanni Fain et Johnson

Rhinonyssus strandtmanni Fain & Johnson, 1966: 25

Type locality: 5 km S of Wooster, Ohio

Distribution: Rwanda, USA (Ohio)

Hosts: Charadrius tricollaris, Ch. vociferus (Charadriiformes: Charadriidae)

Rhinonyssus tringae Fain

Rhinonyssus tringae Fain, 1963b: 96; Černy & Dusbábek, 1970: 485; Amaral & Reboucas, 1974: 55; Butenko, 1984: 31; de Rojas, Mora et al., 2002: 676; de Rojas et al., 2007: 1041

Type locality: Astrida (= Butare), Rwanda

Distribution: Cuba, Russia, Rwanda, Spain

Hosts: Tringa glareola, T. melanoleuca, T. ochropus, T. totanus (Charadriiformes: Scolopacidae)

Rhinonyssus vanellochettusiae Butenko

Rhinonyssus vanellochettusiae Butenko, 1984: 43

Type locality: Imam-Baba

Distribution: Turkmenistan

Host: Vanellochettusia leucura (Charadriiformes: Charadriidae)

Rhinonyssus vanellus Fain

Rhinonyssus afribyx vanellus Fain, 1972: 362

Type locality: Brasschaat nr Anvers, Belgium

Rhinonyssus vanellus Butenko, 1973: 1872; Sartbaev, 1979: 56

Type locality: Novosibirskaya Obl., Karasukskiy Rayon

Distribution: Belgium, Kazakhstan, Kirghizstan, Russia

Host: Vanellus vanellus (Charadriiformes: Charadriidae)

Rhinonyssus waterstoni Hirst

Rhinonyssus waterstoni Hirst, 1921: 359; Castro, 1948: 256; Strandtmann, 1956: 134; Fain, 1957: 44; Butenko, 1969: 208, 1984: 34; Dimov, 2018: 143

Sternostomum waterstoni: Vitzthum, 1935: 569; Bregetova, 1951: 116; 1956: 198; Turk, 1953: 13

Type locality: Shetland Is.

Distribution: Germany, Russia, Shetland Is., USA

Hosts: Alca torda, Uria aalge, U. lomyia (Charadriiformes: Alcidae)

Subfam. Ptilonyssinae (Castro) Fain

Ptilonyssinae (Castro, 1948) Fain, 1957: 155

Ptilonyssidae: Castro, 1948

Type genus: Ptilonyssus Berlese et Trouessart, 1889

Genus Passeronyssus Fain

Passeronyssus Fain, 1960: 110

Type species: Ptilonyssus viduae Fain, 1956

Passeronyssus kittacinclae Fain

Passeronyssus kittacinclae Fain, 1964: 300

Type locality: Zoo Anvers

Distribution: South Asia

Host: Kittacincla malabarica (Passeriformes: Muscicapidae)

Passeronyssus megaluri Sakakibara

Passeronyssus megaluri Sakakibara, 1968: 305

Type locality: West Sepik Distr., Telefomin

Distribution: New Guinea

Host: ? Megalurus timoriensis (Passeriformes: Megaluridae)

Passeronyssus urieli do Amaral

Passeronyssus urieli do Amaral, 1973: 68

Type locality: Cipó, Est de São Paulo

Distribution: Brazil

Host: Myiarchus ferox australis (Passeriformes: Tyrannidae)

Passeronyssus viduae (Fain)

Ptilonyssus viduae Fain, 1956a: 147

Passeronyssus viduae (Fain): Fain, 1960: 110

Type locality: Astrida (= Butare)

Distribution: Rwanda

Host: Vidua macroura (Passeriformes: Viduidae)

Passeronyssus zeferinoi Amaral

Passeronyssus zeferinoi Amaral, 1973: 73

Type locality: Hanhaem, Est. de São Paulo

Distribution:Brazil

Host: Cacicus haemorrhow (Passeriformes: Icteridae)

Genus Locustellonyssus Bregetova

Locustellonyssus Bregetova, 1965: 1093

Type species: Locustellonyssus amurensis Bregetova, 1965

Locustellonyssus amurensis Bregetova

Locustellonyssus amurensis Bregetova, 1965: 1093

Type locality: Amuro-Zeyskoe Plato (Amurskaya obl.)

Distribution: Russia

Host: Locustella fasciolata (Passeriformes: Megaluridae)

Locustellonyssus sibiricus Butenko et Stanyukovich

Locustellonyssus sibiricus Butenko & Stanyukovich, 2001: 526

Type locality: Manatka, Tomskaya Oblast

Distribution: Russia

Host: Locustella certhiola (Passeriformes: Megaluridae)

Genus Pipronyssus Fain et Aitken

Pipronyssus Fain & Aitken, 1967: 36

Type species: Pipronyssus manaci Fain et Aitken, 1967

Pipronyssus manaci Fain et Aitken

Pipronyssus manaci Fain & Aitken, 1967: 36; 1970: 349

Type locality: Elgin Trace, Vega de Oropouche

Distribution: Brazil, Trinidad

Hosts: Pipra erythrocephala, P. iris, P. pipra (Passeriformes: Pipridae)

Genus Ptilonyssus Berlese et Trouessart

Ptilonyssus Berlese & Trouessart, 1889: 128

Type species: Ptilonyssus echinatus Berlese et Trouessart, 1889

Neonyssus Hirst, 1922: 771

Type species: Neonyssus intermedius Hirst, 1921

Rhinonyssoides Hirst, 1922: 770 – syn. by Zumpt & Till, 1955: 69

Type species: Rhinonyssoides trouessarti Hirst, 1922

Neonyssoides Hirst, 1923: 976

Type species: Rhinonyssus (Neonyssoides) nucifragae Hirst, 1923

Flavionyssus Castro, 1948: 266 – syn. by Zumpt & Till, 1955: 69

Type species: Ptilonyssus (Flavionyssus) rabelloi Castro, 1948

Rochanyssus Castro, 1948: 272 – syn. by Zumpt & Till, 1955: 69

Type species: Ptilonyssus (Rochanyssus) werneri Castro, 1948

Paraneonyssus Castro, 1948: 274 – syn. by Zumpt & Till, 1955: 69

Type species: Neonyssus (Paraneonyssus) enriettii Castro, 1948

Travanyssus Castro, 1948: 276 – syn. by Zumpt & Till, 1955: 69

Type species: Neonyssus (Travanyssus) paranensis Castro, 1948

Rhinacarus Castro, 1948: 257

Type species: Rhinonyssus (Rhinacarus) angrensis Castro, 1948

[Vitznyssus Castro, 1948: 277 – syn. by Zumpt & Till, 1955: 69, but restaured by Butenko, 1977]

Type species: Dermanyssus nitzschi Giebel, 1871

Cas Baker et Wharton, 1952: 81

Type species: Rhinonyssus (Rhinacarus) angrensis Castro, 1948

Hapalognatha Butenko, 1959: 17; 1960: 1490

Type species: Hapalognatha prima Butenko, 1959 (= Ptilonyssus motacillae Fain, 1956)

Periglischrodes Baker & Delfinado, 1964: 589 – Syn. by Bregetova, 1967: 135

Type species: Periglischrodes gressitti Baker et Delfinado, 1964 – transferred to Rhinonyssidae by Prasad (1969a)

Ptilonyssoides Vitzthum, 1935

Type species: Ptilonyssoides triscutatus Vitzthum, 1935

Spizonyssus Bregetova, 1967

Type species: Neonyssus (Spizonyssus) pygmaeus Bregetova, 1967

Ptilonyssus acanthopneustes Stanyukovich et Butenko

Ptilonyssus acanthopneustes Stanyukovich & Butenko, 2003: 37

Type locality: Tyumenskaya Oblast

Distribution: Russia

Host: Phylloscopus borealis (Passeriformes: Phylloscopidae)

Ptilonyssus acrocephali Fain

Ptilonyssus acrocephali Fain, 1964a: 55; Domrow, 1969b: 358; 1988: 868; Pence, 1972; 1975: 23; Fain, Sixl & Moritsch, 1974: 3; Spicer, 1987: 261; Knee, Proctor & Galloway, 2008: 369; Knee & Galloway, 2017: 95

Type locality: Deurne (Anvers, Belgium)

Distribution: Australia, Austria, Belgium, Canada, USA

Hosts: Acrocephalus arundinaceus, A. australis, A. schoenobaenus, A. s. scirpaceus (Passeriformes: Acrocephalidae), Regulus calendula (Passeriformes: Regulidae)

Ptilonyssus afroturdi Fain

Ptilonyssus afroturdi Fain, 1962c: 130

Type locality: Akanyaru

Distribution: Rwanda

Host: Turdus olivaceus graueri (Passeriformes: Turdidae)

Ptilonyssus agelaii Fain

Ptilonyssus agelaii Fain & Aitken, 1967: 7

Type locality: Biche-Ortoire Lagoon, Biche

Distribution: Trinidad

Host: Agelaius icterocephalus (Passeriformes: Icteridae)

Ptilonyssus ailuroedi Domrow

Ptilonyssus ailuroedi Domrow, 1964a: 31; 1964b: 348; 1964d: 619; 1988: 868

Type locality: Mt. Glorious, S.E. Queensland

Distribution: Australia

Host: Ailuroedus crassirostris (Passeriformes: Ptilonorhynchidae)

Ptilonyssus alaudae (Butenko et Stanyukovich)

Neonyssus (Otocorinyssus) alaudae Butenko & Stanyukovich, 2001: 520

Astridiella alaudae (Butenko):

Type locality: Turkmenistan

Distribution: Turkmenistan

Host: Alauda arvensis (Passeriformes: Alaudidae)

Ptilonyssus amarali Fain

Ptilonyssus amarali Fain, 1964b: 7; Fain & Aitken, 1967: 6

Type locality: Anvers Zoo

Distribution: Brazil, Trinidad

Hosts: Cyanerpes caeruleus, C. cyanea (Passeriformes: Thraupidae)

Ptilonyssus amazonicus Fain

Ptilonyssus amazonicus Fain, 1972: 358

Type locality: Redondo Lake, nr. Manaos

Distribution: Brazil

Host: Aves

Ptilonyssus ammomani Stanyukovich et Butenko

Ptilonyssus ammomani Stanyukovich & Butenko, 2003: 32

Type locality: Karagan, Ashhabad Region

Distribution: Turkmenistan

Host: Ammomanes deserti (Passeriformes: Alaudidae)

Ptilonyssus andropadi Fain

Ptilonyssus andropadi Fain, 1956a: 141; Strandtmann, 1960: 144

Type locality: Rugege Forest (Rwanda)

Distribution: Malaysia (Sabah), Rwanda

Hosts: Andropadus latirostris eugenius, Hypsipetes flavala connectens, Phyllastrephus fischeri sucosus (Passeriformes: Pycnonotidae)

Ptilonyssus angrensis (Castro)

Rhinonyssus (Rhinacarus) angrensis Castro, 1948: 257

Cas angrensis: Strandtmann & Wharton, 1958: 169

Sternostoma angrensis (Castro): Zumpt & Till, 1955: 91; Fain, 1957: 84; Amaral, 1968: 122; Sartbaev, 1983: 84

Ptilonyssus angrensis: Domrow, 1969b: 348; 1988: 868; Pence, 1973: 887; Knee, Proctor & Galloway, 2008: 369

Type locality: Fazenda Japuiba, Angra dos Reis, Brazil

Distribution: Australia, Brazil, Canada, Kirghizstan, USA

Hosts: Hirundo aethiopica, H. neoxena, Progne chalybea domestica, P. subis, Ptyonoprogne rupestris, Riparia riparia (Passeriformes: Hirundinidae)

Ptilonyssus angustirostris Fain et Aitken

Ptilonyssus angustirostris Fain & Aitken, 1968: 12; 1970: 353

Type locality: APEG Forest

Distribution: Brazil

Host: Glyphorynchus spirurus (Passeriformes: Furnariidae, Dendrocolapinae)

Ptilonyssus anthi Stanyukovich et Butenko

Ptilonyssus anthi Stanyukovich & Butenko, 2003: 43

Type locality: Ryazanskaya Oblast

Distribution: Russia

Host: Anthus trivialis (Passeriformes: Motacillidae)

Ptilonyssus astridae Fain

Ptilonyssus astridae Fain, 1956a: 141; 1957b: 98; Fain & Lukoschus, 1979: 10; Domrow, 1988: 868; 1992: 1605; Fain, Herin & Puylaert, 1977: 386

Paraneonyssus astridae Strandtmann & Wharton, 1958: 176

Type locality: Astrida (= Butare), Rwanda

Distribution: Australia, Cameroon, Rwanda

Hosts: Donacola castaneothorax (Passeriformes: Ploceidae), Euschistospiza dybowskii, Lagonosticta senegala ruberrima, L. rubricata congica, Poephila personata (Passeriformes: Estrildidae), Vidua macroura (Passeriformes: Viduidae)

Ptilonyssus aureliani Fain

Ptilonyssus aureliani Fain, 1956a: 146; 1957b: 109; Fain, Herin & Puylaert, 1977: 386

Type locality: Astrida (= Butare, Rwanda)

Distribution: Cameroon, Rwanda

Hosts: Camaroptera brevicaudata, Prinia subflava graueri (Passeriformes: Cisticolidae)

Ptilonyssus balimoensis Sakakibara

Ptilonyssus balimoensis Sakakibara, 1968a: 17; Domrow, 1969: 368; 1988: 869

Type locality: Western District, Balimo (Papua New Guinea)

Distribution: Australia, Papua New Guinea

Hosts: Meliphaga macleayana, Xanthotis flaviventer (Passeriformes: Meliphagidae)

Ptilonyssus belemensis Fain et Aitken

Ptilonyssus belemensis Fain & Aitken, 1968: 4

Type locality: Mucambo Forest

Distribution: Brazil

Host: unknown (Passeriformes: ? Furnariidae, Dendrocolaptinae)

Ptilonyssus bombycillae Fain

Ptilonyssus bombycillae Fain, 1972: 357; Rodríguez, Úbeda & Guevara, 1993: 13; Knee, Proctor & Galloway, 2008: 369

Ptilonyssus bombycillae Pence, 1973c: 360 – type from Bombycilla cedrorum, New Orleans, La. (USA); Spicer, 1978: 891

Type locality: Waarschoot

Distribution: Belgium, Canada, Spain, USA

Hosts: Bombycilla cedrorum, B. garrulus (Passeriformes: Bombycillidae)

Ptilonyssus bradypteri (Fain)

Passeronyssus bradypteri Fain, 1962a: 143

Ptilonyssus bradypteri: Domrow, 1969b: 358; 1988: 869

Type locality: Akanyaru (Rwanda)

Distribution: Australia, Rwanda

Hosts: Bradypterus baboecala centralis (Passeriformes: Megaluridae), Cinclorhamphus mathewsi (Passeriformes: Sylviidae)

Ptilonyssus buloloensis Sakakibara

Ptilonyssus buloloensis Sakakibara, 1968c: 426; Domrow, 1988: 869

Type locality: Bulolo River, Papua New Guinea

Distribution: Australia, Papua New Guinea

Host: Manucodia (Phonygammus) keraudrenii (Passeriformes: Paradisaeidae)

Ptilonyssus cacici Fain

Ptilonyssus cacici Fain, 1964b: 9

Type locality: Anvers Zoo

Distribution: South America

Host: Cacicus sp. (Passeriformes: Icteridae)

Ptilonyssus calamocichlae Fain

Ptilonyssus calamocichlae Fain, 1956a: 142

Type locality: Akanyaru

Distribution: Rwanda

Host: Acrocephalus [Calamocichla] rufescens foxi (Passeriformes: Acrocephalidae)

Ptilonyssus calamocichlae hippolaisi Fain

Ptilonyssus calamocichlae hippolaisi Fain, 1963c: 78

Type locality: Akanyaru

Distribution: Rwanda

Host: Hippolais icterina (Passeriformes: Acrocephalidae)

Ptilonyssus calandrellae (Bregetova)

Neonyssus (Otocorinyssus) calandrellae Bregetova, 1967: 129

Astridiella calandrellae (Bregetova): Úbeda et al., 1986: 167

Type locality: at Zeravshan River, 15 km upstream Penjikent (Tajikistan)

Distribution: Georgia, Spain, Tadjikistan

Host: Calandrella cinerea (Passeriformes: Alaudidae)

Ptilonyssus callinectoides (Brooks et Strandtmann)

Tyranninyssus callinectoides Brooks & Strandtmann, 1960: 420; Hyland & Moorhouse, 1970: 53

Ptilonyssus callinectoides (Brooks et Strandtmann): Pence, 1972c: 790; Knee, Proctor & Galloway, 2008: 369

Type locality: Briscoe County, Texas

Distribution: Canada, Mexico, USA (La., Texas)

Hosts: Myiarchus cinerascens, M. crinitus, M. tyrannulus (Passeriformes: Tyrannidae)

Ptilonyssus calvaria Knee

Ptilonyssus calvaria Knee, 2008: 350; Knee & Galloway, 2017: 93

Type locality: Winnipeg (Manitoba)

Distribution: Canada (Manitoba, Alberta)

Hosts: Passerculus sandwicensis, Spizella passerina (Passeriformes: Passerellidae)

Ptilonyssus camptostoma Hyland et Moorhouse

Ptilonyssus camptostoma Hyland & Moorhouse, 1970: 44

Type locality: Tlacotalpan (Veracruz)

Distribution: Mexico

Host: Camptostoma imberbe (Passeriformes: Tyrannidae)

Ptilonyssus capensis Zumpt et Till

Ptilonyssus capensis Zumpt & Till, 1955: 72; Fain, 1956a: 138; 1957b: 89; Butenko, 1965: 132; Úbeda-Ontíveros & Guevara-Benítez, 1978

Neonyssus capensis (Zumpt et Till): Shumilo & Lunkashu, 1970: 19; Sartbaev, 1983: 84

Paraneonyssus capensis: Strandtmann & Wharton, 1958: 176

Type locality: Cape Province

Distribution: Moldavia, Kirghizstan, Russia, Rwanda, South Africa, Spain

Hosts: Anthus campestris, A. pratensis, A. spinoletta, A. trivialis, Macronyx capensis, M. croceus, Motacilla alba (Passeriformes: Motacillidae)

Ptilonyssus capitatus (Strandtmann)

Paraneonyssus capitatus Strandtmann, 1956: 133; Strandtmann & Wharton, 1958: 177

Astridiella capitatus (Str.): Fain, 1959: 21; Butenko, 1965: 132

Ptilonyssus capitatus Domrow, 1969b: 339; 1988: 869

Type locality: Tooele, Utah

Distribution: Australia, Russia, Rwanda, South Africa, USA (Utah)

Hosts: Alauda arvensis, Mirafra javanica, Otocoris alpestris (Passeriformes: Alaudidae), Teloporus dohertyi, T. zeylonus (Passeriformes: Malaconotidae)

Ptilonyssus carduelis Fain

Ptilonyssus carduelis Fain, 1962b: 253; Wilson & Haas, 1980: 545; Úbeda-Ontíveros, Guevara-Benítez & Morillas-Marquez, 1985: 272; Knee, Proctor & Galloway, 2008: 368

Neonyssus (Fringilonyssus) carduelis (Fain): Bregetova, 1967: 131; Shumilo & Lunkashu, 1970: 20; Beron, 1975: 187; Sartbaev, 1983: 84

Type locality: Anvers (Belgium)

Distribution: Belgium, Bulgaria, Canada, Kirghizstan, Moldavia, Russia, Spain, Tajikistan, USA (Alaska); Australia (introduced)

Hosts: Acanthis flavirostris, Carduelis [Acanthis] c. cannabina, Carpodacus erythrinus, C. flammea, C. rhodochlamys, C. rubicilla, Leucosticte brandti, Loxia curvirostra, L. leucoptera, Pinicola enucleator (Passeriformes: Fringillidae)

Ptilonyssus cerchneis Fain

Ptilonyssus cerchneis Fain, 1957b: 133; Strandtmann, 1962: 100; Bregetova, 1964: 5; Domrow, 1965b: 457; 1969b: 374; 1988: 869; Butenko, 1984: 156; Knee, Proctor & Galloway, 2008: 368; Knee & Galloway, 2017: 95

Rhinonyssoides cerchneis, Strandtmann & Wharton, 1958: 172

Type locality: Rugege (Rwanda)

Distribution: Australia, Canada, Russia, Rwanda, USA

Hosts: Falco berigora, F. cenchroides, F. naumanni, F. sparverius, F. tinnunculus, F. [Cerchneis] tinnunculus rufescens (Falconiformes: Falconidae), Turdus migratorius (Passeriformes: Turdidae)

Ptilonyssus cercomacrae Fain et Aitken

Ptilonyssus cercomacrae Fain & Aitken, 1970: 390

Type locality: APEG (Varsea), IPEAN, Belém

Distribution: Brazil

Host: Cercomacra tyrannina (Passeriformes: Formicariidae)

Ptilonyssus certhiae Fain et Bafort

Ptilonyssus certhiae Fain & Bafort, 1963b: 483; Fain, Sixl & Moritsch, 1974: 3

Type locality: St Laureins (Belgium)

Distribution: Austria, Belgium

Host: Certhia b. brachydactyla (Passeriformes: Certhiidae)

Ptilonyssus certhiaxicola Fain

Ptilonyssus certhiaxicola Fain, 1964: 31; Fain & Aitken, 1967: 6; 1970: 356

Type locality: Anvers Zoo

Distribution: Brazil, Trinidad

Hosts: Certhiaxis cinnamomea, Synallaxis rutilans (Passeriformes: Furnariidae)

Ptilonyssus certhilaudae Fain

Ptilonyssus certhilaudae Fain, 1972: 360

Type locality: Bloemhof, Transvaal

Distribution: South Africa

Host: Certhilauda albofasciata (Passeriformes: Alaudidae)

Ptilonyssus chalybaedomesticae Amaral

Ptilonyssus chalybaedomesticae Amaral, 1967: 169; 1968: 121; Černý & Dusbábek, 1970: 482

Type locality: Angra dos Reis (Rio de Janeiro)

Distribution: Brazil

Host: Progne chalybea domestica (Passeriformes: Hirundinidae)

Ptilonyssus chiroxiphiae Fain et Aitken

Ptilonyssus chiroxiphiae Fain & Aitken, 1970: 361

Type locality: APEG (Station A), IPEAN, Belém

Distribution: Brazil

Host: Chiroxiphia pareola (Passeriformes: Pipridae)

Ptilonyssus chloris Fain

Ptilonyssus carduelis chloris Fain, 1962b: 257

Neonyssus chloris (Fain): Shumilo & Lunkashu, 1970: 20

Ptilonyssus (Frigilonyssus) carduelis chloris Fain: Bregetova, 1967: 130

Ptilonyssus chloris Fain: Rodríguez, Úbeda, Guevara & Ariza, 1989: 152

Type locality: Anvers (Belgium)

Distribution: Belgium, Moldavia, Russia, Spain

Host: Chloris chloris (Passeriformes: Fringillidae)

Ptilonyssus chlorocichlae Fain

Ptilonyssus chlorocichlae Fain, 1956a: 142

Type locality: Akanyaru

Distribution: Rwanda

Host: Chlorcichla flavicollis pallidigula (Passeriformes: Pycnonotidae)

Ptilonyssus chloropsicola Fain

Ptilonyssus chloropsicola Fain, 1966: 90

Type locality: Anvers (Zoo)

Distribution: Southeast Asia

Host: Chloropsis cochinchinensis (Passeriformes: Chloropseidae)

Ptilonyssus cinnyricincli Fain

Ptilonyssus cinnyricincli Fain, 1962c: 132

Type locality: Buzi

Distribution: Mozambique

Host: Cinnyricinclus leucogaster (Passeriformes: Sturnidae)

Ptilonyssus cinnyris Zumpt et Till

Ptilonyssus cinnyris Zumpt & Till, 1955: 78; Fain, 1956a: 146; 1957b: 86; Nadchatram, McClure & Chong, 1964: 112; Domrow, 1969: 374; 1988: 869

Rhinonyssoides cinnyris Strandtmann & Wharton, 1958: 172

Type locality: Cape Prov. (South Africa)

Distribution: Australia, Malaysia, Rwanda, South Africa

Hosts: Arachnothera longirostris, Chalcomitra senegalensis aequatorialis, Cinnyris afra, C. regius kivuensis, Cyrtostomus frenatus, Nectarinia kilimensis (Passeriformes: Nectariniidae)

Ptilonyssus cisticolarum Fain

Ptilonyssus cisticolarum Fain, 1959a: 26; 1960: 109

Type locality: Rustenburg, Transvaal

Distribution: Botswana, Rwanda, South Africa

Hosts: Apalis flavida caniceps, Cisticola robusta nuchalis, C. lais (Passeriformes: Cisticolidae)

Ptilonyssus coccothraustis Fain et Bafort

Ptilonyssus coccothraustis Fain & Bafort, 1963b: 477; Sixl, 1969: 218; Fain, Sixl & Moritsch, 1974: 3

Neonyssus coccothraustis: Bregetova, 1967: 130; Shumilo & Lunkashu, 1970: 20

Type locality: Anvers (Belgium)

Distribution: Austria, Belgium, Canada, Georgia, Kyrghizstan, Moldavia, Ukraine

Hosts: Cocothraustes coccothraustes, C. vespertinus (Passeriformes: Fringillidae)

Ptilonyssus colluricinclae Domrow

Ptilonyssus colluricinclae Domrow, 1964d: 596; 1969b: 339; 1979d: 494; 1988: 870

Type locality: Brisbane (Qu.)

Distribution: Australia

Hosts: Colluricincla phaea (= harmonica), C. megarhyncha, Pachycephala griseiceps, P. pectoralis (Passeriformes: Pachycephalidae)

Ptilonyssus colopteryx Fain et Aitken

Ptilonyssus colopteryx Fain & Aitken, 1971: 32

Type locality: APEG Forest, Station A (IPEAN)

Distribution: Brazil

Host: Colopteryx galeatus (Passeriformes: Tyrannidae)

Ptilonyssus condylocoxa Fain et Lukoschus

Ptilonyssus condylocoxa Fain & Lukoschus, 1979: 11; Koch, 1986: 60; Domrow, 1988: 870

Type locality: Beverley Springs, Western Australia

Distribution: Australia

Host: Ramsayornis fasciatus (Passeriformes: Meliphagidae)

Ptilonyssus conopophagae Fain et Aitken

Ptilonyssus conopophagae Fain & Aitken, 1970: 378

Type locality: Mosqueiro Ferry, Benavides, Belém

Distribution: Brazil

Host: Conopophaga roberti (Passeriformes: Conopophagidae)

Ptilonyssus conopophilae Fain et Lukoschus

Ptilonyssus conopophilae Fain & Lukoschus, 1979: 13; Koch, 1986: 60; 1988: 870

Type locality: Napier Downs, Western Australia

Distribution: Australia

Hosts: Conopophila rufogularis, Grantiella picta (Passeriformes: Meliphagidae), ? Taeniopygia [Poephila guttata] castanotis (Passeriformes: Estrildidae)

Ptilonyssus corcoracis Domrow

Ptilonyssus corcoracis Domrow, 1969: 345; 1988: 870

Type locality: Condamine, Charleville

Distribution: Australia

Host: Corcorax melanorhamphus (Passeriformes: Corcoracidae)

Ptilonyssus corvi Pence

Ptilonyssus corvi Pence, 1972c: 791

Type locality: Laplace, Louisiana

Distribution: USA (La.)

Host: Corvus brachyrhynchos (Passeriformes: Corvidae)

Ptilonyssus corythopicola Fain et Aitken

Ptilonyssus corythopicola Fain & Aitken, 1970: 372

Type locality: Mosqueiro Ferry, Benavides, Belém

Distribution: Brazil

Host: Corythopis torquata (Passeriformes: Conopophagidae)

Ptilonyssus cractici Domrow

Ptilonyssus cractici Domrow, 1964d: 598, 616; 1966b: 196; 1969b: 349; 1972: 100; 1979: 497; Fain & Lukoschus, 1979: 10

Type locality: Brisbane (Qu.), Condamine, Logan Village (Australia)

Distribution: Australia, New Zealand

Hosts: Cracticus mentalis, C. nigrogularis, C. quoyi, C. torquatus (Passeriformes: Cracticidae), Dacelo gigas (Coraciiformes: Alcedinidae), Gymnorhina hypoleuca, G. tibicen, Lalage tricolor (Passeriformes: Compephagidae), Strepera graculina (Passeriformes: Artamidae)

Ptilonyssus cyanosylviae Butenko et Lavrovskaya

Ptilonyssus cyanosylviae Butenko & Lavrovskaya, 1980: 530

Type locality: Novosibirskaya Oblast

Distribution: Russia

Host: Cyanosylvia svecica (Passeriformes: Muscicapidae)

Ptilonyssus daptrii Fain

Ptilonyssus daptrii Fain, 1966: 88

Type locality: Rio Preto da Eva

Distribution: Brazil

Host: Daptrius ater (Falconiformes: Falconidae)

Ptilonyssus degtiarevae Dimov et Mironov

Ptilonyssus degtiarevae Dimov & Mironov, 2012: 169

Type locality: Gavrilovo Village, Leningradskaya Obl.

Distribution: Russia

Host: Passer domesticus (Passeriformes: Passeridae)

Ptilonyssus dendrocittae (Sakakibara et Strandtmann)

Paraneonyssus dendrocitti Sakakibara & Strandtmann, 1968: 43

Ptilonyssus dendrocittae (Sakakibara & Strandtmann)

Type locality: Nan-T‘ou Hsien: Pu-Li

Distribution: Taiwan

Host: Dendrocitta [Crypsirina] occipitalis (Passeriformes: Corvidae)

Ptilonyssus dendrocolapticola Fain et Aitken

Ptilonyssus dendrocolapticola Fain & Aitken, 1968: 8

Type locality: APEG Forest

Distribution: Brazil

Host: Dendrocolaptes certhia (Passeriformes: Furnariidae, Dendrocolaptinae)

Ptilonyssus desfontainei Zumpt et Till

Ptilonyssus desfontainei Zumpt & Till, 1955: 71; Fain, 1957b: 84

Type locality: Transvaal

Distribution: South Africa

Host: Thamnolaea cinnamomeiventris (Passeriformes: Turdidae)

Ptilonyssus diadori Butenko et Lavrovskaya

Ptilonyssus diadori Butenko & Lavrovskaya, 1980: 530

Type locality: Russia

Distribution: Russia

Host: Aves

Ptilonyssus dicaei Domrow

Ptilonyssus dicaei Domrow, 1966a: 201; 1967a: 217; 1969b: 366; 1988: 870

Type locality: Mt. Jukes, near Mackay, N.Q.

Distribution: Australia

Host: Dicaeum hirundinaceum (Passeriformes: Dicaeidae)

Ptilonyssus dicruri Fain

Ptilonyssus dicruri Fain, 1956a: 137; Domrow, 1969b: 348; 1988: 871; Domrow & Nadchatram, 1978: 87; Feider & Mironescu, 1980: 61

Ptilonyssoides dicruri: Strandtmann & Wharton, 1958: 170; Strandtmann, 1960: 140

Passeronyssus dicruri (Fain): Fain, 1960: 110; Domrow, 1965b: 451

Type locality: Astrida (= Butare), Rwanda

Distribution: Australia, Botswana, Malaysia, Papua New Guinea, Rwanda, Thailand

Hosts: Chibia bracteata, Dicrurus adsimilis, D. hottentottus, D. leucophaeus stigmatops, D. macrocercus cathoecus, D. remifer (Passeriformes: Dicruridae)

Ptilonyssus dioptrornis Fain

Ptilonyssus dioptrornis Fain, 1956a: 137; 1959a: 26; Domrow, 1969b: 347; 1988: 871; Domrow & Nadchatram, 1978: 86; Kadosaka, Kaneko & Asanuma, 1983: 221

Ptilonyssoides dioptrornis Strandtmann & Wharton, 1958: 170

Passeronyssus dioptrornis (Fain): Fain, 1960: 110

Type locality: Rugege Forest (Rwanda)

Distribution: Australia, Japan, Malaysia, Rwanda, South Africa

Hosts: Cossypha [Bessonornis] archeri, C. dichroa, Dioptrornis fischeri toruensis (Passeriformes: Turdidae), Copsychus malabaricus, C. saularis musicus, Eopsaltria capito, Muscicapa c. cyanomelana, M. rufigastra (Passeriformes: Muscicapidae)

Ptilonyssus dolicaspis Feider et Mironescu

Ptilonyssus dolicaspis Feider & Mironescu, 1980: 59

Type locality: West Squak

Distribution: Papua New Guinea

Host: Megalurus timonensis (Passeriformes: Megaluridae)

Ptilonyssus domrowi Feider et Mironescu

Ptilonyssus domrowi Feider & Mironescu, 1980: 57

Type locality: Tambul

Distribution: Papua New Guinea

Host: Paramythia montium (Passeriformes: Paramythiidae)

Ptilonyssus donatoi Pereira et Castro

Ptilonyssus (Rhinonyssoides) donatoi Pereira & Castro, 1949: 224

Ptilonyssus donatoi: Zumpt & Till, 1955: 88; Fain, 1957b: 84; Amaral, 1968: 121

Type locality: Estado de Sao Paulo

Distribution: Brazil

Host: Coragyps atratus foetens (Cathartiformes: Catarthidae)

Ptilonyssus dryoscopi Zumpt et Till

Ptilonyssus dryoscopi Zumpt & Till, 1955: 73; Fain, 1956a: 139; 1957b: 84

Paraneonyssus dryoscopi (Zumpt et Till): Strandtmann & Wharton, 1958: 177

Type locality: Cape Prov. (South Africa)

Distribution: Rwanda, South Africa

Hosts: Dryoscopus cubla, Tchagra senegala (Passeriformes: Malaconotidae)

Ptilonyssus echinatus Berlese et Trouessart

Ptilonyssus echinatus Berlese & Trouessart, 1889: 9; Vitzthum, 1935: 578; Pereira & Castro, 1949: 219; Zumpt & Till, 1955: 74; Fain, 1956: 136; 1956: 114; 1957b: 137; 1957c: 218; 1960: 108; 1962b: 262; Strandtmann & Wharton, 1958: 179; Strandtmann, 1960: 146; George, 1961: 114; Domrow, 1964d: 607; 1969b: 349, 423; 1988: 871; Fain & Lukoschus, 1972: 90; Fain, Sixl & Moritsch, 1974: 1; Beron, 1975: 186; Scherbinina & Butenko, 1981: 70; Spicer, 1987: 261; Knee, Proctor & Galloway, 2008: 369; Huhta, 2016: 148 (S. incert.).

Ptilonyssoides echinatus (Berlese et Trouessart): Shumilo & Lunkashu, 1970: 29

Hapalognatha platytrichia Butenko, 1959: 17; 1960: 1494. – Type: Tatarstan (Russia), from Hirundo rustica (syn. by Fain, 1962b: 262)

Type locality: Germany

Distribution: Australia, Austria, Bulgaria, Cameroon, Canada, Cuba, ? Finland, Germany, Moldavia, Russia, South Africa, Spain, Suriname, Thailand, Trinidad, Turkmenistan, Ukraine, USA

Hosts: Atticora melanoleuca, Delichon urbica, Hirundo neoxena, H. nigrita, H. pyrrhonota, H. rustica, H. rustica erythrogaster, H. rustica gutturalis, Hylochelidon ariel, H. nigricans, Riparia riparia, Petrochelidon pyrrhonota, P. spilodera, Stelgidopteryx ruficollis, Tachycineta bicolor (Passeriformes: Hirundinidae)

Ptilonyssus elaeniae Fain et Aitken

Ptilonyssus elaeniae Fain & Aitken, 1967: 11

Type locality: La Horquette Trace, Tumpuna Road

Distribution: Trinidad

Host: Elaenia flavogaster (Passeriformes: Tyrannidae)

Ptilonyssus elbeli (Strandtmann)

Cas elbeli Strandtmann, 1960: 137

Ptilonyssus elbeli (Strandtmann): Domrow, 1968; Feider & Mironescu, 1978: 266; Sartbaev, 1981: 130 [Cas elbelli]

Type locality: Thailand

Distribution: Azerbaidjan, Belgium, Kirghizstan, Romania, Thailand

Hosts: Acridotheres tristis, Sturnus vulgaris (Passeriformes: Sturnidae)

Ptilonyssus elongatus Fain

Ptilonyssus elongatus Fain, 1964e:123

Type locality: Astrida (= Butare)

Distribution: Rwanda

Hosts: Apalis flavida caniceps, Prinia leucopogon reichenowi (Passeriformes: Cisticolidae)

Ptilonyssus emberizae Fain

Ptilonyssus emberizae Fain, 1956a: 140; 1957b: 95; 1964a; Sixl, 1969: 218; Fain, Sixl & Moritsch, 1974: 2; Domrow, 1972: 99; 1988: 871; Kadosaka, Kaneko & Asanuma, 1983: 221; Guevara Benítez, Úbeda Ontíveros & Morillas Marquez, 1986a: 75

Paraneonyssus emberizae Strandtmann & Wharton, 1958: 177

Fain & Lukoschus, 1979: 10

Neonyssus emberizae (Fain): Shumilo & Lunkashu, 1970: 21; Scherbinina & Butenko, 1981: 70

Type locality: Astrida (= Butare), Rwanda

Distribution: Australia, Austria, Belgium, Moldavia, New Zealand, Spain, Russia, Rwanda, Turkmenistan, Ukraine

Hosts: Emberiza bruniceps, E. cia, E. citrinella, E. flaviventris, E. schoeniclus, E. sulphurata (Passeriformes: Emberizidae), Lonchura pectoralis, Steganopleura bichenovii (Passeriformes: Estrildidae), Fringilla coelebs (Passeriformes: Fringillidae), Plectrophenax nivalis (Passeriformes: Calcariidae), Sturnus vulgaris (Passeriformes: Sturnidae)

Ptilonyssus enicuri Fain et Nadchatram

Passeronyssus enicuri Fain & Nadchatram, 1962: 276

Ptilonyssus enicuri Fain et Nadchatram: Kadosaka, Kaneko & Asanuma, 1983: 222

Type locality: Malaysia

Distribution: Japan, Malaysia

Hosts: Enicurus schistaceus, Erithacus a. akahige (Passeriformes: Turdidae)

Ptilonyssus enrietti Castro

Neonyssus (Paraneonyssus) enrietti Castro, 1948: 274; Pereira & Castro, 1949: 227

Ptilonyssus enrietti (Castro): Zumpt & Till, 1955: 89; Fain, 1957b: 89

Type locality: Brazil

Distribution: Brazil

Host: Phylloscartes ventralis ventralis (Passeriformes: Tyrannidae)

Ptilonyssus eremophilae Butenko et Lavrovskaya

Ptilonyssus eremophilae Butenko & Lavrovskaya, 1980: 526; Sartbaev, 1983: 84

Type locality: Ryazanskaya Obl. (Russia)

Distribution: Kirghizstan, Russia

Host: Eremophila alpestris (Passeriformes: Alaudidae)

Ptilonyssus estrildicola Fain

Ptilonyssus estrildicola Fain, 1959a: 22; Domrow, 1965: 453; Sakakibara, 1968b: 309

Type locality: Astrida (= Butare), Rwanda

Distribution: Australia, Botswana, Papua New Guinea, Rwanda, South Africa

Hosts: Amadina erythrocephala, Batis capensis, Granatina granatina, Lagonosticta rhodopareia, L. rubricata, Pytilia melba, Taeniopygia castanotis, Uraeginthus angolensis, U. bengalus ugandae (Passeriformes: Estrildidae)

Ptilonyssus euroturdi Fain et Hyland

Ptilonyssus euroturdi Fain & Hyland, 1963a: 381; Fain & Bafort, 1963b: 485; Fain, 1966: 7; Feider & Mironescu, 1968; Shumilo & Lunkashu, 1970: 29; Pence, 1972: 1162; Fain, Sixl & Moritsch, 1974: 3; Sartbaev, 1983: 85 ; Spicer, 1984: 800; Kadosaka, Kaneko & Asanuma, 1987: 36; Rodríguez, Úbeda & Guevara, 1990: 301; De Rojas et al., 2001: 961; Knee, Proctor & Galloway, 2008: 369; Dimov, 2016; 2018: 88

Ptilonyssus euroturdi mimicola Fain & Hyland, 1963a: 384 – Type: Kalamazoo Co., Michigan, from Dumetella carolinensis (syn. by Spicer, 1984: 800)

Type locality: Lauwe nr. Bruges, Belgium

Distribution: Austria, Belgium, Canada, Guatemala, Japan, Kirghizstan, Moldavia, New Zealand, Romania, Russia, Spain, Trinidad, Ukraine, USA

Hosts: Catharus dryas, Turdus e. ericetorum, T. fumigatus, T. iliacus, T. merula, T. migratorius, T. naumanni euromus, T. nudigenus, T. philomelus, T. pilaris, T. ruficollis, T. v. viscivorus (Passeriformes: Turdidae), Dumetella carolinensis (Passeriformes: Mimidae), Sturnus vulgaris (Passeriformes: Sturnidae)

Ptilonyssus faini (Strandtmann)

Ptilonyssoides faini Strandtmann, 1960: 142

Ptilonyssus faini: Domrow, 1968: 348

Type locality: Trus Madi Mt, Pampang, Ulu Kaingaran, North Borneo

Distribution: Malaysia (Borneo)

Host: Dicrurus hottentotus borneensis (Passeriformes: Dicruridae)

Ptilonyssus fluvicolae (Fain et Aitken)

Neotyranninyssus fluvicolae Fain et Aitken, 1967: 29; Hyland & Moorehouse, 1970: 53

Ptilonyssus fluvicolae (Fain et Aitken)

Type locality: Nariva Road, 3 mile post (Trinidad)

Distribution: Mexico, Trinidad

Hosts: Fluvicola pica, Pyrocephalus rubinus (Passeriformes: Tyrannidae)

Ptilonyssus formicarii Fain et Aitken

Ptilonyssus formicarii Fain & Aitken, 1970: 364

Type locality: Mosqueiro Ferry, Benavides, Belém

Distribution: Brazil

Host: Formicarius analis (Passeriformes: Formicariidae)

Ptilonyssus formicarii colmae Fain et Aitken

Ptilonyssus formicarii colmae Fain & Aitken, 1970: 368

Type locality: Mosqueiro Ferry, Benavides, Belém

Distribution: Brazil

Host: Formicarius colmae (Passeriformes: Formicariidae)

Ptilonyssus fringillae Fain et Sixl

Ptilonyssus fringillae Fain & Sixl, 1971: 90; Fain, Sixl & Moritsch, 1974: 4; Úbeda-Ontíveros & Guevara-Benítez, 1978

Type locality: Brunnsee, Styria; paratypes from Belgium

Distribution: Austria, Belgium, Spain

Hosts: Anthus pratensis (Passeriformes: Motacillidae); Regulus regulus (Passeriformes: Regulidae)

Ptilonyssus fringillicola Fain

Ptilonyssus fringillicola Fain, 1959a: 28; 1962b: 262; Butenko, 1965: 132; Shumilo & Lunkashu, 1970: 30; Fain, Sixl & Moritsch, 1974: 2; Fain, Herin & Pulyaert, 1977: 387

Ptilonyssus pyrrhulae Feider, 1962: 43 – Type: Romania, from Pyrrhula pyrrhula (syn. by Fain, Sixl & Moritsch, 1974: 2)

Type locality: Rugege Forest (Rwanda)

Distribution: Austria, Belgium, Cameroon, Kirghizstan, Romania, Russia, Rwanda, South Africa, Ukraine

Hosts: Carduelis c. carduelis, Pyrrhula pyrrhula, P. pyrrhula coccinea, Serinus atrogularis, S. mozambicus, S. [Poliospiza] striolata kivuensis (Passeriformes: Fringillidae), Emberiza flaviventris (Passeriformes: Emberizidae)

Ptilonyssus gerygonae Domrow

Ptilonyssus sp. Domrow, 1967a: 217

Ptilonyssus gerygonae Domrow, 1969b: 363; 1979d: 500; 1988: 871

Type locality: Queensland

Distribution: Australia

Hosts: Acanthiza reguloides; Gerygone palpebrosa (Passeriformes: Acanthizidae)

Ptilonyssus gilcolladoi Úbeda, Rodríguez, Guevara et Rojas

Ptilonyssus gilcolladoi Úbeda, Rodríguez, Guevara & Rojas, 1989: 157

Type locality: Prov. de Granada

Distribution: Spain

Host: Cettia cetti (Passeriformes: Cettiidae)

Ptilonyssus gliciphilae Domrow

Ptilonyssus gliciphilae Domrow, 1966a: 204; 1967a: 217; 1969b: 370; 1979d: 504; 1988: 872

Type locality: Chelona nr. Sarina, Esk

Distribution: Australia

Hosts: Gliciphila indistincta, Myzomela pectoralis (Passeriformes: Meliphagidae)

Ptilonyssus grallinae Domrow

Ptilonyssus grallinae Domrow, 1964d: 611, 618; 1969b: 366; 1979d: 500; 1988: 872; Fain & Lukoschus, 1979: 10

Type locality: Brisbane, Condamine (Qu.)

Distribution: Australia

Host: Grallina cyanoleuca (Passeriformes: Monarchidae)

Ptilonyssus gressitti (Baker et Delfinado)

Periglischrodes gressitti Baker et Delfinado, 1964: 589

Ptilonyssus gressitti (Baker et Delfinado): Bregetova, 1967: 135

Type locality: Bulolo

Distribution: Papua New Guinea

Host: «Chiroptera indet. » (according to Bregetova, 1967, it is a chance inclusion in a collection of spinturnicid mites, the real host should be a bird)

Ptilonyssus hirsti (de Castro et Pereira)

Neonyssus hirsti de Castro & Pereira, 1947: 129

Neonyssus (Paraneonyssus) hirsti: Černy & Dusbábek, 1970: 483

Ptilonyssus (Paraneonyssus) hirsti: Pereira & Castro, 1949: 227; Porter & Strandtmann, 1952: 393

Paraneonyssus hirsti Strandtmann & Wharton, 1958: 177; Feider, 1962: 58;1964: 268

Ptilonyssus hirsti: Zumpt & Till, 1955: 70; Fain, 1956a: 140; 1957b: 86; 1963: 170; Domrow, 1964d: 608, 1969b: 341; 1988: 872; Sixl, 1969: 218; Amaral, 1968: 119; Butenko, 1971; Fain, Sixl & Moritsch, 1974: 1; Pence, 1975: 24; Fain, Herin & Pulyaert, 1977: 387; Úbeda-Ontíveros & Guevara-Benítez, 1978; Shterbinina & Butenko, 1981; Knee, Proctor & Galloway, 2008: 370; Moodi et al., 2014: 3; Dimov, 2016; 2018: 85

Type locality: Portugal and North America

Distribution: Australia, Austria, Brazil, Cameroon, Canada, Cuba, Hawaii Is., Iran, New Guinea, Portugal, Romania, Rwanda, South Africa, Spain, Thailand, Turkmenistan, USA

Hosts: Parus major (Passeriformes: Paridae), Passer domesticus, P. griseus, P. indicus, P. montanus, P. ugandae griseus, Petronia dentata (Passeriformes: Passeridae), Fringilla coelebs (Passeriformes: Fringillidae)

Ptilonyssus hiyodori Kadosaka, Kaneko et Asanuma

Ptilonyssus hiyodori Kadosaka, Kaneko & Asanuma, 1987: 36

Type locality: Japan

Distribution: Japan

Host: Hypsipetes amaurotis amaurotis (Passeriformes: Pycnonotidae)

Ptilonyssus hoffmannae Luz-Zamudio

Ptilonyssus hoffmannae Luz-Zamudio, 1984: 113

Type locality: Mexico

Distribution: Mexico

Host: Carpodacus mexicanus centralis (Passeriformes: Fringillidae)

Ptilonyssus hoseini (Fain et Aitken)

Passeronyssus hoseini Fain & Aitken, 1967: 21

Passeronyssus havanensis (Fain et Aitken): Pence, 1972c: 793; Černy & Dusbábek, 1970: 486

Ptilonyssus hoseini (Fain et Aitken): Pence, 1972c: 793; 1973b: 876; Spicer, 1977b: 498; 1987: 261

Type locality: Saint Isidore-Plum Road, Biche (Trinidad)

Distribution: Cuba, Trinidad, USA

Hosts: Hylophylax poecilonota, Phlegopsis nigromaculata (Passeriformes: Thamnophilidae), Cyclarchis gujanensis, Vireo bellii, V. flavifrons, V. griseus, V. solitarius (Passeriformes: Vireonidae)

Ptilonyssus hylophylax Fain et Aitken

Ptilonyssus hylophylax Fain & Aitken, 1970: 383

Type locality: Pirelli Plantation, Marituba; Mosqueiro Ferry, Benevides

Distribution: Brazil

Hosts: Hylophylax poecilonota, Phlegopsis nigromaculata (Passeriformes: Formicariidae)

Ptilonyssus hypocnemoides Fain et Aitken

Ptilonyssus hypocnemoides Fain & Aitken, 1970: 389

Type locality: APEG Forest (Station A), Mucambo Forest, IPEAN, Belém

Distribution: Brazil

Host: Hypocnemoides maculicauda (Passeriformes: Formicariidae)

Ptilonyssus icteridius Strandtmann et Furman

Ptilonyssus icteridius Strandtmann & Furman, 1956: 167; Fain, 1957b: 96; Fain & Aitken, 1967: 5; Pence, 1972c: 794; 1975: 24; Cooper, Madden & Crites, 1975: 262; Spicer, 1977b: 498; 1978: 891, 1987: 261; Spicer, 1978: 891; Knee, Proctor & Galloway, 2008: 368; Moodi et al., 2014: 3; Knee & Galloway, 2017: 93

Paraneonyssus icteridius (Str. et Furman): Zemskaya & Ilienko, 1958: 476

Ptilonyssus (Paraneonyssus) icteridius (Str. et Furman): Wallace & Olsen, 1966

Neonyssus (Paraneonyssus) icteridius (Str. et Furman): Černý & Dusbábek, 1970: 483

Type locality: USA

Distribution: Canada, Iran, Russia, Trinidad, USA

Hosts: Agelaius phoeniceus, Euphagus carolinus, E. cyanocephalus, Icterus galbula, I. nigrogularis, Molothrus ater californicus, Quiscalus lugubris, Qu. quiscula, Sturnella magna, S. neglecta, Xanthocephalus xanthocephalus (Passeriformes: Icteridae), Passer domesticus, P. montanus (Passeriformes: Passeridae), Porzana carolina (Gruiformes: Rallidae), Piranga ludoviciana, Spiza americana (Passeriformes: Cardinalidae), Myiarchus crinitus (Passeriformes: Tyrannidae), Carpodacus mexicanus (Passeriformes: Fringillidae)

Ptilonyssus indicatoris Fain

Ptilonyssus indicatoris Fain, 1957b: 134

Type locality: Akanyaru

Distribution: Rwanda

Host: Indicator indicator (Piciformes: Indicatoridae)

Ptilonyssus inflatipalpus Fain et Aitken

Ptilonyssus inflatipalpus Fain & Aitken, 1970: 396

Type locality: Mosqueiro Ferry, Benevides

Distribution: Brazil

Host: Myrmornis torquata (Passeriformes: Formicariidae)

Ptilonyssus inornatus Fain et Aitken

Ptilonyssus inornatus Fain & Aitken, 1968: 14; 1970: 352

Type locality: APEG Forest

Distribution: Brazil

Hosts: Myrmotherula axillaris, M. longipennis, M. menetriesi (Passeriformes: Formicariidae)

Ptilonyssus insularis cubanus Černý

Ptilonyssus insularis cubanus Černý, 1969: 231

Ptilonyssus insularis Černý & Dusbábek, 1970: 482

Type locality: Aguacate, Mantingua, Baracoa, prov. of Oriente

Distribution: Cuba

Host: Dives atroviolaceus (Passeriformes: Icteridae)

Ptilonyssus insularis insularis Černý

Ptilonyssus insularis insularis Černý, 1969: 230

Ptilonyssus insularis Černý & Dusbábek, 1970: 482

Type locality: Cayo Piedra, Isla de Pinos

Distribution: Cuba (Pinos)

Host: Quiscalus niger caribaeus (Passeriformes: Icteridae)

Ptilonyssus intermedius (Hirst)

Neonyssus intermedius Hirst, 1922: 771; Vitzthum, 1935: 578; Castro, 1948: 268; Pereira & Castro, 1949: 228; Zumpt & Patterson, 1951: 77; Zumpt & Till, 1955: 62; Fain, 1957b: 52

Ptilonyssus intermedius (Hirst): Fain & Hyland, 1962: 341

Type locality: Madagascar

Distribution: Madagascar, Russia

Host: Aves («on a bird from Madagascar«)

Ptilonyssus isakovae Butenko et Lavrovskaya

Ptilonyssus isakovae Butenko & Lavrovskaya, 1983: 113

Type locality: Kirghizstan (Ottuk Vill., Issykkul Lake)

Distribution: Kirghizstan

Host: Phylloscopus inornatus (Passeriformes: Phylloscopidae)

Ptilonyssus japuibensis Castro

Ptilonyssus japuibensis Castro, 1948: 264; Pereira & Castro, 1949: 219; Zumpt & Till, 1955: 89; Fain, 1957b: 103; George, 1961: 127; Fain & Aitken, 1967: 3; Amaral, 1968: 120; Hyland & Moorhouse, 1970: 44; Fain & Aitken, 1970: 353; Pence, 1973b: 877; Knee, Proctor & Galloway, 2008: 368; Knee & Galloway, 2017: 93

Type locality: Fazenda Japuiba, Angra dos Reis (Estado do Rio de Janeiro)

Distribution: Brazil, Canada, Mexico, Trinidad, USA

Hosts: Rhamphocelus carbo, Rh. bresilius dorsalis, Sporophila americana, Tachyphonus rufus, T. surinamus, Volatinia jacarina (Passeriformes: Thraupidae), Pheucticus ludovicianus (Passeriformes: Cardinalidae), Junco hiemalis, Pipilo erythrophthalmus, Spizella palida, Zonotrichia albicollis (Passeriformes: Passerellidae), Molothrus ater, Quiscalus quiscala (Passeriformes: Icteridae), Empidonax alnorum (Passeriformes: Tyrannidae)

Ptilonyssus japuibensis cyanocompsae Fain et Aitken

Ptilonyssus japuibensis cyanocompsae Fain & Aitken, 1970: 354

Type locality: Mosqueiro Ferry, Benavides, Belém

Distribution: Brazil

Host: Cyanocompsa cyanoides (Passeriformes: Thraupidae)

Ptilonyssus langei Butenko

Ptilonyssus langei Butenko, 1972: 1074

Type locality: Azerbaidjan, Kazakhstan, Russia

Distribution: Azerbaidjan, Kazakhstan, Russia

Host: Sturnus vulgaris (Passeriformes: Sturnidae)

Ptilonyssus lanii Zumpt et Till

Ptilonyssus lanii Zumpt & Till, 1955: 71; Fain, 1956: 143; 1957c: 217; George, 1961: 119; Shumilo & Lunkashu, 1970: 30; Domrow & Nadchatram, 1978: 87; Sartbaev, 1981: 131

Ptilonyssus angustatus Feider & Mironescu, 1969: 21 – Type from Romania, on Lanius collurio.

Type locality: Transvaal

Distribution: Kirghizstan, Malaysia, Moldavia, Romania, Russia, South Africa, Ukraine, USA

Hosts: Lanius collaris, L. collurio, L. cristatus, L. ludovicianus, L. minor (Passeriformes: Laniidae); Sturnus vulgaris (Passeriformes: Sturnidae), Coracias garrulus (Coraciiformes: Coraciidae)

Ptilonyssus lobatus Strandtmann

Ptilonyssus lobatus Strandtmann, 1960: 147; Fain & Nadchatram, 1962: 279

Type locality: Khao Yoi, Phet Buri (Thailand)

Distribution: Malaysia, Thailand

Hosts: Sturnia sturnina, Sturnus javanicus grandis, S. tristis tristis (Passeriformes: Sturnidae)

Ptilonyssus lovottiae Dimov et Mironov

Ptilonyssus lovottiae Dimov & Mironov, 2012: 171

Type locality: Konzj, Leningrad Province

Distribution: Russia

Host: Passer montanus (Passeriformes: Passeridae)

Ptilonyssus ludovicianus Černý

Ptilonyssus ludovicianus Černý, 1969: 228; Černý & Dusbábek, 1970: 482; Knee & Galloway, 2017: 93

Type locality: Soledad, prov. of Las Villas, Botanical Garden

Distribution: Canada, Cuba

Hosts: Pheucticus ludovicianus, Passerina cyanea (Passeriformes: Cardinalidae)

Ptilonyssus lusciniae (Fain)

Passeronyssus lusciniae Fain, 1962: 140; 1963a: 174; Úbeda, Rodríguez & Guevara, 2000

Ptilonyssus lusciniae (Fain): Shumilo & Lunkashu, 1970: 31

Type locality: Belgium

Distribution: Belgium, Malaysia, Moldavia, Rwanda, Spain

Hosts: Luscinia c. cyanae, L. luscinia, Myrmecocychla nigra, Phoenicurus phoenicurus (Passeriformes: Muscicapidae)

Ptilonyssus lymozemae Domrow

Ptilonyssus lymozemae Domrow, 1965b: 453; 1969b: 371; 1979d: 504; 1988: 872

Type locality: Samford, Logan Village

Distribution: Australia

Host: Myzomela sanguinolenta (Passeriformes: Meliphagidae)

Ptilonyssus macclurei Fain

Ptilonyssus macclurei Fain, 1963c: 72; Domrow, 1964d: 609, 617; 1967: 217; 1969b: 366; 1988: 872

Type locality: Mitchell River Mission, Queensland (???)

Distribution: Australia, Malaysia ?

Host: Rhipidura leucophrys (Passeriformes: Muscicapidae)

Ptilonyssus madagascariensis Grétillat, Capron et Brygoo

Ptilonyssus madagascariensis Grétillat, Capron & Brygoo, 1959: 381

Type locality: Tananarive

Distribution: Madagascar

Host: Hypsipetes [Ixocincla] madagascariensis (Passeriformes: Pycnonotidae)

Ptilonyssus malacoptilae (Fain et Aitken)(new comb.)

Passeronyssus malacoptilae Fain & Aitken, 1971: 40

Type locality: Ananindeua

Distribution: Brazil

Host: Malacoptila rufa (Piciformes: Bucconidae)

Ptilonyssus malaysiae Fain

Ptilonyssus malaysiae Fain, 1964c: 65

Type locality: Bukit Lagong, Selangor

Distribution: Malaysia

Host: Irena puella malayensis (Passeriformes: Irenidae)

Ptilonyssus maluri Domrow

Ptilonyssus maluri Domrow, 1965b: 451; 1969b: 339; 1979d: 493; 1988: 872

Type locality: Esk

Distribution: Australia

Host: Malurus melanocephalus (Passeriformes: Maluridae)

Ptilonyssus mariacastroae Fain

Ptilonyssus mariacastroae Fain, 1961b: 48; Fain & Aitkin, 1967: 6; Spicer, 1977b: 498

Type locality: Anvers Zoo (Brazil)

Distribution: Brazil, Mexico, Trinidad

Hosts: Amazilia chionopectus, A. yucatanensis, Chrysolampis mosquitus, Saucerottia tobaci (Apodiformes: Trochilidae)

Ptilonyssus melanocoryphae (Bregetova)

Neonyssus (Otocorinyssus) melanocoryphae Bregetova, 1967: 127

Astridiella melanocoryphae (Bregetova): Úbeda & Morillas Márquez, 1979: 128

Type locality: at Zeravshan River, 15 km upstream Penjikent

Distribution: Tadjikistan

Host: Melanocorypha bimaculata (Passeriformes: Alaudidae)

Ptilonyssus meliphagae Domrow

Ptilonyssus meliphagae Domrow, 1964d: 606; 1967a: 217; 1969b: 372; 1979: 508; 1988: 873

Type locality: Mt. Cotton, Queensland

Distribution: Australia

Hosts: Meliphaga chrysops; Melithreptus albogularis (Passeriformes: Meliphagidae)

Ptilonyssus melissae Spicer

Ptilonyssus melissae Spicer, 1977a: 597; 1978: 891; 1987: 261

Type locality: Diamond-A Ranch, Hood Co., Fort Worth, Tarrant Co., Texas

Distribution: USA (Texas, Montana, Alasca)

Hosts: Carpodacus mexicanus, C. purpureus, Leucosticte tephrocotis, Pinicola enucleator (Passeriformes: Fringillidae)

Ptilonyssus melittophagi Fain

Ptilonyssus melittophagi Fain, 1956a: 136; 1959a: 26

Type locality: Astrida (= Butare), Rwanda

Distribution: Rwanda, South Africa

Hosts: Melittophagus lafresnayi oreobates, M. pusillus (Passeriformes: Meropidae)

Ptilonyssus microecae Domrow

Ptilonyssus microecae Domrow, 1966a: 196; 1967: 217; 69b: 356; 1988: 873

Type locality: Esk

Distribution: Australia

Hosts: Microeca fascinans; M. flavigaster (Passeriformes: Muscicapidae)

Ptilonyssus mimi George

Ptilonyssus mimi George, 1961: 120

Type locality: Kingsville, Texas

Distribution: USA (Texas)

Host: Mimus polyglottos (Passeriformes: Mimidae)

Ptilonyssus mironovi Dimov

Ptilonyssus mironovi Dimov, 2012: 27

Type locality: Lomonosov, Leningrad Province

Distribution: Russia

Host: Parus caeruleus (Passeriformes: Paridae)

Ptilonyssus missimensis Sakakibara

Ptilonyssus missimensis Sakakibara, 1968c: 429

Type locality: Mt. Missim, Morobe District

Distribution: Papua New Guinea

Host: “Honey eater” (Passeriformes: Meliphagidae)

Ptilonyssus monarchae Domrow

Ptilonyssus monarchae Domrow, 1969b: 360; 1988: 873

Type locality: Innisfail

Distribution: Australia

Host: Monarcha trivirgata (Passeriformes: Monarchidae)

Ptilonyssus montifringillae Butenko et Lavrovskaya

Ptilonyssus montifringillae Butenko & Lavrovskaya, 1983: 118

Type locality: Jaz, Issykkul Region

Distribution: Kirghizstan

Host: Montifringilla nivalis (Passeriformes: Passeridae)

Ptilonyssus morofskyi Hyland

Ptilonyssus morofskyi Hyland, 1963: 146; Fain, 1964a: 61; Pence, 1972; 1975: 25; Spicer, 1977a: 600; 1978: 892; Guevara & Úbeda, 1980: 333; Knee, Proctor & Galloway, 2008: 368; Knee & Galloway, 2017: 93

Neonyssus (Spizonyssus) morofskyi (Hyland): Bregetova, 1967: 131; Černý & Dusbábek, 1970: 482

Neonyssus morofskyi (Hyland): Shumilo & Lunkashu, 1970: 22

Type locality: Kalamazoo Co., Michigan

Distribution: Belgium, Canada, Cuba, Moldavia, Spain, USA

Hosts: Carduelis [Spinus] tristis, C. [Acanthis] cannabina, C. flammea, Loxia curvirostra, Serinus serinus (Passeriformes: Fringillidae), Calcarius ornatus, Plectrophenax nivalis (Passeriformes: Calcariidae), Aimophila ruficeps, Ammodramus lecontei, Ammospiza maritima, Calamospiza melanocorys, Junco caniceps, J. hyemalis, Melospiza georgiana, M. lincolni, M. melodia, Passerella iliaca, Passerculus sandwichensis, Pipilo erythrophthalmus, Pooecetes gramineus, Spizella pusilla, Zonotrichia albicollis, Z. leucophrys, Z. querula (Passeriformes: Passerellidae), Dendroica coronata, D. pinus, Mniotilta varia, Geothlypis trichas, Seiurus motacilla, Wilsonia pusilla (Passeriformes: Parulidae)

Ptilonyssus motacillae Fain

Ptilonyssus motacillae Fain, 1956a: 143; 1957b: 137; 1959a: 24; 1960: 109; 1962a: 128; Strandtmann & Wharton, 1958: 181; Domrow, 1964d: 600; 1965b: 460; 1967: 217; 1969b: 352; 1979: 499; 1988: 873; Shumilo & Lunkashu, 1970: 31; Fain, Sixl & Moritsch, 1974: 2; Pence, 1975: 25; Sartbaev, 1983: 84; Rodriguez, Úbeda & Guevara, 1993: 13 ; Butenko, Stanyukovich & Dimov, 2010: 15; Dimov, 2018: 87

Hapalognatha prima Butenko, 1959: 17; 1960: 1490

Ptilonyssus motacilla: Butenko, 1965

Ptilonyssus estrildicola taeniopygiae Fain, 1963b: 74; Fain, Herin & Puylaert, 1977: 386

Ptilonyssus estrildicola Fain, 1959: 22; Domrow, 1965b: 453

Ptilonyssus fringillicola Fain, 1959: 28

Ptilonyssus cinnyricincti Fain, 1962: 132

Type locality: Astrida (= Butare), Rwanda

Distribution: Australia, Austria, Botswana, Kirghizstan, Moldavia, Russia, Rwanda, South Africa, Spain, Ukraine, USA

Hosts: Anthus australis, A. spinoletta, A. trivialis, Ficedula albicollis, Motacilla aguimp vidua, M. alba, M alba baicalensis, M. capensis wellsi, M. cinerea, M. citreola, M. flava, M. tschutschensis macronyx (Passeriformes: Motacillidae), Lonchura punctulata, L. [Eodice] malabarica, Taeniopygia castanotis (Passeriformes: Estrildidae), Pachycephala pectoralis, P. rufiventris (Passeriformes: Pachycephalidae), Petroica rosea (Passeriformes: Petroicidae), Cercomela familiaris modesta, Erithacus rubecula, Erythropygia coryphaeus, Luscinia svecica, Muscicapa striata, Myrmecocichla formicivora, Oenanthe isabellina, O. oenanthe, Phoenicurus erythronotus, Ph. motacillae (Passeriformes: Muscicapidae), Cyanosylvia svecica, Saxicola rubetra, S. torquata axillaris (Passeriformes: Turdidae), Mirafra africanoides (Passeriformes: Alaudidae), Acridotheres tristis (Passeriformes: Sturnidae), Panurus biarmicus (Passeriformes: Panuridae), Leucosticte nemoricola, Serinus pusillus (Passeriformes: Fringillidae), Scotocerca inquieta (Passeriformes: Cisticolidae), Perdix daurica (Galliformes: Phasianidae)

Ptilonyssus motacillae phoenicuri Fain

Ptilonyssus motacillae phoenicuri Fain, 1966: 90; Shumilo & Lunkashu, 1970: 32

Type locality: Strée (Hainaut), Belgium

Distribution: Belgium, Ukraine

Hosts: Phoenicurus ochruros, Ph. o. gibraltariensis (Passeriformes: Muscicapidae)

Ptilonyssus muscicapae Bregetova

Ptilonyssus muscicapae Bregetova, 1970: 59; Shumilo & Lunkashu, 1970: 32

Type locality: Ribachiy, Kaliningradskaya Obl. (Russia)

Distribution: Moldavia, Russia, Turkmenistan

Hosts: Muscicapa latirostris, M. sibirica, M. striata (Passeriformes: Muscicapidae)

Ptilonyssus muscicapoides Butenko et Lavrovskaya

Ptilonyssus muscicapoides Butenko & Lavrovskaya, 1980b: 351

Type locality: Russia

Distribution: Russia

Host: Ficedula hypoleuca (Passeriformes: Muscicapidae)

Ptilonyssus myiobii Fain et Aitken

Ptilonyssus myiobii Fain & Aitken, 1970: 356

Type locality: Mosqueiro Ferry, Benavides, Belém

Distribution: Brazil

Host: Myiobius barbatus (Passeriformes: Tyrannidae)

Ptilonyssus myrmotherulae Fain et Aitken

Ptilonyssus myrmotherulae Fain & Aitken, 1968: 16; 1970: 356

Type locality: APEG Forest, Belém

Distribution: Brazil

Host: Myrmotherula hauxwelli (Passeriformes: Formicariidae)

Ptilonyssus myzanthae Domrow

Ptilonyssus myzanthae Domrow, 1964d: 603, 617; 1969b: 367; 1979: 500; 1988: 873

Type locality: Condamine, Qu.

Distribution: Australia

Hosts: Anthochaera chrysoptera, Myzantha flavigula, M. melanocephala (Passeriformes: Meliphagidae)

Ptilonyssus myzomelae Domrow

Ptilonyssus myzomelae Domrow, 1965b: 455; 1969b: 370; 1979d: 502; 1988: 874

Type locality: Samford

Distribution: Australia

Hosts: Myzomela obscura, M. sanguinolenta (Passeriformes: Meliphagidae)

Ptilonyssus nadchatrami Fain

Ptilonyssus nadchatrami Fain, 1964c: 63

Type locality: Bukit Lagong, Selangor

Distribution: Malaysia

Hosts: Pycnonotus brunneus, P. e. erythrophthalmus (Passeriformes: Pycnonotidae)

Ptilonyssus neochmiae Domrow

Ptilonyssus neochmiae Domrow, 1969b: 341, 423; 1988: 874; 1992: 1605

Type locality: Innisfail

Distribution: Australia

Hosts: Neochmia albiventer, N. phaeton (Passeriformes: Ploceidae), Erythrura gouldiae, Poephila acuticauda (Passeriformes: Estrildidae)

Ptilonyssus nicatoris Fain

Ptilonyssus nicatoris Fain, 1958b: 294

Type locality: Kivu

Distribution: DR Congo

Host: Nicator chloris (Passeriformes: Pycnonotidae)

Ptilonyssus nitzschi (Giebel)(nomen dubium, see Fain, 1957d)

Dermanyssus nitzschi Giebel, 1871: 32

Neonyssus (Vitznyssus) nitzschi: Castro, 1948: 277

Ptilonyssus nitzschi: Zumpt & Till, 1955: 69; Spicer, 1978: 892

Type locality: ? Europe

Distribution: Australia, [Europe], USA

Hosts: Caprimulgus carolinensis, C. macrurus, Phalaenoptilus nuttallii (Caprimulgiformes: Caprimulgidae)

Ptilonyssus nivalis Knee

Ptilonyssus nivalis Knee, 2008: 359

Type locality: Oak Hammock Marsh, Manitoba

Distribution: Canada

Host: Plectrophenax nivalis (Passeriformes: Fringillidae)

Ptilonyssus novaeguineae (Hirst)

Rhinonyssus novae-guineae Hirst, 1921: 769

Passeronyssus novae-guineae (Hirst): Fain, 1960b: 314

Ptilonyssus novae-guineae (Hirst): Fain & Hyland, 1962: 346

Ptilonyssus novaeguineae (Hirst): Sakakibara, 1968; Domrow, 1969: 374; Feider & Mironescu, 1980: 61

Type locality: Papua New Guinea

Distribution: Papua New Guinea

Hosts: Paradisaea reggiana, Ptiloris magnificus, P. paradiseus, P. victoriae (Passeriformes: Paradisaeidae)

Ptilonyssus nucifragae (Hirst)

Rhinonyssus (Neonyssoides) nucifragae Hirst, 1923: 975; Bregetova, 1965: 133

Neonyssus nucifragae: Bregetova, 1953: 332; Fain, 1957: 52; Isakova, 1965: 89

Ptilonyssus nucifragae (Hirst): Fain, 1960b: 311

Neonyssoides nucifragae (Hirst): Bregetova, 1967: 133; Shumilo & Lunkashu, 1970:

Type locality: Heilsberg

Distribution: [East Prussia – formerly in Germany], Kirghizstan, Latvia, Russia, Ukraine

Hosts: Garrulus glandarius, Nucifraga caryocatactes, Perisoreus infaustus (Passeriformes: Corvidae)

Ptilonyssus nudus Berlese et Trouessart

Ptilonyssus nudus Berlese & Trouessart, 1889: 126; Berlese, 1892: 124; Oudemans, 1905: 237; Hirst, 1916: 73; Vitzthum, 1935: 578; Nordberg, 1936: 49; Castro & Pereira, 1947: 127; Bregetova, 1951: 118; Porter & Strandtmann, 1952: 393; Turk, 1953: 13; Zumpt & Till, 1955: 79; Fain, 1956a: 144; 1957b: 100; 1963: 171; George, 1961: 116; Butenko, 1965; Isakova, 1965: 82; 1968; Domrow, 1969: 372; Shumilo & Lunkashu, 1970: 33; Pence, 1975: 109; Fain, Herin & Pulyaert, 1977: 387; Knee, Proctor & Galloway, 2008: 369; Huhta, 2016: 148 (Sp. incert.), Dimov, 2018: 93

Type locality: Europe (France, Italy)

Distribution: [Australia], Cameroon, Canada, England, ?Finland, France, Germany, Holland, Italy, Moldavia, Portugal, Russia, Rwanda, USA

Hosts: Passer domesticus, P. griseus, P. montanus (Passeriformes: Passeridae); also there are published findings on Hypochera funerea nigerrima, Vidua macroura (Passeriformes: Viduidae)(Fain, 1956) and on Emberiza cirlus (Pass.: Emberizidae) (Berlese & Trouessart, 1889), and Fringilla coelebs (from Vitzthum, 1935), but Fain (1957b) thinks, that this information is doubtful. Also findings on Parus atricapilus, P. major (Passeriformes: Paridae), Sturnus vulgaris (Passeriformes: Sturnidae), Hirundo rustica, Riparia riparia (Passeriformes: Hirundinidae), Carpodacus roseus (Passeriformes: Fringillidae)

Ptilonyssus ohioensis Fain et Johnston

Ptilonyssus ohioensis Fain & Johnston, 1966: 29

Type locality: Ohio

Distribution: USA (Ohio)

Host: Cathartes aura (Cathartiformes: Cathartidae)

Ptilonyssus olaoi Pereira et Castro

Ptilonyssus (Ptilonyssus) olaoi Pereira & Castro, 1949: 219

Ptilonyssus olaoi Pereira et Castro: Zumpt & Till, 1955: 88; Fain, 1957: 104; Strandtmann, 1960: 146; Amaral, 1968: 121

Type locality: Boracéa (Estado de Sao Paulo)

Distribution: Brazil

Host: Ampelion cucullatus (Passeriformes: Cotingidae)

Ptilonyssus orientalis (Ewing)

Sommatericola orientalis Ewing, 1933: 13

Ptilonyssus orientalis (Ewing): Standtmann, 1956: 136

Type locality: Bangkok

Distribution: Thailand

Host: Lanius nigriceps longicaudatus (Passeriformes: Laniidae)

Ptilonyssus orioli Fain

Ptilonyssus oriolus Fain, 1956a: 144; 1962b: 269; Butenko, 1969; Shumilo & Lunkashu, 1970: 33

Type locality: Akanyaru (Rwanda)

Distribution: Belgium, Moldavia, Rwanda, Ukraine

Hosts: Oriolus larvatus angolensis, O. o. oriolus (Passeriformes: Oriolidae)

Ptilonyssus orthonychus Domrow

Ptilonyssus orthonychus Domrow, 1969b: 354; 1988: 874

Type locality: Wilson’s Peak, Queensland

Distribution: Australia

Host: Orthonyx temminckii (Passeriformes: Orthonychidae)

Ptilonyssus paradisaeus (Sakakibara)

Ptilonyssoides paradisaeus Sakakibara, 1968: 298

Ptilonyssus paradisaeus (Sakakibara)

Type locality: Morobe District, Wau, Nakata DRidge, 1515 m

Distribution: Papua New Guinea

Host: Paradisornis rudolphi (Passeriformes: Paradisaeidae)

Ptilonyssus paranensis (Castro)

Neonyssus (Travanyssus) paranensis Castro, 1948: 276; Pereira & Castro, 1949; Zumpt & Till, 1955: 90

Ptilonyssus (Travanyssus) paranensis (Castro): Pereira & Castro, 1949: 227

Ptilonyssus paranensis: Zumpt & Till, 1955: 90; Fain, 1957b: 100; Amaral, 1968: 120

Type locality: Estado de Parana

Distribution: Brazil

Host: Elaenia flavogaster (Passeriformes: Tyrannidae)

Ptilonyssus pari Fain et Hyland

Ptilonyssus pari Fain & Hyland, 1963a: 375; Fain & Bafort, 1963b: 485; Butenko, 1965: 131; Sixl, 1969: 218; Sixl & Reich, 1971: 401; Butenko & Stolbov, 1971; Pence, 1972c: 793; 1975: 90; Fain, Sixl & Moritsch, 1974: 2; Spicer, 1978: 892; Kadosaka et al., 1983; Rodriguez, Úbeda, Guevara & Ariza, 1989: 152; Dimov, 2016; 2018: 81

Neonyssus pari (Fain et Hyland): Shumilo & Lunkashu, 1970: 23

Type locality: Shiawansee, Michigan (USA), Fosse, etc. (Belgium)

Distribution: Austria, Belgium, Japan, Moldavia, Russia, Spain, Ukraine, USA (La., Nevada)

Hosts: Parus ater, P. atricapillus, P. atricapillus subrhenanus, P. caeruleus, P. carolinensis, P. cristatus mitratus, P. gambeli, P. insularis, P. m. major, P. montanus (Passeriformes: Paridae), Sitta europaea (Sittidae)

Ptilonyssus pastoris Butenko

Ptilonyssus pastoris Butenko, 1972: 1075

Type locality: Mihaylovka Vill., Dzhambulskiy Rayon, Kazakhstan

Distribution: Kazakhstan

Host: Pastor roseus (Passeriformes: Sturnidae)

Ptilonyssus pericrocoti (Sakakibara)

Paraneonyssus pericrocoti Sakakibara, 1967: 597

Ptilonyssus pericrocoti (Sakakibara)

Type locality: Liukuei

Distribution: Taiwan

Host: Pericrocotus roseus divaricatus (Passeriformes: Campephagidae)

Ptilonyssus perisorei George

Ptilonyssus perisorei George, 1961: 108; Spicer, 1978: 892; Wilson & Haas, 1980: 545

Type locality: Santa Fe, New Mexico

Distribution: Canada, USA (Alaska, New Mexico, Utah)

Hosts: Aphelocoma coerulescens, Cyanocitta stelleri, Perisoreus canadensis (Passeriformes: Corvidae)

Ptilonyssus phainopeplae George

Ptilonyssus phainopeplae George, 1961: 108; Pence, 1975: 25

Type locality: Val Verde Co., Texas

Distribution: USA (Texas)

Host: Phainopepla nitens (Passeriformes: Ptilogonatidae)

Ptilonyssus philemoni Domrow

Ptilonyssus philemoni Domrow, 1964d: 600, 617; 1965b: 460; 1967: 218; 1969b: 367; 1979: 502; 1988: 874

Ptilonyssus pentagonicus Fain & Lukoschus, 1979: 18; 1986: 60. Type loc.: Mount Hart (Western Australia), ex Manorina flavigula (syn. by Domrow, 1987)

Type locality: Logan Village, Condamine, Qu.

Distribution: Australia

Hosts: Entomyzon cyanotis; Manorina flavigula, Meliphaga gracilis, M. notata, Myzantha flavigula, Philemon citreogularis, P. corniculatis, Plectorhyncha lanceolata (Passeriformes: Meliphagidae)

Ptilonyssus phyllastrephi Zumpt et Till

Ptilonyssus phyllastrephi Zumpt & Till, 1955: 71; Fain, 1957b: 104

Type locality: Cape Prov.

Distribution: South Africa

Host: Phyllastrephus terrestris (Passeriformes: Pycnonotidae)

Ptilonyssus phylloscopi Fain

Ptilonyssus phylloscopi Fain, 1962b: 257; Isakova, 1965; Guevara-Benítez & Úbeda Ontíveros, 1980a: 231

Type locality: Anvers (Belgium)

Distribution: Belgium, Kirghizstan, Russia, Spain

Hosts: Phylloscopus collybita, Ph. griseolus, Ph. inornatus, Ph. t. trochilus (Passeriformes: Phylloscopidae)

Ptilonyssus pinicola Knee

Ptilonyssus pinicola Knee, 2008: 359

Type locality: Coyote Lake Nature Sanctuary, Alberta

Distribution: Canada

Host: Pinicola enucleator (Passeriformes: Fringillidae)

Ptilonyssus pipromorphae Fain et Aitken

Ptilonyssus pipromorphae Fain & Aitken, 1967: 13; 1968: 3; 1970: 355

Type locality: Morne Poui Road, Aripo Valley

Distribution: Brazil, Trinidad

Hosts: Pipromorpha macconnelli, P. oleaginea (Passeriformes: Tyrannidae)

Ptilonyssus pirangae (Černý)

Neonyssus (Paraneonyssus) pirangae Černý, 1969: 232; Černý & Dusbábek, 1970: 484

Ptilonyssus pirangae: Pence, 1972c: 793; 1973: 887; Knee, Proctor & Galloway, 2008: 368; Moodi, 2014: 3

Type locality: Havana, Botanical Garden

Distribution: Canada, Cuba, Iran

Hosts: Piranga ludoviciana, P. rubra rubra (Passeriformes: Thraupidae), Parus bicolor (Passeriformes: Paridae)

Ptilonyssus pittae Domrow

Ptilonyssus pittae Domrow, 1964a: 26; 1964d: 618; 1969d: 349; 1979: 495; 1988: 874

Type locality: Upper Brookfield, S.E. Queensland

Distribution: Australia

Host: Pitta versicolor (Passeriformes: Pittidae)

Ptilonyssus platypsaris Fain et Aitken

Ptilonyssus platypsaris Fain & Aitken, 1971: 34

Type locality: APEG Forest, Station A (IPEAN)

Distribution: Brazil

Host: Platypsaris minor (Passeriformes: Cotingidae)

Ptilonyssus platyrhinchus Fain et Aitken

Ptilonyssus platyrhinchus Fain & Aitken, 1970: 374

Type locality: Mosqueiro Ferry, Benavides, Belém

Distribution: Brazil

Host: Platyrhinchus saturatus (Passeriformes: Tyrannidae)

Ptilonyssus plesiotypicus Knee

Ptilonyssus plesiotypicus Knee, 2008: 367

Type locality: Alberta

Distribution: Canada

Host: Carpodacus purpureus (Passeriformes: Fringillidae)

Ptilonyssus ploceanus Fain

Ptilonyssus ploceanus Fain, 1956a: 139; 1957b: 89; Fain, Herin & Puylaert, 1977: 388; Lindholm, Venter & Ueckermann, 1998: 148

Type locality: Akanyaru (Rwanda)

Distribution: Cameroon, Rwanda, South Africa

Hosts: Anaplectes rubriceps leuconotus, Euplectes ardeus, E. hordeaceus, E. macrourus, E. orix, E. o. nigrifrons, Hyphanturgus ocularius, Ploceus cucullatus, P. vitellinus, Quelea erythrops, Textor atrogularis (Passeriformes: Ploceidae), Petronia dentata (Passeriformes: Passeridae), Serinus mozambicus (Passeriformes: Fringillidae); Chrysococcyx caprius (Cuculiformes: Cuculidae)

Ptilonyssus prunellae Fain et Bafort

Ptilonyssus prunellae Fain & Bafort, 1963b: 480; 1966: 7; Beron, 1975: 186; Guevara-Benítez & Úbeda Ontíveros, 1978a: 259; Sartbaev, 1983: 84

Type locality: Anvers, etc. (Belgium)

Distribution: Belgium, Bulgaria, Kirghizstan, Spain

Hosts: Prunella fulvescens, P. m. modularis (Passeriformes: Prunellidae)

Ptilonyssus psalidoprocnei Fain

Ptilonyssus psalidoprocnei Fain, 1956a: 146

Type locality: Astrida (= Butare)

Distribution: Rwanda

Host: Psalidoprocne albiceps (Passeriformes: Hirundinidae)

Ptilonyssus psaltriparus Spicer

Ptilonyssus psaltriparus Spicer, 1978: 892

Type locality: San Pablo Reservoir, Contra Costa Co., California

Distribution: USA (California)

Host: Psaltriparus minimus (Passeriformes: Paridae)

Ptilonyssus pseudothymanzae Fain et Lukoschus

Ptilonyssus pseudothymanzae Fain & Lukoschus, 1979: 15; Koch, 1986: 60; Domrow, 1988: 874

Type locality: Western Australia (The Kimberley)

Distribution: Australia (WA)

Hosts: Meliphaga flavescens, Melithreptus albogularis (Passeriformes: Meliphagidae)

Ptilonyssus psophodae Domrow

Ptilonyssus psophodae Domrow, 1964a: 29; 1964d: 618; 1969b: 366; 1988: 874

Type locality: Upper Brookfield, S.E. Queensland

Distribution: Australia

Host: Psophodes olivaceus (Passeriformes: Cinclosomatidae)

Ptilonyssus ptyonoprognes Butenko et Lavrovskaya

Ptilonyssus ptyonoprognes Butenko & Lavrovskaya, 1983: 116

Type locality: Kirghizstan

Distribution: Kirghizstan

Host: Ptyonoprogne rupestris (Passeriformes: Hirundinidae)

Ptilonyssus pycnonoti Fain

Ptilonyssus pycnonoti Fain, 1956a: 146; 1957b: 109; 1959a: 24; 1960: 109

Type locality: Astrida (= Butare)

Distribution: Rwanda, South Africa

Hosts: Pycnonotus barbarus tricolor, P. nigricans (Passeriformes: Pycnonotidae)

Ptilonyssus pycnonoti malayi Fain

Ptilonyssus pycnonoti malayi Fain, 1963: 78

Type locality: Bukit Mandol, Selangor

Distribution: Malaysia

Host: Pycnonotus plumosus (Passeriformes: Pycnonotidae)

Ptilonyssus pygiptilae Fain et Aiken

Ptilonyssus pygiptilae Fain & Aiken, 1971: 37

Type locality: Ananindeua

Distribution: Brazil

Host: Pygiptila stellaris (Passeriformes: Thamnophilidae)

Ptilonyssus pygmaeus (Bregetova)

Neonyssus (Spizonyssus) pygmaeus Bregetova, 1967: 131

Neonyssus pygmaeus Bregetova: Shumilo & Lunkashu, 1970: 22

Ptilonyssus pygmaeus (Bregetova): Domrow, 1969: 341

Type locality: Kaliningradskaya Onl. (Russia), 10 km S Penjikent

Distribution: Moldavia, Russia, Tadjikistan

Hosts: Carduelis carduelis, C. carduelis subcaniceps, Spinus spinus (Passeriformes: Fringillidae)

Ptilonyssus pyriglenae Fain et Aitken

Ptilonyssus pyriglenae Fain & Aitken, 1968: 18; 1970: 355

Type locality: Mucambo Forest

Distribution: Brazil

Host: Pyriglena leucoptera (Passeriformes: Formicariidae)

Ptilonyssus pyrrhulinus Stanyukovich et Butenko

Ptilonyssus pyrrhulinus Stanyukovich & Butenko, 2003: 38

Type locality: Ryazanskaya Oblast

Distribution: Russia

Host: Pyrrhula pyrrhula (Passeriformes: Fringillidae)

Ptilonyssus rabelloi (Castro)

Ptilonyssus (Flavionyssus) rabelloi Castro, 1948: 266; Pereira & Castro, 1949: 226

Ptilonyssus rabelloi (Castro): Zumpt & Till, 1955: 88; Fain, 1957: 109; Amaral, 1968: 120

Type locality: Juquiá (Estado de São Paulo)

Distribution: Brazil

Host: Sittasomus griseicapillus sylviellus (Passeriformes: Furnariidae, Dendrocolaptinae)

Ptilonyssus radovskyi Feider et Mironescu

Ptilonyssus radovskyi Feider & Mironescu, 1980: 55

Type locality: Wan Creek, Tambula

Distribution: Papua New Guinea

Hosts: Peneothello cyanus, Tregellasia leucops (Passeriformes: Petroicidae), Amblyornis macgregori (Passeriformes: Ptilonorhynchidae)

Ptilonyssus reguli Fain et Sixl

Ptilonyssus reguli Fain & Sixl, 1969: 264; Fain, Sixl & Moritsch, 1974: 4; Rodriguez, Úbeda & Guevara, 1990: 301

Type locality: Autal Graz (Austria)

Distribution: Austria, Spain

Hosts: Regulus ignicapillus, R. regulus (Passeriformes: Regulidae)

Ptilonyssus richmondenae George

Ptilonyssus richmondenae George, 1961: 122; Pence, 1975: 25; Hilario-Perez & Dowling, 2018: 296

Type locality: Liberty Hill, Texas

Distribution: USA (Texas)

Hosts: Richmondena cardinalis (Passeriformes: Fringillidae), Molothrus ater (Passeriformes: Icteridae)

Ptilonyssus rhipidurae Domrow

Ptilonyssus rhipidurae Domrow, 1966a: 199; 1969b: 366; 1988: 875

Type locality: Esk

Distribution: Australia

Host: Rhipidura fuliginosa (Passeriformes: Muscicapidae)

Ptilonyssus ripariae Stanyukovich et Butenko

Ptilonyssus ripariae Stanyukovich & Butenko, 2003: 35

Type locality: Pologrudino Village, Tyumenskaya Oblast

Distribution: Russia

Host: Riparia riparia (Passeriformes: Hirundinidae)

Ptilonyssus ruandae Fain

Ptilonyssus ruandae Fain, 1956b: 395; 1957b: 109; 1960: 110; Strandtmann & Wharton, 1958: 182; Domrow, 1964d: 608; 1969b: 366; 1988: 875; Kadosaka, Kaneko & Asanuma, 1987: 33

Ptilonyssus ruandae alcippei Fain & Nadchatram, 1962: 279; Nadchatram, McClure

& Chong, 1964: 104 – Type: Malaysia, Mont Brinchang (1700 m), Pahang, from Alcippe nipalensis peracensis; McClure, Ratanaworadhan et al., 1973: 35

Type locality of Ptilonyssus ruandae Fain: Akanyaru (Rwanda)

Distribution: Australia, Botswana, Japan, Malaysia, Rwanda

Hosts: Parisoma sucaeruleum (Passeriformes: Sylviidae), Zosterops lateralis, Z. palpebrosa japonica (Passeriformes: Zosteropidae), Alcippe nipalensis peracensis (Passeriformes: Timaliidae)

Ptilonyssus ruandae sylviae Fain

Ptilonyssus ruandae sylviae Fain, 1963a: 172; Fain, Sixl & Moritsch, 1974: 2

Type locality: Clermont (Namur), Strée (Hainaut)

Distribution: Austria, Belgium

Hosts: Sylvia a. atricapilla, S. c. communis (Passeriformes: Sylviidae)

Ptilonyssus sairae Castro

Ptilonyssus (Ptilonyssus) sairae Castro, 1948: 260; Pereira & Castro, 1949: 219 Ptilonyssus sairae: Zumpt & Till, 1955: 89; Fain, 1956a: 144; 1957b: 102; George, 1961: 125; Fain & Aitken, 1967: 4; 1968: 3; Amaral, 1968: 120; Spicer, 1977a: 598; 1977b: 498; 1978: 892; 1984: 799; 1987: 261; Kadosaka, Kaneko & Asanuma, 1987: 35; Morelli & Spicer, 2007: 183; Mascarenhas, Coimbra et al., 2011: 507; Knee & Galloway, 2017: 94–95; Dimov, 2016; 2018: 89

Ptilonyssus constrictus Ford, 1961: 5 – type from Dendroica petechia, Seiurus noveboracensis (Passeriformes: Parulidae), Trinidad, USA (syn. by Pence, 1973b: 877)

Ptilonyssus constrictus longisetosus Černy & Dusbábek, 1970: 481 – type from Seiurus motacilla (Passeriformes: Parulidae), Havana, Cuba

Ptilonyssus garridoi Cruz, 1971: 1

Type locality of Ptilonyssus (Ptilonyssus) sairae Castro: Brazil

Distribution: Brazil, Canada, Cuba, Guatemala, Japan, Mexico, Russia, Trinidad, USA (incl. Alasca)

Hosts: Emberiza schoeniclus (Passeriformes: Emberizidae), Aimophila ruficeps, Ammodramus savannarum, Arremon taciturnus, Chondestes grammacus, Junco caniceps, J. hyemalis, Melospiza melodia, Paroaria coronata, Pooecetes gramineus, Spizella passerina, S. pallida, S. pusilla, Zonotrichia querula (Passeriformes: Passerellidae), Chlorospingus ophthalmicus, Tangara gyrola, T. seledon, Tiaris fuliginosa (Passeriformes: Thraupidae), Dendroica auduboni, D. coronata, D. erithacorides, D. petechia, D. pinus, Geothlypis philadelphia, Muniotitla varia, Oreothlypis peregrina, O. ruficapilla, Parula americana, Seiurus noveboracensis, Setophaga pensylvanica, S. fusca, Vermivora chrysoptera, V. celata, V. ruficapilla, Wilsonia pusilla (Passeriformes: Parulidae), Passerina ciris, Guiraca caerulea (Passeriformes: Cardinalidae), Parus carolinensis, P. major (Passeriformes: Paridae), Carduelis pinus, Carduelis [Spinus] tristis, Carduelis chlorus, Chloris sinica minor (Passeriformes: Fringillidae), Polioptila albiloris (Passeriformes: Polioptilidae), Euphagus cynocephalus, Molothrus ater (Passeriformes: Icteridae)

P. S. Pence & Casto (1976) consider Ptilonyssus japuibensis Castro, P. japuibensis cyanocompsae Fain et Aitken, P. agelaii Fain, P. teretistris Černy, P. ludovicianus Černy and P. garridoi Cruz, but this was not followed by the Canadian and some other authors.

Ptilonyssus salpinctis George

Ptilonyssus salpinctis George, 1961: 116

Type locality: Brisko Co., Texas

Distribution: USA (Texas)

Host: Salpinctes obsoletus (Passeriformes: Troglodytidae)

Ptilonyssus saltator Fain et Aiken

Ptilonyssus saltator Fain & Aiken, 1971: 37

Type locality: Ananindeua

Distribution: Brazil

Host: Saltator maximus (Passeriformes: Fringillidae)

Ptilonyssus schoenobaeni Fain, Sixl et Moritsch

Ptilonyssus schoenobaeni Fain, Sixl & Moritsch, 1974: 4

Type locality: Illmitz, Burgenland

Distribution: Austria

Host: Acrocephalus schoenobaenus (Passeriformes: Acrocephalidae)

Ptilonyssus schumili Butenko et Lavrovskaya

Ptilonyssus schumili Butenko & Lavrovskaya, 1980: 522; Scherbinina & Butenko, 1981: 70; Dimov, 2016; 2018: 86

Type locality: Ryazanskaya Obl., Okskiy Zap.

Distribution: Russia, Kazahstan, Kirghizstan, Turkmenistan

Hosts: Alauda arvensis, Galerida cristata (Passeriformes: Alaudidae)

Ptilonyssus sclateriae Fain et Aitken

Ptilonyssus schoenobaeni Fain & Aitken, 1970: 401

Type locality: Mucambo Forest, IPEAN, Belém

Distribution: Brazil

Host: Sclateria naevia (Passeriformes: Thamnophilidae)

Ptilonyssus sclerurus Fain et Aitken

Ptilonyssus sclerurus Fain & Aitken, 1970: 395

Type locality: Pirelli Plantation, Marituba

Distribution: Brazil

Host: Sclerurus rufigularis (Passeriformes: Furnariidae)

Ptilonyssus serini Fain

Ptilonyssus serini Fain, 1956a: 139; 1957b: 90

Type locality: Akanyaru, Rwanda

Distribution: Rwanda

Hosts: Serinus canicollis sassii, S. citrinelloides frontalis, S. mozambicus barbatus (Passeriformes: Fringillidae)

Ptilonyssus setosae Domrow

Ptilonyssus setosae Domrow, 1969b: 362; 1988: 875

Type locality: Ella Bay, Queensland

Distribution: Australia

Hosts: Rhipidura rufifrons, R. setosa (Passeriformes: Muscicapidae)

Ptilonyssus sialiae George

Ptilonyssus sialiae George, 1961: 122

Type locality: Lubbock Co., Texas

Distribution: USA (Texas)

Host: Sialia currucoides (Passeriformes: Turdidae)

Ptilonyssus sittae Fain

Ptilonyssus sittae Fain, 1965: 158; 1966: 117; Domrow, 1969b: 348; 1988: 875; Butenko, 1969; Sixl, 1969: 218; Shumilo & Lunkashu, 1970: 33; Sixl & Reich, 1971: 401; Fain, Sixl & Moritsch, 1974: 3

Ptilonyssus strandtmannianus Feider & Mironescu, 1969: 644 (syn. by Domrow, 1972: 575)

Ptilonyssus bregetovae Feider & Mironescu, 1969: 656 (syn. by Domrow, 1972: 575)

Ptilonyssus pelmaspis Feider & Mironescu, 1969: 656 (syn. by Domrow, 1972: 575)

Ptilonyssus maxvachoni Feider & Mironescu, 1970: 17 (syn. by Domrow, 1972: 575)

Type locality: Kappellen, nr. Antwerp (Belgium)

Distribution: Australia, Austria, Belgium, Moldavia, Romania, Ukraine

Hosts: Climacteris picumnus (Passeriformes: Climacteridae), Sitta europaea (Passeriformes: Sittidae), Parus major (Passeriformes: Paridae)

Ptilonyssus souzai Pereira et Castro

Ptilonyssus (Rhinonyssoides) souzai Pereira & Castro, 1949: 84; Zumpt & Till, 1955: 88

Ptilonyssus souzai: Fain, 1957b: 84; Fain & Johnston, 1966: 29

Type locality: Brazil

Distribution: Brazil

Hosts: Milvago chimachima chimachima (Falconiformes: Falconidae, Polyborinae), Rupornis magnirostris magniplumis (Accipitriformes: Accipitridae)

Ptilonyssus sphecotheris Domrow

Ptilonyssus sphecotheris Domrow, 1964d: 613; 1967: 218; 1969b: 372; 1988: 875

Type locality: Brisbane, Qu.

Distribution: Australia

Hosts: Chlamydera nuchalis (Passeriformes: Ptilonorhynchidae), Sphecotheres flaviventris, S. vieillotii (Passeriformes: Oriolidae)

Ptilonyssus spini Stanyukovich et Butenko

Ptilonyssus spini Stanyukovich & Butenko, 2003: 34; Dimov, 2018: 92

Type locality: Ribachiy, Kaliningradskaya Oblast

Distribution: Russia (Kaliningrad)

Host: Spinus spinus (Passeriformes: Fringillidae)

Ptilonyssus spinosus (Brooks et Strandtmann)

Tyranninyssus spinosus Brooks & Strandtmann, 1960: 421; Hyland, 1961: 279; Černý & Dusbábek, 1970: 484

Ptilonyssus spinosus (Brooks et Strandtmann): Mendes et al., 2014: 480

Type locality: Lubbock County, Texas

Distribution: Brazil, Cuba (incl. Pinos), Puerto Rico, USA (Michigan, Texas)

Hosts: Machetornis rixosa, Muscivora forficata, Pitangus sulphuratus, Tyrannus c. caudifasciatus, T. c. flavescens, T. dominicensis, T. melancholicus, T. tyranni, T. tyrannus, T. verticalis (Passeriformes: Tyrannidae)

Ptilonyssus steganurae Fain

Ptilonyssus steganurae Fain, 1967: 150

Type locality: Zoo Anvers

Distribution: Central Africa

Host: Vidua [Steganura] paradisea (Passeriformes: Viduidae)

Ptilonyssus sternostomicus Fain et Aitken

Ptilonyssus sternostomicus Fain & Aitken, 1968: 20; 1970: 353

Type locality: APEG Forest

Distribution: Brazil

Host: Schiffornis turdinus (Passeriformes: Pipridae)

Ptilonyssus stomioperae Domrow

Ptilonyssus stomioperae Domrow, 1966a: 206; 1967a: 217; 1969b: 317; 1979d: 504; 1988: 876

Type locality: Magnificent Creek, Mitchell River

Distribution: Australia

Hosts: Meliphaga flava, Stomiopera unicolor (Passeriformes: Meliphagidae)

Ptilonyssus strandtmanni Fain

Ptilonyssus strandtmanni Fain, 1956a: 148; 1957b: 86; Fain, Herin & Pulyaert, 1977: 387; Guevara-Benítez & Úbeda Ontíveros, 1980a: 231; Sartbaev, 1983: 84; Butenko, 1984: 157

Type locality: Astrida (= Butare), Rwanda

Distribution: Cameroon, Kirghizstan, Russia, Rwanda, Spain

Hosts: Apus affinis, A. apus, A. caffer streubeli (Apodiformes: Apodidae)

Ptilonyssus stresemanni Vitzthum

Ptilonyssus stresemanni Vitzthum, 1935: 579; Castro, 1948: 260; Pereira & Castro, 1949: 219; Zumpt & Till, 1955: 89; Fain, 1957b: 101; 1961a: 511

Type locality: Brazil

Distribution: Brazil

Host: Procnias albus (Passeriformes: Cotingidae)

Ptilonyssus struthideae Domrow

Ptilonyssus struthideae Domrow, 1969d: 343; 1988: 876

Type locality: Condamine, Charleville, Qu.

Distribution: Australia

Host: Struthidea cinerea (Passeriformes: Grallinidae)

Ptilonyssus sturnopastoris Fain

Ptilonyssus sturnopastoris Fain, 1963d: 1; Domrow, 1969b: 372; 1988: 876

Type locality: N. Queensland (described from Anvers, Belgium)

Distribution: Australia

Hosts: Aplonis metallica, Sturnopastor capensis (Passeriformes: Sturnidae)

Ptilonyssus sylviicola Stanyukovich et Butenko

Ptilonyssus sylviicola Stanyukovich & Butenko, 2003: 40

Type locality: Tatarstan

Distribution: Russia (Tatarstan)

Host: Sylvia communis (Passeriformes: Fringillidae)

Ptilonyssus tachycinetae George

Ptilonyssus tachycinetae George, 1961: 112; Butenko, 1965: 132

Type locality: Florida City, Florida

Distribution: Russia, USA (Florida)

Hosts: Riparia riparia, Tachycineta bicolor (Passeriformes: Hirundinidae)

Ptilonyssus terenotricci Fain et Aitken

Ptilonyssus terenotricci Fain & Aitken, 1970: 357

Type locality: Mosqueiro Ferry, Benavides, Belém

Distribution: Brazil

Host: Terenotriccus erythrurus (Passeriformes: Tyrannidae)

Ptilonyssus teretistris Černý

Ptilonyssus teretistris Černý, 1969: 227; Černý & Dusbábek, 1970: 482

Type locality: El Dorado, prov. of Las Villas

Distribution: Cuba

Host: Teretistris fornsi (Passeriformes: Parulidae)

Ptilonyssus terpsiphonei Fain

Ptilonyssus terpsiphonei Fain, 1956a: 145; 1957b: 137; 1959a: 22; Strandtmann & Wharton, 1958: 182; Domrow, 1965b: 453; 1969b: 356; 1988: 876; Sakakibara, 1968b: 309

Type locality: Astrida (= Butare), Rwanda

Distribution: Australia, Papua New Guinea, Rwanda, South Africa

Hosts: Carterornis leucotis, Myagra rubecula, Piesorhynchus alecto, Seisura inquieta (Passeriformes: Muscicapidae), Arses kaupi, Monarcha melanopsis, M. trivirgata, Terpsiphone viridis, T. viridis kivuensis (Passeriformes: Monarchidae)

Ptilonyssus thamnomanes Fain et Aitken

Ptilonyssus thamnomanes Fain & Aitken, 1970: 374

Type locality: Mosqueiro Ferry, Benavides, Belém

Distribution: Brazil

Host: Thamnomanes caesius (Passeriformes: Formicariidae)

Ptilonyssus thamnophili Fain et Aitken

Ptilonyssus thamnophili Fain & Aitken, 1967: 9; 1970: 356

Type locality: Elgin Trace, Vega de Oropouche (Trinidad)

Distribution: Brazil, Trinidad

Hosts: Thamnophilus aethiops, Th. doliatus (Passeriformes: Thamnophilidae), Platyrinchus saturatus (Passeriformes: Tyrannidae)

Ptilonyssus thryothori Pence

Ptilonyssus thryothori Pence, 1972d: 1165; 1973b: 876

Type locality: Laplace, La.

Distribution: USA (Louisiana)

Host: Thryothorus ludovicianus (Passeriformes: Troglodytidae)

Ptilonyssus thymanzae Domrow

Ptilonyssus thymanzae Domrow, 1964d: 604, 617; 1966a: 204, 1969b: 317; 1979: 505; 1988: 876

Ptilonyssus sp. Domrow, 1965b: 548

Type locality: Condamine, Qu.

Distribution: Australia

Hosts: Acanthagenys rufogularis, Anthochaera chrysoptera, Meliphaga chrysops, M. fusca, M. lewinii, M. melanops, M. notata, M. penicillata, Myzantha flavigula, M. melanocephala (Passeriformes: Meliphagidae)

Ptilonyssus tillae Fain

Ptilonyssus tillae Fain, 1959a: 25

Type locality: Sabi Sand Wildtuin, near Newington, Transvaal

Distribution: Rwanda, South Africa

Hosts: Lamprocolius chalybeus, Lamprotornis australis, L. mevesii (Passeriformes: Sturnidae)

Ptilonyssus toxostomae Pence

Ptilonyssus toxostomae Pence, 1972d: 1163; 1973b: 876; Spicer, 1987: 261

Type locality: Louisiana, Covington

Distribution: USA (Louisiana)

Host: Toxostoma rufum (Passeriformes: Mimidae)

Ptilonyssus traubi Strandtmann

Ptilonyssus traubi Strandtmann, 1960: 144

Type locality: Kinabalu Mt., Tenompak (Sabah)

Distribution: Malaysia (Sabah)

Host: Aves indet.

Ptilonyssus travassosfilhoi (Castro)

Neonyssus (Paraneonyssus) travassosfilhoi Castro, 1948: 275

Ptilonyssus (Paraneonyssus) travassosfilhoi (Castro): Pereira & Castro, 1949: 227

Ptilonyssus travassosfilhoi (Castro): Zumpt & Till, 1955: 88 ; Fain, 1957b: 88; Fain & Aitken, 1970: 352

Type locality: Juquiá (Estado de São Paulo)

Distribution: Brazil

Hosts: Philydor lichtensteini (Passeriformes: Furnariidae, Dendrocolaptinae), Automolus infuscatus (Passeriformes: Furnariidae)

Ptilonyssus triscutatus (Vitzthum)

Ptilonyssoides triscutatus Vitzthum, 1935: 581; Strandtmann & Wharton, 1958: 170; Shumilo & Lunkashu, 1970: 28; Butenko, 1984: 161

Neonyssus (Ptilonyssoides) triscutatus (Vitzthum): Castro, 1948: 276

Neonyssus triscutatus (Vitzthum): Zumpt & Till, 1955: 68

Ptilonyssus triscutatus Fain, 1956b: 136; 1957b: 121; Nadchatram, McClure & Chong, 1964: 104; Domrow, 1966b: 196; 1967: 217; 1969b: 347; 1979: 494; 1988: 877; Fain, Herin & Puylaert, 1977: 386; Scherbinina & Butenko, 1981: 70

Type locality: Europe

Distribution: Australia, Cameroon, Kazakhstan, Malaysia, Moldavia, Russia, Rwanda, Turkmenistan, Ukraine

Hosts: Melittophagus bullocki, Merops apiaster, M. nubicus, M. ornatus, M. persicus, M. superciliosus, M. viridis (Coraciiformes: Meropidae)

Ptilonyssus troglodytis Fain

Ptilonyssus troglodytis Fain, 1964a: 60; Knee, Proctor & Galloway, 2008: 370

Type locality: Bois de Oisquercq (Brabant)

Distribution: Belgium, Canada

Host: Troglodytes troglodytes (Passeriformes: Troglodytidae)

Ptilonyssus trouessarti (Hirst)

Rhinonyssoides trouessarti Hirst, 1921: 770; Strandtmann & Wharton, 1958: 172; syn. with Ptilonyssus by Zumpt & Till, 1955: 69

Ptilonyssus trouessarti: Zumpt & Till, 1955: 69; Fain & Hyland, 1962: 346; Domrow, 1964d: 608; 1969b: 373; 1979: 510; 1988: 877; Sakakibara, 1968b: 302

Type locality: Australia

Distribution: Australia, Papua New Guinea

Hosts: Oriolus flavocinctus, O. sagittatus; type from «Sphecotheres maxillaris» (Passeriformes: Oriolidae)

Ptilonyssus trouessarti pseudotrouessarti Sakakibara

Ptilonyssus trouessarti pseudotrouessarti Sakakibara, 1968b: 305

Type locality: Bulolo River

Distribution: Papua New Guinea

Host: “Leatherneck” (Tropidorhynchus sordidus – Meliphagidae)

Ptilonyssus tyrannus (Brooks et Strandtmann)

Tyranninyssus tyrannus Brooks & Strandtmann, 1960: 419; Hyland & Moorhouse, 1970: 48

Tyranninyssus tyrannus caribaeus Černý, 1969: 233. Type: Cuba, Santo Tomas, Cienaga de Zapata, prov. Las Villas, from Contopus caribaeus caribaeus

Tyranninyssus tyrannisoides Brooks & Strandtmann, 1960: 420. Type: Lamb County, Texas, from Empidonax minimus, Sayornis sayus (Passeriformes: Tyrannidae). Syn. by Hyland & Moorhouse, 1970: 48

Ptilonyssus tyrannus Pence, 1972: 790; Knee, Proctor & Galloway, 2008: 370; Knee & Galloway, 2017: 95

Type locality: Lamb County, Texas

Distribution: Guatemala, Mexico, Canada, USA (Texas)

Hosts: Contopus sordlidulus, C. caribaeus caribaeus, Empidonax alnorum, E. flavescens, E. flaviventris, E. minimus, E. traillii, Mitrephanes phaeocercus, Nuttallornis borealis, Sayornis phoebe, S. sayus, Tyrannus melancholicus (Passeriformes: Tyrannidae)

Ptilonyssus viduicola Fain

Ptilonyssus viduicola Fain, 1962c: 135; Fain, Herin & Puylaert, 1977: 388

Type locality: Akanyaru (Rwanda)

Distribution: Cameroon, Rwanda

Host: Vidua macroura (Passeriformes: Viduidae)

Ptilonyssus vireonis (Dusbábek)

Passeronyssus vireonis Dusbábek, 1969: 220; Černy & Dusbábek, 1970: 486

Ptilonyssus vireonis (Dusbábek): Pence, 1972c: 793; Knee, Proctor & Galloway, 2008: 370

Type locality: Havana

Distribution: Canada, Cuba, USA (La.)

Hosts: Vireo o. olivaceus, V. solitarius (Passeriformes: Vireonidae)

Ptilonyssus vossi Spicer

Ptilonyssus vossi Spicer, 1977a: 595; 1987: 261

Type locality: Fort Worth, Tarrant Co., Texas

Distribution: USA (Texas)

Host: Troglodytes aedon (Passeriformes: Troglodytidae)

Ptilonyssus walterberghi Fain

Ptilonyssus walterberghi Fain, 1958: 3

Type locality: Anvers Zoo

Distribution: Japan

Host: Suthora webbiana (Passeriformes: Paridae)

Ptilonyssus werneri (Castro)

Neonyssus (Rochanyssus) werneri Castro, 1948: 272; Pereira & Castro, 1949: 226

Ptilonyssus (Rochanyssus) werneri (Castro): Pereira & Castro, 1949: 226

Ptilonyssus werneri (Castro): Zumpt & Till, 1955: 90; Fain, 1957b: 86; Fain & Aitken, 1967: 3; 1970: 355; Amaral, 1968: 120

Type locality: Botucatû (Estado de São Paulo)

Distribution: Brazil, Trinidad

Hosts: Elaenia flavogaster, Е. obscura sordida (Passeriformes: Tyrannidae), Gnorimopsar chopi chopi, Sturnella [Leistis militares] superciliaris (Passeriformes: Icteridae)

Ptilonyssus wilsoni Sakakibara

Ptilonyssus wilsoni Sakakibara, 1967: 599; Tenorio, 1976: 19; Domrow & Nadchatram, 1978: 87

Type locality: Enarotali, NW New Guinea

Distribution: Malaysia, Papua New Guinea

Hosts: Apus affinis, Collocalia esculenta (Apodiformes: Apodidae)

Ptilonyssus xenops Fain et Aitken

Ptilonyssus xenops Fain & Aitken, 1968: 8; 1971: 30

Type locality: APEG Forest

Distribution: Brazil

Host: Xenops minutus (Passeriformes: Furnariidae)

Ptilonyssus xyphorhynchus Fain et Aitken

Ptilonyssus xyphorhynchus Fain & Aitken, 1970: 369

Type locality: Mosqueiro Ferry, Benavides, Belém

Distribution: Brazil

Host: Xyphorhynchus spixii (Passeriformes: Dendrocolapidae)

Ptilonyssus zeferinoi Amaral

Ptilonyssus zeferinoi Amaral, 1968: 107

Type locality: Estado de São Paulo

Distribution: Brazil

Hosts: Gnorimopsar chopi chopi, Sturnella [Leistis militares] superciliaris (Passeriformes: Icteridae)

Ptilonyssus zumpti Fain

Ptilonyssus zumpti Fain, 1956b: 138; 1957b: 91

Type locality: Akanyaru (Rwanda)

Distribution: Rwanda

Host: Amblyospiza albifrons montana (Passeriformes: Ploceidae)

Genus Sternostoma Berlese et Trouessart

Sternostoma Berlese & Trouessart, 1889: 126

Type species: Sternostoma cryptorhynchum Berlese et Trouessart, 1889

Sternostomum Vitzthum, 1935: 569

Type species: Sternostomum technaui Vitzthum, 1935

Agapornyssus Gretillat et al., 1959: 376

Type species: Agapornyssus faini Grétillat, Capron & Brygoo, 1959

Sternostomoides Bregetova, 1965: 709 – syn. by Domrow, 1969: 374

Type species: Sternostomum technaui Vitzthum, 1935

Rhinosterna Fain, 1964: 125 – syn. by Domrow, 1969: 374

Type species: Rhinosterna aymarae Fain, 1964

Sternoecius Fain et Aitken, 1967: 24 – syn. by Domrow, 1969: 374

Type species: Sternoecius piprae Fain et Aitken, 1967

Sternostoma alexfaini Feider et Mironescu

Sternostoma alexfaini Feider & Mironescu, 1969: 32

Type locality: Sinaia

Distribution: Romania

Host: Lanius excubitor (Passeriformes: Laniidae)

Sternostoma artami Feider et Mironescu

Sternostoma artami Feider & Mironescu, 1982: 87

Type locality: Papua New Guinea

Distribution: Papua New Guinea

Host: Artamus leucorhynchus (Passeriformes: Artamidae)

Sternostoma augei Amaral

Sternostoma augei Amaral, 1962: 69; Pence & Casto, 1975b: 362

Type locality: Itaporanga (Estado do São Paulo)

Distribution: Brazil, USA (Texas)

Host: Speotyto cunicularia grallaria (Strigiformes: Strigidae)

Sternostoma aymarae (Fain)

Rhinosterna aymarae Fain, 1964e: 126

Sternostoma aymarae (Fain): Domrow, 1969: 374

Type locality: Argentina

Distribution: Argentina

Host: Amoropsittaca aymara (Psittaciformes: Psittacidae)

Sternostoma batis Fain

Sternostoma laniorum var. batis Fain, 1957b: 77

Sternostoma batis Fain: Butenko, 1965; Fain & Aitken, 1968: 22; 1970: 349; Sartbaev, 1983: 84; Úbeda, Rodríguez & Guevara, 2000

Type locality: Akanyaru (Rwanda)

Distribution: Brazil, Kirghizstan, Russia, Rwanda, Spain

Hosts: Alauda arvensis, Eremophila alpestris (Passeriformes: Alaudidae), Batis molitor puella, Oenanthe oenanthe (Passeriformes: Muscicapidae), Xiphorhynchos spixii (Passeriformes: Furnariidae, Dendrocolaptinae), Pyriglena leucoptera (Passeriformes: Furnariidae), Loxia curvirostra (Passeriformes: Fringillidae)

Sternostoma boydae Strandtmann (nom. emend.)

Sternostoma boydi Strandtmann, 1951: 138 (named after Elizabeth Boyd); Zumpt & Till, 1955: 86; Fain, 1956a: 151; 1957b: 66; 1960c: 95; 1963a: 176; Fain & Aitken, 1967: 16; Fain & Holland, 1972: 32; Pence, 1972: 781; Feider & Mironescu, 1972: 326; Spicer, 1978: 894; Sartbaev, 1983: 84; Butenko, 1984: 84; Knee, 2008: 9

Type locality: Galveston, Texas, USA

Distribution: Australia, Belgium, Canada, Kazakhstan, Kirghizstan, Russia (incl. Far East), Trinidad, USA, Venezuela, Zambia

Hosts: Actitis macularia, Arenaria interpres, Calidris alba, Crocethia alba, Ereunetes pusillus, Limosa lapponica, Numenius arquatus, Phalaropus lobatus, Philomachus pugnax, Tringa [Totanus] erythropus, T. nebularia, T. ochropus, T. totanus, Rhyacophilus glareola (Charadriiformes: Scolopacidae), Chlidonias leucoptera, Onychophon [Sterna] fuscatus fuscatus, Sterna hirundo (Charadriiformes: Sternidae), Larus atricilla, L. californicus, L. delawarensis, L. glaucescens, L. occidentalis (Charadriiformes: Laridae)

Sternostoma bruxellarum Fain

Sternostoma bruxellarum Fain, 1961b: 53; Butenko, 1965; Sixl, 1969; Shumilo & Lunkashu, 1970: 24; Beron, 1975: 7

Type locality: Liège Prov. and Brussels

Distribution: Austria, Belgium, Bulgaria, Moldavia, Russia, Ukraine

Host: Sturnus vulgaris (Passeriformes: Sturnidae)

Sternostoma callithrix Fain et Aitken

Sternostoma callithrix Fain & Aitken, 1970: 343

Type locality: Ananindeua, Para

Distribution: Brazil

Host: Empidonomus varius (Passeriformes: Tyrannidae)

Sternostoma christinae Guevara – Benítez, Lopez-Roman et Úbeda – Ontíveros

Sternostoma christinae Guevara – Benítez, Lopez-Roman & Úbeda – Ontíveros, 1974: 161

Type locality: Granada

Distribution: Spain

Host: Phylloscopus collybita (Passeriformes: Phylloscopidae)

Sternostoma cisticolae Fain

Sternostoma cisticolae Fain, 1957c: 218; 1959a: 32

Type locality: Debeete

Distribution: Botswana, South Africa

Hosts: Apalis flavida, Cisticola chiniana (Passeriformes: Cisticolidae)

Sternostoma clementei Amaral

Sternostoma clementei Amaral, 1968: 114

Type locality: Boracéa, Estado de Sao Paulo

Distribution: Brazil

Host: Knipolegus cyanirostris (Passeriformes: Tyrannidae)

Sternostoma colii Fain

Sternostoma colii Fain, 1956b: 392; 1957b: 79

Type locality: Astrida (= Butare)

Distribution: Rwanda

Host: Colius striatus kiwuensis (Coliiformes: Coliidae)

Sternostoma constricta Feider et Mironescu

Sternostoma constricta Feider & Mironescu, 1982: 87

Type locality: Papua New Guinea

Distribution: Papua New Guinea

Host: Paradisaea reggiana (Passeriformes: Paradisaeidae)

Sternostoma cooremani Fain

Sternostoma cooremani Fain, 1956a: 154; 1957b: 72; Strandtmann & Wharton, 1958: 166; Domrow, 1965b: 448; 1966b: 192; 1969b: 376; Shumilo & Lunkashu, 1970: 24; Fain, Herin & Puylaert, 1977: 392; Sartbaev, 1981: 130; Butenko, 1984: 151

Type locality: Astrida (= Butare), Rwanda

Distribution: Australia, Cameroon, Kirghizstan, Malaysia, Moldavia, Russia, Rwanda

Hosts: Dacelo gigas, Melittophagus lafresnayi oreobates, Merops apiaster, M. nubicoides, M. orientalis, M. ornatus (Coraciiformes: Meropidae)

Sternostoma cooremani halcyoni Fain et Nadchatram

Sternostoma cooremani halcyoni Fain & Nadchatram, 1962: 280

Sternostoma cooremani: McClure, Ratanaworadhan et al., 1973: 37

Type locality: Rantau Panjang, Selangor

Distribution: Malaysia

Host: Halcyon chloris humii (Coraciiformes: Halcyonidae)

Sternostoma cordiscutata Feider et Mironescu

Sternostoma cordiscutata Feider & Mironescu, 1982: 87

Type locality: Papua New Guinea

Distribution: Papua New Guinea

Host: Chalcites meyeri (Cuculiformes: Cuculidae)

Sternostoma crotophagae Pence et Casto

Sternostoma crotophagae Pence & Casto, 1975: 364

Type locality: La Salle Co., Millett, Texas

Distribution: USA (Texas)

Host: Crotophaga sulcirostris (Cuculiformes: Cuculidae)

Sternostoma cryptorhynchum Berlese et Trouessart

Sternostoma cryptorhynchum Berlese & Trouessart, 1889: 128; Vitzthum, 1935: 571; Castro, 1948: 257; Zumpt & Till, 1955: 86; Fain, 1956a: 152; 1957b: 70; Domrow, 1969: 379; Fain, Hehn & Puylaert, 1977: 392; Knee, Proctor & Galloway, 2008: 369

Type locality: Europe

Distribution: France, Cameroon, Canada

Hosts: Clyptospiza monteiri (Passeriformes: Estrildidae), Passer domesticus (Passeriformes: Passeridae), Pinicola enucleator (Passeriformes: Fringillidae)

Sternostoma cuculorum Fain

Sternostoma cuculorum Fain, 1956a: 155; 1957b: 74; 1957c: 218; Strandtmann & Wharton, 1958: 166; Domrow, 1965b: 449; 1969b: 376; 1979: 510; Shumilo & Lunkashu, 1970: 25; McClure, Ratanaworadhan et al., 1973: 37; Butenko, 1984: 155

Sternostoma cuculorum var. urolestis Fain, 1959a: 32 – South Africa, ex Urolestes melanoleucus (Laniidae)

Sternostoma laniorum Fain: Strandtmann & Wharton, 1958: 167; Domrow, 1965b: 449; 1966b: 193; Shumilo & Lunkashu, 1970: 26

Sternostoma zosteropus Domrow, 1966a: 194

Type locality: Gakoma [Rwanda]

Distribution: Australia, Botswana, Canada, Malaysia, Moldavia, Russia, Rwanda, Ukraine

Hosts: Arses kaupi, Cacomantis pyrrhophanus, C. variolosus, Colluricincla megarhyncha (Passeriformes: Artamidae), Eopsaltria capito, Eudynamys orientalis, Monarcha trivirgata, Myiagra rubecula, Oreoica gutturalis (Passeriformes: Falcunculidae), Rhipidura leucophrys (Passeriformes: Rhipiduridae), Zosterops lateralis (Passeriformes: Zosteropidae), Chrysococcyx caprius, Clamator levaillanti, Cuculus canorus, C. solitarius, Phaenicophaeus chlorophaeus (Cuculiformes: Cuculidae); Urolestes melanoleucus (Passeriformes: Laniidae), Yuhina zantholeuca (Passeriformes: Timaliidae), Sturnus sturninus (Passeriformes: Sturnidae), Catharus ustulatus (Passeriformes: Turdidae)

Sternostoma cuculorum korolevae Bregetova et Slavoshevskaya

Sternostoma cuculorum korolevae Bregetova & Slavoshevskaya, 1983: 234

Type locality: Altai, Troitskiy Rayon, Malaya rechka Vill.

Distribution: Russia

Host: Cuculus sp. (Cuculiformes: Cuculidae)

Sternostoma darlingi Spicer

Sternostoma darlingi Spicer, 1984: 795

Type locality: Guatemala

Distribution: Guatemala

Host: Mitrephanes phaeocercus (Passeriformes: Tyrannidae)

Sternostoma dumetellae Pence

Sternostoma dumetellae Pence, 1972b: 785; Spicer, 1987: 261

Type locality: Laplace, La.

Distribution: USA

Host: Dumetella carolinensis (Passeriformes: Mimidae)

Sternostoma dureni Fain

Sternostoma dureni Fain, 1956a: 153; 1957b: 68; Strandtmann & Wharton, 1958: 166; Domrow, 1969b: 376; Dimov, 2016; 2018: 62

Type locality: Astrida (= Butare), Rwanda

Distribution: Australia, Russia, Rwanda

Hosts: Oreocincla lunulata, Turdoides jardinei emini, T. melanops sharpei, Turdus merula, T. olivaceus graueri (Passeriformes: Turdidae)

Sternostoma epistomata Feider et Mironescu

Sternostoma epistomata Feider & Mironescu, 1973: 333; Butenko, 1984: 139

Sternostoma chlidoniadis Butenko, 1974: 1089 – type: Novosibirskaya obl., Zdvinskiy rayon, Malie Chany Lake, from Chlidonias nigra (syn. by Butenko, 1984)

Type locality: Tulcea (Romania)

Distribution: Romania, Russia

Hosts: Chlidonias hybrida, Ch. nigra (Charadriiformes: Sternidae)

Sternostoma eurocephali Fain

Sternostoma eurocephali Fain, 1960: 111

Type locality: Gravelotte (Transvaal)

Distribution: South Africa

Host: Eurocephalus anguitimens (Passeriformes: Laniidae)

Sternostoma ficedulae Fain et Sixl

Sternostoma eurocephali Fain & Sixl, 1971: 89; Fain, Sixl & Moritsch, 1974: 6

Type locality: Brunnsee (Styria)

Distribution: Austria

Host: Ficedula albicollis (Passeriformes: Motacillidae)

Sternostoma francolini Fain

Sternostoma francolini Fain, 1960c: 95; 1961a: 518; Sartbaev, 1979; 1983: 83; Butenko, 1984: 153

Type locality: Lochinvar, nr. Monze (Zambia)

Distribution: Belgium, Kazakhstan, Kyrghizstan, Russia, Zambia

Hosts: Alectoris chukar, Francolinus coqui, Perdix perdix perdix (Galliformes: Phasianidae)

Sternostoma fulicae Fain et Bafort

Sternostoma fulicae Fain & Bafort, 1963: 474; Domrow, 1969: 380; Feider & Mironescu, 1973: 159; Butenko, 1976: 77, 1984: 149; Sartbaev, 1981: 130; de Rojas et al., 2001: 961; de Rojas, Mora et al., 2002: 676

Type locality: Belgium

Distribution: Australia, Azerbaidjan, Belgium, Kirghizstan, Romania, Russia, Spain

Hosts: Fulica atra, Gallinula chloropus (Gruiformes: Rallidae)

Sternostoma furmani Strandtmann

Sternostoma furmani Strandtmann, 1960: 132

Type locality: Trus Madi Mt., Pampang, Uli Kaingaran, North Borneo

Distribution: Malaysia (Borneo)

Host: Arborophila brunneopectus erythrophys (Galliformes: Phasianidae)

Sternostoma galowayi Knee

Sternostoma galowayi Knee, 2018

Type locality: Winnipeg, Manitoba

Distribution: Canada

Host: Eremophila alpestris (Passeriformes: Alaudidae)

Sternostoma giganteum Fain

Sternostoma giganteum Fain, 1962c: 148

Type locality: Kisenyi

Distribution: Rwanda

Host: Batis molitor puella (Passeriformes: Muscicapidae)

Sternostoma gliciphilae Domrow

Sternostoma gliciphilae Domrow, 1966a: 192; 1969b: 377; Fain & Lukoschus, 1979: 19

Type locality: Esk

Distribution: Australia

Hosts: Gliciphila indistincta, Ramsayornis fasciatus (Passeriformes: Meliphagidae)

Sternostoma graculi (Butenko)(new comb.)

Sternostomoides graculi Butenko, 1999: 47

Type locality: Kyrghizstan

Distribution: Kyrghizstan

Host: Pyrrhocorax [Graculus] graculus (Passeriformes: Corvidae)

Sternostoma guevarai Guevara-Benítez, Úbeda-Ontíveros et Cutillas-Barrios

Sternostoma guevarai Guevara-Benítez, Úbeda-Ontíveros & Cutillas-Barrios, 1979: 29

Type locality: Granada

Distribution: Spain

Host: Alectoris rufa (Galliformes: Phasianidae)

Sternostoma hedonophilum Fain et Aitken

Sternostoma hedonophilum Fain & Aitken, 1970: 348

Type locality: Mosqueiro Ferry, Benavides, Belém

Distribution: Brazil

Host: Platyrhinchus saturatus (Passeriformes: Tyrannidae)

Sternostoma hirundinis Fain

Sternostoma hirundinis Fain, 1956b: 154; 1957b: 72; 1959a: 31; Shumilo & Lunkashu, 1970: 25; Černy & Dusbábek, 1970: 485; Pence, 1973a: 359; Spicer, 1977b: 498; 1987: 261

Type locality: Astrida (= Butare), Rwanda

Distribution: Cuba, Mexico, Moldavia, Rwanda, South Africa, Ukraine, USA (La.)

Hosts: Hirundo rustica, H. r. erythrogaster, H. smithi, Progne subis, Psalidoprocne albiceps, Riparia riparia, Tachycineta albilina (Passeriformes: Hirundinidae), Bombycilla cedrorum (Passeriformes: Bombycillidae), Cisticola aberrans (Passeriformes: Cisticolidae)

Sternostoma hirundo Fain, Herin et Puylaert

Sternostoma hirundo Fain, Herin & Puylaert, 1977: 389

Type locality: Chirundu (South Africa); paratypes from Cameroon and Rwanda

Distribution: Cameroon, Rwanda, South Africa

Hosts: Hirundo aethiopica, H. rustica, H. smithi (Passeriformes: Hirundinidae)

Sternostoma hutsoni Furman

Sternostoma hutsoni Furman, 1957: 477; Fain & Aitken, 1967: 17; 1971: 28; Spicer, 1984: 799

Type locality: Buttowillow, Kern County, California

Distribution: Brazil, Guatemala, Trinidad, USA (California)

Hosts: Catharus dryas, Hylocichla u. ustulata, Turdus albicollis, T. nudigenis, T. rufiventris (Passeriformes: Turdidae)

Sternostoma hylandi Fain et Johnston

Sternostoma hylandi Fain & Johnston, 1966: 32; Shumilo & Lunkashu, 1970: 27; Fain, Sixl & Moritsch, 1974: 6; Butenko, 1984: 148; Knee, Proctor & Galloway, 2008: 370

Type locality: 5 km S of Wooster, Ohio

Distribution: Austria, Canada, Russia, USA (Ohio)

Hosts: Dendrocopos leucotos, D. major, D. [Picoides] pubescens (Piciformes: Picidae)

Sternostoma inflatum Fain

Sternostoma inflatum Fain, 1963c: 83

Type locality: Akanyaru

Distribution: Rwanda

Host: Turdoides jardinei emini (Passeriformes: Timaliidae)

Sternostoma isabelae Ubuda – Ontíveros et Guevara – Benítez

Sternostoma isabelae Ubuda -Ontíveros & Guevara – Benítez, 1980: 366

Type locality: Moraleda de Zafayona (Granada)

Distribution: Spain

Host: Calandrella cinerea (Passeriformes: Alaudidae)

Sternostoma kelloggi Hyland et Clark

Sternostoma kelloggi Hyland & Clark, 1959: 223; Černy & Dusbábek, 1970: 485; Pence, 1972a: 781; 1975: 22

Type locality: Kalamazoo County, Michigan

Distribution: Cuba, USA (Louisiana, Michigan)

Hosts: Dumetella carolinensis, Mimus polyglottos orpheus, Toxostoma rufum (Passeriformes: Mimidae)

Sternostoma kodrensis Shumilo et Lunkashu

Sternostoma kodrensis Shumilo & Lunkashu, 1970: 25

Type locality: Shepot and Tarashani Villages

Distribution: Ukraine

Host: Troglodytes troglodytes (Passeriformes: Troglodytidae)

Sternostoma lagonostictae Fain

Sternostoma lagonostictae Fain, 1956b: 155; 1957b: 72

Type locality: Akanyaru

Distribution: Rwanda

Host: Lagonosticta rubricata congica (Passeriformes: Estrildidae)

Sternostoma longisetosae Hyland

Sternostoma longisetosae Hyland, 1961: 280; Hyland & Moorhouse, 1970: 56; Knee, Proctor & Galloway, 2008: 369

Sternostoma callithrix Fain & Aitken, 1970: 348 – Type from Empidonomus varius, Ananindeua, Pará, Brazil (syn. by Pence & Casto, 1975b)

Type locality: Michigan (USA)

Distribution: Brazil, Canada, Mexico, USA

Hosts: Empidonax alnorum, E. traillii, E. vireonis, Muscivora tyrannus, Pyrocephalus rubinus, Pitangus sulphuratus, Tyrannus tyrannus (Passeriformes: Tyrannidae)

Sternostoma loxiae Fain

Sternostoma loxiae Fain, 1965: 159; 1966: 120; Knee & Proctor, 2010

Type locality: Belgium

Distribution: Belgium, Canada, England

Hosts: Loxia curvirostra (Passeriformes: Fringillidae), Sialia currucoides (Passeriformes: Turdidae)

Sternostoma marchae Dimov

Sternostoma marchae Dimov, 2012: 225; 2018: 59

Type locality: Saint Petersburg, Leningradskaya Obl.

Distribution: Russia

Host: cage Serinus canaria (Passeriformes: Fringillidae)

Sternostoma minutus (Bregetova)

Sternostomum minutus Bregetova, 1950: 1007

Type locality: Vostochniy Murman, Zapovednik “Sem ostrovov”

Distribution: Russia

Host: Charadrius hiaticula (Charadriiformes: Charadriidae)

Sternostoma mortelmansi Fain et Bastin

Sternostoma mortelmansi Fain & Bastin, 1959: 116

Type locality: Anvers Zoo

Distribution: Brazil

Host: Apodiformes – Trochilidae gen. sp.

Sternostoma nectarinia Fain

Sternostoma nectarinia Fain, 1956a: 152; 1957b: 68; Strandtmann, 1960: 135; Fain, Herin & Puylaert, 1977: 392

Type locality: Astrida (= Butare), Rwanda

Distribution: Cameroon, DR Congo, Rwanda, Thailand

Hosts: Chalcomitra senegalensis aequatorialis, Cinnyris cupreus septentrionalis, Dicaeum concolor olivaceum, Nectarinia pulchella, N. purpureiventris (Passeriformes: Nectariniidae)

Sternostoma neosittae Domrow

Sternostoma sp. Domrow, 1967a: 217

Sternostoma neosittae Domrow, 1969b: 377

Type locality: Kowanyama (Queensland)

Distribution: Australia

Host: Neositta striata (Passeriformes: Neosittidae)

Sternostoma numerovi (Butenko)(new comb.)

Sternostomoides numerovi Butenko, 1999: 49

Type locality: Guinea

Distribution: Guinea

Host: Eremopteryx leucotis (Passeriformes: Alaudidae)

Sternostoma opistaspis Feider et Mironescu

Sternostoma opistaspis Feider & Mironescu, 1982: 87

Type locality: Papua New Guinea

Distribution: Papua New Guinea

Host: Cracticus cassicus (Passeriformes: Cracticidae)

Sternostoma paddae Fain

Sternostoma paddae Fain, 1958a: 8; Fain & Bafort, 1963a: 3; Domrow, 1969: 379

Type locality: Antverp Zoo

Distribution: South Asia

Host: Padda oryzivora (Passeriformes: Estrildidae)

Sternostoma pastor Fain

Sternostoma pastor Fain, 1967: 152; Feider & Mironescu, 1978: 265

Type locality: Antverp Zoo

Distribution: Belgium, Romania

Host: Sturnus roseus (Passeriformes: Sturnidae)

Sternostoma pencei Spicer

Sternostoma pencei Spicer, 1984: 797

Type locality: Baja Verapaz, 2 km E, 2.4 km S Purulhá

Distribution: Guatemala, USA (Missouri)

Host: Empidonax flavescens (Passeriformes: Tyrannidae)

Sternostoma piprae (Fain et Aitken)

Sternoecius piprae Fain & Aitken, 1967: 24; 1968: 3

Sternostoma piprae (Fain et Aitken): Domrow, 1969: 374

Type locality: Corneillac Estate, Vega de Oropouche (Trinidad)

Distribution: Brazil, Trinidad

Hosts: Manacus manacus, Pipra erythrocephala (Passeriformes: Pipridae)

Sternostoma pirangae Pence

Sternostoma pirangae Pence, 1973b: 874; Spicer, 1984: 799; Mascarenhas, Coimbra et al., 2011: 507

Sternostoma tangarae Fain & Aitken (in part), Pence, 1972: 785

Type locality: Mandeville. La.

Distribution: Brazil, Guatemala, USA (Louisiana, Texas)

Hosts: Piranga rubra (Passeriformes: Cardinalidae), Chlorospingus ophthalmicus (Passeriformes: Thraupidae), Paroaria coronata (Passeriformes: Emberizidae)

Sternostoma porteri Hyland

Sternostoma porteri Hyland, 1963: 149; Fain & Johnston, 1966: 31; Pence, 1972: 786; 1973: 886; Pence & Casto, 1975b: 362; Butenko, 1984: 145; Knee, Proctor & Galloway, 2008: 370

Type locality: Kalamazoo Co., Michigan

Distribution: Azerbajdjan, Canada, Ukraine, USA (Michigan, Ohio, Texas)

Hosts: Centurus aurifrons, Colaptes auratus, Picoides pubescens, P. villosus, Picus viridis, Sphyrapicus varius (Piciformes: Picidae)

Sternostoma quiscali Fain et Aitken

Sternostoma quiscali Fain & Aitken, 1967: 18; Černy & Dusbábek, 1970: 485

Type locality: El Socorro Estate Road, La Paille Village, San Juan

Distribution: Cuba, Trinidad

Hosts: Quiscalus lugubris, Qu. niger caribaeus (Passeriformes: Icteridae)

Sternostoma sayornis Pence et Casto

Sternostoma sayornis Pence & Casto, 1975b: 366

Type locality: Texas, Presidio Co., Candelaria

Distribution: USA (Texas)

Host: Sayornis nigricans (Passeriformes: Tyrannidae)

Sternostoma schiffornis Fain et Aitken

Sternostoma schiffornis Fain & Aitken, 1970: 343

Type locality: Mosqueiro Ferry, Benavides, Belém

Distribution: Brazil

Host: Schiffornis turdinus (Passeriformes: Pipridae)

Sternostoma setifer Knee

Sternostoma setifer Knee, 2008: 367

Type locality: Delta Marsh, Manitoba

Distribution: Canada

Host: Empidonax minimus (Passeriformes: Tyrannidae)

Sternostoma sialiphilus Hyland et Ford

Sternostoma sialiphilus Hyland & Ford, 1960: 101; Knee, Proctor & Galloway, 2008: 369

Type locality: Kalamazoo County, Michigan

Distribution: Canada, USA (Michigan)

Hosts: Sialia sialis (Passeriformes: Turdidae), Riparia riparia (Passeriformes: Hirundinidae)

Sternostoma sinense Fain et Bafort

Sternostoma sinense Fain & Bafort, 1963c: 7; Fain, Herin & Puylaert, 1977: 392

Type locality: Anvers Zoo

Distribution: Cameroon; type comes from Asian bird (Leiothrix lutea)

Hosts: Phoeniculus purpureus (Coraciiformes: Phoeniculidae), Quelea erythropus, Q. quelea (Passeriformes: Ploceidae), Leiothrix lutea (Passeriformes: Timaliidae)

Sternostoma sternahirundo Butenko

Sternostoma sternahirundo Butenko, 1974: 1090; 1984: 142; De Rojas et al., 961

Type locality: Novosibirskaya Obl., Zdvinskiy rayon, Malie Chany Lake (Russia)

Distribution: Russia, Spain

Hosts: Sterna hirundo (Charadriiformes: Sternidae), Larus argentatus (Charadriiformes: Laridae)

Sternostoma straeleni Fain

Sternostoma straeleni Fain, 1958c: 9

Type locality: Vitshumbi (Virunga Nat. Park)

Distribution: D.R. Congo

Host: Cinnyricinclus leucogaster (Passeriformes: Sturnidae)

Sternostoma strandtmanni Furman

Sternostoma strandtmanni Furman, 1957: 476; Fain & Aitken, 1967: 18; 1971: 28; de Rojas, Mora et al., 2002: 676

Type locality: California

Distribution: Brazil, Spain, Trinidad, USA

Hosts: Agelaius icterocephalus, A. tricolor, Icterus nigrogularis, Chrysomus ruficapillus, Leistes militaris, Molothrus sp., Quiscalus lugubris (Passeriformes: Icteridae); Tringa totanus (Charadriiformas: Scolopacidae)

Sternostoma strigitis Butenko

Sternostoma strigitis Butenko, 1976: 303; 1984: 143

Type locality: Ryazanskaya Obl.

Distribution: Russia

Host: Asio otus (Strigiformes: Strigidae)

Sternostoma sturnicola Fain

Sternostoma sturnicola Fain, 1956a: 151; 1957b: 65; 1959a: 31; 1961b: 52; Strandtmann & Wharton, 1958: 168; Fain, Herin & Puylaert, 1977: 392

Type locality: Astrida (= Butare), Rwanda

Distribution: Cameroon, Botswana, Rwanda

Hosts: Buphagus africanus (Passeriformes: Buphagidae), Lamprocolius chalybaeus, L. purpureus, Lamprotornis purpuropterus (Passeriformes: Sturnidae)

Sternostoma tangarae Fain et Aitken

Sternostoma tangarae Fain & Aitken, 1967: 20; Pence, 1972b: 785

Type locality: La Fortune Estate, Vega de Oropouche

Distribution: Trinidad, USA

Hosts: Piranga rubra (Passeriformes: Cardinalidae), Tangara mexicana (Passeriformes: Thraupidae)

Sternostoma technaui (Vitzthum)

Sternostomum technaui Vitzthum, 1935: 569

Sternostoma technaui: Zumpt & Till, 1955: 91; Fain, 1957b: 62; Furman, 1957: 482; Strandtmann & Wharton, 1958: 168; Domrow, 1969b: 379; Pence, 1972b: 786; Sartbaev, 1983: 84; Kadosaka, Kaneko & Asanuma, 1987: 38; Knee, Proctor & Galloway, 2008: 370

Sternostomoides technaui (Vitzthum): Bregetova, 1965: 709

Rhinonyssus technaui: Castro, 1948: 257

Sternostoma turdi Zumpt & Till, 1955: 85; Fain, 1956: 151; 1957b: 64; 1959a: 31; 1962: 264; 1963: 176; Furman, 1957: 481; Fain & Aitken, 1967: 17; Fain, Sixl & Moritsch, 1974: 5; Úbeda-Ontíveros & Guevara-Benítez, 1979: 291; de Rojas et al., 2001: 961; de Rojas, Mora et al., 2002: 676. – Type: South Africa. Syn. by Domrow, 1969: 380

Sternostomoides turdi (Zumpt et Till): Bregetova, 1965: 712; Shumilo & Lunkashu, 1970: 23

Sternostoma spatulatum Furman, 1957: 480 – type: Buttowillow, Kern County, California, from Hylocichla u. ustulata (Turdidae) ; Knee & Galloway, 2017: 95.

Syn. by Domrow, 1969: 380

Sternostomoides spatulatum (Furman): Bregetova, 1965: 712

Sternostomoides spatulatus (Furman): Černy & Dusbábek, 1970: 486

Sternostoma borceanum Feider & Mironescu, 1968: 105. Type: Romania, from Turdus ericetorum, T. pilaris, T. torcuatus (syn. by Fain, Sixl & Moritsch, 1974: 6); Feider & Mironescu, 1978: 267, from Sturnus vulgaris, Sf. Ghorghe, Tulcea

Sternostomoides orlandoi Dusbábek, 1969: 225; Černy & Dusbábek, 1970: 486 –

Type locality: Santa Fé, Prov. of Havana, Cuba, from Mimocichla plumbea rubripes, M. plumbea schistacea (Passeriformes: Turdidae). Syn. by Kadosaka, Kaneko & Asanuma, 1987: 38

Type locality of Sternostoma technaui: Europe

Distribution: Australia, Austria, Belgium, Brazil, Canada, Germany, Japan, Kirghizstan, Moldavia, Romania, Russia, Rwanda, South Africa, Spain, Tadjikistan, Trinidad, Ukraine, USA

Hosts: Cinclus cinclus aquaticus, C. pallasi (Passeriformes: Cinclidae), Hylocichla u. ustulata, Monticola saxatilis, Oreocincla dauma, O. lunulata, Turdus abyssinicus baraka, T. ericetorum, T. fumigatus, T. Iliacus, T. libonyanus, T. merula, T. migratorius, T. olivaceus, T. olivaceus graueri, T. philomelos, T. pilaris, T. ruficollis, T. viscivorus (Passeriformes: Turdidae), Sturnus vulgaris (Passeriformes: Sturnidae)

Sternostoma thienponti Fain

Sternostoma thienponti Fain, 1956a: 152; 1957b: 68; 1960: 111; Strandtmann & Wharton, 1958: 168; Domrow, 1965b: 449; 1966b: 194; 1969b: 375

Sternostoma thieponti [sic] Fain: Strandtmann, 1960: 135

Type locality: Astrida (= Butare), Rwanda

Distribution: Australia, Botswana, Rwanda, Thailand

Hosts: Dicrurus adsimilis, D. macrocercus cathoecus (Passeriformes: Dicruridae), Cracticus nigrogularis, C. quoyi (Passeriformes: Cracticidae), Gymnorhina tibicen (Passeriformes: Artamidae)

Sternostoma tracheacolum Lawrence

Sternostoma tracheacolum Lawrence, 1948: 366; Zumpt & Till, 1955: 86; Fain, 1956a: 152; 1957b: 65; Strandtmann & Wharton, 1958: 168; Strandtmann, 1960: 137; Fain & Hyland, 1962: 404; Amaral, 1962: 72; Murray, 1966: 262; Domrow, 1966a: 194; 1969b: 377, 423; Mathey, 1967: 777; Shumilo & Lunkashu, 1970: 26; Hyland & Moorhouse, 1970: 56; Fain & Aitken, 1971: 28; Pence, 1972b: 785; McClure, Ratanaworadhan et al., 1973: 37; Fain, Sixl & Moritsch, 1974: 5; Blasco Sabio & Portus Vinyeta, 1974: 37; Guevara-Benítez & Úbeda-Ontíveros, 1974: 176; Knee & Proctor, 2010; Knee & Galloway, 2017: 94

Sternostoma meddai Lombardini, 1953: 187 – Italy (syn. by Fain & Hyland, 1962)

Sternostoma castroae Fain, 1956b: 393 – Rwanda, ex Macronyx croceus (syn. by Fain, 1957)

Agapornyssus faini Grétillat, Capron & Brygoo, 1959: 375 – Madagascar, ex Agapornis cana (syn. by Fain & Hyland, 1962)

Type locality: Pietermaritzburg (South Africa)

Distribution: Australia, Austria, Belgium, Brazil, Canada, DR Congo, France, Great Britain, Italy (incl. Sardinia), Kirghizstan, Madagascar, Mexico, Moldavia, Russia, South Africa, Spain, Thailand, Trinidad, Uruguay, USA

Hosts: Coereba flaveola (Passeriformes: Coerebidae), Chloebia [Poephila] gouldiae, (Passeriformes: Estrildidae), Arachnothera longirostris, Nectarinia jugularis flammaxillaris (Nectariniidae), Sylvia melanocephala (Sylviidae), Chloris chloris, Coccothraustes coccothraustes, Serinus canaria (Fringillidae), Hirundo rustica, Iridoprocne bicolor, Riparia riparia (Hirundinidae), Dendrocincla merula (Furnariidae, Dendrocolaptinae), Cinnyricinclus leucogaster verreauxi (Sturnidae), Myiarchus crinitus, Rhynchocyclus olivaceus, Tyrannus melancholicus (Tyrannidae), Seiurus noveboracensis (Parulidae), Parus major (Paridae), Molothrus ater, Sturnella magna (Icteridae), Thraupidae, Melopsittacus undulatus (Psittaciformes: Psittacidae), Macronyx croceus (Passeriformes: Motacillidae), Melospiza melodia, Pipilo fuscus, Spizella pusilla, Sturnella magna (Passeriformes: Emberizidae), Cardinalis [Pyrrhuloxia] sinuatus, Passerina cyanea (Passeriformes: Cardinalidae), Acrocephalus arundinaceus (Passeriformes: Acrocephalidae)

Sternostoma tyrannus Brooks et Strandtmann

Sternostoma tyrannus Brooks & Strandtmann, 1960: 418; Pence, 1975: 23

Type locality: Lamb County, Texas

Distribution: USA (Texas)

Host: Nuttallornis borealis (Passeriformes: Tyrannidae)

Sternostoma ubedai Úbeda – Ontíveros et Guevara – Benítez

Sternostoma ubedai Úbeda – Ontíveros & Guevara – Benítez, 1981: 176

Type locality: Prov. de Granada

Distribution: Spain

Host: Sylvia melanocephala (Passeriformes: Sylviidae)

Sternostoma ziegleri Feider et Mironescu

Sternostoma ziegleri Feider & Mironescu, 1982: 87

Type locality: Papua New Guinea

Distribution: Papua New Guinea

Host: Cacomantis variolosus (Cuculiformes: Cuculidae)

Sternostoma zini Dimov et Knee

Sternostoma zini Dimov & Knee, 2012: 139

Type locality: Viritsa, Leningrad Province

Distribution: Russia

Host: Cuculus canorus (Cuculiformes: Cuculidae)

Genus Tinaminyssus Strandtmann et Wharton

Tinaminyssus Strandtmann & Wharton, 1958: 161

Type species: Neonyssus (Ptilonyssoides) trappi Peraira et de Castro, 1949: 229

Mesonyssus Fain, 1960: 313 (syn. by Domrow, 1969: 305)

Type species: Neonyssus treronis Fain, 1956: 394

Mesonyssoides Fain & Nadchatram, 1962: 271 (syn. by Domrow, 1969: 305)

Type species: Mesonyssoides malayi Fain & Nadchatram, 1962: 272

Mesonyssoides Strandtmann & Clifford, 1962: 723 (syn. with Mesonyssus by Wilson, 1964b: 357; with Tinaminyssus by Domrow, 1969: 305)

Type species: Mesonyssoides ixoreus Strandtmann & Clifford, 1962: 723

Psittonyssus Fain, 1963: 69 (syn. by Domrow, 1969: 305)

Type species: Psittonyssus baforti Fain, 1963: 70

Falconyssus Fain, 1966: 85 (syn. by Domrow, 1969: 305)

Type species: Falconyssus elani Fain, 1966: 86

Tinaminyssus alexfaini (Amaral)(new comb.)

Mesonyssus alexfaini Amaral, 1967: 135

Type locality: Itaporanga, Est. de São Paulo

Distribution: Brazil

Host: Leptotila verreauxi (Columbiformes: Columbidae)

Tinaminyssus alisteri (Wilson)(new comb.)

Mesonyssus alisteri Wilson, 1964b: 370

Type locality: Morobe District, Wau, Bulolo River

Distribution: Papua New Guinea

Host: «king parrot« (Psittaciformes)

Tinaminyssus angustus (Wilson)

Mesonyssus angustus Wilson, 1966: 607

Tinaminyssus angustus (Wilson): Domrow

Type locality: Morobe District: Wau: Bulolo

Distribution: Papua New Guinea

Tinaminyssus aprosmicti (Domrow)

Mesonyssoides aprosmicti Domrow, 1964b: 26; Fain & Lukoschus, 1979: 20

Mesonyssoides platycerci Domrow, 1964b: 27 (syn. by Domrow, 1969: 306)

Mesonyssus aposmicti (Domrow): Domrow, 1967a: 217; Wilson, 1968: 395

Tinaminyssus aprosmicti Domrow, 1969b: 306; 1979d: 488; 1987: 883

Type locality: Condamine, Kowanyama

Distribution: Australia

Hosts: Aprosmictus erythropterus, A. scapularis, Barnardius barnardi, Platycercus adscitus, Psephotus haematonotus, P. varius (Psittaciformes: Psittaculidae)

Tinaminyssus baforti (Fain)

Psittonyssus baforti Fain, 1963: 70

Tinaminyssus baforti (Fain): Domrow

Type locality: Avicentra (Anvers)

Distribution: East Africa

Host: Psittacula baforti (Psittaciformes: Psittaculidae)

Tinaminyssus belopolskii (Bregetova)

Neonyssus belopolskii Bregetova, 1950: 1005; 1951: 119; 1953: 332; Fain, 1957b: 53; Strandtmann & Wharton, 1958: 162; Mitchell, 1961: 105

Neonyssus ardeae Zumpt & Till, 1955: 63 – type: Benoni, Transvaal, from Ardea melanocephala; Fain, 1956a: 133; 1957b: 137; 1959a: 33 (syn. by Strandtmann, 1956: 137)

Neonyssus belepolskii: Strandtmann, 1956: 137

Mesonyssus belopolskii Fain, 1961a: 515; 1962d: 306; Domrow, 1965b: 433; 1966b: 191; 1967a: 216; Wilson, 1968a: 213; Černy & Dusbábek, 1970: 486; Butenko, 1984: 91

Mesonyssus belopolskii nycticoracis Fain, 1961a: 516; Butenko, 1984: 93. – Type: Astrida (= Butare) Rwanda, from Nycticorax nycticorax (syn. by Domrow, 1969)

Neonyssus ixobrychi Fain, 1956a: 134; Butenko, 1965: 131. – Type: Astrida (= Butare), Rwanda, from Ixobrychus minutus (syn. by Domrow, 1969: 311, but restaured by Butenko, 1984)

Neonyssus marcandrei Gretillat, Capron & Brygoo, 1959: 379 – Type: Lac Itasy, Madagascar, from Melanophyx ardesiaea (Ardeidae). Syn. by Fain, 1961a: 516.

Mesonyssus minuti Feider & Mironescu, 1974: 764 – type: Iasi, Ciric (syn. by Domrow, 1979: 513)

Tinaminyssus belopolskii Domrow, 1969b: 311; 1987: 883; Pence, 1972a: 162; Bernardon, Müller & Mascarenhas, 2013: 674.

Type locality: Russia (Sudzuhinskiy Goszapovednik)

Distribution: Australia, Botswana, Brazil, Cuba, Madagascar, Philippines (Mindanao), Russia, Rwanda, South Africa, Spain, Canary Isl., USA

Hosts: Ardea cinerea, A. melanocephala, A. purpurea manillensis, Bubulcus ibis, Butorides striata, B. virescens, Egretta garzetta, E. thula thula, Florida caerulea, Hydranassa tricolor, Ixobrychus minutus, Leucophoyx thula, Melanophyx ardesiaea, Notophoyx novaehollandiae, N. picata, Nycticorax nycticorax, Syrigma sibilatrix (Pelicaniformes: Ardeidae)

Tinaminyssus bubulci (Zumpt et Till)

Neonyssus bubulci Zumpt & Till, 1955: 66; Fain, 1956a: 134; 1957: 54; 1959: 33; 1960: 108; Černy & Dusbábek, 1970: 486 (Cuba). – Type: Johannesburg, South Africa, from Bubulcus ibis Syn. by Domrow (1969), but revalidated by Pence (1972)

Mesonyssus bubulci (Zumpt et Till): Černy & Dusbábek, 1970: 486 (Cuba); Butenko, 1984: 99

Tinaminyssus bubulci (Zumpt et Till): Pence, 1972: 160; 1973: 883; Rodríguez-Braza, Guevara-Benítez & Úbeda-Ontíveros, 1990: 73; de Rojas et al., 2002

Type locality: Johannesburg (South Africa)

Distribution: Cuba, South Africa

Hosts: Bubulcus ibis, Egretta intermedia brachyrhyncha (Pelecaniformes: Ardeidae)

Tinaminyssus buteonis (Fain)(new comb.)

Neonyssus buteonis Fain, 1956a: 135; 1957b: 51; 1959a: 32

Mesonyssus buteonis Fain, 1962d: 306

Falconyssus buteonis (Fain): Fain, 1966: 87

Type locality: Astrida (= Butare), Rwanda

Distribution: Rwanda, South Africa

Hosts: Buteo rufofuscus augur, Lophaetus occipitalis (Accipitriformes: Accipitridae); Coracias caudata (Coraciiformes: Coraciidae)

Tinaminyssus camargoi (Amaral) (new comb.)

Mesonyssus camargoi Amaral, 1962: 145

Type locality: Pirapora, Est. de São Paulo

Distribution: Brazil

Host: Penelope sp. (Galliformes: Cracidae)

Tinaminyssus carapachibeyus Dusbábek

Tinaminyssus carapachibeyus Dusbábek, 1969: 216; Černy & Dusbábek, 1970: 487

Type locality: Carapachibey, Isla de Pinos

Distribution: Cuba (Isla de Pinos)

Host: Geotrygon chrysia (Columbiformes: Columbidae)

Tinaminyssus castroae (Amaral)(new comb.)

Neonyssus castroae Amaral, 1963: 9

Type locality: Itaporanga, Est. de São Paulo

Distribution: Brazil

Hosts: Oreopeleia montana and Leptotila verreauxi (Columbiformes: Columbidae)

Tinaminyssus chiarellii Amaral et Baquer

Tinaminyssus chiarellii Amaral & Baquer, 1963: 65; Hyland & Moorhouse, 1970: 54

Type locality: Zoo in São Paulo

Distribution: Brazil, Mexico

Hosts: Cyanocorax chrysops chrysops (Passeriformes: Corvidae), Pitangus sulfuratus (Passeriformes: Tyrannidae)

Tinaminyssus columbae (Crossley)

Neonyssus (Neonyssus) columbae Crossley, 1950: 309

Neonyssus columbae Crossley: Zumpt & Till, 1955: 67; Fain, 1956a: 134; Strandtmann & Wharton, 1958: 162

Mesonyssus columbae (Crossley): Fain, 1962: 306; Foulk & Matthysse, 1965: 127; Domrow, 1965b: 440; Wilson, 1964: 368, 1966b: 1211; Černy, 1970: 244; Fain, Sixl & Moritsch, 1974: 7; Butenko, 1969: 208, 1984: 114; Dimov, 2016; 2018: 177

Tinaminyssus columbae (Crossley): Domrow, 1969b: 313; 1987: 884; Pence, 1975: 21; Kadosaka, Kaneko & Asanuma, 1987; Úbeda et al., 2003: 68; Knee, Proctor & Galloway, 2008: 368

Type locality: Alabama

Distribution: Australia, Austria, Belgium, Canada, Czech Rep., Japan, Russia, Rwanda, Spain, Ukraine, USA (Alabama, Georgia, Kentucky)

Hosts: Columba guinea guinea, C. livia, C. livia domestica, Ocyphaps lophotes, Streptopelia o. orientalis, S. semitorquata, Zenaida macroura (Columbiformes: Columbidae), Milvus migrans, M. tenebrosus (Accipitriformes: Accipitridae)

Tinaminyssus crypturelli Fain

Tinaminyssus crypturelli Fain, 1967: 153

Type locality: Anvers Zoo

Distribution: South America

Host: Crypturellus tataupa (Tinamiformes: Tinamidae)

Tinaminyssus cunhai (Amaral)(new comb.)

Mesonyssus cunhai Amaral, 1963: 103

Type locality: Itaporanga, Est. de São Paulo

Distribution: Brazil

Host: Leptotila verreauxi (Columbiformes: Columbidae)

Tinaminyssus daceloae (Domrow)

Mesonyssus daceloae Domrow, 1965b: 445

Falconyssus daceloae (Domrow): Fain, 1966: 87

Tinaminyssus daceloae Domrow, 1969b: 322; 1979d: 492; 1983: 884

Type locality: Samford, Esk

Distribution: Australia

Hosts: Dacelo gigas, D. leachii (Coraciiformes: Halcyonidae)

Tinaminyssus egrettae (Butenko)(new comb.)

Mesonyssus egrettae Butenko, 1984: 96

Type locality: Krasnodarskiy Kray, Primorsko-Ahtarskiy Rayon

Distribution: Russia

Host: Egretta garzetta (Pelicaniformes: Ardeidae)

Tinaminyssus elani (Fain)(new comb.)

Falconyssus elani Fain, 1966c: 86

Type locality: Luluabourg (now Kananga)

Distribution: DR Congo

Host: Elanus coeruleus (Accipitriformes: Accipitridae)

Tinaminyssus epileus (Wilson)

Mesonyssus epileus Wilson, 1964b: 358; 1965: 638; 1968a: 213; Tenorio, 1976

Tinaminyssus epileus (Wilson): Domrow, 1969: 312

Type locality: Finschhafen

Distribution: Papua New Guinea

Host: Aviceda subcristata (Accipitriformes: Accipitridae)

Tinaminyssus geopeliae (Fain)

Mesonyssus geopeliae Fain, 1964b: 33; Domrow, 1965b: 440; 1966b: 192; Wilson, 1966b: 1211; Fain & Lukoschus, 1979: 20

Tinaminyssus geopeliae Domrow, 1969b: 315, 1979d: 490; 1987: 884; McClure, Ratanaworadhan et al., 1973; 35

Type locality: Anvers Zoo (Malaysia)

Distribution: Australia, Hawaii, Malaysia, Papua New Guinea, Philipppines

Hosts: Geopelia cuneata, G. humeralis, G. placida, G. striata striata (Columbiformes: Columbidae)

Tinaminyssus geotrygoni (Dusbábek)

Mesonyssus geotrygoni Dusbábek, 1969: 218; Černy & Dusbábek, 1970: 487

Tinaminyssus geotrygoni: Pence, 1975: 21

Type locality: Carapachibey, Isla de Pinos

Distribution: Cuba (Pinos)

Host: Geotrygon chrysia (Columbiformes: Columbidae)

Tinaminyssus gerschi (Feider et Mironescu) (new comb.)

Mesonyssus gerschi Feider & Mironescu, 1974: 752

Type locality: Delta of Danube

Distribution: Romania

Host: Ardea purpurea (Pelecaniformes: Ardeidae)

Tinaminyssus gourae (Wilson)(new comb.)

Mesonyssus gourae Wilson, 1964b: 361

Type locality: NW New Guinea, Boemi River

Distribution: Papua New Guinea

Hosts: Ducula pinon, Goura cristata (Columbiformes: Columbidae)

Tinaminyssus halcyonus (Domrow)

Mesonyssus halcyonus Domrow, 1965b: 443; Wilson, 1966a: 609; 1968a: 213

Falconyssus halcyonus: Fain, 1966: 86

Tinaminyssus halcyonus Domrow, 1969b: 321, 1972: 99; 1979: 492; 1987: 884; McClure, Ratanaworadhan et al., 1973: 35

Type locality: Logan Village, Brisbane

Distribution: Australia, New Zealand, Papua New Guinea, Philippines

Hosts: Halcyon chloris collaris, H. macleayii, H. pyrrhopygius, H. sanctus (Coraciiformes: Halcyonidae)

Tinaminyssus hirtus (Wilson)

Mesonyssus hirtus Wilson, 1966b: 1210; 1968a: 215

Tinaminyssus hirtus Domrow, 1969b: 313; 1987: 884; McClure, Ratanaworadhan et al., 1973: 35

Type locality: Siaton, Kandugay, Negros Oriental, Philippines

Distribution: Australia, Papua New Guinea, Philippines

Hosts: Chalcophaps chrysochlora, Ch. indica indica, Ch. stephani (Columbiformes: Columbidae)

Tinaminyssus ixobrychi (Fain)

Neonyssus ixobrychi Fain, 1956a: 154; 1957b: 54; Butenko, 1965: 131

Mesonychus ixobrychi (Fain): Feider & Mironescu, 1974: 751; Butenko, 1984: 94

Tinaminyssus ixobrychi (Fain): Domrow, 1969: 311

Mesonyssus minuti Feder & Mironescu, 1974: 764 (syn. by Butenko, 1984: 95)

Type locality: Astrida [Butare], Rwanda

Distribution: Moldavia, Kirghizstan, Rwanda, Russia, South Africa

Host: Ixobrychus minutus (Pelicaniformes: Ardeidae)

Tinaminyssus ixoreus (Strandtmann et Clifford)

Mesonyssoides ixoreus Strandtmann & Clifford, 1962: 723; Isakova, 1965

Tinaminyssus ixoreus (Strandtmann et Clifford): Pence, 1975: 21; Spicer, 1984: 800

Type locality: Lost Horse Canyon, Ravalli County, Montana

Distribution: Canada, Guatemala, Russia, USA (Montana)

Hosts: Catharus dryas, Ixoreus naevius (Passeriformes: Turdidae)

Tinaminyssus juxtamelloi Pence et Canaris

Tinaminyssus juxtamelloi Pence & Canaris, 1976: 116

Type locality: Mimbres, Luna Co., New Mexico

Distribution: USA (New Mexico)

Host: Columba fasciata (Columbiformes: Columbidae)

Tinaminyssus kakatuae (Domrow)

Mesonyssoides kakatuae Domrow, 1964b: 25

Mesonyssus kakatuae Domrow, 1966b: 191; 1967a: 217; Wilson, 1966: 759

Tinaminyssus kakatuae Domrow, 1969b: 306; 1987: 885

Type locality: Kowanyama, Condamine

Distribution: Australia

Hosts: Calyptorhynchus banksi, Cacatua roseicapilla, C. sulphurea occidentalis, Nymphicus [Leptolophus] hollandicus (Psittaciformes: Cacatuidae)

Tinaminyssus macropygiae (Wilson)

Mesonyssus macropygiae Wilson, 1966a: 607; 1968b: 215

Tinaminyssus macropygiae Domrow, 1969b: 319;1987: 885

Type locality: NW New Guinea, Oransbari

Distribution: Australia, Papua New Guinea, Philippines

Hosts: Macropygia amboinensis, M. phasianella (Columbiformes: Columbidae)

Tinaminyssus malayi (Fain et Nadchatram)

Mesonyssoides malayi Fain & Nadchatram, 1962: 272

Tinaminyssus malayi (Fain et Nadchatram): McClure, Ratanaworadhan et al., 1973: 35

Type locality: Mont Brinchang (1700 m), Pahang

Distribution: Malaysia

Host: Cyanops (= Megalaima) franklini minor (Piciformes: Capitonidae)

Tinaminyssus marciae (Amaral)(new comb.)

Mesonyssus marciae Amaral, 1966: 153

Type locality: Boracéa, São Paulo

Distribution: Brazil

Host: Leuchocloris albicolis (Apodiformes: Trochilidae)

Tinaminyssus megaloprepiae Domrow

Tinaminyssus megaloprepiae Domrow, 1969b: 313; 1987: 885

Type locality: Maalan

Distribution: Australia

Host: Megaloprepia magnifica (Columbiformes: Columbidae)

Tinaminyssus melloi (de Castro)

Neonyssus (Neonyssus) melloi de Castro, 1948: 270

Mesonyssus melloi (de Castro): Fain, 1962d: 306; Domrow, 1965b: 434; 1966b: 192; Amaral, 1968: 122; Butenko, 1969; Shumilo & Lunkashu, 1970: 16; Fain, Sixl & Moritsch, 1974: 7; Sartbaev, 1982: 131; Rodriguez-Braza, 1987: 175C; Dimov, 2018: 174

Neonyssus hirsutus Feider, 1962: 58. – Type: Romania, from Columba livia (syn. by Domrow, 1969: 312)

Neonyssus melloi: Zumpt & Till, 1955: 68; Fain, 1957b: 137; 1959a: 33; Strandtmann & Wharton, 1958: 163

Tinaminyssus melloi: Domrow, 1969b: 312 ; 1972: 99; 1987: 885; Zamudio, 1988: 205; Kadosaka, Kaneko & Asanuma, 1987; Úbeda et al., 2003: 66; Knee, Proctor & Galloway, 2008: 368

Type locality: Brazil

Distribution: Australia, Austria, Canary Is., Canada, Czech Rep., Brazil, Hungary, Japan, Kirghizstan, Moldavia, New Zealand, Romania, Russia, South Africa, Spain, Ukraine, USA

Hosts: Columba livia domestica, C. norfolciensis, C. oenas, C. palumbus, Leucosarcia melanoleuca, Stigmatopelis senegalensis, Streptopelia capicola, S. chinensis, S. o. orientalis, Zenaida macroura (Columbiformes: Columbidae), Passer domesticus (Passeriformes: Passeridae)

Tinaminyssus metropeliae (Fain)(new comb.)

Mesonyssus metropeliae Fain, 1972: 361

Type locality: Zoo Anvers

Distribution: Zoo Anvers

Host: Metropelia caeciliae (Columbiformes: Columbidae)

Tinaminyssus milvi (Fain)

Mesonyssus milvi Fain, 1962: 149

Tinaminyssus milvi (Fain): Domrow, 1969: 305

Type locality:

Distribution:

Host: Milvus (Accipitriformes: Accipitridae)

Tinaminyssus minisetosum (Butenko)

Mesonyssus minisetosum Butenko, 1984: 115

Tinaminyssus minisetosum (Butenko): Úbeda et al., 2003: 70

Type locality: Kirghizstan, Alayskaya Dolina (Valley)

Distribution: Kirghizstan, Spain

Host: Columba palumbus (Columbiformes: Columbidae)

Tinaminyssus myristicivorae Domrow

Mesonyssus sp. “A” Domrow, 1967a: 215

Tinaminyssus myristicivorae Domrow, 1969b: 316, 1979d: 490; 1987: 885

Type locality: Kowanyama

Distribution: Australia

Host: Myristicivora spilorrhoa (Columbiformes: Columbidae)

Tinaminyssus navajasi (Pereira et Castro)

Neonyssus (Ptilonyssoides) navajasi Pereira & Castro, 1949: 231

Neonyssus navajasi Pereira et Castro: Zumpt & Till, 1955: 87; Fain, 1957b: 57

Mesonyssus navajasi: Fain, 1962d: 306; Amaral, 1968: 122

Tinaminyssus navajasi: Strandtmann & Wharton, 1958: 161

Type locality: Estados de São Paulo and Mato Grosso

Distribution: Brazil

Host: Nothura maculosa (Tinamiformes: Tinamidae)

Tinaminyssus neoixobrychi Pence

Tinaminyssus neoixobrychi Pence, 1972a: 165; 1975: 22

Type locality: Louisiana, Norco

Distribution: USA

Host: Ixobrychus exilis (Pelicaniformes: Ardeidae)

Tinaminyssus ocyphabus (Domrow)

Mesonyssus ocyphabus Domrow, 1965b: 437

Tinaminyssus ocyphabus Domrow, 1969b: 312; 1987: 885

Type locality: Condamine, Samford

Distribution: Australia

Hosts: Geophaps scripta, Ocyphaps lophotes (Columbiformes: Columbidae).

Tinaminyssus oenae (Fain)(new comb.)

Mesonyssus oenae Fain, 1963c: 61

Type locality: Anvers Zoo

Distribution: South Africa

Host: Oena c. capensis (Columbiformes: Columbidae)

Tinaminyssus peleiae (Wilson)(new comb.)

Mesonyssus pileiae Wilson, 1966a: 609

Type locality: Leyte I., Santa Cruz, Mahaplag

Distribution: Philippines

Host: Phapitreron brevirostris (Columbiformes: Columbidae)

Tinaminyssus phabus (Domrow)

Mesonyssus phabus Domrow, 1965b: 435

Tinaminyssus phabus Domrow, 1969b: 317

Type locality: Esk

Distribution: Australia

Host: Phaps chalcoptera (Columbiformes: Columbidae)

Tinaminyssus phalliger (Fain)(new comb.)

Psittanyssus phalliger Fain, 1965: 160

Type locality: Zoo Anvers

Distribution: Indonesia

Host: Psittacula alexandri (Psittaciformes: Psittaculidae)

Tinaminyssus phassae (Wilson)(new comb.)

Mesonyssus phassae Wilson, 1966: 602

Type locality: Kaohsiung Hsien, Liukuei

Distribution: Taiwan

Host: Sphenurus sieboldii sororius (Columbiformes: Columbidae)

Tinaminyssus prionituri (Wilson)(new comb.)

Mesonyssus prionituri Wilson, 1966: 759; 1968: 397

Type locality: Philippines (Leyte, Palawan)

Distribution: Philippines (Leyte, Palawan)

Hosts: Prioniturus [flavicans] montanus, P. [discurus] platenae, P. discurus whiteheadi (Psittaciformes: Psittaculidae)

Tinaminyssus pteroclesi (Fain)(new comb.)

Mesonyssus pteroclesi Fain, 1963c: 63

Type locality: Lochinvar

Distribution: South Africa

Host: Pterocles gutturalis (Pteroclidiformes: Pteroclididae)

Tinaminyssus ptilinopi (Wilson)

Mesonyssus ptilinopi Wilson, 1964b: 366; 1965: 638; 1966: 1212; 1968: 221; Tenorio, 1976

Tinaminyssus ptilinopi: Domrow, 1969b: 315; McClure, Ratanaworadhan et al., 1973: 36

Type locality: SE New Guinea (Papua), Cape Kileton

Distribution: Australia, Malaysia, Papua New Guinea, Philippines, Taiwan

Hosts: Ptilinopus coronulatus, P. ornatus, P. perlatus, P. superbus (Columbiformes: Columbidae)

Tinaminyssus sartbaevi (Butenko)

Mesonyssus sartbaevi Butenko, 1984: 111

Tinaminyssus sartbaevi (Butenko): Úbeda et al., 2003: 68

Type locality: Kirghizstan, Alayskaya Dolina (Valley)

Distribution: Kirghizstan, Spain

Host: Columba palumbus (Columbiformes: Columbidae)

Tinaminyssus saucerottiae (Fain et Aitken)

Trochilonyssus saucerottiae Fain & Aitken, 1967: 32

Tinaminyssus saucerottiae (Fain et Aitken): Fain & Aitken, 1970: 410

Type locality: La Horquette Trace, Tumpuna Road, Trinidad

Distribution: Trinidad

Hosts: Campylopterus largipennis, Saucerottia tobaci (Apodiformes: Trochilidae)

Tinaminyssus schoutedeni (Fain)

Neonyssus schoutedeni Fain, 1956a: 134; 1957b: 49

Mesonyssus schoutedeni Fain, 1960c: 92

Falconyssus schoutedeni (Fain): Fain, 1966: 86

Tinaminyssus schoutedeni: Domrow & Nadchatram, 1978: 85

Type locality: Astrida (= Butare), Rwanda

Distribution: Malaysia, Rwanda, Zambia

Hosts: Halcyon leucocephala, H. smyrnensis (Coraciiformes: Halcyonidae)

Tinaminyssus senotrusovae (Butenko)(new comb.)

Mesonyssus senotrusovae Butenko, 1984: 103

Type locality: Tyumenskaya Obl., Yalutorovskiy Rayon (Russia)

Distribution: Kazakhstan, Russia

Host: Streptopelia turtur (Columbiformes: Columbidae)

Tinaminyssus serraoi (de Castro)(new comb.)

Neonyssus (Neonyssus) serraoi de Castro, 1948: 270; Zumpt & Till, 1955: 87; Fain, 1957b: 52

Mesonyssus serraoi (de Castro): Fain, 1962d: 306; Amaral, 1968: 122

Type locality: Brazil

Distribution: Brazil

Host: Rhynchotus rufescens (Tinamiformes: Tinamidae)

Tinaminyssus squamosus (Vitzthum)

Rhinonyssus squamosus Vitzthum, 1935: 567; Pereira & de Castro, 1949: 222; Zumpt & Till, 1955: 88

Neonyssus squamosus (Vitzthum): Fain, 1957b: 48

Rhinonyssoides squamosus: Strandtmann & Wharton, 1958: 173

Mesonyssus squamosus: Fain, 1961: 45; 1962: 305

Falconyssus squamosus: Fain & Aitken, 1969: 407

Tinaminyssus squamosus: Domrow

Type locality: West Indies

Distribution: West Indies, Brazil

Hosts: Florisuga melivora, Sericotes holosericeus (Apodiformes: Trochilidae)

Tinaminyssus streptopeliae (Fain)(new comb.)

Mesonyssus melloi streptopeliae Fain, 1962d: 310; Shumilo & Lunkashu, 1970: 17

Mesonyssus streptopeliae: Fain, Herin & Puylaert, 1977: 392; Sartbaev, 1983: 86

Type locality: Musha (near Astrida, Rwanda)

Tinaminyssus melloi streptopeliae: Rodriguez-Braza, Guevara-Benítez & Úbeda-Ontíveros, 1990: 73

Distribution: Cameroon, Kirghizstan, Moldavia, Rwanda, Spain, Ukraine

Hosts: Columba livia, Streptopelia orientalis, S. s. semitorquata, S. turtur (Columbiformes: Columbidae)

Tinaminyssus streptopelioides (Butenko)

Mesonyssus streptopelioides Butenko, 1984: 107

Tinaminyssus streptopelioides (Butenko): de Rojas et al., 2002:

Type locality: Azerbaydzhan (Kizil-Agach Reserve)

Distribution: Azerbaydzhan, Kazakhstan, Russia, Turkmenistan

Host: Streptopelia turtur (Columbiformes: Columbidae)

Tinaminyssus tanysipterae (Wilson)

Mesonyssus tanysipterae Wilson, 1966a: 605

Tinaminyssus tanysipterae Domrow, 1979: 490; 1987: 886

Type locality: NW New Guinea, Oransbary

Distribution: Australia, Papua New Guinea

Hosts: Tanysiptera danae, T. galatea, T. sylvia (Coraciiformes: Halcyonidae)

Tinaminyssus tetae (Wilson)(new comb.)

Mesonyssus tetae Wilson, 1964b: 605; Tenorio, 1976

Type locality: NW New Guinea, Enarotali, 1900 m

Distribution: Papua New Guinea

Host: green fruit dove (Ptilinopus purpuratus)(Columbiformes: Columbidae)

Tinaminyssus tinamicola (Fain)(new comb.)

Mesonyssus tinamicola Fain, 1963c: 66

Type locality: Anvers Zoo

Distribution: Argentina

Host: Eudromia elegans (Tinamiformes: Tinamidae, Nothurinae)

Tinaminyssus trappi (Pereira et Castro)

Neonyssus (Ptilonyssoides) trappi Pereira & Castro, 1949: 229

Mesonyssus trappi: Fain, 1962d: 306; Amaral, 1968: 122

Tinaminyssus trappi: Strandtmann & Wharton, 1958: 161

Type locality: Boracéa (Estado de São Paulo)

Distribution: Brazil

Host: Tinamus solitarius (Tinamiformes: Tinamidae)

Tinaminyssus treronis (Fain)(new comb.)

Neonyssus treronis Fain, 1956b: 394; 1957b: 56

Mesonyssus treronis Fain, 1962d: 314

Type locality: Akanyaru

Distribution: Rwanda

Host: Treron c. calva (Columbiformes: Columbidae)

Tinaminyssus triangulus (Strandtmann)

Neonyssus triangulus Strandtmann, 1961: 323

Mesonyssus triangulus (Strandtmann): Fain, 1962: 307

Tinaminyssus triangulus (Strandtmann): Pence, 1975: 22

Type locality: Edinburg, Texas

Distribution: USA (Texas)

Host: Zenaida asiatica (Columbiformes: Columbidae)

Tinaminyssus trichoglossi (Domrow)

Mesonyssoides trichoglossi Domrow, 1964: 29

Mesonyssus trichoglossi Wilson, 1964b: 378; 1966b: 191; 1967a: 217

Mesonyssus neopsittaci Wilson, 1964: 372; 1966: 759; 1968: 397 – Type: Enarotali, 1900 m, Bulldog Road, 2440 m, from many species of parrots (syn. by Domrow, 1969: 311)

Mesonyssus domicellae Wilson, 1964b: 375; 1968b: 397 – Type: May River, Papua New Guinea, from Domicella lory (syn. by Domrow, 1969: 311)

Mesonyssus chalcopsyttae Wilson, 1968b: 395 – Type: Oriomo River, Western District, Papua New Guinea, from Chalcopsytta scintillata (syn. by Domrow, 1969: 311)

Tinaminyssus trichoglossi Domrow, 1969b: 311; 1979d: 488; 1987: 886

Type locality: Logan & Tamborine Villages, Queensland

Distribution: Australia, Papua New Guinea

Hosts: Chalcopsitta scintillata, Charmosina papou, Glossopsitta pusilla, Micropsitta bruijnii, Neopsittacus musschenbroekii, N. pullicauda, Pseudeos fuscata, Psittacella modesta, Trichoglossus chlorolepidotus, T. haematodus, T. moluccanus (Psittaciformes: Psittaculidae)

Tinaminyssus turturi (Fain)(new comb.)

Mesonyssus turturi Fain, 1962d: 314

Type locality: Astrida (= Butare)

Distribution: Rwanda

Host: Turtur afer (Columbiformes: Columbidae)

Tinaminyssus welchi Domrow

Mesonyssus sp. “B” Domrow, 1967a: 215

Tinaminyssus welchi Domrow, 1969b: 319; 1987: 886

Type locality: Kowanyama

Distribution: Australia

Host: Ducula [Myristicivora] spilorrhoa (Columbiformes: Columbidae, Treroninae)

Tinaminyssus zenaidurae (Crossley)

Neonyssus zenaidurae Crossley, 1952: 386; Zumpt & Till, 1955: 87; Fain, 1956b: 394; 1957b: 55

Mesonyssus zenaidurae (Crossley): Fain, 1962: 314; Wilson, 1964: 369; Fain & Aitken, 1967: 39

Tinaminyssus zenaidurae: Pence, 1973a: 359; 1975: 22; Powders & Coffey, 1983: 107; Knee, Proctor & Galloway, 2008: 368

Type locality: Dickens Co., Texas

Distribution: Canada, Trinidad, USA (Louisiana, Oklahoma, Texas, Alabama, California, Mississippi)

Hosts: Columbigallina passerina, C. talpacoti, Zenaida [Zenaidura] macroura (Columbiformes: Columbidae)

Tinaminyssus zumpti (Fain)

Mesonyssus zumpti Fain, 1960: 92

Falconyssus zumpti (Fain): Fain, 1966: 86

Type locality: Rooiberg, Transvaal

Distribution: South Africa

Host: Halcyon chelicuti (Coraciiformes: Halcyonidae)

Genus Trochilonyssus Fain et Aitken

Trochilonyssus Fain & Aitken, 1967: 32

Type species: Trochilonyssus trinitatis Fain et Aitken, 1967

Trochilonyssus trinitatis Fain et Aitken

Trochilonyssus trinitatis Fain et Aitken, 1967: 32

Type locality: Esperanza Estate, Vega de Oropouche

Distribution: Trinidad

Host: Glaucis hirsuta (Apodiformes: Trochilidae)

Genus Tyranninyssus Brooks et Strandtmann

Tyranninyssus Brooks & Strandtmann, 1960: 418

Type species: Tyranninyssus tyrannus Brooks et Strandtmann, 1960

Tyranninyssus attilae Fain et Aitken

Tyranninyssus attilae Fain & Aitken, 1970: 405; 1971: 43

Type locality: Mosqueiro Ferry, Benavides, Belém

Distribution: Brazil

Hosts: Attila spadiceus (Passeriformes: Cotingidae); Myiozetetes cayenensis (Passeriformes: Tyrannidae)

Tyranninyssus myiarchi Cruz

Tyranninyssus myiarchi Cruz, 1971: 3

Type locality: Cayo Real

Distribution: Cuba

Host: Myiarchus stolidus (Passeriformes: Tyrannidae)

Tyranninyssus myiophobi Fain et Aitken

Tyranninyssus myiophobi Fain & Aitken, 1967: 27

Type locality: Fort Read

Distribution: Trinidad

Host: Myiophobus fasciatus (Passeriformes: Tyrannidae)

Tyranninyssus tyrannus Brooks et Strandtmann

Tyranninyssus tyrannus Brooks & Strandtmann, 1960: 418

Type locality:

Distribution:

Host: (Passeriformes: Tyrannidae)

Genus Vitznyssus Castro

Neonyssus (Vitznyssus) Castro, 1948: 277

Type species: Dermanyssus nitzschi Giebel, 1871

Astridiella Fain, 1957d: 148

Type species: Ptilonyssus scotornis Fain, 1956

Vitznyssus afrotis (Fain)

Astridiella afrotis Fain, 1966: 83

Vitznyssus afrotis (Fain): Butenko, 1984; Knee, 2018

Type locality: Mafeking

Distribution: South Africa

Host: Afrotis afra (Otidiformes: Otididae)

Vitznyssus caprimulgi (Fain)

Astridiella caprimulgi Fain, 1957d: 96

Type locality: Akanyaru (Rwanda)

Distribution: Rwanda, Botswana

Hosts: Caprimulgus europaeus, Scotornis fossii welwitschi, Cosmetornis vexillarius (Caprimulgiformes: Caprimulgidae)

Vitznyssus erici Knee

Vitznyssus erici Knee, 2018: 10

Type locality: Winnipeg, Manitoba

Distribution: Canada

Host: Chordeiles minor (Caprimulgiformes: Caprimulgidae)

Vitznyssus neotis (Fain)

Astridiella neotis Fain, 1957d: 98; 1960: 108; 1966: 83

Vitznyssus neotis: Strandtmann & Wharton, 1958: 175

Type locality: Astrida (= Butare)

Distribution: Botswana, Rwanda

Hosts: Choriotis kori, Neotis cafra (Gruiformes: Otididae)

Vitznyssus scotornis (Fain)

Ptilonyssus scotornis Fain, 1956a: 148

Astridiella scotornis (Fain): Fain, 1957d: 95; 1957: 148

Vitznyssus scotornis (Fain): Butenko, 1984: 119

Type locality: Astrida (= Butare), Rwanda

Distribution: Rwanda, Suriname

Hosts: Caprimulgus europaeus, C. nigrescens, C. tristigma, C. tristis, Scotornis fossii welwitschi (Caprimulgiformes: Caprimulgidae)

Vitznyssus tetragis Butenko

Vitznyssus vitzthumi Butenko, 1969: 209; 1971b: 260

Vitznyssus tetragis Butenko, 1977: 371; 1984: 122

Type locality: Azerbaidjan

Distribution: Azerbaidjan

Host: Tetrax tetrax (Otidiformes: Otididae)

Vitznyssus tsachevi Dimov et De Rojas

Vitznyssus tsachevi Dimov & De Rojas, 2012: 126

Type locality: Pikalyovo, Leningrad Prov.

Distribution: Russia

Host: Caprimulgus europaeus (Caprimulgiformes: Caprimulgidae)

Vitznyssus vitzthumi (Fain)

Rhinonyssus nitzschi Vitzthum, 1935: 572 (non Giebel, 1871)

Astridiella vitzthumi Fain, 1957d: 95; 1957b: 127; 1961: 513; 1966: 83

Vitznyssus vitzthumi: Butenko, 1969: 209; 1971b: 260; 1984: 121

Type locality: Europe

Distribution: Europe

Host: Otis tarda (Otidiformes: Otididae)

Taxa inquirenda (genus and species)

Tympanospinctus pardoxus Berlese, 1918: 117

Distribution of Rhinonyssidae

EUROPE

Austria (Fain & Sixl, 1969, 1971; Sixl, 1969, 1972; Sixl & Reich, 1971; Fain, Sixl & Moritsch, 1974) – Ptilonyssus acrocephali, P. certhiae, P. coccothraustis, P. echinatus, P. emberizae, P. euroturdi, P. fringillae, P. fringillicola, P. hirsti, P. motacillae, P. pari, P. ruandae sylviae, P. sittae, P. reguli, P. schoenobaeni, Rallinyssus caudistigmus, Sternostoma batis, S. bruxellarum, S. ficedulae, S. hylandi, S. laniorum, S. tracheacolum, S. technaui (=turdi), Tinaminyssus columbae, T. melloi

Azerbaidjan (Butenko, 1965, 1973a, 1971b, 1976, 1984, 2001; Butenko & Stolbov, 1971, 1972) – Ptilonyssus triscutatus, Ptilonyssus [Cas] elbeli, Rallinyssus caudistigmus (= caspicus), R. gallinulae, Rhinonyssus alberti, Rh. minutus, Rh. neglectus, Rh. podicipedus (= tschomgae), Rh. rhinolethrum (= nyrocae), Sternostoma bruxellarum, S. fulicae, Vitznyssus [Astridiella] tetragis

Belgium (Fain, 1961, 1962b, 1963a, 1963b, 1964a, 1965, 1966, 1972; Fain & Bafort, 1963b; Fain & Hyland, 1963a; Fain & Sixl, 1971) – Ptilonyssus acrocephali, P. bombycillae, P. carduelis, P. chloris, P. coccothraustis, P. echinatus, P. emberizae, P. euroturdi, P. fringillae, P. fringillicola, P. hirsti, P. [Passeronyssus] lusciniae, P. morofskyi, P. motacillae, P. nudus, P. orioli, P. pari, P. phylloscopi, P. prunellae, P. ruandae sylviae, P. sittae, P. troglodytis, Rallinyssus gallinulae, R. verheyeni, Rhinoecius oti, Rhinonyssus [afribyx] vanellus, Rh. coniventris, Rh. echinipes, Sternostoma bruxellarum, S. francolini, S. fulicae, S. loxiae, S. technaui, S. tracheacolum, Tinaminyssus columbae, T. melloi

Bulgaria (Beron, 1975) – Ptilonyssus carduelis, P. echinatus, P. prunellae, Sternostoma bruxellarum

Czech Rep. (Černy, 1970) – Tinaminyssus columbae, T. hirsutus, T. melloi, T. streptopeliae

Denmark

Greenland (Trägårdh, 1904b) – Rhinonyssus levinseni

Finland (Huhta, 2016) – Ptilonyssus echinatus, P. nudus (as Species incertae)

France (contin.) (Berlese & Trouessart, 1889; Gretillat, 1961; Jolivet, 1975) – Ptilonyssus nudus, Rallinyssus strandtmanni, Rhinonyssus himantopus, Rh. rhinolethrum, Sternostoma cryptorhynchum, S. tracheacolum

Georgia (Bregetova, 1967) – Ptilonyssus calandrellae

Germany (Berlese & Trouessart, 1889; Bregetova, 1953) – Ptilonyssus echinatus, Rhinonyssus neglectus, Rh. waterstoni, Sternostoma caledonicum

Great Britain (Hirst, 1916, 1921, Turk, 1953; Bygrave, 1980) – Ptilonyssus nudus, Rhinonyssus coniventris (= neglectus, = echinipes), Sternostoma caledonicum, S. tracheacolum, S. waterstoni

Holland (van Eyndhoven, 1964) – Ptilonyssus nudus, Rhinonyssus levinseni, Rh. rhinolethrum

Hungary (Rozsa, 1990) – Tinaminyssus melloi

Italy (contin.) (Lombardini, 1953) – Ptilonyssus nudus, Sternostoma tracheacolum (= S. meddai)

Latvia (Bregetova, 1953, 1967; Butenko, 2001) – Ptilonyssus nucifragae, Sternostoma bruxellarum

Moldavia (Shumilo, 1960, 1965, 1968; Lunkashu, 1966; Shumilo & Dementieva, 1963; Shumilo & Lunkashu, 1970) – Ptilonyssus carduelis, P. chloris, P. coccothraustis, P. echinatus, P. emberizae, P. euroturdi, P. hirsti, P. lanii, P. lusciniae, P. morofskyi, P. motacillae, P. muscicapae, P. orioli, P. pari, P. pygmaeus, P. sittae, P. triscutatus, Sternostoma bruxellarum, S. cooremani, S. cuculorum, S. hirundinis, S. technaui (= turdi), S. tracheacolum, Tinaminyssus melloi, T. streptopeliae

Poland (Szeleszczuk & Kruszewicz, 1987) – Sternostoma tracheacolum

Portugal (Castro & Pereira, 1947; Ribeiro-Diogo, Lino de Sousa & Santos-Amoroso, 1973) – Ptilonyssus hirsti, Sternostoma tracheacolum

Romania (Feider, 1962; Feider & Mironescu, 1962, 1968, 1969, 1970, 1972a, 1972b, 1973a, 1973b, 1973c, 1974a, 1974b, 1978) – Ptilonyssus elbeli, P. euroturdi, P. fringillicola (= pyrrhulae), P. hirsti, P. sittae, Rallinyssus caudistigmus, Rhinonyssus alberti, Rh. podicipedus, Rh. rhinolethrum, Sternostoma epistomata, S. fulicae, S. pastor, S. technaui (= borceanum), Tinaminyssus gerschi

Russia (all, continent.) (Belopolskaya, 1947, 1952; Bregetova, 1950, 1951, 1953, 1956, 1964, 1965a, 1965b, 1965c, 1967a, 1967b, 1970; Zemskaya & Ilienko, 1958; Butenko, 1959, 1960a, 1960b, 1962a, 1962b, 1964, 1965, 1968, 1969, 1971a, 1971b, 1971c, 1972, 1973a, 1973b, 1973c, 1974a, 1974b, 1984, 1999, 2001; Butenko & Priklonskiy, 1964; Isakova, 1963, 1965, 1966, 1968; Butenko & Stolbov, 1971; Ivanov & Alifanov, 1972; Butenko & Lavrovskaya, 1980, 1983, 1984a, 1984b, 1985, 1988; Bregetova & Slavoshevskaya, 1983; Butenko, Stanyukovich & Lavrovskaya, 1996; Stanyukovich & Butenko, 1998, 2003, 2004; Butenko & Stanyukovich, 1997, 1999, 2001; Butenko, 2003; Butenko, Stanyukovich & Dimov, 2009; Dimov, Butenko & Stanyukovich, 2009; Dimov, 2010, 2011, 2012, 2013a, 2013b, 2016, 2018; Dimov & Knee, 2012; Dimov & De Rojas, 2012; Dimov & Mironov, 2012; Dimov & Mascarenas, 2012; Dimov & Spicer, 2013) – Larinyssus benoiti, L. iohanssenae, L. orbicularis, L. substerna, Locustellonyssus amurensis, L. sibiricus, Ptilonyssus acanthopneustes, P. anthi, P. capensis, P. capitatus, P. carduelis, P. chloris, P. cerchneis, P. cyanosylviae, P. degtiarevae, P. echinatus (= Hapalognatha platytrichia), P. [Cas] elbeli, P. emberizae, P. euroturdi, P. fringillicola, P. hirsti, P. icteridius, P. lanii, P. lovottiae, P. mironovi, P. motacillae (= Hapalognatha prima), P. muscicapae, P. muscicapoides, P. nucifragae, P. nudus, P. pari, P. phylloscopi, P. [Neonyssus (Spizonyssus)] pygmaeus, P. pyrrhulinus, P. ripariae, P. sairae, P. schumili, P. spini, P. sylviicola, P. tachycinetae, P. triscutatus, Rallinyssus caudistigmus (= caspicus), R. verheyeni, Rhinoecius aegolii, Rh. alifanovi, Rh. brikinboricus, Rh. nycteae, Rh. subbisetosus, Rhinonyssus alberti, Rh. bregetovae, Rh. caledonicus, Rh. colymbicola, Rh. coniventris, Rh. dobromiri, Rh. echinipes, Rh. himantopus, Rh. kadrae, Rh. minutus, Rh. neglectus, Rh. pluvialis, Rh. podicipedus, Rh. polyocephali, Rh. polystictae, Rh. rhinolethrum (= anatidae = nyrocae), Rh. tringae, Rh. vanellus, Rh. waterstoni, Sternostoma batis, S. boydae, S. bruxellarum, S. cooremani, S. cuculorum, S. dureni, S. epistomata (= chlidoniadis), S. francolini, S. fulicae, S. hylandi, S. marchae, S. sternahirundo, S. strandtmanni, S. strigiformes, S. technaui (= turdi), S. zini; Tinaminyssus belopolskii, T. columbae, T. egrettae, T. ixoreus, T. melloi, Vitznyssus tsachevi (here are listed only some of the Russian Rhinonyssidae, which are more than140 sp.)

Serbia (Kulišić, 1988) – Tinaminyssus melloi

Spain (cont.) (Blasco Sabio & Portús Vinyeta, 1974; Guevara-Benítez, Lopez & Úbeda-Ontíveros, 1974; Guevara-Benítez & Úbeda-Ontíveros, 1974, 1975a, 1975b, 1978a, 1978b, 1978c, 1978d, 1978d; 1979, 1980a, 1980b; Guevara-Benítez, Úbeda-Ontíveros & Cutillas-Barrios, 1979; Úbeda-Ontíveros, Guevara- Benítez & Guevara Pozo, 1978; Guevara-Benítez, Úbeda-Ontíveros & Guevara- Pozo, 1978; Úbeda-Ontíveros & Guevara – Benítez, 1978, 1980, 1981, 1985, 2000; Úbeda-Ontíveros, Guevara – Benítez & Morillas-Marquez, 1983, 1985; Úbeda-Ontíveros, Guevara-Benítez & Morillas-Marquez, 1986; Guevara-Benítez, Úbeda-Ontíveros & Morillas- Marquez, 1986a, 1986b; Úbeda, Rodriguez, Guevara & Rojas, 1989; Rodriguez Braza et al., 1989; Rodriguez, Úbeda & Guevara, 1990, 1991, 1993; Úbeda, Rodriguez & Guevara, 2000; de Rojas, Mora, Úbeda et al., 2001; de Rojas, Mora et al., 2002; Lozano et al., 2003; Úbeda, de Rojas et al., 2003; de Rojas, Úbeda et al., 2007) – Ptilonyssus bombycillae, P. [Astridiella] calandrellae, P. capensis, P. carduelis, P. chloris, P. echinatus, P. emberizae, P. euroturdi, P. fringillae, P. gilcolladoi, P. hirsti, P. lusciniae, P. morofskyi, P. motacillae, P. pari, P. phylloscopi, P. prunellae, P. reguli, P. strandtmanni, Rallinyssus caudistigmus, Rhinoecius bisetosus, Rhinonyssus echinipes, Rh. neglectus, Rh. tringae, Rh. vanellus, Sternostoma christinae, S. fulicae, S. guevarai, S. isabelae, S. sternahirundo, S. technaui (= turdi), S. tracheacolum, S. ubedai, Tinaminyssus columbae, T. melloi, T. minisetosum, T. sartbaevi, T. streptopeliae

Canary Is. (Úbeda-Ontiveros, Guevara-Benítez & Rodriguez-Braza, 1987; Rodriguez-Braza, Guevara-Benítez & Úbeda-Ontiveros, 1990) – Tinaminyssus bubulci, T. melloi

Ukraine (Shumilo & Lunkashu, 1970, Butenko, 1984) – Rallinyssus caudistigmus, Ptilonyssus coccothraustis, P. echinatus, P. emberizae, P. euroturdi, P. fringillicola, P. lanii, P. motacillae, P. motacillae phoenicuri, P. nucifragae, P. orioli, P. pari, P. sittae, P. triscutatus, Rhinonysus rhinolethrum (= nyrocae), Sternostoma bruxellarum, S. cuculorum, S. hirundinis, S. kodrensis, S. technaui (= turdi), Tinaminyssus columbae, T. melloi, T. streptopeliae

AFRICA

Botswana (Fain, 1956, 1958b, 1959а, 1960d, 1966) – Passeronyssus dicruri, Ptilonyssus cisticolarum, P. lanii, P. motacillae, Ruandanyssus buboensis, Sternostoma colii, S. cuculorum (= laniorum), S. thienponti, Tinaminyssus bubulci, Vitznyssus [Astridiella] caprimulgi, V.[Astridiella] neotis

Cameroon (Fain, 1960a, 1960d; Fain, Herin & Puylaert, 1977) – Ptilonyssus angrensis, P. astridae, P. aureliani, P. echinatus, P. estrildicola, P. fringillicola, P. hirsti, P. lanii, P. nudus, P. ploceanus, P. strandtmanni, P. taeniopygiae, P. triscutatus, P. viduicola, Sternostoma colii, S. cooremani, S. cryptorhynchum, S. hirundo, S. nectarinia, S. sinense, S. sturnicola, Tinaminyssus streptopeliae

Congo DR (Fain, 1957b, 1958b, 1966c) – Larinyssus orbicularis, Ptilonyssus nicatoris, Rhinonyssus afribyx, Rh. himantopus, Rh. rhinolethrum, Sternostoma nectarinia, S. straeleni, Tinaminyssus elani

Egypt (Selim, El-Kasaby & El-Refaii, 1968) – Tinaminyssus melloi

Guinea (Butenko, 1999) – Sternostoma numerovi

Madagascar (Hirst, 1922; Grétillat, Capron & Brygoo, 1959) – Ptilonyssus intermedius, P. madagascariensis, Sternostoma tracheacolum (= Agapornyssus faini)

Mozambique (Fain, 1958b, 1962c) – Ptilonyssus cinnyricincli, Ruandanyssus buboensi

Rwanda (Fain, 1956b, 1957a, 1957b, 1959a, 1962c, 1962d, 1963c; 1964e; Fain, Herin & Puylaert, 1977) – Larinyssus orbicularis, Passeronyssus dicruri, P. dioptrornis, Ptilonyssus afroturdi, P. aureliani, P. bradypteri, P. calamocichlae hippolaisi, P. capensis, P. cinnyris, P. cisticolarum, P. elongatus, P. hirsti, P. lusciniae, P. melittophagi, P. nudus, P. orioli, P. psalidoprocnei, P. pycnonoti, P. ruandae, P. serini, P. strandtmanni, P. terpsiphonei, P. tillae, P. triscutatus, P. viduae, P. viduicola, P. zumpti, Rallinyssus congolensis, R. limnocoracis, Rhinoecius tytonis, Rhinonyssus apus, Rh. afribyx, Rh. coniventris, Rh. himantopus, Rh. poliocephali, Rh. rhinolethrum, Rh. tringae, Ruandanyssus terpsiphonei, Sternostoma batis, S. boydae, S. colii, S. cooremani, S. cuculorum, S. cuculorum korolevae, S. dureni, S. giganteum, S. hirundinis, S. hirundo, S. inflatum, S. lagonostictae, S. nectarinia, S. sturnicola, S. technaui (= turdi), S. thienponti, S. tracheacolum (= castroae), Tinaminyssus belopolskii (= Neonyssus ardeae), T. streptopeliae, T. treronis, Vitznyssus scotornis

South Africa (Zumpt & Till, 1951, 1955; Fain, 1957b, 1959a, 1960a, 1960c, 1960d, 1963c, 1966, 1972; Fain, Herin & Puylaert, 1977; Lindholm, Venter & Ueckermann, 1998) – Ptilonyssus capensis, P. capitatus, P. certhilaudae, P. cinnyris, P. cisticolarum, P. dioptrornis, P. dryoscopi, P. echinatus, P. estrildicola, P. fringillicola, P. hirsti, P. lanii, P. melittophagi, P. motacillae, P. ploceanus, P. pycnonoti, P. terpsiphonei, P. tillae, Rallinyssus caudistigmus, Rhinoecius tytonis, Rhinonyssus himantopus, Sternostoma cisticolae, S. cuculorum var. urolestis, S. eurocephali, S. hirundo, S. technaui (= turdi), Tinaminyssus belopolskii (= ardeae), T. bubulci, T. buteonis, T. melloi, T. oenae, T. zumpti, Vitznyssus [Astridiella] afrotis

Zambia (Fain, 1960c; Mwase & Baker, 2006) – Larinyssus orbicularis, Rhinonyssus himantopus, Rh. rhinolethrum, Ruandanyssus terpsiphonei, Sternostoma boydae, S. francolini, Tinaminyssus schoutedeni, T. zumpti

East Arica – Tinaminyssus baforti

ASIA

Afghanistan (Butenko, 1973a) – Rhinonyssus podicipedus (= tschomgae)

India (Kapoor & Kaur, 1975) – varia

Indonesia (Fain, 1965; Wilson, 1966) – Tinaminyssus kakatuae, T. phalliger

Iran (Moodi et al., 2014) – Ptilonyssus hirsti, P. icteridius, P. pirangae

Japan (Kaneko, Matsudaira & Prince Masahito, 1978; Ehara, 1980; Kadosaka, Kaneko & Asanuma, 1983, 1987) – Ptilonyssus euroturdi, P. hiyodori, P. emberizae, P. enicuri, P. pari, P. ruandae, P. sairae, Rallinyssus caudistigmus, Rhinonyssus himantopus, Rh. rhinolethrum, Ruandanyssus terpsiphonei, Sternostoma technaui, Tinaminyssus columbae, T. melloi

Kazakhstan (Butenko, Senotrusova & Sema, 1970; Butenko, 1972, 1973a, 1973b, 1973c, 1974, 1984, 2001; Butenko & Lavrovskaya, 1980) – Charadrinyssus [Larinyssus] charadrius, Ptilonyssus langei, P. pastoris, P. schumili, P. triscutatus, Rallinyssus caudistigmus, Rhinonyssus alberti, Rh. bregetovae, Rh. chettusiae, Rh. himantopus, Rh. neglectus, Rh. rhinolethrum (=nyrocae), Rh. shcherbininae, Rh. vanellus, Sternostoma boydae, S. bruxellarum, S. francolini

Kirghizstan (Sartbaev, 1961, 1977, 1979, 1981; Sartbaev & Kumushaliev, 1982; Bregetova, 1967; Butenko & Lavrovskaya, 1980, 1983; Butenko, 1984, 1999) – Larinyssus benoiti, Ptilonyssus [Cas] angrensis, P. capensis, P. cerchneis, P. [Cas] elbeli, P. eremophilae, P. euroturdi, P. fringillicola, P. isakovae, P. lanii, P. montifringillae, P. motacillae, P. nucifragae, P. phylloscopi, P. prunellae, P. ptyonoprognes, P. schumili, P. strandtmanni, Rallinyssus caspicus, R. caudistigmus, R. verheyeni, Rhinonyssus himantopus, Rh. minutus, Rh. rhinolethrum [anatidae], Rh. shcherbininae, Rh. vanellus, Sternostoma batis, S. boydae, S. cooremani, S. francolini, S. fulicae, S. graculi, S. technaui, S. tracheacolum, Tinaminyssus melloi, T. streptopeliae, Vitznyssus [Astridiella] caprimulgi

Malaysia (Strandtmann, 1960; Fain & Nadchatram, 1962; Nadchatram, McClure & Chong, 1964; Fain, 1964c; McClure, Ratanaworadhan et al., 1973; Domrow & Nadchatram, 1978) – Ptilonyssus andropadi, P. cinnyris, P. dioptrornis, P. dicruri, P. enicuri, P. geopeliae, P. lanii, P. lobatus, P. lusciniae, P. macclurei, P. malaysiae, P. motacillae, P. nadchatrami, P. ploceanus, P. pycnonoti, P. ruandae, P. traubi, P. triscutatus, P. wilsoni, Rallinyssus [Rallinyssoides] limnocoracis, Sternostoma cooremani halcyoni, S. cuculorum, S. furmani, S. tracheacolum, Tinaminyssus malayi, T. ptilinopi, T. schoutedeni

Philippines (Wilson, 1966, 1967a, 1968a; Domrow, 1969; McClure, Ratanaworadhan et al., 1973) – Rallinyssus amaurornis, R. congolensis, R. porzanae, R. rallinae, Tinaminyssus geopeliae, T. halcyonus, T. hirtus, T. macropygiae, T. peleiae, T. prionituri, T. ptilinopi

Taiwan (Maa & Kuo, 1965; Wilson, 1965, 1966a; Sakakibara, 1967, 1968; Sakakibara & Strandtmann, 1968; McClure, Ratanaworadhan et al., 1973) – Ptilonyssus [Paraneonyssus] dendrocittae, P. [Paraneonyssus] pericrocoti, Rallinyssus amaurornis, Tinaminyssus phassae, T. ptilinopi

Tajikistan (Bregetova, 1965, 1967) – Ptilonyssus [Astridiella] calandrellae, P. melanocoryphae, Ptilonyssus [Neonyssus (Spizonyssus)] pygmaeus, Sternostoma technaui

Thailand (Ewing, 1933; Strandtmann, 1960; Wilson, 1968c; Boonkong & Meckvichai, 1987) – Ptilonyssus [Passeronyssus] dicruri, P. dioptrornis, P. echinatus, P. [Cas] elbeli, P. faini, P. hirsti, P. lobatus, P. [Vitznyssus] nitzschi, P. [Sommatericola] orientalis, P. orioli, Rhinoecus bisetosus, Rh. cavannus, Rhinonyssus rhinolethrum, Sternostoma batis, S. bruxellarum (sub Cas elbeli), S. nectarinia, S. thienponti, S. tracheacolum

Turkmenistan (Bregetova, 1965, 1970; Butenko & Shcherbinina, 1976; Shcherbinina & Butenko, 1981; Butenko, 1984; Butenko & Stanyukovich, 2001; Stanyukovich & Butenko, 2003) – Ptilonyssus [Neonyssus] alaudae, Ptilonyssus ammomani, P. echinatus, P. emberizae, P. hirsti, P. muscicapae, P. schumili, P. triscutatus, Rallinyssus caudistigmus, Rhinoecius subbisetosus, Rhinonyssus himantopus, Rh. minutus, Rh. shcherbininae, Rh. vanellochettusiae, Tinaminyssus streptopelioides

Uzbekistan (Butenko, 1971a, 1984) – Rhinonyssus podicipedus, Rh. rhinolethrum (= subrhinolethrum)

NORTH AMERICA

Canada (George, 1961; Pence, 1975; Ballard & Ring, 1979; Hood & Welch, 1980; Knee, 2008; Knee, Proctor & Galloway, 2008; Knee & Proctor, 2010; Knee & Galloway, 2017; Knee, 2018) – Ptilonyssus acrocephali, P. angrensis, P. bombycillae, P. callinectoides, P. calvaria, P. carduelis, P. cerchneis, P. coccothraustis, P. echinatus, P. euiroturdi, P. hirsti, P. icteridius, P. japuibensis, P. ludovicianus, P. morofskyi, P. nivalis, P. nudus, P. perisorei, P. pinicola, P. pirangae, P. plesiotypicus, P. porteri, P. sairae, P. troglodytis, P. tyrannus, P. vireonis, Rallinyssus caudistigmus, R. sorae, R. verheyeni, Rhinoecius aegolii, Rh. alifanovi, Rh. brikinboricus, Rh. cooremani, Rh. grandis, Rh. nycteae, Rhinonyssus colymbicola, Rh. coniventris, Rh. pluvialis, Rh. rhinolethrum, Rh. strandtmanni, Sternostoma boydae, S. cryptorhynchum, S. cuculorum (= laniorum), S. gallowayi, S. guevarai, S. hylandi, S. longisetosae, S. loxiae, S. setifer, S. sialiphilus, S. technaui, S. tracheacolum, Tinaminyssus columbae, T. melloi, T. zinaidurae, Vitznyssus erici (56)

Mexico (Hyland & Moorehouse, 1970; Spicer 1977b; Luz Zamudio, 1984) – Ptilonyssus callinectoides, P. camptostoma, P. hoffmannae, P. japuibensis, P. fluvicolae, P. mariacastroae, P. saiorae, P. tyrannus, Sternostoma hirundinis, S. longisetosa, S. tracheacolum, Tinaminyssus chiarellii, Tyranninyssus. spinosus

USA (Strandtmann, 1948, 1951, 1952, 1956a, 1956b, 1959, 1961b, 1962; Strandtmann & Furman, 1956; Furman, 1957; Owen, 1958; Hyland & Clark, 1959; Brooks & Strandtmann, 1960; Hyland & Ford, 1960; Mitchell & Rhodes, 1960; George, 1961; Hyland, 1961, 1962; Kennedy & Strandtmann, 1962; Mitchell, 1963a, 1963b; Foulk & Matthysse, 1965; Spory, 1965; Fain & Johnson, 1966; Wilson, 1958, 1968a, 1980; Brown, 1971; Cooper & Troutman, 1974; Pence, 1972a, 1972b, 1972c, 1972d, 1973a, 1973b, 1973c, 1975; Pence & Casto, 1975a, 1975b, 1976; Cooper, Madden & Crites, 1975; Pence & Canaris, 1976; Spicer, 1977a, 1977b, 1978, 1979, 1984, 1987; Pence & Young, 1979; Wilson & Haas, 1980; Powders & Coffey, 1983; Hilario-Pérez & Dowling, 2018) – Larinyssus orbicularis, Ptilonyssus alberti, P. callinectoides, P. capitatus, P. carduelis, P. cerchneis, P. echinatus, P. euroturdi, P. hoseini, P. icteridius, P. japuibensis, P. lanii, P. melissae, P. mimi, P. morofskyi, P. motacillae, P. nudus, P. ohioensis, P. perisorei, P. phainopeplae, P. psaltriparus, P. richmondenae, P. sairae (= constrictus), P. salpinctis, P. sialiae, P. tachycinetae, P. thryothori, P. toxostomae, P. vireonis, P. vossi, Rallinyssus caudistigmus, R. sorae, R. verheyeni, Rhinoecius bisetosus, Rh. cooremani, Rh. grandis, Rhinonyssus alberti, Rh. caledonicus, Rh. coniventris, Rh. himantopus, Rh. pluvialis, Rh. podilymbi, Rh. rhinolethrum, Rh. strandtmanni, Sternostoma crotophagae, S. dumetellae, S. hutsoni, S. hylandi, S. kelloggi, S. longisetosae, S. pencei, S. pirangae, S. porteri, S. sayornis, S. strandtmanni, S. technaui (= spatulatum), S. tracheacolum, S. tyrannus, Tinaminyssus belopolskii, T. columbae, T. melloi, Tyranninyssus callinectoides, T. juxtamelloi, T. spinosus, T. triangulus, T. tyrannus

Alaska (Strandtmann, 1956; Wilson & Haas, 1980) – Ptilonyssus carduelis, P. perisorei, P. sairae, Rhinonyssus coniventris

Hawaii Is. (Tenorio, Denmark & Swift, 1985; van Riper & van Riper III, 1985) – Ptilonyssus hirsti, Rhinonyssus coniventris, Sternostoma tracheacolum, Tinaminyssus geopeliae

SOUTH and CENTRAL AMERICA

Argentina (Fain, 1963c, 1964e; Kun & Vega, 2018; Wilson, 1970) – Rallinyssus caudistigma, Rhinonyssus rhinolethrum, Sternostoma (= Rhinosterna) aymarae, Tinaminussus tinamicola

Brazil (Vitzthum, 1935; Castro & Pereira, 1947; Castro, 1948; Pereira & Castro, 1949; Amaral, 1962a, 1962b, 1962c, 1963, 1963b, 1964b, 1966, 1967a, 1967b, 1967c, 1967d, 1968, 1973; Amaral & Baquer, 1963; Fain & Aitken, 1968, 1969, 1970, 1971; Fain, 1964, 1972; Amaral & Reboucas, 1974; Sinkoc, 2006; Mascarenhas & Brum, 2006; Paulsen, 2006; Mascarenhas et al., 2007; Mascarenhas et al., 2009; Mascarenhas, Coimbra et al., 2011; Dimov & Mascarenhas, 2012; Coimbra et al., 2012 ; Mascarenhas, Coimbra, Afonso, Sincoc, Müller & Brum, 2012; Mascarenhas, Pesenti, Coimbra, Lambrecht & Müller, 2013; Bernardon et al., 2012; Bernardon, Müller & Mascarenhas, 2013, 2017; Mendes, Mascarenhas, Sinkoc & Müller, 2014; Sinkoc, Brum & Müller, 2016; Gastal et al., 2018; Santos et al., 2018; Mascarenhas et al., 2018; Gastal & Mascarenhas, 2018; Silva da Silva, Scheer & Müller, 2018; Bernardon, Mascarenhas, Pereira Jr. & Müller, 2018) – Larinyssus orbicularis, Pipronyssus manaci, Ptilonyssus amarali, P. amazonicus, P. angustirostris, P. cercomacrae, P. certhiaxicola, P. chalybaedomesticae, P. chiroxiphiae, P. colopteryx, P. conopophagae, P. corythopicola, P. donatoi, P. enrietti, P. formicarii, P. formicarii colmae, P. hirsti, P. hylophylax, P. hypocnemoides, P. icteridius, P. inflatipalpus, P. inornatus, P. japuibensis, P. japuibensis cyanocompsae, P. myiobii, P. myrmotherulae, P. olaioi, P. paranensis, P. pipromorphae, P. platyrhynchus, P. psaltriparus, P. pyriglenae, P. rabelloi, P. sairae, P. sclateriae, P. sclerurus, P. souzai, P. spinosus, P. sternostomicus, P. stresemanni, P. terenotricci, P. thamnomanes, P. thamnophili, P. travassosfilhoi, P. werneri, P. xiphorhynchus, P. zeferinoi, Rhinonyssus rhinolethrum, R. sphenisci, Sternostoma batis, S. boydae, S. hedonophilum, S. longisetosae (= callithrix), S. mortelmansi, S. piprae, S. pirangae, S. strandtmanni, S. tracheacolum, Tinaminyssus belopolskii, T. cunhai, T. marciae, T. saucerottiae, T. squamosus, Tyranninyssus attilae – 61 (108)

Chile (contin.) (Fain, 1964) – Rhinonyssus belenopteri

Cuba (Černý, 1967, 1969 ; Dusbábek, 1969; Dusbábek & Černý, 1969; Černý & Dusbábek, 1970; Cruz, 1971) – Larinyssus orbicularis, Ptilonyssus chalybeaedomesticae, P. constrictus constrictus, P. c. longisetosus, P. echinatus, P. euroturdi, P. icteridius, P. hirsti, P. hoseini (= Passeronyssus havanensis), P. insularis insularis, P. insularis cubanus, P. ludovicianus, P. mimicola, P. morofskyi, P. pirangae, P. sairae, P. teretistris, P. [Passeronyssus] vireonis, Rhinonyssus coniventris, Rh. himantopus, Rh. rhinolethrum, Rh. spinactitis, Rh. tringae, Sternostoma hirundinis, S. kelloggi, S. orlandoi, S. quiscali, S. spatulatus, Tinaminyssus belopolskii, T. bubulci, T. carapachibeyus, T. geotrygoni, Tyranninyssus myiarchi, T. spinosus, T. tyrannus (= T. t. caribaeus)

Guatemala (Spicer, 1979, 1984) – Ptilonyssus euroturdi (= P. e. mimicola), P. sairae, P. tyrannus, Sternostoma darlingi, S. hutsoni, S. pencei, S. pirangae, Tinaminyssus ixoreus

Peru (Fain & Mortelmans, 1959) – Rhinonyssus sphenisci

Puerto Rico (Brooks & Strandtmann, 1960) – Tyranninyssus spinosus

Suriname (Fain & Lukoschus, 1972) – Ptilonyssus [Astridiella] scotornis, Ptilonyssus echinatus

Trinidad y Tobago (Fain & Aitken, 1967) – Neotyranninyssus fluvicolae, Passeronyssus hoseini, Pipronyssus manaci, Ptilonyssus agelaii, P. amarali, P. constrictus, P. elaeniae, P. echinatus, P. euroturdi, P. certhiaxicola, P. icteridius, P. japuibensis, P. mariacastroae, P. pipromorphae, P. sairae, P. serini, P. thamnophili, P. werneri, Sternoecius piprae, Sternostoma boydae, S. hutsoni, S. quiscali, S. strandtmanni, S. tangarae, S. technaui (= turdi), S. tracheacolum, Tinaminyssus zenaidurae, Trochilonyssus saucerottiae, T. trinitatis, Tyranninyssus myiophobi

Uruguay (Cassamagnaghi, 1952) – Sternostoma tracheacolum

Venezuela (Fain & Holland, 1972) – Larinyssus sterna, Sternostoma boydae

South America (Fain, 1967) – Tinaminyssus crypturelli

West Indies (Fain & Aitken, 1969) – Tinaminyssus squamosus

OCEANIA

Australia (continent)(Domrow, 1964a, 1964b, 1964c, 1965, 1966a, 1967a, 1969, 1979, 1987, 1992; Fain & Lukoshus, 1979; Halliday, 1998) – Larinyssus benoiti, L. orbicularis, Ptilonyssus acrocephali, P. ailuroedi, P. angrensis, P. astridae, P. balimoensis, P. bradypteri, P. buloloensis, P. capitatus, P. carduelis, P. cerchneis, P. cinnyris, P. colluricinclae, P. condylocoxa, P. conopophilae, P. corcoracis, P. cractici, P. dicaei, P. dicruri, P. dioptornis, P. echinatus, P. emberizae, P. gerygonae, P. gliciphilae, P. grallinae, P. hirsti, P. lymozemae, P. macclurei, P. maluri, P. meliphagae, P. microecae, P. monarchae, P. motacillae, P. myzanthae, P. myzomelae, P. neochmiae, P. ohioensis, P. orthonychus, P. pentagonicus (= P. philemoni), P. pittae, P. pseudothymanzae, P. psophodae, P. pygmaeus, P. rhipidurae, P. ruandae, P. setosae, P. sittae, P. sphecotheris, P. stomioperae, P. struthideae, P. sturnopastoris, P. terpsiphonei, P. thymanzae, P. triscutatus, P. trouessarti, Rallinyssus caudistigmus, R. congolensis, R. gallinulae, Rhinoecius cooremani, Rhinonyssus coniventris, Rh. himantopus, Rh. minutus, Rh. poliocephali, Rh. rhinolethrum, Ruandanyssus artami, R. terpsiphonei, Sternostoma cooremani, S. cuculorum, S. dureni, S. gliciphilae, S. neosittae, S. paddae, S. technaui, S. thienponti, S. tracheacolum, Tinamynyssus aprosmicti, T. belopolskii, T. columbae, T. daceloae, T. geopeliae, T. halcyonus, T. hirtus, T. kakatuae, T. macropygiae, T. megaloprepiae, T. melloi, T. myristicivorae, T. ocyphabus, T. phabus, T. tanysipterae, T. trichoglossi, T. welchi

New Zealand (Ramsay, 1970; Domrow, 1972) – Ptilonyssus cractici, P. emberizae, P. euroturdi, Rallinyssus gallinulae, Rhinonyssus rhinolethrum, Tinaminyssus halcyonis, T. melloi

Papua New Guinea (Baker & Delfinado, 1964; Wilson, 1964b, 1965, 1966a, 1968; Sakakibara, 1967, 1968a, 1968b, 1968c; Tenorio, 1976; Domrow, 1979; Feider & Mironescu, 1980, 1982) – Ptilonyssus balimoensis, P. buloloensis, P. colluricinclae, P. cychramus, P. dicruri, P. dolicaspis, P. domrowi, P. estrildicola, P. gressitti, P. missimensis, P. nitzschi, P. novaeguineae, P. paradisaeus, P. radovskyi, P. terpsiphonei, P. trouessarti, P. trouessarti pseudotrouessarti, P. wilsoni, Rallinyssus cychramus, R. gallinulae (= rallus), Sternostoma artami, S. constricta, S. cordiscutata, S. opistaspis, S. ziegleri, Tinaminyssus alisteri, T. angustus, T. epileus, T. gourae, T. halcyonus, T. macropygiae, T. ptilinopi, T. tanysipterae, T. tetae

ANTARCTICA (Gastal, Mascarenhas, Vanstreels & Ruas, 2018; Wilson, 1967b, 1970, 1971) – Rhinonyssus rhinolethrum, Rh. sphenisci schelli, Rh. sphenisci

Hosts of Rhinonyssidae

Aves

Accipitriformes

Fam. Accipitridae

Aviceda subcristata – Tinaminyssus epileus

Buteo rufofuscus augur – Tinaminyssus buteonis

Elanus coeruleus – Tinaminyssus elani

Lophaetus occipitalis – Tinaminyssus buteonis

Milvus tenebrosus – Tinaminyssus columbae

Rupornis magnirostris magniplumis – Ptilonyssus souzai

Accipitriformes – Tinaminyssus epileus

Anseriformes

Fam. Anatidae

Aix sponsa – Rhinonyssus rhinolethrum

Alopochen aegyptiacus – Rhinonyssus rhinolethrum

Anas acuta – Rhinonyssus rhinolethrum (= anatidae)

A. [Mareca] americana – Rhinonyssus rhinolethrum

A. carolinensis – Rhinonyssus rhinolethrum

A. clypeata – Rhinonyssus rhinolethrum (= anatidae)

A. crecca – Rhinonyssus rhinolethrum (= anatidae= subrhinolethrum)

A. discors – Rhinonyssus rhinolethrum

A. erythrorhyncha – Rhinonyssus rhinolethrum

A. falcata – Rhinonyssus rhinolethrum (= anatidae)

A. georgicus – Rhinonyssus rhinolethrum

A. gibberifrons – Rhinonyssus levinseni

A. penelope – Rhinonyssus rhinolethrum (= anatidae= subrhinolethrum)

A. platyrhynchos – Rhinonyssus levinseni, Rh. rhinolethrum (= anatidae = subrhinolethrum), Rh. kadrae

A. poecilorhyncha zonorhyncha – Rhinonyssus rhinolethrum

A. querquedula – Rhinonyssus rhinolethrum (= anatidae), Rh. levinseni

A. strepera – Rhinonyssus rhinolethrum (= anatidae)

A. superciliosa – Rhinonyssus rhinolethrum

Anser albifrons – Rhinonyssus rhinolethrum

A. anser – Rhinonyssus levinseni, Rh. rhinolethrum

A. erythropus – Rhinonyssus rhinolethrum

A. rossii – Rhinonyssus rhinolethrum

Aythya affinis – Rhinonyssus rhinolethrum

A. baeri – Rhinonyssus rhinolethrum (= nyrocae)

A. ferina – Rhinonyssus rhinolethrum (= anatidae)

A. fuligula – Rhinonyssus rhinolethrum (= nyrocae = anatidae)

A. marila – Rhinonyssus marilae

A. nyroca – Rhinonyssus rhinolethrum (= nyrocae)

Branta canadensis – Rhinonyssus rhinolethrum

B. hutchinsii – Rhinonyssus rhinolethrum

Bucephala albeola – Rhinonyssus rhinolethrum

B. clangula – Rhinonyssus rhinolethrum

Callonetta leucophrys – Rhinonyssus rhinolethrum

Casarca ferruginea – Rhinonyssus rhinolethrum

Chen hyperborea – Rhinonyssus rhinolethrum

Ch. coerulescens – Rhinonyssus rhinolethrum

Clangula hyemalis – Rhinonyssus clangulae, Rh. rhinolethrum

Cygnus columbianus – Rhinonyssus rhinolethrum

C. melanocoryphus – Rhinonyssus rhinolethrum

Dendrocygna arcuata – Rhinonyssus rhinolethrum

D. bicolor – Rhinonyssus rhinolethrum

D. viduata – Rhinonyssus rhinolethrum

Melanitta deglandi – Rhinonyssus rhinolethrum

Mergus albellus – Rhinonyssus rhinolethrum

M. merganser – Rhinonyssus rhinolethrum

Metopiana peposaca – Rhinonyssus rhinolethrum

Netta [Metopiana] peposaca – Rhinonyssus rhinolethrum

N. rufina – Rhinonyssus rhinolethrum (= nyrocae)

Plectropterus gambenis – Rhinonyssus rhinolethrum

Polysticta stelleri – Rhinonyssus levinseni, Rh. polystictae

Sarkidiornis melanotos – Rhinonyssus rhinolethrum

Somateria fischeri – Rhinonyssus levinseni

S. mollissima – Rhinonyssus levinseni, Rh. polystictae

S. spectabilis – Rhinonyssus levinseni

Spatula clypeata – Rhinonyssus rhinolethrum

Tadorna tadorna – Rhinonyssus rhinolethrum

Apodiformes

Fam. Apodidae

Apus affinis – Ptilonyssus strandtmanni, P. wilsoni

A. apus – Ptilonyssus strandtmanni

A. caffer streubeli – Ptilonyssus strandtmanni, Rhinonyssus apus

Collocalia esculenta – Ptilonyssus wilsoni

Fam. Trochilidae

Amazilia chionopectus – Ptilonyssus mariacastroae

A. yucatanensis – Ptilonyssus mariacastroae

Campylopterus largipennis – Tinaminyssus saucerottiae

Chrysolampis mosquitus – Ptilonyssus mariacastroae

Florisuga mellivora – Tinaminyssus squamosus

Glaucis hirsuta – Trochilonyssus trinitatis

Saucerottia tobaci – Ptilonyssus mariacastroae, Tinaminyssus saucerottiae

Sericotes hoplosericeus – Tinaminyssus squamosus

Trochilidae – Sternostoma mortelmansi

Caprimulgiformes

Fam. Caprimulgidae

Caprimulgus carolinensis –Vitznyssus [Astridiella] niztschi

C. europaeus – Vitznyssus caprimulgi, V. scotornis, Vitznyssus tsachevi

C. macrurus bimaculatus – Vitznyssus [Astridiella] nitzschi

C. nigrescens – Vitznyssus [Astridiella] scotornis

C. tristigma – Vitznyssus [Astridiella] scotornis

C. tristis – Vitznyssus [Astridiella] scotornis

Chordeiles minor – Ptilonyssus sp., Vitznyssus erici

Phalaenoptilus nuttalii – Vitznyssus [Astridiella] nitzschi

Scotornis fossii welwitschii – Vitznyssus caprimulgi, V. scotornis

Cathartiformes

Fam. Catarthidae

Cathartes aura – Ptilonyssus ohioensis

Coragyps atratus foetens – Ptilonyssus donatoi

Charadriiformes

Fam. Alcidae

Cerorhinca monocerata – Rhinonyssus caledonicus

Uria grylle – Rhinonyssus caledonicus

Fam. Charadriidae

Belenopterus chilensis – Rhinonyssus belenopteri

Charadrius alexandrinus – Rhinonyssus coniventris, Rh. echinipes, Rh. minutus, Rh. shcherbininae

Ch. dubius – Rhinonyssus dubius

Ch. hiaticula – Rhinonyssus coniventris, Rh. echinipes, Rh. minutus, Sternostoma minutus

Ch. melanops – Rhinonyssus himantopus

Ch. melodus – Rhinonyssus echinipes

Ch. morinellus – Charadrinyssus [Larinyssus] charadrius

Ch.[Leucopolius] pecuarius – Rhinonyssus minutus leucopolius

Ch. placidus japonicus – Rhinonyssus himatopus

Ch. pluvialis – Pluvialis dominica

Ch. tricollaris – Rhinonyssus himantopus, Rh. strandtmanni

Ch. vociferus – Rhinonyssus strandtmanni

Ch. wilsonia – Rhinonyssus echinipes

Chettusia gregaria – Rhinonyssus chettusiae

Erythrogonys cinctus – Rhinonyssus himantopus

Hoplopterus [Vanellus] armatus – Rhinonyssus sp.

Lobibyx miles – Rhinonyssus himantopus

L. novaehollandiae – Rhinonyssus himantopus

Pluvialis dominica – Rhinonyssus pluvialis

P. squatarola – Rhinonyssus echinipes

Vanellochettusia leucura – Rhinonyssus vanellochettusiae

Vanellus [sub syn. Hemiparra] crassirostris – Rhinonyssus himantopus

Vanellus [Afribyx] senegallus – Rhinonyssus afribyx

Vanellus [Stephanibyx] lugubris – Rhinonyssus afribyx

V. vanellus – Rhinonyssus dobromiri, Rh. vanellus

Fam. Glareolidae

Galachrysia cinerea – Larinyssus benoiti

Glareola nordmanni – Larinyssus benoiti

G. pratincola – Larinyssus benoiti, L. orbicularis

Stiltia isabella – Larinyssus benoiti

Fam. Laridae

Larus argentatus – Larinyssus orbicularis, Sternostoma sternahirundo

L. atricilla – Larinyssus orbicularis, Sternostoma boydae

L. californicus – Sternostoma boydae

L. delawarensis – Larinyssus orbicularis, Sternostoma boydae

L. dominicanus – Larinyssus orbicularis

L. glaucescens – Sternostoma boydae

L. minutus – Larinyssus orbicularis

L. novaehollandiae – Larinyssus orbicularis

L. occidentalis – Sternostoma boydae

Thalasseus [sandvicensis] acuflavidus – Sternostoma sp. (hutsoni ?), Larynyssus orbicularis

Fam. Rynchopidae

Rynchops nigra – Larinyssus orbicularis

Fam. Sternidae

Anous minutus – Larinyssus orbicularis

Chlidonias hybrida – Sternostoma boydae, S. epistomata

Ch. leucoptera – Larinyssus orbicularis, Sternostoma boydae

Ch. nigra – Larinyssus orbicularis, Sternostoma epistomata

Gelochelidon nilotica – Larinyssus orbicularis

Onychoprion [Sterna] fuscatus fuscatus – Larinyssus sterna, Sternostoma boydae

Sterna fuscata – Larinyssus orbicularis

S. hirundinacea – Larinyssus orbicularis

S. hirundo – Larinyssus iohanssenae, L. orbicularis, L. substerna, Sternostoma sternahirundo

S. maxima – Larinyssus orbicularis

Thalasseus [Sterna] bergii – Larinyssus orbicularis

Thalasseus superciliaris – Sternostoma boydae

Fam. Recurvirostridae

Himantopus himantopus – Rhinonyssus himantopus

H. leucocephalus – Rhinonyssus himantopus

H. mexicanus – Rhinonyssus himantopus

Recurvirostra americana – Rhinonyssus himantopus

Fam. Scolopacidae

Actitis macularia – Rhinonyssus spinactitis, Sternostoma boydae

Arenaria interpres – Rhinonyssus coniventris, Sternostoma boydae

Calidris alpina – Rhinonyssus neglectus

C. canutus – Rhinonyssus coniventris, Rh. neglectus

C. maritima – Rhinonyssus coniventris

Crocethia alba – Sternostoma boydae

Ereunetes pusillus – Sternostoma boydae

Phalaropus lobatus – Sternostoma boydae

Philomachus pugnax – Sternostoma boydae

Rhyacophilus glareola – Sternostoma boydae

Tringa erythropus – Sternostoma boydae

T. glareola – Rhinonyssus neglectus, Rh. tringae

T. melanoleuca – Rhinonyssus tringae

Tringa [Totanus] nebularia – Sternostoma boydae

T. ochropus – Rhinonyssus neglectus, Rh. tringae, Sternostoma boydae

T. totanus – Rhinonyssus tringae, Sternostoma boydae, S. strandtmanni

Coliiformes

Fam. Coliidae

Colius striatus kiwuensis – Sternostoma colii

Columbiformes

Fam. Columbidae

Chalcophaps chrysochlora – Tinaminyssus hirtus

Ch. indica indica – Tinaminyssus hirtus

Ch. stephani – Tinaminyssus hirtus

Columba fasciata – Tinaminyssus juxtamelloi

C. livia – Tinaminyssus columbae, T. melloi, T. streptopeliae

C. livia domestica – Tinaminyssus columbae

C. norfolciensis – Tinaminyssus melloi

C. oenas – Tinaminyssus melloi

C. palumbus – Tinaminyssus melloi, T. minisetosum, T. sartbaevi

Columbigallina passerina – Tinaminyssus zenaidurae

C. talpacoti – Tinaminyssus zenaidurae

Columbina picula –

Ducula pinon – Tinaminyssus gourae

D. spilorrhoa – Tinaminyssus welchi

Geopelia cuneata – Tinaminyssus geopeliae

G. humeralis – Tinaminyssus geopeliae

G. placida – Tinaminyssus geopeliae

G. striata striata – Tinaminyssus geopeliae

Geophaps scripta – Tinaminyssus ocyphabus

Geotrygon chrysia – Tinaminyssus carapachibeyus, T. geotrygoni

Goura cristata – Tinaminyssus gourae

Leptotila verreauxi – Tinaminyssus cunhai, T. alexfaini

Leucosarcia melanoleuca – Tinaminyssus melloi

Macropygia phasianella – Tinaminyssus macropygiae

Megaloprepia magnifica – Tinaminyssus megaloprepiae

Metropelia caeciliae – Tinaminyssus metropeliae

Myristicivora spilorrhoa – Tinaminyssus myristicivorae, T. welchi

Ocyphaps lophotes – Tinaminyssus columbiae, T. ocyphabus

Oena c. capensis – Tinaminyssus oenae

Oreopeleia montana – Tinaminyssus castroae

Phapitreron brevirostris – Tinaminyssus peleiae

Phaps chalcoptera – Tinaminyssus phabus

Ptilinopus coronulatus – Tinaminyssus ptilinopi

P. ornatus – Tinaminyssus ptilinopi

P. perlatus – Tinaminyssus ptilinopi

P. superbus – Tinaminyssus ptilinopi

P. purpuratus – Tinaminyssus tetae

Sphenurus sieboldii sororius – Tinaminyssus phassae

Streptopelia capicola – Tinaminyssus melloi

S. chinensis – Tinaminyssus melloi

S. d. decaocto – Tinaminyssus streptopeliae

S. o. orientalis – Tinaminyssus columbae, T. melloi, T. streptopeliae

S. semitorquata – Tinaminyssus columbae, T. streptopeliae

S. turtur – Tinaminyssus senotrisovae, T. streptopeliae, T. streptopelioides

Treron c. calva – Tinaminyssus treronis

Turtur afer – Tinaminyssus turturi

Zenaida asiatica – Tinaminyssus triangulus

Z. macroura – Tinaminyssus zenaidurae, T. melloi

Coraciiformes

Fam. Coraciidae

Coracias caudata – Tinaminyssus buteonis

C. garrulus – Ptilonyssus lanii

Fam. Halcyonidae

Dacelo gigas – Sternostoma cooremani, Tinaminyssus daceloae

D. leachii – Sternostoma cooremani, Tinaminyssus daceloae

Halcyon chelicuti – Tinaminyssus zumpti

H. chloris – Tinaminyssus halcyonus

H. chloris humii – Sternostoma cooremani halcyoni

H. leucocephala – Tinaminyssus schoutedeni

H. macleayii – Tinaminyssus halcyonus

H. pyrrhopygius – Tinaminyssus halcyonus

H. sanctus – Tinaminyssus halcyonus

H. smyrnensis – Tinaminyssus schoutedeni

Melittophagus bullocki – Ptilonyssus triscutatus

M. lafresnayi oreobates – Ptilonyssus melittophagi, Sternostoma cooremani

M. pusillus – Ptilonyssus melittophagi

Merops apiaster – Ptilonyssus triscutatus, Sternostoma cooremani

M. nubicoides – Sternostoma cooremani

M. nubicus – Ptilonyssus triscutatus

M. orientalis – Sternostoma cooremani

M. ornatus – Sternostoma cooremani

M. persicus – Ptilonyssus triscutatus

M. superciliosus – Ptilonyssus triscutatus

M. viridis – Ptilonyssus triscutatus

Tanysiptera danae – Tinaminyssus tanysipterae

T. galatea – Tinaminyssus tanysipterae

T. sylvia – Tinaminyssus tanysipterae

Fam. Phoeniculidae

Phoeniculus purpureus – Sternostoma sinense

Cuculiformes

Fam. Cuculidae

Cacomantis variolosus – Sternostoma ziegleri

Chalcites meyeri – Sternostoma cordiscutata

Chrysococcyx caprius – Ptilonyssus ploceanus, Sternostoma cuculorum

Clamator levaillanti – Sternostoma cuculorum

Crotophaga sulcifrons [Crotophagidae ?] – Sternostoma crotophagae

Cuculus canorus – Sternostoma cuculorum, S. zini

C. solitarius – Sternostoma cuculorum

Phaenicophaeus chlorophaeus – Sternostoma cuculorum

Falconiformes

Fam. Falconidae

Falco berigora – Ptilonyssus cerchneis

F. cenchroides – Ptilonyssus cerchneis

F. naumanni – Ptilonyssus cerchneis

F. sparverius – Ptilonyssus cerchneis

F. tinnunculus – Ptilonyssus cerchneis

F. [Cerchneis] tinnunculus rufescens – Ptilonyssus cerchneis

Milvago chimachima chimachima – Ptilonyssus souzai

Galliformes

Fam. Phasianidae

Alectoris chukar – Sternostoma francolini

A. rufa – Sternostoma guevarai

Arborophila brunneopectus erythrophys – Sternostoma furmani

Francolinus coqui – Sternostoma francolini

Perdix daurica – Ptilonyssus motacillae, Sternostoma francolini

P. perdix – Sternostoma francolini, S. guevarai

Fam. Cracidae

Penelope sp. – Tinaminyssus camargoi

Gruiformes

Fam. Rallidae

Amaurornis phoenicurus chinensis – Rallinyssus amaurornis

Fulica americana – Rallinyssus caudistigmus, Rhinonyssus rhinolethrum

F. armillata – Rallinyssus caudistigmus

F. atra – Rallinyssus caudistigmus, Sternostoma fulicae

F. cristata – Rallinyssus caudistigmus

Gallinula chloropus – Rallinyssus caudistigmus (= caspicus), R. gallinulae, Sternostoma fulicae, S. strandtmanni

G. chloropus indica – Rallinyssus caudistigmus

Hypotaenidia philippensis – Rallinyssus gallinulae

Ortygonax nigricans – Rallinyssus trappi

Poliolimnas cinereus collingwoodi – Rallinyssus amaurornis

Porphyrio porphyrio (= melanotus, = poliocephalus) – Rallinyssus gallinulae

Porzana carolina – Ptilonyssus icteridius, Rallinyssus sorae

P. f. fusca – Rallinyssus limnocoracis

P. parva – Rallinyssus gallinulae

P. pusilla – Rallinyssus gallinulae

Porzana tabuensis filipina – Rallinyssus porzanae

Rallina eurizonoides eurizonoides – Rallinyssus rallinae

Rallus aquaticus – Rallinyssus verheyeni

R. elegans – Rallinyssus caudistigmus, R. verheyeni

R. limicola – Rallinyssus verheyeni

R. pectoralis – Rallinyssus gallinulae

Otidiformes

Fam. Otididae

Afrotis afra afra – Vitznyssus [Astridiella] afrotis

Choriotis kori – Vitznyssus [Astridiella] neotis

Neotis cafra – Vitznyssus [Astridiella] neotis

Otis tarda – Vitznyssus vitzthumi, Rhinonyssus minutus

Tetrax tetrax – Vitnyssus tetragis

Passeriformes

Fam. Acanthizidae

Acanthiza reguloides – Ptilonyssus gerygonae

Gerygone palpebrosa – Ptilonyssus gerygonae

Fam. Acrocephalidae

Acrocephalus arundinaceus – Ptilonyssus acrocephali, Sternostoma tracheacolum

A. australis – Ptilonyssus acrocephali

A. schoenobaenus – Ptilonyssus acrocephali, P. schoenobaeni

A. scirpaceus – Ptilonyssus acrocephali

Fam. Alaudidae

Alauda arvensis – Ptilonyssus [Astridiella] alaudae, Ptilonyssus capitatus, P. schumili, Sternostoma batis

Ammomanes deserti – Ptilonyssus ammomani

Calandrella cinerea – Ptilonyssus [Astridiella] calandrellae, Sternostoma isabelae

Eremophila alpestris – Ptilonyssus capitatus, P. eremophilae, Sternostoma batis, S. gallowayi

Galerida cristata = Ptilonyssus schumili

Melanocorypha bimaculata – Ptilonyssus [Astridiella] melanocoryphae

Fam. Bombycillidae

Bombycilla cedrorum – Ptilonyssus bombycillae

B. garrulus – Ptilonyssus bombycillae

Fam. Calcariidae

Calcarius ornatus – Ptilonyssus morofskyi

C. l. lapponicus – Ptilonyssus emberizae

Plectrophenax nivalis – Ptilonyssus emberizae, P. morofskyi, P. nivalis

Fam. Campephagidae

Pericrocotus roseus divaricatus – Ptilonyssus [= Paraneonyssus] pericrocoti, Ruandanyssus terpsiphone

Fam. Cardinalidae

Passerina cyanea – Ptilonyssus ludovicianus

Pheucticus ludovicianus – Ptilonyssus japuibensis, P. ludovicianus

Fam. Cettiidae

Cettia cetti – Ptilonyssus gilcolladoi

Fam. Chloropseidae

Chloropsis cochinchinensis – Ptilonyssus chloropsicola

Fam. Cisticolidae

Apalis flavida caniceps – Ptilonyssus elongatus

Apalis flavida – Sternostoma cisticolae

Camaroptera brevicaudata – Ptilonyssus aureliani

Prinia leucopogon reichenowi – Ptilonyssus elongatus

Scotocerca inquieta – Ptilonyssus motacillae

Fam. Corvidae

Aphelocoma coerulescens – Ptilonyssus perisorei

Cyanocitta stelleri – Ptilonyssus perisorei

Garrulus glandarius – Ptilonyssus nucifragae

Nucifraga caryocatactes – Ptilonyssus nucifragae

Perisoreus canadensis – Ptilonyssus perisorei

P. infaustus – Ptilonyssus nucifragae

Fam. Dicruridae

Chibia bracteata – Ptilonyssus dicruri

Dicrurus adsimilis – Ptilonyssus dicruri

D. hottentottus – Ptilonyssus dicruri

D. leucophaeus stigmatops – Ptilonyssus dicruri

D. macrocercus cathoecus – Ptilonyssus dicruri

D. remifer – Ptilonyssus dicruri

Fam. Emberizidae

Calidris a. alpine – Ptilonyssus emberizae

Cyanocompsa brissonii – Sternostoma tracheacolum

Emberiza bruniceps – Ptilonyssus emberizae

E. c. calandra – Ptilonyssus emberizae

E. cia – Ptilonyssus emberizae

E. c. citrinella – Ptilonyssus emberizae

E. flaviventris – Ptilonyssus fringillicola

E. hortulana – Ptilonyssus emberizae

E. schoeniclus – Ptilonyssus emberizae, P. sairae

E. spodocephala personata – Ptilonyssus emberizae

E. sulphurata – Ptilonyssus emberizae

Paroaria coronata – Ptilonyssus sairae, Sternostoma pirangae

Pheucticus ludovicianus – Ptilonyssus japuibensis

Ph. melanocephalus – Ptilonyssus sp.

Piranga ludoviciana – Ptilonyssus pirangae

Fam. Estrildidae

Amadina erythrocephala – Ptilonyssus estrildicola

Batis capensis – Ptilonyssus estrildicola

Erythrura gouldiae – Ptilonyssus neochmiae

Euschistospiza dybovskii – Ptilonyssus astridae

Lagonosticta rhodopareia – Ptilonyssus estrildicola

L. rubricata – Ptilonyssus estrildicola

Lonchura pectoralis – Ptilonyssus emberizae

Poephila acuticauda – Ptilonyssus neochmiae

P. personata – Ptilonyssus astridae

Pytilia melba – Ptilonyssus estrildicola

Taeniopygia castanotis – Ptilonyssus estrildicola

Uraeginthus angolensis – Ptilonyssus estrildicola

U. bengalus ugandae – Ptilonyssus estrildicola

Fam. Fringillidae

Carduelis [Acanthis] flavirostris – Ptilonyssus carduelis

C. [Acanthis] c. cannabina – Ptilonyssus carduelis, P. morofskyi

C. carduelis – Ptilonyssus motacillae

C. c. carduelis – Ptilonyssus fringillicola, P. pygmaeus

C. flammea – Ptilonyssus carduelis, P. morofskyi

C. c. subcaniceps – Ptilonyssus pygmaeus

C. tristis – Ptylonyssus morofrskyi, P. spini

Carpodacus erythrinus – Ptilonyssus carduelis

C. mexicanus – Ptilonyssus icteridius

C. mexicanus centralis – Ptilonyssus hoffmannae

C. purpureus – Ptilonyssus plesiotypicus

C. rhodochlamys – Ptilonyssus carduelis

C. roseus – Ptilonyssus nudus

C. rubicilla – Ptilonyssus carduelis

Chloris chloris – Ptilonyssus chloris, P. pygmaeus, Sternostoma tracheacolum

Chloris sinica minor – Ptilonyssus sairae

Cocothraustes coccothraustes – Ptilonysus coccothraustis, Sternostoma tracheacolum

C. vespertinus – Ptilonysus coccothraustis

Fringilla coelebs – Ptilonyssus emberizae, P. fringillae, P. hirsti

Leucosticte brandti – Ptilonyssus carduelis

L. nemoricola – Ptilonyssus motacillae

Loxia curvirostra – Ptilonyssus carduelis, P. morofskii, Sternostoma batis, S. cryptorhyncum, S. loxiae

L. leucoptera – Ptilonyssus carduelis, Sternostoma cryptorhyncum

Pinicola enucleator – Ptilonyssus carduelis, P. pinicola, Sternostoma cryptorhynchum

Pyrrhula pyrrhula – Ptilonyssus fringillicola

P. pyrrhula coccinea – Ptilonyssus fringillicola

Richmondena cardinalis – Ptilonyssus richmondenae

Serinus atrogularis – Ptilonyssus fringillicola

S. canaria – Ptilonyssus fringillicola, Sternostoma marchae, S. tracheacolum

S. pusillus – Ptilonyssus motacillae

S. serinus – Ptilonyssus morofskyi

Spinus spinus – Ptilonyssus spini

S. [Poliospiza] striolata kivuensis – Ptilonyssus fringillicola

Fam. Furnariidae

Certhiaxis cinnamomea – Ptilonyssus certhiaxicola

Dendrocolaptes certhia – Ptilonyssus dendrocolapticola

Glyphorynchus spirurus – Ptilonyssus angustirostris

Myrmotherula axillaris – Ptilonyssus inornatus

Myrmotherula hauxwelli – Ptilonyssus myrmotherulae

M. longipennis – Ptilonyssus inornatus

M. menetriesi – Ptilonyssus inornatus

Philydor lichtensteini – Ptilonyssus travassosfilhoi

Sittasomus griseicapillus sylviellus – Ptilonyssus rabeloi

Synallaxis rutilans – Ptilonyssus certhiaxicola

Xenops minutus – Ptilonyssus xenops

Xiphorhynchos spixii – Ptilonyssus xyphorhynchus

Fam. Grallinidae

Grallina cyanoleuca – Ptilonyssus grallinae

Struthidea cinerea – Ptilonyssus struthideae

Fam. Hirundinidae

Atticora melanoleuca – Ptilonyssus echinatus

Delichon urbica – Ptilonyssus echinatus, Sternostoma tracheacolum

Hirundo aethiopica – Ptilonyssus echinatus, P. angrensis, Sternostoma hirundo

H. neoxena – Ptilonyssus angrensis

H. nigrita – Ptilonyssus echinatus

H. pyrrhonota – Ptilonyssus echinatus

H. rustica – Ptilonyssus echinatus [=Hapalognatha prima], [Hapalognatha platytrichia], Ptilonyssus emberizae, Sternostoma hirundinis, S. hirundo

H. smithi – Sternostoma hirundinis, S. hirundo

Hylochelidon ariel – Ptilonyssus echinatus

H. nigricans – Ptilonyssus echinatus

Iridoprocne bicolor – Sternostoma tracheacolum

Petrochelidon pyrrhonota – Ptilonyssus echinatus

P. spilodera – Ptilonyssus echinatus

Progne chalybea domestica – Ptilonyssus chalybeaedomesticae, P. angrensis

P. subis – Ptilonyssus angrensis, Sternostoma hirundinis

Psalidoprocne albiceps – Ptilonyssus psalidoprocnei, Sternostoma hirundinis

Ptyonoprogne rupestris – Ptilonyssus [Cas] angrensis

Riparia riparia – Ptilonyssus [Cas] angrensis, P. echinatus, P. ripariae, P. sialiphilus, Sternostoma hirundinis, S. tracheacolum

Stelgidopteryx ruficollis – Ptilonyssus echinatus

Tachycineta albilina – Sternostoma hirundinis

T. bicolor – Ptilonyssus echinatus, P. tachycinetae

Fam. Icteridae

Agelaius icterocephalus – Ptilonyssus agelaii, Sternostoma strandtmanni

A. phoeniceus – Ptilonyssus icteridius

A. tricolor – Sternostoma strandtmanni, S. tracheacolum

Cacicus haemorrhow – Passeronyssus zeferinoi

Chrysomus ruficapillus – Ptilonyssus icteridius, P. sairae, Sternostoma strandtmanni

Dives atroviolaceus – Ptilonyssus insularis cubanus

Euphagus cyanocephalus – Ptilonyssus echinatus

E. carolinus – Ptilonyssus icteridius

Gnorimopsar chopi chopi – Ptilonyssus zeferinoi

Icterus bullocki – Sternostoma tracheacolum

I. galbula – Ptilonyssus icteridius

I. nigrogularis – Ptilonyssus icteridius, Sternostoma strandtmanni

Molothrus ater – Ptilonyssus icteridius, P. japuibensis, P. richmondenae, Sternostoma tracheacolum

Molothrus ater californicus – Ptilonyssus icteridius

Quiscalus lugubris – Ptilonyssus icteridius, Sternostoma quiscali, S. strandtmanni

Qu. niger caribaeus – Ptilonyssus insularis insularis, Sternostoma quiscali

Qu. quiscala – Ptilonyssus icteridius, P. japuibensis, Sternostoma tracheacolum

Sturnella [Leistis militares] superciliaris – Ptilonyssus zeferinoi

Sturnella magna – Ptilonyssus icteridius, Sternostoma tracheacolum

S. neglecta – Ptilonyssus icteridius

Xanthocephalus xanthocephalus – Ptilonyssus icteridius

Fam. Irenidae

Irena puella malayensis – Ptilonyssus malaysiae

Fam. Laniidae

Eurocephalus anguitimens – Sternostoma eurocephali

Lanius collaris – Ptilonyssus lanii

L. collurio – Ptilonyssus lanii, Sternostoma cuculorum [laniorum]

L. cristatus – Ptilonyssus lanii

L. ludovicianus – Ptilonyssus lanii, Sternostoma cuculorum [laniorum]

L. minor – Ptilonyssus lanii, Sternostoma cuculorum [laniorum]

L. nigriceps longicaudatus – Ptilonyssus orientalis

Urolestes melanoleucus – Sternostoma cuculorum urolestis

Fam. Malaconotidae

Dryoscopus cubla – Ptilonyssus dryoscopi

Tchagra senegala – Ptilonyssus dryoscopi

Teloporus dohertyi – Ptilonyssus capitatus

T. zeylonus – Ptilonyssus capitatus

Fam. Maluridae

Malurus melanocephalus – Ptilonyssus maluri

Fam. Megaluridae

Bradypterus baboecala centralis – Ptilonyssus bradypteri

Locustella certhiola – Locustellonyssus sibiricus

L. fasciolata – Locustellonyssus amurensis

Megalurus timoriensis – Ptilonyssus dolicaspis, Passeronyssus megaluri

Fam. Meliphagidae

Anthochaera chrysoptera – Ptilonyssus myzanthae

Conopophila rufogularis – Ptilonyssus conopophilae

Entomyzon cyanotis – Ptilonyssus philemoni

Grantiella picta – Ptilonyssus conopophilae

Gliciphila indistincta – Sternostoma gliciphilae

Manorina flavigula – Ptilonyssus pentagonicus

Meliphaga chrysops – Ptilonyssus meliphagae

M. flava – Ptilonyssus stomioperae

M. gracilis – Ptilonyssus philemoni

M. notata – Ptilonyssus philemoni

Melithreptus albogularis – Ptilonyssus meliphagae

Myzantha flavigula – Ptilonyssus myzanthae, P. philemoni

M. melanocephala – Ptilonyssus myzanthae

Philemon corniculatis – Ptilonyssus philemoni

Ph. citreogularis – Ptilonyssus philemoni

Plectorhyncha lanceolata – Ptilonyssus philemoni

Ramsayornis fasciatus – Ptilonyssus condyloxa, Sternostoma gliciphilae

Stomiopera unicolor – Ptilonyssus stomioperae

“Honey eater” – Ptilonyssus balimoensis, P. missimensis

Fam. Mimidae

Dumetella carolinensis – Ptilonyssus euroturdi, Sternostoma dumetellae, S. kelloggi

Mimus polyglottos – Ptilonyssus mimi, Sternostoma kelloggi

Toxostoma rufum – Ptilonyssus toxostomae, Sternostoma kelloggi

Fam. Monarchidae

Arses kaupi – Ptilonyssus terpsiphonei

Myiagra rubecula – Ruandanyssus terpsiphonei

Monarcha melanopsis – Ptilonyssus terpsiphonei

M. trivirgata – Ptilonyssus monarchae, Ptilonyssus terpsiphonei, Ruandanyssus terpsiphonei

Terpsiphone viridis kivuensis – Ptilonyssus terpsiphonei, Ruandanyssus terpsiphonei

Fam. Motacillidae

Anthus australis – Ptilonyssus motacillae

A. campestris – Ptilonyssus capensis

A. pratensis – Ptilonyssus capensis, P. fringillae

A. spinoletta – Ptilonyssus capensis, P. motacillae

A. trivialis – Ptilonyssus capensis, P. motacillae

Ficedula albicollis – Ptilonyssus motacillae, Sternostoma ficedulae

F. hypoleuca – Ptilonyssus muscicapoides

Macronyx capensis – Ptilonyssus capensis

M. croceus – Ptilonyssus capensis, Sternostoma tracheacolum (= S. castroae)

Motacilla aguimp vidua – Ptilonyssus motacillae

M. alba – Ptilonyssus capensis, P. motacillae, [Hapalognatha langei]

M. capensis wellsi – Ptilonyssus motacillae

M. cinerea – Ptilonyssus motacillae

M. citreola – Ptilonyssus motacillae

M. flava – Ptilonyssus motacillae

M. tschutschensis macronyx – Ptilonyssus motacillae

Fam. Muscicapidae

Batis molitor puella – Sternostoma batis, S. giganteum

Carterornis leucotis – Ptilonyssus terpsiphonei

Cercomela familiaris modesta – Ptilonyssus motacillae

Copsychus malabaricus – Ptilonyssus dioptrornis

C. saularis musicus – Ptilonyssus dioptrornis

Cossypha [Bessonornis] archeri – Ptilonyssus dioptrornis

C. dichroa – Ptilonyssus dioptrornis

Cyanosylvia svecica – Ptilonyssus cyanosylviae

Dioptrornis fischeri toruensis – Ptilonyssus dioptrornis

Eopsaltria capito – Ptilonyssus dioptrornis

Erithacus rubecula – Ptilonyssus motacillae

Erythropygia coryphaeus – Ptilonyssus motacillae

Kittacincla malabarica – Passeronyssus kittacinclae

Luscinia c. cyanae – Ptilonyssus lusciniae

L. luscinia – Ptilonyssus lusciniae

L. svecica – Ptilonyssus motacillae

Muscicapa cyanomelana cyanomelana – Ptilonyssus dioptrornis

M. latirostris – Ptilonyssus muscicapae

M. rufigastra – Ptilonyssus dioptrornis

M. sibirica – Ptilonyssus muscicapae

M. striata – Ptilonyssus motacillae, P. muscicapae

Myagra rubecula – Ptilonyssus terpsiphonei

Myrmecocichla formicivora – Ptilonyssus motacillae

M. nigra – Ptilonyssus lusciniae

Oenanthe isabellina – Ptilonyssus motacillae

O. oenanthe – Ptilonyssus motacillae, Sternostoma batis

Phoenicurus erythronotus – Ptilonyssus motacillae

Ph. ochruros – Ptilonyssus motacillae phoenicuri

Piesorhynchus alecto – Ptilonyssus terpsiphonei

Seisura inquieta – Ptilonyssus terpsiphonei

Fam. Nectariniidae

Arachnothera longirostris – Ptilonyssus cinnyris, Sternostoma tracheacolum

Chalcomitra senegalensis aequatorialis – Ptilonyssus cinnyris

Cinnyris afra – Ptilonyssus cinnyris

C. regius kivuensis – Ptilonyssus cinnyris

C. cupreus septentrionalis – Sternostoma nectarinia

Chalcomitra senegalensis aequatorialis – Sternostoma nectarinia

Cyrtostomus frenatus – Ptilonyssus cinnyris

Dicaeum concolor olivaceum – Sternostoma nectarinia

Nectarinia jugularis flammaxillaris – Sternostoma tracheacolum

N. kilimensis – Ptilonyssus cinnyris

N. pulchella – Sternostoma nectarinia

N. purpureiventris – Sternostoma nectarinia

Fam. Neosittidae

Neositta striata – Sternostoma neosittae

Fam. Oriolidae

Oriolus flavocinctus – Ptilonyssus trouessarti

O. larvatus angolensis – Ptilonyssus orioli

O. oriolus – Ptilonyssus orioli

O. sagittatus – Ptilonyssus trouessarti

Sphecotheres flaviventris – Ptilonyssus sphecotheris

S. vieillotii – Ptilonyssus sphecotheris

Fam. Or thonychidae

Orthonyx temminckii – Ptilonyssus orthonychus

Fam. Pachycephalidae

Colluricincla harmonica – Ptilonyssus colluricinclae

C. megarhyncha – Ptilonyssus colluricinclae

Pachycephala griseiceps – Ptilonyssus colluricinclae, Ruandanyssus terpsiphonei

P. pectoralis – Ptilonyssus colluricinclae, P. motacillae, Ruandanyssus terpsiphonei

P. rufiventris – Ptilonyssus motacillae, Ruandanyssus terpsiphonei

Fam. Panuridae

Panurus biarmicus – Ptilonyssus motacillae

Fam. Paradisaeidae

Manucodia (Phonygammus) keraudrenii – Ptilonyssus buloloensis

Paradisaea reggiana – Ptilonyssus novaeguineae, Sternostoma constricta

Paradisornis rudolphi – Ptilonyssus paradisaeus

Ptiloris magnificus – Ptilonyssus novaeguineae

P. paradiseus – Ptilonyssus novaeguineae

P. victoriae – Ptilonyssus novaeguineae

Fam. Paramythiidae

Paramythia montium – Ptilonyssus domrowi

Fam. Paridae

Parus ater – Ptilonyssus pari

Parus ater insularis – Ptilonyssus pari

P. atricapillus subrhenanus – Ptilonyssus pari

P. bicolor – Ptilonyssus pirangae

P. carolinensis – Ptilonyssus sairae

P. caeruleus – Ptilonyssus mironovi, P. pari

P. cristatus mitratus – Ptilonyssus pari

P. gambeli – Ptilonyssus pari

P. insularis – Ptilonyssus pari

P. m. major – Ptilonyssus pari, P. sairae, P. sittae, Sternostoma tracheacolum

P. m. minor – Ptilonyssus pari

P. montanus – Ptilonyssus pari

P. varius varius – Ptilonyssus pari

Psaltriparus minimus – Ptilonyssus psaltriparus

Suthora webbiana – Ptilonyssus webbiana

Fam. Parulidae

Dendroica auduboni – Ptilonyssus sairae

D. coronata – Ptilonyssus japuibensis, P. morofskyi, P. sairae

D. erithacorides – Ptilonyssus sairae

D. magnolia – Ptilonyssus japuibensis

D. petechia – Ptilonyssus japuibensis, P. sairae (= constrictus), Sternostoma loxiae

D. pinus – Ptilonyssus japuibensis, P. morofskyi, P. sairae

D. striata – Ptilonyssus japuibensis

D. tigrina – Ptilonyssus japuibensis

Geothlypis philadelphia – Ptilonyssus sairae

G. trichas – Ptilonyssus morofskyi, P. sairae

Muniotilta varia – Ptilonyssus morofskyi, P. sairae, Sternostoma tracheacolum

Myioborus miniatus – Ptilonyssus sairae

Oreothlypis peregrina – Ptilonyssus sairae

O. ruficapilla – Ptilonyssus sairae

Seiurus aurocapillus – Ptilonyssus japuibensis

S. motacilla – Ptilonyssus morofskyi

S. noveboracensis – Ptilonyssus japuibensis, P. sairae (= constrictus), Sternostoma tracheacolum

Setophaga ruticilla – Ptilonyssus japuibensis, P. morofskyi

Teretistris fornsi – Ptilonyssus teretistris

Vermivora celata – Ptilonyssus japuibensis, P. sairae

V. peregrina – Ptilonyssus japuibensis, P. morofskyi

V. ruficapilla – Ptilonyssus sairae

Wilsonia canadensis – Ptilonyssus japuibensis

W. pusilla – Ptilonyssus morofskyi, P. sairae

Fam. Passerellidae

Calamospiza melanocorys – Ptilonyssus morofskyi

Junco hiemalis – Ptilonyssus japuibensis, P. morofskyi

Melospiza georgiana – Ptilonyssus morofskyi

M. lincolnii – Ptilonyssus morofskyi

M. melodia – Ptilonyssus morofskyi

Passerella iliaca – Ptilonyssus morofskyi

Spizella pallida – Ptilonyssus japuibensis

S. passerina – Ptilonyssus calvaria

Zonotrichia albicollis – Ptilonyssus japuibensis

Fam. Passeridae

Montifringilla nivalis – Ptilonyssus montifringillae

Passer domesticus – Ptilonyssus degtiarevae, P. hirsti, P. icteridius, P. nudus, Sternostoma cryptorhynchum, S. tracheacolum, Tinaminyssus melloi

P. griseus – Ptilonyssus hirsti, P. nudus

P. indicus – Ptolonyssus hirsti

P. montanus – Ptilonyssus hirsti, P. icteridius

P. ugandae griseus – Ptilonyssus hirsti

Petronia dentata – Ptilonyssus hirsti, P. ploceanus

Fam. Petroicidae

Petroica rosea – Ptilonyssus motacillae

Peneothello cyanus – Ptilonyssus radovskyi

Tregellasia leucops – Ptilonyssus radovskyi

Fam. Phylloscopidae

Phylloscopus borealis – Ptilonyssus acanthopneustes

Ph. collybita – Ptilonyssus phylloscopi, Sternostoma christinae

Ph. griseolus – Ptilonyssus phylloscopi

Ph. inornatus – Ptilonyssus isakovae, P. phylloscopi

Ph. t. trochilus – Ptilonyssus phylloscopi

Fam. Pipridae

Chiroxiphia pareola – Ptilonyssus chiroxiphiae

Manacus candei – Pellonyssus marui

M. manacus – Sternostoma piprae

Pipra erythrocephala – Pipronyssus manaci, Sternostoma [Sternoecius] piprae

P. iris – Pipronyssus manaci

P. pipra – Pipronyssus manaci

Schiffornis turdinus – Ptilonyssus sternostomicus, Sternostoma schiffornis

Fam. Pittidae

Pitta versicolor – Ptilonyssus pittae

Fam. Ploceidae

Amblyospiza albifrons montana – Ptilonyssus zumpti

Anaplectes rubriceps leuconotus – Ptilonyssus ploceanus

Euplectes ardeus – Ptilonyssus ploceanus

E. hordeaceus – Ptilonyssus ploceanus

E. macrourus – Ptilonyssus ploceanus

E. orix nigrifrons – Ptilonyssus ploceanus

Hyphanturgus ocularius – Ptilonyssus ploceanus

Neochmia albiventer – Ptilonyssus neochmiae

N. phaeton – Ptilonyssus neochmiae

Ploceus cucullatus – Ptilonyssus ploceanus

P. vitellinus – Ptilonyssus ploceanus

Poephila bichenovii – Ptilonyssus emberizae

P. gouldiae – Ptilonyssus emberizae

Quelea erythrops – Ptilonyssus ploceanus, Sternostoma sinense

Q. quelea – Sternostoma sinense

Textor atrogularis – Ptilonyssus ploceanus

Fam. Polioptilidae

Polioptila albiloris – Ptilonyssus sairae

Fam. Prunellidae

Prunella fulvescens – Ptilonyssus prunellae

P. m. modularis – Ptilonyssus prunellae

Fam. Ptilogonatidae

Phainopepla nitens – Ptilonyssus phainopeplae

Fam. Ptilonorhynchidae

Ailuroedus crassirostris – Ptilonyssus ailuroedi

Amblyornis macgregori – Ptilonyssus radovskyi

Chlamydera nuchalis – Ptilonyssus sphecotheris

Fam. Pycnonotidae

Andropadus latirostris eugenius – Ptilonyssus andropadi

Chlorocichla flavicollis pallidigula – Ptilonyssus chlorocichlae

Hypsipetes amaurotis amaurotis – Ptilonyssus hiyodori

H. flavala connectens – Ptilonyssus andropadi

H. [Isocincla] madagascariensis – Ptilonyssus madagascariensis

Phyllastrephus fischeri sucosus – Ptilonyssus andropadi

Ph. terrestris – Ptilonyssus phyllastrephi

Pycnonotus barbarus tricolor – Ptilonyssus pycnonoti

P. brunneus – Ptilonyssus nadchatrami

P. e. erythrophthalmus – Ptilonyssus nadchatrami

P. nigricans – Ptilonyssus pycnonoti

P. plumosus – Ptilonyssus pycnonoti malayi

Fam. Regulidae

Regulus calendula – Ptilonyssus acrocephali

R. ignicapillus – Ptilonyssus reguli

R. regulus – Ptilonyssus reguli

Fam. Rhipiduridae

Rhipidura leucophrys – Sternostoma cuculorum

Fam. Sittidae

Sitta europaea – Ptilonyssus pari, P. sittae

Fam. Sturnidae

Acridotheres tristis – Ptilonyssus elbeli, P. motacillae, Sternostoma bruxellarum

Aplonis metallica – Ptilonyssus sturnopastoris

Cinnyricinclus leucogaster – Ptilonyssus cinnyricincli, Sternostoma straeleni

C. leucogaster verreauxi – Sternostoma tracheacolum

Lamprocolius chalybaeus – Ptilonyssus tillae, Sternostoma sturnicola

L. purpureus – Sternostoma sturnicola

Lamprotornis australis – Ptilonyssus tillae

L. mevesii – Ptilonyssus tillae

L. purpuropterus – Sternostoma sturnicola

Pastor roseus – Ptilonyssus pastoris, Sternostoma bruxellarum

Sturnia sturnina – Ptilonyssus lobatus

Sturnopastor capensis – Ptilonyssus sturnopastoris

Sturnus javanicus grandis – Ptilonyssus elbeli, P. lobatus

S. nigricollis – Sternostoma bruxellarum

S. sturninus – Sternostoma cuculorum

S. tristis tristis – Ptilonyssus elbeli, P. lobatus, Sternostoma bruxellarum

S. vulgaris – Ptilonyssus elbeli, P. emberizae, P. euroturdi, P. langei, P. lanii, Sternostoma bruxellarum, S. technaui (= borceanum)

Fam. Sylviidae

Cinclorhamphus mathewsi – Ptilonyssus bradypteri

Hippolais icterina – Ptilonyssus calamocichlae hippolaisi

Parisoma sucaeruleum – Ptilonyssus ruanda

Sylvia a. atricapilla – Ptilonyssus ruandae sylviae

S. communis – Ptilonyssus ruandae sylviae, P. sylviicola

S. melanocephala – Ptilonyssus ubedai

Fam. Thamnophilidae

Hylophylax poecilonota – Ptilonyssus hoseini, P. hylophylax

Phlegopsis nigromaculata – Ptilonyssus hoseini, P. hylophylax

Pygiptila stellaris – Ptilonyssus pygiptilae

Sclateria naevia – Ptilonyssus sclateriae

Thamnophilus aethiops – Ptilonyssus thamnophili

Th. doliatus – Ptilonyssus thamnophili

Fam. Thraupidae

Chlorospingus ophthalmicus – Ptilonyssus pirangae, P. sairae

Cyanerpes caeruleus – Ptilonyssus amarali

C. cyanea – Ptilonyssus amarali

Piranga ludoviciana – Ptilonyssus icterius, P. pirangae

Procnias albus – Ptilonyssus stresemanni

Rhamphocelus bresilius dorsalis – Ptilonyssus japuibensis

Rh. carbo centralis – Ptilonyssus japuibensis

Sporophila americana – Ptilonyssus japuibensis

Tachyphonus rufus – Ptilonyssus japuibensis

T. surinamus – Ptilonyssus japuibensis

Tangara gyrola – Ptilonyssus sairae

T. mexicana – Ptilonyssus sp., Sternostoma tangarae

T. seledon – Ptilonyssus sairae

Thraupis palmarum – Ptilonyssus sp.

Volatinia jacarina – Ptilonyssus japuibensis

Fam. Timaliidae

Alcippe nipalensis – Ptilonyssus ruandae alcippei

Leiothrix lutea – Sternostoma sinense

Nothura maculosa – Tinaminyssus navajasi

Rhynchotus rufescens – Tinaminyssus serraoi

Tinamus solitarius – Tinaminyssus trappi

Turdoides jardinei emini – Sternostoma inflatum

Yuhina zantholeuca – Sternostoma cuculorum

Fam. Troglodytidae

Salpinctes obsoletus – Ptilonyssus salpinctis

Thryothorus ludovicianus – Ptilonyssus thryothori

Troglodytes aedon – Ptilonyssus vossi

T. troglodytes – Ptilonyssus troglodytis, Sternostoma kodrensis

Fam. Turdidae

Catharus dryas – Ptilonyssus euroturdi, Sternostoma hutsoni, Tinaminyssus ixoreus

C. minimus – Sternostoma spatulatum (= ? S. technaui)

C. [Hylocichla] mustelinus – Ptilonyssus euroturdi

C. [Hylocichla] ustulatus – Sternostoma cuculorum (= laniorum), Sternostoma hutsoni, S. technaui (= Sternostomoides spatulatum), Pteronyssus icteridius

Cyanosylvia svecica – Ptilonyssus motacillae

Enicurus schistaceus – Ptilonyssus enicuri

Erithacus akahige akahige – Ptilonyssus enicuri

Ixoreus naevius – Tinaminyssus ixoreus

Mimocichla plumbea – Ptilonyssus euroturdi

M. plumbea rubripes – Sternostoma orlandoi

M. plumbea schistacea – Sternostoma orlandoi

Monticola saxatilis – Sternostoma technaui (= Sternostomoides turdi)

Oreocincla dauma – Sternostoma technaui (= Sternostomoides turdi)

O. lunulata – Sternostoma dureni, S. technaui

Saxicola rubetra – Ptilonyssus motacillae

S. torquata axillaris – Ptilonyssus motacillae

Sialia currucoides – Ptilonyssus loxiae, P. sialiae

S. sialis – Sternostoma sialiphilus

Thamnolaea cinnamomeiventris – Ptilonyssus desfontainei

Turdoides melanops sharpei – Sternostoma dureni

T. jardinei emini – Sternostoma dureni

Turdus albicollis – Sternostoma hutsoni

T. dauma toratugumi – Rhinonyssus himantopus

T. e. ericetorum – Ptilonyssus euroturdi, Sternostoma technaui (= turdi = borceanum)

T. fumigatus – Ptilonyssus euroturdi, Sternostoma dureni, S. technaui (= Sternostomoides turdi)

T. iliacus – Ptilonyssus euroturdi, Sternostoma technaui (= S. turdi)

T. merula – Ptilonyssus euroturdi, S. technaui (= Sternostomoides turdi)

T. migratorius – Ptilonyssus euroturdi, P. cerchneis, Sternostoma technaui

T. musicus – Sternostoma technaui (= Sternostomoides turdi)

T. naumanni euromus – Ptilonyssus euroturdi, Sternostoma technaui

T. nudigenis – Ptilonyssus euroturdi, Sternostoma hutsoni, Sternostoma technaui (= Sternostomoides turdi)

T. olivaceus – Sternostoma technaui (= Sternostomoides turdi)

T. olivaceus graueri – Ptilonyssus afroturdi, Sternostoma dureni

T. philomelos – Ptilonyssus euroturdi, Sternostoma technaui (= Sternostomoides turdi)

T. pilaris – Sternostoma technaui (= Sternostomoides turdi), Ptilonyssus euroturdi

T. ruficollis – Ptilonyssus euroturdi, Sternostoma technaui (= Sternostomoides turdi)

T. rufiventris – Sternostoma hutsoni

T. torquatus – Sternostoma technaui (= Sternostomoides turdi = borceanum)

T. viscivorus – Sternostoma technaui (= Sternostomoides turdi), Ptilonyssus euroturdi

T. v. viscivorus – Ptilonyssus euroturdi, Sternostoma technaui (= Sternostomoides turdi)

Fam. Tyrannidae

Camptostoma imberbe – Ptilonyssus camptostoma

Colopteryx galeatus – Ptilonyssus colopteryx

Contopus caribaeus caribaeus – Ptilonyssus tyrannus [Tyranninyssus tyrannus caribaeus]

C. sordlidulus – Ptilonyssus tyrannus

Elaenia flavogaster – Ptilonyssus elaeniae, P. paranensis, P. werneri

E. obscura – Ptilonyssus werneri

Empidonax alnorum – Sternostoma longisetosae, Ptilonyssus tyrannus

E. flavescens – Sternostoma pencei

E. flaviventris – Ptilonyssus tyrannus

E. minimus – Ptilonyssus tyrannus [tyrannisoides], Sternostoma setifer

E. traillii – Sternostoma longisetosae, Ptilonyssus tyrannus

E. virescens – Sternostoma longisetosae

Empidonomus varius – Sternostoma longisetosae (= callithrix)

Fluvicola pica – Ptilonyssus flavicolae

Knipolegus cyanirostris – Sternostoma clementei

Machetornis rixosa – Ptilonyssus spinosus

Mitrephanes phaeocercus – Sternostoma darlingi

Muscivora forficata – Tyranninyssus spinosus

M. tyrannus – Sternostoma longisetosa

Myiarchus cinerascens – Ptilonyssus callinectoides

M. crinitus – Ptilonyssus callinectoides, P. icteridius, Sternostoma tracheacolum

M. ferox australis – Passeronyssus urieli

M. stolidus – Ptilonyssus myiarchi

M. tyrannulus – Ptilonyssus callinectoides

Myiobius barbatus – Ptilonyssus myiobii

Myiophobus fasciatus – Tyranninyssus myiophobi

Myiozetetes cayenensis – Tyranninyssus attilae

Nuttallornis borealis – Sternostoma tyrannus, Ptilonyssus tyrannus

Phylloscartes ventralis – Ptilonyssus enrietti

Pipromorpha macconnelli – Ptilonyssus pipromorphae

P. oleaginea – Ptilonyssus pipromorphae

Pitangus sulphuratus – Ptilonyssus spinosus, Sternostoma longisetosae

Platyrhinchus saturatus – Ptilonyssus platyrhinchus, P. thamnophili, Sternostoma hedonophilum

Pyrocephalus rubinus – Ptilonyssus fluvicolae, Sternostoma longisetosae

Rhynchocyclus olivaceus – Sternostoma tracheacolum

Sayornis nigricans – Sternostoma sayornis

S. phoebe – Sternostoma longisetosa

S. sayus – Tyranninyssus tyrannus [tyrannisoides]

Terenotriccus erythrurus – Ptilonyssus terenotricci

Tyrannus dominicensis – Ptilonyssus spinosus

T. melancholicus – Ptilonyssus spinosus, P. tyrannus, Sternostoma tracheacolum

T. tyrannus – Ptilonyssus spinosus, Ptilonyssus sp., Sternostoma longisetosa

T. verticalis – Ptilonyssus spinosus

Fam. Viduidae

Hypochera funerea nigerrima – Ptilonyssus lanii, P. nudus

Vidua macroura – Ptilonyssus astridae, P. nudus, P. viduicola

V. [Steganura] paradisea – Ptilonyssus steganurae

Fam. Vireonidae

Cyclarchis gujanensis – Ptilonyssus [Passeronyssus] hoseini

Vireo bellii – Ptilonyssus hoseini

V. flavifrons – Ptilonyssus vireonis

V. griseus – Ptilonyssus hoseini

V. olivaceus – Ptilonyssus vireonis

V. solitarius – Ptilonyssus vireonis

Fam. Zosteropidae

Zosterops lateralis – Sternostoma cuculorum (= zosteropus)

Z. palpebrosa japonica – Ptilonyssus ruandae

Pelecaniformes

Fam. Ardeidae

Ardea cinerea – Tinaminyssus belopolskii

A. melanocephala – Tinaminyssus belopolskii

A. purpurea – Tinaminyssus gerschi

A. purpurea manillensis – Tinaminyssus belopolskii

Bubulcus ibis – Tinaminyssus belopolskii, T. bubulci

Butorides striata – Tinaminyssus belopolskii

Butorides virescens – Tinaminyssus belopolskii

E. garzetta – Tinaminyssus belopolskii (= Neonyssus ardeae), T. egrettae

E. intermedia brachyrhyncha – Tinaminyssus bubulci

E. thula thula – Tinaminyssus belopolskii

Florida caerulea – Tinaminyssus belopolskii

Hydranassa tricolor – Tinaminyssus belopolskii

Ixobrychus minutus – Tinaminyssus ixobrychi

I. exilis – Tinaminyssus neoixobrychi

Leucophoyx thula – Tinaminyssus belopolskii

Melanophyx ardesiaea – Tinaminyssus belopolskii (= Neonyssus marcandrei)

Nycticorax nycticorax – Tinaminyssus belopolskii (= T. b. nycticoracis)

Syrigma sibilatrix – Tinaminyssus belopolskii

Piciformes

Fam. Bucconidae

Malacoptila rufa – Ptilonyssus malacoptilae

Fam. Capitonidae

Cyanops (= Megalaima) franklini minor – Tinominyssus malayi

Fam. Indicatoridae

Indicator indicator – Ptilonyssus indicatoris

Fam. Picidae

Centurus aurifrons – Sternostoma porteri

Colaptes auratus – Sternostoma porteri

Dendrocopos leucotos – Sternostoma hylandi

D. major – Sternostoma hylandi

D. pubescens – Sternostoma hylandi

Picoides pubescens – Sternostoma hylandi, Sternostoma porteri

P. villosus – Sternostoma porteri

Picus viridis – Sternostoma porteri

Sphyrapicus varius – Sternostoma porteri

Podicipediformes

Fam. Podicipedidae

Podiceps auritus – Rhinonyssus alberti, Rh. colymbicola

P. caspicus – Rhinonyssus alberti

P. cristatus – Rhinonyssus alberti, Rh. podicipedus

P. griseigena – Rhinonyssus alberti, Rh. colymbicola

P. nigricollis – Rhinonyssus colymbicola

P. ruficollis – Rallinyssus verheyeni, Rhinonyssus alberti, Rh. poliocephali

Podilymbus podiceps – Rhinonyssus podylimbi

Psittaciformes

Fam. Psittacidae

Psilopsiagon [Amoropsittaca] aymara – Sternostoma [= Rhinosterna] aymarae

Melopsittacus undulatus – Sternostoma tracheacolum

Fam. Psittaculidae

Aprosmictus erythropterus – Tinaminyssus aprosmicti

A. scapularis – Tinaminyssus aprosmicti

Barnardius barnardi – Tinaminyssus aprosmicti

Chalcopsitta scintillata – Tinaminyssus chalcoscittae

Charmosina papou – Tinaminyssus trichoglossi

Glossopsitta pusilla – Tinaminyssus trichoglossi

Neopsittacus pullicauda – Tinaminyssus neopsittaci

Platycercus adscitus – Tinaminyssus aprosmicti

Prioniturus [flavicans] montanus – Tinaminyssus prianituri

P. [discurus] platenae – Tinaminyssus prianituri

P. discurus whiteheadi – Tinaminyssus prianituri

Pseudeos fuscata – Tinaminyssus trichoglossi

Psittacella brehmi – Tinaminyssus neopsittaci

P. modesta – Tinaminyssus neopsittaci

Psittacula cyanocephala – Tinaminyssus psittaculae

P. krameri – Tinaminyssus baforti

Trichoglossus chlorolepidotus – Tinaminyssus trichoglossi

T. haematodus – Tinaminyssus trichoglossi

T. moluccanus – Tinaminyssus trichoglossi

King parrot (Alisterus ?) – Tinaminyssus alisteri

Fam. Cacatuidae

Cacatua roseicapilla – Tinaminyssus kakatuae

C. sulphurea occidentalis – Tinaminyssus kakatuae

Calyptorhynchus banksi – Tinaminyssus kakatuae

Pteroclidiformes

Fam. Pteroclididae

Pterocles gutturalis – Tinaminyssus pteroclesi

Sphenisciformes

Fam. Spheniscidae

Pygoscelis adeliae – Rhinonyssus sphenisci schelli

P. papua – Rhinonyssus sphenisci schelli

Spheniscus humboldti – Rhinonyssus sphenisci

S. magellanicus – Rhinonyssus sphenisci

Strigiformes

Fam. Strigidae

Aegolius acadicus – Rhinoecius aegolii

A. funereus – Rhinoecius aegolii

Asio capensis – Rhinoecius africanus

A. flammeus – Rhinoecius alifanovi

A. otus – Rhinoecius brikinboricus, Rh. oti, Sternostoma strigitis

Athene noctua – Rhinoecius bisetosus, Rh. subbisetosus

Bubo africanus – Zumptnyssus buboensis

B. scandiacus [= Nyctea scandiaca] – Rhinoecius nycteae

B. virginianus – Rhinoecius grandis

Glaucidium cuculoides brugeli – Rhinoecius bisetosus

Ninox connivens – Rhinoecius cooremani

N. novaeseelandiae – Rhinoecius cooremani

Otus bakkamoena – Rhinoecius cavannus

Pulsatrix melanonota koeniswaldiana – Rhinoecius grandis

Speotyto cunicularia – Rhinoecius bisetosus, Sternostoma augei

Strix nebulosa – Rhinoecius cooremani

S. varia – Rhinoecius cooremani

Surnia ulula – Rhinoecius aegolii

Fam. Tytonidae

Tyto alba affinis – Rhinoecius tytonis

Tinamiformes

Fam. Tinamidae

Crypturellus tataupa – Tinaminyssus crypturelli

Eudromia elegans – Tinaminyssus tinamicola

Nothura maculosa – Tinaminyssus navajasi

Rhynchotus rufescens – Tinaminyssus serraoi

Tinamus solitarius – Tinaminyssus trappi

Bibliography of Rhinonyssidae

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Amaral V. 1966. Mesonyssus marciae n. sp. (Acarina: Rhinonyssidae Vitz., 1935) parasita do beija-flor (Leuchocloris albicolis, Vieilot – Trochilidae). – Arqu. Inst. Biol., 33(4): 149–155.

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Amaral V. 1967b. Ptilonyssus chalybeaedomesticae n. sp. (Acarina: Rhinonyssidae Vitz., 1935) parasita da Andorinha azul (Progne chalybea domestica – Hirundinidae). – Arqu. Inst. Biol., 34(3): 169–174.

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Amaral V. 1967d. Ptilonyssus echinatus subsp. taperaefuscae n. subsp. (Acarina: Rhinonyssidae) lista de hospedeiros Hirundinidae. – Arqu. Inst. Biol., 34(3): 199–211.

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Berlese A., E. L. Trouessart. 1889. Diagnoses d’acariens nouveaux ou peu connus. – Bull. Biblioth. Sci. Ouest., 2(2): 126–130.

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Beron P. 1975. Contribution to the study of nasal mites (fam. Cytoditidae, Halarachnidae and Rhinonyssidae) in Bulgaria. – Folia parasitologica (Praha), 22: 185–188.

Bhat H.R., S.M. Kulkarni, A.C. Mishra. 1983. Records of Mesostigmata, Ereynetidae and Pterygosomidae (Acarina) in Western Himalayas, Sikkim and hill districts of West Bengal. – Journal Bombay Nat. Hist. Society, 80: 91–110.

Blasco Sabio C., M. Portus Vinyeta. 1974. Acariasis respiratoria en canaries españoles por Sternostoma tracheacolum Lawrence, 1948. Aportatión al tratamiento. – Revista Iberica de Parasitologia, 34(1–2): 33–47.

Boonkong S., W. Meckvichai. 1987. Arthropod parasites of the tree sparrow (Passer montanus L., 1758) in Bangkok, Thailand. – J. Sci. Soc. Thailand, 13: 231–237.

Bregetova N. G. 1950. New species of endoparasitic acarina of the family Rhinonyssidae (Gamasoidea, Acarina). – Acad. Nauk SSSR Dokl., 71: 1005–1008.

Bregetova N. G. 1951. [Mites parasitizing the nasal cavities of birds]. – Parasit. Sbornik, Zool. Inst. AS USSR, 13: 111–119 (in Russian).

[Bregetova N.G.]. 1953. [On the fauna of the gamasid mites of the Far East]. – Parazitologicheskiy sbornik, 15: 302–338 (in Russian).

[Bregetova N. G.] Брегетова Н.Г. 1956. [Gamasid mites (Gamasoidea)] Гамазовые клещи (Gamasoidea). –[Moscow – Leningrad] Москва – Ленинград: 247 pp.

Bregetova N. G. 1964. Some problems of evolution of the rhinonyssid mites. – First Int. Congr. Parasitol. Leningrad, 1–7.

Bregetova N. G. 1965a. Generic grouping of the rhinonyssid mites (Mesostigmata, Rhinonyssidae), parasites of birds. I. The new genus Sternostomoides. – Ent. Obozr., 44: 709–713.

Bregetova N. G. 1965b. Generic groups of rhinonyssid mites (Mesostigmata, Rhinonyssidae) – birds parasites. 2. New genus Locustellonyssus. – Zool. Zh., 44: 1092–1097.

Bregetova N. G. 1965c. A new species of parasitic mites of the genus Rhinoecius Cooreman (Mesostigmata: Rhinonyssidae). – Rev. Ent. U.R.S.S., 44(1): 210–216.

Bregetova N. G. 1967a. The generic groups of rhinonyssid mites-parasites of birds. III. Genera Neonyssus Hirst, 1921 and Neonyssoides Hirst, 1923. – Parazit. Sb., 23: 124–136 (in Russian, summ. Engl.).

Bregetova N. G. 1967b. The ontogeny of gamasoid mites as a basis for the formation of their natural system. – Parazitologiya, 1: 465–479 (in Russian).

Bregetova N. G. 1970. Mites from the nasal cavity of flycatchers of the genus Muscicapa. – Parazitologiya, 4(1): 59–62.

Bregetova N. G. 1971. Some considerations on the system and phylogeny of gamasid mites. – Proc. 3rd Intern. Congr. Acarol., pp. 263–267.

Bregetova N. G., L.V. Slavoshevskaya. 1983. A new subspecies of rhinonyssid mites of the genus Sternostoma (Gamasina) from Cuculus sp. of the Altai Territory. – Parazitologiya, 17(3): 234–239 (in Russian, summ. Engl.).

Brooks D. L.1959. The nasal mites of some west Texas flycatchers. Thesis MSc, 51 p.

Brooks D. L., W. Strandtmann. 1960. The nasal mites (Acarina) of some west Texas flycatchers (Tyrannidae). – J. Parasitol., 46(4): 418–432.

Brown N. Sandra. 1971. A survey of the Arthropod parasites of pigeons (Columba livia) in Boston. – J. Parasitol., 57(6): 1379–1380.

Brown N.S., G.I. Wilson. 1975. A comparison of the ectoparasites of the house sparrow (Passer domesticus) from North America and Europe. – The American Midland Naturalist, 94(1): 154–165.

Bush L. E. 1963. Incidence of nasal mites in different age classes of herring gulls (Larus argentatus). – J. Parasit., 49: 525.

Butenko O. M. 1960. New species of mites belonging to the family Rhinonyssidae parasitizing nasal cavity of birds. – Zool. Zhurnal., 39(10): 1489–1496.

[Butenko O. M.] 1962. [A review of the gamasoid mites, connected with birds. Tr. Okskogo Goszapovednika, 7] Обзор гамазоидных клещей, связанных с птицами. – Труды Окс. госзаповедника, 7: 353–385 (in Russian).

Butenko O. M. 1964. On the ecology of the mite Rhinonyssus rhynolethrum (Trouessart, 1895)(Rhinonyssidae, Gamasoidea). – Zool. Zh., 43: 274–276.

Butenko O. M. 1965. Tick [sic!] species parasitizing nasal cavities of birds, which are new for the USSR fauna. – Zool. Zh., 44(1): 131–133 (in Russian, summ. Engl.).

Butenko O. M. 1968. A method of collection of mites from the nasal cavity of living birds Rhinonyssidae. – Zoologicheskiy Zhurnal, 47(3): 451–452.

Butenko O. M. 1969. Zoogeographic analysis of mites of the family Rhinonyssidae (Gamasoidea, Parasitiformes) of the USSR fauna. – Zool. Zh., 48(2): 483–494 (in Russian, summ. English).

Butenko O. M. 1971a. A contribution to the systematic status of mites of the family Rhinonyssidae (Gamasoidea) [parasites of Anseriformes]. – Zoologicheskiy Zhurnal, 50(4): 483–494 (in Russian, summ. English).

[Butenko O. M.] 1971b. [Materials on the fauna of bird mites of Rhinonyssidae family (Gamasoidea) of Lenkoran lowland]. – Trudi Okskogo Gos. Zapovednika: 259–261 (in Russian).

[Butenko O. M.] 1971c. A summary of the knowledge of the nasal mites of birds of the Okskii Reserve Area. – Trudy Okskogo Gosudarstvennogo Zapovednika, 8: 204–223.

Butenko O. M. 1972. Mites of the genus Ptilonyssus parasitizing on Starlings. – Zoologicheskiy Zhurnal, 51(7): 1073–1077(in Russian, summ. English).

Butenko O. M. 1973a. Rhinonyssid mites (Gamasoidea) parasites of Grebes. – Parazitologiya (St. Petersburg), 7(3): 244–246 (in Russian, summ. English).

Butenko O. M. 1973b. A new species of the genus Larinyssus (Gamasoidea Rhinonyssidae). – Zoologicheskiy Zhurnal, 52(6): 951–953 (in Russian, summ. English).

Butenko O. M. 1973c. Two new species of the genus Rhinonyssus (Gamasoidea, Rhinonyssidae). - Zoologicheskiy Zhurnal, 52(12): 1872–1875 (in Russian, summ. English).

Butenko O. M. 1974a. A new species of parasitic mite of the genus Rhinonyssus (Gamasoidea Rhinonyssidae). – Zoologicheskiy Zhurnal, 53(3): 464–466.

Butenko O. M. 1974b. Two new species of the genus Sternostoma (Gamasoidea, Rhinonyssidae) from terns. – Zoologicheskiy Zhurnal, 53(7): 1089–1091 (in Russian, summ. Engl.).

Butenko O. M. 1975. On the taxonomic position of the mites of the genus Larinyssus Strandtmann (Gamasoidea, Rhinonyssidae), parasites of Laridae. – Parazitologiya (Leningrad), 9(2): 175–182 (in Russian, summ. English).

Butenko O. M. 1976a. Rhinonyssids (Gamasoidea: Rhinonyssidae), parasites of the rails from the region of the Lenkoran hibernation site. – Parazitologiya (Leningrad), 10(1): 72–77 (in Russian, summ. English).

Butenko O. M. 1976b. New species of the Rhinonyssid mites (Gamasoidea, Rhinonyssidae), parasitic in owls. – Parazitologiya (Leningrad), 10(4): 303–309 (in Russian, summ. English).

Butenko O. M. 1977. Morphology and systematics of the genus Vitznyssus Castro (Gamasoidea). – Zoologicheskiy Zhurnal, 56: 368–374 (in Russian, summ. English).

[Butenko O. M.] Бутенко О. М. 1984. [Nasal mites of Non-passerine birds of the Soviet Union]. – Zool. Inst. Acad. Sciences (Leningrad), 187 pp. (in Russian).

Butenko O. M. 1999. Two new species of mites of the genus Sternostomoides Bregetova, 1965 (Gamasoidea: Rhinonyssidae). – Acarina, 7: 47–51.

Butenko O. M., K. I. Lavrovskaya. 1980a. New species of rhinonyssids of the genus Ptilonyssus s. str. (Gamasoidea, Parasitiformes) parasitic in Larks. -Parazitologiya (St. Petersburg), 14(6): 522–531 (in Russian, summ. Engl.).

Butenko O. M., K. I. Lavrovskaya. 1980b. A new species of Rhinonyssids (Gamasoidea: Rhinonyssidae) from the nasal cavity of Ficedula hypoleuca (Pall.). – Parazitologiya (Leningrad), 14(4): 351–354 (in Russian, summ. English).

[Butenko O. M., K. I. Lavrovskaya]. Бутенко О.М., Лавровская К.И.1983. [New species of Rhinonyssid mites in Kirghizstan] Новые виды клещей-ринониссид в Киргизии // [Entomological research in Kirghizstan] Энтомол. иссл. в Киргизии., 1983, вып. 16, [Frunze, Ilim Publ. House]Фрунзе, изд. “Илим” c. 113–125 (in Russian).

[Butenko O. M., K. I. Lavrovskaya] Бутенко О.М., Лавровская К.И. 1984a. [Rhinonyssid mites of genus Ptilonyssus (Gamasoidea, Rhinonissidae), from Lanius] Клещи-ринониссиды рода Ptilonyssus (Gamasoidea, Rhinonyssidae), паразитирующие у сорокопутов. Научные основы охраны и рационального использования птиц: [Proceedings of Oka State reserve, Ryazan] Труды Окского госуд. заповедника. – Рязань, 1984 – Вып. 15. – p. 199–224. (in Russian).

[Butenko O. M., K. I. Lavrovskaya] Бутенко О.М., Лавровская К.И. 1984b.[Fauna of Rhinonyssidae on Passeriform birds. Genus Ptilonyssus s.str.] Фауна клещей-ринониссид паразитов воробьиных птиц., 1. Клещи р. Ptilonyssus s.str. Деп. ПИК ВИНИТИ, 1984, – 162 с. (in Russian).

[Butenko O. M., K. I. Lavrovskaya] Бутенко О.М., Лавровская К.И. 1985. [On the population structure and the distribution of numbers of Rhinonyssid mites] Материалы по структуре популяции и распределению её численности у клещей-ринониссид. Тез. докл. 5 Всес. акарологич. совещ., 1985, Фрунзе, изд-во “Илим”. С. 49–50. (in Russian).

[Butenko O. M., K. I. Lavrovskaya] Бутенко О.М., Лавровская К.И. 1988. [On the joint parasitizing of Rhinonyssidae (Gamasoidea)]. О совместном паразитировании клещей-ринониссид (Gamasoidea). Матер. Х Конф. Украинск. общ-ва паразитол., 1988, 3: 12. (in Russian).

[Butenko O. M., K. I. Lavrovskaya, M. K. Stanyukovich] Бутенко О.М., Лавровская К.И., Станюкович М.К. 1997. [Gamasoidea, Rhinonyssidae from Anseriformes in Russia and the adjacent countries] Kлещи-ринониссиды (Gamasoidea, Rhinonyssidae) пластинчатоклювых птиц России и сопредельных стран. – Пробл. энтомологии в России, т. 1, С-Пб. p. 56–57.

[Butenko O.M., S.G. Priklonskiy], Бутенко О.М., С.Г. Приклонский. 1964. [To the problem of the fauna and ecology of the gamasid mites of birds] К вопросу о фауне и экологии гамазовых клещей птиц. In: Сборник паразитологических работ. Труды Астраханского заповедника, вып.9. [Papers of Astrahan Reserve].

[Butenko O. M., V.N. Senotrusova, A.M. Sema] Бутенко О.М., Сенотрусова В.Н., Сёма А.М. 1970. [On the fauna of the mite family Rhinonyssidae in South Kazakhstan] Материалы по фауне клещей сем. Rhinonyssidae Южного Казахстана. Тез.докл. 2 Всес. совещ. по теоретической и прикладной акарoлогии. 1970, Киев, изд. “Наукова думка”. С. 86–87. (in Russian).

[Butenko O.M., O. H. Shterbinina] Бутенко О.М., Щербинина О.Х. 1976. [On the fauna of the Gamasid mites of Rhinonyssidae family on birds in Turkmenistan] Материалы по фауне гамазовых клещей сем. Rhinonyssidae птиц Туркменистана. Тез. докл. Всес.научн.конф. по комплексн. изуч. и освоен. пустын. терр. СССР, секция 4, Ашхабад, изд. “Илым”. – С. 58. (in Russian).

Butenko O. M., M. K. Stanyukovich. 1999. The distribution of rhinonyssid mites (Acari, Gamasina, Rhinonyssidae) in the nasal cavities of some anatid birds. – Ekologija, 2: 52–56.

Butenko O. M., M. K. Stanyukovich. 2001. New species of the Rhinonyssid mites (Gamasina: Rhinonyssidae) from birds of Russia and neighbouring countries – Parazitologiya, 35(6): 519–530.

Butenko O.M., M.K. Stanyukovich, I. Dimov. 2009. Rhinonyssid mites of the genus Ptilonyssus Berlese et Trouessart, 1889 (Gamasina: Rhinonyssidae) from nasal cavities of passerines (Aves: Passeriformes) from Russia and adjacent countries. VIIIth Nat. Conf. of Parasitology, 23–26 September 2009, Varna. Abstracts.

Butenko O.M., M.K. Stanyukovich, I. Dimov. 2012. On the composite species Ptilonyssus motacillae Fain, 1956 (Gamasina: Rhinonyssidae) from nasal cavities of passerines (Aves: Passeriformes). – Archimed, J. of Science and Practice, 12

Butenko O.M., M.K. Stanyukovich, I.D. Dimov. 2010. Co-parasitism of rhinonyssid mites (Parasitiformes: Gamasina, Rhinonyssidae) in the nasal cavities of birds. – British Society for Parasitology. 2010

Бутенко О.М., М.К. Станюкович, К.И. Лавровская. 1996. [Gamasoidea, Rhinonyssidae of Anseriformes, Anatidae of Russia and the neibouring countries] Клещи-ринониссиды (Gamasoidea, Rhinonyssidae) утиных птиц (Anseriformes, Anatidae) России и сопредельных стран. – [Ed. MOIP, Moskow] Матер. научн. конф. “Систематика, таксономия и фауна паразитов” 1996, М., изд. МОИП: 25–26.

[Butenko O. M., N.M. Stolbov] Бутенко О.М., Н.М. Столбов. 1971. [Gamasid mites of fam. Rhinonyssidae, parasitizing birds in the southern areas of Tyumen region] Гамазовые клещи сем. Rhinonyssidae (Gamasoidea, Parasitiformes), паразитирующие у птиц в южных районах Тюменской области. – [Proc. of the Okskiy Reserve] Тр. Окского госуд. заповедника. [Moskow] Москва, 257–259 (in Russian).

[Butenko O. M., L.T. Yurlov] Бутенко О.М., Юрлов К.Т. 1981. [To the characteristics of the relations between the Rhinonyssidae mites with birds of order Anseriformes in the forest-steppe of Western Siberia] К характеристике взаимоотношений клещей- ринониссид с птицами отряда гусеобразных в лесостепи Западной Сибири. – [Ecology and biocenotic connections of the migrating birds in Western Siberia] Экология и биоценотические связи перелетных птиц Западной Сибири. – [Novosibirsk, Ed. Nauka] Новосибирск: Наука, 245–251 (in Russian).

[Butenko O. M., L.T. Yurlov, N.M. Stolbov] Бутенко О.М., Л.Т. Юрлов, Н.М. Столбов. 1972. [Distribution by hosts of the birds mites of the family Rhinonyssidae] Распределение по хозяевам птичьих клещей семейств Rhinonyssidae (Gamasoidea). – In: [Transcontinental connections of migrating birds and their role in the spreading of arboviruses] Трансконтинентальные связи перелетных птиц и их роль в распространении арбовирусов. [Novosibirsk] Новосибирск: 370–375 (in Russian).

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Carmen B.S., P.V. Montserrat. 1974. Respiratory acariasis in spanish canaries by Sternostoma tracheacolum Lawrence, 1948. Contribution to treatment. – Rev. Iber. Parasitol., 34: 33–48.

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The Mite Family Spinturnicidae

Parasites on bats (Chiroptera), living during all stages on the wing or tail membranes. The species of the genus Ancystropus are specialized to live on the eyelids and eye canthi and the Meristaspis – on the eye canthi and on the wing membranes.

The „Pediculus vespertilionis“ of Linnaeus (1758) was invalidated in 1936, but has been used often in the later years as „Spinturnix vespertilionis“. The genus Spinturnix von Heyden, 1826 is the largest in the family, containing 50 out of 113 valid species in the family.

The first major research on these mites was done by Kolenati (1856–1860), than by Oudemans, who described in 1902 the subfamily Spinturnicinae (within Parasitidae) and raised it in a family Spinturnicidae (Oudemans, 1916). Until 1960 many new taxa (including the valid genera Spinturnix von Heyden, Ancystropus Kol., Meristaspis Kol., Periglischrus Kol. and 37 out of the present day 113 valid species have been described by von Heyden (1826), C.L. Koch (1826, 1839, 1841), Kolenati (1856, 1857), Canestrini (1884), Banks (1902), Oudemans (1902, 1910), Hirst (1923, 1927), De Meillon & Lavoipierre (1944), Hoffmann (1944), Till (1958), Turk (1950), Radford (1947, 1951), Zumpt (1951), Zumpt & Till (1954), Hiregaudar & Ball (1955), Tibbetts (1957), together with many synonyms.

The revision of Rudnick (1960) is the modern treatment of the family. Included are 37 species of Spinturnicidae (including five newly described) of seven genera, three of which new (Eyndhovenia, Paraperiglischrus and Paraspinturnix). After this revision appeared, many contributions were made by Advani & Vazirani, Allred, Andreyko, A. Baker, Benoit, Beron, Bhat, Bobkova, Delfinado & Baker, Dias, Domrow, Dubovchenko, Dusbábek, Estrada-Peña, Fain, Ferenc, Furman, Gu & Wang, Haitlinger, Herin & Tipton, Juvara, Machado-Allison, Morales-Malacara, Nadchatram, Pan & Teng, Pinchuk, Prasad, Rybin, Seo, Stanyukovich, Till, Uchikawa, Vshivkov, Wharton, Wilson, Ye & Ma, and others. Important series of publications were made by J. Deunff and his co-laborators, including descriptions of several new species. Since 1960 (including Benoit, 1959) have been described 63 valid new species and six new genera: Cameronieta Machado-Allison, Emballonuria Uchikawa, Zhang, O’Connor et Klompen, Mesoperiglischrus Dusbábek, Parameristaspis Advani et Vazirani, Oncoscelus Delfinado et Baker and Periglischrodes Baker et Delfinado (= Ptilonyssus). The genus Spinturnix was thoroughly reviewed by Uchikawa, Zhang, O‘Connor & Klompen (1994).

So far Spinturnicidae have been recorded on bats from 13 families in all continents: Pteropodidae, Emballonuridae, Hipposideridae, Megadermatidae, Molossidae, Mormoopidae, Natalidae, Noctilionidae, Nycteridae, Phyllostomatidae, Rhinolophidae, Rhinopomatidae, Vespertilionidae.

The facts, that the last valid genera have been described in 1968 and than in 1994 indicate that we already know in broad lines the systematics and the distribution of Spinturnicidae in the world.

The Composition of Spinturnicidae

Fam. Spinturnicidae Oudemans, 1902 – 12 gen., 110 sp.

Ancystropus Kolenati, 1856 – 7 sp.

Cameronieta Machado-Allison, 1965 – 7 sp.

Emballonuria Uchikawa, Zhang, O’Connor et Klompen, 1994 – 1 sp.

Eyndhovenia Rudnick, 1960 – 2 sp.

Meristaspis Kolenati, 1857 – 7 sp.

Mesoperiglischrus Dusbábek, 1968 – 2 sp.

Oncoscelus Delfinado et Baker, 1963 – 1 sp.

Parameristaspis Advani et Vazirani, 1981 – 1 sp.

Paraperiglischrus Rudnick, 1960 – 4 sp.

Paraspinturnix Rudnick, 1960 – 1 sp.

Periglischrus Kolenati, 1857 – 30 sp.

Spinturnix von Heyden, 1826 – 50 sp.

Catalogue of Spinturnicidae

Family Spinturnicidae Oudemans

Pteroptida Kolenati, 1857: 59

Pteroptidae Murray, 1877: 175

Pteroptina Kramer, 1882: 392

Periglischrinae Benoit: 383

Spinturnicinae Oudemans, 1902: 69

Spinturnicidae Oudemans, 1916: 7

Type genus: Spinturnix von Heyden, 1826

Genus Ancystropus Kolenati

Ancystropus Kolenati, 1856: 25

Type species: Ancystropus zeleborii Kolenati, 1856

Ancystropus aequatorialis Estrada-Peña, Ballesta et Ibanez

Ancystropus aequatorialis Estrada-Peña, Ballesta & Ibanez, 1992: 309

Type locality: Bioko I.

Distribution: Equatorial Guinea

Host: Rousettus aegyptiacus (Chiroptera: Pteropodidae)

Ancystropus aethiopicus Hirst

Ancystropus aethiopicus Hirst, 1923: 979; Radford, 1942: 67; 1950: 376; Benoit, 1959b: 23; Dusbábek & Bergmans, 1980: 66

Spinturnix aethiopicus: Zumpt, 1951: 80

Ancytropus aetheopicus [sic!]: Hiregaudar & Bal, 1956: 90

Type locality: Zanzibar

Distribution: Angola, DR Congo, Gambia, Nigeria, Tanzania (Zanzibar), Uganda

Hosts: Epomophorus labiatus minor, E. wahlbergi haldemani, Hypsignathus monstrosus, Micropteropus pusillus (Chiroptera: Pteropodidae)

Ancystropus eonycteris Delfinado et Baker

Ancystropus eonycteris Delfinado & Baker, 1963: 917; Uchikawa & Kobayashi, 1978: 250; Cuy, 1979: 164; Uchikawa, 1990: 39; Corpuz-Raros, 1993: 184; Ahamad et al., 2013: 140; Corpuz-Raros & Lit, 2015: 33

Ancystropus nakatae Prasad, 1969a: 535; Fain, 2002: 68 – Type: Negros Oriental, Camp Look Out, Valencia, from Eonycteris spelaea glandifera (syn. by Domrow)

Type locality of A. eonycteris: Central Cave, Cotabato, Mindanao

Distribution: Indonesia (Java), Malaysia, Philippines, Thailand

Hosts: Eonycteris robusta, E. spelaea glandifera, Rousettus leschenaulti (Chiroptera: Pteropodidae)

Ancystropus leleupi Benoit

Ancystropus leleupi Benoit, 1959a: 106

Type locality: Katanga, grottes de Kakontwe

Distribution: DR Congo

Host: Rousettus aegyptiacus (Chiroptera: Pteropodidae)

Ancystropus notopteris Uchikawa

Ancystropus notopteris Uchikawa, 1990: 31

Type locality: Colombo Cave, Suva

Distribution: Fiji, New Caledonia, Vanuatu (New Hebrides)

Hosts: Notopteris macdonaldi, N. neocaledonica (Chiroptera: Pteropodidae)

Ancystropus taprobanius (Turk)

Meristaspis taprobanius Turk, 1950: 73; Chow, Thevasagayam & Tharumrajah, 1954: 12

Ancystropus taprobanius (Turk): Rudnick, 1960: 177; Domrow, 1972: 571; Prasad, 1974: 68; Domrow & Nadchatram, 1978: 90; Uchikawa, 1990: 41; Durden, Peterson et al., 1990: 53; Corpuz-Raros & Lit, 2015: 33

Ancytropus [sic] indica Hiregaudar & Bal, 1955: 221 (nomen nudum)

Ancytropus [sic] indicus Hiregaudar & Bal, 1956: 92 – typus from the Jogeshwari and Kanheri caves, Bombay (syn. by Domrow, 1972: 571)

Ancystropus indicus Hiregaudar & Bal: Rudnick, 1960: 176

Ancystropus rudnicki Baker & Delfinado, 1964: 577, from Papua New Guinea, on Rousettus amplexicaudatus; Prasad, 1969c: 672 (syn. by Domrow, 1972: 571

Type locality: Rattota, Matale District

Distribution: India, Indonesia (Ceram, Java), Laos, Malaysia, Nepal, Papua New Guinea, Philippines, Sri Lanka, Thailand

Hosts: Cynopterus brachyotus, C. luzoniensis, Rousettus amplexicaudatus, R. leschenaulti, R. seminudus, R. stresemanni (Chiroptera: Pteropodidae), Scotophilus kuhli (Chiroptera: Vespertilionidae)

Ancystropus zeleborii Kolenati

Ancystropus zeleborii Kolenati, 1856: 25; Costa, 1966: 80; 1967: 106; Pan & Teng, 1973: 84; Whitaker, Baccus, Cleveland & Gu, 1990: 53; Uchikawa, 1990: 39; Durden, Peterson et al., 1990: 53; Corpuz-Raros & Lit, 2015: 33

Leiostaspis zeleborii: Kolenati, 1857: 60; 1860: 580

Ancystropus zelebori: Hirst, 1923: 978; Radford, 1942: 67; 1947: 305; 1950: 376

Spinturnix zelebori: Zumpt, 1951: 80

Ancystropus palawanensis Delfinado & Baker, 1963: 915; – type: Philippines, Busuanga I., 6 km SE San Nicolas, Palawan, from Cynopterus brachyotis and Rousettus amplexicaudatus; Baker & Delfinado, 1964: 579

Type locality of A. zeleborii: Egypt

Distribution: China, DR Congo, Cyprus, Egypt, India, Indonesia (Ceram), Israel, Laos, Malaysia (Sabah, Sarawak, Selangor), Nepal, Papua New Guinea, Philippines, Solomon Islands, Thailand, Uganda, Vietnam

Hosts: Acerodon jubatus, Aethalops alecto, Alionycteris paucidentata, Balionycteris maculata, Cynopterus brachyotis, C. luzoniensis, C. sphinx, Dobsonia viridis champani, Haplonycteris fischeri, Penthetor lucasi, Ptenochirus jagori, Rousettus aegyptiacus, R. amplexicaudatus, R. leschenaulti, R. palawanensis, R. spelaea (Chiroptera: Pteropodidae) , (?) Lavia frons rex (Chiroptera: Megadermatidae), Rhinopoma microphyllum (Chiroptera: Rhinopomatidae)

Genus Cameronieta Machado-Allison

Cameronieta Machado-Allison, 1965b: 244

Type species: Cameronieta thomasi Machado-Allison, 1965

Cameronieta almaensis Almeida, Gettinger et Gardner

Cameronieta almaensis: Almeida, Gettinger & Gardner, 2016: 212

Type locality: Serra das Almas Reserve, State of Ceára

Distribution: Brazil

Host: Pteronotus parnelli (Chiroptera: Mormoopidae)

Cameronieta elongata Furman

Periglischrus elongatus Furman, 1966: 140

Cameronieta elongatus: Herrin & Tipton, 1975: 7; Palacios -Vargas et al., 1985: 140

Type locality: Mt. Tamana Caves, Trinidad

Distribution: Mexico, Panama, Trinidad, Venezuela

Hosts: Pteronotus parnelli (Chiroptera: Mormoopidae), Chilonycteris rubiginosa fusca (Chiroptera: Vespertilionidae)

Cameronieta machadoi Dusbábek

Periglischrus strandtmanni: Dusbábek, 1968: 5 (pro parte)

Cameronieta machadoi Dusbábek, 1967: 150

Type locality: Cueva de los Lagos, Serro de la Guanabana, Isla de Pinos

Distribution: Cuba (Isl. Pinos)

Host: Chilonycteris macleayi macleayi (Chiroptera: Phyllostomatidae)

Cameronieta strandtmanni (Tibbetts)

Periglischrus strandtmanni Tibbetts, 1957: 14; Rudnick, 1960: 199; Dusbábek, 1968: 5; Kingston, Villa & Lopez-Forment, 1971: 928

Cameronieta strandtmanni (Tibbetts): Dusbábek, 1967: 156; Herrin & Tipton, 1975: 6; Whitaker & Morales-Malacara, 2005: 551

Type locality: Frio Cave, Uvalde County, Texas

Distribution: Cuba, Mexico, USA (Texas), Venezuela

Hosts: [Chilonycteris fuliginosa torrei, Ch. macleayi macleayi, Ch. parnelli bothi, quated by mistake by Dusbábek, 1968, see the species Cameronieta tibbetsi, C. machadoi, C. torrei], Mormoops blainvillei, M. megalophylla senicula (Chiroptera: Mormoopidae)

Cameronieta thomasi Machado-Allison

Cameronieta thomasi Machado-Allison, 1965b: 244; 1965c: 243; Machado-Allison & Antequera, 1969: 376; Herrin & Tipton, 1975: 9; Gannon & Willig, 1994: 282

Type locality: Cueva Walter Dupuy, Estado Miranda (Venezuela)

Distribution: Colombia, Mexico, Puerto Rico, Venezuela

Hosts: Pteronotus parnellii, P. rubiginosa, P. suapurensis (Chiroptera: Mormoopidae)

Cameronieta tibbettsi Dusbábek

Periglischrus strandtmanni Dusbábek, 1968: 5 (pro parte)

Cameronieta tibbettsi Dusbábek, 1967: 156

Type locality: Cueva de William Palmer, Guanajay, prov. Pinar del Rio

Distribution: Cuba

Host: Pteronotus parnelli boothi (Chiroptera: Mormoopidae)

Cameronieta torrei Dusbábek

Periglischrus strandtmanni Dusbábek, 1968: 5 (pro parte)

Cameronieta torrei Dusbábek, 1967: 153

Type locality: Cueva Colon, Mayajigua, prov. Las Villas

Distribution: Cuba

Host: Pteronotus (Chilonycteris) quadridens fuliginosus (Chiroptera: Mormoopidae)

Cameronieta torrei dusbabeki Deunff, Whitaker et Kurta

Cameronieta torrei dusbabeki Deunff, Whitaker & Kurta, 2007: 179

Type locality: Culebrones Cave, 7 km southwest of Arecibo

Distribution: Puerto Rico

Host: Pteronotus (Chilonycteris) quadridens fuliginosus (Chiroptera: Mormoopidae)

Genus Emballonuria Uchikawa, Zhang, O’Connor et Klompen

Emballonuria Uchikawa, Zhang, O’Connor & Klompen, 1994: 289

Type species: Spinturnix orientalis Turk, 1950

Emballonuria orientalis (Turk)

Spinturnix orientalis Turk, 1950: 73; Chow, Thevasagayam & Tharumrajah, 1954: 126; Rudnick, 1960: 225

Emballonuria orientalis (Turk): Uchikawa, Zhang, O’Connor & Klompen, 1994: 290

Type locality: Mousekande, Gammaduwa, Ceylon

Distribution: Sri Lanka

Host: Saccolaimus [Taphozous] saccolaimus (Chiroptera: Emballonuridae)

Genus Eyndhovenia Rudnick

Eyndhovenia Rudnick, 1960: 187

Type species: Pteroptus euryalis Canestrini, 1884

Eyndhovenia brachypus Sun, Wang et Wang

Eyndhovenia brachypus Sun, Wang & Wang, 1986: 194

Type locality: Fujian, Wuyi Mountains Nature Preserve

Distribution: China

Host: Rhinolophus rouxi (Chiroptera: Rhinolophidae)

Eyndhovenia euryalis (Canestrini)

Pteroptus euryalis Canestrini, 1884: 1657; Berlese, 1892: 124

Spinturnix euryalis Hirst, 1927: 323; Turk & Turk, 1952: 485; Turk, 1953: 13; Krochko, 1995: 83

Spinturnix omahonyi Turk, 1945: 800

Eyndhovenia euryalis (Canestrini): Rudnick, 1960: 188; many other authors

Eyndhovia [sic] euryalis ahi Baker & Delfinado, 1964: 585 (syn. by Domrow, 1972: 567; syn. with Spinturnix oudemansi Eyndhoven, 1941 by Uchikawa & Dusbábek, 1978: 257)

Type locality: Toscana (Italy)

Distribution: Algeria, Armenia, Australia, Bosnia and Herzegovina, Belgium, Bulgaria, China, Crimea, England, France (incl. Corsica), Germany, Greece (incl. Creta), Holland, Israel, Italy, Japan, Kenya, Kirghizstan, Macedonia, Montenegro, Morocco, Poland, Romania, Serbia, Slovakia, South Africa, Spain, Sudan, Tadjikistan, Thailand, Turkey, Ukraine

Hosts: Rhinolophus axillaris, Rh. blasii, Rh. clivosus augur, Rh. euryale, Rh. ferrumequinum, Rh. hipposideros, Rh. megachyllus, Rh. mehelyi, Rh. rouxi sinicus (Chiroptera: Rhinolophidae), Eptesicus serotinus, Myotis blythi, M. emarginatus, Pipistrellus pipistrellus, P. gaisleri (Chiroptera: Vespertilionidae), Miniopterus schreibersi (Chiroptera: Miniopteridae); Hipposideros larvatus (Chiroptera: Hipposideridae)

Eyndhovenia euryalis oudemansi (van Eyndhoven)

Eyndhovenia oudemansi van Eyndhoven, 1941: 54 (syn. by Rudnick, 1960: 188), but later restaured.

Eyndhovenia euryalis oudemansi (van Eyndhoven): Krištofik, Piksa & Sahanowicz, 2012: 102; Orlova, Stanyukovich & Orlov, 2015: 26; Orlova & Orlov, 2019: 320

Type locality: Mergelsteinbrüche, St. Pietersberg, Limburg

Distribution: Netherlands

Hosts: Rhinolophus ferrumequinum, Rh. hipposideros, Rh. euryale, Rh. mehelyi (Chiroptera: Rhinolophidae), Miniopterus schreibersi (Miniopteridae), Pipistrellus pipistrellus, Eptesicus serotinus, Myotis blythus

Eyndhovenia euryalis cornuti Uchikawa et Dusbábek

Eyndhovenia euryalis cornuti Uchikawa & Dusbábek, 1978: 247

Type locality: Shojo-do, Kashiwazaki, Niigata Pref.

Distribution: Japan

Host: Rhinolophus cornutus (Chiroptera: Rhinolophidae)

Genus Meristaspis Kolenati

Meristaspis Kolenati, 1857: 60

Type species: Pteroptus lateralis Kolenati, 1856 (by subsequent designation, Vitzthum, 1932)

Meristaspis calcarata (Hirst)

Ancystropus (Meristaspis) calcaratus Hirst, 1923: 983

Meristaspis calcarata (Hirst): Domrow, 1972: 568; 1974; 1979: 321; 1988: 889; Wilson & Wodzicki, 1977; Durden, Peterson et al., 1990: 53

Meristaspis calcaratus: Rudnick, 1960: 183; Delfinado & Baker, 1963: 908; Wilson, 1967: 144; Uchikawa & Wada, 1979: 121; Fain, 2002: 68

Type locality: Rook Island

Distribution: Australia, Indonesia (Ceram), Japan (Ryukyu’s), Madagascar, Mariana Is. (Saipan, Guam), Niue Is., Palau, Papua New Guinea, Philippines, Rook Is., Vanuatu (New Hebrides)

Hosts: Pteropus alecto, P. dasymallus, P. geddiei, P. hypomelenas (?), P. m. mariannus, P. poliocephalus, P. pselaphon, P. rufus, P. scapulatus, P. speciosus, P. tablensis, P. t. tonganus, P. vampyrus (Chiroptera: Pteropodidae)

Meristaspis jordani (Radford)

Ancystropus jordani Radford, 1947: 309

Meristaspis jordani (Radford): Baker & Delfinado, 1964: 581; Prasad, 1969c: 666; Domrow, 1972: 569; Durden, Peterson et al., 1990: 53

Meristaspis jordani philippinensis Prasad, 1969a: 539; 1969b: 672 – type from Negros Oriental, Malindog, San Antonio, Sibulan, on Harpyonycteris whiteheadi (syn. by Domrow, 1972: 569); Fain, 2002: 69

Ancystropus lavellanus Radford, 1951: 97 – type from Vela Lavella, Solomon Islands on (?) Pteropus lavellanus (syn. by Domrow, 1972: 569)

Meristaspis lavellanus: Rudnick, 1960: 187

Type locality: Koelawi Paloe, W. Sulawesi

Distribution: India, Indonesia (Ceram, Sulawesi), Papua New Guinea (Bismarck Arch., Manus Is.), Philippines, Solomon Islands

Hosts: a bat (?Pteropus lavellanus), Cynopterus luzoniensis, Dobsonia moluccensis, Harpyonycteris whiteheadi (Chiroptera: Pteropodidae)

Meristaspis kenyaensis (Radford)

Ancystropus kenyaensis Radford, 1947: 305; 1950b: 376

Spinturnix pipistrellus Radford (not pipistrelli Kolenati, 1859), 1951: 99 – type: Njala, Sierra Leone, from Pipistrellus stampflii (syn. by Rudnick, 1960: 185)

Meristaspis kenyaensis (Radford): Rudnick, 1960: 185; Dusbábek & Bergmans, 1980: 66

Type locality: Kaimosi (Kenya)

Distribution: Kenya, Nigeria, Sierra Leone

Hosts: Eidolon helvum (Chiroptera: Pteropodidae), Pipistrellus stampflii (Chiroptera: Vespertilionidae)

Meristaspis lateralis (Kolenati)

Pteroptus lateralis Kolenati, 1856: 29

Meristaspis lateralis (Kolenati): Kolenati, 1857: 60; 1859: 84; Vitzthum, 1932: 36; 1941: 776; Radford, 1950: 39; Baker & Wharton, 1952: 63; Hiregaudar & Bal, 1955: 221; 1956: 86; Rudnick, 1960: 180; Delfinado & Baker, 1963: 911; Baker & Delfinado, 1964: 581; Costa, 1966: 80; 1967: 106; Prasad, 1969c: 668; 1974: 69; Domrow, 1972: 568; Pan & Teng, 1973: 84; Domrow & Nadchatram, 1978: 89; Uchikawa & Kobayashi, 1978: 251; Uchikawa & Suzuki, 1980: 16; Durden, Peterson et al., 1990: 53; Fain, 2002: 68;

Pteroptus punctolyra: Kolenati, 1856: 28 – type: Egypt, from Rhinopoma microphyllum (syn. by Rudnick, 1960)

Pteroptus roseus: Kolenati, 1856: 28 – Egypt, on Rhinopoma microphyllum (syn. by Rudnick, 1960)

Tinoglischrus punctolyra: Kolenati, 1857: 60; 1859: 83; Vitzthum, 1932: 37

Ancystropus (Meristaspis) lateralis: Hirst, 1923: 985; Stiles & Nolan, 1931: 629; Radford, 1942: 67; 1950: 376; 1954: 309

Meristaspis lateralis ceylonicus Turk, 1950: 73 – type: Rattota, Matale Distr., Ceylon

from Rousettus seminudus; Hiregaudar & Bal, 1956: 86; syn. by Rudnick, 1960: 180

Spinturnix lateralis: Zumpt, 1951: 80 (in part)

Meristaspis lateralis ceylanicus: Chow, Thevasagayam & Tharumrajah, 1954: 126

Meristaspis taprobanius Turk (in part, male only), 1950: 73 – type: Rattota, Matale Distr., Ceylon, from Rousettus seminudus; Chow, Thevasagayam & Tharumrajah, 1954: 126 Chow, Thevasagayam & Tharumrajah, 1954: 126 – syn. by Rudnick, 1960: 180

Type locality: Egypt

Distribution: China, Cyprus, Egypt (= Pteroptus roseus), India, Indonesia, Israel, Laos, Malaysia, New Guinea, Philippines, Sri Lanka, Thailand, Yemen

Hosts: Cynopterus brachyotus, C. b. luzoniensis, Eonycteris spelaea, E. s. glandifera, E. robusta, Macroglossus minimus lagochilus, Rousettus aegyptiacus, R. amplexicaudatus, R. leschenaulti, R. seminudus, R. stresemanni (Chiroptera: Pteropodidae), Rhinopoma microphyllum (Chiroptera: Rhinopomatidae)

Meristaspis macroglossi (Hirst)

Ancystropus (Meristaspis) macroglossi Hirst, 1923: 981; Domrow, 1972: 568

Meristaspis macroglossi (Hirst): Domrow, 1979: 322; Domrow & Nadchatram, 1978: 89

Meristaspis dusbabeki Baker & Delfinado, 1964: 583; Prasad, 1969: 666; Tenorio, 1976: 19 (syn. by Domrow, 1972: 568)

Type locality: Gilolo (Papua New Guinea)

Distribution: Malaysia, Papua New Guinea, Solomon Islands

Hosts: Cynopterus horsfieldi, Macroglossus minimus lagochilus, M. minimus, Nyctimene albiventer papuanus, Syconycteris crassa papuana (Chiroptera: Pteropodidae)

Meristaspis mindanaoensis Delfinado et Baker

Meristaspis mindanaoensis Delfinado & Baker, 1963: 910; Baker & Delfinado, 1964: 579; Domrow, 1967: 775; 1972: 568; Prasad, 1969: 670; Uchikawa & Kobayashi, 1978: 251; Cuy, 1979: 178; Corpuz-Raros, 1993: 184; Corpuz-Raros & Lit, 2015: 34

Meristaspis hardyi Prasad, 1969: 535, from Solomon Is., Guadalcanal, Nini Creek, Roroni, ex Nyctimene major (syn. by Domrow, 1972: 568)

Type locality of M. mindanaoensis: Mt. Malindang, 1666 m, Masawan, Mutya, Mindanao

Distribution: Australia, Malaysia, Papua New Guinea, Philippines, Solomon Islands, Thailand

Hosts: Cynopterus brachyotus, Eonycteris spelaea, Macroglossus minimus lagochilus, Megaerops ecaudata, Nyctimene aello, N. albiventer, N. draconilla, N. major, N. robinsoni, Paranyctimene raptor, Prioniturus malindangensis, Rousettus amplexicaudatus, R. leschenaulti, Syconycteris crassa papuana (Chiroptera: Pteropodidae)

Genus Mesoperiglischrus Dusbábek

Mesoperiglischrus Dusbábek, 1968: 12

Type species: Mesoperiglischrus nyctiellinus Dusbábek, 1968

Mesoperiglischrus natali (Furman)

Periglischrus natali Furman, 1966: 128; Herrin & Tipton, 1975: 60; Palacios-Vargas et al., 1985: 140

Mesoperiglischrus natali (Furman): Dusbábek, 1968: 18; Morales-Malacara, 2001: 194

Type locality: San Lorenzo Caves, Fort Sherman, Canal Zone, Panama

Distribution: Brazil, Mexico, Panama, Trinidad, Venezuela

Hosts: Natalus stramineus, N. mexicanus saturatus, N. tumidirostris haymani (Chiroptera: Natalidae)

Mesoperiglischrus nyctiellinus Dusbábek

Mesoperiglischrus nyctiellinus Dusbábek, 1968: 13

Type locality: Hotel Las Cuevas, Trinidad, Prov. de Las Villas, Cuba

Distribution: Cuba

Host: Natalus (Nyctiellus) lepidus (Chiroptera: Natalidae)

Genus Oncoscelus Delfinado et Baker

Oncoscelus Delfinado & Baker, 1963: 913

Type species: Ancystropus kanheri Hiregaudar et Bal, 1956

Oncoscelus kanheri (Hiregaudar et Bal)

Ancytropus [sic !] kanheri Hiregaudar & Bal, 1955: 221; 1956: 95

Ancystropus kanheri Hiregaudar et Bal: Rudnick, 1960: 177; Domrow, 1972: 571; Uchikawa, 1990: 36; Corpuz-Raros & Lit, 2015: 33

Oncoscelus kanheri (Hiregaudar et Bal): Delfinado & Baker, 1963: 914; Baker & Delfinado, 1964: 579; Prasad, 1969: 676; Seneviratna, Chandrica Fernando & Udagama-Randeniya, 2009: 998

Type locality: the Jogeshwari, Kanheri and Elephanta caves, Bombay

Distribution: India, Indonesia (Papua, Sumba), Malaysia, Philippines (Palawan), Papua New Guinea, Thailand, Solomon Islands, Sri Lanka

Hosts: Eonycteris spelaea, Rousettus amplexicaudatus, R. leschenaulti, R. stresemanni (Chiroptera: Pteropodidae)

Genus Parameristaspis Advani et Vazirani

Parameristaspis Advani & Vazirani, 1981: 81

Type species: Parameristaspis delfinadoi Advani et Vazirani, 1981

Parameristaspis delfinadoae Advani et Vazirani (nomen emend.)

Parameristaspis delfinadoi Advani & Vazirani, 1981: 72

Type locality: India

Distribution: India

Hosts: Chiroptera

Genus Paraperiglischrus Rudnick

Paraperiglischrus Rudnick, 1960: 191

Type species: Pteroptus rhinolophinus C.L. Koch, 1841

Paraperiglischrus hipposideros Baker et Delfinado

Paraperiglischrus hipposideros Baker & Delfinado, 1964: 589; Prasad, 1969: 676; Domrow, 1972: 572; Cuy, 1979: 171; Corpuz-Raros, 1993: 184; Gu & Wang, 1988; Seneviratna, Chandrica Fernando & Udagama-Randeniya, 2009: 998 ; Corpuz-Raros & Lit, 2015: 34

Paraperiglischrus analis Pan & Teng, 1973: 83. Type: Fukien Prov., from Hipposideros p. pratti (syn. by Uchikawa, 1979a)

Type locality: New Britain: Keravat (Papua New Guinea); paratypes from New Guinea

Distribution: China, Japan (Iriomote Is.), Papua New Guinea (incl. New Britain), Thailand, Borneo, Philippins, Sri Lanka

Hosts: Hipposideros armiger, H. diadema griseus, H. lavatus, H. pratti pratti, H. turpis (Chiroptera: Hipposideridae), Cynopterus sphynx, Phileter sp. (Chiroptera: Pteropodidae)

Paraperiglischrus rhinolophinus (C.L. Koch)

Pteroptus rhinolophinus C.L. Koch, 1841: f. 38; 1842: 126

Periglischrus rhinolophinus (C.L. Koch): Turk, 1945: 798; 1953: 14

Periglischrus rhinolophus Walckenaer & Gervais, 1847: 546

Pteroptus euryalis Berlese (in part, not Canestrini, 1884), 1889: 3, f.54–3; 1892: 27; 1921: 93

Pteroptus interruptus Kolenati, 1856: 29; 1856: 189

Periglischrus interruptus Kolenati, 1857: 60; 1859: 79; 1860: 153; Rollinat & Trouessart, 1897: 135; Oudemans, 1903: 137 (in part); 1910: 72; Hirst, 1923: 987; Vitzthum, 1932 (in part): 37; Stiles & Nolan, 1931: 633; van Eyndhoven, 1940: 36; 1941: 47; 1950: 1009; Radford, 1942: 67; 1950: 370; Starkoff, 1957: 126; Lombardini, 1958

Pteroptus hipposideros Kolenati, 1856: 29; 1856: 191

Periglischrus hipposideros Kolenati, 1857: 60; 1857: 135; 1859: 82; 1860: 147; C.L. Koch, 1865: 274; Rollinat & Trouessart, 1897: 136; Oudemans, 1903: 137; 1910: 72; Stiles & Nolan, 1931: 633; Radford, 1942: 67; 1950: 376

Periglischrus asema Kolenati, 1857: 60; 1857: 135; 1859: 81; C.L. Koch, 1865: 264; Murray, 1877: 177; Rollinat & Trouessart, 1897: 136; Oudemans, 1903: 137; Stiles & Nolan, 1931: 633; van Eyndhoven, 1941: 53; Radford, 1942: 67; 1950: 376

Periglischrus glutinimargo Kolenati, 1859: 80; 1860: 150; Oudemans, 1903: 137; Stiles & Nolan, 1931: 633; Radford, 1942: 67; 1950: 376

Periglischrus africanus Zumpt, 1950: 93 – type: Sterkfontein Caves, Transvaal, from Rhinolophus clivosus augur; syn. by Rudnick, 1960: 193; Till, 1958: 256

Periglischrus africana Hiregaudar et Bal (for africanus Zumpt), 1956: 104

Periglischrus rhinolophi Hiregaudar & Bal, 1955: 221 – type: Tulsi Lake Tunnel, Bombay, from Rhinolophus rouxi; 1956: 104; syn. by Rudnick, 1960: 193

Paraperiglischrus rhinolophinus (C.L. Koch): Rudnick, 1960: 192; Dusbábek, 1964a: 6; Beron, 1968: 158; 1973: 125; 1974: 67; Domrow, 1972: 572; Uchikawa & Kobayashi, 1978: 252; Pan & Teng, 1980: 84; Rybin, 1983: 356; Gu & Wang, 1988; Durden, Peterson et al., 1990: 53; Imaz, Aihartza & Totorika, 1999: 24; Krištofík & Danko, 2012: 171

Type locality: Europe

Distribution: Australia, Bulgaria, Cameroon, China, Croatia, Greece (incl. Creta), Holland, Indonesia, Italy, Japan, Egypt, France (incl. Corsica), Great Britain, India, Ireland, Italy, Kenya, Kirghizstan, Korea, Laos, Macedonia, Malaysia, Serbia, Slovakia, South Africa, Spain, Sri Lanka, Thailand

Hosts: Rhinolophus blasii, Rh. clivosus augur, Rh. c. cornutus, Rh. euryale, Rh. f. ferrumequinum, Rh. ferrumequinum nippon, Rh. hipposideros, Rh. hipposideros minutus, Rh. megaphyllus, Rh. mehelyi, Rh. rouxi sinicus, Rh. shamelli, Rh. yunaensis (Chiroptera: Rhinolophidae), Barbastella barbastellus (Chiroptera: Vespertilionidae), Miniopterus schreibersi (Chiroptera: Miniopteridae), Rousettus amplexicaudatus (Chiroptera: Pteropodidae)

Paraperiglischrus ruber Estrada-Peña, Ballestra et Ibanez

Paraperiglischrus ruber Estrada-Peña, Ballestra & Ibanez, 1992: 311

Type locality: São Tomé and Príncipe

Distribution: São Tomé and Príncipe

Host: Hipposideros ruber (Chiroptera: Hipposideridae)

Paraperiglischrus sternalis Petrova et Taskaeva

Paraperiglischrus sternalis Petrova & Taskaeva, 1975: 59; Uchikawa, 1979a: 12

Type locality: China

Distribution: China, Japan

Host: Rhinolophus ferrumequinum, Rh. f. mikado (Chiroptera: Rhinolophidae)

Genus Periglischrus Kolenati

Periglischrus Kolenati, 1857: 60

Type species: Periglischrus caligus Kolenati, 1857

Periglischrus acutisternus Machado-Allison

Periglischrus acutisternus Machado-Allison, 1964: 200; 1965: 263; Herrin & Tipton, 1966: 28; Machado-Allison & Antequera, 1969: 374; 1971: 3; Herrin & Tipton, 1975: 28; Morales-Malacara, 2001: 194; Dantas -Torres et al., 2009; Almeida et al., 2011: 755; Gettinger, 2018: 3

Periglischrus tiptoni Furman, 1966: 147 – type: Panama, Panama City, from: Phyllostomus elongatus, Ph. hastatus panamensis, Ph. h. hastatus, Ph. d. discolor, Trachops cirrhosus (Phyllostomatidae), Myotis chiloensis (Vespertilionidae)

Type locality: Caripito, Monagas (Venezuela)

Distribution: Brazil, Colombia, Mexico, Panama, Peru, Trinidad, Venezuela

Hosts: Artibeus cinereus, A. fuliginosus, A. jamaicensis, Carolia perspicillata, Chiroderma villosum, Desmodus rotundus, Mimon bennetti, Phyllostomus concolor, Ph. discolor, Ph. elongatus, Ph. hastatus panamensis, Ph. h. hastatus, Sturnira lilium, Trachops cirrhosa, Uroderma magnirostrum, Vampyrops sp. (Chiroptera: Phyllostomatidae); Pteronotus parnelli, P. suapurensis (Chiroptera: Mormoopidae), Molossops planirostris (Chiroptera: Molossidae), Myotis albescens, M. chiloensis (Chiroptera: Vespertilionidae)

Periglischrus caligus Kolenati

Periglischrus caligus Kolenati, 1857: 60; 1859: 79; Oudemans, 1903: 135; Vitzthum, 1932: 36; 1941: 776; Stiles & Nolan, 1931: 633; Radford, 1942: 67; 1950a: 39; 1950b: 376; Baker & Wharton, 1952: 63; Tibbets, 1957: 13; Rudnick, 1960: 196; Furman, 1966: 142; Dusbábek & Lukoschus, 1971: 149; Herrin & Tipton, 1975: 40; Palacios-Vargas et al., 1985: 140; Gettinger & Gribel, 1989; Sheeler Gordon & Owen, 1999; Morales-Malacara, 2001: 194; Mendoza-Uribe & Chavez-Chorocco, 2003; Dick et al., 2007: 373; Silva et al., 2009: 470

Type locality: Suriname

Distribution: Bolivia, Brazil, Mexico, Panama, Peru, Suriname, Venezuela

Hosts: Artibeus planirostris, Glossophaga leachi, G. longirostris, G. morenoi, G. soricina, Platyrrhinus lineatus (Chiroptera: Phyllostomatidae, Phyllostomatinae)

Periglischrus cubanus Dusbábek

Periglischrus cubanus Dusbábek, 1968: 8; Gannon & Willig, 1994: 282; Morales-Malacara, 2001: 194

Type locality: Cueva de Santa Catalina, Camarioca, Prov. de Matanzas, Cuba

Distribution: Cuba

Hosts: Brachyphylla cavernarum, B. nana, Erophylla s. sezecorni, Phylonycteris poeyi (Chiroptera: Phyllostomatidae, Phyllostomatinae)

Periglischrus delfinadoae Dusbábek

Periglischrus delfinadoae Dusbábek, 1967: 527; 1968: 6; Morales-Malacara, 2001: 194

Type locality: Soroa, Prov. de Pinar del Rio, Cuba

Distribution: Cuba, Mexico

Hosts: Macrotus waterhousi, M. w. minor (Chiroptera: Phyllostomatidae, Macrotinae)

Periglischrus dusbabeki Machado-Alison et Antequera

Periglischrus dusbabeki Machado-Allison & Antequera, 1971: 9; Morales-Malacara, 2001: 194

Type locality: Yaracuy State, 19 km NW Urama

Distribution: Venezuela

Host: Mimon crenulatum (Chiroptera: Phyllostomatidae)

Periglischrus eurysternus Morales-Malacara, Juan et Juste

Periglischrus eurysternus Morales-Malacara, Juan & Juste, 2002: 304

Type locality: Mexico

Distribution: Mexico, Panama

Host: Tonatia saurophila (Chiroptera: Phyllostomatidae)

Periglischrus gameroi Machado-Alison et Antequera

Periglischrus gameroi Machado-Allison & Antequera, 1971: 6; Herrin & Tipton, 1975: 21; Morales-Malacara, 2001: 194; Gettinger, 2018: 3

Type locality: Trujillo State, 18 km N Valera (El Cenizo)

Distribution: Mexico, Peru, Venezuela

Host: Lonchorhina aurita (Chiroptera: Phyllostomatidae)

Periglischrus grandisoma Herrin et Tipton

Periglischrus grandisoma Herrin & Tipton, 1975: 30; Morales-Malacara, 2001: 194; Gettinger, 2018

Type locality: San Juan Rio Maniplare (155 m), 163 km ESE Pto Ayacucho, T.F. Amazonas, Venezuela

Distribution: Peru, Venezuela

Host: Phylloderma stenops (Chiroptera: Phyllostomatidae)

Periglischrus herrerai Machado-Allison

Periglischrus herrerai Machado-Allison, 1965a: 282; Machado-Allison & Antequera, 1969: 375; Herrin & Tipton, 1975: 55; Morales-Malacara, 2001: 195; Mendoza-Uribe & Chavez-Chorocco, 2003; Rojas et al., 2008: 185; Almeida et al., 2011: 755; Gettinger, 2018: 3

Periglischrus desmodi Furman, 1966: 139 – Type: St. Patrick’s Estate, Arima Valley, Trinidad (syn. by Furman, 1966: 166)

Type locality: Caripito, Monagas (Venezuela)

Distribution: Brazil, Colombia, Costa Rica, Mexico, Panama, Peru, Trinidad, Venezuela

Hosts: Artibeus planirostris (Chiroptera: Phyllostomatidae), Desmodus rotundus (Chiroptera: Phyllostomatidae, Desmodontinae); Myotis nigricans (Chiroptera: Vespertilionidae)

Periglischrus hopkinsi Machado-Allison

Periglischrus hopkinsi Machado-Allison, 1965a: 275; Herrin & Tipton, 1975: 53; Morales-Malacara, 2001: 195; Gettinger, 2018: 3

Type locality: Boca de Villacoa, Bolivar

Distribution: Peru, Venezuela

Hosts: Lionycteris spurrelli, Lonchophylla robusta (Chiroptera: Phyllostomatidae)

Periglischrus iheringi Oudemans

Periglischrus iheringi Oudemans, 1902: 38; Stiles & Nolan, 1931: 633; Rudnick, 1960: 197; Machado-Allison, 1964: 194; Machado-Allison & Antequera, 1969: 373; Furman, 1966: 135; Dusbábek, 1970b: 275; Dusbábek & Lukoschus, 1971: 149; Hoffmann, de Barrera & Méndez, 1972: 152; Herrin & Tipton, 1975: 58; Whitaker & Brian Abrell, 1987; Gannon & Willig, 1994: 282; Sheeler – Gordon & Owen, 1999: 92; Morales-Malacara, 2001: 195; Mendoza-Uribe & Chavez-Chorocco, 2003; Dick et al., 2007: 373; Dantas -Torres et al., 2009; Silva et al., 2009: 470; Almeida et al., 2011: 755; Reeves et al., 2016: 1218; Ramírez-Martínez et al., 2016: 312 ; Gettinger, 2018: 3

Periglischrus jheringi Oudemans: Oudemans, 1903: 135; Banks, 1915: 73; Radford, 1942: 67; 1950: 376

Periglischrus meridensis Hirst, 1927: 335 – type from Artibeus jamaicensis, Merida, Venezuela (syn. by Rudnick, 1960: 197); Allen, 1940: 303; Radford, 1942: 67; 1950: 376

Spinturnix ewingia Wharton, 1938: 139 – type from Artibeus jamaicensis yucatanicus, Second cave on San Roque Road, Yucatan, Mexico (syn. by Rudnick, 1960: 197); Radford, 1942: 66; 1950: 375

Spinturnix artibiensis Radford, 1951: 97 – type from Artibeus lituratus palmarum, Trinidad, British West Indies (syn. by Rudnick, 1960: 197)

Type locality: São Paulo, Brazil

Distribution: Bolivia, Brazil, Colombia, Cuba, Guatemala, Honduras, Mexico, Panama, Paraguay, Peru, Puerto Rico, Surinam, Trinidad, Venezuela, Virgin Islands

Hosts: Artibeus cinereus aztecus, A. cinereus, A. fimbriatus, A. hirsutus, A. intermedius, A. jamaicensis, A. lituratus, A. litturatus fallax, A. lituratus palmarum, A. obscurus, A. planirostris, A. toltecus, Chiroderma salvini, Dermanura azteca, D. phaeotis, D. tolteka, Enchisthenes hartii, Glossophaga soricina, Lophostoma silviculum, Phyllostomus discolor, Platyrrhinus helleri, P. lineatus, Stenoderma rufum, S. rufum darioi, Sturnira lilium, Uroderma bilobatum, Vampyressa pusilla, Vampyrodes major, Vampyrops helleri, V. lineatus, V. vittatus (Chiroptera: Phyllostomatidae, Phyllostomatinae), Desmodus rotundus, D. r. murinus (Chiroptera: Phyllostomatidae, Desmodontinae), Molossus ater (Chiroptera: Molossidae), Noctilio albiventris (Chiroptera: Noctilionidae)

Periglischrus leptosternus Morales-Malacara et López-Ortega

Periglischrus leptosternus Morales-Malacara & López-Ortega, 2001: 153; Ramírez-Martínez et al., 2016: 312

Type locality: Central Mexico

Distribution: Mexico

Host: Choeronycteris mexicana (Chiroptera: Phyllostomatidae)

Periglischrus micronycteridis Furman

Periglischrus micronycteridis Furman, 1966: 147; Herrin & Tipton, 1975: 21; Morales-Malacara, 2001: 194; Gettinger, 2018: 4; Graciolli, Guerrero & Catzeflis, 2019: 8

Type locality: Borinquen Highway (Canal Zone, Panama)

Distribution: Brazil, French Guiana, Panama, Peru, Trinidad, Venezuela

Hosts: Micronycteris megalotis microtis, M. m. megalotis, M. minuta, Trinycteris nicefori (Chiroptera: Phyllostomatidae, Micronycterinae)

Periglischrus moucheti Till

Periglischrus moucheti Till, 1958: 252

Type locality: Cameroon, Yaounde

Distribution: Cameroon, DR Congo

Hosts: Hipposideros caffer centralis, H. caffer guineensis (Chiroptera: Hipposideridae)

Periglischrus nycteris Till

Periglischrus nycteris Till, 1958: 255

Type locality: Banco Forest, Ivory Coast

Distribution: Ivory Coast

Host: Nycteris arge (Chiroptera: Nycteridae)

Periglischrus ojastii Machado-Allison

Periglischrus ojastii Machado-Allison, 1964: 197; Machado-Allison & Antequera, 1969: 374; Sheeler Gordon & Owen, 1999: 92; Morales-Malacara, 2001: 195; Presley, 2012: 61; Ramírez-Martínez et al., 2016: 312

Periglischrus ojasti: Dick, Gettinger & Gardner, 2007: 373; Dantas -Torres et al., 2009; Gettinger, 2018: 4

Periglischrus aitkeni Furman, 1966: 137

Type locality: Estado Monagas, Caripito

Distribution: Bolivia, Brazil, Colombia, Mexico, Panama, Peru, Venezuela

Hosts: Carollia perspicillata, Sturnira erythromos, S. lilium, S. ludovici, S. ororaphilum, S. parvidens (Chiroptera: Phyllostomatidae, Phyllostomatinae)

Periglischrus paracaligus Herrin et Tipton

Periglischrus paracaligus Herrin & Tipton, 1975: 41; Morales-Malacara, 2001: 195

Type locality: nr. Cojoro, 36 km NNE Paraguaipoa, Zulia, Venezuela

Distribution: Mexico, Venezuela

Hosts: Leptonycteris curasoae, L. nivalis (Chiroptera: Phyllostomatidae)

Periglischrus paracutisternus Machado-Allison et Antequera

Periglischrus paracutisternus Machado-Allison & Antequera, 1971: 13; Herrin & Tipton, 1975: 27; Morales-Malacara, 2001: 194; Gettinger, 2018: 4

Type locality: Venezuela

Distribution: Mexico, Peru, Venezuela

Hosts: Anoura geoffroyi, Rhogeessa tumida, Trachops cirrhosis (Chiroptera: Phyllostomatidae, Phyllostomatinae)

Periglischrus paratorrealbai Herrin et Tipton

Periglischrus paratorrealbai Herrin & Tipton, 1975: 36; Morales-Malacara, 2001: 194

Type locality: Boca Mavaca, 84 km SSE Esmeralda (138 m), T.F. Amazonas, Venezuela

Distribution: Venezuela

Hosts: Phylloderma stenops, Tonatia bidens (Chiroptera: Phyllostomatidae)

Periglischrus paravargasi Herrin et Tipton

Periglischrus paravargasi Herrin & Tipton, 1975: 46; Morales-Malacara, 2001: 195; Gettinger, 2018: 4; Graciolli, Guerrero & Catzeflis, 2019: 8

Type locality: 2 km SW Altamira (620 m), Barinas, Venezuela

Distribution: Brazil, French Guiana, Peru, Venezuela

Hosts: Anoura caudifera, A. geoffroyi, Phyllostomus discolor (Chiroptera: Phyllostomatidae)

Periglischrus parvus Machado-Allison

Periglischrus parvus Machado – Allison, 1964: 195; 1965: 268; Herrin & Tipton, 1975; Morales-Malacara, 2001: 194

Type locality: Estado Monagas, Caripito

Distribution: Venezuela

Hosts: Micronycteris hirsuta, M. megalotis, M. nicefori (Chiroptera: Phyllostomatidae)

Periglischrus ramirezi Machado-Allison et Antequera

Periglischrus ramirezi Machado-Allison & Antequera, 1971: 3; Herrin & Tipton, 1975: 19; Morales-Malacara, 2001: 194; Gettinger, 2018: 4

Type locality: Bolivar State, 59 km SE El Dorado (El Mónaco)

Distribution: Peru, Venezuela

Host: Rhinophylla pumilio (Chiroptera: Phyllostomatidae)

Periglischrus setosus Machado-Alison

Periglischrus setosus Machado-Alison, 1964: 199; Machado-Allison & Antequera, 1969: 375

Type locality: Valle de Caripe, Monagas (Venezuela)

Distribution: Columbia, Venezuela

Host: Glossophaga soricina (Chiroptera: Phyllostomatidae)

Periglischrus squamosus Machado-Alison

Periglischrus squamosus Machado-Alison, 1965a: 279

Type locality: Estado Miranda, Quebrada de Caurimare (Venezuela)

Distribution: Venezuela

Hosts: Anoura caudifera, A. cultrata, A. geoffroyi (Chiroptera: Phyllostomatidae)

Periglischrus steresotrichus Morales-Malacara et Juste

Periglischrus steresotrichus Morales-Malacara & Juste, 2002: 299

Type locality: Mexico

Distribution: Mexico

Host: Tonatia evotis (Chiroptera: Phyllostomatidae)

Periglischrus tonatii Herrin et Tipton

Periglischrus tonatii Herrin & Tipton, 1975: 23; Morales-Malacara, 2001: 194; Gettinger, 2018: 4

Type locality: 25 km S Pto. Ayacucho (114 m), Venezuela

Distribution: Brazil, Peru, Venezuela

Hosts: Tonatia bidens, T. brasiliensis, T. carrikeri, T. silvicola (Chiroptera: Phyllostomatidae). One chance finding on Desmodus rotundus, one on Lophostoma silviculum.

Periglischrus torrealbai Machado-Alison

Periglischrus torrealbai Machado-Alison, 1965a: 276; Machado-Allison & Antequera, 1969: 373; 1971: 3; Yunker et al., 1975: 252; Morales-Malacara, 2001: 194; Almeida et al., 2018: 256; Gettinger, 2018: 4

Periglischrus inflatiseta Furman, 1966: 134.

Type locality: Valle de Caripe, Monagas (Venezuela)

Distribution: Brazil, Colombia, Panama, Peru, Venezuela

Hosts: Lophostoma silviculum, Phyllostomus discolor, Ph. elongatus, Ph. hastatus, ? Sturnira lilium (Chiroptera: Phyllostomatidae)

Periglischrus triaenopsis Benoit

Periglischrus triaenopsis Benoit, 1961: 397

Type locality: Tanga, Sigi River

Distribution: Tanzania

Host: Triaenops afer (Chiroptera: Hipposideridae)

Periglischrus vargasi Hoffmann

Periglischrus vargasi Hoffmann, 1944: 91; Baker & Wharton, 1952: 62; Tibbetts, 1957: 14; Rudnick, 1960: 199; Furman, 1966: 143; Palacios -Vargas et al., 1985: 140; Gannon & Willig, 1994: 282; Sheeler – Gordon & Owen, 1999: 92; Morales-Malacara, 2001: 195; Silva et al., 2009: 470; Gettinger, 2018: 5; Ramírez-Martínez et al., 2016: 312

Type locality: Yerbabuena, Guerrero, Mexico

Distribution: Brazil, Guatemala, Mexico, Panama, Peru, Puerto Rico, Venezuela, USA (Texas)

Hosts: Anoura caudifera, A. cultrata, A. geoffroyi, A. g. lasiopyga, Artibeus jamaicensis, Leptonycteris yerbabuenae, L. sanborni, Macrotus californicus, Monophyllus redmani, Sturnira lilium, Trachops cirrhosus (Chiroptera: Phyllostomatidae)

Periglischrus zuluensis Till

Periglischrus zuliuensis Till, 1958: 250

Type locality: Between Tete Pan and the Pongola River, Ubombo District, Zululand

Distribution: South Africa

Host: Nycteris thebaica capensis (Chiroptera: Nycteridae)

Genus Spinturnix von Heyden

Spinturnix von Heyden, 1826: 612

Type species: Pteroptus myoti Kolenati, 1856 (designated by Opinion 128 of the International Commission on Zoological Nomenclature – 1936)

Pteroptus Dufour, 1832: 98.

Type: Pteroptus vespertilionis Dufour

Celeripes Walckenaer et Gervais, 1844: 227

Diplostaspis Kolenati, 1857: 60.

Type: D. nattererii Kolenati

Monostaspis Kolenati, 1857: 60

Heterostaspis Kolenati, 1859: 157

Paraspinturnix Rudnick, 1960: 231 – syn. by Domrow, 1972: 556

Type species: Paraspinturnix globosus Rudnick, 1960

Spinturnix abyssinica Hirst

Spinturnix abyssinicus Hirst, 1927: 333; Stiles & Nolan, 1931: 630; Radford, 1942: 66; 1950: 375; Zumpt, 1951: 80; Zumpt & Till, 1954: 216; Rudnick, 1960: 216

Type locality: Addis-Ababa

Distribution: Cameroon, Ethiopia

Host: Scotophilus leucogaster (Chiroptera: Vespertilionidae)

Spinturnix acuminata acuminata (C.L. Koch)

Monostaspis arcuatus: Kolenati, 1857: 60

Spinturnix noctulae Oudemans, 1910: 68.

Type: Germany, from Nyctalus noctula

Pteroptus acuminatus C.L. Koch, 1836: 21

Spinturnix acuminata (C.L. Koch): Karataş & Çakır, 2004

Spinturnix acuminatus (C.L. Koch): Turk, 1953: 13; Vshivkov, 1963: 325; Costa, 1966: 80; 1967: 106; Dusbábek, 1970: 62; Boldori, 1970; Mahnert, 1971: 927; Pan & Teng, 1973: 86; Mehl, 1979: 33; Haitlinger & Ruprecht, 1982: 171; Rybin, 1983: 356; Stanjukovich, 1990: 195; Medvedev et al., 1991: 28; Estrada-Peña, Balcells & Serra-Cobo, 1991: 257; Krochko, 1995: 83; Haitlinger & Walter, 1997: 104; Haitlinger & Łupicki, 2008: 125; Ahamad et al., 2013: 140

Spinturnix acuminata acuminata (C.L. Koch): Uchikawa et al., 1994: 296

Type locality: Europe

Distribution: Afghanistan, Austria, Azerbaijan, Belgium, Bulgaria, China, Denmark, England, Estonia, Germany, Holland, India, Israel, Italy, Japan, Kazakhstan, Kirghizstan, Malaysia, Moldavia, Morocco, Norway, Poland, Romania, Russia, Slovakia, Spain, Sri Lanka, Turkey, Ukraine

Hosts: Barbastella leucomelas, Eptesicus serotinus, Hypsugo savii, Murina leucogaster, Myotis blythi, M. dasycneme, M. daubentoni, M. myotis, Nyctalus lasiopterus, N. leisleri, N. noctula, Pipistrellus kuhli, P. pipistrellus, Scotophilus heathii (Chiroptera: Vespertilionidae)

Spinturnix acuminata helvetiae Deunff, Keller et Aellen

Spinturnix acuminatus acuminatus (Koch, 1836) sensu Pinchuk (pro parte), 1971

Spinturnix helvetiae Deunff, Keller & Aellen, 1986: 805; Stanyukovich, 1997; Ferenc & Myslajek, 2003: 278; Krištofík & Danko, 2012: 173

Spinturnix acuminata helvetiae Deunff, Keller et Aellen: Uchikawa et al., 1994: 296

Type locality: Col de Bretolet, Valais

Distribution: Belarus, Germany, Moldavia, Poland, Slovakia, Switzerland

Hosts: Myotis daubentoni, Nyctalus leisleri (Chiroptera: Vespertilionidae)

Spinturnix aelleni Benoit

Spinturnix aelleni Benoit, 1959b: 24

Type locality: Lambaréné

Distribution: Gabon

Host: Eptesicus tenuipinnis (Chiroptera: Vespertilionidae)

Spinturnix americana (Banks)

Pteroptus americanus Banks, 1902: 173; Vitzthum, 1932: 37

Spinturnix americanus (Banks): Banks, 1915: 72; van Eyndhoven, 1941: 53; Rudnick, 1960: 218; Whitaker & Easterla, 1975; Herrin & Tipton, 1975; Pence, Knox Jones & Knipping, 1981; Whitaker, 1982: 193; Whitaker & Maser, 1985: 74; Dick, Gannon, Little & Patrick, 2003: 815; Poissant & Broders, 2008: 515; Presley, 2012: 61; Ahamad et al., 2013: 140; Lima Silva & Graciolli, 2013: 175; Gettinger, 2018: 5; Graciolli, Guerrero & Catzeflis, 2019: 8

Pteroptus echinipes Banks, 1910: 5 – type from Myotis lucifugus, Homer, New York (syn. by Rudnick, 1960: 218)

Spinturnix araguensis Vitzthum, 1932: 34 – type from Myotis nigricans, Puerto de la Cruz, Aragua, Venezuela (syn. by Rudnick, 1960: 218)

Spinturnix iowae Keegan, 1943 – type from Myotis lucifugus, Milford, Iowa (syn. by Rudnick, 1960: 218); Radford, 1950: 375

Spinturnix carloshoffmanni Hoffmann, 1944: 185 – type: Cerro del Xitle, Tlalpan, Mexico, from Natalus m. mexicanus (Natalidae); Radford, 1950: 375; Baker & Wharton, 1952: 62; Rudnick, 1960: 222

Spinturnix grossus: Kohls & Jellison (not Banks, 1910), 1948: 117

Spinturnix americana (Banks): Uchikawa et al., 1994: 291

Type locality: a cave in Indiana

Distribution: Brazil, Canada, French Guiana, Mexico, Panama, Paraguay, Peru, USA (incl. Alaska), Venezuela; Malaysia (!?)

Hosts: Antrozous pallidus, Eptesicus fuscus, Myotis albescens, M. chiloensis, M. dominicensis, M. evotis, M. keaysi, M. keenii, M. lucifugus, M. n. nigricans, M. septentrionalis, M. simus, M. sodalis, M. thysanodes, M. velifer incautus, M. velifer velifer, M. volans, M. yumanensis, M. yumanensis saturatus, M. yumanensis sociabilis, Corynorhynus mexicanus (Chiroptera: Vespertilionidae), Natalus mexicanus mexicanus (Chiroptera: Natalidae); Eumops glaucinus (Chiroptera: Molossidae); Saccopteryx canescens (Chiroptera: Emballonuridae)

Spinturnix bakeri Rudnick

Spinturnix bakeri Rudnick, 1960: 226; Dusbábek, 1968: 2; Herrin & Tipton, 1975: 63; Gannon & Willig, 1994: 282; Ahamad et al., 2013: 140; Gettinger, 2018: 5

Type locality: Sunol, Alameda County, California

Distribution: Canada, Cuba, Peru, Puerto Rico, USA (Arkansas, California, Kansas, Minnesota, Oregon, Pennsylvania), Venezuela; Malaysia (!?)

Hosts: Eptesicus fuscus bernardinus, E. fuscus pallidus, E. fuscus dutertreus, E. fuscus fuscus, E. montosus, Myotis keaysi, M. volans (Chiroptera: Vespertilionidae)

Spinturnix banksi Rudnick

Spinturnix banksi Rudnick, 1960: 221

Type locality: Bone Cave, eight miles north of Batesville, Independence Co., Arkansas

Distribution: USA (Arkansas, Illinois, Oklahoma, Tennessee)

Host: Myotis grisescens (Chiroptera: Vespertilionidae)

Spinturnix bechsteini Deunff, Walter, Bellido et Volleth

Spinturnix bechsteini Deunff, Walter, Bellido & Volleth, 2004: 826; Reckardt & Kerth, 2009; Bruyndonckx, Henry et al., 2009: 3581; Haitlinger & Piksa, 2012: 15; Krištofík & Danko, 2012: 172

Type locality: Ebrach, Steigerwald, Bavaria

Distribution: Belgium, Bulgaria, France, Germany, Italy, Poland, Slovakia, Tunisia

Host: Myotis bechsteini (Chiroptera: Vespertilionidae)

Spinturnix bregetovae Stanyukovich

Spinturnix bregetovae Stanyukovich, 1996: 106; Orlova, Kazakov et al., 2016: 3; Scheffler et al., 2012: 144

Type locality: Primorsky Kray, Suputinsky Reserve (Russia)

Distribution: Mongolia, Russia (Far East)

Hosts: Myotis gracilis, M. ikonnikovi, M. sibiricus (Chiroptera: Vespertilionidae)

Spinturnix brevisetosa Gu et Wang

Spinturnix brevisetosus Gu & Wang, 1984: 165

Type locality: Xingyi County

Distribution: China

Host: bats (Chiroptera)

Spinturnix chiengmai Prasad

Spinturnix chiengmai Prasad, 1970: 661

Type locality: Huai, Mae Sanam, Hod, Chiengmai

Distribution: Thailand

Host: bat (Chiroptera)

Spinturnix dasycnemi (Kolenati)

Pteroptus dasycnemi Kolenati, 1856: 177; 1856: 26 ; Berlese, 1892: 28; Stiles & Nolan, 1931: 631

Diplostaspis dasycnemi: Kolenati, 1857: 60; 1859: 159; 1860: 106; C.L. Коch, 1865: 201

Spinturnix dasycnemi: Buitendijk, 1945: 308; Estrada-Peña & Sanchez, 1989: 107; Uchikawa et al., 1994: 291; Krištofík & Danko, 2012: 173

Type locality: Europe

Distribution: Holland, Italy, Poland, Slovakia

Host: Myotis dasycneme (Chiroptera: Vespertilionidae)

Spinturnix daubentonii (Kolenati)

Diplostaspis daubentonii Kolenati, 1857: 60; Stiles & Nolan, 1931: 631; Uchikawa et al., 1994: 291

Diplostaspis stellata Kolenati, 1859: 165

Spinturnix daubentonii, Neumann, 1942: 54; 1943: 106

Spinturnix andegavinus Deunff, 1977: 607; Estrada-Peña, 1990: 95; Balcells & Serra-Cobo, 1991: 257; Dietz & Walter, 1995: 451; Haitlinger & Walter, 1997: 102; Lučan, 2006: 147; Scheffler, 2016: 377

Type locality of Diplostaspis daubentonii: Europe

Type locality of S. andegavinus: Maine-et-Loire, caves de Chanzelles, de Guédéniau (France); paratypes from other places in France, Portugal and Spain

Distribution: Czech Rep., France, Germany, Mongolia, Poland, Portugal, Slovakia, Spain

Hosts: Myotis myotis, M. nattereri, M. dasycneme, M. daubentoni (Chiroptera: Vespertilionidae)

Spinturnix delacruzi Estrada-Peña, Ballestra et Ibanez

Spinturnix delacruzi Estrada-Peña, Ballestra & Ibanez, 1992: 307

Type locality: Niefang

Distribution: Equatorial Guinea

Host: Rousettus aegyptiacus (Chiroptera: Pteropodidae)

Spinturnix domrowi Deunff et Volleth

Spinturnix domrowi Deunff & Volleth, 1987: 112

Type locality: Kuala Lumpur

Distribution: Malaysia

Host: Glischropus tylopus (Chiroptera: Vespertilionidae)

Spinturnix emarginata (Kolenati)

Pteroptus emarginatus Kolenati, 1856: 28; Berlese, 1892: 28; Stiles & Nolan, 1931: 631

Diplostaspis emarginatus: Kolenati, 1857: 60

Diplostaspis ciliata Kolenati, 1859: 164; 1860: 115; C.L. Koch, 1865: 178

Spinturnix emarginatus Kolenati: Rudnick, 1960: 233; Dusbábek, 1964a: 8; Beron, 1973: 126; 1974: 67; Deunff, 1977: 605; Estrada-Peña, Balcells & Serra-Cobo, 1991: 257; Orlova & Orlov, 2018: 329; 2019: 237

Spinturnix emarginata Kolenati: Uchikawa et al., 1994: 291; Krištofík, Krzysztof & Sahanowicz, 2012: 103; Krištofík & Danko, 2012: 173

Type locality: Europe

Distribution: Bulgaria, Crimea, Czech Rep., France, Holland, Hungary, Poland, Serbia, Slovakia, Spain, Tadjikistan

Hosts: Myotis emarginatus (Chiroptera: Vespertilionidae), Rhinolophus euryale (Chiroptera: Rhinolophidae)

Spinturnix eptesici Domrow

Spinturnix eptesici Domrow, 1972: 561; 1979: 318; 1988: 890

Type locality: Queensland, Mt. Iron Pot

Distribution: Australia

Hosts: Vespadelus [Eptesicus] douglasorum, V. [E.] pumilus, Nyctophilus arnhemensis (Chiroptera: Vespertilionidae), Taphozous georgianus (Chiroptera: Emballonuridae)

Spinturnix faini Benoit

Spinturnix faini Benoit, 1959b: 25

Type locality: Ituri, Mont Wago nr. Blukwa

Distribution: DR Congo

Host: Eptesicus rendalli (Chiroptera: Vespertilionidae)

Spinturnix globosa (Rudnick)

Paraspinturnix globosus Rudnick, 1960: 231; Allred & Beck, 1966: 9; Furman, 1966: 153; Whitaker, 1973: 1149

Paraspinturnix globosa Rudnick: Whitaker, Deunff & Belwood, 2000: 445

Spinturnix globosus (Rudnick): [Domrow, 1972: 556]; Whitaker, 1982: 192; Whitaker & Maser, 1985: 74

Type locality: Nickajack Cave, three miles south of Jasper, Marion Co., Tennessee

Distribution: Panama, USA (Tennessee, Indiana, Utah, Arizona, Oklahoma, Oregon)

Hosts: Myotis chiloensis or M. n. nigricans, M. sodalis, M. volans (Chiroptera: Vespertilionidae), Glossophaga soricina (Chiroptera: Phyllostomatidae)

Spinturnix intecta Dusbábek et Bergmans

Spinturnix intectus Dusbábek & Bergmans, 1980: 67

Type locality: Ibadan

Distribution: Nigeria

Host: Scotophilus dinganii (Chiroptera: Vespertilionidae)

Spinturnix kolenatii Oudemans

Pteroptus arcuatus: Kolenati, 1856: 163. According to Runick, 1960, arcuatus Koch should not be used in Spinturnicidae.

Pteroptus carnifex Kolenati, 1856: 28

Diplostaspis carnifex Kolenati, 1857: 60

Spinturnix carnifex: Stiles & Nolan, 1931: 631

Spinturnix kolenatii Oudemans, 1910: 68

Spinturnix murinus: Allen, 1940: 303

Pteroptus carnifex Kolenati (not C.L. Koch, 1839, not Oudemans, 1902), 1856: 28; Berlese, 1892: 28; Huhta, 2016: 146

Diplostaspis carnifex (Kolenati)(not C.L. Koch, 1839, not Oudemans, 1902), 1857: 60; 1859: 158; 1860: 51; C. Koch, 1865: 209

Diplostaspis nilssonii Kolenati, 1857

Spinturnix carnifex: Stiles & Nolan (in part, not C.L. Koch, 1839, not Oudemans, 1902), 1931: 631

Spinturnix vespertilionis, Oudemans (not of other authors), 1903: 28; 1904: 72; 1931: 109; Vitzthum, 1932: 3; Buitendijk, 1945: 308

Spinturnix kolenatii Oudemans, 1910: 68; Stiles & Nolan, 1931: 631; van Eyndhoven, 1947: 136; Rudnick, 1960: 223; Dusbábek, 1970a: 62; 1970b: 44; Uchikawa & Wada, 1979: 123; Scheffler et al., 2010:

Spinturnix kolenatii Oudemans: Neumann, 1942: 54; Vshivkov, 1963: 325; Mrciak & Nyholm, 1967: 13; Haitlinger & Ruprecht, 1977: 333; 1985: 616; Rybin, 1983: 359; Estrada-Peña, Balcells & Serra-Cobo, 1991: 257; Medvedev et al., 1991: 28; Orlov & Orlova, 2010: 41; Krištofík & Danko, 2012: 173

Spinturnix kolenati: Nikitchenko, 1995: 88; Scheffler et al., 2016: 401

Type locality: Europe

Distribution: Afghanistan, England, Finland, Germany, Holland, Hungary, Japan, Kirghizstan, Mongolia, Norway, Poland, Slovakia, Russia, Spain, Ukraine

Hosts: Eptesicus nilssoni, E. parvus, E. s. serotinus, E. turcomanus, Myotis blythi, M. brandti, M. daubentoni, M. mystacinus, Nyctalus noctula, Pipistrellus nathusii, P. pipistrellus, Plecotus auritus, Vespertilio murinus (Chiroptera: Vespertilionidae)

Spinturnix kolenatoides Ye et Ma

Spinturnix kolenatoides Ye & Ma, 1996: 421

Type locality: Yining City, Xinjiang Uygur

Distribution: China

Host: “bat”(Chiroptera)

Spinturnix lawrencei Zumpt

Spinturnix lawrencei Zumpt, 1951: 80; Zumpt & Till, 1954: 216; Rudnick, 1960: 210; Uchikawa et al., 1994

Type locality: Noodsberg Caves near Pietermaritzburg, Natal

Distribution: Kenya, South Africa

Host: Myotis tricolor (Chiroptera: Vespertilionidae)

Spinturnix loricata Domrow

Spinturnix loricata Domrow, 1972: 565; 1988: 890

Type locality: S. Australia, Naracoorte, Bat Cave

Distribution: Australia (S.A.)

Host: Miniopterus schreibersi (Chiroptera: Miniopteridae)

Spinturnix maedai Uchikawa et Wada

Spinturnix maedai Uchikawa & Wada, 1979: 123; Uchikawa et al., 1994: 296

Type locality: Misato, Nagano Pref.

Distribution: Japan

Hosts: Murina ussuriensis, M. hilgendorfi (Chiroptera: Vespertilionidae)

Spinturnix mexicana Rudnick

Spinturnix mexicanus Rudnick, 1960: 230

Type locality: Isla Blanca, four miles west of Guaymas, Sonora, Mexico

Distribution: Mexico

Host: Pizonyx vivesi (Chiroptera: Vespertilionidae)

Spinturnix multisetosa Rudnick

Spinturnix multisetosus Rudnick, 1960: 209

Type locality: Namoroka, grotte Ambohimirija

Distribution: Madagascar

Host: Myotis goudotii (Chiroptera: Vespertilionidae)

Spinturnix myoti (Kolenati)

Pteroptus murinus Walckenaer & Gervais (not Thompson, 1935, not Allen, 1940), 1847: 545; Berlese, 1892: 28 (syn. by Rudnick, 1960: 204; see synonymy of Spinturnix vespertilionis sensu Sp. myoti in Oudemans, 1910: 68–69)

Pteroptus myotis Kolenati, 1856: 181

Pteroptus myoti Kolenati, 1856: 26; Haller, 1882: 303; Berlese, 1892: 28; Jablonowski, 1918: 3; Stiles & Nolan, 1931: 631

Diplostaspis myoti Kolenati, 1857: 60; 1859: 156; 1860: 122; C.L. Koch, 1865: 158; Stiles & Nolan, 1931: 631

Spinturnix myoti (Kolenati): Int. Comm. on Zool. Nom., 1936, Smiths. Misc. Coll. (Publ. 3395), 73(8): 29; Baker & Wharton, 1952: 63; Vshivkov, 1963: 325; Dusbábek, 1964: 6; 1966: 43; Beron & Kolebinova, 1964: 232; Costa, 1966: 80; 1967: 106; Beron, 1968: 158; 1973: 126; 1974: 67; Mahnert, 1971: 927); Uchikawa & Wada, 1979: 121; Mehl, 1979: 33; Stanjukovich, 1990: 193; Estrada-Peña, Balcells & Serra-Cobo, 1991: 257; Haitlinger & Walter, 1997: 104; Bobkova, 2002: 79; Lourenço & Palmeirim, 2007: 163; Orlova et al., 2011: 137; Krištofík & Danko, 2012: 174; Orlova, Kazakov et al., 2016: 4; Bendgeddou et al., 2017; Orlova & Orlov, 2018: 320

Pteroptus capaccinii Kolenati, 1856: 26

Diplostaspis nattererii Kolenati, 1857: 60

Diplostaspis nattereri: C.L. Koch, 1865: 173

Heterostaspis octastigma Kolenati, 1859: 167

Heterostaspis octostigma [sic!]: Vitzthum, 1932: 37

Pteroptus grossus Banks, 1910: 6

Spinturnix vespertilionis Hirst, 1927 (not of other authors)

Type locality: Europe

Distribution: Algeria, Austria, Bosnia and Herzehovina, Bulgaria, Crimea, Estonia, Finland, France, Great Britain, Hungary, India (Jammu & Kashmir), Israel, Japan, Kirghizstan, Macedonia, Mongolia, Montenegro, Norway, Poland, Portugal, Russia, Serbia, Spain, Turkey, Ukraine

Hosts: Barbastella barbastellus, Myotis blythi, M. brandti, M. capaccinii, M. emarginatus, M. dasycneme, M. daubentoni, M. ikonnikovi, M. macrodactylus, M. myotis, M. nattereri, M. petax, M. punicus, Nyctalus noctula, Pipistrellus nathusii, Plecotus auritus (Chiroptera: Vespertilionidae); Miniopterus schreibersi (Chiroptera: Miniopteridae); Rhinolophus euryale, Rh. hipposideros, Rh. ferrumequinum (Chiroptera: Rhinolophidae)

Spinturnix mystacina (Kolenati)

Diplostaspis mystacinus Kolenati (not mystacina, Oudemans, 1902), 1857: 60; Oudemans, 1902: 70

Diplostaspis mystacina Kolenati (not Oudemans, 1902) , 1859: 163; 1860: 96; C.L. Koch, 1865: 187

Spinturnix mystacinus (Kolenati): Oudemans (not mystacina Oudemans, 1902), 1910: 68; Buitendijk, 1945: 308; Rudnick, 1960: 207; Dusbábek, 1966: 43; Beron & Kolebinova, 1964: 234; Beron, 1968: 158; 1973: 126; Deunff, 1977: 613; Uchikawa & Wada, 1979: 124; Mehl, 1979: 33; Cicolani & Manilla, 1980: 32; Stanjukovich, 1990: 195; Stanjukovich & Malinovskiy, 1992; Haitlinger & Walter, 1997: 105; Lanza, 1999: 153; Scheffler, 2016: 401

Spinturnix mystacina (Kolenati): Stiles & Nolan (in part, not Oudemans, 1902), 1931: 631; Uchikawa et al., 1994; Bruyndonckx et al., 2009; Krištofík & Danko, 2012: 174

Spinturnix murinus Thompson, 1935 (not Walckenaer & Gervais, 1847)

Type locality: Europe

Distribution: Bulgaria, Czech Rep., Finland, France, Germany, Great Britain, Holland, Italy, Japan, Kazakhstan, Kirghizstan, Tadjikistan, Moldavia, Mongolia, Morocco, Norway, Poland, Romania, Russia, Slovakia, Spain

Hosts: Eptesicus gobiensis, E. serotinus, Hypsugo alaschanicus, Myotis alcathoe, M. aurascens, M. brandti, M. dasycneme, M. gracilis, M. hosonoi, M. ikonnikovi, M. myotis, M. mystacinus, M. nattereri, Nyctalus noctula, Plecotus auritus, Vespertilio murinus (Chiroptera: Vespertilionidae)

Spinturnix mystacina brandti Haitlinger

Spinturnix mystacina brandti Haitlinger, 1978: 483

Type locality: Wojcieszów Górny

Distribution: Poland

Hosts: Myotis brandti, M. mystacinus (Chiroptera: Vespertilionidae)

Spinturnix nobleti Deunff, Volleth, Keller et Aellen

Spinturnix nobleti Deunff, Volleth, Keller & Aellen, 1990: 478; Uchikawa et al., 1994: 296; Scheffler et al., 2012: 145

Type locality: Ortolo (Corsica)

Distribution: Corsica (France), France cont., Italy, Mongolia, Switzerland

Hosts: Hypsugo alaschanicus, H. savii (Chiroptera: Vespertilionidae)

Spinturnix novaehollandiae Hirst

Spinturnix novae – hollandiae Hirst, 1931: 564; van Eyndhoven, 1941: 54; Radford, 1942: 66; 1950: 375; Rudnick, 1960: 234

Spinturnix novaehollandiae Hirst: Domrow, 1959; 1972: 558; 1979: 318; 1988: Green, 1989; Uchikawa et al., 1994: 291

Spinturnix antipodianus Hirst, 1931: 563 – type: Cairns Distr., N. Queensland, Chiroptera indet.; van Eyndhoven, 1941: 54; Radford, 1942: 66; 1950: 375; Domrow, 1959; Rudnick, 1960: 233 – syn. by Domrow, 1988: 890

Spinturnix queenslandicus Prasad, 1969b: 577 – type: Border Waterhole, N.T., North of Camooweal, Queensland, from Chalinolobus gouldii venatoris – syn. by Domrow, 1972: 558

Type locality: Lucindale, South Australia

Distribution: Australia (S.A., W.A., N.S.W.)

Hosts: Chalinolobus dwyeri, Ch. gouldi, Ch. morio, Ch. picatus, Myotis adversus, Nycticeius greyi, N. rueppelli, Nyctophilus geoffroyi, Vespadelus [Eptesicus] pumilus (Chiroptera: Vespertilionidae); Miniopterus schreibersi (Chiroptera: Miniopteridae)

Spinturnix nudata Allred

Spinturnix nudatus Allred, 1969: 237

Type locality: Gupis, Kohighzar, West Pakistan

Distribution: Pakistan

Host: Chiroptera indet.

Spinturnix orri Rudnick

Spinturnix orri Rudnick, 1960: 228; Whitaker & Maser, 1985: 74

Spinturnix orii: Uchikawa et al., 1994: 296

Type locality: Cowell, Contra Costa Co., California

Distribution: Mexico, USA (California, Texas, Utah, Oregon)

Hosts: Antrozous dubiaquerqus, A. pallidus pacificus, A. p. cantwelli, A. p. pallidus, A. p. minor (Chiroptera: Vespertilionidae, Antrozoinae)

Spinturnix paracuminata Baker et Delfinado

Spinturnix paracuminatus Baker & Delfinado, 1964: 574

Spinturnix paracuminata Baker et Delfinado: Domrow, 1972: 561

Type locality: NE New Guinea, Suanimbu, E of Maprik

Distribution: Papua New Guinea

Hosts: Miniopterus sp.(Chiroptera: Miniopteridae); Myotis adversus, Pipistrellus angulatus (New Ireland), P. papuanus (Chiroptera: Vespertilionidae)

Spinturnix pindarensis Bhat

Spinturnix pindarensis Bhat, 1973: 471

Type locality: Uttar Pradesh, Dwali, 2770 m, Pindar Valley, Almora District

Distribution: India

Host: Myotis siligorensis (Chiroptera: Vespertilionidae)

Spinturnix plecotina (C.L. Koch)

Pteroptus plecotinus C.L. Koch, 1839: 23 (24); 1842: 128; Walckenaer & Gervais, 1847: 546; Berlese, 1892: 27

Pteroptus plecutinus [sic] C.L. Koch, 1839: 23

Pteroptus transversus Kolenati, 1856: 184; 1856: 26; Berlese, 1892: 28

Diplostaspis transversa Kolenati, 1859: 184

Diplostaspis transversa Kolenati, 1859: 164; 1860: 131; Stiles & Nolan, 1931: 632-

Spinturnix plecotinus (C.L. Koch): Oudemans, 1910: 67; 1936: 350; Hirst, 1927: 330; Vitzthum, 1929: 29; Stiles & Nolan, 1931: 632; van Eyndhoven, 1941: XXVIII; 1950: 376; Neumann, 1942: 54; Turk, 1945: 799; 1953: 13; Buitendijk, 1945: 308; Rudnick, 1960: 208; Cooreman, 1960a: 258; 1960b: 2; Dusbábek, 1964: 7; Beron, 1973: 127; Mitchell & Punzo, 1976; Deunff, 1977: 613; Haitlinger, 1978: 475; Mehl, 1979: 33; Uchikawa & Wada, 1979: 124; Gu et al., 1986: 336; Estrada-Peña et al., 1989; Stanjukovich, 1990: 195; Haitlinger & Walter, 1997: 103; Bobkova, 2002: 79; Scheffler et al., 2010: 376; 2012: 144; 2016: 401; Huhta, 2016: 147

Spinturnix plecoti Oudemans, 1902: 48 – type: Germany, from Plecotus auritus; 1903: 32; 1904: 72; Stiles & Nolan, 1931: 632

Spinturnix plecotina: Uchikawa et al., 1994: 291; Lanza, 1999: 153; Imaz, Aihartza & Totorika, 1999: 24; Toprak, 2009; Krištofík & Danko, 2012: 175

Type locality: Europe

Distribution: Afghanistan, Bulgaria, China, Czech Rep., Estonia, Finland, France, Germany, Great Britain, Holland, Ireland, Japan, Latvia, Mongolia, Morocco, Norway, Poland, Romania, Russia, Slovakia, Spain, Canary Islands, Taiwan, Turkey, Ukraine

Hosts: Barbastella barbastellus, B. leucomelas, Eptesicus nilssoni, E. serotinus, Myotis brandti, M. daubentoni, M. nattereri, Nyctalus noctula, Plecotus auritus, P. auritus sacrimontis, P. austriacus, ?P. kozlovi, P. ognevi, P. tenerifae (Chiroptera: Vespertilionidae), Rhinolophus euryale, Rh. ferrumequinum, Rh. lepidus (Chiroptera: Rhinolophidae)

Spinturnix psi (Kolenati)

Pteroptus psi Kolenati, 1856: 182

Diplostaspis psi Kolenati, 1857: 60; 1859: 161; 1860: 127

Spinturnix psi (Kolenati): Oudemans, 1925; Hirst, 1927; Hiregaudar & Bal, 1955, 1956; Rudnick, 1960: 211; Domrow, 1962: 281; Dusbábek, 1964b: 392; 1970: 62; Beron & Kolebinova, 1964: 232; Beron, 1973: 126; 1974: 67; Costa, 1966: 80; 1967: 106; Dusbábek, 1970a: 62; Pan & Teng, 1973: 86; Deunff, 1977: 615; Domrow & Nadchatram, 1978: 88; Uchikawa & Wada, 1979: 124; Whitaker, Baccus, Cleveland & Gu, 1990; Durden, Peterson et al., 1990: 53; Uchikawa et al., 1994: 299; Imaz, Aihartza & Totorika, 1999: 24; Lourenço & Palmeirim, 2007: 163; Krištofík & Danko, 2012: 175; Huhta, 2016: 147

Spinturnix amboinensis Oudemans, 1925: 31 – Type: Ambon, Moluccas, from Microchiroptera (syn. by Rudnick, 1960: 211)

Type locality: Serbia

Distribution: Afghanistan, Algeria, Australia, Bosnia and Herzegovina, Bulgaria, China, Crimea, E. Timor, France (incl. Corsica), Hungary, India, Indonesia, Israel, Italy (Sardinia), Japan, Korea, Malaysia, Montenegro, Morocco, Nepal, Papua New Guinea, Philippines, Portugal, Romania, Russia, Serbia, Slovakia, Spain, South Africa, Taiwan, Timor-Leste, Turkey

Hosts: Miniopterus australis, M. fuliginosus, M. macrocneme, M. macrodens, M. magnater, M. majori, M. medius, M. oceanensis, M. pusillus, M. schreibersi, M. solomonensis, M. tibialis, M. tristis (Chiroptera: Miniopteridae); Myotis brandti, M. capaccinii, M. myotis, Pipistrellus javanicus (Chiroptera: Vespertilionidae), Rhinolophus euryale, Rh. megaphyllus (Chiroptera: Rhinolophidae)

Spinturnix punctata (Sundevall)

Pteroptus punctatus Sundevall, 1833: 37; Stiles & Nolan, 1931: 632; Oudemans, 1936: 359

Spinturnix punctatus: van Eyndhoven, 1943: xliv; Rudnick, 1960: 232 (sp. inquir.)

Pteroptus barbastelli Kolenati, 1856: 187; 1856: 27; Berlese, 1892: 28; Stiles & Nolan, 1931: 630

Diplostaspis barbastelli: Kolenati, 1857: 60; 1859: 162; 1860: 138; C. Koch, 1865: 256

Spinturnix acuminatus bohemicus: Dusbábek, 1962: 366; 1964b: 390; 1970; Beron, 1965: 148 (syn. by Dusbábek, 1964: 390)

Spinturnix acuminatus barbastelli: Dusbábek, 1964b: 390; 1970; 1972; Pinchuk, 1971: 98

Spinturnix punctata: Uchikawa et al., 1994; Deunff, Keller & Aellen, 1996: 199; Krištofík & Danko, 2012: 175; Beron (in print)

Spinturnix barbastelli (Kolenati): Haitlinger, 1978: 477; Stanjukovich, 1990: 195; Medvedev et al., 1991: 28

Type locality: Sweden

Distribution: Bulgaria, France (Corsica), Germany, Kirghizstan, Lituania, Poland, Russia, Slovakia, Spain, Sweden, Switzerland, Turkey

Hosts: Barbastella barbastellus, B. leucomelas darjelingensis, Hypsugo savii (Chiroptera: Vespertilionidae)

Spinturnix rudnicki Advani et Vazirani

Spinturnix rudnicki Advani & Vazirani, 1981: 77

Type locality: Radjastan, Broach Dist.

Distribution: India

Host: Scotophilus h. heathi (Chiroptera: Vespertilionidae)

Spinturnix scotophili Zumpt et Till

Spinturnix scotophili Zumpt & Till, 1954: 214; Dusbábek & Bergmans, 1980: 66

Type locality: Kataluro, 20 miles west of Torit

Distribution: Cameroon, Nigeria, Sudan

Hosts: Scotophilus dinganii, S. leucogaster, S. nigrita (= murinoflavus) (Chiroptera: Vespertilionidae)

Spinturnix scotophili repens Dusbábek et Bergmans

Spinturnix scotophili repens Dusbábek & Bergmans, 1980: 66

Type locality: Agege, 20 km north of Lagos

Distribution: Nigeria

Host: Scotophilus nigrita (Chiroptera: Vespertilionidae)

Spinturnix scuticornis Dusbábek

Spinturnix scuticornis Dusbábek, 1970a: 62; Uchikawa et al., 1994: 294

Type locality: Jalalabad (Afghanistan)

Distribution: Afghanistan, Taiwan

Hosts: Miniopterus schreibersi (Chiroptera: Miniopteridae); Myotis longipes, M. adersus (Chiroptera: Vespertilionidae)

Spinturnix semilunaris DeMeillon et Lavoipierre

Spinturnix semilunaris DeMeillon & Lavoipierre, 1944: 65; Lavoipierre, 1946: 79; Radford, 1950: 376; Zumpt, 1950: 92; 1951: 80; Rudnick, 1960: 212; Anciaux de Faveaux, 1965: 31; Nader & Kock, 1987; Uchikawa et al., 1994: 301

Type locality: Fountains, Pretoria, Transvaal

Distribution: Angola, Central African Republic, DR Congo, Ethiopia, Kenya, Saudi Arabia, South Africa, Sudan, Tanzania, Uganda

Hosts: Miniopterus inflatus, M. minor, M. [schreibersi] natalensis, M. schreibersi, (Chiroptera: Miniopteridae), Mops condylura niveiventris (Chiroptera: Molossidae), Rhinolophus h. hildebrandti (Chiroptera: Rhinolophidae)

Spinturnix setosus Pan et Teng

Spinturnix setosus Pan & Teng, 1973: 85

Type locality: Fukien Prov.

Distribution: China

Host: Myotis sp. (Chiroptera: Vespertilionidae)

Spinturnix sinica Gu et Wang

Spinturnix sinicus Gu & Wang, 1984: 163

Type locality: Zhijin County, Guizhou Prov.

Distribution: China

Host: Ia io longimana (Chiroptera: Vespertilionidae)

Spinturnix subacuminata Furman

Spinturnix subacuminatus Furman, 1966: 151; Herrin & Tipton, 1975: 65

Spinturnix subacuminata Furman: Uchikawa et al., 1994: 296

Type locality: Bocas del Toro Province

Distribution: Panama, Venezuela

Hosts: Rhogeessa minutilla, Rh. tumida (Chiroptera: Vespertilionidae)

Spinturnix surinamensis Dusbábek et Lukoschus

Spinturnix surinamensis Dusbábek & Lukoschus, 1971: 150; Herrin & Tipton, 1975: 64; Graciolli, Guerrero & Catzeflis, 2019: 8

Type locality: Lelydorp (Suriname)

Distribution: French Guiana, Suriname, Venezuela

Hosts: Eptesicus andinus, E. brasiliensis, E. furinalis, E. melanopterus (Chiroptera: Vespertilionidae)

Spinturnix tibetensis Teng

Spinturnix tibetensis Teng, 1981: 465

Type locality: Tibet

Distribution: Tibet (China), Negros (Philippines)

Host: Pipistrellus javanicus (Chiroptera: Vespertilionidae)

Spinturnix traubi Morales-Malacara et Lopez

Spinturnix traubi Morales-Malacara & Lopez, 1998: 543

Type locality: Tlaxcala

Distribution: Mexico

Host: Corynorhinus mexicanus (Chiroptera: Vespertilionidae)

Spinturnix tylonycterisi Deunff et Volleth

Spinturnix tylonycterisi Deunff & Volleth, 1989: 186

Type locality: nr. Kuala-Lumpur

Distribution: Malaysia

Hosts: Tylonycteris pachypus, T. robustula (Chiroptera: Vespertilionidae)

Spinturnix uchikawai Orlova, Zhigalin & Zhigalina

Spinturnix uchikawai Orlova, Zhigalin & Zhigalina, 2015: 27

Type locality: Kunashir Isl.

Distribution: Russia (Kuril Islands)

Host: Myotis macrodactylus (Chiroptera: Vespertilionidae)

Spinturnix walkerae Zumpt et Till

Spinturnix walkerae Zumpt & Till, 1954: 216; Dusbábek & Bergmans, 1980: 66

Spinturnix walkeri Zumpt et Till: Whitaker & Mumford, 1978: 634

Type locality: Muhuroni (Kenya)

Distribution: Kenya, Nigeria

Hosts: Eptesicus rendalli, Myotis bocagei cupreolus, Pipistrellus nanus nanus, Scotoecus albofuscus, Scotophilus dinganii (Chiroptera: Vespertilionidae)

Spinturnix wilsoni Prasad

Spinturnix wilsoni Prasad, 1969b: 573; Domrow, 1972: 557

Type locality: Limestone Cave, Putei, Papua

Distribution: Papua New Guinea

Hosts: Myotis adversus, Pipistrellus papuanus (Chiroptera: Vespertilionidae)

Species dubiae or inquirendae

Spinturnix aviaton Seo

Spinturnix aviaton Seo, 1966: 47

Type locality: North Korea

Distribution: North Korea

Host: (Chiroptera: Vespertilionidae)

Spinturnix carnificina (Kolenati)

carnificina Kolenati, 1839

Spinturnix carnificina: Uchikawa et al., 1994: 297

Type locality: Europe

Distribution: Japan, Romania

Hosts: Eptesicus japonensis, E. nilssoni, E. serotinus (Chiroptera: Vespertilionidae)

Spinturnix spix (Kolenati)

spix Kolenati, 1856

Spinturnix spix (Kolenati)

Spinturnix verutus Delfinado & Baker, 1963: 906 – type: Philippines (syn. by Prasad, 1969)

Type locality: Europe

Distribution: Europe, Philippines

Hosts: bats (Chiroptera: Vespertilionidae)

Spinturnix sp

– ex Myotis frater – Mongolia (Scheffler et al., 2012: 145)

Species inquirendae

Pteroptus heteropus Berlese, 1910: 263 – according to Rudnick (1960, p.233) this vagely described species (from Pteropus edulis, “Batavia”, Java) may be a Meristaspis.

Spinturnix vespertilionis lanzai Lombardini, 1944: 5 - Described from Rhinolophus sp., this species should eventually change its name (“vespertilionis” being invalid).

Nomina nuda (after Rudnick, 1960)

Acarus lutescens Müller, 1776

Pteroptus pilifer Neumann, 1883

Acarus pictus Olfers, 1816

Pteroptus discolor Kolenati, 1856

Diplostaspis discolor, Kolenati, 1857

Pteroptus lyratus Kolenati, 1856

Tristaspis conspersa Kolenati, 1859

Pteroptus conspersus Kolenati, 1857

Tristaspis conspersus Kolenati, 1856

Monostaspis hexastigma Kolenati, 1857

Heterostaspis hexastigma, Kolenati, 1859

Monostaspis nathusii Kolenati, 1857

Diplostaspis nilssonii Kolenati, 1857

Diplostaspis atratula, Kolenati, 1859

Diplostaspis bechsteinii Kolenati, 1860

Tinoglischrus audouinii Murray, 1877

Meristaspis mülleri Kolenati, 1857

Periglischrus miniopteri Joseph, 1882

In the revision of Rudnick (1960, p.235) are listed many names containing the word “vespertilionis”, suppressed and out of use in the taxonomy of Spinturnicidae (Opinion 128 of the International Commission on Zoological Nomenclature, 1936).

A new genus and species, Microspinturnix koreicus, recorded from North Korea by Seo Tai Whan (1966), also seems undescribed (described in Korean in Sen Tai Whan & Chai Tak Jen, 1966, p.47 - 48).

Distribution of Spinturnicidae

EUROPE

Armenia (Ohandjanian, 1961, 1970; Arutunjan & Ohandjanian, 1974, 1983) – Eyndhovenia euryale, E. euryale oudemansi, Spinturnix myoti

Austria (Mahnert, 1971) – Spinturnix acuminata, S. myoti

Azerbaijan (Gadjiev & Dubovchenko, 1966, 1967) – Eyndhovenia euryalis, Paraperiglischrus rhinolophinus, Spinturnix acuminata, S. kolenatii, S. multisetosus, S. myoti, S. psi

Belarus (Arzamasov & Kurskov, 1962; Arzamasov, 1968; Stanyukovich, 1997) – Spinturnix [vespertilionis], S. acuminata helvetiae

Belgium (Leruth, 1939; Bruyndonckx, Henry et al., 2009) – Spinturnix «murinus Walckenaer», Spinturnix bechsteini

Bulgaria (Dusbábek, 1964; Beron & Kolebinova, 1964; Beron, 1968, 1970, 1974, in print; Scheffler, 2011) – Eyndhovenia euryalis, Paraperiglischrus rhinolophinus, Spinturnix acuminata, S. bechsteini, S. emarginata, S. myoti, S. mystacina, S. nobleti, S. plecotina, S. psi

Bosnia and Herzegowina (Burazerović et al., 2018) – Eyndhovenia euryalis, Spinturnix myoti, S. psi

Croatia (Hirst, 1927) – Paraperiglischrus rhinolophinus, Spinturnix myoti

Czech Rep. (Mrciak & Rosicky, 1956; Dusbábek, 1962; Lučan, 2006) – Spinturnix myoti, S. daubentonii (= andegavinus), S. emarginata, S. mystacina

Denmark (Haarløv, 1943) – Spinturnix acuminata

Estonia (Stanjukovich, 1990) – Spinturnix acuminata, S. kolenatii, S. myoti, S. plecotina

Finland (Mrciak & Nyholm, 1967; Huhta, 2016) – Spinturnix carnifex (published sub kolenatii), S. myoti, S. mystacina, S. plecotina, S. psi

France (contin.) (Oudemans, 1902; Hirst, 1927; Beron, 1971; Deunff, 1977; Deunff, Volleth, Keller & Aellen, 1990; Deunff et al., 2004) – Eyndhovenia euryalis euryalis, E. euryalis oudemansi, Paraperiglischrus rhinolophinus, Spinturnix bechsteini, S. daubentoni (= andegavinus), S. emarginata, S. myoti, S. mystacina, S. nobleti, S. plecotina, S. psi

Corsica (Hirst, 1927; van Eyndhoven, 1941; Beron, 1971; Deunff, Volleth, Keller & Aellen, 1990; Deunff, Keller & Aellen, 1996; Bruyndonckx et al., 2010) – Eyndhovenia euryalis, Paraperiglischrus rhinolophinus, Spinturnix myoti, S. nobleti, S. psi, S. punctata

Georgia (Rekk, 1976) – Spinturnix carnifex

Germany (C.L. Koch, 1836; Oudemans, 1904, 1910; Rudnick, 1960; Uchikawa et al., 1994; Dietz & Walter, 1995; Haitlinger & Walter, 1997; Deunff et al., 2004; Rupp, Zahn & Ludwig, 2004; Heddergott, 2008; Frank et al., 2015) – Spinturnix acuminata (= abominabilis, = noctulae), S. acuminata helvetiae, S. bechsteini, S. daubentonii (= andegavinus), S. kolenatii, S. myoti, S. mystacina, S. plecotina, S. punctata

Great Britain (Hirst, 1927; Turk, 1945; Turk & Turk, 1952; Rudnick, 1960; Jessop, 1993; Baker & Craven, 2003; Dodds, 2009) – Eyndhovenia euryalis, Paraperiglischrus rhinolophinus, Spinturnix acuminata, S. kolenatii, S. myoti, S. mystacina, S. plecotina

Greece

Crete (Beron, 1970) – Eyndhovenia euryalis, Paraperiglischrus rhinolophinus

Holland (van Eyndhoven, 1941; Buitendijk, 1945; Estrada-Peña & Sanchez, 1989; Uchikawa et al., 1994) – Eyndhovenia euryalis oudemansi, Paraperiglischrus rhinolophinus, Spinturnix acuminata acuminata, S. dasycnemi, S. plecotina

Hungary (Jablonowski, 1918; Mahunka & Molnos, 1962; Beron, 1965) – Eyndhovenia euryalis, Paraperiglischrus rhinolophinus, Spinturnix acuminata, S. emarginata, S. kolenatii, S. myoti, S. psi

Ireland (Turk, 1946) – Paraperiglischrus rhinolophinus, Spinturnix plecotina

Italy (contin.)(Hirst, 1927; Starkoff, 1957; Uchikawa et al., 1994; Lanza, 1999; Deunff et al., 2004) – Eyndhovenia euryalis, Paraperiglischrus rhinolophinus (= Periglischrus interruptus), Spinturnix acuminata, S. bechsteini, S. myoti, S. mystacina, S. nobleti

Sicily (Willmann, 1955) – Spinturnix “vespertilionis” (= myoti)

Sardinia (Uchikawa et al., 1994; Bruyndonckx et al., 2010) – Spinturnix myoti, S. psi

Latvia (Stanjukovich, 1990; Salmane, 2001; Jaunbauere, Salmane & Spungis, 2008; Salmane & Brumelis, 2010) – Spinturnix kolenatii, S. myoti, S. plecotina

Lituania (Stanjukovich, 1990) – Spinturnix punctata (=barbastelli), S. kolenatii

Macedonia (Beron, 1974) – Eyndhovenia euryalis, Paraperiglischrus rhinolophinus, Spinturnix myoti

Moldavia (Andreyko, 1968; [Andreyko, Skvorzov & Pinchuk], 1968; Andreyko & Shumilo, 1970; Pintchuk, 1971, 1976; Stanyukovich, 1997) – Paraperiglischrus rhinolophinus, Spinturnix a. acuminata, S. a. helvetiae, S. carnifex, S. myoti, S. mystacina, S. plecotina, S. punctata

Montenegro (Burazerović et al., 2018) – Eyndhovenia euryalis, Spinturnix myoti, S. psi

Norway (Mehl, 1979) – Spinturnix acuminata, S. kolenatii, S. myoti, S. mystacina, S. plecotina

Poland (Kozłowski, 1955; Kozłowski & Kosiel, 1961; Haitlinger, 1978; Haitlinger & Ruprecht, 1977, 1982, 1985, 1992; Ferenc & Skoracki, 2000; Ferenc & Myslajek, 2003; Haitlinger & Łupicki, 2008; Piksa, Siuda & Skwarek, 2011; Orlova & Zapart, 2012; Haitlinger & Piksa, 2012; Krištofík, Piksa & Sahanowicz, 2012; Sahanowicz & Ciechanowski, 2013; Szubert – Kruszyńska et al., 2009; Postawa & Szubert-Kruszyńska, 2010, 2014) – Eyndhovenia euryalis oudemansi, Spinturnix acuminata acuminata, S. a. helvetiae, S. barbastelli, S. bechsteini, S. dasycnemi, S. emarginata, S. kolenatii, S. myoti, S. mystacina, S. mystacina brandti, S. plecotina

Portugal (cont.) (Deunff, 1977; Dias, 1984; Lourenço & Palmeirim, 2007, 2008) – Spinturnix daubentonii (= andegavina), S. myoti, S. psi

Romania (Hirst, 1927 [sub «Hungary»], Beron & Kolebinova, 1964; Juvara, 1967; Uchikawa et al., 1994; Pocora et al., 2013) – Eyndhovenia euryalis, Paraperiglischrus rhinolophinus, Spinturnix acuminata, S. myoti, S. mystacina, S. psi ; Sp. dubia: S. carnificina

Russia (all, continent.) (Zhovtiy et al., 1962; Yamilianov & Vysokovsky, 1962; Tagiltzev, 1971; Rybin, 1983; Medvedev et al., 1991; Stanjukovich, 1990, 1996, 1997; Uchikawa et al., 1994; Orlova, 2011, 2013, 2014; Orlov, Chistiakov et al., 2014; Orlov, Kapitonov et al., 2011; Orlov & Orlova, 2010; Orlova & Orlov, 2011, 2015; Orlova, Orlov & Kshnyacev, 2012; Orlova, Orlov & Zhigalin, 2014; Orlova & Zapart, 2012; Orlova, Zhigalin & Zhigalina, 2015; Orlova, Zhigalin & Khritankov, 2015; Orlova, Zhigalin, Orlov, Kruskop & Bogdanov, 2015; Orlova, Stanyukovich & Orlov, 2015; Orlova, Kazakov et al., 2016) – Spinturnix acuminata acuminata, S. bregetovae, S. kolenatii, S. myoti, S. mystacina, S. plecotina, S. psi, S. punctata (= barbastelli)

Crimea (Vshivkov, 1963; Orlova & Orlov, 2018, 2019) – Eyndhovenia euryalis, Spinturnix emarginatus, S. myoti

Kuril Isl. (Orlova, Zhigalin & Zhigalina, 2015) – Spinturnix uchikawai

Serbia (Rudnick, 1960; Burazerović et al., 2018) – Eyndhovenia euryalis, Paraperiglischrus rhinolophinus, Spinturnix еmarginatus, S. myoti, S. psi

Slovakia (Dusbábek, 1962; Beron, 1965; Uchikawa & Dusbábek, 1978; Uchikawa et al., 1994; Kaňuch et al., 2005; Uhrin et al., 2006, 2010; Danko, Kristin & Krištofík, 2010; Krištofík & Danko, 2012) – Eyndhovenia euryale euryale, E. euryalis oudemansi, Paraperiglischrus rhinolophinus, Spinturnix acuminata helvetiae, S. bechsteini, S. carnifex, S. daubentonii (= andegavinus), S. dasycnemi, S. emarginata, S. kolenatii, S. myoti, S. mystacina, S. plecotina, S. psi, S. punctata (= S. acuminatus bohemicus, S. a. barbastelli)

Slovenia (Novak, 2005) – Spinturnix sp.

Spain (cont.) (Deunff, 1977; Quintero-Alonso, Pereira-Lorenzo & Sammartin-Duran, 1987; Estrada-Peña et al., 1989; Peribañez-Lopez, Sánchez-Acedo & Estrada-Peña, 1989; Estrada-Peña, Ibáñez & Trujillo, 1990; Estrada-Peña, 1990; Estrada-Peña, Balcells & Serra-Cobo, 1991; Imaz, Aihartza & Totorika, 1999) – Eyndhovenia euryalis euryalis, E. euryalis oudemansi, Paraperiglischrus rhinolophinus, S. daubentonii (= andegavina), S. acuminata, S. emarginata, S. kolenatii, S. myoti, S. mystacina, S. plecotina, S. psi

Balears (Estrada-Peña et al., 1989) – Spinturnix myoti, S. psi

Sweden (Ryberg, 1947; Sundevall, 1833) – Spinturnix punctata

Switzerland (Deunff, Keller & Aellen, 1986; Deunff, Volleth, Keller & Aellen, 1990) – Spinturnix acuminata helvetiae, S. nobleti

Turkey (all) (Beron, 1974; Uchikawa et al., 1994; Karatas & Çakır, 2004; Ciček, Stanyukovich, Yagci, Aktaš & Karaer, 2007; Toprak, 2009; Postawa, Furman, Oztunc & Çoraman, 2014) – Ancystropus, Eyndhovenia euryalis, Spinturnix acuminata, S. myoti, S. psi, S. punctata

Ukraine (Uchikawa et al., 1994; Nikitchenko, 1995; Krochko, 1995; Bobkova, 2002, 2004, 2005; Pocora et al., 2013) – Eyndhovenia euryalis oudemansi, Spinturnix acuminata, «S. carnifex», S. kolenatii, S. myoti, S. plecotina, S. psi

AFRICA

Algeria (Anciaux de Faveaux, 1976; Deunff, 1977; Uchikawa et al., 1994) – Eyndhovenia euryalis, Spinturnix myoti, S. psi

Angola (Uchikawa et al., 1994) – Ancystropus aethiopicus, Spinturnix semilunaris

Cameroon (Aellen, 1952; Till, 1958; Taufflieb & Mouchet, 1959; Dusbábek & Bergmans, 1980) – Paraperiglischrus rhinolophinus (= Periglischrus africanus), P. moucheti, Spinturnix aethiopica, S. kenyaensis, S. scotophili

Congo DR (Benoit, 1959, 1959b; Anciaux de Faveaux, 1965) – Ancystropus aethiopicus, A. leleupi, Spinturnix faini, S. semilunaris

Egypt (Kolenati, 1856; Rudnick, 1960) – Meristaspis lateralis (= Pteroptus punctolyra, = P. roseus)

Equatorial Guinea (Estrada-Peña, Ballesta & Ibanez, 1992) – Paraperiglischrus ruber, Spinturnix delacruzi

Ethiopia (Hirst, 1927; Uchikawa et al., 1994) – Spinturnix abyssinica, S. semilunaris

Gabon (Benoit, 1959b) – Spinturnix aelleni

Gambia – Ancystropus aethiopicus

Ivory Coast (Till, 1958) – Periglischrus nycteris

Kenya (Radford, 1947; Zumpt & Till, 1954; Whitaker & Mumford, 1978; Uchikawa et al., 1994) – Meristaspis kenyaensis, Paraperiglischrus rhinolophinus, Spinturnix lawrencei, S. semilunaris, S. walkerae

Liberia (Rudnick, 1960) – Meristaspis kenyaensis

Madagascar (Zumpt, 1950; Rudnick, 1960) – Meristaspis calcarata, M. kenyaensis, Spinturnix multisetosus

Morocco (Hirst, 1927; Estrada-Peña, Ibáñez & Trujillo, 1990; Bruyndonckx et al., 2010) – Eyndhovenia euryalis euryalis, E. euryalis oudemansi, Spinturnix acuminata, S. myoti, S. mystacina, S. plecotina, S. psi

Nigeria (Dusbábek & Bergmans, 1980) – Meristaspis kenyaensis, Ancystropus aethiopicus, Spinturnix intecta, S. scotophili, S. scotophili repens, S. walkerae

Sierra Leone (Radford, 1951; Rudnick, 1960) – Meristaspis kenyaensis (= Spinturnix pipistrellus)

South Africa (DeMeillon & Lavoipierre, 1944; Lavoipierre, 1944; Zumpt, 1950a, 1950b, 1951; Till, 1958; Uchikawa et al., 1994) – Paraperiglischrus rhinolophinus, Periglischrus zuluensis, Spinturnix lawrencei, S. semilunaris, S. psi

Sudan (Zumpt & Till, 1954; Uchikawa et al., 1994) – Spinturnix scotophili, S. semilunaris

Tanzania (Benoit, 1961; Uchikawa et al., 1994) – Periglischrus triaenopsis, Spinturnix semilunaris

Zanzibar (Hirst, 1923) – Ancystropus aethiopicus

Tunisia (Deunff, 1977; Uchikawa et al., 1994; Deunff et al., 2004) – Spinturnix bechsteini, S. myoti, S. psi

Uganda (Rudnick, 1960; Uchikawa et al., 1994) – Ancystropus aethiopicus, A. zeleborii, Spinturnix semilunaris

Zambia (Mwase & Baker, 2006) – Spinturnix walkerae

ASIA

Afghanistan (Cooreman, 1960b; Dusbábek, 1970; Uchikawa et al., 1994) – Spinturnix acuminata, S. kolenatii, S. plecotina, S. psi, S. scuticornis, S. «vespertilionis»

China (Pan & Teng, 1973; Petrova & Taskaeva, 1975; Teng, 1981; Gu & Wang, 1984; Sun, Wang & Wang, 1986; Wang, Sun & Wang, 1986; Whitaker, Baccus, Cleveland & Gu, 1990; Tian & Gu, 1992; Ye & Ma, 1996; Ren, T.-g. & Guo, X.-g, 2008) – Eyndhovenia brachypus, E. euryalis, Paraperiglischrus hipposideros, P. rhinolophinus, P. sternalis, Spinturnix brevisetosa, S. kolenatoides, S. plecotina, S. psi, S. sinica, S. tibetensis

Cyprus (Hirst, 1923; Rudnick, 1960) – Ancystropus zeleborii, Meristaspis lateralis

India (Hiregaudar & Bal, 1955, 1956; Rudnick, 1960; Bhat, 1973; Sharma & Sharma, 1978; Advani & Vazirani, 1981; Corpuz-Raros & Lit, 2015) – Ancystropus taprobanius, A. zeleborii, Meristaspis lateralis, Eyndhovenia euryale, Oncoscelus kanheri, Parameristaspis delfinadoae, Paraperiglischrus rhinolophinus, Spinturnix myoti, S. pindarensis, S. psi, S. rudnicki

Indonesia (Berlese, 1910; Hirst, 1923; Oudemans, 1914, 1915, 1925; Vitzthum, 1926; Radford, 1947; Rudnick, 1960; Delfinado & Baker, 1963; Durden, Peterson et al., 1990; Peterson et al., 1990; Uchikawa et al., 1994; Corpuz-Raros & Lit, 2015) – Ancystropus eonycteris, A. taprobanius, A. zeleborii, Emballonuria orientalis, Meristaspis calcarata, M. jordani, M. lateralis, M. macroglossi, Oncoscelus kanheri, Paraperiglischrus rhinolophinus, Spinturnix acuminata (= javensis), S. psi (= amboinensis). Sp. inquirendae: Pteroptus heteropus [Meristaspis ?] (Batavia, Java, on Pteropus edulis – Berlese, 1910: 263)

Iran (Sharifi et al., 2008; Malek-Hosseini et al., 2016) – Spinturnix sp.

Israel and Palestina (Hirst, 1923; Shulov, 1957; Rudnick, 1960; Costa, 1966, 1967) – Ancystropus zeleborii, Eyndhovenia euryalis, Meristaspis lateralis, Spinturnix acuminata, S. myoti, S. psi

Japan (Yamashita & Haga, 1954; Wada & Sasa, 1966; Sasa & Wada, 1968; Kawashima, 1973; Takada, 1975; Wada, 1977; Uchikawa & Dusbábek, 1978; Uchikawa, 1974, 1975, 1979, 1990; Uchikawa & Wada, 1979; Uchikawa et al., 1994) – Eyndhovenia euryalis cornuti, E. euryalis oudemansi, Meristaspis calcarata, Paraperiglischrus hipposideros, P. rhinolophinus (= strandtmanni), P. sternalis, Spinturnix acuminata acuminata, S. kolenatii, S. maedai, S. myoti, S. mystacina, S. plecotina, S. psi ; Sp. dubia: S. carnificina

Kazakhstan (Senotrusova, 1987; Stanyukovich, 1997) – Spinturnix acuminata, S. mystacina, S. myoti

Kirghizstan (Rybin, 1966, 1977, 1983; Kuchuk, 1984; Rybin, Horáček & Červený, 1989) – Eyndhovenia euryalis, Paraperiglischrus rhinolophinus, Periglischrus sp., Spinturnix acuminata, S. barbastelli, S. kolenatii, S. myoti, S. mystacina, S. punctata (= barbastelli)

Korea (Rudnick, 1960; Sou Tai Whan, 1966; Sou Tai Whan, Chai Tak Jen, 1966; Kim & Lee, 1990; Kim, Lee & Kang, 1992; Kim, Hai & Bying, 1992) – Eyndhovenia euryalis oudemansi [=ahi], Paraperiglischrus rhinolophinus, Microspinturnix koreicus, Spinturnix aviaton, S. plecotina, S. psi, [S. vespertilionis]

Laos (Prasad, 1969c; Corpuz-Raros & Lit, 2015) – Ancystropus taprobanius, A. zeleborii, Meristaspis lateralis, Paraperiglischrus rhinolophinus

Malaysia (Delfinado & Baker, 1963; Beck, 1971; Domrow, 1972; Domrow & Nadchatram, 1978; Deunff & Volleth, 1987, 1989; Uchikawa et al., 1994) – Ancystropus eonycteris, A. taprobanius, A. zeleborii, …Meristaspis lateralis, M. macroglossi, M. mindanaoensis, Oncoscelus kanheri, Paraperiglischrus hipposideros, P. rhinolophinus, Spinturnix domrowi, S. psi, S. tylonycterisi; published also « Spinturnix acuminatus, S. bakeri and S. americanus » by Ahamad et al., 2013 (unreliable !)

Mongolia (Zhovtiy et al., 1962; Dusbábek, 1966; Scheffler et al., 2010, 2012, 2016) – Spinturnix [vespertilionis], S. andegavinus, S. “frater”, S. kolenatii, S. maedai, S. myoti, S. mystacina, S. plecotina

Nepal (Mitchell & Punzo, 1976; Uchikawa et al., 1994; Dahal & Thapa, 2011; Corpus- Raros & Lit, 2015) – Ancystropus taprobanius, Meristaspis lateralis, Spinturnix plecotina, S. psi

Pakistan (Allred, 1969) – Spinturnix nudata, S. plecotina

Philippines (Delfinado, 1960; Delfinado & Baker, 1963; Prasad, 1969a, 1969b; Uchikawa et al., 1994; Fain, 2002; Corpuz-Raros, 1993; Corpuz-Raros & Lit, 2015) – Ancystropus eonycteris (= nakatae), A. taprobanius (= rudnicki, = indicus), A. zeleborii (= palawanensis), Emballonuria orientalis, Meristaspis calcarata, M. jordani (syn. ssp. philippinensis), M. lateralis, M. mindanaoensis, Oncoscelus kahneri, Spinturnix psi, S. spix (= verutus), S. tibetensis

Saudi Arabia (Nader & Kock, 1987) – Spinturnix semilunaris

Solomon Islands (Prasad, 1869; Corpuz-Raros & Lit, 2015) – Ancystropus zeleborii, Meristaspis jordani, M. mindanaoensis (= M. hardyi Prasad)

Sri Lanka (Turk, 1950; Chow, Thevasagayam & Tharumrajah, 1954; Rudnick, 1960; Uchikawa et al., 1994; Seneviratna, Chandrica Fernando & Udagama-Randeniya, 2009) – Ancystropus taprobanius, Emballonuria orientalis, Meristaspis lateralis (= M. l. ceylonicus), Oncoscelus kanheri, Paraperiglischrus hipposideros, P. rhinolophinus, Spinturnix acuminata, S. psi

Tadjikistan (Stanyukovich, 1997; Stanjukovich & Malinovskiy, 1992) – Spinturnix emarginata, S. mystacina

Taiwan (Rudnick, 1960; Baker & Delfinado, 1964; Uchikawa et al., 1994) – Spinturnix psi, S. scuticornis

Timor-Leste (Corpuz-Raros & Lit, 2015) – Meristaspis lateralis, Spinturnix psi

Thailand (Rudnick, 1960; Hill & McNeely, 1975; Uchikawa & Kobayashi, 1978; Uchikawa, 1979a, 1990; Uchikawa & Suzuki, 1980; Corpuz-Raros & Lit, 2015) – Ancystropus eonycteris, A. taprobanius, A. zeleborii, Eyndhovenia euryalis, Meristaspis lateralis, M. mindanaoensis, Oncoscelus kanheri, Paraperiglischrus hipposideros (= analis), P. rhinolophinus

Vietnam (Baker & Delfinado, 1964) – Ancystropus zeleborii

Yemen (Radford, 1954) – Meristaspis lateralis

NORTH AMERICA

Canada (Dymond, 1938; Spencer, 1940, 1941; Judd, 1953; Rudnick, 1960; Poissant & Broders, 2008; Czenze & Broders, 2011) – Spinturnix americana (= S. carloshoffmanni), S. bakeri

Mexico (Wharton, 1938; Hoffmann, 1944; Kohls & Jellison, 1948; Rudnick, 1960; Casebeer, 1966; Kingston, Vill & Lopez-Forment, 1971; Hoffmann, de Barrera, Méndez, 1972; Bassols de Barrera, 1979, 1981; Morales-Malacara, 1980, 1982, 1996, 2001; Palacios-Vargas & Morales-Malacara, 1983; Palacios-Vargas, 1985; Palacios-Vargas et al., 1985; Morales-Malacara & López – W., 1990, 1998; Uchikawa et al., 1994; Morales-Malacara & López – Ortega, 1994, 1998, 2001, 2002; Sheeler – Gordon, 1996; Sheeler – Gordon & Owen, 1999; Morales-Malacara & Juste, 2002; Villegas – Guzman et al., 2005; Colín-Martínez & García-Estrada, 2016; Ramírez Martínez et al., 2016; Morales-Malacara et al., 2018) – Cameronieta elongata, C. strandtmanni, C. thomasi, Mesoperiglischrus natali, Periglischrus acutisternus, P. caligus, P. delfinadoae, P. eurysternus, P. gameroi, P. herrerai, P. iheringi, P. leptosternus, P. ojastii, P. paracustisternus, P. steresotrichus, P. tonatii, P. vargasi, Spinturnix americana (= carloshoffmanni), S. mexicana, S. orri, S. traubi

USA (Banks, 1902, 1910; Rysgaard, 1942; Keegan, 1943; Kohls & Jellison, 1948; Holdenried, Evans & Longanecker, 1951; Augustson & Wood, 1953; Tibbetts, 1957; Jameson, 1959; Rudnick, 1960; Allred & Beck, 1966; Tipton & Saunders, 1971; Whitaker, 1973; Whitaker & Willson, 1968, 1974; Whitaker & Easterla, 1975; Whitaker & Maser, 1985; Whitaker, Deunff & Belwood, 2000; Ritzi, Ammerman, Dixon & Richerson, 2001; Dick, Gannon, Little & Patrick, 2003; Reeves et al., 2007) – Cameronieta strandtmanni, Periglischrus vargasi, Spinturnix americana (= echinipes = iowae = carloshoffmanni = grossus), S. bakeri, S. banksi, S. globosa [=Paraspinturnix globosus], S. myoti, S. orri

SOUTH and CENTRAL AMERICA

Bolivia (Dick, Gettinger & Gardner, 2007) – Periglischrus caligus, P. iheringi, P. ojastii

Brazil (Kolenati, 1857; Oudemans, 1902, 1903; Rudnick, 1960; Confalonieri, 1976; Gettinger & Gribel, 1989; Azevedo, Linardi & Coutinho, 2002; Almeida et al., 2007; Dantas -Torres et al., 2009; Silva et al., 2009; Silva & Graciolli, 2013; Almeida et al., 2011, 2015, 2016, 2018; Lima Silva & Graciolli, 2013; Almeida, Gettinger & Gardner, 2016) – Cameronieta almaensis, Mesoperiglischrus natali, Periglischrus acutisternus, P. caligus, P. iheringi, P. herrerai, P. micronycteridis, P. ojastii, P. paravargasi, P. tonatii, P. torrealbai, P. vargasi, Radfordiella desmodi, Spinturnix americana

Colombia (Rudnick, 1960; Furman, 1966; Machado-Allison & Antequera, 1969; Tamsitt & Fox, 1970) – Cameronieta thomasi, Periglischrus acutisternus, P. herrerai, P. iheringi, P. ojastii, P. setosus, P. torrealbai, P. vargasi

Costa Rica (Deunff, Whitaker & Kurta, 2007; Rojas et al., 2008) – Cameronieta torrei dusbabeki, Periglischrus herrerai

Cuba (Rudnick, 1960; Dusbábek, 1967a, 1967b, 1968, 1969, 1970b) – Cameronieta machadoi, C. strandtmanni, C. tibbettsi, C. torrei, Mesoperiglischrus nyctiellinus, Periglischrus cubanus, P. iheringi, P. vargasi, Spinturnix bakeri

Dominica (Pence, Knox Jones & Kniopping, 1981) – Spinturnix americana

Dominican Republic – Periglischrus iheringi

French Guiana (Graciolli, Guerrero & Catzeflis, 2019) – Periglischrus micronycteridis, P. paravargasi, Spinturnix americana, S. surinamensis

Guatemala (Rudnick, 1960) – Periglischrus iheringi, P. vargasi

Honduras (Rudnick, 1960) – Periglischrus iheringi

Panama (Furman, 1966; Yunker et al., 1975; Morales-Malacara, 2001; Morales-Malacara & Juste, 2002) – Mesoperiglischrus natali, Cameronieta elongata, C. strandtmanni, Periglischrus ojastii (= aitkeni), P. caligus, P. eurysternus, P. herrerai (= desmodi), P. torrealbai (= inflatiseta), P. iheringi, P. micronycteridis, P. acutisternus (= tiptoni), P. vargasi, Spinturnix americana, S. subacuminata, Paraperiglischrus globosus

Paraguay (Rudnick, 1960; Whitaker & Brian Abrell, 1987; Presley & Willig, 2008; Presley, 2004, 2012) – Periglischrus iheringi, Spinturnix americana

Peru (Guerrero, 1996; Morales-Malacara, 2001; Mendoza-Uribe & Chavez – Chorocco, 2003; Gettinger, 2018) – Periglischrus acutisternus, P. caligus, P. gameroi, P. grandisoma, P. herrerai, P. hopkinsi, P. iheringi, P. micronycteridis, P. ojastii, P. paracutisternus, P. paravargasi, P. ramirezi, P. tonatii, P. torrealbai, Spinturnix americana, S. bakeri

Puerto Rico (Rudnick, 1960; Tamsitt & Fox, 1970; Gannon & Willig, 1994; Deunff, Whitaker & Kurta, 2007, 2011; Kurta, Whitaker, Wrenn & Soto-Centeno, 2007) – Cameronieta tomasi, C. torrei dusbabeki, Periglischrus cubanus, P. iheringi, P. vargasi, Spinturnix bakeri

Suriname (Kolenati, 1857; Dusbábek & Lukoschus, 1971) – Periglischrus caligus, P. iheringi, Spinturnix surinamensis

Trinidad and Tobago (Machado-Allison, 1964; Furman, 1966) – Cameronieta elongata, Mesoperiglischrus natali, Periglischrus iheringi, P. herrerai (= desmodi), P. micronycteridis, P. vargasi

Venezuela (Hirst, 1927; Vitzthum, 1932; Machado-Allison, 1964, 1965a, 1965b; Furman, 1966; Machado-Allison & Antequera, 1971; Herrin & Tipton, 1975; Morales-Malacara, 2001) – Cameronieta elongata, C. thomasi, Periglischrus caligus, P. dusbabeki, P. grandisoma, P. herrerai, P. hopkinsi, P. iheringi (= P. meridensis), P. natali, P. ojastii, P. paracaligus, P. paracutisternus, P. paratorrealbai, P. paravargasi, P. parvus, P. ramirezi, P. setosus, P. squamosus, P. tonatii, P. torrealbai, P. vargasi, Spinturnix americana (= araguensis), S. subacuminata, S. surinamensis

Virgin Islands (Machado-Allison, 1964) – Periglischrus iheringi

Saint Kitts I. (Reevs et al., 2016) – Periglischrus iheringi

OCEANIA

Australia (continent) (Prasad, 1969; Domrow, 1959, 1962, 1967, 1972, 1979, 1988, 1992; Uchikawa et al., 1994; Halliday, 1998) – Eyndhovenia euryalis, Meristaspis calcarata, M. macroglossi, M. mindanaoensis, Paraperiglischrus rhinolophinus, Spinturnix eptesici, S. loricata, S. novaehollandiae (= antipodianus, = queenslandicus), S. psi

Tasmania (Green, 1989) – Spinturnix novaehollandiae

Fiji (Uchikawa, 1990) – Ancystropus notopteris

New Caledonia (?Uchikawa, 1990) – Ancystropus notopteris

Niue Is. (Wilson & Wodzicki, 1977) – Meristaspis calcarata

Papua New Guinea (Hirst, 1923; Rudnick, 1960; Baker & Delfinado, 1964; Prasad, 1969; Domrow, 1972; Greig – Smith, 1975; Uchikawa et al., 1994) – Ancystropus taprobanius, A. zeleborii, Meristaspis calcarata, M. macroglossi, M. mindanaoensis, Oncoscelus kanheri, Periglischrodes gressitti, Spinturnix paracuminata, S. psi, S. wilsoni

Solomon Islands (Radford, 1951; Domrow, 1972; Uchikawa et al., 1994) – Meristaspis lavellanus, M. macroglossi (= dusbabeki), Oncoscelus kanheri, Spinturnix psi

Vanuatu (New Hebrides)(Rudnick, 1960; Domrow, 1974; Uchikawa, 1990) – Ancystropus notopteris, Meristaspis calcarata

Hosts of Spinturnicidae

(the families here do not follow the new arrangement in suborders)

Chiroptera

Fam. Pteropodidae

Acerodon jubatus – Ancystropus zeleborii

Aethalops alecto – Ancystropus zeleborii

Alionycteris paucidentata – Ancystropus zeleborii

Balionycteris maculata – Ancystropus zeleborii

Cynopterus brachyotis – Ancystropus taprobanius (= rudnicki), A. zeleborii (= palawanensis), Meristaspis lateralis, M. mindanaoensis

C. horsfieldi – Meristaspis macroglossi

C. luzoniensis – Ancystropus taprobanius, Z. zeleborii, Meristaspis jordani, M. lateralis

C. sphinx – Ancystropus zeleborii (?), Paraperiglischrus hipposideros (= analis)

Dobsonia moluccensis – Meristaspis calcarata, M. jordani

D. viridis champani – Ancystropus zeleborii

Eidolon helvum – Meristaspis kenyaensis

E. sebaeum – Meristaspis lateralis

Eonycteris robusta – Ancystropus eonycteris, Meristaspis lateralis

E. spelaea – Ancystropus eonycteris, Meristaspis lateralis, M. mindanaoensis, Oncoscelus kanheri

E. spelaea glandifera – Ancystropus eonycteris (= ?nakatae), Meristaspis lateralis

Epomophorus labiatus minor – Ancystropus aethiopicus

E. wahlbergi haldemani – Ancystropus aethiopicus

Haplonycteris fischeri – Ancystropus zeleborii

Harpyonycteris whiteheadi – Meristaspis jordani (syn. ssp. philippinensis)

Hypsignathus monstrosus – Ancystropus aethiopicus

Macroglossus minimus – Meristaspis macroglossi

M. minimus lagochilus – Meristaspis lateralis, M. mindanaoensis, M. macroglossi (= dusbabeki)

M. sobrinus – Meristaspis spp.

Megaerops ecaudatus – Meristaspis mindanaoensis

Micropteropus pusillus – Ancystropus aethiopicus

Notopteris macdonaldi – Ancystropus notopteris

N. neocaledonica – Ancystropus notopteris

Nyctimene albiventer papuanus – Meristaspis macroglossi (= dusbabeki), M. mindanaoensis

N. aello – Meristaspis mindanaoensis

N. draconylla – Meristaspis mindanaoensis

N. major – Meristaspis mindanaoensis (sub “Meristaspis hardyi Prasad”)

N. robinsoni – Meristaspis mindanaoensis

Paranyctimene raptor – Meristaspis mindanaoensis

Penthetor lucasi – Ancystropus zeleborii

Prioniturus malindangensis – Meristaspis mindanaoensis

Ptenochirus jagori – Ancystropus zeleborii

Pteropus alecto – Meristaspis calcarata

P. dasymallus – Meristaspis calcarata

P. geddiei – Meristaspis calcarata

P. hypomelanus – Meristaspis calcarata

P. lavellanus ? – Meristaspis jordani (= lavellanus)

P. m. mariannus – Meristaspis calcarata

P. polyocephalus – Meristaspis calcarata

P. pselaphon – Meristaspis calcarata

P. rufus – Meristaspis calcarata

P. scapulatus – Meristaspis calcarata

P. speciosus – Meristaspis calcarata

P. tablensis – Meristaspis calcarata

P. tonganus tonganus – Meristaspis calcarata

P. vampyrus – Meristaspis calcarata

Rousettus aegyptiacus – Ancystropus aequatorialis, A. leleupi, A. zeleborii, Meristaspis lateralis, Spinturnix delacruzi

R. amplexicaudatus – Ancystropus taprobanius, A. eonycteris, A. zeleborii (= palawanensis), Meristaspis lateralis, M. mindanaoensis, Oncoscelus kanheri

R. axillaris – Eyndhovenia euryalis

R. leschenaulti – Ancystropus eonycteris, A. taprobanius (= indicus), A. zeleborii, Meristaspis lateralis, M. mindanaoensis, Oncoscelus kanheri

R. palawanensis – Ancystropus zeleborii

R. seminudus – Ancystropus taprobanius, Meristaspis lateralis

R. spelaea – Ancystropus zeleborii

R. stresemanni – Ancystropus taprobanius, A. zeleborii, Meristaspis lateralis, Oncoscelus kanheri

Syconycteris crassa papuana – Meristaspis macroglossi, M. mindanaoensis

Fam. Emballonuridae

Saccolaimus saccolaimus – Emballonuria [Spinturnix] orientalis

Saccopteryx canescens – Spinturnix americana

Taphozous georgianus – Spinturnix eptesici

Fam. Hipposideridae

Hipposideros a. armiger – Paraperiglischrus hipposideros

H. caffer – Periglischrus africanus

H. caffer centralis – Periglischrus moucheti

H. caffer guineensis – Periglischrus moucheti

H. diadema griseus – Paraperiglischrus hipposideros

H. lavatus – Eyndhovenia euryalis, Paraperiglischrus hipposideros

H. pratti pratti – Paraperiglischrus hipposideros

H. ridleyi – Paraperiglischrus rhinolophinus

H. ruber – Paraperiglischrus ruber

H. turpis – Paraperiglischrus hipposideros

Hipposideros sp. – Paraperiglischrus hipposideros

Triaenops afer – Periglischrus triaenopsis

Fam. Megadermatidae

(?) Lavia frons rex – Ancystropus zeleborii

Fam. Miniopteridae

Miniopterus australis – Spinturnix psi

M. eschscholzi – Spinturnix psi

M. fuliginosus – Spinturnix psi

M. inflatus – Spinturnix semilunaris

M. magnater – Spinturnix psi

M. macrocneme – Spinturnix psi

M. macrodens – Spinturnix psi

M. majori – Spinturnix psi

M. medius – Spinturnix psi

M. minor – Spinturnix semilunaris

M. natalensis – Spinturnix semilunaris

M. oceanensis – Spinturnix psi

M. p. pusillus – Spinturnix psi

M. schreibersi – Eyndhovenia euryalis, Paraperiglischrus rhinolophinus, Spinturnix loricata, S. myoti, S. novaehollandiae, S. psi, S. scuticornis, S. semilunaris

M. solomonensis – Spinturnix psi

M. tibialis – Spinturnix psi

Miniopterus sp. – Spinturnix paracuminata

Fam. Molossidae

Eumops glaucinus – Spinturnix americana

Molossus ater – Periglischrus iheringi

Molossops planirostris – Periglischrus acutisternus

Mops conylura niveiventer – Spinturnix semilunaris

Tadarida brasiliensis – Spinturnix sp.

Fam. Mormoopidae

Mormoops blainvillei – Cameronieta strandtmanni

M. megalophylla – Cameronieta strandtmanni

M. megalophylla senicula – Cameronieta strandtmanni

Pteronotus davyi – Cameronieta thomasi

P. parnellii – Cameronieta elongata, C. thomasi, Periglischrus acutisternus

P. parnelli boothi – Cameronieta tibbettsi

P. psilotis – Cameronieta thomasi

P. rubiginosa – Cameronieta thomasi

P. rubiginosa fusca – Cameronieta elongata, C. thomasi

P. suapurensis– Cameronieta thomasi, Periglischrus acutisternus

P. (Chilonycteris) macleayi macleayi – Cameronieta machadoi

P. (Chilonycteris) quadridens fuliginosus – Cameronieta torrei, C. torrei dusbabeki

Fam. Natalidae

Natalus (Nyctiellus) lepidus – Mesoperiglischrus nyctiellinus

N. m. mexicanus – Spinturnix americana (= carloshoffmanni)

N. mexicanus saturatus – Mesoperiglischrus natali

Natalus stramineus – Mesoperiglischrus natali

N. tumidirostris haymani – Mesoperiglischrus natali

Fam. Noctilionidae

Noctilio albiventris – Periglischrus iheringi

N. leporinus – Periglischrus ojastii (= aitkeni)

Fam. Nycteridae

Nycteris arge – Periglischrus nycteris

N. thebaica capensis – Periglischrus zuluensis

Fam. Phyllostomatidae

Anoura caudifera – Periglischrus paravargasi, P. squamosus, P. vargasi

A. cultrata – Periglischrus squamosus, P. vargasi

A. geoffroyi – Periglischrus paracutisternus, P. paravargasi, P. squamosus, P. vargasi

A. geoffroyi lasiopyga – Periglischrus vargasi

Artibeus cinereus aztecus – Periglischrus iheringi

A. cinereus – Periglischrus acutisternus

A. fimbriatus – Periglischrus iheringi

A. fraterculus – Periglischrus iheringi

A. fuliginosus – Periglischrus acutisternus

A. hirsutus – Periglischrus iheringi

A. intermedius – Periglischrus iheringi

A. jamaicensis – Periglischrus acutisternus

A. j. jamaicensis – Periglischrus iheringi, P. vargasi

A. j. parvipes – Periglischrus iheringi

A. j. yucatanicus – Periglischrus iheringi

A. lituratus – Periglischrus iheringi

A. lituratus palmarum – Periglischrus iheringi

A. obscurus – Periglischrus iheringi

A. planirostris – Periglischrus caligus, P. herrerai, P. iheringi

A. toltecus – Periglischrus iheringi

Brachyphylla cavernarum – Periglischrus cubanus

B. nana – Periglischrus cubanus

Carollia perspicillata –Periglischrus acutisternus, P. ojastii

Chiroderma salvini – Periglischrus iheringi

Ch. villosum – Periglischrus acutisternus

Choeronycteris mexicana – Periglischrus leptosternus

Dermanura azteca – Periglischrus iheringi

D. phaeotis – Periglischrus iheringi

D. tolteca – Periglischrus iheringi

Desmodus rotundus – Periglischrus acutisternus, P. herrerai, P. iheringi

D. rotundus murinus – Periglischrus iheringi

Enchisthenes hartii – Periglischrus iheringi

Erophylla sezecorni sezecorni – Periglischrus cubanus

Glossophaga leachi – Periglischrus caligus

G. longirostris – Periglischrus caligus

G. morenoi – Periglischrus caligus

G. soricina – Spinturnix [Paraspinturnix] globosa, Periglischrus caligus, P. Iheringi, P. setosus

Leptonycteris curasoae – Periglischrus paracaligus

L. nivalis – Periglischrus paracaligus, P. vargasi

L. yerbabuenae – Periglischrus vargasi

L. sanborni – Periglischrus vargasi

Lionycteris spurelli – Periglischrus hopkinsi

Lonchorhina aurita – Periglischrus gameroi

Lonchophylla robusta – Periglischrus hopkinsi

Lophostoma silviculum – Periglischrus iheringi, P. tonatii, P. torrealbai

Macrotus californicus – Periglischrus vargasi

M. waterhousi – Cameronieta thomasi, Periglischrus delfinadoae, P. vargasi

M. w. minor – Periglischrus delfinadoae

Micronycteris hirsuta – Periglischrus parvus

M. megalotis – Periglischrus micronycteridis, P. parvus

M. megalotis microtis – Periglischrus micronycteridis

M. minuta – Periglischrus micronycteridis

M. nicefori – Periglischrus parvus

Mimon bennetti – Periglischrus acutisternus

M. crenulatum – Periglischrus dusbabeki

Monophyllus redmani – Periglischrus vargasi

Phylloderma stenops – Periglischrus grandisoma, P. paratorrealbai

Phyllonycteris poeyi – Periglischrus cubanus

Phyllostomus concolor – Periglischrus acutisternus

Ph. discolor – Periglischrus acutisternus, P. iheringi, P. paravargasi, P. torrealbai

Ph. elongatus – Periglischrus acutisternus, P. torrealbai

Ph. hastatus panamensis – Periglischrus acutisternus

Ph. h. hastatus – Periglischrus acutisternus, P. torrealbai

Platyrrhinus helleri – Periglischrus iheringi

P. lineatus – Periglischrus iheringi

Rhinophylla pumilio – Periglischrus ramirezi

Rhogeessa minutilla – Spinturnix subacuminata

Rh. tumida – Periglischrus paracutisternus, Spinturnix subacuminata

Stenoderma rufum – Periglischrus iheringi

Stenoderma rufum darioi – Periglischrus iheringi

Sturnira erythromos – Periglischrus ojastii

S. lilium – Periglischrus acutisternus, P. herrerai, P. iheringi, P. ojastii, P. vargasi

S. ludovici – Periglischrus acutisternus, P. iheringi, P. ojastii

S. ororaphilum – Periglischrus ojastii

S. parvidens – Periglischrus ojastii

Tonatia bidens – Periglischrus paratorrealbai, P. tonatii

T. brasiliensis – Periglischrus tonatii

T. carrikeri – Periglischrus tonatii

T. evotis – Periglischrus steresotrichus

T. saurophila – Periglischrus eurysternus

T. silvicola – Periglischrus tonatii

Trachops cirrhosus – Periglischrus acutisternus (= tiptoni), P. paracutisternus, P. vargasi

Trinycteris nicefori – Periglischrus micronycteridis

Uroderma bilobatum – Periglischrus iheringi

U. magnirostrum – Periglischrus acutisternus

Vampyressa pusilla – Periglischrus iheringi

Vampyrodes caraccioli – Periglischrus iheringi

V. major – Periglischrus iheringi

Vampyrops helleri – Periglischrus iheringi

V. lineatus – Periglischrus iheringi

V. vittatus – Periglischrus iheringi

Vampyrops sp. – Periglischrus acutisternus

Fam. Rhinolophidae

Rhinolophus axillaris – Eyndhovenia euryalis

Rh. affinis – Paraperiglischrus rhinolophinus

Rh. blasii – Eyndhovenia euryalis, Paraperiglischrus rhinolophinus

Rh. clivosus augur – Eyndhovenia euryalis, Paraperiglischrus rhinolophinus

Rh. cornutus – Eyndhovenia euryalis cornuti, Paraperiglischrus rhinolophinus

Rh. euryale – Eyndhovenia euryalis, Paraperiglischrus rhinolophinus, Spinturnix myoti, S. plecotina, S. psi

Rh. ferrumequinum – Eyndhovenia euryalis oudemansi, Paraperiglischrus rhinolophinus, Spinturnix myoti, S. plecotina, S. psi

Rh. ferrumequinum mikado – Paraperiglischrus rhinolophinus, P. sternalis

Rh. ferrumequinum nippon – Eyndhovenia euryalis, Paraperiglischrus rhinolophinus

Rh. h. hildebrandti – Spinturnix semilunaris

Rh. hipposideros – Eyndhovenia euryalis, Paraperiglischrus rhinolophinus

Rh. hipposideros minutus – Eyndhovenia euryalis, Paraperiglischrus rhinolophinus

Rh. lepidus – Spinturnix plecotina

Rh. megaphyllus – Eyndhovenia euryalis, Paraperiglischrus rhinolophinus, Spinturnix psi

Rh. mehelyi – Eyndhovenia euryalis, Paraperiglischrus rhinolophinus

Rh. pearsoni – Spinturnix sp.

Rh. refulgens – Spinturnix spp., the other mites in the paper of Ahamad et al. (2013) doubtful (American species of parasites of Pteropodidae)

Rh. rouxi – Paraperiglischrus rhinolophinus, Eyndhovenia brachypus

Rh. rouxi sinicus – Paraperiglischrus rhinolophinus, Eyndhovenia euryalis

Rh. shamelli – Paraperiglischrus rhinolophinus

Rh. stheno – Spinturnix spp.

Rh. yunaensis – Paraperiglischrus rhinolophinus

Rhinolophus sp. – Paraperiglischrus strandtmanni

Fam. Rhinopomatidae

Rhinopoma microphyllum – Ancystropus zeleborii, Meristaspis lateralis

Fam. Vespertilionidae

Antrozous dubiaquerqus – Spinturnix orri

A. pallidus – Spinturnix americana, S. orri

A. pallidus pacificus – Spinturnix orri

A. p. cantwelli – Spinturnix orri

A. p. pallidus – Spinturnix orri

A. p. minor – Spinturnix orri

Barbastella barbastellus – Paraperiglischrus rhinolophinus, Spinturnix acuminata acuminata, S. myoti, S. punctata (barbastelli)

B. leucomelas – Spinturnix a. acuminata, S. plecotina, S. punctata

Chalinolobus dwyeri – Spinturnix novaehollandiae

Ch. gouldi – Spinturnix novaehollandiae

Ch. g. venatoris – Spinturnix novaehollandiae (=queenslandicus)

Ch. morio – Spinturnix novaehollandiae

Ch. picatus – Spinturnix novaehollandiae

Eptesicus andensis – Spinturnix surinamensis

E. brasiliensis – Spinturnix surinamensis

E. furinalis – Spinturnix surinamensis

E. fuscus – Spinturnix americana, S. bakeri

E. fuscus pallidus – Spinturnix bakeri

E. fuscus bernardinus – Spinturnix bakeri

E. fuscus dutertreus – Spinturnix bakeri

E. gobiensis – Spinturnix kolenatii, S. myoti, S. mystacina

E. japonensis – Spinturnix carnificina

E. melanopterus – Spinturnix surinamensis

E. montosus – Spinturnix bakeri

E. nilssoni – Spinturnix carnificina, S. kolenatii, S. plecotina

E. parvus – Spinturnix kolenatii

E. rendalli – Spinturnix faini, S. walkerae

E. serotinus – Eyndhovenia euryalis oudemansi, Spinturnix acuminata, S. carnificina, S. kolenatii, S. plecotina

E. tenuipinnis – Spinturnix aelleni

E. turcomanus – Spinturnix kolenatii

Glischropus tylopus – Spinturnix domrowi

Hypsugo alaschanicus – Spinturnix mystacina, S. nobleti

H. savii – Spinturnix a. acuminata, S. myoti, S. nobleti

Ia io longimana – Spinturnix sinica

Murina aurata – Spinturnix maedai

Murina ussuriensis – Spinturnix maedai

M. leucogaster – Spinturnix acuminata

M. hilgendorfi – Spinturnix maedai

Myotis adversus – Spinturnix novaehollandiae, S. scuticornis, S. wilsoni

M. albescens – Periglischrus acutisternus, Spinturnix americana

M. alcathoe – Spinturnix mystacina

M. aurascens – Spinturnix mystacina

M. bechsteini – Spinturnix bechsteini

M. blythi – Eyndhovenia euryalis oudemansi, Spinturnix acuminata, S. mystacina, S. myoti

M. bocagei cupreolus – Spinturnix walkerae

M. brandti – Spinturnix kolenatii, S. myoti, S. mystacina, S. plecotina, S. psi

M. capaccinii – Spinturnix myoti, S. psi

M. chiloensis – Spinturnix americana, Paraspinturnix globosa, Periglischrus acutisternus (= tiptoni)

M. dasycneme – Spinturnix acuminata, S. myoti, S. daubentoni, S. dasycnemi

M. daubentoni – Spinturnix daubentonii (= andegavinus), S. myoti, S. plecotina

M. dominicensis – Spinturnix americana

M. emarginatus – Eyndhovenia euryalis, Spinturnix emarginata, S. myoti

M. frater – Spinturnix sp.

M. evotis – Spinturnix americana

M. goudotii – Spinturnix multisetosa

M. gracilis – Spinturnix bregetovae, S. mystacina

M. grisescens – Spinturnix banksi

M. hosonoi – Spinturnix mystacina

M. ikonnikovi – Spinturnix bregetovae, S. myoti, S. mystacina

M. keaysi – Spinturnix americana, S. bakeri

M. keenii – Spinturnix americana

M. longipes – Spinturnix scuticornis

M. lucifugus – Spinturnix americana

M. macrodactylus – Spinturnix myoti, S. uchikawai

M. myotis – Spinturnix a. acuminata, S. daubentoni, S. myoti, S. psi

M. mystacinus – Spinturnix myoti, S. mystacina, Steatonyssus mongolicus, S. murinus

M. nattereri – Spinturnix myoti, S. mystacina, S. plecotina

M. «nattereri« – Spinturnix myoti

M. nigricans – ? S. [Paraspinturnix] globosa, Periglischrus tiptoni, Spinturnix americana

M. petax – Spinturnix daubentonii (= andegavinus), S. myoti-complex

M. punicus – Spinturnix myoti

M. sibiricus – Spinturnix bregetovae, S. mystacina

M. siligorensis – Spinturnix pindarensis

M. simus – Spinturnix americana

M. sodalis – Spinturnix americana, S. [Paraspinturnix] globosa

M. thysanodes – Spinturnix americana (= carloshoffmanni)

M. tricolor – Spinturnix lawrencei

M. velifer incautus – Spinturnix americana (= carloshoffmanni)

M. velifer velifer – Spinturnix americana (= carloshoffmanni)

M. volans – Spinturnix americanus, S. bakeri, S. [Paraspinturnix] globosa

M. yumanensis – Spinturnix americana

M. yumanensis saturatus – Spinturnix americana

M. yumanensis sociabilis – Spinturnix americana

Myotis sp. – Spinturnix setosus

Nyctalus lasiopterus aviator – Spinturnix acuminata

N. leisleri – Spinturnix acuminatus, S. acuminata helvetiae

N. noctula – Spinturnix acuminata acuminata, S. plecotina

Nycticeius greyi – Spinturnix novaehollandiae

Nyctophilus arnemensis – Spinturnix eptesici

Pipistrellus angulatus – Spinturnix paracuminata

P. babu – Spinturnix sp.

P. gaisleri – Eyndhovenia euryalis, Spinturnix myoti

P. javanicus – Spinturnix psi

P. kuhli – Spinturnix acuminata

P. nanus – Spinturnix walkerae

P. nathusii – Spinturnix myoti

P. papuanus – Spinturnix wilsoni

P. pipistrellus – Eyndhovenia euryalis oudemansi, Spinturnix acuminata

P. stampfIii – Meristaspis kenyaensis

Pizonyx vivesi – Spinturnix mexicana

Plecotus auritus – Spinturnix kolenatii, S. myoti, S. mystacina, S. plecotina

Plecotus auritus sacrimontis – Spinturnix plecotina

P. austriacus – Spinturnix plecotina

P. kozlovi – Spinturnix plecotina

P. ognevi – Spinturnix plecotina

Corynorhinus mexicanus – Spinturnix americana, S. traubi

Rhogeessa tumida – Spinturnix subacuminata

Scotoecus albofuscus – Spinturnix walkerae

Scotophilus dinganii – Spinturnix intecta, S. scotophili, Spinturnix scotophili repens, S. walkerae

S. heathii – Spinturnix a. acuminata

S. kuhli – Ancystropus taprobanius, Spinturnix sp.

S. leucogaster – Spinturnix abyssinica, Spinturnix scotophili

S. murinoflavus – Spinturnix scotophili

S. nigrita – Spinturnix scotophili repens

Tylonycteris pachypus – Spinturnix tylonycteris

T. robustula – Spinturnix tylonycteris

Vespadelus [Eptesicus] douglasorum – Spinturnix eptesici

V.[E.] pumilus – Spinturnix eptesici

Vespertilio murinus – Spinturnix kolenatii

« bat« 1 (Chiroptera indet.) – Spinturnix novaehollandiae

« bat« 2 (Chiroptera indet.) – Spinturnix abyssinica

« bat« 3 (Chiroptera indet.) – Spinturnix spix

« bat« 4 (Chiroptera indet.) – Spinturnix nudata

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New combinations

Sternostoma numerovi (Butenko) new comb. for Sternostomoides numerovi Butenko, 1999: 49

Tinaminyssus alisteri (Wilson) new comb. for Mesonyssus alisteri Wilson, 1964b: 370

Tinaminyssus buteonis (Fain) new comb. for Neonyssus buteonis Fain, 1956a: 135 (Mesonyssus buteonis Fain, 1962d: 306)

Tinaminyssus alexfaini (Amaral) new comb. for Mesonyssus alexfaini Amaral, 1967: 135

Tinaminyssus camargoi (Amaral) new comb. for Mesonyssus camargoi Amaral, 1962: 145

Tinaminyssus castroae (Amaral) new comb. for Mesonyssus castroae Amaral, 1963: 9

Tinaminyssus cunhai (Amaral) new comb. for Mesonyssus cunhai Amaral, 1963: 103

Tinaminyssus egrettae (Butenko) new comb. for Mesonyssus egrettae Butenko, 1984: 96

Tinaminyssus elani (Fain) new comb. for Falconyssus elani Fain, 1966c: 86

Tinaminyssus gerschi (Feider et Mironescu) new comb. for Mesonyssus gerschi Feider et Mironescu, 1974: 752

Tinaminyssus gourae (Wilson) new comb. for Mesonyssus gourae Wilson, 1964b: 361

Tinaminyssus marciae (Amaral) new comb. for Mesonyssus marciae Amaral, 1974: 752

Tinaminyssus metropeliae (Amaral) new comb. for Mesonyssus metropeliae Fain, 1972: 351

Tinaminyssus oenae (Fain) new comb. for Mesonyssus oenae Fain, 1963c: 61

Tinaminyssus peleiae (Wilson) new comb. for Mesonyssus peleiae Wilson, 1966a: 609

Tinaminyssus phassae (Wilson) new comb. for Mesonyssus phassae Wilson, 1966a: 602

Tinaminyssus prionituri (Wilson) new comb. for Mesonyssus prionituri Wilson, 1966: 759

Tinaminyssus pteroclesi (Fain) new comb. for Mesonyssus pteroclesi Fain, 1963c: 63

Tinaminyssus schoutedeni (Fain) new comb. for Neonyssus schoutedeni Fain, 1956a: 134

Tinaminyssus senotrusovae (Butenko) new comb. for Mesonyssus senotrusovae Butenko, 1984: 103

Tinaminyssus serraoi (de Castro) new comb. for Neonyssus (Neonyssus) serraoi de Castro, 1948: 27

Tinaminyssus squamosus (Vitzthum) new comb. for Rhinonyssus squamosus Vitzthum, 1935: 567

Tinaminyssus streptopeliae (Fain) new comb. for Mesonyssus melloi streptopeliae Fain, 1962d: 310

Tinaminyssus streptopelioides (Butenko) new comb. for Mesonyssus streptopelioides Butenko, 1984: 107

Tinaminyssus tetae (Wilson) new comb. for Mesonyssus tetae Wilson, 1964b: 605

Tinaminyssus tinamicola (Fain) new comb. for Mesonyssus tinamicola Fain, 1963c: 66

Tinaminyssus trappi (Pereira et Castro) new comb. for Neonyssus (Ptilonyssoides) trappi Pereira & Castro, 1949: 229

Tinaminyssus treronis (Fain) new comb. for Neonyssus treronis Fain, 1956b: 394; 1957: 56

Tinaminyssus turturi (Fain) new comb. for Mesonyssus turturi Fain, 1962d: 314

Acknowledgements

Several specialists have provided advice, literature and all kinds of support over an extended period of time. They are kindly asked to accept our sincere gratitude. Among them are L. van der Hammen, M. Koyumjieva, V. Černy, G.W. Krantz, B. Haliday, J. Deunff, A. Estrada-Peña, II. Juvara – Balş, N. Bregetova, A. Fain, O.M. Butenko, G.L. van Eyndhoven, P. Lehtinen, M. Naudo, M. K. Stanjukovich, R.W. Strandtmann, J.O. Whitaker, N. Wilson, M. Mrciak, F. Radovsky, F. N. Vshivkov, T.A. Dubovchenko, F. Dusbábek, R.F. Lawrence, A. Baker, L.M. Pinchuk, R. Haitlinger, R. Domrow, J. de la Cruz, M. Heddergott, N. Wilson, A.B. Hoffmann, C.D. Radford, K. Uchikawa, J.B. Morales-Malacara, M. Anciaux de Faveaux, D. Pence, P. Benoit, F. Zumpt, I. Dimov.